treatments-xml/data/D3/3B/87/D33B87B4FF88FFAF57EF398B81AA7023.xml
2024-06-21 12:52:59 +02:00

194 lines
28 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="6632330FDF4D2CE7D87EC724E89DF8D9" ID-DOI="10.5281/zenodo.5359739" ID-ISSN="2345-7600" ID-Zenodo-Dep="5359739" ID-ZooBank="8F3A63EE-E132-4E18-8C58-C7034BFDA4A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630228788084" checkinUser="valdenar" docAuthor="Ahyong, Shane T." docDate="2018" docId="D33B87B4FF88FFAF57EF398B81AA7023" docLanguage="en" docName="66rbz538-571.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Austrotheres Ahyong 2018, new genus" docType="treatment" docUuid="A73A338E-0805-44BB-98DE-C9E230808F9E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="547" masterDocId="2F02FFCCFF80FFA6540A3F7684687116" masterDocTitle="Revision of Ostracotheres H. Milne Edwards, 1853 (Crustacea: Brachyura: Pinnotheridae)" masterLastPageNumber="571" masterPageNumber="538" pageNumber="546" updateTime="1699192615081" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="C8411265FB18488EB14F1D4A27F6BD98" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0AB126D910199F94C2990631E8C69FCA">
<mods:title id="C5C519C6A8B239A29F8350339FA39D17">Revision of Ostracotheres H. Milne Edwards, 1853 (Crustacea: Brachyura: Pinnotheridae)</mods:title>
</mods:titleInfo>
<mods:name id="85FF2F07D5C6E25E31E0BF4F139C64F5" type="personal">
<mods:role id="E5212285B6FF35DF4D3CF49C08A163C0">
<mods:roleTerm id="151FFE129EB491CA356B8AD93074CE39">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6253BD44BA2F85117759A6F0CDADA579">Ahyong, Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="DF5BD93FB669D69FB4468378B760CDD1">text</mods:typeOfResource>
<mods:relatedItem id="825F62A71082416B28BC8E8ED7617D25" type="host">
<mods:titleInfo id="78C3242CA07892B5C9E1899BD04A77B0">
<mods:title id="5B6F644E73F0ED7ECCFBBAC733A8CC51">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="3B3841733E1B945D4E44A843940703BE">
<mods:date id="32124F5178F8C15C956305A7C9AE0047">2018</mods:date>
<mods:detail id="43151148D386C17727267408F45B1E00" type="pubDate">
<mods:number id="D8872942AFAFBFDC6D5975821F3B7D82">2018-09-21</mods:number>
</mods:detail>
<mods:detail id="1C62971A41EC6B253ED07D66EFF3C599" type="volume">
<mods:number id="BD1E30B45BFFE501FB05563E74881286">66</mods:number>
</mods:detail>
<mods:extent id="6DA28F3D0B363B283DD72F4FF7543BB9" unit="page">
<mods:start id="94B10A43FA16FE8469578367BB86131F">538</mods:start>
<mods:end id="2D33FD95F46C6BDD64A7B2FEA7B7C36C">571</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CE2C8322040EFF89F85B9C082F9BFCFF">journal article</mods:classification>
<mods:identifier id="2E42FE455BDC3BDF21E4B20D40E43349" type="DOI">10.5281/zenodo.5359739</mods:identifier>
<mods:identifier id="3C29522FADC8E07090E7194392CFC6C8" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="1251616497EACE4C429F0FD97AF99A10" type="Zenodo-Dep">5359739</mods:identifier>
<mods:identifier id="A1A780828BF95BCCD89680AF55703A6F" type="ZooBank">8F3A63EE-E132-4E18-8C58-C7034BFDA4A0</mods:identifier>
</mods:mods>
<treatment id="D33B87B4FF88FFAF57EF398B81AA7023" LSID="urn:lsid:zoobank.org:act:A73A338E-0805-44BB-98DE-C9E230808F9E" httpUri="http://treatment.plazi.org/id/D33B87B4FF88FFAF57EF398B81AA7023" lastPageId="9" lastPageNumber="547" pageId="8" pageNumber="546">
