treatments-xml/data/05/23/87/052387C7FF8A927CFF12FD2C1114452E.xml

<document id="5DF3DF6E2A776949EBA6B4081A17A748" ID-CLB-Dataset="300924" ID-DOI="10.11646/zootaxa.5481.4.2" ID-GBIF-Dataset="1cf8602e-9bd3-4672-97c1-5cd45c4cca29" ID-ISSN="1175-5326" ID-Zenodo-Dep="12780207" ID-ZooBank="998A6EAC-8DF0-4F3C-A636-DD1D3649CCB3" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1721351591594" checkinUser="plazi" docAuthor="Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Helmick, Ericka E. &amp; Bahder, Brian W." docDate="2024" docId="052387C7FF8A927CFF12FD2C1114452E" docLanguage="en" docName="zootaxa.5481.4.2.pdf" docOrigin="Zootaxa 5481 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Myxia villaloboi Bahder &amp; Bartlett 2024, sp. n." docType="treatment" docVersion="4" lastPageNumber="451" masterDocId="F91AFFBFFF8F9277FF85FFD51432474F" masterDocTitle="A new species of planthopper in the genus Myxia (Hemiptera: Auchenorrhyncha Fulgoromorpha: Delphacoidea: Cixiidae) from the cloud forest in Costa Rica" masterLastPageNumber="452" masterPageNumber="440" pageNumber="445" updateTime="1723061625864" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="39DDA95CDAC29510AD0CB0D3FBBBCF0A">A new species of planthopper in the genus Myxia (Hemiptera: Auchenorrhyncha Fulgoromorpha: Delphacoidea: Cixiidae) from the cloud forest in Costa Rica</mods:title>
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<mods:namePart id="BC255383BCAC2B8D06100A9DCDA8CE9F">Barrantes Barrantes, Edwin A.</mods:namePart>
<mods:affiliation id="95C9240CED1F2C1B32D56E8E48D8FB20">Universidad of Costa Rica - Sede San Ramón, Departmento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica</mods:affiliation>
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<mods:namePart id="327EB218156F7A40C7B6947E09A110F0">Zumbado Echavarria, Marco A.</mods:namePart>
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<mods:namePart id="1D4DE5279735468899028B571A5E88B9">Bartlett, Charles R.</mods:namePart>
<mods:affiliation id="C56127AB82FB689816D6D9457195EB3B">University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA</mods:affiliation>
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<mods:namePart id="2D327366DD7C88DD3F2B937B9747D167">Helmick, Ericka E.</mods:namePart>
<mods:affiliation id="3BC521AC9D285FA31B4FB838A7865931">University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA</mods:affiliation>
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<mods:namePart id="9A2060C12D18B3BBD114FE1FEBF93DCF">Bahder, Brian W.</mods:namePart>
<mods:affiliation id="5C79C637DAC0B18DC12F6EB418665A34">University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA</mods:affiliation>
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<subSubSection id="C590655AFF8A9272FF12FD2C16A0445C" box="[151,658,761,788]" pageId="5" pageNumber="445" type="nomenclature">
<paragraph id="8D3536D1FF8A9272FF12FD2C16A0445C" blockId="5.[151,658,761,824]" box="[151,658,761,788]" pageId="5" pageNumber="445">
<heading id="D67D81BDFF8A9272FF12FD2C16A0445C" bold="true" box="[151,658,761,788]" fontSize="11" level="1" pageId="5" pageNumber="445" reason="1">
<taxonomicName id="4A8A4D52FF8A9272FF12FD2C167A445C" authority="Bahder &amp; Bartlett" authorityName="Bahder &amp; Bartlett" authorityYear="2024" box="[151,584,761,788]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="445" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. nov.">Myxia villaloboi Bahder &amp; Bartlett</taxonomicName>
<taxonomicNameLabel id="A4CD57B8FF8A9272FDCAFD2C16A0445C" box="[591,658,761,787]" pageId="5" pageNumber="445" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="8D3536D1FF8A9272FF12FCC8151D4478" blockId="5.[151,658,761,824]" box="[151,303,797,824]" pageId="5" pageNumber="445">
(
<figureCitation id="15B12A54FF8A9272FF1AFCC815154478" box="[159,295,797,824]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStartId-0="6.[152,255,1236,1260]" captionStartId-1="6.[152,255,1862,1886]" captionStartId-2="7.[152,255,1817,1841]" captionStartId-3="8.[152,255,1924,1948]" captionStartId-4="9.[152,255,884,908]" captionTargetBox-0="[151,1436,181,1213]" captionTargetBox-1="[151,1429,1291,1832]" captionTargetBox-2="[151,1425,921,1785]" captionTargetBox-3="[151,1436,993,1898]" captionTargetBox-4="[151,1435,181,856]" captionTargetId-0="figure-17@6.[151,1436,181,1213]" captionTargetId-1="figure-42@6.[151,1435,1285,1838]" captionTargetId-2="figure-180@7.[151,1436,921,1794]" captionTargetId-3="figure-370@8.[151,1436,993,1901]" captionTargetId-4="figure-24@9.[151,1435,181,861]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="FIGURE 3. Adult male habitus of Myxia villaloboi sp. n.; A) lateral view and B) dorsal view." captionText-1="FIGURE 4. Adult male Myxia villaloboi sp. n.; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm." captionText-2="FIGURE 5. Adult male Myxia villaloboi sp. n. forewing venation; wing venation ter54minology following Bourgoin et al. (2015); black text = vein, italic text = crossvein, green text = cell." captionText-3="FIGURE 6. Adult male Myxia villaloboi sp. n. genitalia; A) lateral view, B) ventral view and C) dorsal view." captionText-4="FIGURE 7. Aedeagus of Myxia villaloboi sp. n.; A) left lateral view, B) right later view, C) ventral view, and D) dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.12780213" figureDoi-1="http://doi.org/10.5281/zenodo.12780215" figureDoi-2="http://doi.org/10.5281/zenodo.12780217" figureDoi-3="http://doi.org/10.5281/zenodo.12780219" figureDoi-4="http://doi.org/10.5281/zenodo.12780221" httpUri-0="https://zenodo.org/record/12780213/files/figure.png" httpUri-1="https://zenodo.org/record/12780215/files/figure.png" httpUri-2="https://zenodo.org/record/12780217/files/figure.png" httpUri-3="https://zenodo.org/record/12780219/files/figure.png" httpUri-4="https://zenodo.org/record/12780221/files/figure.png" pageId="5" pageNumber="445">Figures 3–7</figureCitation>
)
</paragraph>
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<subSubSection id="C590655AFF8A9272FF12FCB017404430" box="[151,882,869,895]" pageId="5" pageNumber="445" type="materials_examined">
<paragraph id="8D3536D1FF8A9272FF12FCB017404430" blockId="5.[151,1437,869,1976]" box="[151,882,869,895]" pageId="5" pageNumber="445">
<emphasis id="BFFEEAC3FF8A9272FF12FCB015044430" bold="true" box="[151,310,869,895]" pageId="5" pageNumber="445">Type locality.</emphasis>
Hotel Villa Blanca, 
<collectingRegion id="4F4EF833FF8A9272FD9CFCB016D64430" box="[537,740,869,895]" country="Costa Rica" name="Alajuela" pageId="5" pageNumber="445">Alajuela Province</collectingRegion>
, 
<collectingCountry id="F59D7641FF8A9272FD75FCB0175C4430" box="[752,878,869,895]" name="Costa Rica" pageId="5" pageNumber="445">Costa Rica</collectingCountry>
.
</paragraph>
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<subSubSection id="C590655AFF8A9272FF42FC5C10774334" pageId="5" pageNumber="445" type="diagnosis">
<paragraph id="8D3536D1FF8A9272FF42FC5C10774334" blockId="5.[151,1437,869,1976]" pageId="5" pageNumber="445">
<emphasis id="BFFEEAC3FF8A9272FF42FC5C157044EC" bold="true" box="[199,322,905,931]" pageId="5" pageNumber="445">Diagnosis.</emphasis>
Moderate-sized (~ 
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) pale species (pale green 
<emphasis id="BFFEEAC3FF8A9272FC1EFC5F17DA44EC" box="[923,1000,906,931]" italics="true" pageId="5" pageNumber="445">in vivo</emphasis>
, light yellow preserved), with colored patch above frontoclypeal suture bordered with white. Head in dorsal view with transverse carinae on vertex formed by joined subapical carinae before apex of eyes, with large depressed areolet between subapical transverse carina and head apex. Male terminalia with pygofer bearing a large subtriangular medioventral process, taller than width at base. Aedeagal shaft, bearing one subapical process (otherwise devoid of processes), endosoma with 2 elongate subapical processes (ventral process about twice as long as dorsal process). Anal tube short and stout in lateral view with triangular ventral projection near midlength, apex curved ventrad and rounded.
</paragraph>
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<subSubSection id="C590655AFF8A927EFF42FB50172E41AA" lastPageId="9" lastPageNumber="449" pageId="5" pageNumber="445" type="description">
<paragraph id="8D3536D1FF8A9272FF42FB5016EC4244" blockId="5.[151,1437,869,1976]" pageId="5" pageNumber="445">
<emphasis id="BFFEEAC3FF8A9272FF42FB50156B43D0" bold="true" box="[199,345,1157,1183]" pageId="5" pageNumber="445">Description.</emphasis>
<emphasis id="BFFEEAC3FF8A9272FEE6FB50159643D0" box="[355,420,1157,1183]" italics="true" pageId="5" pageNumber="445">Color</emphasis>
. General body color in males pale green 
<emphasis id="BFFEEAC3FF8A9272FC0AFB5317EC43D0" box="[911,990,1158,1183]" italics="true" pageId="5" pageNumber="445">in vivo</emphasis>
, yellow after storage in ethanol, paler ventrally (
<figureCitation id="15B12A54FF8A9272FE8CFB7C157F438C" box="[265,333,1193,1219]" captionStart="FIGURE 3" captionStartId="6.[152,255,1236,1260]" captionTargetBox="[151,1436,181,1213]" captionTargetId="figure-17@6.[151,1436,181,1213]" captionTargetPageId="6" captionText="FIGURE 3. Adult male habitus of Myxia villaloboi sp. n.; A) lateral view and B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12780213" httpUri="https://zenodo.org/record/12780213/files/figure.png" pageId="5" pageNumber="445">Fig. 3</figureCitation>
). Frons pale with narrow orange (in preserved specimens) patch at fastigium and large, bell-shaped orange patch from frontoclypeal suture approximately to frons midpoint surrounded at top left and right sides by white; clypeus white. Wings transparent, unmarked.
