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<document ID-DOI="10.3853/j.2201-4349.69.2017.1653" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237800" ID-ZooBank="68F315FF-3FEB-410E-96EC-5F494510F440" approvalRequired="65" approvalRequired_for_taxonomicNames="63" approvalRequired_for_treatments="2" checkinTime="1629535407606" checkinUser="felipe" docAuthor="Parnaby, Harry E., Ingleby, Sandy &amp; Divljan, Anja" docDate="2017" docId="03DD87C8FFDA73561887FB64FB0790F3" docLanguage="en" docName="RecAustMus.69.5.277-420.pdf" docOrigin="Records of the Australian Museum 69 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Rattus mordax subsp. hageni Troughton 1937" docType="treatment" docVersion="4" lastPageNumber="376" masterDocId="FFE4FFB0FFB973351832FFEBFFD5973C" masterDocTitle="Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney" masterLastPageNumber="420" masterPageNumber="277" pageNumber="376" updateTime="1674245310755" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Parnaby, Harry E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ingleby, Sandy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Divljan, Anja</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-10-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>69</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>277</mods:start>
<mods:end>420</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3853/j.2201-4349.69.2017.1653</mods:identifier>
<mods:identifier type="ISSN">2201-4349</mods:identifier>
<mods:identifier type="Zenodo-Dep">5237800</mods:identifier>
<mods:identifier type="ZooBank">68F315FF-3FEB-410E-96EC-5F494510F440</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03DD87C8FFDA73561887FB64FB0790F3" httpUri="http://treatment.plazi.org/id/03DD87C8FFDA73561887FB64FB0790F3" lastPageNumber="376" pageId="99" pageNumber="376">
<subSubSection box="[181,689,1167,1194]" pageId="99" pageNumber="376" type="nomenclature">
<paragraph blockId="99.[181,689,1167,1194]" box="[181,689,1167,1194]" pageId="99" pageNumber="376">
<heading box="[181,689,1167,1194]" centered="true" fontSize="11" level="2" pageId="99" pageNumber="376" reason="2">
<taxonomicName authority="Troughton, 1937 a" authorityName="Troughton" authorityYear="1937" box="[181,689,1167,1194]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="subSpecies" species="mordax" subSpecies="hageni">
<emphasis bold="true" box="[181,689,1167,1194]" pageId="99" pageNumber="376">
<emphasis bold="true" box="[181,456,1167,1194]" italics="true" pageId="99" pageNumber="376">Rattus mordax hageni</emphasis>
<bibRefCitation author="Troughton, E" box="[463,689,1167,1193]" pageId="99" pageNumber="376" pagination="117 - 127" refId="ref147573" refString="Troughton, E. L. G. 1937 a. Descriptions of some New Guinea mammals. Records of the Australian Museum 20 (2): 117 - 127. https: // doi. org / 10.3853 / j. 0067 - 1975.20.1937.254" type="journal article" year="1937">Troughton, 1937a</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,594,1208,1230]" pageId="99" pageNumber="376" type="reference_group">
<paragraph blockId="99.[151,594,1208,1230]" box="[151,594,1208,1230]" pageId="99" pageNumber="376">
<emphasis box="[151,302,1208,1230]" italics="true" pageId="99" pageNumber="376">Rec. Aust. Mus.</emphasis>
20(2): 120. (
<date box="[426,583,1208,1230]" pageId="99" pageNumber="376" value="1937-08-27">27 August 1937</date>
).
</paragraph>
</subSubSection>
<subSubSection box="[121,573,1246,1270]" pageId="99" pageNumber="376" type="vernacular_names">
<paragraph blockId="99.[121,573,1246,1270]" box="[121,573,1246,1270]" pageId="99" pageNumber="376">
<emphasis bold="true" box="[121,295,1246,1270]" pageId="99" pageNumber="376">Common name</emphasis>
. Steins New
<collectingCountry box="[445,522,1246,1270]" name="Guinea-Bissau" pageId="99" pageNumber="376">Guinea</collectingCountry>
Rat.
</paragraph>
</subSubSection>
<subSubSection pageId="99" pageNumber="376" type="description">
<paragraph blockId="99.[119,749,1286,1516]" pageId="99" pageNumber="376">
<emphasis bold="true" box="[121,291,1286,1310]" pageId="99" pageNumber="376">Current name</emphasis>
.
