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134 lines
16 KiB
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<document id="9694DF6746CE0001219B79E390420BF0" ID-DOI="10.5281/zenodo.13643699" ID-ISSN="1732-2421" ID-Zenodo-Dep="13643699" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723142205826" checkinUser="felipe" docAuthor="Soriano, Carmen & Delclòs, Xavier" docDate="2006" docId="455787BCFF992457FFDEF8E06A8DF9CD" docLanguage="en" docName="ActaPalaeontolPol.51.1.185-200.pdf" docOrigin="Acta Palaeontologica Polonica 51 (1)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Tetraphalerus ponomarenkoi Soriano & Delclòs, 2006, sp. nov." docType="treatment" docVersion="3" lastPageNumber="186" masterDocId="B96EFFC4FF982456FFB8FF876E36FFDB" masterDocTitle="New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family" masterLastPageNumber="200" masterPageNumber="185" pageNumber="186" updateTime="1725365340826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7717409B8A37882C8FAC2CAE7ECA14B2">New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family</mods:title>
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<mods:namePart id="E86D6BE1CC20ECC59218B223365A88C2">Soriano, Carmen</mods:namePart>
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<mods:roleTerm id="8C166658D76013334A24F69442176E23">Author</mods:roleTerm>
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<mods:namePart id="125470EE5D57AEFA2F097980691358D9">Delclòs, Xavier</mods:namePart>
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<mods:typeOfResource id="945322AECDF98F99A646989C2A3BBBD8">text</mods:typeOfResource>
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<mods:titleInfo id="4DD1936C931B65B8BEC03677DB910D8F">
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<mods:title id="B14AC596F69B5D64CC2949F40FB7740D">Acta Palaeontologica Polonica</mods:title>
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</mods:titleInfo>
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<mods:part id="D22D9F032AD697A2711F2CDEBFAEBEA0">
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<mods:date id="192932942FFC03A512787A184A5F38AD">2006</mods:date>
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<mods:number id="258A3BE14004E5F5099FFD177E8CF927">51</mods:number>
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<mods:number id="A1D510C3305F29568663B53A77AD2F15">1</mods:number>
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<mods:identifier id="DFCA4DA9EEFD61DDED630E8D146D2CCA" type="DOI">10.5281/zenodo.13643699</mods:identifier>
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<mods:identifier id="F07CF2A474630D8B1E6383BCE63CD9D0" type="ISSN">1732-2421</mods:identifier>
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<treatment id="455787BCFF992457FFDEF8E06A8DF9CD" ID-DOI="http://doi.org/10.5281/zenodo.13643705" ID-Zenodo-Dep="13643705" LSID="urn:lsid:plazi:treatment:455787BCFF992457FFDEF8E06A8DF9CD" httpUri="http://treatment.plazi.org/id/455787BCFF992457FFDEF8E06A8DF9CD" lastPageNumber="186" pageId="1" pageNumber="186">
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<subSubSection id="85E46521FF992457FFDEF8E06C7EF859" box="[102,584,1894,1923]" pageId="1" pageNumber="186" type="nomenclature">
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<paragraph id="CD4136AAFF992457FFDEF8E06C7EF859" blockId="1.[102,584,1894,1957]" box="[102,584,1894,1923]" pageId="1" pageNumber="186">
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<heading id="960981C6FF992457FFDEF8E06C7EF859" box="[102,584,1894,1923]" fontSize="12" level="2" pageId="1" pageNumber="186" reason="9">