</paragraph>
<paragraph id="8D3536D1FF8A9272FF42FAC3117641D8" blockId="5.[151,1437,869,1976]" pageId="5" pageNumber="445">
<emphasis id="BFFEEAC3FF8A9272FF42FAC31504427F" box="[199,310,1302,1328]" italics="true" pageId="5" pageNumber="445">Structure.</emphasis>
Body length males (
<emphasis id="BFFEEAC3FF8A9272FDA8FAC316094260" box="[557,571,1302,1327]" italics="true" pageId="5" pageNumber="445">n</emphasis>
= 3): 
<quantity id="4A729B34FF8A9272FD0FFAC01703427F" box="[650,817,1301,1328]" metricMagnitude="-3" metricUnit="m" metricValue="6.035" metricValueMax="6.05" metricValueMin="6.02" pageId="5" pageNumber="445" unit="mm" value="6.035" valueMax="6.05" valueMin="6.02">6.02–6.05 mm</quantity>
with wings; 
<quantity id="4A729B34FF8A9272FC48FAC01046427F" box="[973,1140,1301,1328]" metricMagnitude="-3" metricUnit="m" metricValue="4.234999999999999" metricValueMax="4.25" metricValueMin="4.22" pageId="5" pageNumber="445" unit="mm" value="4.234999999999999" valueMax="4.25" valueMin="4.22">4.22–4.25 mm</quantity>
without wings (
<tableCitation id="C008036AFF8A9272FAB6FAC011BC4260" box="[1331,1422,1301,1327]" captionStart="TABLE 3" captionStartId="7.[152,244,151,175]" captionTargetPageId="7" captionText="TABLE 3. Biometric data for Myxia villaloboi sp. n. (in mm)." httpUri="http://table.plazi.org/id/D9F56659FF889270FF1DFF42170547FF" pageId="5" pageNumber="445" tableUuid="D9F56659FF889270FF1DFF42170547FF">Table 3</tableCitation>
). Head. Vertex in dorsal view widest posteriorly, narrowing to near anterior margin of eyes; subapical carina meeting transversely on vertex (at narrowest part of vertex) near anterior portion of eyes resulting in distinct transverse suture, ahead of which the disc is concave forming a broad concave areolet within lateral carinae of vertex between subapical transverse carina and carina at fastigium; anterior margin of head sinuate with acute midpoint, posterior margin angulately incised with small, acute indentation at midpoint (
<figureCitation id="15B12A54FF8A9272FC01FA1C17EE42AC" box="[900,988,1481,1507]" captionStart="FIGURE 4" captionStartId="6.[152,255,1862,1886]" captionTargetBox="[151,1429,1291,1832]" captionTargetId="figure-42@6.[151,1435,1285,1838]" captionTargetPageId="6" captionText="FIGURE 4. Adult male Myxia villaloboi sp. n.; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12780215" httpUri="https://zenodo.org/record/12780215/files/figure.png" pageId="5" pageNumber="445">Fig. 4A</figureCitation>
). Head in lateral view generally rounded, slightly angulate at fastigium, projecting slightly beyond eyes, lateral ocelli conspicuous at ventroanterior margin of eyes, anterior and dorsal to antennae. Antennae short with scape ringlike, pedicle ovoid, situated approximately at midpoint between eye and frontoclypeal suture (
<figureCitation id="15B12A54FF8A9272FD61F9E0170D4100" box="[740,831,1589,1615]" captionStart="FIGURE 4" captionStartId="6.[152,255,1862,1886]" captionTargetBox="[151,1429,1291,1832]" captionTargetId="figure-42@6.[151,1435,1285,1838]" captionTargetPageId="6" captionText="FIGURE 4. Adult male Myxia villaloboi sp. n.; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12780215" httpUri="https://zenodo.org/record/12780215/files/figure.png" pageId="5" pageNumber="445">Fig. 4B</figureCitation>
). In frontal view, frons narrowest dorsally, widest at frontoclypeal suture, dorsal margin concave, lateral margins arched laterad from dorsal margin, median carina present. Median ocellus conspicuous just above frontoclypeal suture (
<figureCitation id="15B12A54FF8A9272FC21F9A817CF41D8" box="[932,1021,1661,1687]" captionStart="FIGURE 4" captionStartId="6.[152,255,1862,1886]" captionTargetBox="[151,1429,1291,1832]" captionTargetId="figure-42@6.[151,1435,1285,1838]" captionTargetPageId="6" captionText="FIGURE 4. Adult male Myxia villaloboi sp. n.; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12780215" httpUri="https://zenodo.org/record/12780215/files/figure.png" pageId="5" pageNumber="445">Fig. 4C</figureCitation>
). Frontoclypeal suture linear.
</paragraph>
<paragraph id="8D3536D1FF8A9272FF42F974104A40F8" blockId="5.[151,1437,869,1976]" pageId="5" pageNumber="445">
Thorax. Pronotum in dorsal view (
<figureCitation id="15B12A54FF8A9272FDCEF97416AF41F4" box="[587,669,1697,1723]" captionStart="FIGURE 4" captionStartId="6.[152,255,1862,1886]" captionTargetBox="[151,1429,1291,1832]" captionTargetId="figure-42@6.[151,1435,1285,1838]" captionTargetPageId="6" captionText="FIGURE 4. Adult male Myxia villaloboi sp. n.; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12780215" httpUri="https://zenodo.org/record/12780215/files/figure.png" pageId="5" pageNumber="445">Fig 4A</figureCitation>
) very short medially, expanding laterally, anterior margin hidden by posterior margin of head, posterior margin broadly concave; lateral carinae arising near midpoint (
<figureCitation id="15B12A54FF8A9272FB62F91011704190" box="[1255,1346,1733,1759]" captionStart="FIGURE 4" captionStartId="6.[152,255,1862,1886]" captionTargetBox="[151,1429,1291,1832]" captionTargetId="figure-42@6.[151,1435,1285,1838]" captionTargetPageId="6" captionText="FIGURE 4. Adult male Myxia villaloboi sp. n.; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12780215" httpUri="https://zenodo.org/record/12780215/files/figure.png" pageId="5" pageNumber="445">Fig. 4A</figureCitation>
); lateral margin in dorsal view formed by lateral carinae extending between eye and tegula; in lateral view, dorsal portion of pronotum declining anteriorly; paradiscal region broad, extending ventrad about to level of antennae, anterior and posterior margins sinuate, carinae near anterior margin (by extension of lateral carinae of disc), ventral margin sinuate extending to sharp point at ventrocaudal margin, lateral carinae terminating at ventral margin (
<figureCitation id="15B12A54FF8A9272FAB1F88011BD4020" box="[1332,1423,1877,1903]" captionStart="FIGURE 4" captionStartId="6.[152,255,1862,1886]" captionTargetBox="[151,1429,1291,1832]" captionTargetId="figure-42@6.[151,1435,1285,1838]" captionTargetPageId="6" captionText="FIGURE 4. Adult male Myxia villaloboi sp. n.; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12780215" httpUri="https://zenodo.org/record/12780215/files/figure.png" pageId="5" pageNumber="445">Fig. 4B</figureCitation>
). Mesonotum in dorsal view about as long at midpoint as wide at widest point (near posterior margin of tegulae); tricarinate, carinae subparallel and complete (or nearly so) to posterior margin (
<figureCitation id="15B12A54FF8A9272FB94F848105940F8" box="[1041,1131,1949,1975]" captionStart="FIGURE 4" captionStartId="6.[152,255,1862,1886]" captionTargetBox="[151,1429,1291,1832]" captionTargetId="figure-42@6.[151,1435,1285,1838]" captionTargetPageId="6" captionText="FIGURE 4. Adult male Myxia villaloboi sp. n.; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12780215" httpUri="https://zenodo.org/record/12780215/files/figure.png" pageId="5" pageNumber="445">Fig. 4A</figureCitation>
).
</paragraph>
<caption id="D9F56659FF899271FF1DFB01106543A2" ID-DOI="http://doi.org/10.5281/zenodo.12780213" ID-Zenodo-Dep="12780213" box="[152,1111,1236,1261]" httpUri="https://zenodo.org/record/12780213/files/figure.png" pageId="6" pageNumber="446" startId="6.[152,255,1236,1260]" targetBox="[151,1436,181,1213]" targetPageId="6" targetType="figure">
<paragraph id="8D3536D1FF899271FF1DFB01106543A2" blockId="6.[152,1111,1236,1261]" box="[152,1111,1236,1261]" pageId="6" pageNumber="446">
<emphasis id="BFFEEAC3FF899271FF1DFB01152A43A2" bold="true" box="[152,280,1236,1261]" pageId="6" pageNumber="446">FIGURE 3.</emphasis>
Adult male habitus of 
<taxonomicName id="4A8A4D52FF899271FD84FB00169A43A2" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[513,680,1237,1261]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. nov.">
<emphasis id="BFFEEAC3FF899271FD84FB00169A43A2" box="[513,680,1237,1261]" italics="true" pageId="6" pageNumber="446">Myxia villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF899271FD2BFB0016D543A2" bold="true" box="[686,743,1237,1261]" pageId="6" pageNumber="446">
<taxonomicNameLabel id="A4CD57B8FF899271FD2BFB0016D543A2" box="[686,743,1237,1261]" pageId="6" pageNumber="446" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; A) lateral view and B) dorsal view.