<subSubSection id="13886529FF88FFAE57EF398B809A7603" box="[997,1266,1789,1813]" pageId="8" pageNumber="546" type="nomenclature">
<paragraph id="5B2D36A2FF88FFAE57EF398B809A7603" blockId="8.[997,1266,1789,1813]" box="[997,1266,1789,1813]" pageId="8" pageNumber="546">
<heading id="006581CEFF88FFAE57EF398B809A7603" bold="true" box="[997,1266,1789,1813]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="546" reason="2">
<emphasis id="69E6EAB0FF88FFAE57EF398B809A7603" bold="true" box="[997,1266,1789,1813]" pageId="8" pageNumber="546">
<taxonomicName id="9C924D21FF88FFAE57EF398B801A7603" ID-CoL="847PS" authority="Ahyong, 2018" authorityName="Ahyong" authorityYear="2018" box="[997,1138,1789,1813]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="546" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="69E6EAB0FF88FFAE57EF398B801A7603" bold="true" box="[997,1138,1789,1813]" italics="true" pageId="8" pageNumber="546">Austrotheres</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72D557CBFF88FFAE5071398B809A7603" box="[1147,1266,1789,1813]" pageId="8" pageNumber="546" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13886529FF88FFAF5727384B867673C3" lastPageId="9" lastPageNumber="547" pageId="8" pageNumber="546" type="diagnosis">
<paragraph id="5B2D36A2FF88FFAF5727384B867673C3" blockId="8.[812,1452,1853,2102]" lastBlockId="9.[136,775,189,725]" lastPageId="9" lastPageNumber="547" pageId="8" pageNumber="546">
<emphasis id="69E6EAB0FF88FFAE5727384B87C97643" bold="true" box="[813,929,1853,1877]" pageId="8" pageNumber="546">Diagnosis.</emphasis>
<emphasis id="69E6EAB0FF88FFAE57A2384B87907643" box="[936,1016,1853,1877]" italics="true" pageId="8" pageNumber="546">Female</emphasis>
: Carapace subcircular to sub-hexagonal, as long as or slightly longer than wide, posterior margin rounded, slightly projecting; sclerotised, hard; longitudinally vaulted, with weak to distinct epigastric ridge; front slightly produced anteriorly, subtruncate; anterolateral margins well defined, lateral surfaces near vertical; dorsal surface smooth or granulated, regions poorly to moderately defined, glabrous. Eyes visible in dorsal view. Antennal flagellum reaching beyond mid-height of eye but not dorsal margin of orbit. Maxilliped 3 ischiomerus completely fused, without trace of suture; palp 2-segmented; carpus shorter than propodus; propodus spatuliform, widened distally; exopod flagellum 2-segmented, distally setose. Chelipeds equal, stout, robust, dactylus and pollex with simple, pointed tips. Pereopods 25 symmetrical from left to right, slightly compressed, ovate in cross-section; dactyli with simple, spiniform apices, subequal or pereopod 5 dactylus longer. Pereopod 5 articulating with body at same level as pereopod 4. Abdomen with 6 free somites and telson; in adults, widest at somites 4 and 5, covering bases of walking legs. Gonopore simple, ovate, on sternite 5 near mesial end of sternite 5/6 suture.
<emphasis id="69E6EAB0FF89FFAF56C33D4B87687343" box="[713,768,573,597]" italics="true" pageId="9" pageNumber="547">Male</emphasis>
: similar to female, though smaller, carapace posterior margin flattened; G1 straight, flattened, distinctly narrowing at distal 1/4, with distinct step followed by lanceolate tip oriented along main axis; G2 without exopod.
