538 lines
80 KiB
XML
538 lines
80 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1184.113498" ID-Pensoft-Pub="1313-2970-1184-65" ID-Pensoft-UUID="CB3CAF6A267D510A893D6754710069F8" ID-ZooBank="0F9F839C577C4AF59206D3F0BD067BD2" ModsDocID="1313-2970-1184-65" checkinTime="1699992652139" checkinUser="pensoft" docAuthor="Antic, Dragan & Akkari, Nesrine" docDate="2023" docId="92000BE4075158B5BAD888FBEC86BD8C" docLanguage="en" docName="ZooKeys 1184: 65-80" docOrigin="ZooKeys 1184" docPubDate="2023-11-14" docSource="http://dx.doi.org/10.3897/zookeys.1184.113498" docTitle="Inversotyphlus ammirandus Antić & Akkari 2023, sp. nov." docType="treatment" docUuid="BCE36811-7013-4F70-B8DB-4CED5E055FD2" docUuidSource="ZooBank" docVersion="1" id="CB3CAF6A267D510A893D6754710069F8" lastPageNumber="65" masterDocId="CB3CAF6A267D510A893D6754710069F8" masterDocTitle="The genus Inversotyphlus Strasser, 1962, stat. nov. and Inversotyphlus ammirandus sp. nov., a new bizarre, highly modified troglobiotic millipede (Diplopoda, Julida, Julidae) from Albania, Balkan Peninsula" masterLastPageNumber="80" masterPageNumber="65" pageNumber="65" updateTime="1699992652139" updateUser="pensoft">
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<mods:title id="E8145D5A0A4E7D05633F41AFE8E518D1">The genus Inversotyphlus Strasser, 1962, stat. nov. and Inversotyphlus ammirandus sp. nov., a new bizarre, highly modified troglobiotic millipede (Diplopoda, Julida, Julidae) from Albania, Balkan Peninsula</mods:title>
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<mods:name id="1F5E0DD2573901FBF753934D582FC54E" type="personal">
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<mods:roleTerm id="C0457EDFD4A9AA30565A79AF9485FBCA">Author</mods:roleTerm>
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<mods:namePart id="9C91BD5E3AF7555E7ADCE83E9A95FC3B">Antic, Dragan</mods:namePart>
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<mods:nameIdentifier id="E3A3CF54374BAFD595760C08BDFE9ACA" type="ORCID">https://orcid.org/0000-0002-1231-4213</mods:nameIdentifier>
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<mods:affiliation id="5FDCEAF36BF5C89EEEB8AE13BF5C4CB0">University of Belgrade-Faculty of Biology, Institute of Zoology, Center for Biospeleology, Studentski trg 16, 11000 Belgrade, Serbia & Serbian Biospeleological Society, Trg Dositeja Obradovica 2, 21000 Novi Sad, Serbia</mods:affiliation>
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<mods:nameIdentifier id="ECA8178D15ECC0CCDD093C34F590788C" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="AB19D6BDC719D1D8B012E772FABAC099">Akkari, Nesrine</mods:namePart>
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<mods:nameIdentifier id="87DD4A9B961E5F01E24177705CABDA7D" type="ORCID">https://orcid.org/0000-0001-5019-4833</mods:nameIdentifier>
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<mods:affiliation id="E3DEF44EE68436F9C8AE9BC0EFA1A2F2">3 rd Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:typeOfResource id="49F3D7E40FA9F53338BB342B3AB4E12A">text</mods:typeOfResource>
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<mods:title id="CCE24B6681696555674F65493C412C86">ZooKeys</mods:title>
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<mods:date id="1C8D0E5853B860228741D7398BF523CD">2023</mods:date>
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<mods:number id="A9D25D2392B2C33C09A9E935DD7AEDE9">2023-11-14</mods:number>
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<mods:number id="51FFAD83D3F0EDA86CF73FA8EC860DD9">1184</mods:number>
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<mods:url id="EFEBF58345A89181978405E77A445186">http://dx.doi.org/10.3897/zookeys.1184.113498</mods:url>
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<mods:classification id="804373A6D8E6D29264F5CF3BFEC1C1E2">journal article</mods:classification>
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<mods:identifier id="02078B8D6D8188F6BDC80E743E775F40" type="DOI">http://dx.doi.org/10.3897/zookeys.1184.113498</mods:identifier>
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<treatment id="92000BE4075158B5BAD888FBEC86BD8C" LSID="urn:lsid:zoobank.org:act:BCE36811-7013-4F70-B8DB-4CED5E055FD2" httpUri="http://treatment.plazi.org/id/92000BE4075158B5BAD888FBEC86BD8C" lastPageNumber="65" pageId="0" pageNumber="65">
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<subSubSection id="92212C50481953F0165C2B53BC53CD77" pageId="0" pageNumber="65" type="nomenclature">
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<paragraph id="191F36A47989373DF3879FBAFF6EF7E1" pageId="0" pageNumber="65">
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<taxonomicName id="AFC58D5159841734AFDBA9C93958D9E4" LSID="https://zoobank.org/BCE36811-7013-4F70-B8DB-4CED5E055FD2" authority="Antić & Akkari, 2023" authorityName="Antić & Akkari" authorityYear="2023" class="Diplopoda" family="Julidae" genus="Inversotyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Inversotyphlus ammirandus" order="Julida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="ammirandus" status="sp. nov.">Inversotyphlus ammirandus</taxonomicName>
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<taxonomicNameLabel id="A71EA6DBFB3DA34E47865853A9E3E635" pageId="0" pageNumber="65">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="7D041D66CDD6747126C1128B8B5769B5" pageId="0" pageNumber="65" type="description">
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<paragraph id="53AC50EE3A2A401BEA8F359A4A394167" pageId="0" pageNumber="65">
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<figureCitation id="D4C4061D581C15B5F872A3D6BBEAF526" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">Figs 1</figureCitation>
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<figureCitation id="B9C80A7BCA060C9819A9630E0C3A37DA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), head A anterodorsal view B dorsal view C anterior part of head, anterodorsal view D lateral view E anterior view F head without gnathochilarium, dorsal view G head without gnathochilarium and right mandible, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.113498.figure2" httpUri="https://binary.pensoft.net/fig/936069" pageId="0" pageNumber="65">, 2</figureCitation>
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<figureCitation id="7EA81BEE67AEC8E613C6C027DA8C458D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), mouthparts A gnathochilarium, ventral view B right anterior part of head, showing antennomere 1 and 2, labral lobe, mandible and stipital bundle C right mandible, ventral view D right mandible, dorsal view. Abbreviations (mandible): et external tooth, it internal tooth, mp molar plate, pl pectinate lamellae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure3" httpUri="https://binary.pensoft.net/fig/936070" pageId="0" pageNumber="65">, 3</figureCitation>
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<figureCitation id="4BFE7AF614869C2DBA67DC8946F38380" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">, 4</figureCitation>
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<figureCitation id="51308665E90C38701A9ED70D043E9518" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C penis, posterior view D right gonopods, mesal view E left gonopods, lateral view F right promere, posterior view G right mesomere, anterior view. Abbreviations: m mesomere, ml mesal lobe, o opisthomere, p promere, s solenomere, t telopodite, v velum. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1184.113498.figure5" httpUri="https://binary.pensoft.net/fig/936072" pageId="0" pageNumber="65">, 5</figureCitation>
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<figureCitation id="F9E04490421BC5E9D739CFFFE2CF3B6C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Gonopods of the members of the genus Inversotyphlus Strasser, 1962 stat. nov., mesal views A I. ammirandus sp. nov., holotype ♂ (NHMW MY 10380), right gonopods (flipped) B I. clavatus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods C I. edentulus (Attems, 1951), comb. nov., ♂ from Njegos Cave (IZB), left gonopods D I. opisthonodus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods E I. gellianae (Makarov & Rađa, 2006), comb. nov., holotype ♂ (IZB), left gonopods F I. gracilis (Antic, 2018), comb. nov., paratype ♂ (IZB), left gonopods G I. lobifer (Attems, 1951), comb. nov., ♂ from bitumen mine Minjera, island of Brac (IZB), left gonopods. B, D-G modified after Antic et al. (2018). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure6" httpUri="https://binary.pensoft.net/fig/936073" pageId="0" pageNumber="65">, 6A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6EFB4774D70300FA763E0CE1F4BA508E" pageId="0" pageNumber="65" type="diagnosis">
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<paragraph id="D68DE42909F61AA9D3DD2EC5F4D4A47C" pageId="0" pageNumber="65">Diagnosis.</paragraph>
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<paragraph id="39DCB514D48B4735677621746C8886F8" pageId="0" pageNumber="65">
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The new species is distinguished from all congeners and other members of
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<taxonomicName id="B56C05BA6854FFAD477D44C8EE9F36C1" authorityName="Verhoeff" authorityYear="1930" lsidName="" pageId="0" pageNumber="65" rank="tribe" tribe="Typhloiulini">Typhloiulini</taxonomicName>
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by its highly modified head, which is characterised by having two labral lobes with a wide incision between to accommodate the lingual palps, a unique gnathochilarium that is distally very wide, bearing a distolateral bundle of long setae on the stipites and distal (anterior) rows of long setae on both the stipites and lamellae linguales (such structures are absent in all congeners). In addition, the body rings are strongly vaulted and bear very long metazonal setae; the tarsal claws are very long.