</paragraph>
</caption>
<caption id="D9F56659FF899271FF1DF89317B040CE" ID-DOI="http://doi.org/10.5281/zenodo.12780215" ID-Zenodo-Dep="12780215" httpUri="https://zenodo.org/record/12780215/files/figure.png" pageId="6" pageNumber="446" startId="6.[152,255,1862,1886]" targetBox="[151,1429,1291,1832]" targetPageId="6" targetType="figure">
<paragraph id="8D3536D1FF899271FF1DF89317B040CE" blockId="6.[151,1436,1862,1922]" pageId="6" pageNumber="446">
<emphasis id="BFFEEAC3FF899271FF1DF89315284011" bold="true" box="[152,282,1862,1886]" pageId="6" pageNumber="446">FIGURE 4.</emphasis>
Adult male 
<taxonomicName id="4A8A4D52FF899271FE19F89216714011" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[412,579,1862,1886]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="446" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. nov.">
<emphasis id="BFFEEAC3FF899271FE19F89216714011" box="[412,579,1862,1886]" italics="true" pageId="6" pageNumber="446">Myxia villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF899271FDCEF89316B74011" bold="true" box="[587,645,1862,1886]" pageId="6" pageNumber="446">
<taxonomicNameLabel id="A4CD57B8FF899271FDCEF89316B74011" box="[587,645,1862,1886]" pageId="6" pageNumber="446" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm.
</paragraph>
</caption>
<caption id="D9F56659FF889270FF1DFF42170547FF" ID-Table-UUID="D9F56659FF889270FF1DFF42170547FF" box="[152,823,151,176]" httpUri="http://table.plazi.org/id/D9F56659FF889270FF1DFF42170547FF" pageId="7" pageNumber="447" startId="7.[152,244,151,175]" targetBox="[163,1387,192,860]" targetIsTable="true" targetPageId="7" targetType="table">
<paragraph id="8D3536D1FF889270FF1DFF42170547FF" blockId="7.[152,823,151,176]" box="[152,823,151,176]" pageId="7" pageNumber="447">
<emphasis id="BFFEEAC3FF889270FF1DFF42153D47E0" bold="true" box="[152,271,151,175]" pageId="7" pageNumber="447">TABLE 3.</emphasis>
Biometric data for 
<taxonomicName id="4A8A4D52FF889270FE65FF4D16BD47E0" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[480,655,151,175]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. nov.">
<emphasis id="BFFEEAC3FF889270FE65FF4D16BD47E0" box="[480,655,151,175]" italics="true" pageId="7" pageNumber="447">Myxia villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF889270FD13FF4D16E047FF" bold="true" box="[662,722,152,176]" pageId="7" pageNumber="447">
<taxonomicNameLabel id="A4CD57B8FF889270FD13FF4D16E047FF" box="[662,722,152,176]" pageId="7" pageNumber="447" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(in mm).
</paragraph>
</caption>
<paragraph id="8D3536D1FF889270FF26FF1511634413" pageId="7" pageNumber="447">
<table id="FF8AC471FF886D88FF26FF1511594413" box="[163,1387,192,860]" gridcols="5" gridrows="17" pageId="7" pageNumber="447">
<tr id="33BA3493FF886D88FF26FF1511594797" box="[163,1387,192,216]" gridrow="0" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FF1515894797" box="[163,443,192,216]" gridcol="0" gridrow="0" pageId="7" pageNumber="447">Character</th>
<th id="706B5DEFFF886D88FDF1FF1517F64797" box="[628,964,192,216]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="7" pageNumber="447">
Male (
<emphasis id="BFFEEAC3FF889270FC97FF1417024797" box="[786,816,193,216]" italics="true" pageId="7" pageNumber="447">n=</emphasis>
2)
</th>
<th id="706B5DEFFF886D88FBABFF1511594797" box="[1070,1387,192,216]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="7" pageNumber="447">
Female (
<emphasis id="BFFEEAC3FF889270FB5CFF1410D44797" box="[1241,1254,193,216]" italics="true" pageId="7" pageNumber="447">n</emphasis>
=3)
</th>
</tr>
<tr id="33BA3493FF886D88FF26FF3D1159464E" box="[163,1387,232,257]" gridrow="1" pageId="7" pageNumber="447" rowspan-0="1">
<td id="706B5DEFFF886D88FDF1FF3D16F2464E" box="[628,704,232,257]" gridcol="1" gridrow="1" pageId="7" pageNumber="447">Range</td>
<td id="706B5DEFFF886D88FCB2FF3D17F6464E" box="[823,964,232,257]" gridcol="2" gridrow="1" pageId="7" pageNumber="447">Average ± SE</td>
<td id="706B5DEFFF886D88FBABFF3D1049464E" box="[1070,1147,232,257]" gridcol="3" gridrow="1" pageId="7" pageNumber="447">Range</td>
<td id="706B5DEFFF886D88FB69FF3D1159464E" box="[1260,1387,232,257]" gridcol="4" gridrow="1" pageId="7" pageNumber="447">Average±SE</td>
</tr>
<tr id="33BA3493FF886D88FF26FEC511594667" box="[163,1387,272,296]" gridrow="2" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FEC515894667" box="[163,443,272,296]" gridcol="0" gridrow="2" pageId="7" pageNumber="447">Body length, with wings</th>
<td id="706B5DEFFF886D88FDF1FEC516F24667" box="[628,704,272,296]" gridcol="1" gridrow="2" pageId="7" pageNumber="447">5.4–5.6</td>
<td id="706B5DEFFF886D88FCB2FEC517F64667" box="[823,964,272,296]" gridcol="2" gridrow="2" pageId="7" pageNumber="447">5.5±0.1</td>
<td id="706B5DEFFF886D88FBABFEC510494667" box="[1070,1147,272,296]" gridcol="3" gridrow="2" pageId="7" pageNumber="447">6.7–6.9</td>
<td id="706B5DEFFF886D88FB69FEC511594667" box="[1260,1387,272,296]" gridcol="4" gridrow="2" pageId="7" pageNumber="447">6.8±0.1</td>
</tr>
<tr id="33BA3493FF886D88FF26FEED1159461E" box="[163,1387,312,337]" gridrow="3" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FEED1589461E" box="[163,443,312,337]" gridcol="0" gridrow="3" pageId="7" pageNumber="447">Body length, no wings</th>
<td id="706B5DEFFF886D88FDF1FEED16F2461E" box="[628,704,312,337]" gridcol="1" gridrow="3" pageId="7" pageNumber="447">3.4–3.6</td>
<td id="706B5DEFFF886D88FCB2FEED17F6461E" box="[823,964,312,337]" gridcol="2" gridrow="3" pageId="7" pageNumber="447">3.5±0.1</td>
<td id="706B5DEFFF886D88FBABFEED1049461E" box="[1070,1147,312,337]" gridcol="3" gridrow="3" pageId="7" pageNumber="447">4.6–4.8</td>
<td id="706B5DEFFF886D88FB69FEED1159461E" box="[1260,1387,312,337]" gridcol="4" gridrow="3" pageId="7" pageNumber="447">4.7±0.1</td>
</tr>
<tr id="33BA3493FF886D88FF26FEB411594636" box="[163,1387,353,377]" gridrow="4" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FEB415894636" box="[163,443,353,377]" gridcol="0" gridrow="4" pageId="7" pageNumber="447">Forewing length</th>
<td id="706B5DEFFF886D88FDF1FEB416F24636" box="[628,704,353,377]" gridcol="1" gridrow="4" pageId="7" pageNumber="447">5.7–5.8</td>
<td id="706B5DEFFF886D88FCB2FEB417F64636" box="[823,964,353,377]" gridcol="2" gridrow="4" pageId="7" pageNumber="447">5.8±0.1</td>
<td id="706B5DEFFF886D88FBABFEB410494636" box="[1070,1147,353,377]" gridcol="3" gridrow="4" pageId="7" pageNumber="447">6.9–7.1</td>
<td id="706B5DEFFF886D88FB69FEB411594636" box="[1260,1387,353,377]" gridcol="4" gridrow="4" pageId="7" pageNumber="447">7.0±0.1</td>
</tr>
<tr id="33BA3493FF886D88FF26FE5C115946EE" box="[163,1387,393,417]" gridrow="5" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FE5C158946EE" box="[163,443,393,417]" gridcol="0" gridrow="5" pageId="7" pageNumber="447">Vertex length</th>
<td id="706B5DEFFF886D88FDF1FE5C16F246EE" box="[628,704,393,417]" gridcol="1" gridrow="5" pageId="7" pageNumber="447">0.5–0.5</td>
<td id="706B5DEFFF886D88FCB2FE5C17F646EE" box="[823,964,393,417]" gridcol="2" gridrow="5" pageId="7" pageNumber="447">0.5±0.0</td>
<td id="706B5DEFFF886D88FBABFE5C104946EE" box="[1070,1147,393,417]" gridcol="3" gridrow="5" pageId="7" pageNumber="447">0.6–0.6</td>
<td id="706B5DEFFF886D88FB69FE5C115946EE" box="[1260,1387,393,417]" gridcol="4" gridrow="5" pageId="7" pageNumber="447">0.6±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FE6411594686" box="[163,1387,433,457]" gridrow="6" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FE6415894686" box="[163,443,433,457]" gridcol="0" gridrow="6" pageId="7" pageNumber="447">Vertex width, basal margin</th>
<td id="706B5DEFFF886D88FDF1FE6416F24686" box="[628,704,433,457]" gridcol="1" gridrow="6" pageId="7" pageNumber="447">0.5–0.5</td>
<td id="706B5DEFFF886D88FCB2FE6417F64686" box="[823,964,433,457]" gridcol="2" gridrow="6" pageId="7" pageNumber="447">0.5±0.0</td>
<td id="706B5DEFFF886D88FBABFE6410494686" box="[1070,1147,433,457]" gridcol="3" gridrow="6" pageId="7" pageNumber="447">0.6–0.6</td>
<td id="706B5DEFFF886D88FB69FE6411594686" box="[1260,1387,433,457]" gridcol="4" gridrow="6" pageId="7" pageNumber="447">0.6±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FE0C115946BD" box="[163,1387,473,498]" gridrow="7" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FE0C158946BD" box="[163,443,473,498]" gridcol="0" gridrow="7" pageId="7" pageNumber="447">Vertex width, distal margin</th>
<td id="706B5DEFFF886D88FDF1FE0C16F246BD" box="[628,704,473,498]" gridcol="1" gridrow="7" pageId="7" pageNumber="447">0.2–0.2</td>
<td id="706B5DEFFF886D88FCB2FE0C17F646BD" box="[823,964,473,498]" gridcol="2" gridrow="7" pageId="7" pageNumber="447">0.2±0.0</td>
<td id="706B5DEFFF886D88FBABFE0C104946BD" box="[1070,1147,473,498]" gridcol="3" gridrow="7" pageId="7" pageNumber="447">0.3–0.3</td>
<td id="706B5DEFFF886D88FB69FE0C115946BD" box="[1260,1387,473,498]" gridcol="4" gridrow="7" pageId="7" pageNumber="447">0.3±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FDD711594555" box="[163,1387,514,538]" gridrow="8" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FDD715894555" box="[163,443,514,538]" gridcol="0" gridrow="8" pageId="7" pageNumber="447">Pronotum length, midline</th>