</paragraph>
</subSubSection>
<subSubSection id="13886529FF89FFAF54823D8B85097223" pageId="9" pageNumber="547" type="type_taxon">
<paragraph id="5B2D36A2FF89FFAF54823D8B85097223" blockId="9.[136,774,765,821]" pageId="9" pageNumber="547">
<emphasis id="69E6EAB0FF89FFAF54823D8B85407203" bold="true" box="[136,296,765,789]" pageId="9" pageNumber="547">
<typeStatus id="84298800FF89FFAF54823D8B84AC7203" box="[136,196,765,789]" pageId="9" pageNumber="547">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="9C924D21FF89FFAF55393D8B862C7203" ID-CoL="84CB5" authority="Ahyong, 2018" authorityName="Ahyong" authorityYear="2018" box="[307,580,765,789]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="species" species="pregenzeri" status="sp. nov.">
<emphasis id="69E6EAB0FF89FFAF55393D8B862C7203" box="[307,580,765,789]" italics="true" pageId="9" pageNumber="547">Austrotheres pregenzeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72D557CBFF89FFAF565A3D8886B57203" box="[592,733,765,789]" pageId="9" pageNumber="547" rank="species">new species</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="13886529FF89FFAF54823C2B861E7523" pageId="9" pageNumber="547" type="etymology">
<paragraph id="5B2D36A2FF89FFAF54823C2B849472C3" blockId="9.[136,776,861,981]" pageId="9" pageNumber="547">
<emphasis id="69E6EAB0FF89FFAF54823C2B85797263" bold="true" box="[136,273,861,885]" pageId="9" pageNumber="547">Etymology.</emphasis>
An arbitrary combination of of the Latin,
<taxonomicName id="9C924D21FF89FFAF54833C0B848F7283" box="[137,231,893,917]" class="Bivalvia" family="Mytilidae" genus="Modiolaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="547" phylum="Mollusca" rank="species" species="australis">
<emphasis id="69E6EAB0FF89FFAF54833C0B848F7283" box="[137,231,893,917]" italics="true" pageId="9" pageNumber="547">australis</emphasis>
</taxonomicName>
, southern, and the suffix -
<emphasis id="69E6EAB0FF89FFAF56183C0B863C7283" box="[530,596,893,917]" italics="true" pageId="9" pageNumber="547">theres</emphasis>
, alluding to the southern distribution of the species of the genus. Gender: masculine.
</paragraph>
<paragraph id="5B2D36A2FF89FFAF54823C8B861E7523" blockId="9.[136,775,1021,1077]" pageId="9" pageNumber="547">
<emphasis id="69E6EAB0FF89FFAF54823C8B854C7503" bold="true" box="[136,292,1021,1045]" pageId="9" pageNumber="547">Composition.</emphasis>
<taxonomicName id="9C924D21FF89FFAF55243C8B876F7503" authority="(Baker, 1907)" authorityName="Ahyong" authorityYear="2018" baseAuthorityName="Baker" baseAuthorityYear="1907" box="[302,775,1021,1045]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="species" species="holothuriensis" status="comb. nov.">
<emphasis id="69E6EAB0FF89FFAF55243C8B86097503" box="[302,609,1021,1045]" italics="true" pageId="9" pageNumber="547">Austrotheres holothuriensis</emphasis>
(
<bibRefCitation id="3F034B53FF89FFAF56793C8B86967503" author="Baker WH" box="[627,766,1021,1045]" pageId="9" pageNumber="547" pagination="173 - 191" refId="ref28134" refString="Baker WH (1907) Notes on South Australian Decapoda Crustacea. Part V. Transactions, Proceedings and Report of the Royal Society of South Australia, 31: 173 - 191." type="journal article" year="1907">Baker, 1907</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="72D557CBFF89FFAF54823B68852D7523" box="[136,325,1053,1077]" pageId="9" pageNumber="547" rank="species">new combination</taxonomicNameLabel>
,
<taxonomicName id="9C924D21FF89FFAF55583B68858B7523" authority="Ahyong, 2018" authorityName="Ahyong" authorityYear="2018" box="[338,483,1054,1077]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="species" species="pregenzeri" status="sp. nov.">