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</paragraph>
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<paragraph id="79111B6AF248D9232A5AC13D2C95302B" pageId="0" pageNumber="65">
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Concerning the gonopods, the new species is characterised by a slightly higher mesomere compared to the promere (vs mesomere lower than promere in
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<taxonomicName id="26A838EAEBA5360C0C4592C9C48669C7" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. opisthonodus" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="opisthonodus">
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<emphasis id="9263812A930B1F17A80175774A783DC2" italics="true" pageId="0" pageNumber="65">I. opisthonodus</emphasis>
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</taxonomicName>
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comb. nov. and
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<taxonomicName id="5D66EB01403FC4F48E4585FDDB0482ED" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. gracilis" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="gracilis">
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<emphasis id="9A99C79C31926232E20956FE12B922CE" italics="true" pageId="0" pageNumber="65">I. gracilis</emphasis>
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</taxonomicName>
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comb. nov., or mesomere much higher than promere in
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<taxonomicName id="54F80EDE13D3B43040239FA7C0A4D2E1" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. lobifer" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="lobifer">
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<emphasis id="20470822C7BD8282D2EFD2D3ED09344B" italics="true" pageId="0" pageNumber="65">I. lobifer</emphasis>
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</taxonomicName>
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comb. nov. and
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<taxonomicName id="EB270E650AD29885D2896DFA2B5BE545" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. clavatus" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="clavatus">
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<emphasis id="91D0470216C041BD3B7F730F016668B5" italics="true" pageId="0" pageNumber="65">I. clavatus</emphasis>
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</taxonomicName>
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comb. nov., or mesomere and promere of equal height in
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<taxonomicName id="3821A88846B009CBD93D00C860EDFE6B" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. edentulus" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="edentulus">
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<emphasis id="51EF9BD2D16A262E7CBDB086FE275755" italics="true" pageId="0" pageNumber="65">I. edentulus</emphasis>
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</taxonomicName>
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comb. nov.). Except in the new species, a mesomere slightly higher than the promere is found only in
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<taxonomicName id="C7709C5835252E086129FB23784BFE13" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. gellianae" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="gellianae">
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<emphasis id="CC61ED4C0D26648E05BD612D5723A5B6" italics="true" pageId="0" pageNumber="65">I. gellianae</emphasis>
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</taxonomicName>
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comb. nov., but
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<taxonomicName id="31E7A1042872D988F777C20FF3F8271C" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. ammirandus" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="ammirandus">
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<emphasis id="CB9B5A125C00BEE01270A386BDD2BA20" italics="true" pageId="0" pageNumber="65">I. ammirandus</emphasis>
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</taxonomicName>
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sp. nov. differs from this and all other members of
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<taxonomicName id="98992AED389EC3F8A19B482B356B4C48" authorityName="Antić & Akkari" authorityYear="2023" class="Diplopoda" family="Julidae" genus="Inversotyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Inversotyphlus" order="Julida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis id="8F7ACE780FA0D8483B79C70A526F3316" italics="true" pageId="0" pageNumber="65">Inversotyphlus</emphasis>
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</taxonomicName>
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stat. nov. in the presence of a spoon-shaped mesomere. Further differences in the gonopods can be seen in Fig.
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<figureCitation id="16B106BDE4984A3B8A739B0C9CE27078" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Gonopods of the members of the genus Inversotyphlus Strasser, 1962 stat. nov., mesal views A I. ammirandus sp. nov., holotype ♂ (NHMW MY 10380), right gonopods (flipped) B I. clavatus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods C I. edentulus (Attems, 1951), comb. nov., ♂ from Njegos Cave (IZB), left gonopods D I. opisthonodus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods E I. gellianae (Makarov & Rađa, 2006), comb. nov., holotype ♂ (IZB), left gonopods F I. gracilis (Antic, 2018), comb. nov., paratype ♂ (IZB), left gonopods G I. lobifer (Attems, 1951), comb. nov., ♂ from bitumen mine Minjera, island of Brac (IZB), left gonopods. B, D-G modified after Antic et al. (2018). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure6" httpUri="https://binary.pensoft.net/fig/936073" pageId="0" pageNumber="65">6</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="24F6475225547DE856FEB1910E17A3CA" pageId="0" pageNumber="65" type="name">
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<paragraph id="80C57DAAEDBF5280466CF655496EE1EE" pageId="0" pageNumber="65">Name.</paragraph>
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<paragraph id="B4C003F5EED0912401552DA0ACC2A237" pageId="0" pageNumber="65">
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From the Latin
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<emphasis id="8EC4EB41A422BE437301D5A8242F0510" italics="true" pageId="0" pageNumber="65">ammirandus</emphasis>
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(= wonderful, admirable, astonishing, remarkable, extraordinary), reflecting the impressive appearance of this bizarre creature. Moreover, this species name is dedicated to all speleologists and speleobiologists around the world who risk their lives exploring deep caves, which makes them admirable too. Adjective in masculine gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="7ED441BA5B0D64B62A7841C1E5F1159B" pageId="0" pageNumber="65" type="materials_examined">
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<paragraph id="90CD8B16333476DF0830315B8D8574BA" pageId="0" pageNumber="65">Material examined.</paragraph>
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<paragraph id="F2C8DB28C8AF1DEF1791096DC867ED49" pageId="0" pageNumber="65">
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<materialsCitation id="814ADB7E20BD1F8A697AB91517E80DB1" collectingDate="2013-09-08" collectorName="T. Cukovic" country="Albania" county="Malesi" elevation="460" location="Boge village" municipality="Madhe Municipality" specimenCount="♂" stateProvince="Shkoder County" typeStatus="Holotype">
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<emphasis id="866339F93E6878AF837D4233F119EE7A" bold="true" italics="true" pageId="0" pageNumber="65">
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<typeStatus id="48C1D9A7E30C2C870A91A18CB34E8C39">Holotype</typeStatus>
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</emphasis>
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<specimenCount id="06BA63FF68EB62444D8EA7B5519B50CC">♂</specimenCount>
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(NHMW MY10380),
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<collectingCountry id="AB53FD2745A8E2A8C262E81B9D875EB9" name="Albania">Albania</collectingCountry>
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,
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<collectingRegion id="634740A33A0F7D36CEFEC6AB75EFB697" country="Albania" name="Shkoder">
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<normalizedToken id="93342991FF5093B9AAEF40143B5311E4" originalValue="Shkodër">Shkoder</normalizedToken>
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County
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</collectingRegion>
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,
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<normalizedToken id="6DFB156E0C3FF1221DACEF27D1AF274C" originalValue="Malësi">
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<collectingCounty id="32DF89848B277E0F8C36E958A3CBCB23">Malesi</collectingCounty>
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</normalizedToken>
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e
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<collectingMunicipality id="B53CEDF41A39B0A0EE455DCDCF09147C">Madhe Municipality</collectingMunicipality>
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,
|
||
<location id="F37F8686CDB96B9C2B84BF988E95FE3E" LSID="urn:lsid:plazi:treatment:92000BE4075158B5BAD888FBEC86BD8C:F37F8686CDB96B9C2B84BF988E95FE3E" country="Albania" county="Malesi" municipality="Madhe Municipality" name="Boge village" stateProvince="Shkoder County">
|
||
<normalizedToken id="E563FFE21FCDE70908DBADA2D8F44F3E" originalValue="Bogë">Boge</normalizedToken>
|
||
village
|
||
</location>
|
||
,
|
||
<location id="B46A71E9B2B335B83AA7712E35DA225D" LSID="urn:lsid:plazi:treatment:92000BE4075158B5BAD888FBEC86BD8C:B46A71E9B2B335B83AA7712E35DA225D" country="Albania" county="Malesi" municipality="Madhe Municipality" name="Prokletije Mountain range" stateProvince="Shkoder County">Prokletije Mountain range</location>
|
||
,
|
||
<location id="CB831467B172AF1F290110606DB39213" LSID="urn:lsid:plazi:treatment:92000BE4075158B5BAD888FBEC86BD8C:CB831467B172AF1F290110606DB39213" country="Albania" county="Malesi" municipality="Madhe Municipality" name="Radohimes (Radohines) mountain" stateProvince="Shkoder County">
|
||
<normalizedToken id="9D7CC1B29949181CFEC9CD536952C5B8" originalValue="Radohimës">Radohimes</normalizedToken>
|
||
(
|
||
<normalizedToken id="06D7448A98F5A0655B5EE3674167F95D" originalValue="Radohinës">Radohines</normalizedToken>
|
||
) mountain
|
||
</location>
|
||
group,
|
||
<location id="187B31CF78F5391C3393590295BE86BB" LSID="urn:lsid:plazi:treatment:92000BE4075158B5BAD888FBEC86BD8C:187B31CF78F5391C3393590295BE86BB" country="Albania" county="Malesi" municipality="Madhe Municipality" name="Cave Ru" stateProvince="Shkoder County">Cave Ru</location>
|
||
, -
|
||
<quantity id="7B2142BEFAE262C94C609F870B7AA763" metricMagnitude="2" metricUnit="m" metricValue="4.6" unit="m" value="460.0">
|
||
<elevation id="FA54A9EE28C745A5D754541948D4E993" metricMagnitude="2" metricUnit="m" metricValue="4.6" unit="m" value="460.0">460 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate id="16E45C78CE03FBD2000A74F5D25C8B76" value="2013-09-08">8 September 2013</collectingDate>
|
||
,
|
||
<collectorName id="905BE34B91617A1F7CE2FA29FB845762">
|
||
T.