<td id="706B5DEFFF886D88FDF1FDD716F24555" box="[628,704,514,538]" gridcol="1" gridrow="8" pageId="7" pageNumber="447">0.1–0.1</td>
<td id="706B5DEFFF886D88FCB2FDD717F64555" box="[823,964,514,538]" gridcol="2" gridrow="8" pageId="7" pageNumber="447">0.1±0.0</td>
<td id="706B5DEFFF886D88FBABFDD710494555" box="[1070,1147,514,538]" gridcol="3" gridrow="8" pageId="7" pageNumber="447">0.2–0.2</td>
<td id="706B5DEFFF886D88FB69FDD711594555" box="[1260,1387,514,538]" gridcol="4" gridrow="8" pageId="7" pageNumber="447">0.2±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FDFF1159450D" box="[163,1387,554,578]" gridrow="9" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FDFF1589450D" box="[163,443,554,578]" gridcol="0" gridrow="9" pageId="7" pageNumber="447">Mesonotum length, midline</th>
<td id="706B5DEFFF886D88FDF1FDFF16F2450D" box="[628,704,554,578]" gridcol="1" gridrow="9" pageId="7" pageNumber="447">1.1–1.1</td>
<td id="706B5DEFFF886D88FCB2FDFF17F6450D" box="[823,964,554,578]" gridcol="2" gridrow="9" pageId="7" pageNumber="447">1.1±0.0</td>
<td id="706B5DEFFF886D88FBABFDFF1049450D" box="[1070,1147,554,578]" gridcol="3" gridrow="9" pageId="7" pageNumber="447">1.3–1.3</td>
<td id="706B5DEFFF886D88FB69FDFF1159450D" box="[1260,1387,554,578]" gridcol="4" gridrow="9" pageId="7" pageNumber="447">1.3±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FD8711594525" box="[163,1387,594,618]" gridrow="10" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FD8715894525" box="[163,443,594,618]" gridcol="0" gridrow="10" pageId="7" pageNumber="447">Mesonotum width</th>
<td id="706B5DEFFF886D88FDF1FD8716F24525" box="[628,704,594,618]" gridcol="1" gridrow="10" pageId="7" pageNumber="447">1.1–1.1</td>
<td id="706B5DEFFF886D88FCB2FD8717F64525" box="[823,964,594,618]" gridcol="2" gridrow="10" pageId="7" pageNumber="447">1.1±0.0</td>
<td id="706B5DEFFF886D88FBABFD8710494525" box="[1070,1147,594,618]" gridcol="3" gridrow="10" pageId="7" pageNumber="447">1.3–1.3</td>
<td id="706B5DEFFF886D88FB69FD8711594525" box="[1260,1387,594,618]" gridcol="4" gridrow="10" pageId="7" pageNumber="447">1.3±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FDAF115945DC" box="[163,1387,634,659]" gridrow="11" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FDAF158945DC" box="[163,443,634,659]" gridcol="0" gridrow="11" pageId="7" pageNumber="447">Frons width, dorsal margin</th>
<td id="706B5DEFFF886D88FDF1FDAF16F245DC" box="[628,704,634,659]" gridcol="1" gridrow="11" pageId="7" pageNumber="447">0.2–0.2</td>
<td id="706B5DEFFF886D88FCB2FDAF17F645DC" box="[823,964,634,659]" gridcol="2" gridrow="11" pageId="7" pageNumber="447">0.2±0.0</td>
<td id="706B5DEFFF886D88FBABFDAF104945DC" box="[1070,1147,634,659]" gridcol="3" gridrow="11" pageId="7" pageNumber="447">0.3–0.3</td>
<td id="706B5DEFFF886D88FB69FDAF115945DC" box="[1260,1387,634,659]" gridcol="4" gridrow="11" pageId="7" pageNumber="447">0.3±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FD76115945F4" box="[163,1387,675,699]" gridrow="12" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FD76158945F4" box="[163,443,675,699]" gridcol="0" gridrow="12" pageId="7" pageNumber="447">Frons width, clypeal suture</th>
<td id="706B5DEFFF886D88FDF1FD7616F245F4" box="[628,704,675,699]" gridcol="1" gridrow="12" pageId="7" pageNumber="447">0.5–0.5</td>
<td id="706B5DEFFF886D88FCB2FD7617F645F4" box="[823,964,675,699]" gridcol="2" gridrow="12" pageId="7" pageNumber="447">0.5±0.0</td>
<td id="706B5DEFFF886D88FBABFD76104945F4" box="[1070,1147,675,699]" gridcol="3" gridrow="12" pageId="7" pageNumber="447">0.6–0.6</td>
<td id="706B5DEFFF886D88FB69FD76115945F4" box="[1260,1387,675,699]" gridcol="4" gridrow="12" pageId="7" pageNumber="447">0.6±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FD1E115945AC" box="[163,1387,715,739]" gridrow="13" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FD1E158945AC" box="[163,443,715,739]" gridcol="0" gridrow="13" pageId="7" pageNumber="447">Frons width, widest</th>
<td id="706B5DEFFF886D88FDF1FD1E16F245AC" box="[628,704,715,739]" gridcol="1" gridrow="13" pageId="7" pageNumber="447">0.5–0.5</td>
<td id="706B5DEFFF886D88FCB2FD1E17F645AC" box="[823,964,715,739]" gridcol="2" gridrow="13" pageId="7" pageNumber="447">0.5±0.0</td>
<td id="706B5DEFFF886D88FBABFD1E104945AC" box="[1070,1147,715,739]" gridcol="3" gridrow="13" pageId="7" pageNumber="447">0.6–0.6</td>
<td id="706B5DEFFF886D88FB69FD1E115945AC" box="[1260,1387,715,739]" gridcol="4" gridrow="13" pageId="7" pageNumber="447">0.6±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FD2611594444" box="[163,1387,755,779]" gridrow="14" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FD2615894444" box="[163,443,755,779]" gridcol="0" gridrow="14" pageId="7" pageNumber="447">Frons width, narrowest</th>
<td id="706B5DEFFF886D88FDF1FD2616F24444" box="[628,704,755,779]" gridcol="1" gridrow="14" pageId="7" pageNumber="447">0.2–0.2</td>
<td id="706B5DEFFF886D88FCB2FD2617F64444" box="[823,964,755,779]" gridcol="2" gridrow="14" pageId="7" pageNumber="447">0.2±0.0</td>
<td id="706B5DEFFF886D88FBABFD2610494444" box="[1070,1147,755,779]" gridcol="3" gridrow="14" pageId="7" pageNumber="447">0.3–0.3</td>
<td id="706B5DEFFF886D88FB69FD2611594444" box="[1260,1387,755,779]" gridcol="4" gridrow="14" pageId="7" pageNumber="447">0.3±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FCCE1159447C" box="[163,1387,795,819]" gridrow="15" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FCCE1589447C" box="[163,443,795,819]" gridcol="0" gridrow="15" pageId="7" pageNumber="447">Frons length, midline</th>
<td id="706B5DEFFF886D88FDF1FCCE16F2447C" box="[628,704,795,819]" gridcol="1" gridrow="15" pageId="7" pageNumber="447">0.5–0.5</td>
<td id="706B5DEFFF886D88FCB2FCCE17F6447C" box="[823,964,795,819]" gridcol="2" gridrow="15" pageId="7" pageNumber="447">0.5±0.0</td>
<td id="706B5DEFFF886D88FBABFCCE1049447C" box="[1070,1147,795,819]" gridcol="3" gridrow="15" pageId="7" pageNumber="447">0.6–0.6</td>
<td id="706B5DEFFF886D88FB69FCCE1159447C" box="[1260,1387,795,819]" gridcol="4" gridrow="15" pageId="7" pageNumber="447">0.6±0.0</td>
</tr>
<tr id="33BA3493FF886D88FF26FC9111594413" box="[163,1387,836,860]" gridrow="16" pageId="7" pageNumber="447">
<th id="706B5DEFFF886D88FF26FC9115894413" box="[163,443,836,860]" gridcol="0" gridrow="16" pageId="7" pageNumber="447">Clypeus length</th>
<td id="706B5DEFFF886D88FDF1FC9116F24413" box="[628,704,836,860]" gridcol="1" gridrow="16" pageId="7" pageNumber="447">0.4–0.4</td>
<td id="706B5DEFFF886D88FCB2FC9117F64413" box="[823,964,836,860]" gridcol="2" gridrow="16" pageId="7" pageNumber="447">0.4±0.0</td>
<td id="706B5DEFFF886D88FBABFC9110494413" box="[1070,1147,836,860]" gridcol="3" gridrow="16" pageId="7" pageNumber="447">0.5–0.5</td>
<td id="706B5DEFFF886D88FB69FC9111594413" box="[1260,1387,836,860]" gridcol="4" gridrow="16" pageId="7" pageNumber="447">0.5±0.0</td>
</tr>
</table>
</paragraph>
<caption id="D9F56659FF889270FF1DF8CC1702401A" ID-DOI="http://doi.org/10.5281/zenodo.12780217" ID-Zenodo-Dep="12780217" httpUri="https://zenodo.org/record/12780217/files/figure.png" pageId="7" pageNumber="447" startId="7.[152,255,1817,1841]" targetBox="[151,1425,921,1785]" targetPageId="7" targetType="figure">
<paragraph id="8D3536D1FF889270FF1DF8CC1702401A" blockId="7.[151,1437,1817,1878]" pageId="7" pageNumber="447">
<emphasis id="BFFEEAC3FF889270FF1DF8CC1528407D" bold="true" box="[152,282,1817,1842]" pageId="7" pageNumber="447">FIGURE 5.</emphasis>
Adult male 
<taxonomicName id="4A8A4D52FF889270FE1BF8CF1675407E" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[414,583,1817,1841]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="447" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. nov.">
<emphasis id="BFFEEAC3FF889270FE1BF8CF1675407E" box="[414,583,1817,1841]" italics="true" pageId="7" pageNumber="447">Myxia villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF889270FDCAF8CF16B8407D" bold="true" box="[591,650,1818,1842]" pageId="7" pageNumber="447">
<taxonomicNameLabel id="A4CD57B8FF889270FDCAF8CF16B8407D" box="[591,650,1818,1842]" pageId="7" pageNumber="447" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
forewing venation; wing venation ter54minology following 
<bibRefCitation id="E91B4B20FF889270FA85F8CC14EF401A" author="Bourgoin, T. &amp; Wang, R. R. &amp; Asche, M. &amp; Hoch, H. &amp; Soulier-Perkins, A. &amp; Stroinski, A. &amp; Yap, S. &amp; Szwedo, J." pageId="7" pageNumber="447" pagination="63 - 77" refId="ref5564" refString="Bourgoin, T., Wang, R. R., Asche, M., Hoch, H., Soulier-Perkins, A., Stroinski, A., Yap, S. &amp; Szwedo, J. (2015) From micropterism to hyperpterism: recognition strategy and standardized homology-driven terminology of the forewing venation patterns in planthoppers (Hemiptera: Fulgoromorpha). Zoomorphology, 134 (1), 63 - 77. https: // doi. org / 10.1007 / s 00435 - 014 - 0243 - 6" type="journal article" year="2015">
Bourgoin 
<emphasis id="BFFEEAC3FF889270FAEDF8CF11AF407E" box="[1384,1437,1817,1841]" italics="true" pageId="7" pageNumber="447">et al.</emphasis>
(2015)
</bibRefCitation>
; black text = vein, italic text = crossvein, green text = cell.