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<taxonomicName id="0AFE4D29FF992457FFDEF8E06FEAF858" authorityName="Soriano & Delclòs" authorityYear="2006" box="[102,476,1895,1923]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="ponomarenkoi" status="sp. nov.">
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<emphasis id="FF8AEAB8FF992457FFDEF8E06FEAF858" box="[102,476,1895,1923]" italics="true" pageId="1" pageNumber="186">Tetraphalerus ponomarenkoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E4B957C3FF992457FE5CF8E16C7EF859" box="[484,584,1894,1922]" pageId="1" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="85E46521FF992457FFDEF8176A8DF9CD" pageId="1" pageNumber="186" type="description">
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<paragraph id="CD4136AAFF992457FFDEF8176EDFF87E" blockId="1.[102,584,1894,1957]" box="[102,233,1936,1957]" pageId="1" pageNumber="186">
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<figureCitation id="55C52A2FFF992457FFDEF8176E8CF87E" box="[102,186,1936,1957]" captionStart="Fig" captionStartId="2.[102,135,891,911]" captionTargetBox="[136,1459,230,863]" captionTargetId="figure-540@2.[124,1463,225,864]" captionTargetPageId="2" captionText="Fig. 1. Camera lucida drawings of Barremian (Early Cretaceous) ommatin Tetraphalerus from Spain. A. Tetraphalerus ponomarenkoi sp. nov. Holotype MCCM LH 78a, part, Las Hoyas, Cuenca Province, dorsal (A1) and ventral (A2) aspects. B. Tetraphalerus penalveri sp. nov. Holotype DEPGM LP 0058b, counterpart, La Pedrera, Lleida Province, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643701" httpUri="https://zenodo.org/record/13643701/files/figure.png" pageId="1" pageNumber="186">Figs. 1A</figureCitation>
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,
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<figureCitation id="55C52A2FFF992457FF7CF8176ED3F87E" box="[196,229,1936,1957]" captionStart="Fig" captionStartId="3.[102,134,1570,1590]" captionTargetBox="[206,1382,225,1543]" captionTargetId="figure-150@3.[206,1382,225,1543]" captionTargetPageId="3" captionText="Fig. 2. Barremian (Early Cretaceous) ommatins Tetraphalerus, Cionocoleus (both Ommatini), Brochocoleus (Brochocoleini), and Zygadenia (Notocupedini) from Spain. A. Tetraphalerus ponomarenkoi sp. nov. Holotype MCCM LH 78a, part, Las Hoyas, Cuenca Province. B. Tetraphalerus penalveri sp. nov. Holotype DEPGM LP 0058b, counterpart, La Pedrera, Lleida Province. C. Cionocoleus longicapitis sp. nov. Holotype IEI LP 0164G, La Pedrera, Lleida Province. D. Brochocoleus indibili sp. nov. Holotype IEI LC 4636a, part, La Cabrúa, Lleida Province. E. Zygadenia viridis sp. nov. Holotype MCCM LH 1805a, part, Las Hoyas, Cuenca Province. F. Zygadenia oculata sp. nov. Holotype IEI LC 3675a, part, La Cabrúa, Lleida Province. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643703" httpUri="https://zenodo.org/record/13643703/files/figure.png" pageId="1" pageNumber="186">2A</figureCitation>
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.
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</paragraph>
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<paragraph id="CD4136AAFF992457FFDEF83F6EF3F832" blockId="1.[102,776,1976,2025]" pageId="1" pageNumber="186">
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1993
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<taxonomicName id="0AFE4D29FF992457FF24F83F6FB2F816" box="[156,388,1976,1997]" class="Magnoliopsida" family="Malvaceae" genus="Coleoptera" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="1" pageNumber="186" phylum="Tracheophyta" rank="subGenus" subGenus="Cupedidae">Coleoptera (Cupedidae)</taxonomicName>
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; Martínez−Delclòs and Ruíz de Loizaga: fig. 4.