|
||
<normalizedToken id="42FE78F2207A9F9E64C3F92354E6BA8E" originalValue="Čuković">Cukovic</normalizedToken>
|
||
</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AE271F5513EC23800FB2FCA1AE4A0562" pageId="0" pageNumber="65" type="description">
|
||
<paragraph id="B9B5E1DF7E0A77CAF88CB5B99D2243A1" pageId="0" pageNumber="65">Description.</paragraph>
|
||
<paragraph id="B716B3C6468ACD5B6268086C35FE6869" pageId="0" pageNumber="65">
|
||
<emphasis id="A2F40ABD7E93F70D9DCEE31A8CBCDCC4" bold="true" italics="true" pageId="0" pageNumber="65">Number of body rings and size</emphasis>
|
||
: 30 mm long, vertical diameter of largest body ring 1.5 mm, body with 36 podous rings + 1 apodous ring + telson.
|
||
</paragraph>
|
||
<paragraph id="846635486A679CAF909DD90560EFF051" pageId="0" pageNumber="65">
|
||
<emphasis id="7446FE419A78AE5C1943D10B30C7B192" bold="true" italics="true" pageId="0" pageNumber="65">Colouration</emphasis>
|
||
: after a decade in ethanol, generally greyish brown (Fig.
|
||
<figureCitation id="3CAF739D593CAE458D130573291AD0BD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1</figureCitation>
|
||
). Anterior part of body and telson rather yellowish brown (Fig.
|
||
<figureCitation id="91D51481EE551C4ACFAC4B6B09A64122" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1A, B</figureCitation>
|
||
). Antennae and forehead yellowish; legs brownish. Metazonae with dark-brown posterior rings.
|
||
</paragraph>
|
||
<paragraph id="EBAB804A45B98495AA21A0E437CA93EB" pageId="0" pageNumber="65">
|
||
<emphasis id="AF6DC2E5ABDD4D2C4823EEC85E8DC210" bold="true" italics="true" pageId="0" pageNumber="65">Head</emphasis>
|
||
: highly modified (Figs
|
||
<figureCitation id="924D8AD86CF3D452B789DB91609F6E8D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), head A anterodorsal view B dorsal view C anterior part of head, anterodorsal view D lateral view E anterior view F head without gnathochilarium, dorsal view G head without gnathochilarium and right mandible, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.113498.figure2" httpUri="https://binary.pensoft.net/fig/936069" pageId="0" pageNumber="65">2</figureCitation>
|
||
,
|
||
<figureCitation id="86BE4B8EED275D56D6369119DFB091DC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), mouthparts A gnathochilarium, ventral view B right anterior part of head, showing antennomere 1 and 2, labral lobe, mandible and stipital bundle C right mandible, ventral view D right mandible, dorsal view. Abbreviations (mandible): et external tooth, it internal tooth, mp molar plate, pl pectinate lamellae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure3" httpUri="https://binary.pensoft.net/fig/936070" pageId="0" pageNumber="65">3</figureCitation>
|
||
), without ommatidia, with a pair of frontal setae (Fig.
|
||
<figureCitation id="4268DA9DE5D3465E7BA839903EC54B56" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), head A anterodorsal view B dorsal view C anterior part of head, anterodorsal view D lateral view E anterior view F head without gnathochilarium, dorsal view G head without gnathochilarium and right mandible, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.113498.figure2" httpUri="https://binary.pensoft.net/fig/936069" pageId="0" pageNumber="65">2A, E</figureCitation>
|
||
; both fallen off). Labrum considerably shorter than gnathochilarium, with two rounded lobes with a wide inscision between lobes to accomodate lingual palps; no labral teeth; 3+3 short labral and 5+5 long supralabral setae (Figs
|
||
<figureCitation id="9673661AF51E7ABC7AD4D7FAF63D534B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), head A anterodorsal view B dorsal view C anterior part of head, anterodorsal view D lateral view E anterior view F head without gnathochilarium, dorsal view G head without gnathochilarium and right mandible, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.113498.figure2" httpUri="https://binary.pensoft.net/fig/936069" pageId="0" pageNumber="65">2</figureCitation>
|
||
,
|
||
<figureCitation id="D596071B326EE1E3FB3ED08B8F5408B7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), mouthparts A gnathochilarium, ventral view B right anterior part of head, showing antennomere 1 and 2, labral lobe, mandible and stipital bundle C right mandible, ventral view D right mandible, dorsal view. Abbreviations (mandible): et external tooth, it internal tooth, mp molar plate, pl pectinate lamellae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure3" httpUri="https://binary.pensoft.net/fig/936070" pageId="0" pageNumber="65">3B</figureCitation>
|
||
). Gnathochilarium (Fig.