</paragraph>
</caption>
<paragraph id="8D3536D1FF87927FFF42FF42167F46E2" blockId="8.[151,1437,151,969]" pageId="8" pageNumber="448">
Forewings elongate with leading and trailing sides approximately parallel-sided, apex rounded; blade transparent (
<figureCitation id="15B12A54FF87927FFEA0FF6E1559479A" box="[293,363,186,213]" captionStart="FIGURE 5" captionStartId="7.[152,255,1817,1841]" captionTargetBox="[151,1425,921,1785]" captionTargetId="figure-180@7.[151,1436,921,1794]" captionTargetPageId="7" captionText="FIGURE 5. Adult male Myxia villaloboi sp. n. forewing venation; wing venation ter54minology following Bourgoin et al. (2015); black text = vein, italic text = crossvein, green text = cell." figureDoi="http://doi.org/10.5281/zenodo.12780217" httpUri="https://zenodo.org/record/12780217/files/figure.png" pageId="8" pageNumber="448">Fig. 5</figureCitation>
), faintly suffused, veins approximately concolorous bearing setae along veins, and a distinct, whitish stigma; apex of clavus exceeding midlength, Pcu+A1 fused just distad of claval midlength, composite vein reaching wing margin before CuP, combined vein stem Sc+R+M forming long stem from basal cell, fork of M from Sc+R basad of fusion of Pcu+A1; fork of RP from Sc+RA slightly proximad of wing midlength; CuA forked near distal claval margin. Branching pattern RA 1-branched, RP 3-branched, MP 4-branched; CuA 2-branched (distally anastomosed forming small closed ‘procubital cell’ [i.e., Emeljanov 1996]); crossveins 
<emphasis id="BFFEEAC3FF87927FFB05FEBA10A146C7" box="[1152,1171,367,392]" italics="true" pageId="8" pageNumber="448">ir</emphasis>
, 
<emphasis id="BFFEEAC3FF87927FFB26FEBA10F946C7" box="[1187,1227,367,392]" italics="true" pageId="8" pageNumber="448">r-m</emphasis>
, 
<emphasis id="BFFEEAC3FF87927FFB5EFEBA10C546C7" box="[1243,1271,367,392]" italics="true" pageId="8" pageNumber="448">im</emphasis>
, 
<emphasis id="BFFEEAC3FF87927FFA82FEBA110D46C7" box="[1287,1343,367,392]" italics="true" pageId="8" pageNumber="448">m-cu</emphasis>
and 
<emphasis id="BFFEEAC3FF87927FFAFFFEBA11AE46C7" box="[1402,1436,367,392]" italics="true" pageId="8" pageNumber="448">icu</emphasis>
present. Spinulation of hind legs 9-8-8.
</paragraph>
<paragraph id="8D3536D1FF87927FFF42FE6216304486" blockId="8.[151,1437,151,969]" pageId="8" pageNumber="448">
Terminalia. Pygofer in lateral view (
<figureCitation id="15B12A54FF87927FFDECFE6216F6469E" box="[617,708,439,465]" captionStart="FIGURE 6" captionStartId="8.[152,255,1924,1948]" captionTargetBox="[151,1436,993,1898]" captionTargetId="figure-370@8.[151,1436,993,1901]" captionTargetPageId="8" captionText="FIGURE 6. Adult male Myxia villaloboi sp. n. genitalia; A) lateral view, B) ventral view and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12780219" httpUri="https://zenodo.org/record/12780219/files/figure.png" pageId="8" pageNumber="448">Fig. 6A</figureCitation>
) broad, narrowest at dorsal margin, greatly expanded ventrally, ventral margin sinuate, posterior margin convex dorsal, then nearly straight to medioventral lobe at ventral margin; anterior margin concave, sinuate. In ventral view (
<figureCitation id="15B12A54FF87927FFD40FE2A172E4556" box="[709,796,511,537]" captionStart="FIGURE 6" captionStartId="8.[152,255,1924,1948]" captionTargetBox="[151,1436,993,1898]" captionTargetId="figure-370@8.[151,1436,993,1901]" captionTargetPageId="8" captionText="FIGURE 6. Adult male Myxia villaloboi sp. n. genitalia; A) lateral view, B) ventral view and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12780219" httpUri="https://zenodo.org/record/12780219/files/figure.png" pageId="8" pageNumber="448">Fig. 6B</figureCitation>
), medioventral process subtriangular, approximately 1.5x as tall as wide at base. Anal segment in lateral view stout, distally downcurved with rounded apex, ventral margin with triangular projection on lateral margins (
<figureCitation id="15B12A54FF87927FFD0AFD9216DA452E" box="[655,744,583,609]" captionStart="FIGURE 6" captionStartId="8.[152,255,1924,1948]" captionTargetBox="[151,1436,993,1898]" captionTargetId="figure-370@8.[151,1436,993,1901]" captionTargetPageId="8" captionText="FIGURE 6. Adult male Myxia villaloboi sp. n. genitalia; A) lateral view, B) ventral view and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12780219" httpUri="https://zenodo.org/record/12780219/files/figure.png" pageId="8" pageNumber="448">Fig. 6A</figureCitation>
). Gonostyli in lateral view (
<figureCitation id="15B12A54FF87927FFB99FD921041452E" box="[1052,1139,583,609]" captionStart="FIGURE 6" captionStartId="8.[152,255,1924,1948]" captionTargetBox="[151,1436,993,1898]" captionTargetId="figure-370@8.[151,1436,993,1901]" captionTargetPageId="8" captionText="FIGURE 6. Adult male Myxia villaloboi sp. n. genitalia; A) lateral view, B) ventral view and C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12780219" httpUri="https://zenodo.org/record/12780219/files/figure.png" pageId="8" pageNumber="448">Fig. 6A</figureCitation>
) slender, weakly upcurved distally, dorsal and ventral margins slightly sinuate, bearing subapical dorsal projection, apex rounded; in ventral view, curved mesad, gonostyli slender, inner lateral margin concave bearing tooth on inner lateral margin just before apex, laterodistal apex truncate, dorsal knobs visible subapically on lateral margin, extending laterocephalad. Aedeagus (
<figureCitation id="15B12A54FF87927FFE92FD02156E45BE" box="[279,348,727,753]" captionStart="FIGURE 7" captionStartId="9.[152,255,884,908]" captionTargetBox="[151,1435,181,856]" captionTargetId="figure-24@9.[151,1435,181,861]" captionTargetPageId="9" captionText="FIGURE 7. Aedeagus of Myxia villaloboi sp. n.; A) left lateral view, B) right later view, C) ventral view, and D) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12780221" httpUri="https://zenodo.org/record/12780221/files/figure.png" pageId="8" pageNumber="448">Fig. 7</figureCitation>
) simple bearing one small, curved process (A1) arising subapically on left dorsolateral margin, angled dorsad, endosoma large, angled right (ventral view, 
<figureCitation id="15B12A54FF87927FFCA9FD2E17B3445A" box="[812,897,763,789]" captionStart="FIGURE 7" captionStartId="9.[152,255,884,908]" captionTargetBox="[151,1435,181,856]" captionTargetId="figure-24@9.[151,1435,181,861]" captionTargetPageId="9" captionText="FIGURE 7. Aedeagus of Myxia villaloboi sp. n.; A) left lateral view, B) right later view, C) ventral view, and D) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12780221" httpUri="https://zenodo.org/record/12780221/files/figure.png" pageId="8" pageNumber="448">Fig. 7C</figureCitation>
) with two slender elongate processes (E1 &amp; E2) arising subapically, E1 slender, arising on dorsal margin, curved dorsad, E2 slender, angled cephalad, approximately twice as long as E1. Phallobase large, subtending aedeagal left and ventral margins (ventral view, 
<figureCitation id="15B12A54FF87927FFB4BFC9611104412" box="[1230,1314,835,861]" captionStart="FIGURE 7" captionStartId="9.[152,255,884,908]" captionTargetBox="[151,1435,181,856]" captionTargetId="figure-24@9.[151,1435,181,861]" captionTargetPageId="9" captionText="FIGURE 7. Aedeagus of Myxia villaloboi sp. n.; A) left lateral view, B) right later view, C) ventral view, and D) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12780221" httpUri="https://zenodo.org/record/12780221/files/figure.png" pageId="8" pageNumber="448">Fig. 7C</figureCitation>
) one large, blunt process (PB1) arising on the left lateral side, extending to aedeagal apex; second process (PB2) arising on right lateral side, extending along ventral margin of aedeagus, reaching apex, terminating in elongated, slender process, J-curved dorsocephalad (
<figureCitation id="15B12A54FF87927FFE35FC7A15C64486" box="[432,500,943,969]" captionStart="FIGURE 7" captionStartId="9.[152,255,884,908]" captionTargetBox="[151,1435,181,856]" captionTargetId="figure-24@9.[151,1435,181,861]" captionTargetPageId="9" captionText="FIGURE 7. Aedeagus of Myxia villaloboi sp. n.; A) left lateral view, B) right later view, C) ventral view, and D) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12780221" httpUri="https://zenodo.org/record/12780221/files/figure.png" pageId="8" pageNumber="448">Fig. 7</figureCitation>
).