<emphasis id="69E6EAB0FF89FFAF55583B68858B7523" box="[338,483,1054,1077]" italics="true" pageId="9" pageNumber="547">A. pregenzeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72D557CBFF89FFAF55E63B6886197523" box="[492,625,1053,1077]" pageId="9" pageNumber="547" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="13886529FF89FFAF54823B2B81AA7023" pageId="9" pageNumber="547" type="discussion">
<paragraph id="5B2D36A2FF89FFAF54823B2B876F77A3" blockId="9.[136,776,1117,1717]" pageId="9" pageNumber="547">
<emphasis id="69E6EAB0FF89FFAF54823B2B849F7563" bold="true" box="[136,247,1117,1141]" pageId="9" pageNumber="547">Remarks.</emphasis>
<taxonomicName id="9C924D21FF89FFAF54F43B2B85EE7563" authorityName="Ahyong" authorityYear="2018" box="[254,390,1117,1141]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF54F43B2B85EE7563" box="[254,390,1117,1141]" italics="true" pageId="9" pageNumber="547">Austrotheres</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="9C924D21FF89FFAF56593B2B86827563" box="[595,746,1117,1141]" class="Malacostraca" family="Pinnotheridae" genus="Ostracotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF56593B2B86827563" box="[595,746,1117,1141]" italics="true" pageId="9" pageNumber="547">Ostracotheres</emphasis>
</taxonomicName>
in overall carapace shape (rounded to ovate, as long as wide or slightly longer than wide, with well-defined anterolateral margins and near vertical lateral surfaces), the 2-segmental maxilliped 3 flagellum, and similar, leftright symmetrical walking legs, but differs in having a relatively hard (versus firm but poorly sclerotised) carapace with an epigastric ridge, glabrous (versus finely setose) carapace surface, and the form of the G1. Whereas the G1 apex in
<taxonomicName id="9C924D21FF89FFAF567A3A2B876F7463" box="[624,775,1373,1397]" class="Malacostraca" family="Pinnotheridae" genus="Ostracotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF567A3A2B876F7463" box="[624,775,1373,1397]" italics="true" pageId="9" pageNumber="547">Ostracotheres</emphasis>
</taxonomicName>
is blunt with a short, sharp, anterolaterally directed papilla (
<figureCitation id="C3A92A27FF89FFAF549A3AEB856B74A3" box="[144,259,1437,1461]" captionStart="Fig" captionStartId="3.[136,171,1782,1804]" captionTargetBox="[138,1475,386,1767]" captionTargetId="figure-8@3.[132,1481,372,1770]" captionTargetPageId="3" captionText="Fig. 2. Ostracotheres tridacnae (Rüppell, 1830). AI, female, cl 11.8 mm, cw 11.7 mm, Red Sea, MNHN-IU-2016-10949; JK, male, cl 8.0 mm, cw 7.6 mm, Red Sea, MNHN-IU-2016-10950; LO, male, cl 12.1 mm, cw 12.1 mm, Red Sea, NHMD-229248. A, J, dorsal habitus; B, right cheliped, anterior view; CF, right pereopods 25; G, right maxilliped 3; H, carapace, right lateral view; I, cephalothorax, anterior view; K, male abdomen; L, left G1, sternal view; M, left G1, tip, abdominal view; N, left G2, sternal view; O, right pereopod 5 dactylus. Scale: AF, H, J, K = 4.0 mm; G, I, L, N, O = 1.0 mm; M = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5359743" httpUri="https://zenodo.org/record/5359743/files/figure.png" pageId="9" pageNumber="547">Fig. 2L, M</figureCitation>
,
<figureCitation id="C3A92A27FF89FFAF55073AEB854074A3" box="[269,296,1437,1461]" captionStart="Fig" captionStartId="7.[136,171,1837,1859]" captionTargetBox="[139,1478,302,1821]" captionTargetId="figure-8@7.[135,1487,299,1831]" captionTargetPageId="7" captionText="Fig. 4. Ostracotheres cynthiae Nobili, 1906. AH, ovigerous female, cl 7.8 mm, cw 8.4 mm, Gulf of Suez, MNHN B10562; I, female lectotype, cl 6.8 mm, cw 8.0 mm, Djibouti, MNHN-IU-2014-11992; J, male, Gulf of Suez; KN, female, cl 3.8 mm, cw 3.7 mm, Red Sea, NHMW 10062; O, ovigerous female, cl 4.0 mm, cw 4.0 mm, Red Sea, NHMW 10063. A, dorsal habitus; B, left chelipid, anterior view; C, cephalothorax, anterior view; D, left maxilliped 3; EH, right pereopods 25; I, right maxilliped 3; J, left G1, sternal view; K, left pereopod 4 dactylus; L, left pereopod 5 dactylus; M, carapace, right lateral view; NO, carapace front, dorsal view. (DH, J, after Monod, 1938: fig. 25). Scale: A, B = 2.0 mm; C, D, I, J, = 0.5; EH, MO = 1.0 mm; KL = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5359747" httpUri="https://zenodo.org/record/5359747/files/figure.png" pageId="9" pageNumber="547">4J</figureCitation>
), the G
<quantity id="9C6A9B47FF89FFAF557F3AEB85F174A3" box="[373,409,1437,1461]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="547" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="9C924D21FF89FFAF55943AEB864D74A3" authorityName="Ahyong" authorityYear="2018" box="[414,549,1437,1461]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF55943AEB864D74A3" box="[414,549,1437,1461]" italics="true" pageId="9" pageNumber="547">Austrotheres</emphasis>
</taxonomicName>
is distally lanceolate, and approximately in line with the G1 axis (
<figureCitation id="C3A92A27FF89FFAF568D3ACB876A74C3" box="[647,770,1469,1493]" captionStart="Fig" captionStartId="11.[136,171,1835,1857]" captionTargetBox="[137,1478,332,1819]" captionTargetId="figure-8@11.[134,1482,328,1826]" captionTargetPageId="11" captionText="Fig. 6. Austrotheres pregenzeri new species. AJ: female holotype, cl 10.5 mm, cw 10.2 mm, Port Stephens, AM P72226. KQ: male paratype, cl 4.9 mm, cw 4.6 mm, Botany Bay, AM P63040. A, K, dorsal habitus; B, carapace, right lateral view; C, cephalothorax, anterior view; D, M, right maxilliped 3; E, right cheliped, anterior view; FI, right pereopods 25; J, female abdomen; L, left cheliped, anterior view; N, male abdomen; O, right G1, sternal view; P, right G1 tip, mesial view; Q, right G2, sternal view. Scale: AB, J = 2.0 mm; CI, K, L, N = 1.0 mm; M = 0.5 mm; OQ = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5359751" httpUri="https://zenodo.org/record/5359751/files/figure.png" pageId="9" pageNumber="547">Fig. 6O, P</figureCitation>
,
<figureCitation id="C3A92A27FF89FFAF54823AAB84A174E3" box="[136,201,1501,1525]" captionStart="Fig" captionStartId="13.[136,171,991,1013]" captionTargetBox="[137,1472,189,976]" captionTargetId="figure-8@13.[133,1479,184,982]" captionTargetPageId="13" captionText="Fig. 7. Austrotheres holothuriensis (Baker, 1907) new combination. AG, female lectotype, cl 8.3 mm, cw 7.2 mm, Gulf St. Vincent, SAM C1389; HN, male paralectotype, cl 6.2 mm, cw 6.5 mm, Gulf St. Vincent, SAM C1389; OQ, male, cl 5.5 mm, cw 5.2 mm, Shoreham, AM G5724; RS, female, cl 13.6 mm, cw 12.6 mm, Tasmania, AM P4047. A, H, carapace, dorsal view; B, I, left maxilliped 3; C, J, right cheliped, anterior view; D, right pereopod 2; E, K, right pereopod 3; F, L, right pereopod 4; G, M, right pereopod 5; N, male abdomen; O, right G1, sternal view; P, right G1 tip, mesial view; Q, right G2, sternal view; R, carapace, right lateral view; S, cephalothorax, anterior view. A, CH, JM = 2.0 mm; B, I, N = 1.0 mm; OQ = 0.5; R = 2.8 mm; S = 1.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5359753" httpUri="https://zenodo.org/record/5359753/files/figure.png" pageId="9" pageNumber="547">7O, P</figureCitation>
).