|
||
<figureCitation id="ED2F931643B6ECEF956AE43FBA523884" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), mouthparts A gnathochilarium, ventral view B right anterior part of head, showing antennomere 1 and 2, labral lobe, mandible and stipital bundle C right mandible, ventral view D right mandible, dorsal view. Abbreviations (mandible): et external tooth, it internal tooth, mp molar plate, pl pectinate lamellae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure3" httpUri="https://binary.pensoft.net/fig/936070" pageId="0" pageNumber="65">3A</figureCitation>
|
||
) subtrapezoidal, with a very wide distal part; stipites strongly developed, with rounded distolateral margins and characteristic long distal setae in a transverse row and distolateral bundles; lamellae linguales rectangular, with a transverse row of long distal setae, a group of somewhat shorter proximal setae, and a longitudinal row of lateral setae, lingual palps directed dorsad; promentum very short, deltoid. Mandibles (Figs
|
||
<figureCitation id="891D828C4760E0D86DECD7AA640B083A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), head A anterodorsal view B dorsal view C anterior part of head, anterodorsal view D lateral view E anterior view F head without gnathochilarium, dorsal view G head without gnathochilarium and right mandible, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.113498.figure2" httpUri="https://binary.pensoft.net/fig/936069" pageId="0" pageNumber="65">2A-F</figureCitation>
|
||
,
|
||
<figureCitation id="F60F5BCE3AC432134BC91690B0C78C86" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), mouthparts A gnathochilarium, ventral view B right anterior part of head, showing antennomere 1 and 2, labral lobe, mandible and stipital bundle C right mandible, ventral view D right mandible, dorsal view. Abbreviations (mandible): et external tooth, it internal tooth, mp molar plate, pl pectinate lamellae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure3" httpUri="https://binary.pensoft.net/fig/936070" pageId="0" pageNumber="65">3B-D</figureCitation>
|
||
) highly modified, dorsoventrally strongly flattened, external (
|
||
<emphasis id="7A2B5FE0C38347DAAF5CA8CCA5026969" bold="true" pageId="0" pageNumber="65">et</emphasis>
|
||
) and internal tooth (
|
||
<emphasis id="28512ECA0833E652BEEEF8872E9E16E9" bold="true" pageId="0" pageNumber="65">it</emphasis>
|
||
) reduced, four pectinate lamellae (pl) hypetrophied, with very long outher pectinate teeth, molar plate (
|
||
<emphasis id="C663B7D9932B504876B53CDC336B0AA4" bold="true" pageId="0" pageNumber="65">mp</emphasis>
|
||
) reduced. Antennae 2.30 mm long, their length ca 150% of vertical diameter of widest body ring. Length of antennomeres I-VIII (in mm): 0.17 (I), 0.57 (II), 0.40 (III), 0.35 (IV), 0.43 (V), 0.27 (VI), 0.07 (VII) and 0.04 (VIII). Length/width ratio of antennomeres I-VII: 1 (I), 2.8 (II), 2 (III), 2 (IV), 1.7 (V), 1 (VI) and 0.5 (VII). Antennomere I with a few very long anterior setae (Figs
|
||
<figureCitation id="3B3F11A21072EA45E9F3D7594425139C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), head A anterodorsal view B dorsal view C anterior part of head, anterodorsal view D lateral view E anterior view F head without gnathochilarium, dorsal view G head without gnathochilarium and right mandible, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.113498.figure2" httpUri="https://binary.pensoft.net/fig/936069" pageId="0" pageNumber="65">2E</figureCitation>
|
||
,
|
||
<figureCitation id="1D69C4D11604CA2D8B50960A954CAB0B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), mouthparts A gnathochilarium, ventral view B right anterior part of head, showing antennomere 1 and 2, labral lobe, mandible and stipital bundle C right mandible, ventral view D right mandible, dorsal view. Abbreviations (mandible): et external tooth, it internal tooth, mp molar plate, pl pectinate lamellae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure3" httpUri="https://binary.pensoft.net/fig/936070" pageId="0" pageNumber="65">3B</figureCitation>
|
||
); antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
|
||
</paragraph>
|
||
<paragraph id="6794AC030C0627A762C896EEE977DA39" pageId="0" pageNumber="65">
|
||
<emphasis id="0C6ECA42B8921302CFAE7D082B089A7B" bold="true" italics="true" pageId="0" pageNumber="65">Body rings</emphasis>
|
||
: body in general moniliform (Fig.
|
||
<figureCitation id="8CC51CF332597E453BB8362D2FDDA49F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1A</figureCitation>
|
||
). Metazonae strongly vaulted compared to prozonae (Fig.
|
||
<figureCitation id="E5EC6B0DC2F63BE7546B1F6167990EFF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1A, C, E, F</figureCitation>
|
||
). Entire metazonal area with longitudinal striations, striae more numerous and denser below ozopore and rarer and less conspicuous above ozopore (Fig.
|
||
<figureCitation id="9E7EE0E6D9A74CDC1F3A1395063C2418" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1E</figureCitation>
|
||
). Length of metazonal setae ca 45% of vertical diameter of rings, ca 40 per ring (Fig.
|
||
<figureCitation id="2ECF318D7DC52AF2526347E82F97F6C4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1E, F</figureCitation>
|
||
). Posterior margin of metazona thickened dorsolaterally (Fig.
|
||
<figureCitation id="F7FB51F1BE01905945AE093491096AB8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1F</figureCitation>
|
||
). Ozopores behind pro-metazonal suture at ca
|
||
<normalizedToken id="424DFBB46795F73AE334840A14295709" originalValue="½">1/2</normalizedToken>
|
||
of metazonal length (Fig.
|
||
<figureCitation id="704F49B9DBEAB17B664711E9187A0761" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1396AC107A359705D442A7D89ABEEBC9" pageId="0" pageNumber="65">
|
||
<emphasis id="CEF3DB2419A8C11F6AACC74C338F7FF1" bold="true" italics="true" pageId="0" pageNumber="65">Pleurotergum 7</emphasis>
|
||
: ventral margin with very low, poorly developed lobe (Fig.
|
||
<figureCitation id="0D1E5AE062ABC5CDD65874D69A2C6658" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BA2CC40157C0C4E2D30D4E7EEEC00640" pageId="0" pageNumber="65">
|
||
<emphasis id="A717D2F0B1936AC3E552D471FFE967A0" bold="true" italics="true" pageId="0" pageNumber="65">Telson</emphasis>
|
||
: pre-anal ring densely setose, with a short, acuminate epiproct (Fig.
|
||
<figureCitation id="449B2AF1EAA5E5EB6BBC97E88E41CFEB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1G, H</figureCitation>
|
||
). Paraprocts rounded, each with ca 30-35 long setae over entire surface. Hypoproct in form of symetrical subtriangle, covered with ca 15 long setae, without any modifications.
|
||
</paragraph>
|
||
<paragraph id="BCD934B76A8C1DE4F30DC7F908141616" pageId="0" pageNumber="65">
|
||
<emphasis id="E8E826EF429E06834B6A19112A99A701" bold="true" italics="true" pageId="0" pageNumber="65">Legs</emphasis>
|
||
: leg-pair 1 modified, hook-shaped, with three complete podomeres; coxa with one seta; prefemur with five setae; femur, postfemur, and tibiotarsus coalesced, with indications of segmentation; femur with two or three setae; postfemur with one seta; tibiotarsal part with a small distal lobe and four setae; tip slightly tuberculate (Figs
|
||
<figureCitation id="8A5A117942AA841CC544B47887A73E67" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1C</figureCitation>
|
||
,
|
||
<figureCitation id="880EB6BC8F978C67CDB53BE5F44B4E43" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4A, B</figureCitation>
|
||
,
|
||
<figureCitation id="FA7F45E0FA854AED1F665372A343BE0E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C penis, posterior view D right gonopods, mesal view E left gonopods, lateral view F right promere, posterior view G right mesomere, anterior view. Abbreviations: m mesomere, ml mesal lobe, o opisthomere, p promere, s solenomere, t telopodite, v velum. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1184.113498.figure5" httpUri="https://binary.pensoft.net/fig/936072" pageId="0" pageNumber="65">5A, B</figureCitation>
|
||
). Anterior walking legs with an adhesive pad on tibia (Fig.
|
||
<figureCitation id="F4C1A0E56E57A69D3AC20AFFE104619E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4C, E</figureCitation>
|
||
), most pronounced in leg-pair 2 (Fig.
|
||
<figureCitation id="47B50FAF31B5C27A5905EAA791937C9B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4C</figureCitation>
|
||
), gradually disappearing in posterior direction, completely disappearing around mid-body (Fig.
|
||
<figureCitation id="F0C614ED5BA341895E47F17689DA3995" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4F</figureCitation>
|
||
). Length of mid-body legs ca 1.45 times as long as mid-body vertical diameter; tarsus ca 2 times as long as tibia with very long, spinelike ventral seta; apical claws strongly developed, very long, ca 60% of tarsus length (Fig.
|
||
<figureCitation id="01509454A8F14B8FE521F214B6A2303A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4F</figureCitation>
|
||
); no accesory claw.
|
||
</paragraph>
|
||
<caption id="073ABBFEBCAA728EA4EB1B7FCF5A4510" doi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65" start="Figure 4" startId="F4">
|
||
<paragraph id="BEB57041E0C25090F1168A05A1A78068" pageId="0" pageNumber="65">
|
||
<emphasis id="9C333F4397334E05F2A984E26239C72B" bold="true" pageId="0" pageNumber="65">Figure 4.</emphasis>
|
||
<taxonomicName id="2BBDEC5780D08256FA1299404E3AA0EB" authorityName="Antić & Akkari" authorityYear="2023" class="Diplopoda" family="Julidae" genus="Inversotyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Inversotyphlus ammirandus" order="Julida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="ammirandus">
|
||
<emphasis id="CE214EB1F9B493139AFBCC5A6AE2E634" italics="true" pageId="0" pageNumber="65">Inversotyphlus ammirandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype ♂ (NHMW MY10380)
|
||
<emphasis id="A9F73840F80B525823E27836B507F49C" bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
leg-pair 1, anterior view
|
||
<emphasis id="ED031179D9DD019E374AD0FA216D4614" bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
left leg 1, lateral view
|
||
<emphasis id="48B942C550FE2BBFFE6DDE015AD74BEF" bold="true" pageId="0" pageNumber="65">C</emphasis>
|
||
telopodites of leg-pair 2, posterior view
|
||
<emphasis id="96539D9808C0561FD7900A997C32A91A" bold="true" pageId="0" pageNumber="65">D</emphasis>
|
||
penis and coxae of leg-pair 2, posterior view
|
||
<emphasis id="840C97365AE4248C2269BCCF075147BD" bold="true" pageId="0" pageNumber="65">E</emphasis>
|
||
left telopodite 10, anterior view
|
||
<emphasis id="CB898CEE2C6B49B4E162EEEB2BA9EC37" bold="true" pageId="0" pageNumber="65">F</emphasis>
|
||
right mid-body telopodite, anterior view
|
||
<emphasis id="DAE331EA8B8AE77E9234B7E664E2CD20" bold="true" pageId="0" pageNumber="65">G</emphasis>
|
||
right gonopods, mesal view
|
||
<emphasis id="73D68A376CA196EA4673950BA9080797" bold="true" pageId="0" pageNumber="65">H</emphasis>
|
||
left gonopods, posterior view
|
||
<emphasis id="00BFFDBE1A1FE845C4D7A22EB897B44B" bold="true" pageId="0" pageNumber="65">I</emphasis>
|
||
right promere, posterior view
|
||
<emphasis id="E35BDD3EA9A4ADDA0461711D84B3B702" bold="true" pageId="0" pageNumber="65">J</emphasis>
|
||
right mesomere, anterior view. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CD792A930EDBCC6CA296DC3D0A341F94" pageId="0" pageNumber="65">
|
||
<emphasis id="481494D332D19E1F0E83037A72681893" bold="true" italics="true" pageId="0" pageNumber="65">Penis</emphasis>
|
||
: bilobed. Lobes prominent, elongated (Figs
|
||
<figureCitation id="5FC05F7B97DFB0A58D00EDD6F64EA47C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4D</figureCitation>
|
||
,
|
||
<figureCitation id="5FC8413FE6C5B23E603DF030B0180FE6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C penis, posterior view D right gonopods, mesal view E left gonopods, lateral view F right promere, posterior view G right mesomere, anterior view. Abbreviations: m mesomere, ml mesal lobe, o opisthomere, p promere, s solenomere, t telopodite, v velum. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1184.113498.figure5" httpUri="https://binary.pensoft.net/fig/936072" pageId="0" pageNumber="65">5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BFD527CE85BD4E4C4A25D352B3E3EAED" pageId="0" pageNumber="65">
|
||
<emphasis id="7DDFF7385071B94EC8835CC365D1DDD1" bold="true" italics="true" pageId="0" pageNumber="65">Gonopods</emphasis>
|
||
:
|
||
<emphasis id="ABFEB5DC3C9016B22C1797D2A0479EC5" italics="true" pageId="0" pageNumber="65">in situ</emphasis>
|
||
protruding from gonopodal sinus (Fig.
|
||
<figureCitation id="1B48746588A117BAB049587F95D7E5B3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380), habitus A whole animal, dorsal view B anterior part of body, ventral, somewhat lateral view C body segments 1 - 7, with leg-pair 1, walking legs and gonopods, lateral view D right half of pleurotergum 7, lateral view E body rings 7 - 9, dorsolateral view F mid-body rings, lateral view G end of body, lateral view H end of body, dorsal view. Scale bars: 1 mm (A-C, F); 0.5 mm (D, E, G, H)." figureDoi="10.3897/zookeys.1184.113498.figure1" httpUri="https://binary.pensoft.net/fig/936068" pageId="0" pageNumber="65">1B, C</figureCitation>
|
||
). Promere (
|
||
<emphasis id="254571A7394D9CC29B7EA96C06F1EABA" bold="true" pageId="0" pageNumber="65">p</emphasis>
|
||
) and mesomere (
|
||
<emphasis id="6FBA9176849808342C7BE3AE1301F824" bold="true" pageId="0" pageNumber="65">m</emphasis>
|
||
) surpassed by opisthomere (
|
||
<emphasis id="B99E3A505BC0702E1EE35B4C57CDA334" bold="true" pageId="0" pageNumber="65">o</emphasis>
|
||
) (Figs
|
||
<figureCitation id="5CB5407972874443BC445DD21695B79A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4G, H</figureCitation>
|
||
,
|
||
<figureCitation id="021FEFB78B94633B2500F0130F2A9560" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C penis, posterior view D right gonopods, mesal view E left gonopods, lateral view F right promere, posterior view G right mesomere, anterior view. Abbreviations: m mesomere, ml mesal lobe, o opisthomere, p promere, s solenomere, t telopodite, v velum. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1184.113498.figure5" httpUri="https://binary.pensoft.net/fig/936072" pageId="0" pageNumber="65">5D, E</figureCitation>
|
||
,
|
||
<figureCitation id="51CAC1A7802F46E3E6306FCB6946AEDA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Gonopods of the members of the genus Inversotyphlus Strasser, 1962 stat. nov., mesal views A I. ammirandus sp. nov., holotype ♂ (NHMW MY 10380), right gonopods (flipped) B I. clavatus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods C I. edentulus (Attems, 1951), comb. nov., ♂ from Njegos Cave (IZB), left gonopods D I. opisthonodus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods E I. gellianae (Makarov & Rađa, 2006), comb. nov., holotype ♂ (IZB), left gonopods F I. gracilis (Antic, 2018), comb. nov., paratype ♂ (IZB), left gonopods G I. lobifer (Attems, 1951), comb. nov., ♂ from bitumen mine Minjera, island of Brac (IZB), left gonopods. B, D-G modified after Antic et al. (2018). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure6" httpUri="https://binary.pensoft.net/fig/936073" pageId="0" pageNumber="65">6A</figureCitation>
|
||
). Promere (Figs
|
||
<figureCitation id="B0F146033FCED551C1CA6903EEE911BD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4G, I</figureCitation>
|
||
,
|
||
<figureCitation id="B82E6156524E382E269A426DCC858B79" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C penis, posterior view D right gonopods, mesal view E left gonopods, lateral view F right promere, posterior view G right mesomere, anterior view. Abbreviations: m mesomere, ml mesal lobe, o opisthomere, p promere, s solenomere, t telopodite, v velum. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1184.113498.figure5" httpUri="https://binary.pensoft.net/fig/936072" pageId="0" pageNumber="65">5D-F</figureCitation>
|
||
,
|
||
<figureCitation id="E5B64DF63DB9D3E135EADF1D0619065D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Gonopods of the members of the genus Inversotyphlus Strasser, 1962 stat. nov., mesal views A I. ammirandus sp. nov., holotype ♂ (NHMW MY 10380), right gonopods (flipped) B I. clavatus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods C I. edentulus (Attems, 1951), comb. nov., ♂ from Njegos Cave (IZB), left gonopods D I. opisthonodus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods E I. gellianae (Makarov & Rađa, 2006), comb. nov., holotype ♂ (IZB), left gonopods F I. gracilis (Antic, 2018), comb. nov., paratype ♂ (IZB), left gonopods G I. lobifer (Attems, 1951), comb. nov., ♂ from bitumen mine Minjera, island of Brac (IZB), left gonopods. B, D-G modified after Antic et al. (2018). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure6" httpUri="https://binary.pensoft.net/fig/936073" pageId="0" pageNumber="65">6A</figureCitation>
|
||
) somewhat shorter than mesomere; subrectangular in anterior and posterior views; sides almost parallel, mesal margin straight, lateral margin slightly convex; distally microsquamose; mesal lobe (
|
||
<emphasis id="34C8BA408EFB19C25CDFF73FD2B63244" bold="true" pageId="0" pageNumber="65">ml</emphasis>
|
||
) well developed, subtriangular in lateral and mesal views, with one or two long distal setae; telopodite (
|
||