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</paragraph>
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<paragraph id="CD4136AAFF992457FC94FF606A2FFEC3" blockId="1.[812,1486,231,435]" pageId="1" pageNumber="186">
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<materialsCitation id="7D963CF7FF992457FC94FF606A23FEC3" collectorName="La Cierva" country="Spain" location="Serrania de Cuenca" municipality="Cuenca Province" pageId="1" pageNumber="186" specimenCount="1" typeStatus="holotype">
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<emphasis id="FF8AEAB8FF992457FC94FF606D9DFF27" box="[812,939,231,252]" italics="true" pageId="1" pageNumber="186">
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<typeStatus id="12458808FF992457FC94FF606D6DFF27" box="[812,859,231,252]" pageId="1" pageNumber="186">Type</typeStatus>
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locality
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</emphasis>
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: Las Hoyas fossil site,
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<collectorName id="600B537CFF992457FB20FF606ACAFF27" box="[1176,1276,231,252]" pageId="1" pageNumber="186">La Cierva</collectorName>
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,
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<location id="C8216071FF992457FAB0FF606BFFFF27" LSID="urn:lsid:plazi:treatment:455787BCFF992457FFDEF8E06A8DF9CD:C8216071FF992457FAB0FF606BFFFF27" box="[1288,1481,231,252]" country="Spain" municipality="Cuenca Province" name="Serrania de Cuenca" pageId="1" pageNumber="186">Serranía de Cuenca</location>
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,
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<collectingMunicipality id="2D25ACD0FF992457FC94FE846DE4FEC3" box="[812,978,259,280]" pageId="1" pageNumber="186">Cuenca Province</collectingMunicipality>
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,
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<collectingCountry id="B5E9763AFF992457FC63FE846A23FEC3" box="[987,1045,259,280]" name="Spain" pageId="1" pageNumber="186">Spain</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="CD4136AAFF992457FC94FEA06A63FEAF" blockId="1.[812,1486,231,435]" pageId="1" pageNumber="186">
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<emphasis id="FF8AEAB8FF992457FC94FEA06DCFFEE7" box="[812,1017,295,316]" italics="true" pageId="1" pageNumber="186">Stratigraphic horizon</emphasis>
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: Second lithosome of finely laminated limestones of La Huérguina Formation, Lower Cretaceous, Barremian (Fregenal−Martínez and Meléndez 2000).
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</paragraph>
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<paragraph id="CD4136AAFF992457FC94FE056D8FFE68" blockId="1.[812,1486,231,435]" pageId="1" pageNumber="186">
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<emphasis id="FF8AEAB8FF992457FC94FE056A25FE4C" box="[812,1043,386,407]" italics="true" pageId="1" pageNumber="186">Derivation of the name</emphasis>
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: After the Russian palaeoentomologist A.G. Ponomarenko.
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</paragraph>
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<paragraph id="CD4136AAFF992457FC94FE406A18FDE4" blockId="1.[812,1485,454,575]" pageId="1" pageNumber="186">
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<emphasis id="FF8AEAB8FF992457FC94FE406DB8FE04" box="[812,910,455,479]" italics="true" pageId="1" pageNumber="186">Material</emphasis>
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.—
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<materialsCitation id="7D963CF7FF992457FC17FE416A18FDE4" collectionCode="MCCM" collectorName="de Cuenca" country="Spain" location="Serrania de Cuenca" municipality="Cuenca Province" pageId="1" pageNumber="186" specimenCode="LH 78" specimenCount="1" typeStatus="holotype">
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<typeStatus id="12458808FF992457FC17FE416A2FFE04" box="[943,1049,454,479]" pageId="1" pageNumber="186" type="holotype">Holotype</typeStatus>
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<collectionCode id="ABEFAE6FFF992457FB9BFE416A4CFE04" box="[1059,1146,454,479]" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="1" pageNumber="186">MCCM</collectionCode>
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<specimenCode id="9D589ED1FF992457FB3CFE406AE7FE04" box="[1156,1233,454,479]" collectionCode="MCCM" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="1" pageNumber="186">LH 78</specimenCode>
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, part and counterpart, from Las Hoyas fossil site in
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<location id="C8216071FF992457FB3EFE616B53FE24" LSID="urn:lsid:plazi:treatment:455787BCFF992457FFDEF8E06A8DF9CD:C8216071FF992457FB3EFE616B53FE24" box="[1158,1381,486,511]" country="Spain" municipality="Cuenca Province" name="Serrania de Cuenca" pageId="1" pageNumber="186">
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Serranía
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<collectorName id="600B537CFF992457FB55FE616B53FE24" box="[1261,1381,486,511]" pageId="1" pageNumber="186">de Cuenca</collectorName>
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</location>
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(
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<collectingMunicipality id="2D25ACD0FF992457FACFFE616DA4FDC4" pageId="1" pageNumber="186">Cuenca Province</collectingMunicipality>
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,
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<collectingCountry id="B5E9763AFF992457FC18FD816DD3FDC4" box="[928,997,518,543]" name="Spain" pageId="1" pageNumber="186">Spain</collectingCountry>
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). Well preserved specimen, lacking part of antennae and fore legs.