</paragraph>
<caption id="D9F56659FF87927FFF1DF85110C140D3" ID-DOI="http://doi.org/10.5281/zenodo.12780219" ID-Zenodo-Dep="12780219" box="[152,1267,1924,1948]" httpUri="https://zenodo.org/record/12780219/files/figure.png" pageId="8" pageNumber="448" startId="8.[152,255,1924,1948]" targetBox="[151,1436,993,1898]" targetPageId="8" targetType="figure">
<paragraph id="8D3536D1FF87927FFF1DF85110C140D3" blockId="8.[152,1267,1924,1948]" box="[152,1267,1924,1948]" pageId="8" pageNumber="448">
<emphasis id="BFFEEAC3FF87927FFF1DF851152A40D3" bold="true" box="[152,280,1924,1948]" pageId="8" pageNumber="448">FIGURE 6.</emphasis>
Adult male 
<taxonomicName id="4A8A4D52FF87927FFE13F850160F40D3" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[406,573,1924,1948]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="448" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. nov.">
<emphasis id="BFFEEAC3FF87927FFE13F850160F40D3" box="[406,573,1924,1948]" italics="true" pageId="8" pageNumber="448">Myxia villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF87927FFDC6F851164E40D3" bold="true" box="[579,636,1924,1948]" pageId="8" pageNumber="448">
<taxonomicNameLabel id="A4CD57B8FF87927FFDC6F851164E40D3" box="[579,636,1924,1948]" pageId="8" pageNumber="448" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
genitalia; A) lateral view, B) ventral view and C) dorsal view.
</paragraph>
</caption>
<caption id="D9F56659FF86927EFF1DFCA111A544C2" ID-DOI="http://doi.org/10.5281/zenodo.12780221" ID-Zenodo-Dep="12780221" box="[152,1431,884,909]" httpUri="https://zenodo.org/record/12780221/files/figure.png" pageId="9" pageNumber="449" startId="9.[152,255,884,908]" targetBox="[151,1435,181,856]" targetPageId="9" targetType="figure">
<paragraph id="8D3536D1FF86927EFF1DFCA111A544C2" blockId="9.[152,1431,884,909]" box="[152,1431,884,909]" pageId="9" pageNumber="449">
<emphasis id="BFFEEAC3FF86927EFF1DFCA1152A44C2" bold="true" box="[152,280,884,909]" pageId="9" pageNumber="449">FIGURE 7.</emphasis>
Aedeagus of 
<taxonomicName id="4A8A4D52FF86927EFE21FCA0167944C2" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[420,587,885,909]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. nov.">
<emphasis id="BFFEEAC3FF86927EFE21FCA0167944C2" box="[420,587,885,909]" italics="true" pageId="9" pageNumber="449">Myxia villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF86927EFDD4FCA016B844C2" bold="true" box="[593,650,885,909]" pageId="9" pageNumber="449">
<taxonomicNameLabel id="A4CD57B8FF86927EFDD4FCA016B844C2" box="[593,650,885,909]" pageId="9" pageNumber="449" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; A) left lateral view, B) right later view, C) ventral view, and D) dorsal view.
</paragraph>
</caption>
<caption id="D9F56659FF86927EFF1DF9E917CA4136" ID-DOI="http://doi.org/10.5281/zenodo.12780223" ID-Zenodo-Dep="12780223" httpUri="https://zenodo.org/record/12780223/files/figure.png" pageId="9" pageNumber="449" startId="9.[152,255,1596,1620]" targetBox="[157,1430,938,1567]" targetPageId="9" targetType="figure">
<paragraph id="8D3536D1FF86927EFF1DF9E917CA4136" blockId="9.[151,1436,1596,1657]" pageId="9" pageNumber="449">
<emphasis id="BFFEEAC3FF86927EFF1DF9E9152A411A" bold="true" box="[152,280,1596,1621]" pageId="9" pageNumber="449">FIGURE 8.</emphasis>
Maximum Likelihood phylogenetic trees based on 1000 replicates; A) 18S rRNA, B) D9-D10 expansion region of 28S rRNA, C) H3 and D) 5’ region of COI; scale bar = percent nucleotide difference.
</paragraph>
</caption>
<paragraph id="8D3536D1FF86927EFF42F91E172E41AA" blockId="9.[151,1436,1739,1947]" box="[199,796,1739,1765]" pageId="9" pageNumber="449">
<emphasis id="BFFEEAC3FF86927EFF42F91E159141AA" bold="true" box="[199,419,1739,1765]" pageId="9" pageNumber="449">Plant associations.</emphasis>
Unknown; collected at light trap.
</paragraph>
</subSubSection>
<subSubSection id="C590655AFF86927EFF42F93A11024046" box="[199,1328,1775,1801]" pageId="9" pageNumber="449" type="distribution">
<paragraph id="8D3536D1FF86927EFF42F93A11024046" blockId="9.[151,1436,1739,1947]" box="[199,1328,1775,1801]" pageId="9" pageNumber="449">
<emphasis id="BFFEEAC3FF86927EFF42F93A15534046" bold="true" box="[199,353,1775,1801]" pageId="9" pageNumber="449">Distribution.</emphasis>
Hotel Villa Blanca, Los Angeles Cloud Forest Reserve, 
<collectingRegion id="4F4EF833FF86927EFC53F93A10904046" box="[982,1186,1775,1801]" country="Costa Rica" name="Alajuela" pageId="9" pageNumber="449">Alajuela Province</collectingRegion>
, 
<collectingCountry id="F59D7641FF86927EFB2BF93A111E4046" box="[1198,1324,1775,1801]" name="Costa Rica" pageId="9" pageNumber="449">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C590655AFF86927EFF42F8C614DF401E" pageId="9" pageNumber="449" type="etymology">
<paragraph id="8D3536D1FF86927EFF42F8C614DF401E" blockId="9.[151,1436,1739,1947]" pageId="9" pageNumber="449">
<emphasis id="BFFEEAC3FF86927EFF42F8C6157C4062" bold="true" box="[199,334,1811,1837]" pageId="9" pageNumber="449">Etymology.</emphasis>
The specific epithet is a patronym for Jairo Villalobo who helped in coordinating field work at Villa Blanca.
</paragraph>
</subSubSection>
<subSubSection id="C590655AFF86927CFF42F88E15304652" lastPageId="11" lastPageNumber="451" pageId="9" pageNumber="449" type="materials_examined">
<paragraph id="8D3536D1FF86927EFF42F88E110340D6" blockId="9.[151,1436,1739,1947]" pageId="9" pageNumber="449">
<emphasis id="BFFEEAC3FF86927EFF42F88E1582403A" bold="true" box="[199,432,1883,1909]" pageId="9" pageNumber="449">Material examined.</emphasis>
<materialsCitation id="3DE23C8CFF86927EFE32F88E16B340D6" collectingDate="2021-07-19" collectionCode="FLREC" collectorName="Coll. &amp; B. W. Bahder" country="Costa Rica" location="Myxia" municipality="Villa Blanca" pageId="9" pageNumber="449" specimenCount="2" specimenCount-male="2" stateProvince="Alajuela" typeStatus="holotype">
<typeStatus id="52318873FF86927EFE32F88E162D403A" box="[439,543,1883,1909]" pageId="9" pageNumber="449" type="holotype">Holotype</typeStatus>
<specimenCount id="9B8CFD58FF86927EFDA3F88E166E403A" box="[550,604,1883,1909]" pageId="9" pageNumber="449" type="male">male</specimenCount>
“ 
<collectingCountry id="F59D7641FF86927EFDE8F88E16DE403A" box="[621,748,1883,1909]" name="Costa Rica" pageId="9" pageNumber="449">Costa Rica</collectingCountry>
, 
<collectingRegion id="4F4EF833FF86927EFD73F88E1767403A" box="[758,853,1883,1909]" country="Costa Rica" name="Alajuela" pageId="9" pageNumber="449">Alajuela</collectingRegion>
Pr. / 
<collectingMunicipality id="6D51ACABFF86927EFC0AF88E1028403A" box="[911,1050,1883,1909]" country="Costa Rica" name="Villa Blanca" pageId="9" pageNumber="449" stateProvince="Alajuela">Villa Blanca</collectingMunicipality>
/ 
<date id="F9341011FF86927EFBAAF88E108A403A" box="[1071,1208,1883,1909]" pageId="9" pageNumber="449" value="2021-07-19">
<collectingDate id="E970E9F9FF86927EFBAAF88E108A403A" box="[1071,1208,1883,1909]" pageId="9" pageNumber="449" value="2021-07-19">19.VII.2021</collectingDate>
</date>
/ 
<collectorName id="207F5307FF86927EFB48F88E113A403A" box="[1229,1288,1883,1909]" pageId="9" pageNumber="449">Coll.</collectorName>
: 
<collectorName id="207F5307FF86927EFA96F88E11AE403A" box="[1299,1436,1883,1909]" pageId="9" pageNumber="449">B.W.Bahder</collectorName>
// Holotype / 
<emphasis id="BFFEEAC3FF86927EFEA9F855163240D4" box="[300,512,1919,1947]" italics="true" pageId="9" pageNumber="449">
<taxonomicName id="4A8A4D52FF86927EFEA9F85515D640D6" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[300,484,1919,1945]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="species" species="villaloboi">
<location id="8855600AFF86927EFEA9F855154040D6" LSID="urn:lsid:plazi:treatment:052387C7FF8A927CFF12FD2C1114452E:8855600AFF86927EFEA9F855154040D6" box="[300,370,1920,1945]" country="Costa Rica" municipality="Villa Blanca" name="Myxia" pageId="9" pageNumber="449" stateProvince="Alajuela">Myxia</location>
villaloboi
</taxonomicName>
<specimenCount id="9B8CFD58FF86927EFE6EF855163240D4" box="[491,512,1920,1947]" count="1" pageId="9" pageNumber="449" type="male">♂</specimenCount>
</emphasis>
” (
<collectionCode id="EB9BAE14FF86927EFD9EF8AA164B40D6" box="[539,633,1919,1945]" pageId="9" pageNumber="449">FLREC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3DE23C8CFF86927EFD09F8AA111F40D6" box="[652,1325,1919,1945]" collectionCode="FSCA" pageId="9" pageNumber="449" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="52318873FF86927EFD09F8AA16CB40D6" box="[652,761,1919,1945]" pageId="9" pageNumber="449" type="paratype">Paratypes</typeStatus>
<specimenCount id="9B8CFD58FF86927EFC85F8AA177C40D6" box="[768,846,1919,1945]" count="2" pageId="9" pageNumber="449" type="male">2 male</specimenCount>
, 
<specimenCount id="9B8CFD58FF86927EFCDCF8AA17FA40D6" box="[857,968,1919,1945]" count="3" pageId="9" pageNumber="449" type="female">3 females</specimenCount>
, same data as holotype (
<collectionCode id="EB9BAE14FF86927EFB5CF8AA111740D6" box="[1241,1317,1919,1945]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="9" pageNumber="449" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="D9F56659FF85927DFF1DF90910114056" ID-DOI="http://doi.org/10.5281/zenodo.12780225" ID-Zenodo-Dep="12780225" httpUri="https://zenodo.org/record/12780225/files/figure.png" pageId="10" pageNumber="450" startId="10.[152,255,1756,1780]" targetBox="[195,1333,196,1722]" targetPageId="10" targetType="figure">
<paragraph id="8D3536D1FF85927DFF1DF90910114056" blockId="10.[151,1437,1756,1817]" pageId="10" pageNumber="450">
<emphasis id="BFFEEAC3FF85927DFF1DF909152B41BB" bold="true" box="[152,281,1756,1780]" pageId="10" pageNumber="450">FIGURE 9.</emphasis>
Maximum Likelihood consensus tree based on 1000 replicates for concatenated sequence data for the 18S rRNA gene, D9-D10 expansion region of the 28S gene, H3 gene, and 5’ region of the COI gene.
</paragraph>
</caption>
<paragraph id="8D3536D1FF85927CFF42F8BF15304652" blockId="10.[151,1437,1898,2033]" lastBlockId="11.[151,1437,151,609]" lastPageId="11" lastPageNumber="451" pageId="10" pageNumber="450">
<emphasis id="BFFEEAC3FF85927DFF42F8BF154D40CA" bold="true" box="[199,383,1898,1925]" pageId="10" pageNumber="450">Sequence Data.</emphasis>
For the COI gene a 529 bp product was generated, for 18S a 1,363 bp product was generated, for 28S a 774 bp product was generated, and for H3 a 315 bp product was generated (
<tableCitation id="C008036AFF85927DFBC3F85A10AF40E6" box="[1094,1181,1935,1961]" captionStart="TABLE 2" captionStartId="2.[152,244,1590,1614]" captionTargetPageId="2" captionText="TABLE 2. Molecular taxon sampling and GenBank accession numbers for specimens used in analysis (all specimens stored at FLREC)." httpUri="http://table.plazi.org/id/D9F56659FF8D9275FF1DF9E3156E413D" pageId="10" pageNumber="450" tableUuid="D9F56659FF8D9275FF1DF9E3156E413D">Table 2</tableCitation>
). For both COI and H3 phylogenies (
<figureCitation id="15B12A54FF85927DFEABF86615404082" box="[302,370,1971,1997]" captionStart="FIGURE 8" captionStartId="9.[152,255,1596,1620]" captionTargetBox="[157,1430,938,1567]" captionTargetId="figure-55@9.[151,1435,933,1573]" captionTargetPageId="9" captionText="FIGURE 8. Maximum Likelihood phylogenetic trees based on 1000 replicates; A) 18S rRNA, B) D9-D10 expansion region of 28S rRNA, C) H3 and D) 5’ region of COI; scale bar = percent nucleotide difference." figureDoi="http://doi.org/10.5281/zenodo.12780223" httpUri="https://zenodo.org/record/12780223/files/figure.png" pageId="10" pageNumber="450">Fig. 8</figureCitation>
), there was generally low bootstrap support for most relationships, however, both phylogenies based on 18S and 28S show strong bootstrap support for both the monophyly of 
<taxonomicName id="4A8A4D52FF85927DFBB2F802104F40BF" authorityName="Bahder &amp; Bartlett" authorityYear="2019" box="[1079,1149,2007,2032]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF85927DFBB2F802104F40BF" box="[1079,1149,2007,2032]" italics="true" pageId="10" pageNumber="450">Myxia</emphasis>
</taxonomicName>
and the placement of 
<taxonomicName id="4A8A4D52FF85927CFAFBF802153047FE" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" lastPageId="11" lastPageNumber="451" order="Hemiptera" pageId="10" pageNumber="450" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. n.">
<emphasis id="BFFEEAC3FF85927CFAFBF802153047FE" italics="true" lastPageId="11" lastPageNumber="451" pageId="10" pageNumber="450">M. villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF84927CFE8EFF42157E47FE" bold="true" box="[267,332,151,177]" pageId="11" pageNumber="451">
<taxonomicNameLabel id="A4CD57B8FF84927CFE8EFF42157E47FE" box="[267,332,151,177]" pageId="11" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
within 
<taxonomicName id="4A8A4D52FF84927CFE20FF4215D947FF" authorityName="Bahder &amp; Bartlett" authorityYear="2019" box="[421,491,151,176]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFE20FF4215D947FF" box="[421,491,151,176]" italics="true" pageId="11" pageNumber="451">Myxia</emphasis>
</taxonomicName>
(99 and 100 respectively). The consensus tree (
<figureCitation id="15B12A54FF84927CFB8AFF42106647FE" box="[1039,1108,151,177]" captionStart="FIGURE 9" captionStartId="10.[152,255,1756,1780]" captionTargetBox="[195,1333,196,1722]" captionTargetId="figure-17@10.[189,1399,181,1732]" captionTargetPageId="10" captionText="FIGURE 9. Maximum Likelihood consensus tree based on 1000 replicates for concatenated sequence data for the 18S rRNA gene, D9-D10 expansion region of the 28S gene, H3 gene, and 5’ region of the COI gene." figureDoi="http://doi.org/10.5281/zenodo.12780225" httpUri="https://zenodo.org/record/12780225/files/figure.png" pageId="11" pageNumber="451">Fig. 9</figureCitation>
) based on concatenated data from all loci demonstrated strong bootstrap support for the monophyly of 
<taxonomicName id="4A8A4D52FF84927CFC6BFF6E1006479B" authorityName="Bahder &amp; Bartlett" authorityYear="2019" box="[1006,1076,187,212]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFC6BFF6E1006479B" box="[1006,1076,187,212]" italics="true" pageId="11" pageNumber="451">Myxia</emphasis>
</taxonomicName>
(100) with 
<taxonomicName id="4A8A4D52FF84927CFB3BFF6E1163479A" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[1214,1361,187,213]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. n.">
<emphasis id="BFFEEAC3FF84927CFB3BFF6E1163479A" box="[1214,1361,187,213]" italics="true" pageId="11" pageNumber="451">M. villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF84927CFADEFF6E11AF479A" bold="true" box="[1371,1437,187,213]" pageId="11" pageNumber="451">
<taxonomicNameLabel id="A4CD57B8FF84927CFADEFF6E11AF479A" box="[1371,1437,187,213]" pageId="11" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
resolving within this group with strong support for resolving closest to 
<taxonomicName id="4A8A4D52FF84927CFC29FF0A107E47B6" authorityName="Bahder &amp; Bartlett" authorityYear="2021" box="[940,1100,223,249]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="hernandezi">
<emphasis id="BFFEEAC3FF84927CFC29FF0A107E47B6" box="[940,1100,223,249]" italics="true" pageId="11" pageNumber="451">M. hernandezi</emphasis>
</taxonomicName>
(100) relative to all other taxa analyzed.
</paragraph>
</subSubSection>
<subSubSection id="C590655AFF84927CFF42FEF215874556" pageId="11" pageNumber="451" type="discussion">
<paragraph id="8D3536D1FF84927CFF42FEF216DC469E" blockId="11.[151,1437,151,609]" pageId="11" pageNumber="451">
<emphasis id="BFFEEAC3FF84927CFF42FEF21509460E" bold="true" box="[199,315,295,321]" pageId="11" pageNumber="451">Remarks.</emphasis>
The placement of the new species within 
<taxonomicName id="4A8A4D52FF84927CFCA0FEF21759460F" authorityName="Bahder &amp; Bartlett" authorityYear="2019" box="[805,875,295,320]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFCA0FEF21759460F" box="[805,875,295,320]" italics="true" pageId="11" pageNumber="451">Myxia</emphasis>
</taxonomicName>
is supported both by morphology and molecular data. The structure of the head and terminalia are consistent with other 
<taxonomicName id="4A8A4D52FF84927CFC4DFE9E1042462A" box="[968,1136,331,357]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BFFEEAC3FF84927CFC4DFE9E103C462B" box="[968,1038,331,356]" italics="true" pageId="11" pageNumber="451">Myxia</emphasis>
species.