<emphasis id="AF79F66D2AF3799777ECB4C519F368B5" bold="true" pageId="0" pageNumber="65">t</emphasis>
|
||
) smaller than mesal lobe, egg-shaped. Mesomere (Figs
|
||
<figureCitation id="C4426383FBA828092ED8142F614101C1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4G, H, J</figureCitation>
|
||
,
|
||
<figureCitation id="25203C2403A7C8E6B840A4CC9145FC0B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C penis, posterior view D right gonopods, mesal view E left gonopods, lateral view F right promere, posterior view G right mesomere, anterior view. Abbreviations: m mesomere, ml mesal lobe, o opisthomere, p promere, s solenomere, t telopodite, v velum. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1184.113498.figure5" httpUri="https://binary.pensoft.net/fig/936072" pageId="0" pageNumber="65">5D, E, G</figureCitation>
|
||
,
|
||
<figureCitation id="3A165F5B5FCA4D457B95873C1D88EFBF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Gonopods of the members of the genus Inversotyphlus Strasser, 1962 stat. nov., mesal views A I. ammirandus sp. nov., holotype ♂ (NHMW MY 10380), right gonopods (flipped) B I. clavatus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods C I. edentulus (Attems, 1951), comb. nov., ♂ from Njegos Cave (IZB), left gonopods D I. opisthonodus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods E I. gellianae (Makarov & Rađa, 2006), comb. nov., holotype ♂ (IZB), left gonopods F I. gracilis (Antic, 2018), comb. nov., paratype ♂ (IZB), left gonopods G I. lobifer (Attems, 1951), comb. nov., ♂ from bitumen mine Minjera, island of Brac (IZB), left gonopods. B, D-G modified after Antic et al. (2018). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure6" httpUri="https://binary.pensoft.net/fig/936073" pageId="0" pageNumber="65">6A</figureCitation>
|
||
) spoon-shaped in anterior view, slightly sigmoid in lateral and mesal views; anterodistal margin slightly denticulate; anterior side distally concave and microsquamose. Opisthomere (Figs
|
||
<figureCitation id="BD60FC9BC5E27EEA18F4F3F54338C16C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C telopodites of leg-pair 2, posterior view D penis and coxae of leg-pair 2, posterior view E left telopodite 10, anterior view F right mid-body telopodite, anterior view G right gonopods, mesal view H left gonopods, posterior view I right promere, posterior view J right mesomere, anterior view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure4" httpUri="https://binary.pensoft.net/fig/936071" pageId="0" pageNumber="65">4G, H</figureCitation>
|
||
,
|
||
<figureCitation id="EBAFDF859D1FC382365F271745C16EFB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Inversotyphlus ammirandus sp. nov., holotype ♂ (NHMW MY 10380) A leg-pair 1, anterior view B left leg 1, lateral view C penis, posterior view D right gonopods, mesal view E left gonopods, lateral view F right promere, posterior view G right mesomere, anterior view. Abbreviations: m mesomere, ml mesal lobe, o opisthomere, p promere, s solenomere, t telopodite, v velum. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1184.113498.figure5" httpUri="https://binary.pensoft.net/fig/936072" pageId="0" pageNumber="65">5D, E</figureCitation>
|
||
,
|
||
<figureCitation id="CF39A13FE30E9B97123849EACA7D9E41" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Gonopods of the members of the genus Inversotyphlus Strasser, 1962 stat. nov., mesal views A I. ammirandus sp. nov., holotype ♂ (NHMW MY 10380), right gonopods (flipped) B I. clavatus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods C I. edentulus (Attems, 1951), comb. nov., ♂ from Njegos Cave (IZB), left gonopods D I. opisthonodus (Antic, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods E I. gellianae (Makarov & Rađa, 2006), comb. nov., holotype ♂ (IZB), left gonopods F I. gracilis (Antic, 2018), comb. nov., paratype ♂ (IZB), left gonopods G I. lobifer (Attems, 1951), comb. nov., ♂ from bitumen mine Minjera, island of Brac (IZB), left gonopods. B, D-G modified after Antic et al. (2018). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1184.113498.figure6" httpUri="https://binary.pensoft.net/fig/936073" pageId="0" pageNumber="65">6A</figureCitation>
|
||
) straight and slender with distal half slightly directed anteriad; proximomesal spine present; anteromesal lamella poorly developed; velum (
|
||
<emphasis id="907C345EAEA0424B42FDD5AEB6BA49DD" bold="true" pageId="0" pageNumber="65">v</emphasis>
|
||
) unipartite, tapering anterodistad, with fimbriated tip; solenomere (
|
||
<emphasis id="63F017BB48929C423CA8F6AD3DDF066D" bold="true" pageId="0" pageNumber="65">s</emphasis>
|
||
) narrow, tubelike.
|
||
</paragraph>
|
||
<caption id="C392CC881099A57B7A4FA9726F3AB70C" doi="10.3897/zookeys.1184.113498.figure5" httpUri="https://binary.pensoft.net/fig/936072" pageId="0" pageNumber="65" start="Figure 5" startId="F5">
|
||
<paragraph id="A014FBAF7F1255422C3BA9B71F75DF40" pageId="0" pageNumber="65">
|
||
<emphasis id="B75BE43AC83CB87DCDB6C425A3B56E5C" bold="true" pageId="0" pageNumber="65">Figure 5.</emphasis>
|
||
<taxonomicName id="CDA39AE96BBFD7C649F0B6986E685EB2" authorityName="Antić & Akkari" authorityYear="2023" class="Diplopoda" family="Julidae" genus="Inversotyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Inversotyphlus ammirandus" order="Julida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="ammirandus">
|
||
<emphasis id="64FD9CA62BF8193DB77EDDF211CFCEB7" italics="true" pageId="0" pageNumber="65">Inversotyphlus ammirandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype ♂ (NHMW MY10380)
|
||
<emphasis id="135F8A6AE4D09EB9C82DAED660895DF5" bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
leg-pair 1, anterior view
|
||
<emphasis id="0FD2B878403D7F3BA15462A5EC6706F6" bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
left leg 1, lateral view
|
||
<emphasis id="CD7E7D4942665FB6EF9037992ED3B703" bold="true" pageId="0" pageNumber="65">C</emphasis>
|
||
penis, posterior view
|
||
<emphasis id="A13872DD5669B204AA3A9E0C319AEC2F" bold="true" pageId="0" pageNumber="65">D</emphasis>
|
||
right gonopods, mesal view
|
||
<emphasis id="CC5EABD4275B5C6D763CC2A7E31049B9" bold="true" pageId="0" pageNumber="65">E</emphasis>
|
||
left gonopods, lateral view
|
||
<emphasis id="270D70CD26823854FB574898251E51B1" bold="true" pageId="0" pageNumber="65">F</emphasis>
|
||
right promere, posterior view
|
||
<emphasis id="D23F99FD605A74249DB1821E7F6DE1E3" bold="true" pageId="0" pageNumber="65">G</emphasis>
|
||
right mesomere, anterior view. Abbreviations:
|
||
<emphasis id="60BE2DF795A791B9C7DE17083C2464AA" bold="true" pageId="0" pageNumber="65">m</emphasis>
|
||
mesomere,
|
||
<emphasis id="122745FA341F0387A19B941680C943B8" bold="true" pageId="0" pageNumber="65">ml</emphasis>
|
||
mesal lobe,
|
||
<emphasis id="2F6B374DF8659C2A87DD115EC73B2FFD" bold="true" pageId="0" pageNumber="65">o</emphasis>
|
||
opisthomere,
|
||
<emphasis id="547FA86EB95A1D26DAFFF92AF4CC3F34" bold="true" pageId="0" pageNumber="65">p</emphasis>
|
||
promere,
|
||
<emphasis id="772D88261B0A330DC028FBF42FFCB4C5" bold="true" pageId="0" pageNumber="65">s</emphasis>
|
||
solenomere,
|
||
<emphasis id="C73DD3C6ADB11640432B69647F5D8901" bold="true" pageId="0" pageNumber="65">t</emphasis>
|
||
telopodite,
|
||
<emphasis id="185CB9184B9FD90FDE8B79892E9266E9" bold="true" pageId="0" pageNumber="65">v</emphasis>
|
||
velum. Scale bars: 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="09D8D584168BC9D3C33F0B47B8273AC5" doi="10.3897/zookeys.1184.113498.figure6" httpUri="https://binary.pensoft.net/fig/936073" pageId="0" pageNumber="65" start="Figure 6" startId="F6">
|
||
<paragraph id="6288E7D05EF624E8CF628CA6F081E5FA" pageId="0" pageNumber="65">
|
||
<emphasis id="0AEB4E7D776E0C7B5E595B874B9C763A" bold="true" pageId="0" pageNumber="65">Figure 6.</emphasis>
|
||
Gonopods of the members of the genus
|
||
<taxonomicName id="DA03F6DCBE4C156A3070ADA361132CC9" authorityName="Antić & Akkari" authorityYear="2023" class="Diplopoda" family="Julidae" genus="Inversotyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Inversotyphlus" order="Julida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DD5DECD7A6CF106ED09E53B2A73CB0EC" italics="true" pageId="0" pageNumber="65">Inversotyphlus</emphasis>
|
||
</taxonomicName>
|
||
Strasser, 1962 stat. nov., mesal views
|
||
<emphasis id="36B8834B1D43CD5A23AD3A5B5B5BC0D6" bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