<taxonomicName authority="Troughton 1937 a" authorityName="Troughton" authorityYear="1937" box="[309,744,1286,1310]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="subSpecies" species="steini" subSpecies="hageni">
<emphasis box="[309,536,1286,1310]" italics="true" pageId="99" pageNumber="376">Rattus steini hageni</emphasis>
<bibRefCitation author="Troughton, E" box="[547,744,1286,1310]" pageId="99" pageNumber="376" pagination="117 - 127" refId="ref147573" refString="Troughton, E. L. G. 1937 a. Descriptions of some New Guinea mammals. Records of the Australian Museum 20 (2): 117 - 127. https: // doi. org / 10.3853 / j. 0067 - 1975.20.1937.254" type="journal article" year="1937">Troughton 1937a</bibRefCitation>
</taxonomicName>
; following
<bibRefCitation author="Taylor, J" box="[228,427,1316,1340]" pageId="99" pageNumber="376" pagination="177 - 336" refId="ref144998" refString="Taylor, J. M., J. H. Calaby, and H. M. Van Deusen. 1982. A revision of the genus Rattus (Rodentia, Muridae) in the New Guinean region. Bulletin of the American Museum of Natural History 173 (3): 177 - 336." type="journal article" year="1982">
Taylor
<emphasis box="[299,350,1316,1340]" italics="true" pageId="99" pageNumber="376">et al.</emphasis>
(1982)
</bibRefCitation>
and Musser &amp; Carleton (2005) but within
<collectingCountry box="[244,460,1345,1369]" name="Papua New Guinea" pageId="99" pageNumber="376">Papua New Guinea</collectingCountry>
<taxonomicName authority="Rummler, 1935" authorityName="Rummler" authorityYear="1935" box="[473,748,1345,1369]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="species" species="steini">
<emphasis box="[473,565,1346,1369]" italics="true" pageId="99" pageNumber="376">R. steini</emphasis>
<bibRefCitation author="Rummler, H" box="[576,748,1345,1369]" pageId="99" pageNumber="376" pagination="105 - 118" refId="ref143780" refString="Rummler, H. 1935. Neue Muriden aus Neuguinea. Zeitschrift fur Saugetierkunde 10: 105 - 118." type="journal article" year="1935">Rümmler, 1935</bibRefCitation>
</taxonomicName>
includes multiple species requiring resolution, see
<bibRefCitation author="Robins, J &amp; Tintinger, K &amp; Aplin, M &amp; Hingston, E &amp; Matisoo-Smith, D" pageId="99" pageNumber="376" pagination="98002" refId="ref143702" refString="Robins, J. H., V. Tintinger, K. P. Aplin, M. Hingston, E. Matisoo-Smith, D. Penny, and S. D. Lavery. 2014. Phylogenetic species identification in Rattus Highlights rapid radiation and morphological similarity of New Guinean species. PLoS ONE 9 (5): e 98002. https: // doi. org / 10.1371 / journal. pone. 0098002" type="journal article" year="2014">
Robins
<emphasis italics="true" pageId="99" pageNumber="376">et al.</emphasis>
(2014: 14)
</bibRefCitation>
.
<bibRefCitation author="Helgen, K" box="[280,435,1404,1428]" pageId="99" pageNumber="376" pagination="689 - 749" refId="ref132662" refString="Helgen, K. M. 2007. A taxonomic and geographic overview of the mammals of Papua. In The Ecology of Papua, ed. A. J. Marshall and B. M. Beehler, Ecology of Indonesia Series, Vol. VI, Part 1 pp. 689 - 749. Singapore: Periplus Editions." type="book chapter" year="2007">Helgen (2007)</bibRefCitation>
regarded eastern populations of
<taxonomicName authorityName="Rummler" authorityYear="1935" box="[151,239,1434,1457]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="species" species="steini">
<emphasis box="[151,239,1434,1457]" italics="true" pageId="99" pageNumber="376">R. steini</emphasis>
</taxonomicName>
to be a distinct species referable to
<taxonomicName authority="(Rummler, 1935)" baseAuthorityName="Rummler" baseAuthorityYear="1935" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="species" species="foersteri">
<emphasis box="[628,749,1433,1457]" italics="true" pageId="99" pageNumber="376">R. foersteri</emphasis>
(
<bibRefCitation author="Rummler, H" box="[128,302,1462,1486]" pageId="99" pageNumber="376" pagination="105 - 118" refId="ref143780" refString="Rummler, H. 1935. Neue Muriden aus Neuguinea. Zeitschrift fur Saugetierkunde 10: 105 - 118." type="journal article" year="1935">Rümmler, 1935</bibRefCitation>
)
</taxonomicName>
, and placed
<taxonomicName authorityName="Troughton" authorityYear="1937" box="[449,522,1462,1486]" class="Mammalia" family="Muridae" genus="Melomys" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="species" species="hageni">
<emphasis box="[449,522,1462,1486]" italics="true" pageId="99" pageNumber="376">hageni</emphasis>
</taxonomicName>
as a synonym of the latter.
</paragraph>
</subSubSection>
<subSubSection pageId="99" pageNumber="376" type="materials_examined">
<paragraph blockId="99.[119,748,1531,1615]" pageId="99" pageNumber="376">
<emphasis bold="true" box="[121,224,1532,1556]" pageId="99" pageNumber="376">
<typeStatus box="[121,224,1532,1556]" pageId="99" pageNumber="376">Holotype</typeStatus>
</emphasis>
. M.6102 by original designation. Male adult, skull (
<figureCitation box="[127,211,1561,1585]" captionStart="Figure 45" captionStartId="99.[794,857,1683,1705]" captionTargetBox="[799,1414,161,1658]" captionTargetId="figure-538@99.[794,1421,151,1669]" captionTargetPageId="99" captionText="Figure 45. AM M.6102, holotype skull of Rattus mordax hageni Troughton, 1937a. (Photography by Stuart Humphreys)." figureDoi="http://doi.org/10.5281/zenodo.5237896" httpUri="https://zenodo.org/record/5237896/files/figure.png" pageId="99" pageNumber="376">Fig. 45</figureCitation>
), study skin, collected by Dr G. A. M. Heydon, collection date not given, registered
<date box="[509,655,1590,1615]" pageId="99" pageNumber="376" value="1936-05-22">22 May 1936</date>
.
</paragraph>
</subSubSection>
<subSubSection pageId="99" pageNumber="376" type="description">
<paragraph blockId="99.[119,749,1630,1742]" pageId="99" pageNumber="376">
<emphasis bold="true" box="[121,233,1630,1654]" pageId="99" pageNumber="376">Condition</emphasis>
. Cranium complete; right dentary missing angular process; left dentary complete. Study skin: only c.
<quantity box="[691,747,1659,1683]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="99" pageNumber="376" unit="cm" value="6.0">6 cm</quantity>
of proximal end of tail present (probably condition when captured), skin otherwise complete and in good condition.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5237896" ID-Zenodo-Dep="5237896" httpUri="https://zenodo.org/record/5237896/files/figure.png" pageId="99" pageNumber="376" startId="99.[794,857,1683,1705]" targetBox="[799,1414,161,1658]" targetPageId="99">
<paragraph blockId="99.[794,1422,1683,1731]" pageId="99" pageNumber="376">
Figure 45. AM M.6102, holotype skull of
<taxonomicName authority="Troughton, 1937 a" authorityName="Troughton" authorityYear="1937" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="subSpecies" species="mordax" subSpecies="hageni">
<emphasis box="[1209,1422,1683,1705]" italics="true" pageId="99" pageNumber="376">Rattus mordax hageni</emphasis>
<bibRefCitation author="Troughton, E" box="[794,968,1709,1731]" pageId="99" pageNumber="376" pagination="117 - 127" refId="ref147573" refString="Troughton, E. L. G. 1937 a. Descriptions of some New Guinea mammals. Records of the Australian Museum 20 (2): 117 - 127. https: // doi. org / 10.3853 / j. 0067 - 1975.20.1937.254" type="journal article" year="1937">Troughton, 1937a</bibRefCitation>
</taxonomicName>
. (Photography by Stuart Humphreys).
</paragraph>
</caption>
<subSubSection pageId="99" pageNumber="376" type="distribution">
<paragraph blockId="99.[121,749,1758,1841]" pageId="99" pageNumber="376">
<emphasis bold="true" box="[121,277,1758,1782]" pageId="99" pageNumber="376">Type locality</emphasis>
.
<materialsCitation country="Papua New Guinea" elevation="915" location="Upper Wahgi River valley" municipality="Mt Hagen" pageId="99" pageNumber="376" specimenCount="1" stateProvince="Western Highlands">
<location LSID="urn:lsid:plazi:treatment:03DD87C8FFDA73561887FB64FB0790F3:8EAB6005FFDA73561915F934FD8291CA" box="[295,599,1758,1782]" country="Papua New Guinea" municipality="Mt Hagen" name="Upper Wahgi River valley" pageId="99" pageNumber="376" stateProvince="Western Highlands">Upper Wahgi River valley</location>
, from alt. of
<quantity box="[121,242,1787,1812]" metricMagnitude="2" metricUnit="m" metricValue="9.15162" metricValueMax="18.288" metricValueMin="0.01524" pageId="99" pageNumber="376" unit="ft" value="3002.5" valueMax="6000.0" valueMin="5.0">
<elevation box="[121,242,1787,1812]" metricMagnitude="2" metricUnit="m" metricValue="9.15162" metricValueMax="18.288" metricValueMin="0.01524" pageId="99" pageNumber="376" unit="ft" value="3002.5" valueMax="6000.0" valueMin="5.0">56,000 ft.</elevation>
</quantity>
, on the southern slopes of
<collectingMunicipality box="[537,649,1788,1812]" pageId="99" pageNumber="376">Mt Hagen</collectingMunicipality>
,
<collectingRegion country="Papua New Guinea" name="Western Highlands" pageId="99" pageNumber="376">Western Highlands Province</collectingRegion>
,
<collectingCountry box="[347,557,1817,1841]" name="Papua New Guinea" pageId="99" pageNumber="376">Papua New Guinea</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="99" pageNumber="376" type="materials_examined">
<paragraph blockId="99.[119,749,1857,1999]" pageId="99" pageNumber="376">
<emphasis bold="true" box="[121,235,1857,1881]" pageId="99" pageNumber="376">
<typeStatus box="[121,235,1857,1881]" pageId="99" pageNumber="376">Paratypes</typeStatus>
</emphasis>
. (7, 6 by subsequent determination). Details as per
<typeStatus box="[161,257,1886,1910]" pageId="99" pageNumber="376">holotype</typeStatus>
, collection dates not given:
<emphasis bold="true" box="[542,628,1886,1910]" pageId="99" pageNumber="376">M.6103</emphasis>
by original designation, female (
<typeStatus box="[347,438,1916,1940]" pageId="99" pageNumber="376">allotype</typeStatus>
), skull, study skin;
<emphasis bold="true" box="[657,742,1915,1939]" pageId="99" pageNumber="376">M.6104</emphasis>
, female, skull, study skin;
<emphasis bold="true" box="[388,513,1945,1969]" pageId="99" pageNumber="376">M.610507</emphasis>
, all females, bodies in alc.;
<emphasis bold="true" box="[173,298,1974,1998]" pageId="99" pageNumber="376">M.610809</emphasis>
, both young males, bodies in alc.
</paragraph>
</subSubSection>
<subSubSection pageId="99" pageNumber="376" type="discussion">
<paragraph blockId="99.[792,1422,1799,2000]" pageId="99" pageNumber="376">
<emphasis bold="true" box="[794,917,1799,1823]" pageId="99" pageNumber="376">Comments</emphasis>
. Troughton gives registration numbers of the
<typeStatus box="[794,887,1829,1853]" pageId="99" pageNumber="376">holotype</typeStatus>
and
<typeStatus box="[939,1025,1829,1853]" pageId="99" pageNumber="376">allotype</typeStatus>
under his heading “Type specimens” but does not mention any
<typeStatus box="[1072,1273,1858,1882]" isOtherTypeStatus="true" pageId="99" pageNumber="376">additional material</typeStatus>
of this taxon, types or otherwise.A further
<specimenCount box="[1094,1242,1887,1911]" pageId="99" pageNumber="376" type="generic">six specimens</specimenCount>
, all from the type locality and registered at the same time as the
<typeStatus box="[1281,1377,1917,1941]" pageId="99" pageNumber="376">holotype</typeStatus>
, are marked as
<typeStatus box="[907,1009,1946,1970]" pageId="99" pageNumber="376">paratypes</typeStatus>
in the register in Troughtons hand and we accept these as part of the type series.
</paragraph>
</subSubSection>
</treatment>
</document>