</taxonomicName>
The form of the aedeagus and molecular analysis support 
<taxonomicName id="4A8A4D52FF84927CFD83FEBA16A846C6" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[518,666,367,393]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. n.">
<emphasis id="BFFEEAC3FF84927CFD83FEBA16A846C6" box="[518,666,367,393]" italics="true" pageId="11" pageNumber="451">M. villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF84927CFD21FEBA16D546C6" bold="true" box="[676,743,367,393]" pageId="11" pageNumber="451">
<taxonomicNameLabel id="A4CD57B8FF84927CFD21FEBA16D546C6" box="[676,743,367,393]" pageId="11" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
as new. With the current data, 
<taxonomicName id="4A8A4D52FF84927CFBD2FEBA10AF46C7" authorityName="Bahder &amp; Bartlett" authorityYear="2019" box="[1111,1181,367,392]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFBD2FEBA10AF46C7" box="[1111,1181,367,392]" italics="true" pageId="11" pageNumber="451">Myxia</emphasis>
</taxonomicName>
is sister to a clade of (
<taxonomicName id="4A8A4D52FF84927CFF24FE46150746E2" authorityName="Van Duzee" authorityYear="1923" box="[161,309,403,429]" class="Insecta" family="Cixiidae" genus="Nymphocixia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFF24FE46150746E2" box="[161,309,403,429]" italics="true" pageId="11" pageNumber="451">Nymphocixia</emphasis>
</taxonomicName>
+ 
<taxonomicName id="4A8A4D52FF84927CFEC1FE46158546E2" authorityName="Fowler" authorityYear="1904" box="[324,439,403,429]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFEC1FE46158546E2" box="[324,439,403,429]" italics="true" pageId="11" pageNumber="451">Haplaxius</emphasis>
</taxonomicName>
), with (
<taxonomicName id="4A8A4D52FF84927CFD97FE46166D46E2" authorityName="Bahder &amp; Bartlett" authorityYear="2019" box="[530,607,403,429]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">Myxia</taxonomicName>
+(
<taxonomicName id="4A8A4D52FF84927CFDF3FE46173846E2" authorityName="Van Duzee" authorityYear="1923" box="[630,778,403,429]" class="Insecta" family="Cixiidae" genus="Nymphocixia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFDF3FE46173846E2" box="[630,778,403,429]" italics="true" pageId="11" pageNumber="451">Nymphocixia</emphasis>
</taxonomicName>
+ 
<taxonomicName id="4A8A4D52FF84927CFC9FFE4617BF46E2" authorityName="Fowler" authorityYear="1904" box="[794,909,403,429]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFC9FFE4617BF46E2" box="[794,909,403,429]" italics="true" pageId="11" pageNumber="451">Haplaxius</emphasis>
</taxonomicName>
)) sister to 
<taxonomicName id="4A8A4D52FF84927CFB8FFE46105646E2" authorityName="Stal" authorityYear="1862" box="[1034,1124,403,429]" class="Insecta" family="Cixiidae" genus="Oecleus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFB8FFE46105646E2" box="[1034,1124,403,429]" italics="true" pageId="11" pageNumber="451">Oecleus</emphasis>
</taxonomicName>
, a finding that is similar to our past results in the clade (
<bibRefCitation id="E91B4B20FF84927CFE52FE6216D3469E" author="Echavarria, M. A. Z. &amp; Barrantes, E. A. B. &amp; Bartlett, C. R. &amp; Helmick, E. E. &amp; Bahder, B. W." box="[471,737,439,465]" pageId="11" pageNumber="451" pagination="417 - 428" refId="ref5688" refString="Echavarria, M. A. Z., Barrantes, E. A. B., Bartlett, C. R., Helmick, E. E. &amp; Bahder, B. W. (2021 a) A new species of Myxia (Hemiptera: Fulgoroidea: Cixiidae) collected on palms from the Reserva Privada el Silenecio de Los Angeles Cloud Forest in Costa Rica. Zootaxa, 5027 (3), 417 - 428. https: // doi. org / 10.11646 / zootaxa. 4915.3.4" type="journal article" year="2021">
Echavarria 
<emphasis id="BFFEEAC3FF84927CFDDCFE6216A0469E" box="[601,658,439,465]" italics="true" pageId="11" pageNumber="451">et al.</emphasis>
2021a
</bibRefCitation>
).
</paragraph>
<paragraph id="8D3536D1FF84927CFF42FE0E15874556" blockId="11.[151,1437,151,609]" pageId="11" pageNumber="451">
The large areolet on the distal portion of the vertex (
<figureCitation id="15B12A54FF84927CFC93FE0E175C46BA" box="[790,878,475,501]" captionStart="FIGURE 4" captionStartId="6.[152,255,1862,1886]" captionTargetBox="[151,1429,1291,1832]" captionTargetId="figure-42@6.[151,1435,1285,1838]" captionTargetPageId="6" captionText="FIGURE 4. Adult male Myxia villaloboi sp. n.; A) dorsal view of head, pronotum and mesonotum, B) lateral view of head, pronotum and mesonotum and C) frontal view of head; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12780215" httpUri="https://zenodo.org/record/12780215/files/figure.png" pageId="11" pageNumber="451">Fig. 4A</figureCitation>
) is unique to 
<taxonomicName id="4A8A4D52FF84927CFB8FFE0E10A946BA" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Bartlett &amp; Helmick &amp; Bahder" authorityYear="2024" box="[1034,1179,475,501]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="villaloboi" status="sp. n.">
<emphasis id="BFFEEAC3FF84927CFB8FFE0E10A946BA" box="[1034,1179,475,501]" italics="true" pageId="11" pageNumber="451">M. villaloboi</emphasis>
</taxonomicName>
<emphasis id="BFFEEAC3FF84927CFB27FE0E10D146BA" bold="true" box="[1186,1251,475,501]" pageId="11" pageNumber="451">
<taxonomicNameLabel id="A4CD57B8FF84927CFB27FE0E10D146BA" box="[1186,1251,475,501]" pageId="11" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
relative to other species of 
<taxonomicName id="4A8A4D52FF84927CFE88FE2A15614557" authorityName="Bahder &amp; Bartlett" authorityYear="2019" box="[269,339,511,536]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEAC3FF84927CFE88FE2A15614557" box="[269,339,511,536]" italics="true" pageId="11" pageNumber="451">Myxia</emphasis>
</taxonomicName>
(
<figureCitation id="15B12A54FF84927CFEE6FE2A159A4556" box="[355,424,511,537]" captionStart="FIGURE 2" captionStartId="4.[152,255,1669,1693]" captionTargetBox="[159,1427,181,1645]" captionTargetId="figure-17@4.[159,1427,181,1645]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal view of head and thorax of Myxia species; A) M. baynardi, B) M. hernandezi, C) M. delta, D) M. belinda." figureDoi="http://doi.org/10.5281/zenodo.12780211" httpUri="https://zenodo.org/record/12780211/files/figure.png" pageId="11" pageNumber="451">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C590655AFF84927CFF42FDF715CE4573" box="[199,508,546,572]" pageId="11" pageNumber="451" type="materials_examined">
<paragraph id="8D3536D1FF84927CFF42FDF715CE4573" blockId="11.[151,1437,151,609]" box="[199,508,546,572]" pageId="11" pageNumber="451">
<emphasis id="BFFEEAC3FF84927CFF42FDF715CE4573" bold="true" box="[199,508,546,572]" pageId="11" pageNumber="451">Other material examined.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C590655AFF84927CFF42FD921114452E" box="[199,1318,583,609]" pageId="11" pageNumber="451" type="reference_group">
<paragraph id="8D3536D1FF84927CFF42FD921114452E" blockId="11.[151,1437,151,609]" box="[199,1318,583,609]" pageId="11" pageNumber="451">
<taxonomicName id="4A8A4D52FF84927CFF42FD9215CC452E" authority="(Kramer 1979)" baseAuthorityName="Kramer" baseAuthorityYear="1979" box="[199,510,583,609]" class="Insecta" family="Cixiidae" genus="Myxia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="delta">
<emphasis id="BFFEEAC3FF84927CFF42FD92157E452E" box="[199,332,583,609]" italics="true" pageId="11" pageNumber="451">Myxia delta</emphasis>
(
<bibRefCitation id="E91B4B20FF84927CFEDFFD9215C4452E" author="Kramer, J. P." box="[346,502,583,609]" pageId="11" pageNumber="451" pagination="301 - 389" refId="ref6072" refString="Kramer, J. P. (1979) Taxonomic study of the planthopper genus Myndus in the Americas (Homoptera: Fulgoroidea: Cixiidae). Transactions of the American Entomological Society, 105, 301 - 389." type="journal article" year="1979">Kramer 1979</bibRefCitation>
)
</taxonomicName>
: 
<materialsCitation id="3DE23C8CFF84927CFD8CFD92103A452E" box="[521,1032,583,609]" collectionCode="V" country="Costa Rica" location="Manzanillo" pageId="11" pageNumber="451" specimenCount="1" stateProvince="Limon">
<collectingCountry id="F59D7641FF84927CFD8CFD9216B4452E" box="[521,646,583,609]" name="Costa Rica" pageId="11" pageNumber="451">Costa Rica</collectingCountry>
, 
<location id="8855600AFF84927CFD14FD921721452E" LSID="urn:lsid:plazi:treatment:052387C7FF8A927CFF12FD2C1114452E:8855600AFF84927CFD14FD921721452E" box="[657,787,583,609]" country="Costa Rica" name="Manzanillo" pageId="11" pageNumber="451" stateProvince="Limon">Manzanillo</location>
, 
<collectingRegion id="4F4EF833FF84927CFC9AFD9217E4452E" box="[799,982,583,609]" country="Costa Rica" name="Limon" pageId="11" pageNumber="451">Limón Province</collectingRegion>
, 
<date id="F9341011FF84927CFC67FD92103A452E" box="[994,1032,583,609]" pageId="11" pageNumber="451" value="2021-05-07">
7. 
<collectionCode id="EB9BAE14FF84927CFC76FD92103A452E" box="[1011,1032,583,609]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="451" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3DE23C8CFF84927CFB8DFD921111452E" box="[1032,1315,583,609]" collectionCode="FLREC" country="Costa Rica" pageId="11" pageNumber="451" specimenCount="3" specimenCount-male="3" stateProvince="Limon">
.2021 (
<specimenCount id="9B8CFD58FF84927CFBD2FD921081452E" box="[1111,1203,583,609]" count="3" pageId="11" pageNumber="451" type="male">3 Males</specimenCount>
, 
<collectionCode id="EB9BAE14FF84927CFB3BFD921128452E" box="[1214,1306,583,609]" pageId="11" pageNumber="451">FLREC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>