<taxonomicName id="A5F09FD77C474C9A2C84910EE061A584" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. ammirandus" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="ammirandus">
|
||
<emphasis id="FFABCF855F77A9A38BB807EAE9BD2468" italics="true" pageId="0" pageNumber="65">I. ammirandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype ♂ (NHMW MY10380), right gonopods (flipped)
|
||
<emphasis id="C0F24E37A10419D28DB3101D50225F19" bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
<taxonomicName id="7224950F4F39E28BD8BF9CB439B9E22A" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. clavatus" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="clavatus">
|
||
<emphasis id="DD8F71999AE5257146CE0F53AB087E12" italics="true" pageId="0" pageNumber="65">I. clavatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken id="02D7FD7C60F75511A0D8DAD6E6D12E80" originalValue="Antić">Antic</normalizedToken>
|
||
, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods
|
||
<emphasis id="445DA59E6774B60AA466A109F0BEC61B" bold="true" pageId="0" pageNumber="65">C</emphasis>
|
||
<taxonomicName id="79D05CE20DC2A1C82C521C32850FD96F" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. edentulus" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="edentulus">
|
||
<emphasis id="B9438593F612616FF9B03F84491983BB" italics="true" pageId="0" pageNumber="65">I. edentulus</emphasis>
|
||
</taxonomicName>
|
||
(Attems, 1951), comb. nov., ♂ from
|
||
<normalizedToken id="5C51CEB1299F24C3CAAC5FE8B5DAA1F8" originalValue="Njegoš">Njegos</normalizedToken>
|
||
Cave (IZB), left gonopods
|
||
<emphasis id="A5AF0D15C5AAA745AC9992389BC33992" bold="true" pageId="0" pageNumber="65">D</emphasis>
|
||
<taxonomicName id="8296E069E50E0774B70FD0AAEE14EF87" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. opisthonodus" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="opisthonodus">
|
||
<emphasis id="307295BBEDCA7DE177859FB74BC68B42" italics="true" pageId="0" pageNumber="65">I. opisthonodus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken id="912EE8DF63316B1E7BCD4529CBC3FCD0" originalValue="Antić">Antic</normalizedToken>
|
||
, 2018), comb. nov., holotype ♂ (NHMSC), left gonopods
|
||
<emphasis id="A4CF34F2EB5C0B422404762FE2934AD1" bold="true" pageId="0" pageNumber="65">E</emphasis>
|
||
<taxonomicName id="68B21830D72E6EA2EEFCC9351438D634" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. gellianae" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="gellianae">
|
||
<emphasis id="97A9A2C69FD965A4114BABFEC3060DB0" italics="true" pageId="0" pageNumber="65">I. gellianae</emphasis>
|
||
</taxonomicName>
|
||
(Makarov &
|
||
<normalizedToken id="F1656E324805E03FD4BFA07AC3A81BC1" originalValue="Rađa">Rađa</normalizedToken>
|
||
, 2006), comb. nov., holotype ♂ (IZB), left gonopods
|
||
<emphasis id="A706F8CD708B043756BD797FCEE5C616" bold="true" pageId="0" pageNumber="65">F</emphasis>
|
||
<taxonomicName id="F8794C5F63C9A5293CF6CBD52E35C35E" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. gracilis" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="5962B3DFCE21EF9FB447C5389711BC31" italics="true" pageId="0" pageNumber="65">I. gracilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken id="3788DED69DF7D62AE07E831BBA772339" originalValue="Antić">Antic</normalizedToken>
|
||
, 2018), comb. nov., paratype ♂ (IZB), left gonopods
|
||
<emphasis id="65F551EFE8754983625E0B6715B0C0A5" bold="true" pageId="0" pageNumber="65">G</emphasis>
|
||
<taxonomicName id="8BD670D0710110468689ED95A024D0CE" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="I. lobifer" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="lobifer">
|
||
<emphasis id="4251C5FB785730E144CE23F6DA5ED652" italics="true" pageId="0" pageNumber="65">I. lobifer</emphasis>
|
||
</taxonomicName>
|
||
(Attems, 1951), comb. nov., ♂ from bitumen mine Minjera, island of
|
||
<normalizedToken id="ED2D33358B389B023160E9BF779E8BE8" originalValue="Brač">Brac</normalizedToken>
|
||
(IZB), left gonopods.
|
||
<emphasis id="0C4292FC1306E9387680A9D0084FC2DB" bold="true" pageId="0" pageNumber="65">B, D-G</emphasis>
|
||
modified after
|
||
<bibRefCitation id="9B2F160CEEBF3BB690403FF88B863119" DOI="https://doi.org/10.11646/zootaxa.4455.2.2" author="Antic, DZ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="65" pagination="258 - 294" refId="B5" refString="Antic, DZ, Drazina, T, Rađa, T, Lucic, LR, Makarov, SE, 2018. Review of the genus Typhloiulus Latzel, 1884 in the Dinaric region, with a description of four new species and the first description of the male of Typhloiulus insularis Strasser, 1938 (Diplopoda: Julida: Julidae). Zootaxa 4455 (2): 258 - 294, DOI: https://doi.org/10.11646/zootaxa.4455.2.2" title="Review of the genus Typhloiulus Latzel, 1884 in the Dinaric region, with a description of four new species and the first description of the male of Typhloiulus insularis Strasser, 1938 (Diplopoda: Julida: Julidae)." url="https://doi.org/10.11646/zootaxa.4455.2.2" volume="4455" year="2018">
|
||
<normalizedToken id="0FCFA175B6A00F379035A628CCD1295C" originalValue="Antić">Antic</normalizedToken>
|
||
et al. (2018)
|
||
</bibRefCitation>
|
||
. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="4B50C6CA883115A043351DC0776BB0B3" pageId="0" pageNumber="65" type="habitat">
|
||
<paragraph id="A919B64AA50E66167E50594FB1CDA47A" pageId="0" pageNumber="65">Habitat.</paragraph>
|
||
<paragraph id="C1DBAEAD193AEA1F075BF57E923C0653" pageId="0" pageNumber="65">
|
||
The Ru Cave (Fig.
|
||
<figureCitation id="8D5DD156C40107C26A893B4336CA1EB3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ru Cave and surroundings A entrance to cave (photograph: Vladimir Georgiev) B Radohimes massif of the Prokletije Mountains where the cave is located (photograph: Marinko Malenica)." figureDoi="10.3897/zookeys.1184.113498.figure7" httpUri="https://binary.pensoft.net/fig/936074" pageId="0" pageNumber="65">7A</figureCitation>
|
||
) was discovered by Bulgarian speleologists in 2010. It is located at an altitude of 2225 m in the
|
||
<normalizedToken id="639E13A50B46194D5016D688FED593FB" originalValue="Radohimës">Radohimes</normalizedToken>
|
||
massif (Fig.
|
||
<figureCitation id="7C9AF8B4467EB149417509265F3FDB83" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ru Cave and surroundings A entrance to cave (photograph: Vladimir Georgiev) B Radohimes massif of the Prokletije Mountains where the cave is located (photograph: Marinko Malenica)." figureDoi="10.3897/zookeys.1184.113498.figure7" httpUri="https://binary.pensoft.net/fig/936074" pageId="0" pageNumber="65">7B</figureCitation>
|
||
) of the Prokletije Mountains. With an explored depth of 574 m, this cave (length 1300 m) is the second deepest cave known in Albania. It has a predominantly vertical character and is risky in terms of rockfall and flooding. The new species was found in a short horizontal meander whose bottom is covered by water, 460 m below the entrance. In addition to the new species, representatives of
|
||
<taxonomicName id="C65BCD67084E8E8A036BFB3E71F2C7F1" class="Clitellata" family="Lumbricidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Crassiclitellata" pageId="0" pageNumber="65" phylum="Annelida" rank="family">Lumbricidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="F1C99E5D08D2F3185DB4C3DC2564B6EE" authorityName="Fleming" authorityYear="1821" class="Arachnida" family="Leiodidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
|
||
, and
|
||
<taxonomicName id="BDC6F1A0C6683CA13AF7821C7CECD27D" class="Aracnida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="65" rank="order">Opiliones</taxonomicName>
|
||
, as well as
|
||
<taxonomicName id="8170452D18DF84E16C017D03186B64A1" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="order">Chordeumatida</taxonomicName>
|
||
(probably
|
||
<taxonomicName id="3B1983AE394C47139AB947B69278C36D" authorityName="Absolon & Lang" authorityYear="1933" class="Diplopoda" family="Anthogonidae" genus="Macrochaetosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Macrochaetosoma" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="65FC1ED68A17396DC7F55CB5C2A5EC41" italics="true" pageId="0" pageNumber="65">Macrochaetosoma</emphasis>
|
||
</taxonomicName>
|
||
Absolon & Lang, 1933), were also found in the cave at various depths (Tamara
|
||
<normalizedToken id="5DEBBB7EED617A05D9CA19F9A3638CCA" originalValue="Čuković">Cukovic</normalizedToken>
|
||
and Vladimir Georgiev, pers. comm.).
|
||
</paragraph>
|
||
<caption id="C33BB7F4583F9E3D6112EF518C5B281B" doi="10.3897/zookeys.1184.113498.figure7" httpUri="https://binary.pensoft.net/fig/936074" pageId="0" pageNumber="65" start="Figure 7" startId="F7">
|
||
<paragraph id="906EA38261BE2658F90878D49B7478E6" pageId="0" pageNumber="65">
|
||
<emphasis id="553528EED7442C0B49E67707EE740962" bold="true" pageId="0" pageNumber="65">Figure 7.</emphasis>
|
||
Ru Cave and surroundings
|
||
<emphasis id="F5ED426F1136F770732F543E467C71D2" bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
entrance to cave (photograph: Vladimir Georgiev)
|
||
<emphasis id="3AB5E9FB14B396733BB2DCD97E4272DB" bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
<normalizedToken id="55323FACACA7450EC9D774CB029186C2" originalValue="Radohimës">Radohimes</normalizedToken>
|
||
massif of the Prokletije Mountains where the cave is located (photograph: Marinko Malenica).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="134F0901EA2E8EBE69BDEC59F8FE8566" pageId="0" pageNumber="65" type="remarks">
|
||
<paragraph id="CA769448393B97A8DA09CAA8FEC8023D" pageId="0" pageNumber="65">Remarks.</paragraph>
|
||
<paragraph id="BF985AAFF2ECD955A370CA9112C4CFAF" pageId="0" pageNumber="65">
|
||
Besides the chordeumatidan
|
||
<taxonomicName id="BA87962450E008127C364A05529956AA" authorityName="Absolong & Lang" authorityYear="1933" class="Diplopoda" family="Anthogonidae" genus="Macrochaetosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Macrochaetosoma troglomontanum" order="Chordeumatida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="troglomontanum">
|
||
<emphasis id="BB0A04C9ED2BD68CF5FED816D9EBF1A3" italics="true" pageId="0" pageNumber="65">Macrochaetosoma troglomontanum</emphasis>
|
||
</taxonomicName>
|
||
Absolon & Lang, 1933 (see
|
||
<bibRefCitation id="C524114ED020A0D7CB8D4A36CF8B2ACC" author="Absolon, K" pageId="0" pageNumber="65" refId="B1" refString="Absolon, K, Lang, J, 1933. Ueber Macrochaetosoma troglomontana nov. gen. n. sp. (Diplopoda) aus dem suedillyrischen Karstgebiete. Zoologischer Anzeiger 101(5/6): 137-143." title="Ueber Macrochaetosoma troglomontana nov. gen. n. sp. (Diplopoda) aus dem suedillyrischen Karstgebiete. Zoologischer Anzeiger 101 (5 / 6): 137 - 143." year="1933">Absolon and Lang 1933</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="91A5B7B1613D44E8649E5502BCAD8D5E" author="Stoev, P" journalOrPublisher="Historia Naturalis Bulgarica" pageId="0" pageNumber="65" pagination="109 - 110" refId="B16" refString="Stoev, P, 2001. On two myriapods (Chilopoda, Diplopoda) new to the fauna of Albania. Historia Naturalis Bulgarica 13: 109 - 110" title="On two myriapods (Chilopoda, Diplopoda) new to the fauna of Albania." volume="13" year="2001">Stoev 2001</bibRefCitation>
|
||
),
|
||
<taxonomicName id="38DF1192F57586600F93534DB0C2E648" authorityName="Antić & Akkari" authorityYear="2023" class="Diplopoda" family="Julidae" genus="Inversotyphlus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Inversotyphlus ammirandus" order="Julida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="ammirandus">
|
||
<emphasis id="288FD17E4F2552B17853BD0CA58303F0" italics="true" pageId="0" pageNumber="65">Inversotyphlus ammirandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is the second undisputed troglobiotic millipede in the fauna of Albania. In addition, this is one of the rare cave-dwelling julids characterised by modifications of the body and mouthparts for a semiaquatic life and filtering diet, and also the first such millipede in the territory of Albania.
|
||
</paragraph>
|
||
<paragraph id="30E53746D71F46929BA6447CF301DBFF" pageId="0" pageNumber="65">
|
||
It is interesting to note that several ultraspecialized hygropetricolous leptodirinine beetles were recently described from this Albanian part of the
|
||
<normalizedToken id="B0D5B5F486C6EB988C79C288B45A7EA2" originalValue="Prokletiје">Prokletiјe</normalizedToken>
|
||
Mountains (
|
||
<bibRefCitation id="32C59ECB300E7DBF44EB7480814DECAE" author="Giachino, PM" journalOrPublisher="Suplementos del Boletin de la Asociacion espanola de Entomologia" pageId="0" pageNumber="65" pagination="34 - 42" refId="B11" refString="Giachino, PM, Casale, A, 2022. New hygropetricolous Leptodirini from Albania (Coleoptera, Leiodidae, Cholevinae). Suplementos del Boletin de la Asociacion espanola de Entomologia 4: 34 - 42" title="New hygropetricolous Leptodirini from Albania (Coleoptera, Leiodidae, Cholevinae)." volume="4" year="2022">Giachino and Casale 2022</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="93132375CF964AC1968F1DAB359FE5EA" pageId="0" pageNumber="65">
|
||
In addition to the two unequivocal troglobionts mentioned above, there are two other Albanian millipede species that may potentially be troglobiotic,
|
||
<taxonomicName id="0AC2F8946B27C3B493E9F46A7F4A2655" authorityName="Mauries, Golovatch & Stoev" authorityYear="1997" class="Diplopoda" family="Paradoxosomatidae" genus="Metonomastus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Metonomastus petrelensis" order="Polydesmida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="petrelensis">
|
||
<emphasis id="F389016BAEDC9346DAAB90FC14F614EF" italics="true" pageId="0" pageNumber="65">Metonomastus petrelensis</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="2720FCC09A9F575EBB2DA6D8E535AF88" originalValue="Mauriès">Mauries</normalizedToken>
|
||
, Golovatch & Stoev, 1997 and
|
||
<taxonomicName id="D2B4FE7E860C576A16BDB7DBFBD59104" authorityName="Mauries, Golovatch & Stoev" authorityYear="1997" class="Diplopoda" family="Julidae" genus="Typhloiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloiulus beroni" order="Julida" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="beroni">
|
||
<emphasis id="8193B17635EEF8E3FBA849A288FF92DF" italics="true" pageId="0" pageNumber="65">Typhloiulus beroni</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="B42C0DD8C3500E069F8B29B566C67FBD" originalValue="Mauriès">Mauries</normalizedToken>
|
||
, Golovatch & Stoev, 1997, both known from artificial galleries (
|
||
<bibRefCitation id="88F458B651D2A563266DAF44E4EEAADF" author="Mauries, J-P" journalOrPublisher="Zoosystema" pageId="0" pageNumber="65" pagination="255 - 292" refId="B14" refString="Mauries, J-P, Golovatch, SI, Stoev, P, 1997. The millipedes of Albania: recent data, new taxa; systematical, nomenclatural and faunistical review (Myriapoda, Diplopoda). Zoosystema 19 (2-3): 255 - 292" title="The millipedes of Albania: recent data, new taxa; systematical, nomenclatural and faunistical review (Myriapoda, Diplopoda)." volume="19" year="1997">
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<normalizedToken id="D89893597053B5E827958E8F74263673" originalValue="Mauriès">Mauries</normalizedToken>
|
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et al. 1997
|
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</bibRefCitation>
|
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).
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A46075588663782CCE3AC29AF8A06C6A" pageId="0" pageNumber="65" type="distribution">
|
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<paragraph id="5BB6D2186FD4390CDC894B2A93F7E9A2" pageId="0" pageNumber="65">Distribution.</paragraph>
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<paragraph id="E0FDD15F7D2ECD2A2FCD6E424C3F5712" pageId="0" pageNumber="65">So far the new species is only known from a single pit in the Albanian part of the Prokletije Mountains.</paragraph>
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</subSubSection>
|
||
<subSubSection id="FCDB1FF37DFA43C7499CCDD9EE6227E8" pageId="0" pageNumber="65" type="additional material examined">
|
||
<paragraph id="62A14F9B9A626793BDE891DEB1B59A61" pageId="0" pageNumber="65">Additional material examined</paragraph>
|
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</subSubSection>
|
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</treatment>
|
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</document> |