<taxonomicName id="9C924D21FF89FFAF54D73AAB850D74E3" authorityName="Ahyong" authorityYear="2018" box="[221,357,1501,1525]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF54D73AAB850D74E3" box="[221,357,1501,1525]" italics="true" pageId="9" pageNumber="547">Austrotheres</emphasis>
</taxonomicName>
also similar to the monotypic ascidian associate,
<taxonomicName id="9C924D21FF89FFAF54F53A8B86567703" authority="Campos, 1996" authorityName="Campos" authorityYear="1996" box="[255,574,1533,1557]" class="Malacostraca" family="Pinnotheridae" genus="Tunicotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF54F53A8B85F87703" box="[255,400,1533,1557]" italics="true" pageId="9" pageNumber="547">Tunicotheres</emphasis>
<bibRefCitation id="3F034B53FF89FFAF55913A8B86567703" author="Campos E" box="[411,574,1533,1557]" pageId="9" pageNumber="547" pagination="556 - 563" refId="ref28544" refString="Campos E (1996) Partial revision of pinnotherid crab genera with a two-segmented palp on the third maxilliped (Decapoda: Brachyura). Journal of Crustacean Biology, 16: 556 - 563." type="journal article" year="1996">Campos, 1996</bibRefCitation>
</taxonomicName>
(
<typeStatus id="84298800FF89FFAF565B3A8B86EA7703" box="[593,642,1533,1557]" pageId="9" pageNumber="547">type</typeStatus>
species:
<taxonomicName id="9C924D21FF89FFAF56FB3A8885E17723" authority="(Rathbun, 1918)" baseAuthorityName="Rathbun" baseAuthorityYear="1918" class="Malacostraca" family="Pinnotheridae" genus="Tunicotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="species" species="moseri">
<emphasis id="69E6EAB0FF89FFAF56FB3A8884B97723" italics="true" pageId="9" pageNumber="547">T. moseri</emphasis>
(
<bibRefCitation id="3F034B53FF89FFAF54EA396B85EA7723" author="Rathbun MJ" box="[224,386,1565,1589]" pageId="9" pageNumber="547" pagination="1 - 461" refId="ref31995" refString="Rathbun MJ (1918) The grapsoid crabs of America. United States National Museum Bulletin, 97: 1 - 461." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
)
</taxonomicName>
from the western Atlantic. Species of both genera have natatory setae on pereopods 34 and a somewhat polygonal carapace, but
<taxonomicName id="9C924D21FF89FFAF5606392B86FF7763" authorityName="Campos" authorityYear="1996" box="[524,663,1629,1653]" class="Malacostraca" family="Pinnotheridae" genus="Tunicotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF5606392B86FF7763" box="[524,663,1629,1653]" italics="true" pageId="9" pageNumber="547">Tunicotheres</emphasis>
</taxonomicName>
is readily separated by having pereopod 5 articulating above the level of pereopod 4, rather than at the same level as in
<taxonomicName id="9C924D21FF89FFAF567239EB876877A3" authorityName="Ahyong" authorityYear="2018" box="[632,768,1693,1717]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF567239EB876877A3" box="[632,768,1693,1717]" italics="true" pageId="9" pageNumber="547">Austrotheres</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5B2D36A2FF89FFAF548239AB81AA7023" blockId="9.[136,775,1757,2101]" lastBlockId="9.[835,1474,189,309]" pageId="9" pageNumber="547">
Unlike most other pinnotherids, in which females rarely or never leave the host and have weakly sclerotised exoskeletons, both sexes in
<taxonomicName id="9C924D21FF89FFAF55A8386B864F7623" authorityName="Ahyong" authorityYear="2018" box="[418,551,1821,1845]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF55A8386B864F7623" box="[418,551,1821,1845]" italics="true" pageId="9" pageNumber="547">Austrotheres</emphasis>
</taxonomicName>
have a comparatively hard carapace, natatory setae on the walking legs, and apparently frequently venture outside judging from the provenance of material reported below. Such morphology and behaviour in
<taxonomicName id="9C924D21FF89FFAF553638EB85AB76A3" authorityName="Ahyong" authorityYear="2018" box="[316,451,1949,1973]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF553638EB85AB76A3" box="[316,451,1949,1973]" italics="true" pageId="9" pageNumber="547">Austrotheres</emphasis>
</taxonomicName>
is consistent with reproductive strategies exhibited by a number of other pinnotherids, such as
<taxonomicName id="9C924D21FF89FFAF54A038AB859876E3" authority="Campos, 1989" authorityName="Campos" authorityYear="1989" box="[170,496,2013,2037]" class="Malacostraca" family="Pinnotheridae" genus="Tumidotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF54A038AB852A76E3" box="[170,322,2013,2037]" italics="true" pageId="9" pageNumber="547">Tumidotheres</emphasis>
<bibRefCitation id="3F034B53FF89FFAF554738AB859876E3" author="Campos E" box="[333,496,2013,2037]" pageId="9" pageNumber="547" pagination="672 - 679" refId="ref28464" refString="Campos E (1989) Tumidotheres, a new genus for Pinnotheres margarita Smith, 1869 and Pinnotheres maculatus Say, 1818 (Brachyura, Pinnotheridae). Journal of Crustacean Biology, 9: 672 - 679." type="journal article" year="1989">Campos, 1989</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="9C924D21FF89FFAF562438AB85697903" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Pinnotheridae" genus="Fabia" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="species" species="subquadrata">
<emphasis id="69E6EAB0FF89FFAF562438AB876F76E3" box="[558,775,2013,2037]" italics="true" pageId="9" pageNumber="547">Fabia subquadrata</emphasis>
<bibRefCitation id="3F034B53FF89FFAF5482388885697903" author="Dana JD" box="[136,257,2045,2069]" pageId="9" pageNumber="547" pagination="283 - 291" refId="ref28753" refString="Dana JD (1851) On the classification of the Crustacea Grapsoidea. American Journal of Science and Arts, (2) 12: 283 - 291." type="journal article" year="1851">Dana, 1851</bibRefCitation>
</taxonomicName>
from the eastern Pacific, which temporarily leave their host for copulatory swarming (
<bibRefCitation id="3F034B53FF89FFAF561E376B86DD7923" author="Campos E" box="[532,693,2077,2101]" pageId="9" pageNumber="547" pagination="672 - 679" refId="ref28464" refString="Campos E (1989) Tumidotheres, a new genus for Pinnotheres margarita Smith, 1869 and Pinnotheres maculatus Say, 1818 (Brachyura, Pinnotheridae). Journal of Crustacean Biology, 9: 672 - 679." type="journal article" year="1989">Campos, 1989</bibRefCitation>
,
<bibRefCitation id="3F034B53FF89FFAF56C8376B86947923" author="Campos E" box="[706,764,2077,2101]" pageId="9" pageNumber="547" pagination="311 - 329" refId="ref28628" refString="Campos E (2016) The Pinnotheridae of the northeastern Pacific (Alaska to Mexico): zoogeographical remarks and new bivalve hosts (Crustacea, Brachyura, Pinnotheridae). Zootaxa, 4170: 311 - 329." type="journal article" year="2016">2016</bibRefCitation>
). Species of
<taxonomicName id="9C924D21FF89FFAF57B33FCB802971C3" authorityName="Ahyong" authorityYear="2018" box="[953,1089,189,213]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF57B33FCB802971C3" box="[953,1089,189,213]" italics="true" pageId="9" pageNumber="547">Austrotheres</emphasis>
</taxonomicName>
may also have similar reproductive strategies, but too little is presently known of their biology to make reliable inferences about behaviour. Both species of
<taxonomicName id="9C924D21FF89FFAF57493E6B87A37023" authorityName="Ahyong" authorityYear="2018" box="[835,971,285,309]" class="Malacostraca" family="Pinnotheridae" genus="Austrotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="69E6EAB0FF89FFAF57493E6B87A37023" box="[835,971,285,309]" italics="true" pageId="9" pageNumber="547">Austrotheres</emphasis>
</taxonomicName>
occur in the southern half of
<collectingCountry id="23857632FF89FFAF510E3E6B810E7023" box="[1284,1382,285,309]" name="Australia" pageId="9" pageNumber="547">Australia</collectingCountry>
(
<figureCitation id="C3A92A27FF89FFAF517E3E6B81DC7023" box="[1396,1460,285,309]" captionStart="Fig" captionStartId="10.[113,148,859,881]" captionTargetBox="[114,753,190,843]" captionTargetId="figure-705@10.[111,755,186,851]" captionTargetPageId="10" captionText="Fig. 5. Distribution of species of Austrotheres new genus. (■) A. holothuriensis (Baker, 1907) new combination; (●) A. pregenzeri new species." figureDoi="http://doi.org/10.5281/zenodo.5359749" httpUri="https://zenodo.org/record/5359749/files/figure.png" pageId="9" pageNumber="547">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>