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</materialsCitation>
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</paragraph>
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<paragraph id="CD4136AAFF992457FC94FDD46B62FCD1" blockId="1.[812,1485,594,779]" pageId="1" pageNumber="186">
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<emphasis id="FF8AEAB8FF992457FC94FDD46DA8FDB1" box="[812,926,595,618]" italics="true" pageId="1" pageNumber="186">Diagnosis</emphasis>
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.—One of the largest species of the genus. The presence of two prominences in the vertex ascribes this species to
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<taxonomicName id="0AFE4D29FF992457FCC2FD156A5EFD71" authorityName="Heller" authorityYear="1913" box="[890,1128,658,682]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="bruchi">
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<emphasis id="FF8AEAB8FF992457FCC2FD156A5EFD71" box="[890,1128,658,682]" italics="true" pageId="1" pageNumber="186">Tetraphalerus bruchi</emphasis>
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</taxonomicName>
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series. From other large species of this series (
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<taxonomicName id="0AFE4D29FF992457FC74FD346B1AFD10" authority="Ponomarenko, 1997" authorityName="Ponomarenko" authorityYear="1997" box="[972,1324,690,715]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="collaris">
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<emphasis id="FF8AEAB8FF992457FC74FD346A76FD11" box="[972,1088,690,714]" italics="true" pageId="1" pageNumber="186">T. collaris</emphasis>
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<bibRefCitation id="A96F4B5BFF992457FBF0FD356B1AFD10" author="Ponomarenko, A. G." box="[1096,1324,690,715]" pageId="1" pageNumber="186" pagination="45 - 55" refId="ref10494" refString="Ponomarenko, A. G. 1997. New beetles of the Family Cupedidae from the Mesozoic of Mongolia. Ommatini, Mesocupedini, Priacmini. Paleontological Journal 31: 45 - 55." type="journal article" year="1997">Ponomarenko, 1997</bibRefCitation>
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</taxonomicName>
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) differs in the length of the head and pronotum, and the relation between the last abdominal segment and the previous one.
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</paragraph>
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<paragraph id="CD4136AAFF992457FC94FC996A8DF9CD" blockId="1.[812,1485,797,1558]" pageId="1" pageNumber="186">
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<emphasis id="FF8AEAB8FF992457FC94FC996D86FCEE" box="[812,944,798,821]" italics="true" pageId="1" pageNumber="186">Description</emphasis>
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.—Measurements: length
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<quantity id="0A069B4FFF992457FB67FC9A6B7AFCEE" box="[1247,1356,797,822]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="1" pageNumber="186" unit="mm" value="20.4">20.4 mm</quantity>
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; width
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<quantity id="0A069B4FFF992457FA12FC9A6D6DFC8E" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="1" pageNumber="186" unit="mm" value="8.7">8.7 mm</quantity>
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; elytra length
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<quantity id="0A069B4FFF992457FC44FCBA6A52FC8E" box="[1020,1124,829,854]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="186" unit="mm" value="12.7">12.7 mm</quantity>
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. Body large and elongate. Head somewhat elongate, about 1.4 times as long as wide, narrowed roundly after eyes; temples 1.5 times longer than eyes, not projected laterally, vertex with two oval flat prominences, with their posterior edge broadened. Eyes comparatively short. Scape about 1.5 times longer than antennomeres 2–4. Neck−like constriction clear, almost as long as the occipital part of head. Pronotum slightly transverse, about 1.5 times as wide as long, lateral edges flattened, anterior and posterior angles rounded; pronotal disc with two longitudinal grooves. Prosternum transverse. Procoxae rounded and contiguous. Mesosternum slightly transverse, quite wider than longer, with big depression before mesocoxae. Mesocoxae longer than wider, narrowing anteriorly, trochantin rounded. Metasternum transverse, about 1.4 times as wide as long, with longitudinal and transverse sutures clearly expressed. Metacoxae sub−triangular, abruptly narrowed laterally. Abdomen with 5 visible ventrites, last ventrite about 1.4 times as long as previous one. Elytra about 4.5 times as long as wide, epipleural rim narrow, with rows of tubercles. Meso− and metafemora narrowed at 2/3 of its length. Tibiae as long as femora. Body covered with tubercles, larger on head and pronotum, and smaller in abdomen.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |