219 lines
17 KiB
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219 lines
17 KiB
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<document ID-DOI="10.3853/j.2201-4349.69.2017.1653" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237800" ID-ZooBank="68F315FF-3FEB-410E-96EC-5F494510F440" approvalRequired="65" approvalRequired_for_taxonomicNames="63" approvalRequired_for_treatments="2" checkinTime="1629535407606" checkinUser="felipe" docAuthor="Parnaby, Harry E., Ingleby, Sandy & Divljan, Anja" docDate="2017" docId="03DD87C8FFDA73561887FB64FB0790F3" docLanguage="en" docName="RecAustMus.69.5.277-420.pdf" docOrigin="Records of the Australian Museum 69 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Rattus mordax subsp. hageni Troughton 1937" docType="treatment" docVersion="4" lastPageNumber="376" masterDocId="FFE4FFB0FFB973351832FFEBFFD5973C" masterDocTitle="Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney" masterLastPageNumber="420" masterPageNumber="277" pageNumber="376" updateTime="1674245310755" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Parnaby, Harry E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ingleby, Sandy</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Divljan, Anja</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2017-10-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>69</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>277</mods:start>
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<mods:end>420</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3853/j.2201-4349.69.2017.1653</mods:identifier>
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<mods:identifier type="ISSN">2201-4349</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5237800</mods:identifier>
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<mods:identifier type="ZooBank">68F315FF-3FEB-410E-96EC-5F494510F440</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03DD87C8FFDA73561887FB64FB0790F3" httpUri="http://treatment.plazi.org/id/03DD87C8FFDA73561887FB64FB0790F3" lastPageNumber="376" pageId="99" pageNumber="376">
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<subSubSection box="[181,689,1167,1194]" pageId="99" pageNumber="376" type="nomenclature">
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<paragraph blockId="99.[181,689,1167,1194]" box="[181,689,1167,1194]" pageId="99" pageNumber="376">
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<heading box="[181,689,1167,1194]" centered="true" fontSize="11" level="2" pageId="99" pageNumber="376" reason="2">
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<taxonomicName authority="Troughton, 1937 a" authorityName="Troughton" authorityYear="1937" box="[181,689,1167,1194]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="subSpecies" species="mordax" subSpecies="hageni">
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<emphasis bold="true" box="[181,689,1167,1194]" pageId="99" pageNumber="376">
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<emphasis bold="true" box="[181,456,1167,1194]" italics="true" pageId="99" pageNumber="376">Rattus mordax hageni</emphasis>
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<bibRefCitation author="Troughton, E" box="[463,689,1167,1193]" pageId="99" pageNumber="376" pagination="117 - 127" refId="ref147573" refString="Troughton, E. L. G. 1937 a. Descriptions of some New Guinea mammals. Records of the Australian Museum 20 (2): 117 - 127. https: // doi. org / 10.3853 / j. 0067 - 1975.20.1937.254" type="journal article" year="1937">Troughton, 1937a</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,594,1208,1230]" pageId="99" pageNumber="376" type="reference_group">
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<paragraph blockId="99.[151,594,1208,1230]" box="[151,594,1208,1230]" pageId="99" pageNumber="376">
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<emphasis box="[151,302,1208,1230]" italics="true" pageId="99" pageNumber="376">Rec. Aust. Mus.</emphasis>
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20(2): 120. (
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<date box="[426,583,1208,1230]" pageId="99" pageNumber="376" value="1937-08-27">27 August 1937</date>
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).
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</paragraph>
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</subSubSection>
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<subSubSection box="[121,573,1246,1270]" pageId="99" pageNumber="376" type="vernacular_names">
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<paragraph blockId="99.[121,573,1246,1270]" box="[121,573,1246,1270]" pageId="99" pageNumber="376">
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<emphasis bold="true" box="[121,295,1246,1270]" pageId="99" pageNumber="376">Common name</emphasis>
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. Stein’s New
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<collectingCountry box="[445,522,1246,1270]" name="Guinea-Bissau" pageId="99" pageNumber="376">Guinea</collectingCountry>
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Rat.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="99" pageNumber="376" type="description">
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<paragraph blockId="99.[119,749,1286,1516]" pageId="99" pageNumber="376">
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<emphasis bold="true" box="[121,291,1286,1310]" pageId="99" pageNumber="376">Current name</emphasis>
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.
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<taxonomicName authority="Troughton 1937 a" authorityName="Troughton" authorityYear="1937" box="[309,744,1286,1310]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="subSpecies" species="steini" subSpecies="hageni">
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<emphasis box="[309,536,1286,1310]" italics="true" pageId="99" pageNumber="376">Rattus steini hageni</emphasis>
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<bibRefCitation author="Troughton, E" box="[547,744,1286,1310]" pageId="99" pageNumber="376" pagination="117 - 127" refId="ref147573" refString="Troughton, E. L. G. 1937 a. Descriptions of some New Guinea mammals. Records of the Australian Museum 20 (2): 117 - 127. https: // doi. org / 10.3853 / j. 0067 - 1975.20.1937.254" type="journal article" year="1937">Troughton 1937a</bibRefCitation>
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</taxonomicName>
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; following
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<bibRefCitation author="Taylor, J" box="[228,427,1316,1340]" pageId="99" pageNumber="376" pagination="177 - 336" refId="ref144998" refString="Taylor, J. M., J. H. Calaby, and H. M. Van Deusen. 1982. A revision of the genus Rattus (Rodentia, Muridae) in the New Guinean region. Bulletin of the American Museum of Natural History 173 (3): 177 - 336." type="journal article" year="1982">
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Taylor
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<emphasis box="[299,350,1316,1340]" italics="true" pageId="99" pageNumber="376">et al.</emphasis>
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(1982)
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</bibRefCitation>
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and Musser & Carleton (2005) but within
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<collectingCountry box="[244,460,1345,1369]" name="Papua New Guinea" pageId="99" pageNumber="376">Papua New Guinea</collectingCountry>
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<taxonomicName authority="Rummler, 1935" authorityName="Rummler" authorityYear="1935" box="[473,748,1345,1369]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="species" species="steini">
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<emphasis box="[473,565,1346,1369]" italics="true" pageId="99" pageNumber="376">R. steini</emphasis>
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<bibRefCitation author="Rummler, H" box="[576,748,1345,1369]" pageId="99" pageNumber="376" pagination="105 - 118" refId="ref143780" refString="Rummler, H. 1935. Neue Muriden aus Neuguinea. Zeitschrift fur Saugetierkunde 10: 105 - 118." type="journal article" year="1935">Rümmler, 1935</bibRefCitation>
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</taxonomicName>
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includes multiple species requiring resolution, see
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<bibRefCitation author="Robins, J & Tintinger, K & Aplin, M & Hingston, E & Matisoo-Smith, D" pageId="99" pageNumber="376" pagination="98002" refId="ref143702" refString="Robins, J. H., V. Tintinger, K. P. Aplin, M. Hingston, E. Matisoo-Smith, D. Penny, and S. D. Lavery. 2014. Phylogenetic species identification in Rattus Highlights rapid radiation and morphological similarity of New Guinean species. PLoS ONE 9 (5): e 98002. https: // doi. org / 10.1371 / journal. pone. 0098002" type="journal article" year="2014">
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Robins
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<emphasis italics="true" pageId="99" pageNumber="376">et al.</emphasis>
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(2014: 14)
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</bibRefCitation>
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.
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<bibRefCitation author="Helgen, K" box="[280,435,1404,1428]" pageId="99" pageNumber="376" pagination="689 - 749" refId="ref132662" refString="Helgen, K. M. 2007. A taxonomic and geographic overview of the mammals of Papua. In The Ecology of Papua, ed. A. J. Marshall and B. M. Beehler, Ecology of Indonesia Series, Vol. VI, Part 1 pp. 689 - 749. Singapore: Periplus Editions." type="book chapter" year="2007">Helgen (2007)</bibRefCitation>
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regarded eastern populations of
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<taxonomicName authorityName="Rummler" authorityYear="1935" box="[151,239,1434,1457]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="species" species="steini">
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<emphasis box="[151,239,1434,1457]" italics="true" pageId="99" pageNumber="376">R. steini</emphasis>
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</taxonomicName>
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to be a distinct species referable to
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<taxonomicName authority="(Rummler, 1935)" baseAuthorityName="Rummler" baseAuthorityYear="1935" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="species" species="foersteri">
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<emphasis box="[628,749,1433,1457]" italics="true" pageId="99" pageNumber="376">R. foersteri</emphasis>
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(
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<bibRefCitation author="Rummler, H" box="[128,302,1462,1486]" pageId="99" pageNumber="376" pagination="105 - 118" refId="ref143780" refString="Rummler, H. 1935. Neue Muriden aus Neuguinea. Zeitschrift fur Saugetierkunde 10: 105 - 118." type="journal article" year="1935">Rümmler, 1935</bibRefCitation>
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)
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</taxonomicName>
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, and placed
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<taxonomicName authorityName="Troughton" authorityYear="1937" box="[449,522,1462,1486]" class="Mammalia" family="Muridae" genus="Melomys" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="species" species="hageni">
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<emphasis box="[449,522,1462,1486]" italics="true" pageId="99" pageNumber="376">hageni</emphasis>
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</taxonomicName>
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as a synonym of the latter.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="99" pageNumber="376" type="materials_examined">
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<paragraph blockId="99.[119,748,1531,1615]" pageId="99" pageNumber="376">
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<emphasis bold="true" box="[121,224,1532,1556]" pageId="99" pageNumber="376">
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<typeStatus box="[121,224,1532,1556]" pageId="99" pageNumber="376">Holotype</typeStatus>
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</emphasis>
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. M.6102 by original designation. Male adult, skull (
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<figureCitation box="[127,211,1561,1585]" captionStart="Figure 45" captionStartId="99.[794,857,1683,1705]" captionTargetBox="[799,1414,161,1658]" captionTargetId="figure-538@99.[794,1421,151,1669]" captionTargetPageId="99" captionText="Figure 45. AM M.6102, holotype skull of Rattus mordax hageni Troughton, 1937a. (Photography by Stuart Humphreys)." figureDoi="http://doi.org/10.5281/zenodo.5237896" httpUri="https://zenodo.org/record/5237896/files/figure.png" pageId="99" pageNumber="376">Fig. 45</figureCitation>
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), study skin, collected by Dr G. A. M. Heydon, collection date not given, registered
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<date box="[509,655,1590,1615]" pageId="99" pageNumber="376" value="1936-05-22">22 May 1936</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="99" pageNumber="376" type="description">
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<paragraph blockId="99.[119,749,1630,1742]" pageId="99" pageNumber="376">
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<emphasis bold="true" box="[121,233,1630,1654]" pageId="99" pageNumber="376">Condition</emphasis>
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. Cranium complete; right dentary missing angular process; left dentary complete. Study skin: only c.
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<quantity box="[691,747,1659,1683]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="99" pageNumber="376" unit="cm" value="6.0">6 cm</quantity>
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of proximal end of tail present (probably condition when captured), skin otherwise complete and in good condition.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5237896" ID-Zenodo-Dep="5237896" httpUri="https://zenodo.org/record/5237896/files/figure.png" pageId="99" pageNumber="376" startId="99.[794,857,1683,1705]" targetBox="[799,1414,161,1658]" targetPageId="99">
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<paragraph blockId="99.[794,1422,1683,1731]" pageId="99" pageNumber="376">
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Figure 45. AM M.6102, holotype skull of
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<taxonomicName authority="Troughton, 1937 a" authorityName="Troughton" authorityYear="1937" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="99" pageNumber="376" phylum="Chordata" rank="subSpecies" species="mordax" subSpecies="hageni">
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<emphasis box="[1209,1422,1683,1705]" italics="true" pageId="99" pageNumber="376">Rattus mordax hageni</emphasis>
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<bibRefCitation author="Troughton, E" box="[794,968,1709,1731]" pageId="99" pageNumber="376" pagination="117 - 127" refId="ref147573" refString="Troughton, E. L. G. 1937 a. Descriptions of some New Guinea mammals. Records of the Australian Museum 20 (2): 117 - 127. https: // doi. org / 10.3853 / j. 0067 - 1975.20.1937.254" type="journal article" year="1937">Troughton, 1937a</bibRefCitation>
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</taxonomicName>
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. (Photography by Stuart Humphreys).
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</paragraph>
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</caption>
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<subSubSection pageId="99" pageNumber="376" type="distribution">
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<paragraph blockId="99.[121,749,1758,1841]" pageId="99" pageNumber="376">
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<emphasis bold="true" box="[121,277,1758,1782]" pageId="99" pageNumber="376">Type locality</emphasis>
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.
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<materialsCitation country="Papua New Guinea" elevation="915" location="Upper Wahgi River valley" municipality="Mt Hagen" pageId="99" pageNumber="376" specimenCount="1" stateProvince="Western Highlands">
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<location LSID="urn:lsid:plazi:treatment:03DD87C8FFDA73561887FB64FB0790F3:8EAB6005FFDA73561915F934FD8291CA" box="[295,599,1758,1782]" country="Papua New Guinea" municipality="Mt Hagen" name="Upper Wahgi River valley" pageId="99" pageNumber="376" stateProvince="Western Highlands">Upper Wahgi River valley</location>
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, from alt. of
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<quantity box="[121,242,1787,1812]" metricMagnitude="2" metricUnit="m" metricValue="9.15162" metricValueMax="18.288" metricValueMin="0.01524" pageId="99" pageNumber="376" unit="ft" value="3002.5" valueMax="6000.0" valueMin="5.0">
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<elevation box="[121,242,1787,1812]" metricMagnitude="2" metricUnit="m" metricValue="9.15162" metricValueMax="18.288" metricValueMin="0.01524" pageId="99" pageNumber="376" unit="ft" value="3002.5" valueMax="6000.0" valueMin="5.0">5–6,000 ft.</elevation>
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</quantity>
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, on the southern slopes of
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<collectingMunicipality box="[537,649,1788,1812]" pageId="99" pageNumber="376">Mt Hagen</collectingMunicipality>
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,
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<collectingRegion country="Papua New Guinea" name="Western Highlands" pageId="99" pageNumber="376">Western Highlands Province</collectingRegion>
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,
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<collectingCountry box="[347,557,1817,1841]" name="Papua New Guinea" pageId="99" pageNumber="376">Papua New Guinea</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="99" pageNumber="376" type="materials_examined">
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<paragraph blockId="99.[119,749,1857,1999]" pageId="99" pageNumber="376">
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<emphasis bold="true" box="[121,235,1857,1881]" pageId="99" pageNumber="376">
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<typeStatus box="[121,235,1857,1881]" pageId="99" pageNumber="376">Paratypes</typeStatus>
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|
</emphasis>
|
|||
|
|
. (7, 6 by subsequent determination). Details as per
|
|||
|
|
<typeStatus box="[161,257,1886,1910]" pageId="99" pageNumber="376">holotype</typeStatus>
|
|||
|
|
, collection dates not given:
|
|||
|
|
<emphasis bold="true" box="[542,628,1886,1910]" pageId="99" pageNumber="376">M.6103</emphasis>
|
|||
|
|
by original designation, female (
|
|||
|
|
<typeStatus box="[347,438,1916,1940]" pageId="99" pageNumber="376">allotype</typeStatus>
|
|||
|
|
), skull, study skin;
|
|||
|
|
<emphasis bold="true" box="[657,742,1915,1939]" pageId="99" pageNumber="376">M.6104</emphasis>
|
|||
|
|
, female, skull, study skin;
|
|||
|
|
<emphasis bold="true" box="[388,513,1945,1969]" pageId="99" pageNumber="376">M.6105–07</emphasis>
|
|||
|
|
, all females, bodies in alc.;
|
|||
|
|
<emphasis bold="true" box="[173,298,1974,1998]" pageId="99" pageNumber="376">M.6108–09</emphasis>
|
|||
|
|
, both young males, bodies in alc.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="99" pageNumber="376" type="discussion">
|
|||
|
|
<paragraph blockId="99.[792,1422,1799,2000]" pageId="99" pageNumber="376">
|
|||
|
|
<emphasis bold="true" box="[794,917,1799,1823]" pageId="99" pageNumber="376">Comments</emphasis>
|
|||
|
|
. Troughton gives registration numbers of the
|
|||
|
|
<typeStatus box="[794,887,1829,1853]" pageId="99" pageNumber="376">holotype</typeStatus>
|
|||
|
|
and
|
|||
|
|
<typeStatus box="[939,1025,1829,1853]" pageId="99" pageNumber="376">allotype</typeStatus>
|
|||
|
|
under his heading “Type specimens” but does not mention any
|
|||
|
|
<typeStatus box="[1072,1273,1858,1882]" isOtherTypeStatus="true" pageId="99" pageNumber="376">additional material</typeStatus>
|
|||
|
|
of this taxon, types or otherwise.A further
|
|||
|
|
<specimenCount box="[1094,1242,1887,1911]" pageId="99" pageNumber="376" type="generic">six specimens</specimenCount>
|
|||
|
|
, all from the type locality and registered at the same time as the
|
|||
|
|
<typeStatus box="[1281,1377,1917,1941]" pageId="99" pageNumber="376">holotype</typeStatus>
|
|||
|
|
, are marked as
|
|||
|
|
<typeStatus box="[907,1009,1946,1970]" pageId="99" pageNumber="376">paratypes</typeStatus>
|
|||
|
|
in the register in Troughton’s hand and we accept these as part of the type series.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|