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<document id="0CD3B96A82BAD5FEAE76D724CC382CF0" ID-CLB-Dataset="25467" ID-DOI="10.11646/zootaxa.4668.4.4" ID-GBIF-Dataset="f463aada-9f40-4254-84a5-3f86928f35d3" ID-ISSN="1175-5326" ID-Zenodo-Dep="3449886" ID-ZooBank="2FC5E81B-4DC4-491A-B133-28FC7766009B" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568898096991" checkinUser="plazi" docAuthor="Marques, Mariana P., Ceríaco, Luis M. P., Stanley, Edward L., Bandeira, Suzana A., Agarwal, Ishan &amp; Bauer, Aaron M." docDate="2019" docId="BE288794FFD9FF93F4DBFDF581E2FCC8" docLanguage="en" docName="zootaxa.4668.4.4.pdf" docOrigin="Zootaxa 4668 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cordylus phonolithos Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer 2019, sp. nov." docType="treatment" docVersion="15" lastPageNumber="518" masterDocId="4211FFECFFDFFF9CF44CFF988233FFF3" masterDocTitle="A new species of Girdled Lizard (Squamata: Cordylidae) from the Serra da Neve Inselberg, Namibe Province, southwestern Angola" masterLastPageNumber="524" masterPageNumber="503" pageNumber="509" updateTime="1698755778065" updateUser="ExternalLinkService">
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<mods:title id="D02818AFD9F26558B3AF607289C636AB">A new species of Girdled Lizard (Squamata: Cordylidae) from the Serra da Neve Inselberg, Namibe Province, southwestern Angola</mods:title>
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<mods:namePart id="7D8D6BA226D1F365AF9E7125CD12E4C7">Marques, Mariana P.</mods:namePart>
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<mods:namePart id="B58C3A5829A9ADE413C1AC586825B088">Ceríaco, Luis M. P.</mods:namePart>
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<mods:namePart id="AF1BF9F45F4EF7C6229F95B7A04E2109">Stanley, Edward L.</mods:namePart>
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<mods:namePart id="39E1AAF55001848168EB56227726F074">Bandeira, Suzana A.</mods:namePart>
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<mods:namePart id="70510A48A0142C6C3741ABDDDBD5848A">Agarwal, Ishan</mods:namePart>
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<mods:namePart id="7EC0B35EBD979A2980DFDBC04962D01D">Bauer, Aaron M.</mods:namePart>
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<paragraph id="363E3682FFD9FF9AF4DBFDF58033FD74" blockId="6.[151,1012,621,719]" box="[151,512,621,648]" pageId="6" pageNumber="509">
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<taxonomicName id="F1814D01FFD9FF9AF4DBFDF583A8FD7B" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[151,411,621,648]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="509" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD9FF9AF4DBFDF583A8FD7B" bold="true" box="[151,411,621,648]" italics="true" pageId="6" pageNumber="509">Cordylus phonolithos</emphasis>
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<taxonomicNameLabel id="1FC657EBFFD9FF9AF5EFFDF58033FD74" box="[419,512,621,647]" pageId="6" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="363E3682FFD9FF9AF4DBFD0983BCFD58" blockId="6.[151,1012,621,719]" box="[151,399,657,683]" pageId="6" pageNumber="509">
(
<tableCitation id="7B030339FFD9FF9AF4D3FD09832EFD58" box="[159,285,657,683]" captionStart-0="Table 1" captionStart-1="TABLE 2" captionStartId-0="13.[151,214,504,530]" captionStartId-1="14.[151,239,513,539]" captionTargetBox-0="[160,1890,599,1721]" captionTargetId-0="graphics@13.[151,1961,796,1731]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Table 1. Uncorrected pairwise distances between cordylid taxa for the ND2 mitochondrial gene. All values are presented in percentages (%)." captionText-1="TABLE 2. Variation in meristic counts for specimens of Cordylus phonolithos sp. nov., C. namakuiyus, C. machadoi and C. angolensis. All measurements are presented in millimeters (mm). Data of C. namakuiyus retrieved from Stanley et al. (2016); data for C. machadoi retrieved from Stanley et al. (2016) and two additional specimens from this study; data for C. phonolithos sp. nov. and C. angolensis collected in this study" httpUri-0="http://table.plazi.org/id/62FE660AFFD2FF91F4DBFE608446FDE1" httpUri-1="http://table.plazi.org/id/62FE660AFFD1FF92F4DBFD998196FD90" pageId="6" pageNumber="509" tableUuid-0="62FE660AFFD2FF91F4DBFE608446FDE1" tableUuid-1="62FE660AFFD1FF92F4DBFD998196FD90">Tables 12</tableCitation>
;
<figureCitation id="AEBA2A07FFD9FF9AF565FD0983B4FD58" box="[297,391,657,683]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStartId-0="6.[151,250,1938,1963]" captionStartId-1="7.[151,250,1682,1707]" captionStartId-2="8.[151,250,1874,1899]" captionStartId-3="9.[151,250,1868,1893]" captionStartId-4="10.[151,250,1876,1901]" captionTargetBox-0="[151,1436,944,1918]" captionTargetBox-1="[154,1431,178,1660]" captionTargetBox-2="[151,1436,1237,1855]" captionTargetBox-3="[151,1436,805,1848]" captionTargetBox-4="[163,1424,813,1851]" captionTargetId-0="figure@6.[151,1436,944,1918]" captionTargetId-1="figure@7.[151,1436,174,1663]" captionTargetId-2="figure@8.[151,1436,1237,1855]" captionTargetId-3="figure@9.[151,1436,805,1848]" captionTargetId-4="figure@10.[163,1424,813,1851]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="FIGURE 4. Dorsal and ventral whole-body views (left) and Dorsal, lateral, and ventral views of the head of the holotype of Cordylus phonolithos sp. nov. (CAS 263581)." captionText-1="FIGURE 5. Cordylus phonolithos sp. nov. in life (A. Holotype CAS 263581; B. Paratype INBAC:AMB 10272). NDolondolo, Serra da Neve, southwestern Angola (Photos by Luis Ceríaco)." captionText-2="FIGURE 6. Comparison of head scalation between C. machadoi, C. phonolithos sp. nov., C. namakuiyus and C. angolensis." captionText-3="FIGURE 7. Figure showing Cranial and postcranial osteology of CAS 263581 Cordylus phonolithos sp. nov. holotype. An= angular, Art = articular, Bo = Basioccipital, Bpt= Basipterygoid, Co = coronoid, De = dentary, Ec = ectopterygoid, Ep = epipterygoid, F = frontal, Hi = hyperischium, Ho = hypoischium, J = jugal, Mx = maxilla, N = nasal, P = parietal, Pa = Palatine, Pf = postfrontal, Pmx= premaxilla, Po = Postorbital, Pocc = paraoccipital, Pp = prepubic cartilage, Ppmp = posteriomedial process of the parietal, Prf = prefrontal, Pro = prootic, Pt = pterygoid, Q = quadrate, Sq = squamosal, St = supratemporal, Vo = vomer." captionText-4="FIGURE 8. Cranial morphology and osteodermal variation on the different Angolan Cordylus species, colorized to show crosssectional thickness." figureDoi-0="http://doi.org/10.5281/zenodo.3449900" figureDoi-1="http://doi.org/10.5281/zenodo.3449902" figureDoi-2="http://doi.org/10.5281/zenodo.3449910" figureDoi-3="http://doi.org/10.5281/zenodo.3449916" figureDoi-4="http://doi.org/10.5281/zenodo.3449920" httpUri-0="https://zenodo.org/record/3449900/files/figure.png" httpUri-1="https://zenodo.org/record/3449902/files/figure.png" httpUri-2="https://zenodo.org/record/3449910/files/figure.png" httpUri-3="https://zenodo.org/record/3449916/files/figure.png" httpUri-4="https://zenodo.org/record/3449920/files/figure.png" pageId="6" pageNumber="509">Fig. 48</figureCitation>
)
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lsid:
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urn:lsid:zoobank.org:act:
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<paragraph id="363E3682FFD9FF9AF4DBFD62804FFCE2" blockId="6.[151,636,761,786]" box="[151,636,761,786]" pageId="6" pageNumber="509">
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<taxonomicName id="F1814D01FFD9FF9AF4DBFD62831DFCE2" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[151,302,761,785]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="509" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD9FF9AF4DBFD62831DFCE2" box="[151,302,761,785]" italics="true" pageId="6" pageNumber="509">C. namakuiyus</emphasis>
</taxonomicName>
[part]:
<treatmentCitation id="B7201093FFD9FF9AF536FD61804FFCE2" ID-DOI="http://doi.org/10.5281/zenodo.3506980" ID-Zenodo-Dep="3506980" author="Stanley, E. L. &amp; Ceriaco, L. M. P. &amp; Bandeira, S. &amp; Valerio, H. &amp; Bates, M. F. &amp; Branch, W. R." box="[378,636,761,786]" httpUri="http://treatment.plazi.org/id/03D78791FF88424ABC8402C4BEA40CDE" page="209" pageId="6" pageNumber="509" year="2016">
<bibRefCitation id="52104B73FFD9FF9AF536FD61804FFCE2" author="Stanley, E. L. &amp; Ceriaco, L. M. P. &amp; Bandeira, S. &amp; Valerio, H. &amp; Bates, M. F. &amp; Branch, W. R." box="[378,636,761,786]" pageId="6" pageNumber="509" pagination="201 - 226" refId="ref12195" refString="Stanley, E. L., Ceriaco, L. M. P., Bandeira, S., Valerio, H., Bates, M. F. &amp; Branch, W. R. (2016) A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola with the description of a new species from the Pro-Namib desert. Zootaxa, 4061 (3), 201 - 226. https: // doi. org / 10.11646 / zootaxa. 4061.3.1" type="journal article" year="2016">
Stanley
<emphasis id="04F5EA90FFD9FF9AF580FD6283CCFCE2" box="[460,511,761,785]" italics="true" pageId="6" pageNumber="509">et al.</emphasis>
(2016: 209)
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</paragraph>
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<paragraph id="363E3682FFD9FF9AF4DBFCD88685FC52" blockId="6.[151,1436,831,930]" pageId="6" pageNumber="509">
<materialsCitation id="86E93CDFFFD9FF9AF4DBFCD88685FC52" ID-GBIF-Occurrence="2416712413" collectingDate="2016-11-21" collectionCode="AMB" collectorName="L. Ceriaco &amp; Ceriaco, S &amp; I. Agarwal" country="Angola" elevation="752" latitude="13.80678" location="N'Dolondolo" longLatPrecision="1" longitude="13.13507" pageId="6" pageNumber="509" specimenCode="CAS 263581" specimenCount="1" stateProvince="Namibe" typeStatus="holotype">
<emphasis id="04F5EA90FFD9FF9AF4DBFCD88338FCA9" bold="true" box="[151,267,832,858]" pageId="6" pageNumber="509">
<typeStatus id="E93A8820FFD9FF9AF4DBFCD88335FCA9" box="[151,262,832,858]" pageId="6" pageNumber="509" type="holotype">Holotype</typeStatus>
.
</emphasis>
A subadult male (
<specimenCode id="66279EF9FFD9FF9AF590FCD88047FCA9" box="[476,628,831,858]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" pageNumber="509" type="Museum">CAS 263581</specimenCode>
, field number
<collectionCode id="5090AE47FFD9FF9AF754FCD8816BFCA9" box="[792,856,832,858]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="6" pageNumber="509" type="Museum">AMB</collectionCode>
10296;
<figureCitation id="AEBA2A07FFD9FF9AF7FBFCD88606FCA9" box="[951,1077,831,858]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="6.[151,250,1938,1963]" captionStartId-1="7.[151,250,1682,1707]" captionTargetBox-0="[151,1436,944,1918]" captionTargetBox-1="[154,1431,178,1660]" captionTargetId-0="figure@6.[151,1436,944,1918]" captionTargetId-1="figure@7.[151,1436,174,1663]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 4. Dorsal and ventral whole-body views (left) and Dorsal, lateral, and ventral views of the head of the holotype of Cordylus phonolithos sp. nov. (CAS 263581)." captionText-1="FIGURE 5. Cordylus phonolithos sp. nov. in life (A. Holotype CAS 263581; B. Paratype INBAC:AMB 10272). NDolondolo, Serra da Neve, southwestern Angola (Photos by Luis Ceríaco)." figureDoi-0="http://doi.org/10.5281/zenodo.3449900" figureDoi-1="http://doi.org/10.5281/zenodo.3449902" httpUri-0="https://zenodo.org/record/3449900/files/figure.png" httpUri-1="https://zenodo.org/record/3449902/files/figure.png" pageId="6" pageNumber="509">Figs. 45A</figureCitation>
) collected within a crevice in a granite boulder in the vicinity of
<location id="335E6059FFD9FF9AF65EFCFC8094FC8D" LSID="urn:lsid:plazi:treatment:BE288794FFD9FF93F4DBFDF581E2FCC8:335E6059FFD9FF9AF65EFCFC8094FC8D" box="[530,679,868,894]" country="Angola" latitude="13.80678" longLatPrecision="1" longitude="13.13507" name="N'Dolondolo" pageId="6" pageNumber="509" stateProvince="Namibe">NDolondolo</location>
(
<figureCitation id="AEBA2A07FFD9FF9AF6F4FCFC813AFC8D" box="[696,777,868,894]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1928]" captionTargetBox="[166,1424,468,1872]" captionTargetId="figure@4.[151,1434,481,1883]" captionTargetPageId="4" captionText="FIGURE 2. Map showing localities for Cordylus species in Angola. Yellow circle—C. angolensis (type locality: Caconda); Red circle—C. phonolithos sp. nov, (type locality: Serra da Neve); Blue circles—C. machadoi (type locality: Serra da Leba); Pink circles—C. namakuiyus (type locality: Caraculo). Major cities are represented by an orange triangle. Principal plotted localities include both bibliographic (Marques et al. 2018) and museum records." figureDoi="http://doi.org/10.5281/zenodo.3449896" httpUri="https://zenodo.org/record/3449896/files/figure.png" pageId="6" pageNumber="509">Figs. 2</figureCitation>
,
<figureCitation id="AEBA2A07FFD9FF9AF75AFCFC8115FC8D" box="[790,806,868,894]" captionStart="FIGURE 9" captionStartId="11.[151,250,1747,1772]" captionTargetBox="[203,1384,181,1722]" captionTargetId="figure@11.[203,1384,181,1722]" captionTargetPageId="11" captionText="FIGURE 9. Whole body osteodermal variation on the different Angolan Cordylus species, colorized to show cross-sectional thickness." figureDoi="http://doi.org/10.5281/zenodo.3449924" httpUri="https://zenodo.org/record/3449924/files/figure.png" pageId="6" pageNumber="509">9</figureCitation>
),
<collectingRegion id="F445F860FFD9FF9AF771FCFC8637FC8D" box="[829,1028,868,894]" country="Angola" name="Namibe" pageId="6" pageNumber="509">Namibe Province</collectingRegion>
,
<collectingCountry id="4E967612FFD9FF9AF05CFCFC8651FC8D" box="[1040,1122,868,894]" name="Angola" pageId="6" pageNumber="509">Angola</collectingCountry>
[-
<geoCoordinate id="53B55045FFD9FF9AF034FCFB8735FC8E" box="[1144,1286,867,893]" degrees="13.80678" direction="north" orientation="latitude" pageId="6" pageNumber="509" precision="1" value="13.80678">13.80678ºN</geoCoordinate>
,
<geoCoordinate id="53B55045FFD9FF9AF15EFCFB87ABFC8D" box="[1298,1432,867,894]" degrees="13.13507" direction="east" orientation="longitude" pageId="6" pageNumber="509" precision="1" value="13.13507">13.13507ºE</geoCoordinate>
,
<quantity id="F1799B67FFD9FF9AF4DBFC1F82EDFC51" box="[151,222,903,930]" metricMagnitude="2" metricUnit="m" metricValue="7.52" pageId="6" pageNumber="509" unit="m" value="752.0">
<elevation id="BDACD1B1FFD9FF9AF4DBFC1F82EDFC51" box="[151,222,903,930]" metricMagnitude="2" metricUnit="m" metricValue="7.52" pageId="6" pageNumber="509" unit="m" value="752.0">752 m</elevation>
</quantity>
elevation], collected by
<collectorName id="9B745354FFD9FF9AF5BEFC108058FC51" box="[498,619,904,930]" pageId="6" pageNumber="509">L. Ceríaco</collectorName>
, S. Bandeira and
<collectorName id="9B745354FFD9FF9AF763FC10819BFC51" box="[815,936,904,930]" pageId="6" pageNumber="509">I. Agarwal</collectorName>
, on
<date id="423F1042FFD9FF9AF79AFC108680FC52" box="[982,1203,903,930]" pageId="6" pageNumber="509" value="2016-11-21">
<collectingDate id="527BE9AAFFD9FF9AF79AFC108680FC52" box="[982,1203,903,930]" pageId="6" pageNumber="509" value="2016-11-21">21 November 2016</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="62FE660AFFD9FF9AF4DBF80A8059F83C" ID-DOI="http://doi.org/10.5281/zenodo.3449900" ID-Zenodo-Dep="3449900" httpUri="https://zenodo.org/record/3449900/files/figure.png" pageId="6" pageNumber="509" startId="6.[151,250,1938,1963]" targetBox="[151,1436,944,1918]" targetPageId="6">
<paragraph id="363E3682FFD9FF9AF4DBF80A8059F83C" blockId="6.[151,1436,1938,1999]" pageId="6" pageNumber="509">
<emphasis id="04F5EA90FFD9FF9AF4DBF80A833DF858" bold="true" box="[151,270,1938,1963]" pageId="6" pageNumber="509">FIGURE 4</emphasis>
. Dorsal and ventral whole-body views (left) and Dorsal, lateral, and ventral views of the head of the holotype of
<taxonomicName id="F1814D01FFD9FF9AF4DBF82E8342F83D" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[151,369,1974,1998]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="509" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD9FF9AF4DBF82E8342F83D" box="[151,369,1974,1998]" italics="true" pageId="6" pageNumber="509">Cordylus phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD9FF9AF53BF82E83FBF83D" bold="true" box="[375,456,1974,1998]" pageId="6" pageNumber="509">
<taxonomicNameLabel id="1FC657EBFFD9FF9AF53BF82E83FBF83D" box="[375,456,1974,1998]" pageId="6" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CAS 263581).
</paragraph>
</caption>
<caption id="62FE660AFFD8FF9BF4DBF90A8127F93C" ID-DOI="http://doi.org/10.5281/zenodo.3449902" ID-Zenodo-Dep="3449902" httpUri="https://zenodo.org/record/3449902/files/figure.png" pageId="7" pageNumber="510" startId="7.[151,250,1682,1707]" targetBox="[154,1431,178,1660]" targetPageId="7">
<paragraph id="363E3682FFD8FF9BF4DBF90A8127F93C" blockId="7.[151,1436,1682,1743]" pageId="7" pageNumber="510">
<emphasis id="04F5EA90FFD8FF9BF4DBF90A833FF958" bold="true" box="[151,268,1682,1707]" pageId="7" pageNumber="510">FIGURE 5</emphasis>
.
<taxonomicName id="F1814D01FFD8FF9BF554F90B83C2F958" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[280,497,1683,1707]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="510" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD8FF9BF554F90B83C2F958" box="[280,497,1683,1707]" italics="true" pageId="7" pageNumber="510">Cordylus phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD8FF9BF5BBF90B8075F958" bold="true" box="[503,582,1683,1707]" pageId="7" pageNumber="510">
<taxonomicNameLabel id="1FC657EBFFD8FF9BF5BBF90B8075F958" box="[503,582,1683,1707]" pageId="7" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life (
<emphasis id="04F5EA90FFD8FF9BF6D9F90A809DF958" bold="true" box="[661,686,1682,1707]" pageId="7" pageNumber="510">A.</emphasis>
Holotype CAS 263581;
<emphasis id="04F5EA90FFD8FF9BF7EBF90B818DF958" bold="true" box="[935,958,1683,1707]" pageId="7" pageNumber="510">B.</emphasis>
Paratype INBAC: AMB 10272). NDolondolo, Serra da Neve, southwestern Angola (Photos by Luis Ceríaco).
</paragraph>
</caption>
<paragraph id="363E3682FFD8FF9BF48BF8BA87AEF8AC" blockId="7.[151,1437,1825,2032]" pageId="7" pageNumber="510">
<materialsCitation id="86E93CDFFFD8FF9BF48BF8BA87AAF8AC" ID-GBIF-Occurrence="2416712542" collectingDate="2016-11-20" collectionCode="INBAC, AMB" collectorName="L. Ceriaco &amp; Ceriaco, S &amp; I. Agarwal" country="Angola" elevation="752" latitude="13.80678" location="N'Dolondolo" longLatPrecision="1" longitude="13.13507" pageId="7" pageNumber="510" specimenCount="1" stateProvince="Namibe" typeStatus="paratype">
<emphasis id="04F5EA90FFD8FF9BF48BF8BA8308F8CF" bold="true" box="[199,315,1826,1852]" pageId="7" pageNumber="510">
<typeStatus id="E93A8820FFD8FF9BF48BF8BA8305F8CF" box="[199,310,1826,1852]" pageId="7" pageNumber="510" type="paratype">Paratype</typeStatus>
.
</emphasis>
A juvenile specimen (
<collectionCode id="5090AE47FFD8FF9BF67BF8BA80A7F8CF" box="[567,660,1826,1852]" pageId="7" pageNumber="510">INBAC</collectionCode>
:
<collectionCode id="5090AE47FFD8FF9BF6D2F8BA80EDF8CF" box="[670,734,1826,1852]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="7" pageNumber="510" type="Museum">AMB</collectionCode>
10272 [same as field number];
<figureCitation id="AEBA2A07FFD8FF9BF008F8B986AFF8CF" box="[1092,1180,1825,1852]" captionStart="FIGURE 5" captionStartId="7.[151,250,1682,1707]" captionTargetBox="[154,1431,178,1660]" captionTargetId="figure@7.[151,1436,174,1663]" captionTargetPageId="7" captionText="FIGURE 5. Cordylus phonolithos sp. nov. in life (A. Holotype CAS 263581; B. Paratype INBAC:AMB 10272). NDolondolo, Serra da Neve, southwestern Angola (Photos by Luis Ceríaco)." figureDoi="http://doi.org/10.5281/zenodo.3449902" httpUri="https://zenodo.org/record/3449902/files/figure.png" pageId="7" pageNumber="510">Fig. 5B</figureCitation>
), collected at the same locality as the holotype, under a small granite rock on a larger rock, by the same collectors, on
<date id="423F1042FFD8FF9BF0F1F8DE87AAF8AC" box="[1213,1433,1861,1888]" pageId="7" pageNumber="510" value="2016-11-20">
<collectingDate id="527BE9AAFFD8FF9BF0F1F8DE87AAF8AC" box="[1213,1433,1861,1888]" pageId="7" pageNumber="510" value="2016-11-20">20 November 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7E9B6509FFD8FF94F48BF8F18129FB34" lastPageId="8" lastPageNumber="511" pageId="7" pageNumber="510" type="diagnosis">
<paragraph id="363E3682FFD8FF94F48BF8F18129FB34" blockId="7.[151,1437,1825,2032]" lastBlockId="8.[151,1437,153,1224]" lastPageId="8" lastPageNumber="511" pageId="7" pageNumber="510">
<emphasis id="04F5EA90FFD8FF9BF48BF8F18371F877" bold="true" box="[199,322,1897,1924]" pageId="7" pageNumber="510">Diagnosis.</emphasis>
A medium sized
<taxonomicName id="F1814D01FFD8FF9BF644F8F1805DF870" box="[520,622,1897,1923]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="04F5EA90FFD8FF9BF644F8F1805DF870" box="[520,622,1897,1923]" italics="true" pageId="7" pageNumber="510">Cordylus</emphasis>
</taxonomicName>
species, identified to genus by the following combination of characters: fully limbed, strongly depressed triangular head and body, osteoderms present, rhomboidal, imbricate and keeled dorsal scales present, occipitals non-spinose, and spinose caudal and limb scales enlarged (
<bibRefCitation id="52104B73FFD8FF9BF0D8F829871AF83F" author="Branch, W. R." box="[1172,1321,1969,1996]" pageId="7" pageNumber="510" refId="ref9221" refString="Branch, W. R. (1998) Field Guide to Snakes and Other Reptiles of Southern Africa, 3 rd edition. Ralph Curtis Books, Sanibel Island, Florida, 399 pp." type="book" year="1998">Branch 1998</bibRefCitation>
;
<bibRefCitation id="52104B73FFD8FF9BF179F8298376F803" author="Broadley, D. G. &amp; Branch, W. R." pageId="7" pageNumber="510" pagination="9 - 34" refId="ref9678" refString="Broadley, D. G. &amp; Branch, W. R. (2002) A review of the small east African Cordylus (Sauria: Cordylidae), with the description of a new species. African Journal of Herpetology, 51, 9 - 34. https: // doi. org / 10.1080 / 21564574.2002.9635459" type="journal article" year="2002">Broadley &amp; Branch 2002</bibRefCitation>
;
<bibRefCitation id="52104B73FFD8FF9BF503F84D8012F803" author="Stanley, E. L. &amp; Bauer, A. M. &amp; Jackman, T. R. &amp; Branch, W. R. &amp; Mouton, P. le" box="[335,545,2005,2032]" pageId="7" pageNumber="510" pagination="53 - 70" refId="ref12109" refString="Stanley, E. L., Bauer, A. M., Jackman, T. R., Branch, W. R. &amp; Mouton, P. le F. N. (2011) Between a rock and a hard polytomy: Rapid radiation in the rupicolous girdled lizards (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58, 53 - 70. https: // doi. org / 10.1016 / j. ympev. 2010.08.024" type="journal article" year="2011">
Stanley
<emphasis id="04F5EA90FFD8FF9BF5E5F84E83D3F81C" box="[425,480,2005,2031]" italics="true" pageId="7" pageNumber="510">et al.</emphasis>
2011
</bibRefCitation>
).
<taxonomicName id="F1814D01FFD8FF9BF67EF84D8111F81C" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[562,802,2005,2031]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="510" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD8FF9BF67EF84D8111F81C" box="[562,802,2005,2031]" italics="true" pageId="7" pageNumber="510">Cordylus phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD8FF9BF764F84E81B3F803" bold="true" box="[808,896,2006,2032]" pageId="7" pageNumber="510">
<taxonomicNameLabel id="1FC657EBFFD8FF9BF764F84E81B3F803" box="[808,896,2006,2032]" pageId="7" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other species in the genus except for
<taxonomicName id="F1814D01FFD7FF94F4F2FF018030FF40" authority="(Reichenow, 1887)" authorityName="Reichenow" authorityYear="1887" baseAuthorityName="Reichenow" baseAuthorityYear="1887" box="[190,515,153,179]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="vittifer">
<emphasis id="04F5EA90FFD7FF94F4F2FF01831AFF40" box="[190,297,153,179]" italics="true" pageId="8" pageNumber="511">C. vittifer</emphasis>
(Reichenow, 1887)
</taxonomicName>
,
<taxonomicName id="F1814D01FFD7FF94F65DFF0180ACFF40" authorityName="Laurent" authorityYear="1964" box="[529,671,153,179]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F65DFF0180ACFF40" box="[529,671,153,179]" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
and
<emphasis id="04F5EA90FFD7FF94F698FF0181B2FF40" box="[724,897,153,179]" italics="true" pageId="8" pageNumber="511">
<taxonomicName id="F1814D01FFD7FF94F698FF01814EFF40" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[724,893,153,179]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="namakuiyus">C. namakuiyus</taxonomicName>
,
</emphasis>
by the presence (
<emphasis id="04F5EA90FFD7FF94F008FF0286BFFF40" box="[1092,1164,154,179]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
absence) of a transverse row of elongated dorsal scales immediately posterior to occipitals (
<figureCitation id="AEBA2A07FFD7FF94F730FF25818CFF2B" box="[892,959,189,216]" captionStart="FIGURE 6" captionStartId="8.[151,250,1874,1899]" captionTargetBox="[151,1436,1237,1855]" captionTargetId="figure@8.[151,1436,1237,1855]" captionTargetPageId="8" captionText="FIGURE 6. Comparison of head scalation between C. machadoi, C. phonolithos sp. nov., C. namakuiyus and C. angolensis." figureDoi="http://doi.org/10.5281/zenodo.3449910" httpUri="https://zenodo.org/record/3449910/files/figure.png" pageId="8" pageNumber="511">Fig. 6</figureCitation>
). It is distinguished from
<taxonomicName id="F1814D01FFD7FF94F096FF258776FF24" baseAuthorityName="Reichenow" baseAuthorityYear="1887" box="[1242,1349,189,215]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="vittifer">
<emphasis id="04F5EA90FFD7FF94F096FF258776FF24" box="[1242,1349,189,215]" italics="true" pageId="8" pageNumber="511">C. vittifer</emphasis>
</taxonomicName>
by possessing an incomplete row of pre-occipital scales between posterior parietal and occipital scales (
<emphasis id="04F5EA90FFD7FF94F09BFF7A872CFF08" box="[1239,1311,226,251]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
complete), and by having infralabials that are moderately deeply ridged (
<emphasis id="04F5EA90FFD7FF94F716FE9E8191FEEC" box="[858,930,262,287]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
usually smooth). It is distinguished from
<taxonomicName id="F1814D01FFD7FF94F1CFFE9D8336FEB0" authorityName="Laurent" authorityYear="1964" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F1CFFE9D8336FEB0" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
by having a large keyhole-shaped interparietal in contact with frontoparietals and separating the anterior parietals (
<emphasis id="04F5EA90FFD7FF94F54BFED6837CFE94" box="[263,335,334,359]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
small, diamond-shaped interparietal not in contact with frontoparietal and never completely separating the anterior parietals in
<taxonomicName id="F1814D01FFD7FF94F5A5FEE98044FE78" authorityName="Laurent" authorityYear="1964" box="[489,631,369,395]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F5A5FEE98044FE78" box="[489,631,369,395]" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
, see
<figureCitation id="AEBA2A07FFD7FF94F6FCFEE980C0FE7F" box="[688,755,369,396]" captionStart="FIGURE 6" captionStartId="8.[151,250,1874,1899]" captionTargetBox="[151,1436,1237,1855]" captionTargetId="figure@8.[151,1436,1237,1855]" captionTargetPageId="8" captionText="FIGURE 6. Comparison of head scalation between C. machadoi, C. phonolithos sp. nov., C. namakuiyus and C. angolensis." figureDoi="http://doi.org/10.5281/zenodo.3449910" httpUri="https://zenodo.org/record/3449910/files/figure.png" pageId="8" pageNumber="511">Fig. 6</figureCitation>
); having the intrusion of a scale in broad contact with the interparietal and occipitals, thereby separating posterior parietals (
<emphasis id="04F5EA90FFD7FF94F724FE0E8183FE5C" box="[872,944,406,431]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
absence of this scale and posterior parietals in full contact in
<taxonomicName id="F1814D01FFD7FF94F517FE2183D8FE20" authorityName="Laurent" authorityYear="1964" box="[347,491,441,467]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F517FE2183D8FE20" box="[347,491,441,467]" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
, see
<figureCitation id="AEBA2A07FFD7FF94F66AFE218059FE27" box="[550,618,441,468]" captionStart="FIGURE 6" captionStartId="8.[151,250,1874,1899]" captionTargetBox="[151,1436,1237,1855]" captionTargetId="figure@8.[151,1436,1237,1855]" captionTargetPageId="8" captionText="FIGURE 6. Comparison of head scalation between C. machadoi, C. phonolithos sp. nov., C. namakuiyus and C. angolensis." figureDoi="http://doi.org/10.5281/zenodo.3449910" httpUri="https://zenodo.org/record/3449910/files/figure.png" pageId="8" pageNumber="511">Fig. 6</figureCitation>
); a higher number of supralabials (
<quantity id="F1799B67FFD7FF94F7B5FE218615FE20" box="[1017,1062,441,467]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="511" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F062FE2186E7FE20" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[1070,1236,441,467]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD7FF94F062FE2186E7FE20" box="[1070,1236,441,467]" italics="true" pageId="8" pageNumber="511">C. phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD7FF94F090FE218704FE20" bold="true" box="[1244,1335,441,467]" pageId="8" pageNumber="511">
<taxonomicNameLabel id="1FC657EBFFD7FF94F090FE218704FE20" box="[1244,1335,441,467]" pageId="8" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="04F5EA90FFD7FF94F173FE2287B4FE20" box="[1343,1415,442,467]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
<quantity id="F1799B67FFD7FF94F1C2FE20829EFE04" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="511" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F4F8FE458370FE04" authorityName="Laurent" authorityYear="1964" box="[180,323,477,503]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F4F8FE458370FE04" box="[180,323,477,503]" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
); males with fewer caudal scales at the 15
<superScript id="C1F49BCAFFD7FF94F751FE448119FE19" attach="left" box="[797,810,476,490]" fontSize="6" pageId="8" pageNumber="511">th</superScript>
tail whorl (
<quantity id="F1799B67FFD7FF94F7E2FE4581EAFE04" box="[942,985,477,503]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="511" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F793FE4586B7FE04" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[991,1156,477,503]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD7FF94F793FE4586B7FE04" box="[991,1156,477,503]" italics="true" pageId="8" pageNumber="511">C. phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD7FF94F0C7FE4586D6FE04" bold="true" box="[1163,1253,477,503]" pageId="8" pageNumber="511">
<taxonomicNameLabel id="1FC657EBFFD7FF94F0C7FE4586D6FE04" box="[1163,1253,477,503]" pageId="8" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="04F5EA90FFD7FF94F0A0FE468707FE04" box="[1260,1332,478,503]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
<quantity id="F1799B67FFD7FF94F176FE4587AFFE04" box="[1338,1436,477,503]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="8" pageNumber="511" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">1011 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F4DBFD998315FDE8" authorityName="Laurent" authorityYear="1964" box="[151,294,513,539]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F4DBFD998315FDE8" box="[151,294,513,539]" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
); an orange-brown dorsal body coloration (
<emphasis id="04F5EA90FFD7FF94F742FD9A8165FDE8" box="[782,854,514,539]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
darker brown to black in
<taxonomicName id="F1814D01FFD7FF94F039FD998730FDE8" authorityName="Laurent" authorityYear="1964" box="[1141,1283,513,539]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F039FD998730FDE8" box="[1141,1283,513,539]" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
), the absence of dark speckles on throat and ventral body surfaces (
<emphasis id="04F5EA90FFD7FF94F6BCFDBE810BFDCC" box="[752,824,550,575]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
presence in
<taxonomicName id="F1814D01FFD7FF94F788FDBD8660FDCC" authorityName="Laurent" authorityYear="1964" box="[964,1107,549,575]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F788FDBD8660FDCC" box="[964,1107,549,575]" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
) and a reduced, widely separated posteromedial parietal process (similar to
<taxonomicName id="F1814D01FFD7FF94F6E2FDD18174FD90" baseAuthorityName="Bocage" baseAuthorityYear="1895" box="[686,839,585,611]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="04F5EA90FFD7FF94F6E2FDD18174FD90" box="[686,839,585,611]" italics="true" pageId="8" pageNumber="511">C. angolensis</emphasis>
</taxonomicName>
and unlike
<emphasis id="04F5EA90FFD7FF94F787FDD18651FD90" box="[971,1122,585,611]" italics="true" pageId="8" pageNumber="511">
<taxonomicName id="F1814D01FFD7FF94F787FDD1866EFD90" authorityName="Laurent" authorityYear="1964" box="[971,1117,585,611]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">C. machadoi</taxonomicName>
,
</emphasis>
in which it is extensive and forked, and
<taxonomicName id="F1814D01FFD7FF94F56DFDF583FAFD74" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[289,457,621,647]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD7FF94F56DFDF583FAFD74" box="[289,457,621,647]" italics="true" pageId="8" pageNumber="511">C. namakuiyus</emphasis>
</taxonomicName>
, in which it is extended and unbifurcated). It is distinguished from
<taxonomicName id="F1814D01FFD7FF94F083FDF5874BFD74" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[1231,1400,621,647]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD7FF94F083FDF5874BFD74" box="[1231,1400,621,647]" italics="true" pageId="8" pageNumber="511">C. namakuiyus</emphasis>
</taxonomicName>
by having the intrusion of a scale in broad contact with interparietal and occipitals, thereby separating posterior parietals (
<emphasis id="04F5EA90FFD7FF94F497FD2E8310FD3C" box="[219,291,694,719]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
absence of this scale, posterior parietals in full contact, see
<figureCitation id="AEBA2A07FFD7FF94F78EFD2D8636FD23" box="[962,1029,693,720]" captionStart="FIGURE 6" captionStartId="8.[151,250,1874,1899]" captionTargetBox="[151,1436,1237,1855]" captionTargetId="figure@8.[151,1436,1237,1855]" captionTargetPageId="8" captionText="FIGURE 6. Comparison of head scalation between C. machadoi, C. phonolithos sp. nov., C. namakuiyus and C. angolensis." figureDoi="http://doi.org/10.5281/zenodo.3449910" httpUri="https://zenodo.org/record/3449910/files/figure.png" pageId="8" pageNumber="511">Fig. 6</figureCitation>
); a higher number of supralabials (
<quantity id="F1799B67FFD7FF94F1C2FD2D829EFD00" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="511" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F4F8FD41836BFD00" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[180,344,729,755]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD7FF94F4F8FD41836BFD00" box="[180,344,729,755]" italics="true" pageId="8" pageNumber="511">C. phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD7FF94F513FD41838BFD00" bold="true" box="[351,440,729,755]" pageId="8" pageNumber="511">
<taxonomicNameLabel id="1FC657EBFFD7FF94F513FD41838BFD00" box="[351,440,729,755]" pageId="8" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="04F5EA90FFD7FF94F5F3FD428034FD00" box="[447,519,730,755]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
<quantity id="F1799B67FFD7FF94F641FD408067FD00" box="[525,596,728,755]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="8" pageNumber="511" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">45 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F616FD418132FD00" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[602,769,729,755]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD7FF94F616FD418132FD00" box="[602,769,729,755]" italics="true" pageId="8" pageNumber="511">C. namakuiyus</emphasis>
</taxonomicName>
); a higher number of femoral pores (
<quantity id="F1799B67FFD7FF94F0D2FD4186FAFD00" box="[1182,1225,729,755]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="8" pageNumber="511" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F083FD418747FD00" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[1231,1396,729,755]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD7FF94F083FD418747FD00" box="[1231,1396,729,755]" italics="true" pageId="8" pageNumber="511">C. phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD7FF94F137FD4182FBFCE4" bold="true" pageId="8" pageNumber="511">
<taxonomicNameLabel id="1FC657EBFFD7FF94F137FD4182FBFCE4" pageId="8" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="04F5EA90FFD7FF94F49CFD66832BFCE4" box="[208,280,766,791]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
<quantity id="F1799B67FFD7FF94F552FD658352FCE4" box="[286,353,765,791]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="8" pageNumber="511" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4-6 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F524FD65803CFCE4" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[360,527,765,791]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD7FF94F524FD65803CFCE4" box="[360,527,765,791]" italics="true" pageId="8" pageNumber="511">C. namakuiyus</emphasis>
</taxonomicName>
); a higher number of generation glands (
<quantity id="F1799B67FFD7FF94F790FD658673FCE4" box="[988,1088,765,791]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="8" pageNumber="511" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">1617 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F00BFD6586DFFCE4" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[1095,1260,765,791]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD7FF94F00BFD6586DFFCE4" box="[1095,1260,765,791]" italics="true" pageId="8" pageNumber="511">C. phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD7FF94F0B8FD65877EFCE4" bold="true" box="[1268,1357,765,791]" pageId="8" pageNumber="511">
<taxonomicNameLabel id="1FC657EBFFD7FF94F0B8FD65877EFCE4" box="[1268,1357,765,791]" pageId="8" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="04F5EA90FFD7FF94F119FD6687AEFCE4" box="[1365,1437,766,791]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
<quantity id="F1799B67FFD7FF94F4DBFCB982E2FCC8" box="[151,209,801,827]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="8" pageNumber="511" unit="in" value="12.0">12 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F494FCB9834CFCC8" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[216,383,801,827]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD7FF94F494FCB9834CFCC8" box="[216,383,801,827]" italics="true" pageId="8" pageNumber="511">C. namakuiyus</emphasis>
</taxonomicName>
); males with fewer caudal scales at the 15
<superScript id="C1F49BCAFFD7FF94F712FCB88158FCDD" attach="left" box="[862,875,800,814]" fontSize="6" pageId="8" pageNumber="511">th</superScript>
tail whorl (
<quantity id="F1799B67FFD7FF94F7A3FCB98628FCC8" box="[1007,1051,801,827]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="511" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F06EFCB986FBFCC8" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[1058,1224,801,827]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD7FF94F06EFCB986FBFCC8" box="[1058,1224,801,827]" italics="true" pageId="8" pageNumber="511">C. phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD7FF94F09CFCB98719FCC8" bold="true" box="[1232,1322,801,827]" pageId="8" pageNumber="511">
<taxonomicNameLabel id="1FC657EBFFD7FF94F09CFCB98719FCC8" box="[1232,1322,801,827]" pageId="8" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="04F5EA90FFD7FF94F17DFCBA874AFCC8" box="[1329,1401,802,827]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
<quantity id="F1799B67FFD7FF94F1CCFCB9829EFCAC" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="511" unit="in" value="10.0">10 in</quantity>
<taxonomicName id="F1814D01FFD7FF94F4FFFCDD836AFCAC" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[179,345,837,863]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD7FF94F4FFFCDD836AFCAC" box="[179,345,837,863]" italics="true" pageId="8" pageNumber="511">C. namakuiyus</emphasis>
</taxonomicName>
); absence of osteoderms on throat and ventral surfaces (
<emphasis id="04F5EA90FFD7FF94F784FCDE8623FCAC" box="[968,1040,838,863]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
presence), and significantly thicker caudal osteoderms than dorsal osteoderms (resembling
<taxonomicName id="F1814D01FFD7FF94F6B3FCF181BFFC70" authorityName="Laurent" authorityYear="1964" box="[767,908,873,899]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F6B3FCF181BFFC70" box="[767,908,873,899]" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
and
<taxonomicName id="F1814D01FFD7FF94F78EFCF1866AFC70" baseAuthorityName="Bocage" baseAuthorityYear="1895" box="[962,1113,873,899]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="04F5EA90FFD7FF94F78EFCF1866AFC70" box="[962,1113,873,899]" italics="true" pageId="8" pageNumber="511">C. angolensis</emphasis>
</taxonomicName>
); temporal scales are weakly keeled (
<emphasis id="04F5EA90FFD7FF94F4A3FC168304FC54" box="[239,311,910,935]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
strongly keeled). In coloration
<taxonomicName id="F1814D01FFD7FF94F6DEFC158105FC54" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[658,822,909,935]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD7FF94F6DEFC158105FC54" box="[658,822,909,935]" italics="true" pageId="8" pageNumber="511">C. phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD7FF94F770FC1581A7FC54" bold="true" box="[828,916,909,935]" pageId="8" pageNumber="511">
<taxonomicNameLabel id="1FC657EBFFD7FF94F770FC1581A7FC54" box="[828,916,909,935]" pageId="8" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is quite similar to
<taxonomicName id="F1814D01FFD7FF94F02CFC158735FC54" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[1120,1286,909,935]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD7FF94F02CFC158735FC54" box="[1120,1286,909,935]" italics="true" pageId="8" pageNumber="511">C. namakuiyus</emphasis>
</taxonomicName>
, although the new species has a more vibrant coloration (orange-brown
<emphasis id="04F5EA90FFD7FF94F755FC2A8152FC38" box="[793,865,946,971]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
light brown). It is distinguished from
<taxonomicName id="F1814D01FFD7FF94F149FC2987AFFC38" baseAuthorityName="Bocage" baseAuthorityYear="1895" box="[1285,1436,945,971]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="04F5EA90FFD7FF94F149FC2987AFFC38" box="[1285,1436,945,971]" italics="true" pageId="8" pageNumber="511">C. angolensis</emphasis>
</taxonomicName>
by having a large keyhole-shaped interparietal in contact with frontoparietals, thereby separating anterior parietals, with an intrusion of a scale in broad contact with interparietal and occipitals (
<emphasis id="04F5EA90FFD7FF94F7A3FC628604FBE0" box="[1007,1079,1018,1043]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
small, diamond-shaped interparietal not in contact with frontoparietal, thereby never completely separating anterior parietals, posterior parietals in broad contact, see
<figureCitation id="AEBA2A07FFD7FF94F52BFBD98399FBAF" box="[359,426,1089,1116]" captionStart="FIGURE 6" captionStartId="8.[151,250,1874,1899]" captionTargetBox="[151,1436,1237,1855]" captionTargetId="figure@8.[151,1436,1237,1855]" captionTargetPageId="8" captionText="FIGURE 6. Comparison of head scalation between C. machadoi, C. phonolithos sp. nov., C. namakuiyus and C. angolensis." figureDoi="http://doi.org/10.5281/zenodo.3449910" httpUri="https://zenodo.org/record/3449910/files/figure.png" pageId="8" pageNumber="511">Fig. 6</figureCitation>
); fewer ventral transverse scale rows (23
<emphasis id="04F5EA90FFD7FF94F737FBDA81F0FBA8" box="[891,963,1090,1115]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
27), orange-brown dorsal body coloration (
<emphasis id="04F5EA90FFD7FF94F4EDFBFE82DAFB8C" box="[161,233,1126,1151]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
brown with blackish speckles over paler dorsal ground coloration), and by the absence of a longitudinal series of whitish speckles along dorsal surface (
<emphasis id="04F5EA90FFD7FF94F6E6FB1280C1FB50" box="[682,754,1162,1187]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
presence of two longitudinal series of small whitish speckles along dorsum), and presence (
<emphasis id="04F5EA90FFD7FF94F5A5FB368002FB34" box="[489,561,1198,1223]" italics="true" pageId="8" pageNumber="511">versus</emphasis>
absence) of a loreal.
</paragraph>
</subSubSection>
<caption id="62FE660AFFD7FF94F4DBF8CA87B4F899" ID-DOI="http://doi.org/10.5281/zenodo.3449910" ID-Zenodo-Dep="3449910" box="[151,1415,1874,1899]" httpUri="https://zenodo.org/record/3449910/files/figure.png" pageId="8" pageNumber="511" startId="8.[151,250,1874,1899]" targetBox="[151,1436,1237,1855]" targetPageId="8">
<paragraph id="363E3682FFD7FF94F4DBF8CA87B4F899" blockId="8.[151,1415,1874,1899]" box="[151,1415,1874,1899]" pageId="8" pageNumber="511">
<emphasis id="04F5EA90FFD7FF94F4DBF8CA8320F898" bold="true" box="[151,275,1874,1899]" pageId="8" pageNumber="511">FIGURE 6.</emphasis>
Comparison of head scalation between
<taxonomicName id="F1814D01FFD7FF94F6E4F8CB8119F898" authorityName="Laurent" authorityYear="1964" box="[680,810,1875,1899]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD7FF94F6E4F8CB8119F898" box="[680,810,1875,1899]" italics="true" pageId="8" pageNumber="511">C. machadoi</emphasis>
</taxonomicName>
,
<taxonomicName id="F1814D01FFD7FF94F77AF8CB81FFF898" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[822,972,1875,1899]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD7FF94F77AF8CB81FFF898" box="[822,972,1875,1899]" italics="true" pageId="8" pageNumber="511">C. phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD7FF94F79EF8CB8611F898" bold="true" box="[978,1058,1875,1899]" pageId="8" pageNumber="511">
<taxonomicNameLabel id="1FC657EBFFD7FF94F79EF8CB8611F898" box="[978,1058,1875,1899]" pageId="8" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="F1814D01FFD7FF94F07CF8CB86F5F898" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[1072,1222,1875,1899]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="511" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD7FF94F07CF8CB86F5F898" box="[1072,1222,1875,1899]" italics="true" pageId="8" pageNumber="511">C. namakuiyus</emphasis>
</taxonomicName>
and
<taxonomicName id="F1814D01FFD7FF94F0BBF8CB87B2F898" baseAuthorityName="Bocage" baseAuthorityYear="1895" box="[1271,1409,1875,1899]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="517" phylum="Chordata" rank="species" species="angolensis">
C.
<emphasis id="04F5EA90FFD7FF94F159F8CB87B2F898" box="[1301,1409,1875,1899]" italics="true" pageId="8" pageNumber="511">angolensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="7E9B6509FFD6FF93F48BFF008764FEEC" lastPageId="15" lastPageNumber="518" pageId="9" pageNumber="512" type="description">
<paragraph id="363E3682FFD6FF95F48BFF008048FCE4" blockId="9.[151,1437,152,791]" pageId="9" pageNumber="512">
<emphasis id="04F5EA90FFD6FF95F48BFF0083D9FF40" bold="true" box="[199,490,152,179]" pageId="9" pageNumber="512">
Description of
<typeStatus id="E93A8820FFD6FF95F530FF0183D6FF40" box="[380,485,153,179]" pageId="9" pageNumber="512" type="holotype">holotype</typeStatus>
.
</emphasis>
SVL
<quantity id="F1799B67FFD6FF95F67DFF0180A8FF40" box="[561,667,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="7.14" pageId="9" pageNumber="512" unit="mm" value="71.4">71.4 mm</quantity>
. Head and body depressed. Head 1.3 times longer (
<quantity id="F1799B67FFD6FF95F0BDFF018769FF40" box="[1265,1370,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="9" pageNumber="512" unit="mm" value="21.6">21.6 mm</quantity>
) than broad (
<quantity id="F1799B67FFD6FF95F4A4FF248360FF24" box="[232,339,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="9" pageNumber="512" unit="mm" value="16.5">16.5 mm</quantity>
). HH
<quantity id="F1799B67FFD6FF95F5D5FF2583C7FF24" box="[409,500,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="9" pageNumber="512" unit="mm" value="8.7">8.7 mm</quantity>
. SEL
<quantity id="F1799B67FFD6FF95F675FF2580A7FF24" box="[569,660,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="512" unit="mm" value="8.2">8.2 mm</quantity>
. Nasals in median broad contact; entire frontonasal lozenge-shaped, broader than long, separated from frontal by enlarged prefrontals (in median contact, forming a suture), separated from rostral by nasals, separated from loreal by prefrontals; frontal in contact with first and second supraoculars, followed by a pair of frontoparietals in broad, median contact; a distinctive keyhole-shaped interparietal in broad contact with the frontoparietals, separating anterior parietals; intrusion of a scale in broad contact with interparietal and occipitals, thereby separating posterior parietals; right posterior parietal scale is fragmented; parietal window visible; 10 rugose occipital scales; 10 elongated nuchal scales. Four supraoculars and three supraciliaries. Nasals large, with nostril pierced centrally on upper margin. Loreal in contact with preocular, nasal and first two supralabials; three suboculars, well separated from the lip by the third, fourth and fifth supralabials. Rostral twice as broad as deep; supralabials 6; infralabials 6; chin shields 5. Mental twice as broad as long; gulars smooth, enlarged and forming transverse rows posteriorly, with 17 gulars between the posterior extent of the jaws. Dorsal scales rectangular, rugose, strongly and obtusely keeled; dorsals and laterals in 25 transverse and 23 longitudinal rows; ventrals squarish, smooth, in 23 transverse and 17 longitudinal rows. Scales on dorsum of limbs large, strongly keeled and spinose with thin, non-imbricated osteoderms; subdigital lamellae under fourth toe 15; SAL
<quantity id="F1799B67FFD6FF95F0FFFD09872FFD58" box="[1203,1308,657,683]" metricMagnitude="-2" metricUnit="m" metricValue="2.8899999999999997" pageId="9" pageNumber="512" unit="mm" value="28.9">28.9 mm</quantity>
. AGD
<quantity id="F1799B67FFD6FF95F127FD0982F5FD3C" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" pageId="9" pageNumber="512" unit="mm" value="32.4">32.4 mm</quantity>
. HML
<quantity id="F1799B67FFD6FF95F55FFD2D8359FD3C" box="[275,362,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="9" pageNumber="512" unit="mm" value="9.3">9.3 mm</quantity>
, RUL
<quantity id="F1799B67FFD6FF95F5FEFD2D8039FD3C" box="[434,522,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="9" pageNumber="512" unit="mm" value="7.9">7.9 mm</quantity>
; FL
<quantity id="F1799B67FFD6FF95F676FD2D8093FD3C" box="[570,672,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="9" pageNumber="512" unit="mm" value="12.9">12.9 mm</quantity>
; TFL
<quantity id="F1799B67FFD6FF95F6ADFD2D8174FD3C" box="[737,839,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="9" pageNumber="512" unit="mm" value="10.6">10.6 mm</quantity>
; LTL
<quantity id="F1799B67FFD6FF95F7CBFD2D81ECFD3C" box="[903,991,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="9" pageNumber="512" unit="mm" value="9.8">9.8 mm</quantity>
; femoral pores seven; generation glands 1617. Tail with whorls of large, elongate, strongly keeled, spinose and acuminate scales, spines directed posteriorly; largest spines in dorsolateral position.
</paragraph>
<caption id="62FE660AFFD6FF95F4DBF8D487A7F807" ID-DOI="http://doi.org/10.5281/zenodo.3449916" ID-Zenodo-Dep="3449916" httpUri="https://zenodo.org/record/3449916/files/figure.png" pageId="9" pageNumber="512" startId="9.[151,250,1868,1893]" targetBox="[151,1436,805,1848]" targetPageId="9">
<paragraph id="363E3682FFD6FF95F4DBF8D487A7F807" blockId="9.[151,1437,1868,2037]" pageId="9" pageNumber="512">
<emphasis id="04F5EA90FFD6FF95F4DBF8D48327F896" bold="true" box="[151,276,1868,1893]" pageId="9" pageNumber="512">FIGURE 7.</emphasis>
Figure showing Cranial and postcranial osteology of CAS 263581
<taxonomicName id="F1814D01FFD6FF95F781F8D4869BF897" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[973,1192,1868,1892]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="512" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD6FF95F781F8D4869BF897" box="[973,1192,1868,1892]" italics="true" pageId="9" pageNumber="512">Cordylus phonolithos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FC657EBFFD6FF95F0FCF8D58730F897" box="[1200,1283,1869,1893]" pageId="9" pageNumber="512" rank="species">
<emphasis id="04F5EA90FFD6FF95F0FCF8D586CEF896" bold="true" box="[1200,1277,1869,1893]" pageId="9" pageNumber="512">sp. nov</emphasis>
.
</taxonomicNameLabel>
holotype.
<emphasis id="04F5EA90FFD6FF95F13EF8D587BDF897" box="[1394,1422,1869,1892]" italics="true" pageId="9" pageNumber="512">An</emphasis>
= angular,
<emphasis id="04F5EA90FFD6FF95F4BCF8E98323F87B" box="[240,272,1905,1928]" italics="true" pageId="9" pageNumber="512">Art</emphasis>
= articular,
<emphasis id="04F5EA90FFD6FF95F5C1F8E9839AF87B" box="[397,425,1905,1928]" italics="true" pageId="9" pageNumber="512">Bo</emphasis>
= Basioccipital,
<emphasis id="04F5EA90FFD6FF95F614F8E98048F87B" box="[600,635,1905,1928]" italics="true" pageId="9" pageNumber="512">Bpt</emphasis>
= Basipterygoid,
<emphasis id="04F5EA90FFD6FF95F760F8E98179F87B" box="[812,842,1905,1928]" italics="true" pageId="9" pageNumber="512">Co</emphasis>
= coronoid,
<emphasis id="04F5EA90FFD6FF95F782F8E981DFF87B" box="[974,1004,1905,1928]" italics="true" pageId="9" pageNumber="512">De</emphasis>
= dentary,
<emphasis id="04F5EA90FFD6FF95F013F8E98649F87B" box="[1119,1146,1905,1928]" italics="true" pageId="9" pageNumber="512">Ec</emphasis>
= ectopterygoid,
<emphasis id="04F5EA90FFD6FF95F163F8E98778F87B" box="[1327,1355,1905,1928]" italics="true" pageId="9" pageNumber="512">Ep</emphasis>
= epipterygoid,
<emphasis id="04F5EA90FFD6FF95F4B4F80D8334F85F" box="[248,263,1941,1964]" italics="true" pageId="9" pageNumber="512">F</emphasis>
= frontal,
<emphasis id="04F5EA90FFD6FF95F53FF80D83BFF85F" box="[371,396,1941,1964]" italics="true" pageId="9" pageNumber="512">Hi</emphasis>
= hyperischium,
<emphasis id="04F5EA90FFD6FF95F672F80D806EF85F" box="[574,605,1941,1964]" italics="true" pageId="9" pageNumber="512">Ho</emphasis>
= hypoischium,
<emphasis id="04F5EA90FFD6FF95F74BF80D8121F85F" box="[775,786,1941,1964]" italics="true" pageId="9" pageNumber="512">J</emphasis>
= jugal,
<emphasis id="04F5EA90FFD6FF95F721F80D81BEF85F" box="[877,909,1941,1964]" italics="true" pageId="9" pageNumber="512">Mx</emphasis>
= maxilla,
<emphasis id="04F5EA90FFD6FF95F04DF80D8621F85F" box="[1025,1042,1941,1964]" italics="true" pageId="9" pageNumber="512">N</emphasis>
= nasal,
<emphasis id="04F5EA90FFD6FF95F021F80D864FF85F" box="[1133,1148,1941,1964]" italics="true" pageId="9" pageNumber="512">P</emphasis>
= parietal,
<emphasis id="04F5EA90FFD6FF95F0BCF80D873FF85F" box="[1264,1292,1941,1964]" italics="true" pageId="9" pageNumber="512">Pa</emphasis>
= Palatine,
<emphasis id="04F5EA90FFD6FF95F1CAF80D87AEF85E" box="[1414,1437,1941,1965]" italics="true" pageId="9" pageNumber="512">Pf</emphasis>
= postfrontal,
<emphasis id="04F5EA90FFD6FF95F569F8218361F823" box="[293,338,1977,2000]" italics="true" pageId="9" pageNumber="512">Pmx</emphasis>
= premaxilla,
<emphasis id="04F5EA90FFD6FF95F592F82183C9F823" box="[478,506,1977,2000]" italics="true" pageId="9" pageNumber="512">Po</emphasis>
= Postorbital,
<emphasis id="04F5EA90FFD6FF95F6C1F82180F3F823" box="[653,704,1977,2000]" italics="true" pageId="9" pageNumber="512">Pocc</emphasis>
= paraoccipital,
<emphasis id="04F5EA90FFD6FF95F724F82181B7F823" box="[872,900,1977,2000]" italics="true" pageId="9" pageNumber="512">Pp</emphasis>
= prepubic cartilage,
<emphasis id="04F5EA90FFD6FF95F013F82186A9F823" box="[1119,1178,1977,2000]" italics="true" pageId="9" pageNumber="512">Ppmp</emphasis>
= posteriomedial process of the parietal,
<emphasis id="04F5EA90FFD6FF95F57DF8458362F806" box="[305,337,2013,2037]" italics="true" pageId="9" pageNumber="512">Prf</emphasis>
= prefrontal,
<emphasis id="04F5EA90FFD6FF95F590F8458032F807" box="[476,513,2013,2036]" italics="true" pageId="9" pageNumber="512">Pro</emphasis>
= prootic,
<emphasis id="04F5EA90FFD6FF95F63DF84580BBF807" box="[625,648,2013,2036]" italics="true" pageId="9" pageNumber="512">Pt</emphasis>
= pterygoid,
<emphasis id="04F5EA90FFD6FF95F75CF8458111F807" box="[784,802,2013,2036]" italics="true" pageId="9" pageNumber="512">Q</emphasis>
= quadrate,
<emphasis id="04F5EA90FFD6FF95F7EDF8458189F807" box="[929,954,2013,2036]" italics="true" pageId="9" pageNumber="512">Sq</emphasis>
= squamosal,
<emphasis id="04F5EA90FFD6FF95F001F8458652F807" box="[1101,1121,2013,2036]" italics="true" pageId="9" pageNumber="512">St</emphasis>
= supratemporal,
<emphasis id="04F5EA90FFD6FF95F155F8458701F807" box="[1305,1330,2013,2036]" italics="true" pageId="9" pageNumber="512">Vo</emphasis>
= vomer.
</paragraph>
</caption>
<paragraph id="363E3682FFD5FF96F48BFF008166FCE4" blockId="10.[151,1437,152,791]" pageId="10" pageNumber="513">
<emphasis id="04F5EA90FFD5FF96F48BFF008393FF40" bold="true" box="[199,416,152,179]" pageId="10" pageNumber="513">Cranial osteology.</emphasis>
The parietal is pentagonal with two short, well-spaced posteromedial processes bracing a very small, laterally flattened posteriomedial supraoccipital process (
<figureCitation id="AEBA2A07FFD5FF96F7ECFF2581D6FF24" box="[928,997,189,215]" captionStart="FIGURE 7" captionStartId="9.[151,250,1868,1893]" captionTargetBox="[151,1436,805,1848]" captionTargetId="figure@9.[151,1436,805,1848]" captionTargetPageId="9" captionText="FIGURE 7. Figure showing Cranial and postcranial osteology of CAS 263581 Cordylus phonolithos sp. nov. holotype. An= angular, Art = articular, Bo = Basioccipital, Bpt= Basipterygoid, Co = coronoid, De = dentary, Ec = ectopterygoid, Ep = epipterygoid, F = frontal, Hi = hyperischium, Ho = hypoischium, J = jugal, Mx = maxilla, N = nasal, P = parietal, Pa = Palatine, Pf = postfrontal, Pmx= premaxilla, Po = Postorbital, Pocc = paraoccipital, Pp = prepubic cartilage, Ppmp = posteriomedial process of the parietal, Prf = prefrontal, Pro = prootic, Pt = pterygoid, Q = quadrate, Sq = squamosal, St = supratemporal, Vo = vomer." figureDoi="http://doi.org/10.5281/zenodo.3449916" httpUri="https://zenodo.org/record/3449916/files/figure.png" pageId="10" pageNumber="513">Fig. 7</figureCitation>
). The premaxilla is unpaired and bears seven pleurodont teeth and five foramina, with a dorsal process that extends posteriorly to be clasped by the nasals, which themselves insert into an unpaired frontal posteriorly and prefrontals posterolaterally. The maxilla is typically scinciform, with a deeply grooved crista dentalis and 20 pleurodont teeth. A laminar lacrimal lies medial to the facial process of the maxilla, extending from the posterioventral process of the prefrontal to the anterior process of the jugal. No palpebral is present, though the prefrontal has a small, flattened, laterally projecting tubercle that supports the anteriormost superorbital osteoderm in much the same way. The jugal is triangular in cross-section and asymmetrically T-shaped, with a tapering anterior process and a broad, truncated posterior process that extends along and past the posterior edge of the maxilla. Edentate pterygoids extend back to connect with the quadrates, becoming C-shaped in cross-section posterior to the epipterygoid condyle. The squamosal is curved and blade-like, circular in cross-section anteriorly, becoming flattened posteriorly, where it articulates with the cephalic condyle of the quadrate and the supratemporals. Supratemporals are flattened, sickle-shaped and unfused with the paroccipital processes. The bones of the braincase are unfused, suggesting that this individual may be a subadult. The prootic bears an extended alar process, a well-developed, anteriorly expanded crista prootica, and a very weak supratrigeminal process. Basipterygoid processes are well developed and flattened. The lower jaw has a large adductor fossa, a flattened and medially curved retroarticular process, a medially open Meckelian canal and a dentary with a strong subdental shelf, 22 mandibular teeth, and 11 dentary foramina.
</paragraph>
<caption id="62FE660AFFD5FF96F4DBF8CC836FF862" ID-DOI="http://doi.org/10.5281/zenodo.3449920" ID-Zenodo-Dep="3449920" httpUri="https://zenodo.org/record/3449920/files/figure.png" pageId="10" pageNumber="513" startId="10.[151,250,1876,1901]" targetBox="[163,1424,813,1851]" targetPageId="10">
<paragraph id="363E3682FFD5FF96F4DBF8CC836FF862" blockId="10.[151,1436,1876,1937]" pageId="10" pageNumber="513">
<emphasis id="04F5EA90FFD5FF96F4DBF8CC8321F89E" bold="true" box="[151,274,1876,1901]" pageId="10" pageNumber="513">FIGURE 8.</emphasis>
Cranial morphology and osteodermal variation on the different Angolan
<taxonomicName id="F1814D01FFD5FF96F7BDF8CD867EF89E" box="[1009,1101,1877,1901]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="513" phylum="Chordata" rank="genus">
<emphasis id="04F5EA90FFD5FF96F7BDF8CD867EF89E" box="[1009,1101,1877,1901]" italics="true" pageId="10" pageNumber="513">Cordylus</emphasis>
</taxonomicName>
species, colorized to show crosssectional thickness.
</paragraph>
</caption>
<caption id="62FE660AFFD4FF97F4DBF94B82CEF8E3" ID-DOI="http://doi.org/10.5281/zenodo.3449924" ID-Zenodo-Dep="3449924" httpUri="https://zenodo.org/record/3449924/files/figure.png" pageId="11" pageNumber="514" startId="11.[151,250,1747,1772]" targetBox="[203,1384,181,1722]" targetPageId="11">
<paragraph id="363E3682FFD4FF97F4DBF94B82CEF8E3" blockId="11.[151,1437,1747,1808]" pageId="11" pageNumber="514">
<emphasis id="04F5EA90FFD4FF97F4DBF94B8327F91F" bold="true" box="[151,276,1747,1772]" pageId="11" pageNumber="514">FIGURE 9.</emphasis>
Whole body osteodermal variation on the different Angolan
<taxonomicName id="F1814D01FFD4FF97F7C0F94C81DBF91F" box="[908,1000,1748,1772]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="514" phylum="Chordata" rank="genus">
<emphasis id="04F5EA90FFD4FF97F7C0F94C81DBF91F" box="[908,1000,1748,1772]" italics="true" pageId="11" pageNumber="514">Cordylus</emphasis>
</taxonomicName>
species, colorized to show cross-sectional thickness.
</paragraph>
</caption>
<paragraph id="363E3682FFD4FF90F48BF8F98326FE5C" blockId="11.[151,1437,1889,2025]" lastBlockId="12.[151,1437,153,899]" lastPageId="12" lastPageNumber="515" pageId="11" pageNumber="514">
<emphasis id="04F5EA90FFD4FF97F48BF8F983F6F88E" bold="true" box="[199,453,1889,1917]" pageId="11" pageNumber="514">Postcranial osteology</emphasis>
. The
<typeStatus id="E93A8820FFD4FF97F645F8FA8058F88F" box="[521,619,1890,1916]" pageId="11" pageNumber="514" type="holotype">holotype</typeStatus>
has 25 presacral, two sacral and 16 postsacral vertebrae (
<figureCitation id="AEBA2A07FFD4FF97F0B3F8FA8777F88E" box="[1279,1348,1890,1917]" captionStart="FIGURE 7" captionStartId="9.[151,250,1868,1893]" captionTargetBox="[151,1436,805,1848]" captionTargetId="figure@9.[151,1436,805,1848]" captionTargetPageId="9" captionText="FIGURE 7. Figure showing Cranial and postcranial osteology of CAS 263581 Cordylus phonolithos sp. nov. holotype. An= angular, Art = articular, Bo = Basioccipital, Bpt= Basipterygoid, Co = coronoid, De = dentary, Ec = ectopterygoid, Ep = epipterygoid, F = frontal, Hi = hyperischium, Ho = hypoischium, J = jugal, Mx = maxilla, N = nasal, P = parietal, Pa = Palatine, Pf = postfrontal, Pmx= premaxilla, Po = Postorbital, Pocc = paraoccipital, Pp = prepubic cartilage, Ppmp = posteriomedial process of the parietal, Prf = prefrontal, Pro = prootic, Pt = pterygoid, Q = quadrate, Sq = squamosal, St = supratemporal, Vo = vomer." figureDoi="http://doi.org/10.5281/zenodo.3449916" httpUri="https://zenodo.org/record/3449916/files/figure.png" pageId="11" pageNumber="514">Fig. 7</figureCitation>
). There are five cervical ribs, three sternal ribs and two xiphisternal ribs. The asternal ribs are asymmetric, with four long ribs with ossified costal cartilage and six short asternal ribs on the left side and five long and seven short ribs on the right. The first three cervical ribs are distally flattened with bifid cartilaginous projections. The pelvic girdle is well developed and flattened. No iliac tubercle is present. There is a well-developed, ventrally angled pubic tubercle directly anterior to the obturator foramen. Both hypoischium and hyperischium are well developed. Pubic bones are well separated by a bifurcate prepubic cartilage. The sternal plate is broad and lacks a fontanelle. Interclavicle cruciform, clavicles rod-like and flattened dorsally. The epicoracoid is narrow and curved, connecting the scapular ray to the primary and secondary coracoid rays, but not to the anterior process of the scapular. The condyles of long bones are unfused and the metatarsals and metacarpels are not fully developed, suggesting that this individual is not fully adult. Digits display the typical squamate phalangeal arrangement of 2-3-4-5-3 for the manus and 2-3-4-5-4 for the pes.
</paragraph>
<paragraph id="363E3682FFD3FF90F48BFE2081D8FD3C" blockId="12.[151,1437,153,899]" pageId="12" pageNumber="515">
<emphasis id="04F5EA90FFD3FF90F48BFE208365FE20" bold="true" box="[199,342,440,467]" pageId="12" pageNumber="515">Osteoderms</emphasis>
. Scales of the dorsal and temporal regions of the skull and the ventrolateral aspects of the jaws are underlain with rugose osteoderms (
<figureCitation id="AEBA2A07FFD3FF90F615FE458093FE04" box="[601,672,477,503]" captionStart="FIGURE 8" captionStartId="10.[151,250,1876,1901]" captionTargetBox="[163,1424,813,1851]" captionTargetId="figure@10.[163,1424,813,1851]" captionTargetPageId="10" captionText="FIGURE 8. Cranial morphology and osteodermal variation on the different Angolan Cordylus species, colorized to show crosssectional thickness." figureDoi="http://doi.org/10.5281/zenodo.3449920" httpUri="https://zenodo.org/record/3449920/files/figure.png" pageId="12" pageNumber="515">Fig. 8</figureCitation>
). These osteoderms are fused to the proximal parietal, frontal and postorbital bones, although the mesokinetic and metakinetic joints appear unobstructed and flexible. The tail, legs and dorsal and lateral aspects of the body are covered in osteoderms (
<figureCitation id="AEBA2A07FFD3FF90F7E5FDBD81DDFDCC" box="[937,1006,549,575]" captionStart="FIGURE 9" captionStartId="11.[151,250,1747,1772]" captionTargetBox="[203,1384,181,1722]" captionTargetId="figure@11.[203,1384,181,1722]" captionTargetPageId="11" captionText="FIGURE 9. Whole body osteodermal variation on the different Angolan Cordylus species, colorized to show cross-sectional thickness." figureDoi="http://doi.org/10.5281/zenodo.3449924" httpUri="https://zenodo.org/record/3449924/files/figure.png" pageId="12" pageNumber="515">Fig. 9</figureCitation>
). The dorsal and lateral trunk are protected by noncontiguous, rectangular,
<quantity id="F1799B67FFD3FF90F671FDD180F5FD90" box="[573,710,585,611]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="12" pageNumber="515" unit="mm" value="150.0" valueMax="200.0" valueMin="100.0">100-200 µm</quantity>
thick osteoderms that become increasingly keeled and mucronate laterally. The caudal osteoderms are significantly thicker (up to
<quantity id="F1799B67FFD3FF90F724FDF4818FFD74" box="[872,956,620,647]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="515" unit="mm" value="500.0">500µm</quantity>
), sharply spined and arranged in imbricate transverse whorls. The whole limbs are covered by imbricate circular/rhomboid osteoderms, keeled and mucronate dorsally, plate-like ventrally. The gular and ventral regions lack osteoderms.
</paragraph>
<paragraph id="363E3682FFD3FF90F48BFD4080C9FC70" blockId="12.[151,1437,153,899]" pageId="12" pageNumber="515">
<emphasis id="04F5EA90FFD3FF90F48BFD40837CFD00" bold="true" box="[199,335,728,755]" pageId="12" pageNumber="515">Coloration.</emphasis>
Dorsum orange-brown, fading to dirty yellow laterally. Head orange-brown; supralabials and infralabials yellowish; a dark-brown bar extends from the posterior aspect of the eye to the temporals. The base of the tail is brown, with an orange coloration similar to that the laterally that extends towards the tip; dorsum of limbs dark brown. Laterally, a dark-brown line extends from the neck towards the insertion of the forelimbs. The body venter is cream and subcaudal surface is faded orange.
</paragraph>
<caption id="62FE660AFFD3FF90F4DBF8F386E0F877" ID-DOI="http://doi.org/10.5281/zenodo.3449928" ID-Zenodo-Dep="3449928" box="[151,1235,1899,1924]" httpUri="https://zenodo.org/record/3449928/files/figure.png" pageId="12" pageNumber="515" startId="12.[151,250,1899,1924]" targetBox="[157,1432,919,1875]" targetPageId="12">
<paragraph id="363E3682FFD3FF90F4DBF8F386E0F877" blockId="12.[151,1235,1899,1924]" box="[151,1235,1899,1924]" pageId="12" pageNumber="515">
<emphasis id="04F5EA90FFD3FF90F4DBF8F38313F877" bold="true" box="[151,288,1899,1924]" pageId="12" pageNumber="515">FIGURE 10.</emphasis>
Habitat view of the type locality of
<taxonomicName id="F1814D01FFD3FF90F6C3F8F3815AF870" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[655,873,1899,1923]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="515" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD3FF90F6C3F8F3815AF870" box="[655,873,1899,1923]" italics="true" pageId="12" pageNumber="515">Cordylus phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD3FF90F723F8F381F3F870" bold="true" box="[879,960,1899,1923]" pageId="12" pageNumber="515">
<taxonomicNameLabel id="1FC657EBFFD3FF90F723F8F381F3F870" box="[879,960,1899,1923]" pageId="12" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Photo by Ishan Agarwal).
</paragraph>
</caption>
<caption id="62FE660AFFD2FF91F4DBFE608446FDE1" ID-Table-UUID="62FE660AFFD2FF91F4DBFE608446FDE1" box="[151,1653,504,530]" httpUri="http://table.plazi.org/id/62FE660AFFD2FF91F4DBFE608446FDE1" pageId="13" pageNumber="516" targetBox="[160,1890,599,1721]" targetIsTable="true" targetPageId="13">
<paragraph id="363E3682FFD2FF91F4DBFE608446FDE1" blockId="13.[151,1653,504,530]" box="[151,1653,504,530]" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4DBFE6082C3FDE1" bold="true" box="[151,240,504,530]" pageId="13" pageNumber="516">Table 1.</emphasis>
Uncorrected pairwise distances between cordylid taxa for the ND2 mitochondrial gene. All values are presented in percentages (%).
</paragraph>
</caption>
<paragraph id="363E3682FFD2FF91F5D7FD6C8576F94A" pageId="13" pageNumber="516">
<table id="4481C422FFD20063F4ECFDCF8551F94A" box="[160,1890,599,1721]" gridcols="21" gridrows="20" pageId="13" pageNumber="516">
<tr id="88B134C0FFD20063F4ECFDCF8551FCF4" box="[160,1890,599,775]" gridrow="0" pageId="13" pageNumber="516" rowspan-0="1">
<th id="CB605DBCFFD20063F5D7FDCF8381FCF4" box="[411,434,599,775]" gridcol="1" gridrow="0" pageId="13" pageNumber="516">C. aridus</th>
<th id="CB605DBCFFD20063F5F0FDCF8030FCF4" box="[444,515,599,775]" gridcol="2" gridrow="0" pageId="13" pageNumber="516">
C.
<taxonomicName id="F1814D01FFD2FF91F5A0FD1C8030FD18" authorityName="Broadley &amp; Branch" authorityYear="2002" box="[492,515,644,747]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="516" phylum="Chordata" rank="species" species="beraduccii">beraduccii</taxonomicName>
</th>
<th id="CB605DBCFFD20063F641FDCF8060FCF4" box="[525,595,599,775]" gridcol="3" gridrow="0" pageId="13" pageNumber="516">C. cordylus</th>
<th id="CB605DBCFFD20063F611FDCF8097FCF4" box="[605,676,599,775]" gridcol="4" gridrow="0" pageId="13" pageNumber="516">C. imkeae</th>
<th id="CB605DBCFFD20063F6E2FDCF80C6FCF4" box="[686,757,599,775]" gridcol="5" gridrow="0" pageId="13" pageNumber="516">C. jonesii</th>
<th id="CB605DBCFFD20063F6B3FDCF8175FCF4" box="[767,838,599,775]" gridcol="6" gridrow="0" pageId="13" pageNumber="516">
C.
<taxonomicName id="F1814D01FFD2FF91F763FD148175FD18" authorityName="Laurent" authorityYear="1964" box="[815,838,652,747]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="516" phylum="Chordata" rank="species" species="machadoi">machadoi</taxonomicName>
</th>
<th id="CB605DBCFFD20063F71CFDCF81A4FCF4" box="[848,919,599,775]" gridcol="7" gridrow="0" pageId="13" pageNumber="516">C. macropholis</th>
<th id="CB605DBCFFD20063F7EDFDCF81DBFCF4" box="[929,1000,599,775]" gridcol="8" gridrow="0" pageId="13" pageNumber="516">
C.
<taxonomicName id="F1814D01FFD2FF91F79DFDFC81DBFD18" authorityName="Greenbaum, Stanley, Kusamba, Moninga, Goldberg &amp; Bursey" authorityYear="2012" box="[977,1000,612,747]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="516" phylum="Chordata" rank="species" species="marunguensis">marunguensis</taxonomicName>
</th>
<th id="CB605DBCFFD20063F7BEFDCF860AFCF4" box="[1010,1081,599,775]" gridcol="9" gridrow="0" pageId="13" pageNumber="516">C. mclachlani</th>
<th id="CB605DBCFFD20063F00FFDCF86B9FCF4" box="[1091,1162,599,775]" gridcol="10" gridrow="0" pageId="13" pageNumber="516">C. minor</th>
<th id="CB605DBCFFD20063F0D8FDCF86E9FCF4" box="[1172,1242,599,775]" gridcol="11" gridrow="0" pageId="13" pageNumber="516">
C.
<taxonomicName id="F1814D01FFD2FF91F08FFDE086E9FD18" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[1219,1242,632,747]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="516" phylum="Chordata" rank="species" species="namakuiyus">namakuiyus</taxonomicName>
</th>
<th id="CB605DBCFFD20063F0A9FDCF8718FCF4" box="[1253,1323,599,775]" gridcol="12" gridrow="0" pageId="13" pageNumber="516">C. niger</th>
<th id="CB605DBCFFD20063F179FDCF874FFCF4" box="[1333,1404,599,775]" gridcol="13" gridrow="0" pageId="13" pageNumber="516">C. oelofseni</th>
<th id="CB605DBCFFD20063F1CAFDCF87CBFCF4" box="[1414,1528,599,775]" colspan="2" colspanRight="1" gridcol="14" gridrow="0" pageId="13" pageNumber="516">
C.
<taxonomicName id="F1814D01FFD2FF91F1EBFDE1878DFD19" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[1447,1470,633,746]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="516" phylum="Chordata" rank="species" species="phonolithos">
<emphasis id="04F5EA90FFD2FF91F1EBFDE1878DFD19" box="[1447,1470,633,746]" italics="true" pageId="13" pageNumber="516">phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD2FF91F189FD7087EFFCF3" bold="true" box="[1477,1500,744,768]" pageId="13" pageNumber="516">sp</emphasis>
.
<emphasis id="04F5EA90FFD2FF91F189FD2087EFFD2D" bold="true" box="[1477,1500,696,734]" pageId="13" pageNumber="516">nov</emphasis>
.
</th>
<th id="CB605DBCFFD20063F24BFDCF842DFCF4" box="[1543,1566,599,775]" gridcol="16" gridrow="0" pageId="13" pageNumber="516">C. rhodesianus</th>
<th id="CB605DBCFFD20063F264FDCF845CFCF4" box="[1576,1647,599,775]" gridcol="17" gridrow="0" pageId="13" pageNumber="516">C. tropidosternum</th>
<th id="CB605DBCFFD20063F235FDCF84F3FCF4" box="[1657,1728,599,775]" gridcol="18" gridrow="0" pageId="13" pageNumber="516">C. ukingensis</th>
<th id="CB605DBCFFD20063F286FDCF8522FCF4" box="[1738,1809,599,775]" gridcol="19" gridrow="0" pageId="13" pageNumber="516">
C.
<taxonomicName id="F1814D01FFD2FF91F2B6FD328522FD19" baseAuthorityName="Reichenow" baseAuthorityYear="1887" box="[1786,1809,682,746]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="516" phylum="Chordata" rank="species" species="vittifer">vittifer</taxonomicName>
</th>
<th id="CB605DBCFFD20063F357FDCF8551FCF4" box="[1819,1890,599,775]" gridcol="20" gridrow="0" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F307FD6A8551FD01" box="[1867,1890,751,774]" italics="true" pageId="13" pageNumber="516">O.</emphasis>
<taxonomicName id="F1814D01FFD2FF91F307FDF48551FD1A" baseAuthorityName="Boie" baseAuthorityYear="1828" box="[1867,1890,620,745]" class="Reptilia" family="Cordylidae" genus="Ouroborus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="516" phylum="Chordata" rank="species" species="cataphractus">
<emphasis id="04F5EA90FFD2FF91F307FDF48551FD1A" box="[1867,1890,620,745]" italics="true" pageId="13" pageNumber="516">cataphractus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="88B134C0FFD20063F4ECFCB68551FCB6" box="[160,1890,814,837]" gridrow="1" pageId="13" pageNumber="516">
<th id="CB605DBCFFD20063F4ECFCB68551FCB6" box="[160,1890,814,837]" colspan="21" colspanRight="20" gridcol="0" gridrow="1" pageId="13" pageNumber="516">C. aridus</th>
</tr>
<tr id="88B134C0FFD20063F4ECFCC78551FC85" box="[160,1890,863,886]" gridrow="2" pageId="13" pageNumber="516" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="CB605DBCFFD20063F4ECFCC783B3FC85" box="[160,384,863,886]" gridcol="0" gridrow="2" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFCC78310FC85" box="[160,291,863,886]" italics="true" pageId="13" pageNumber="516">C. beraduccii</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FCC78030FC85" box="[444,515,863,886]" gridcol="2" gridrow="2" pageId="13" pageNumber="516">20.1</td>
</tr>
<tr id="88B134C0FFD20063F4ECFC088551FC54" box="[160,1890,912,935]" gridrow="3" pageId="13" pageNumber="516" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="CB605DBCFFD20063F4ECFC0883B3FC54" box="[160,384,912,935]" gridcol="0" gridrow="3" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFC08833DFC54" box="[160,270,912,935]" italics="true" pageId="13" pageNumber="516">C. cordylus</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FC088030FC54" box="[444,515,912,935]" gridcol="2" gridrow="3" pageId="13" pageNumber="516">14.3</td>
<td id="CB605DBCFFD20063F641FC088060FC54" box="[525,595,912,935]" gridcol="3" gridrow="3" pageId="13" pageNumber="516">16</td>
</tr>
<tr id="88B134C0FFD20063F4ECFC598551FC2B" box="[160,1890,961,984]" gridrow="4" pageId="13" pageNumber="516" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="CB605DBCFFD20063F4ECFC5983B3FC2B" box="[160,384,961,984]" gridcol="0" gridrow="4" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFC598333FC2B" box="[160,256,961,984]" italics="true" pageId="13" pageNumber="516">C. imkeae</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FC598030FC2B" box="[444,515,961,984]" gridcol="2" gridrow="4" pageId="13" pageNumber="516">12.2</td>
<td id="CB605DBCFFD20063F641FC598060FC2B" box="[525,595,961,984]" gridcol="3" gridrow="4" pageId="13" pageNumber="516">19.6</td>
<td id="CB605DBCFFD20063F611FC598097FC2B" box="[605,676,961,984]" gridcol="4" gridrow="4" pageId="13" pageNumber="516">11.7</td>
</tr>
<tr id="88B134C0FFD20063F4ECFC6B8551FBF9" box="[160,1890,1011,1034]" gridrow="5" pageId="13" pageNumber="516" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="CB605DBCFFD20063F4ECFC6B83B3FBF9" box="[160,384,1011,1034]" gridcol="0" gridrow="5" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFC6B82CFFBF9" box="[160,252,1011,1034]" italics="true" pageId="13" pageNumber="516">C. jonesii</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FC6B8030FBF9" box="[444,515,1011,1034]" gridcol="2" gridrow="5" pageId="13" pageNumber="516">19.2</td>
<td id="CB605DBCFFD20063F641FC6B8060FBF9" box="[525,595,1011,1034]" gridcol="3" gridrow="5" pageId="13" pageNumber="516">14.7</td>
<td id="CB605DBCFFD20063F611FC6B8097FBF9" box="[605,676,1011,1034]" gridcol="4" gridrow="5" pageId="13" pageNumber="516">15.4</td>
<td id="CB605DBCFFD20063F6E2FC6B80C6FBF9" box="[686,757,1011,1034]" gridcol="5" gridrow="5" pageId="13" pageNumber="516">18.3</td>
</tr>
<tr id="88B134C0FFD20063F4ECFBBC8551FBC8" box="[160,1890,1060,1083]" gridrow="6" pageId="13" pageNumber="516" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="CB605DBCFFD20063F4ECFBBC83B3FBC8" box="[160,384,1060,1083]" gridcol="0" gridrow="6" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFBBC8328FBC8" box="[160,283,1060,1083]" italics="true" pageId="13" pageNumber="516">C. machadoi</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FBBC8030FBC8" box="[444,515,1060,1083]" gridcol="2" gridrow="6" pageId="13" pageNumber="516">18</td>
<td id="CB605DBCFFD20063F641FBBC8060FBC8" box="[525,595,1060,1083]" gridcol="3" gridrow="6" pageId="13" pageNumber="516">15.6</td>
<td id="CB605DBCFFD20063F611FBBC8097FBC8" box="[605,676,1060,1083]" gridcol="4" gridrow="6" pageId="13" pageNumber="516">16.7</td>
<td id="CB605DBCFFD20063F6E2FBBC80C6FBC8" box="[686,757,1060,1083]" gridcol="5" gridrow="6" pageId="13" pageNumber="516">16.9</td>
<td id="CB605DBCFFD20063F6B3FBBC8175FBC8" box="[767,838,1060,1083]" gridcol="6" gridrow="6" pageId="13" pageNumber="516">15.4</td>
</tr>
<tr id="88B134C0FFD20063F4ECFBCD8551FB9F" box="[160,1890,1109,1132]" gridrow="7" pageId="13" pageNumber="516" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1" rowspan-8="1" rowspan-9="1">
<th id="CB605DBCFFD20063F4ECFBCD83B3FB9F" box="[160,384,1109,1132]" gridcol="0" gridrow="7" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFBCD8300FB9F" box="[160,307,1109,1132]" italics="true" pageId="13" pageNumber="516">C. macropholis</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FBCD8030FB9F" box="[444,515,1109,1132]" gridcol="2" gridrow="7" pageId="13" pageNumber="516">16.2</td>
<td id="CB605DBCFFD20063F641FBCD8060FB9F" box="[525,595,1109,1132]" gridcol="3" gridrow="7" pageId="13" pageNumber="516">18.5</td>
<td id="CB605DBCFFD20063F611FBCD8097FB9F" box="[605,676,1109,1132]" gridcol="4" gridrow="7" pageId="13" pageNumber="516">12.4</td>
<td id="CB605DBCFFD20063F6E2FBCD80C6FB9F" box="[686,757,1109,1132]" gridcol="5" gridrow="7" pageId="13" pageNumber="516">13.5</td>
<td id="CB605DBCFFD20063F6B3FBCD8175FB9F" box="[767,838,1109,1132]" gridcol="6" gridrow="7" pageId="13" pageNumber="516">16</td>
<td id="CB605DBCFFD20063F71CFBCD81A4FB9F" box="[848,919,1109,1132]" gridcol="7" gridrow="7" pageId="13" pageNumber="516">16.4</td>
</tr>
<tr id="88B134C0FFD20063F4ECFB1E8551FB6E" box="[160,1890,1158,1181]" gridrow="8" pageId="13" pageNumber="516" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1" rowspan-9="1">
<th id="CB605DBCFFD20063F4ECFB1E83B3FB6E" box="[160,384,1158,1181]" gridcol="0" gridrow="8" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFB1E8370FB6E" box="[160,323,1158,1181]" italics="true" pageId="13" pageNumber="516">C. marunguensis</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FB1E8030FB6E" box="[444,515,1158,1181]" gridcol="2" gridrow="8" pageId="13" pageNumber="516">18.4</td>
<td id="CB605DBCFFD20063F641FB1E8060FB6E" box="[525,595,1158,1181]" gridcol="3" gridrow="8" pageId="13" pageNumber="516">12.1</td>
<td id="CB605DBCFFD20063F611FB1E8097FB6E" box="[605,676,1158,1181]" gridcol="4" gridrow="8" pageId="13" pageNumber="516">14.5</td>
<td id="CB605DBCFFD20063F6E2FB1E80C6FB6E" box="[686,757,1158,1181]" gridcol="5" gridrow="8" pageId="13" pageNumber="516">17.3</td>
<td id="CB605DBCFFD20063F6B3FB1E8175FB6E" box="[767,838,1158,1181]" gridcol="6" gridrow="8" pageId="13" pageNumber="516">15.5</td>
<td id="CB605DBCFFD20063F71CFB1E81A4FB6E" box="[848,919,1158,1181]" gridcol="7" gridrow="8" pageId="13" pageNumber="516">15</td>
<td id="CB605DBCFFD20063F7EDFB1E81DBFB6E" box="[929,1000,1158,1181]" gridcol="8" gridrow="8" pageId="13" pageNumber="516">16.4</td>
</tr>
<tr id="88B134C0FFD20063F4ECFB2F8551FB3D" box="[160,1890,1207,1230]" gridrow="9" pageId="13" pageNumber="516" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECFB2F83B3FB3D" box="[160,384,1207,1230]" gridcol="0" gridrow="9" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFB2F8314FB3D" box="[160,295,1207,1230]" italics="true" pageId="13" pageNumber="516">C. mclachlani</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FB2F8030FB3D" box="[444,515,1207,1230]" gridcol="2" gridrow="9" pageId="13" pageNumber="516">15.6</td>
<td id="CB605DBCFFD20063F641FB2F8060FB3D" box="[525,595,1207,1230]" gridcol="3" gridrow="9" pageId="13" pageNumber="516">18.4</td>
<td id="CB605DBCFFD20063F611FB2F8097FB3D" box="[605,676,1207,1230]" gridcol="4" gridrow="9" pageId="13" pageNumber="516">13.4</td>
<td id="CB605DBCFFD20063F6E2FB2F80C6FB3D" box="[686,757,1207,1230]" gridcol="5" gridrow="9" pageId="13" pageNumber="516">14.7</td>
<td id="CB605DBCFFD20063F6B3FB2F8175FB3D" box="[767,838,1207,1230]" gridcol="6" gridrow="9" pageId="13" pageNumber="516">18.7</td>
<td id="CB605DBCFFD20063F71CFB2F81A4FB3D" box="[848,919,1207,1230]" gridcol="7" gridrow="9" pageId="13" pageNumber="516">18.3</td>
<td id="CB605DBCFFD20063F7EDFB2F81DBFB3D" box="[929,1000,1207,1230]" gridcol="8" gridrow="9" pageId="13" pageNumber="516">13</td>
<td id="CB605DBCFFD20063F7BEFB2F860AFB3D" box="[1010,1081,1207,1230]" gridcol="9" gridrow="9" pageId="13" pageNumber="516">15.4</td>
</tr>
<tr id="88B134C0FFD20063F4ECFB708551FB0C" box="[160,1890,1256,1279]" gridrow="10" pageId="13" pageNumber="516" rowspan-1="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECFB7083B3FB0C" box="[160,384,1256,1279]" gridcol="0" gridrow="10" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFB7082C6FB0C" box="[160,245,1256,1279]" italics="true" pageId="13" pageNumber="516">C. minor</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FB708030FB0C" box="[444,515,1256,1279]" gridcol="2" gridrow="10" pageId="13" pageNumber="516">4.1</td>
<td id="CB605DBCFFD20063F641FB708060FB0C" box="[525,595,1256,1279]" gridcol="3" gridrow="10" pageId="13" pageNumber="516">17.9</td>
<td id="CB605DBCFFD20063F611FB708097FB0C" box="[605,676,1256,1279]" gridcol="4" gridrow="10" pageId="13" pageNumber="516">11.6</td>
<td id="CB605DBCFFD20063F6E2FB7080C6FB0C" box="[686,757,1256,1279]" gridcol="5" gridrow="10" pageId="13" pageNumber="516">9.7</td>
<td id="CB605DBCFFD20063F6B3FB708175FB0C" box="[767,838,1256,1279]" gridcol="6" gridrow="10" pageId="13" pageNumber="516">17</td>
<td id="CB605DBCFFD20063F71CFB7081A4FB0C" box="[848,919,1256,1279]" gridcol="7" gridrow="10" pageId="13" pageNumber="516">16.6</td>
<td id="CB605DBCFFD20063F7EDFB7081DBFB0C" box="[929,1000,1256,1279]" gridcol="8" gridrow="10" pageId="13" pageNumber="516">13.1</td>
<td id="CB605DBCFFD20063F7BEFB70860AFB0C" box="[1010,1081,1256,1279]" gridcol="9" gridrow="10" pageId="13" pageNumber="516">16</td>
<td id="CB605DBCFFD20063F00FFB7086B9FB0C" box="[1091,1162,1256,1279]" gridcol="10" gridrow="10" pageId="13" pageNumber="516">12.3</td>
</tr>
<tr id="88B134C0FFD20063F4ECFA818551FAC3" box="[160,1890,1305,1328]" gridrow="11" pageId="13" pageNumber="516" rowspan-1="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECFA8183B3FAC3" box="[160,384,1305,1328]" gridcol="0" gridrow="11" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFA81831CFAC3" box="[160,303,1305,1328]" italics="true" pageId="13" pageNumber="516">C. namakuiyus</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FA818030FAC3" box="[444,515,1305,1328]" gridcol="2" gridrow="11" pageId="13" pageNumber="516">18</td>
<td id="CB605DBCFFD20063F641FA818060FAC3" box="[525,595,1305,1328]" gridcol="3" gridrow="11" pageId="13" pageNumber="516">15.4</td>
<td id="CB605DBCFFD20063F611FA818097FAC3" box="[605,676,1305,1328]" gridcol="4" gridrow="11" pageId="13" pageNumber="516">15.2</td>
<td id="CB605DBCFFD20063F6E2FA8180C6FAC3" box="[686,757,1305,1328]" gridcol="5" gridrow="11" pageId="13" pageNumber="516">15.8</td>
<td id="CB605DBCFFD20063F6B3FA818175FAC3" box="[767,838,1305,1328]" gridcol="6" gridrow="11" pageId="13" pageNumber="516">14.3</td>
<td id="CB605DBCFFD20063F71CFA8181A4FAC3" box="[848,919,1305,1328]" gridcol="7" gridrow="11" pageId="13" pageNumber="516">5.8</td>
<td id="CB605DBCFFD20063F7EDFA8181DBFAC3" box="[929,1000,1305,1328]" gridcol="8" gridrow="11" pageId="13" pageNumber="516">16.1</td>
<td id="CB605DBCFFD20063F7BEFA81860AFAC3" box="[1010,1081,1305,1328]" gridcol="9" gridrow="11" pageId="13" pageNumber="516">14.7</td>
<td id="CB605DBCFFD20063F00FFA8186B9FAC3" box="[1091,1162,1305,1328]" gridcol="10" gridrow="11" pageId="13" pageNumber="516">17.5</td>
<td id="CB605DBCFFD20063F0D8FA8186E9FAC3" box="[1172,1242,1305,1328]" gridcol="11" gridrow="11" pageId="13" pageNumber="516">15.1</td>
</tr>
<tr id="88B134C0FFD20063F4ECFAD28551FA92" box="[160,1890,1354,1377]" gridrow="12" pageId="13" pageNumber="516" rowspan-1="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECFAD283B3FA92" box="[160,384,1354,1377]" gridcol="0" gridrow="12" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFAD282DCFA92" box="[160,239,1354,1377]" italics="true" pageId="13" pageNumber="516">C. niger</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FAD28030FA92" box="[444,515,1354,1377]" gridcol="2" gridrow="12" pageId="13" pageNumber="516">13.3</td>
<td id="CB605DBCFFD20063F641FAD28060FA92" box="[525,595,1354,1377]" gridcol="3" gridrow="12" pageId="13" pageNumber="516">17.4</td>
<td id="CB605DBCFFD20063F611FAD28097FA92" box="[605,676,1354,1377]" gridcol="4" gridrow="12" pageId="13" pageNumber="516">10</td>
<td id="CB605DBCFFD20063F6E2FAD280C6FA92" box="[686,757,1354,1377]" gridcol="5" gridrow="12" pageId="13" pageNumber="516">12.1</td>
<td id="CB605DBCFFD20063F6B3FAD28175FA92" box="[767,838,1354,1377]" gridcol="6" gridrow="12" pageId="13" pageNumber="516">15.5</td>
<td id="CB605DBCFFD20063F71CFAD281A4FA92" box="[848,919,1354,1377]" gridcol="7" gridrow="12" pageId="13" pageNumber="516">15.6</td>
<td id="CB605DBCFFD20063F7EDFAD281DBFA92" box="[929,1000,1354,1377]" gridcol="8" gridrow="12" pageId="13" pageNumber="516">11.1</td>
<td id="CB605DBCFFD20063F7BEFAD2860AFA92" box="[1010,1081,1354,1377]" gridcol="9" gridrow="12" pageId="13" pageNumber="516">15.7</td>
<td id="CB605DBCFFD20063F00FFAD286B9FA92" box="[1091,1162,1354,1377]" gridcol="10" gridrow="12" pageId="13" pageNumber="516">12</td>
<td id="CB605DBCFFD20063F0D8FAD286E9FA92" box="[1172,1242,1354,1377]" gridcol="11" gridrow="12" pageId="13" pageNumber="516">10.5</td>
<td id="CB605DBCFFD20063F0A9FAD28718FA92" box="[1253,1323,1354,1377]" gridcol="12" gridrow="12" pageId="13" pageNumber="516">14.1</td>
</tr>
<tr id="88B134C0FFD20063F4ECFAE48551FA60" box="[160,1890,1404,1427]" gridrow="13" pageId="13" pageNumber="516" rowspan-1="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECFAE483B3FA60" box="[160,384,1404,1427]" gridcol="0" gridrow="13" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFAE48320FA60" box="[160,275,1404,1427]" italics="true" pageId="13" pageNumber="516">C. oelofseni</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FAE48030FA60" box="[444,515,1404,1427]" gridcol="2" gridrow="13" pageId="13" pageNumber="516">13.9</td>
<td id="CB605DBCFFD20063F641FAE48060FA60" box="[525,595,1404,1427]" gridcol="3" gridrow="13" pageId="13" pageNumber="516">15.8</td>
<td id="CB605DBCFFD20063F611FAE48097FA60" box="[605,676,1404,1427]" gridcol="4" gridrow="13" pageId="13" pageNumber="516">7.1</td>
<td id="CB605DBCFFD20063F6E2FAE480C6FA60" box="[686,757,1404,1427]" gridcol="5" gridrow="13" pageId="13" pageNumber="516">12.1</td>
<td id="CB605DBCFFD20063F6B3FAE48175FA60" box="[767,838,1404,1427]" gridcol="6" gridrow="13" pageId="13" pageNumber="516">15.2</td>
<td id="CB605DBCFFD20063F71CFAE481A4FA60" box="[848,919,1404,1427]" gridcol="7" gridrow="13" pageId="13" pageNumber="516">15.1</td>
<td id="CB605DBCFFD20063F7EDFAE481DBFA60" box="[929,1000,1404,1427]" gridcol="8" gridrow="13" pageId="13" pageNumber="516">10.2</td>
<td id="CB605DBCFFD20063F7BEFAE4860AFA60" box="[1010,1081,1404,1427]" gridcol="9" gridrow="13" pageId="13" pageNumber="516">14.5</td>
<td id="CB605DBCFFD20063F00FFAE486B9FA60" box="[1091,1162,1404,1427]" gridcol="10" gridrow="13" pageId="13" pageNumber="516">12.3</td>
<td id="CB605DBCFFD20063F0D8FAE486E9FA60" box="[1172,1242,1404,1427]" gridcol="11" gridrow="13" pageId="13" pageNumber="516">10.8</td>
<td id="CB605DBCFFD20063F0A9FAE48718FA60" box="[1253,1323,1404,1427]" gridcol="12" gridrow="13" pageId="13" pageNumber="516">14.5</td>
<td id="CB605DBCFFD20063F179FAE4874FFA60" box="[1333,1404,1404,1427]" gridcol="13" gridrow="13" pageId="13" pageNumber="516">9.1</td>
</tr>
<tr id="88B134C0FFD20063F4ECFA358551FA37" box="[160,1890,1453,1476]" gridrow="14" pageId="13" pageNumber="516" rowspan-1="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECFA3583B3FA37" box="[160,384,1453,1476]" gridcol="0" gridrow="14" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFA35831EFA37" box="[160,301,1453,1476]" italics="true" pageId="13" pageNumber="516">C. phonolithos</emphasis>
<emphasis id="04F5EA90FFD2FF91F578FA3583B3FA37" bold="true" box="[308,384,1453,1476]" pageId="13" pageNumber="516">sp. nov.</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FA358030FA37" box="[444,515,1453,1476]" gridcol="2" gridrow="14" pageId="13" pageNumber="516">17.9</td>
<td id="CB605DBCFFD20063F641FA358060FA37" box="[525,595,1453,1476]" gridcol="3" gridrow="14" pageId="13" pageNumber="516">14.9</td>
<td id="CB605DBCFFD20063F611FA358097FA37" box="[605,676,1453,1476]" gridcol="4" gridrow="14" pageId="13" pageNumber="516">16.6</td>
<td id="CB605DBCFFD20063F6E2FA3580C6FA37" box="[686,757,1453,1476]" gridcol="5" gridrow="14" pageId="13" pageNumber="516">15.2</td>
<td id="CB605DBCFFD20063F6B3FA358175FA37" box="[767,838,1453,1476]" gridcol="6" gridrow="14" pageId="13" pageNumber="516">14.8</td>
<td id="CB605DBCFFD20063F71CFA3581A4FA37" box="[848,919,1453,1476]" gridcol="7" gridrow="14" pageId="13" pageNumber="516">7.3</td>
<td id="CB605DBCFFD20063F7EDFA3581DBFA37" box="[929,1000,1453,1476]" gridcol="8" gridrow="14" pageId="13" pageNumber="516">16.8</td>
<td id="CB605DBCFFD20063F7BEFA35860AFA37" box="[1010,1081,1453,1476]" gridcol="9" gridrow="14" pageId="13" pageNumber="516">13.7</td>
<td id="CB605DBCFFD20063F00FFA3586B9FA37" box="[1091,1162,1453,1476]" gridcol="10" gridrow="14" pageId="13" pageNumber="516">18.4</td>
<td id="CB605DBCFFD20063F0D8FA3586E9FA37" box="[1172,1242,1453,1476]" gridcol="11" gridrow="14" pageId="13" pageNumber="516">15</td>
<td id="CB605DBCFFD20063F0A9FA358718FA37" box="[1253,1323,1453,1476]" gridcol="12" gridrow="14" pageId="13" pageNumber="516">6.7</td>
<td id="CB605DBCFFD20063F179FA35874FFA37" box="[1333,1404,1453,1476]" gridcol="13" gridrow="14" pageId="13" pageNumber="516">15.2</td>
<td id="CB605DBCFFD20063F1CAFA358783FA37" box="[1414,1456,1453,1476]" gridcol="14" gridrow="14" pageId="13" pageNumber="516">16.1</td>
</tr>
<tr id="88B134C0FFD20063F4ECFA468551FA06" box="[160,1890,1502,1525]" gridrow="15" pageId="13" pageNumber="516" rowspan-1="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECFA4683B3FA06" box="[160,384,1502,1525]" gridcol="0" gridrow="15" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECFA468302FA06" box="[160,305,1502,1525]" italics="true" pageId="13" pageNumber="516">C. rhodesianus</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0FA468030FA06" box="[444,515,1502,1525]" gridcol="2" gridrow="15" pageId="13" pageNumber="516">20.3</td>
<td id="CB605DBCFFD20063F641FA468060FA06" box="[525,595,1502,1525]" gridcol="3" gridrow="15" pageId="13" pageNumber="516">14.8</td>
<td id="CB605DBCFFD20063F611FA468097FA06" box="[605,676,1502,1525]" gridcol="4" gridrow="15" pageId="13" pageNumber="516">15.4</td>
<td id="CB605DBCFFD20063F6E2FA4680C6FA06" box="[686,757,1502,1525]" gridcol="5" gridrow="15" pageId="13" pageNumber="516">18.5</td>
<td id="CB605DBCFFD20063F6B3FA468175FA06" box="[767,838,1502,1525]" gridcol="6" gridrow="15" pageId="13" pageNumber="516">11.7</td>
<td id="CB605DBCFFD20063F71CFA4681A4FA06" box="[848,919,1502,1525]" gridcol="7" gridrow="15" pageId="13" pageNumber="516">14.1</td>
<td id="CB605DBCFFD20063F7EDFA4681DBFA06" box="[929,1000,1502,1525]" gridcol="8" gridrow="15" pageId="13" pageNumber="516">16</td>
<td id="CB605DBCFFD20063F7BEFA46860AFA06" box="[1010,1081,1502,1525]" gridcol="9" gridrow="15" pageId="13" pageNumber="516">13.8</td>
<td id="CB605DBCFFD20063F00FFA4686B9FA06" box="[1091,1162,1502,1525]" gridcol="10" gridrow="15" pageId="13" pageNumber="516">18.2</td>
<td id="CB605DBCFFD20063F0D8FA4686E9FA06" box="[1172,1242,1502,1525]" gridcol="11" gridrow="15" pageId="13" pageNumber="516">17.8</td>
<td id="CB605DBCFFD20063F0A9FA468718FA06" box="[1253,1323,1502,1525]" gridcol="12" gridrow="15" pageId="13" pageNumber="516">13.8</td>
<td id="CB605DBCFFD20063F179FA46874FFA06" box="[1333,1404,1502,1525]" gridcol="13" gridrow="15" pageId="13" pageNumber="516">15.3</td>
<td id="CB605DBCFFD20063F1CAFA468783FA06" box="[1414,1456,1502,1525]" gridcol="14" gridrow="15" pageId="13" pageNumber="516">15</td>
<td id="CB605DBCFFD20063F19BFA46842DFA06" box="[1495,1566,1502,1525]" colspan="2" colspanRight="1" gridcol="15" gridrow="15" pageId="13" pageNumber="516">13.8</td>
</tr>
<tr id="88B134C0FFD20063F4ECF9978551F9D5" box="[160,1890,1551,1574]" gridrow="16" pageId="13" pageNumber="516" rowspan-1="1" rowspan-18="1" rowspan-19="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECF99783B3F9D5" box="[160,384,1551,1574]" gridcol="0" gridrow="16" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECF997837CF9D5" box="[160,335,1551,1574]" italics="true" pageId="13" pageNumber="516">C. tropidosternum</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0F9978030F9D5" box="[444,515,1551,1574]" gridcol="2" gridrow="16" pageId="13" pageNumber="516">18.7</td>
<td id="CB605DBCFFD20063F641F9978060F9D5" box="[525,595,1551,1574]" gridcol="3" gridrow="16" pageId="13" pageNumber="516">13.1</td>
<td id="CB605DBCFFD20063F611F9978097F9D5" box="[605,676,1551,1574]" gridcol="4" gridrow="16" pageId="13" pageNumber="516">13.8</td>
<td id="CB605DBCFFD20063F6E2F99780C6F9D5" box="[686,757,1551,1574]" gridcol="5" gridrow="16" pageId="13" pageNumber="516">16.8</td>
<td id="CB605DBCFFD20063F6B3F9978175F9D5" box="[767,838,1551,1574]" gridcol="6" gridrow="16" pageId="13" pageNumber="516">14.6</td>
<td id="CB605DBCFFD20063F71CF99781A4F9D5" box="[848,919,1551,1574]" gridcol="7" gridrow="16" pageId="13" pageNumber="516">14.8</td>
<td id="CB605DBCFFD20063F7EDF99781DBF9D5" box="[929,1000,1551,1574]" gridcol="8" gridrow="16" pageId="13" pageNumber="516">16.4</td>
<td id="CB605DBCFFD20063F7BEF997860AF9D5" box="[1010,1081,1551,1574]" gridcol="9" gridrow="16" pageId="13" pageNumber="516">13.6</td>
<td id="CB605DBCFFD20063F00FF99786B9F9D5" box="[1091,1162,1551,1574]" gridcol="10" gridrow="16" pageId="13" pageNumber="516">15.9</td>
<td id="CB605DBCFFD20063F0D8F99786E9F9D5" box="[1172,1242,1551,1574]" gridcol="11" gridrow="16" pageId="13" pageNumber="516">15.9</td>
<td id="CB605DBCFFD20063F0A9F9978718F9D5" box="[1253,1323,1551,1574]" gridcol="12" gridrow="16" pageId="13" pageNumber="516">14.2</td>
<td id="CB605DBCFFD20063F179F997874FF9D5" box="[1333,1404,1551,1574]" gridcol="13" gridrow="16" pageId="13" pageNumber="516">13.6</td>
<td id="CB605DBCFFD20063F1CAF9978783F9D5" box="[1414,1456,1551,1574]" gridcol="14" gridrow="16" pageId="13" pageNumber="516">14.5</td>
<td id="CB605DBCFFD20063F19BF997842DF9D5" box="[1495,1566,1551,1574]" colspan="2" colspanRight="1" gridcol="15" gridrow="16" pageId="13" pageNumber="516">14.9</td>
<td id="CB605DBCFFD20063F264F997845CF9D5" box="[1576,1647,1551,1574]" gridcol="17" gridrow="16" pageId="13" pageNumber="516">13.9</td>
</tr>
<tr id="88B134C0FFD20063F4ECF9D88551F9A4" box="[160,1890,1600,1623]" gridrow="17" pageId="13" pageNumber="516" rowspan-1="1" rowspan-16="1" rowspan-19="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECF9D883B3F9A4" box="[160,384,1600,1623]" gridcol="0" gridrow="17" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECF9D88312F9A4" box="[160,289,1600,1623]" italics="true" pageId="13" pageNumber="516">C. ukingensis</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0F9D88030F9A4" box="[444,515,1600,1623]" gridcol="2" gridrow="17" pageId="13" pageNumber="516">19.3</td>
<td id="CB605DBCFFD20063F641F9D88060F9A4" box="[525,595,1600,1623]" gridcol="3" gridrow="17" pageId="13" pageNumber="516">13.6</td>
<td id="CB605DBCFFD20063F611F9D88097F9A4" box="[605,676,1600,1623]" gridcol="4" gridrow="17" pageId="13" pageNumber="516">15</td>
<td id="CB605DBCFFD20063F6E2F9D880C6F9A4" box="[686,757,1600,1623]" gridcol="5" gridrow="17" pageId="13" pageNumber="516">17</td>
<td id="CB605DBCFFD20063F6B3F9D88175F9A4" box="[767,838,1600,1623]" gridcol="6" gridrow="17" pageId="13" pageNumber="516">15</td>
<td id="CB605DBCFFD20063F71CF9D881A4F9A4" box="[848,919,1600,1623]" gridcol="7" gridrow="17" pageId="13" pageNumber="516">13.6</td>
<td id="CB605DBCFFD20063F7EDF9D881DBF9A4" box="[929,1000,1600,1623]" gridcol="8" gridrow="17" pageId="13" pageNumber="516">14.8</td>
<td id="CB605DBCFFD20063F7BEF9D8860AF9A4" box="[1010,1081,1600,1623]" gridcol="9" gridrow="17" pageId="13" pageNumber="516">10.4</td>
<td id="CB605DBCFFD20063F00FF9D886B9F9A4" box="[1091,1162,1600,1623]" gridcol="10" gridrow="17" pageId="13" pageNumber="516">15.4</td>
<td id="CB605DBCFFD20063F0D8F9D886E9F9A4" box="[1172,1242,1600,1623]" gridcol="11" gridrow="17" pageId="13" pageNumber="516">15.5</td>
<td id="CB605DBCFFD20063F0A9F9D88718F9A4" box="[1253,1323,1600,1623]" gridcol="12" gridrow="17" pageId="13" pageNumber="516">13.7</td>
<td id="CB605DBCFFD20063F179F9D8874FF9A4" box="[1333,1404,1600,1623]" gridcol="13" gridrow="17" pageId="13" pageNumber="516">14.7</td>
<td id="CB605DBCFFD20063F1CAF9D88783F9A4" box="[1414,1456,1600,1623]" gridcol="14" gridrow="17" pageId="13" pageNumber="516">14.5</td>
<td id="CB605DBCFFD20063F19BF9D887CBF9A4" box="[1495,1528,1600,1623]" gridcol="15" gridrow="17" pageId="13" pageNumber="516">14</td>
<td id="CB605DBCFFD20063F264F9D8845CF9A4" box="[1576,1647,1600,1623]" gridcol="17" gridrow="17" pageId="13" pageNumber="516">15.5</td>
<td id="CB605DBCFFD20063F235F9D884F3F9A4" box="[1657,1728,1600,1623]" gridcol="18" gridrow="17" pageId="13" pageNumber="516">14.4</td>
</tr>
<tr id="88B134C0FFD20063F4ECF9E98551F97B" box="[160,1890,1649,1672]" gridrow="18" pageId="13" pageNumber="516" rowspan-1="1" rowspan-20="1">
<th id="CB605DBCFFD20063F4ECF9E983B3F97B" box="[160,384,1649,1672]" gridcol="0" gridrow="18" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECF9E982CFF97B" box="[160,252,1649,1672]" italics="true" pageId="13" pageNumber="516">C. vittifer</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0F9E98030F97B" box="[444,515,1649,1672]" gridcol="2" gridrow="18" pageId="13" pageNumber="516">20</td>
<td id="CB605DBCFFD20063F641F9E98060F97B" box="[525,595,1649,1672]" gridcol="3" gridrow="18" pageId="13" pageNumber="516">14.6</td>
<td id="CB605DBCFFD20063F611F9E98097F97B" box="[605,676,1649,1672]" gridcol="4" gridrow="18" pageId="13" pageNumber="516">16.5</td>
<td id="CB605DBCFFD20063F6E2F9E980C6F97B" box="[686,757,1649,1672]" gridcol="5" gridrow="18" pageId="13" pageNumber="516">18.2</td>
<td id="CB605DBCFFD20063F6B3F9E98175F97B" box="[767,838,1649,1672]" gridcol="6" gridrow="18" pageId="13" pageNumber="516">15</td>
<td id="CB605DBCFFD20063F71CF9E981A4F97B" box="[848,919,1649,1672]" gridcol="7" gridrow="18" pageId="13" pageNumber="516">15.5</td>
<td id="CB605DBCFFD20063F7EDF9E981DBF97B" box="[929,1000,1649,1672]" gridcol="8" gridrow="18" pageId="13" pageNumber="516">18.2</td>
<td id="CB605DBCFFD20063F7BEF9E9860AF97B" box="[1010,1081,1649,1672]" gridcol="9" gridrow="18" pageId="13" pageNumber="516">14.8</td>
<td id="CB605DBCFFD20063F00FF9E986B9F97B" box="[1091,1162,1649,1672]" gridcol="10" gridrow="18" pageId="13" pageNumber="516">17.6</td>
<td id="CB605DBCFFD20063F0D8F9E986E9F97B" box="[1172,1242,1649,1672]" gridcol="11" gridrow="18" pageId="13" pageNumber="516">17.3</td>
<td id="CB605DBCFFD20063F0A9F9E98718F97B" box="[1253,1323,1649,1672]" gridcol="12" gridrow="18" pageId="13" pageNumber="516">14.7</td>
<td id="CB605DBCFFD20063F179F9E9874FF97B" box="[1333,1404,1649,1672]" gridcol="13" gridrow="18" pageId="13" pageNumber="516">16.7</td>
<td id="CB605DBCFFD20063F1CAF9E98783F97B" box="[1414,1456,1649,1672]" gridcol="14" gridrow="18" pageId="13" pageNumber="516">16.1</td>
<td id="CB605DBCFFD20063F19BF9E9842DF97B" box="[1495,1566,1649,1672]" colspan="2" colspanRight="1" gridcol="15" gridrow="18" pageId="13" pageNumber="516">16.8</td>
<td id="CB605DBCFFD20063F264F9E9845CF97B" box="[1576,1647,1649,1672]" gridcol="17" gridrow="18" pageId="13" pageNumber="516">15.2</td>
<td id="CB605DBCFFD20063F235F9E984F3F97B" box="[1657,1728,1649,1672]" gridcol="18" gridrow="18" pageId="13" pageNumber="516">15.1</td>
<td id="CB605DBCFFD20063F286F9E98522F97B" box="[1738,1809,1649,1672]" gridcol="19" gridrow="18" pageId="13" pageNumber="516">16.1</td>
</tr>
<tr id="88B134C0FFD20063F4ECF93A8551F94A" box="[160,1890,1698,1721]" gridrow="19" pageId="13" pageNumber="516" rowspan-1="1">
<th id="CB605DBCFFD20063F4ECF93A83B3F94A" box="[160,384,1698,1721]" gridcol="0" gridrow="19" pageId="13" pageNumber="516">
<emphasis id="04F5EA90FFD2FF91F4ECF93A8308F94A" box="[160,315,1698,1721]" italics="true" pageId="13" pageNumber="516">O. cataphractus</emphasis>
</th>
<td id="CB605DBCFFD20063F5F0F93A8030F94A" box="[444,515,1698,1721]" gridcol="2" gridrow="19" pageId="13" pageNumber="516">20.4</td>
<td id="CB605DBCFFD20063F641F93A8060F94A" box="[525,595,1698,1721]" gridcol="3" gridrow="19" pageId="13" pageNumber="516">17.3</td>
<td id="CB605DBCFFD20063F611F93A8097F94A" box="[605,676,1698,1721]" gridcol="4" gridrow="19" pageId="13" pageNumber="516">15.8</td>
<td id="CB605DBCFFD20063F6E2F93A80C6F94A" box="[686,757,1698,1721]" gridcol="5" gridrow="19" pageId="13" pageNumber="516">14.4</td>
<td id="CB605DBCFFD20063F6B3F93A8175F94A" box="[767,838,1698,1721]" gridcol="6" gridrow="19" pageId="13" pageNumber="516">17.2</td>
<td id="CB605DBCFFD20063F71CF93A81A4F94A" box="[848,919,1698,1721]" gridcol="7" gridrow="19" pageId="13" pageNumber="516">18.1</td>
<td id="CB605DBCFFD20063F7EDF93A81DBF94A" box="[929,1000,1698,1721]" gridcol="8" gridrow="19" pageId="13" pageNumber="516">16</td>
<td id="CB605DBCFFD20063F7BEF93A860AF94A" box="[1010,1081,1698,1721]" gridcol="9" gridrow="19" pageId="13" pageNumber="516">17.6</td>
<td id="CB605DBCFFD20063F00FF93A86B9F94A" box="[1091,1162,1698,1721]" gridcol="10" gridrow="19" pageId="13" pageNumber="516">15.9</td>
<td id="CB605DBCFFD20063F0D8F93A86E9F94A" box="[1172,1242,1698,1721]" gridcol="11" gridrow="19" pageId="13" pageNumber="516">13.6</td>
<td id="CB605DBCFFD20063F0A9F93A8718F94A" box="[1253,1323,1698,1721]" gridcol="12" gridrow="19" pageId="13" pageNumber="516">17</td>
<td id="CB605DBCFFD20063F179F93A874FF94A" box="[1333,1404,1698,1721]" gridcol="13" gridrow="19" pageId="13" pageNumber="516">15</td>
<td id="CB605DBCFFD20063F1CAF93A8783F94A" box="[1414,1456,1698,1721]" gridcol="14" gridrow="19" pageId="13" pageNumber="516">15</td>
<td id="CB605DBCFFD20063F19BF93A842DF94A" box="[1495,1566,1698,1721]" colspan="2" colspanRight="1" gridcol="15" gridrow="19" pageId="13" pageNumber="516">16.1</td>
<td id="CB605DBCFFD20063F264F93A845CF94A" box="[1576,1647,1698,1721]" gridcol="17" gridrow="19" pageId="13" pageNumber="516">14.4</td>
<td id="CB605DBCFFD20063F235F93A84F3F94A" box="[1657,1728,1698,1721]" gridcol="18" gridrow="19" pageId="13" pageNumber="516">16.9</td>
<td id="CB605DBCFFD20063F286F93A8522F94A" box="[1738,1809,1698,1721]" gridcol="19" gridrow="19" pageId="13" pageNumber="516">16.1</td>
<td id="CB605DBCFFD20063F357F93A8551F94A" box="[1819,1890,1698,1721]" gridcol="20" gridrow="19" pageId="13" pageNumber="516">18.5</td>
</tr>
</table>
</paragraph>
<caption id="62FE660AFFD1FF92F4DBFD998196FD90" ID-Table-UUID="62FE660AFFD1FF92F4DBFD998196FD90" httpUri="http://table.plazi.org/id/62FE660AFFD1FF92F4DBFD998196FD90" pageId="14" pageNumber="517" targetBox="[165,2014,622,1677]" targetIsTable="true" targetPageId="14">
<paragraph id="363E3682FFD1FF92F4DBFD998196FD90" blockId="14.[151,2038,513,611]" pageId="14" pageNumber="517">
<emphasis id="04F5EA90FFD1FF92F4DBFD998339FDE8" bold="true" box="[151,266,513,539]" pageId="14" pageNumber="517">TABLE 2.</emphasis>
Variation in meristic counts for specimens of
<taxonomicName id="F1814D01FFD1FF92F6B4FD9981EDFDE8" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[760,990,513,539]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD1FF92F6B4FD9981EDFDE8" box="[760,990,513,539]" italics="true" pageId="14" pageNumber="517">Cordylus phonolithos</emphasis>
</taxonomicName>
<emphasis id="04F5EA90FFD1FF92F7A9FD998608FDE8" bold="true" box="[997,1083,513,539]" pageId="14" pageNumber="517">
<taxonomicNameLabel id="1FC657EBFFD1FF92F7A9FD998608FDE8" box="[997,1083,513,539]" pageId="14" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="04F5EA90FFD1FF92F004FD99874EFDE8" box="[1096,1405,513,539]" italics="true" pageId="14" pageNumber="517">
<taxonomicName id="F1814D01FFD1FF92F004FD9986D9FDE8" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[1096,1258,513,539]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="namakuiyus">C. namakuiyus</taxonomicName>
,
<taxonomicName id="F1814D01FFD1FF92F0B9FD99874EFDE8" authorityName="Laurent" authorityYear="1964" box="[1269,1405,513,539]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="machadoi">C. machadoi</taxonomicName>
</emphasis>
and
<emphasis id="04F5EA90FFD1FF92F1FEFD998478FDE8" box="[1458,1611,513,539]" italics="true" pageId="14" pageNumber="517">
<taxonomicName id="F1814D01FFD1FF92F1FEFD998475FDE8" baseAuthorityName="Bocage" baseAuthorityYear="1895" box="[1458,1606,513,539]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="angolensis">C. angolensis</taxonomicName>
.
</emphasis>
All measurements are presented in millimeters (mm). Data of
<taxonomicName id="F1814D01FFD1FF92F5C2FDBD8018FDCC" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[398,555,549,575]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="namakuiyus">
<emphasis id="04F5EA90FFD1FF92F5C2FDBD8018FDCC" box="[398,555,549,575]" italics="true" pageId="14" pageNumber="517">C. namakuiyus</emphasis>
</taxonomicName>
retrieved from
<bibRefCitation id="52104B73FFD1FF92F683FDBD819AFDCC" author="Stanley, E. L. &amp; Ceriaco, L. M. P. &amp; Bandeira, S. &amp; Valerio, H. &amp; Bates, M. F. &amp; Branch, W. R." box="[719,937,549,575]" pageId="14" pageNumber="517" pagination="201 - 226" refId="ref12195" refString="Stanley, E. L., Ceriaco, L. M. P., Bandeira, S., Valerio, H., Bates, M. F. &amp; Branch, W. R. (2016) A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola with the description of a new species from the Pro-Namib desert. Zootaxa, 4061 (3), 201 - 226. https: // doi. org / 10.11646 / zootaxa. 4061.3.1" type="journal article" year="2016">
Stanley
<emphasis id="04F5EA90FFD1FF92F769FDBD8161FDCC" box="[805,850,549,575]" italics="true" pageId="14" pageNumber="517">et al</emphasis>
. (2016)
</bibRefCitation>
; data for
<taxonomicName id="F1814D01FFD1FF92F045FDBD86A3FDCC" authorityName="Laurent" authorityYear="1964" box="[1033,1168,549,575]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD1FF92F045FDBD86A3FDCC" box="[1033,1168,549,575]" italics="true" pageId="14" pageNumber="517">C. machadoi</emphasis>
</taxonomicName>
retrieved from
<bibRefCitation id="52104B73FFD1FF92F178FDBD8439FDCC" author="Stanley, E. L. &amp; Ceriaco, L. M. P. &amp; Bandeira, S. &amp; Valerio, H. &amp; Bates, M. F. &amp; Branch, W. R." box="[1332,1546,549,575]" pageId="14" pageNumber="517" pagination="201 - 226" refId="ref12195" refString="Stanley, E. L., Ceriaco, L. M. P., Bandeira, S., Valerio, H., Bates, M. F. &amp; Branch, W. R. (2016) A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola with the description of a new species from the Pro-Namib desert. Zootaxa, 4061 (3), 201 - 226. https: // doi. org / 10.11646 / zootaxa. 4061.3.1" type="journal article" year="2016">
Stanley
<emphasis id="04F5EA90FFD1FF92F1C6FDBD878CFDCC" box="[1418,1471,549,575]" italics="true" pageId="14" pageNumber="517">et al.</emphasis>
(2016)
</bibRefCitation>
and two additional specimens from this study; data for
<taxonomicName id="F1814D01FFD1FF92F4BCFDD183BEFD90" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[240,397,585,611]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD1FF92F4BCFDD183BEFD90" box="[240,397,585,611]" italics="true" pageId="14" pageNumber="517">C. phonolithos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FC657EBFFD1FF92F5D8FDD183D8FD90" box="[404,491,585,611]" pageId="14" pageNumber="517" rank="species">
<emphasis id="04F5EA90FFD1FF92F5D8FDD183D0FD90" bold="true" box="[404,483,585,611]" pageId="14" pageNumber="517">sp. nov</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="F1814D01FFD1FF92F652FDD1809CFD90" baseAuthorityName="Bocage" baseAuthorityYear="1895" box="[542,687,585,611]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="04F5EA90FFD1FF92F652FDD1809CFD90" box="[542,687,585,611]" italics="true" pageId="14" pageNumber="517">C. angolensis</emphasis>
</taxonomicName>
collected in this study.
</paragraph>
</caption>
<paragraph id="363E3682FFD1FF92F644FDF687ACF97E" pageId="14" pageNumber="517">
<table id="4481C422FFD10063F4E9FDF685EDF97E" box="[165,2014,622,1677]" gridcols="12" gridrows="27" pageId="14" pageNumber="517">
<tr id="88B134C0FFD10063F4E9FDF685EDFD74" box="[165,2014,622,647]" gridrow="0" pageId="14" pageNumber="517" rowspan-0="1" rowspan-11="1" rowspan-5="1" rowspan-8="1">
<th id="CB605DBCFFD10063F644FDF68117FD74" box="[520,804,622,647]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="14" pageNumber="517">
<taxonomicName id="F1814D01FFD1FF92F644FDF680D1FD74" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[520,738,622,647]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="phonolithos">Cordylus phonolithos</taxonomicName>
</th>
<th id="CB605DBCFFD10063F708FDF6862CFD74" box="[836,1055,622,647]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="14" pageNumber="517">
<taxonomicName id="F1814D01FFD1FF92F708FDF6862CFD74" authorityName="Stanley, Ceriaco, Bandeira, Valerio, Bates &amp; Branch" authorityYear="2016" box="[836,1055,622,647]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="namakuiyus">Cordylus namakuiyus</taxonomicName>
</th>
<th id="CB605DBCFFD10063F092FDF687EAFD74" box="[1246,1497,622,647]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="14" pageNumber="517">
<taxonomicName id="F1814D01FFD1FF92F092FDF68797FD74" authorityName="Laurent" authorityYear="1964" box="[1246,1444,622,647]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="machadoi">
<emphasis id="04F5EA90FFD1FF92F092FDF68797FD74" box="[1246,1444,622,647]" italics="true" pageId="14" pageNumber="517">Cordylus machadoi</emphasis>
</taxonomicName>
</th>
<th id="CB605DBCFFD10063F2CDFDF6855FFD74" box="[1665,1900,622,647]" colspan="2" colspanRight="1" gridcol="9" gridrow="0" pageId="14" pageNumber="517">
<taxonomicName id="F1814D01FFD1FF92F2CDFDF6857CFD74" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[1665,1871,622,647]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="angolensis" status="sp. nov.">Cordylus angolensis</taxonomicName>
</th>
</tr>
<tr id="88B134C0FFD10063F4E9FD0A85EDFD58" box="[165,2014,658,683]" gridrow="1" pageId="14" pageNumber="517" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td id="CB605DBCFFD10063F644FD0A80B1FD58" box="[520,642,658,683]" gridcol="1" gridrow="1" pageId="14" pageNumber="517">
<emphasis id="04F5EA90FFD1FF92F642FD0A806CFD58" bold="true" box="[526,607,658,683]" pageId="14" pageNumber="517">sp. nov.</emphasis>
</td>
</tr>
<tr id="88B134C0FFD10063F4E9FD2B85EDFD3F" box="[165,2014,691,716]" gridrow="2" pageId="14" pageNumber="517" rowspan-0="1">
<td id="CB605DBCFFD10063F644FD2B80B1FD3F" box="[520,642,691,716]" gridcol="1" gridrow="2" pageId="14" pageNumber="517">Male n=1</td>
<td id="CB605DBCFFD10063F6EDFD2B8117FD3F" box="[673,804,691,716]" gridcol="2" gridrow="2" pageId="14" pageNumber="517">
<specimenCount id="2087FD0BFFD1FF92F6EDFD2B80C6FD3F" box="[673,757,691,716]" pageId="14" pageNumber="517" type="juvenile">Juvenile</specimenCount>
n=1
</td>
<td id="CB605DBCFFD10063F708FD2B818FFD3F" box="[836,956,691,716]" gridcol="3" gridrow="2" pageId="14" pageNumber="517">
<specimenCount id="2087FD0BFFD1FF92F708FD2B81BCFD3F" box="[836,911,691,716]" pageId="14" pageNumber="517" type="female">Female</specimenCount>
n=5
</td>
<td id="CB605DBCFFD10063F795FD2B862CFD3F" box="[985,1055,691,716]" gridcol="4" gridrow="2" pageId="14" pageNumber="517">
<specimenCount id="2087FD0BFFD1FF92F795FD2B863EFD3F" box="[985,1037,691,716]" pageId="14" pageNumber="517" type="male">Male</specimenCount>
</td>
<td id="CB605DBCFFD10063F07BFD2B868EFD3F" box="[1079,1213,691,716]" gridcol="5" gridrow="2" pageId="14" pageNumber="517">Juvenile n=4</td>
<td id="CB605DBCFFD10063F092FD2B877CFD3F" box="[1246,1359,691,716]" gridcol="6" gridrow="2" pageId="14" pageNumber="517">
<specimenCount id="2087FD0BFFD1FF92F092FD2B871AFD3F" box="[1246,1321,691,716]" pageId="14" pageNumber="517" type="female">Female</specimenCount>
</td>
<td id="CB605DBCFFD10063F121FD2B87EAFD3F" box="[1389,1497,691,716]" gridcol="7" gridrow="2" pageId="14" pageNumber="517">Male n=5</td>
<td id="CB605DBCFFD10063F1B5FD2B846CFD3F" box="[1529,1631,691,716]" gridcol="8" gridrow="2" pageId="14" pageNumber="517">
<specimenCount id="2087FD0BFFD1FF92F1B5FD2B847EFD3F" box="[1529,1613,691,716]" pageId="14" pageNumber="517" type="juvenile">Juvenile</specimenCount>
</td>
<td id="CB605DBCFFD10063F2CDFD2B853CFD3F" box="[1665,1807,691,716]" gridcol="9" gridrow="2" pageId="14" pageNumber="517">Female n=1</td>
<td id="CB605DBCFFD10063F360FD2B855FFD3F" box="[1836,1900,691,716]" gridcol="10" gridrow="2" pageId="14" pageNumber="517">
<specimenCount id="2087FD0BFFD1FF92F360FD2B8553FD3F" box="[1836,1888,691,716]" pageId="14" pageNumber="517" type="male">Male</specimenCount>
</td>
<td id="CB605DBCFFD10063F3C6FD2B85EDFD3F" box="[1930,2014,691,716]" gridcol="11" gridrow="2" pageId="14" pageNumber="517">Juvenile</td>
</tr>
<tr id="88B134C0FFD10063F4E9FD4F85EDFCE3" box="[165,2014,727,784]" gridrow="3" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FD4F83D8FCE3" box="[165,491,727,784]" gridcol="0" gridrow="3" pageId="14" pageNumber="517">Maximum SVL</th>
<td id="CB605DBCFFD10063F644FD4F80B1FCE3" box="[520,642,727,784]" gridcol="1" gridrow="3" pageId="14" pageNumber="517">
(
<typeStatus id="E93A8820FFD1FF92F643FD4F8043FD03" box="[527,624,727,752]" pageId="14" pageNumber="517" type="holotype">Holotype</typeStatus>
) 71.4
</td>
<td id="CB605DBCFFD10063F6EDFD4F8117FCE3" box="[673,804,727,784]" gridcol="2" gridrow="3" pageId="14" pageNumber="517">
(
<typeStatus id="E93A8820FFD1FF92F6E4FD4F8137FD03" box="[680,772,727,752]" pageId="14" pageNumber="517" type="paratype">Paratype</typeStatus>
) 49.9
</td>
<td id="CB605DBCFFD10063F708FD4F818FFCE3" box="[836,956,727,784]" gridcol="3" gridrow="3" pageId="14" pageNumber="517">101.9</td>
<td id="CB605DBCFFD10063F795FD4F862CFCE3" box="[985,1055,727,784]" gridcol="4" gridrow="3" pageId="14" pageNumber="517">n=1 89.7</td>
<td id="CB605DBCFFD10063F07BFD4F868EFCE3" box="[1079,1213,727,784]" gridcol="5" gridrow="3" pageId="14" pageNumber="517">57.1</td>
<td id="CB605DBCFFD10063F092FD4F877CFCE3" box="[1246,1359,727,784]" gridcol="6" gridrow="3" pageId="14" pageNumber="517">n=6 97.8</td>
<td id="CB605DBCFFD10063F121FD4F87EAFCE3" box="[1389,1497,727,784]" gridcol="7" gridrow="3" pageId="14" pageNumber="517">94.2</td>
<td id="CB605DBCFFD10063F1B5FD4F846CFCE3" box="[1529,1631,727,784]" gridcol="8" gridrow="3" pageId="14" pageNumber="517">n=5 66.7</td>
<td id="CB605DBCFFD10063F2CDFD4F853CFCE3" box="[1665,1807,727,784]" gridcol="9" gridrow="3" pageId="14" pageNumber="517">79.5</td>
<td id="CB605DBCFFD10063F360FD4F855FFCE3" box="[1836,1900,727,784]" gridcol="10" gridrow="3" pageId="14" pageNumber="517">n=1 78.2</td>
<td id="CB605DBCFFD10063F3C6FD4F85EDFCE3" box="[1930,2014,727,784]" gridcol="11" gridrow="3" pageId="14" pageNumber="517">n=1 61.9</td>
</tr>
<tr id="88B134C0FFD10063F4E9FC8585EDFCC5" box="[165,2014,797,822]" gridrow="4" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FC8583D8FCC5" box="[165,491,797,822]" gridcol="0" gridrow="4" pageId="14" pageNumber="517">Chin Shields</th>
<td id="CB605DBCFFD10063F644FC8580B1FCC5" box="[520,642,797,822]" gridcol="1" gridrow="4" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F6EDFC858117FCC5" box="[673,804,797,822]" gridcol="2" gridrow="4" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F708FC85818FFCC5" box="[836,956,797,822]" gridcol="3" gridrow="4" pageId="14" pageNumber="517">56</td>
<td id="CB605DBCFFD10063F795FC85862CFCC5" box="[985,1055,797,822]" gridcol="4" gridrow="4" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F07BFC85868EFCC5" box="[1079,1213,797,822]" gridcol="5" gridrow="4" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F092FC85877CFCC5" box="[1246,1359,797,822]" gridcol="6" gridrow="4" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F121FC8587EAFCC5" box="[1389,1497,797,822]" gridcol="7" gridrow="4" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F1B5FC85846CFCC5" box="[1529,1631,797,822]" gridcol="8" gridrow="4" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F2CDFC85853CFCC5" box="[1665,1807,797,822]" gridcol="9" gridrow="4" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F360FC85855FFCC5" box="[1836,1900,797,822]" gridcol="10" gridrow="4" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F3C6FC8585EDFCC5" box="[1930,2014,797,822]" gridcol="11" gridrow="4" pageId="14" pageNumber="517">5</td>
</tr>
<tr id="88B134C0FFD10063F4E9FCDB85EDFCAF" box="[165,2014,835,860]" gridrow="5" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FCDB83D8FCAF" box="[165,491,835,860]" gridcol="0" gridrow="5" pageId="14" pageNumber="517">Femoral Pores</th>
<td id="CB605DBCFFD10063F644FCDB80B1FCAF" box="[520,642,835,860]" gridcol="1" gridrow="5" pageId="14" pageNumber="517">7</td>
<td id="CB605DBCFFD10063F6EDFCDB8117FCAF" box="[673,804,835,860]" gridcol="2" gridrow="5" pageId="14" pageNumber="517">0</td>
<td id="CB605DBCFFD10063F708FCDB818FFCAF" box="[836,956,835,860]" gridcol="3" gridrow="5" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F795FCDB862CFCAF" box="[985,1055,835,860]" gridcol="4" gridrow="5" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F07BFCDB868EFCAF" box="[1079,1213,835,860]" gridcol="5" gridrow="5" pageId="14" pageNumber="517">06</td>
<td id="CB605DBCFFD10063F092FCDB877CFCAF" box="[1246,1359,835,860]" gridcol="6" gridrow="5" pageId="14" pageNumber="517">67</td>
<td id="CB605DBCFFD10063F121FCDB87EAFCAF" box="[1389,1497,835,860]" gridcol="7" gridrow="5" pageId="14" pageNumber="517">57</td>
<td id="CB605DBCFFD10063F1B5FCDB846CFCAF" box="[1529,1631,835,860]" gridcol="8" gridrow="5" pageId="14" pageNumber="517">56</td>
<td id="CB605DBCFFD10063F2CDFCDB853CFCAF" box="[1665,1807,835,860]" gridcol="9" gridrow="5" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F360FCDB855FFCAF" box="[1836,1900,835,860]" gridcol="10" gridrow="5" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F3C6FCDB85EDFCAF" box="[1930,2014,835,860]" gridcol="11" gridrow="5" pageId="14" pageNumber="517">5</td>
</tr>
<tr id="88B134C0FFD10063F4E9FCF185EDFC71" box="[165,2014,873,898]" gridrow="6" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FCF183D8FC71" box="[165,491,873,898]" gridcol="0" gridrow="6" pageId="14" pageNumber="517">Generation Glands</th>
<td id="CB605DBCFFD10063F644FCF180B1FC71" box="[520,642,873,898]" gridcol="1" gridrow="6" pageId="14" pageNumber="517">1713</td>
<td id="CB605DBCFFD10063F6EDFCF18117FC71" box="[673,804,873,898]" gridcol="2" gridrow="6" pageId="14" pageNumber="517">0</td>
<td id="CB605DBCFFD10063F708FCF1818FFC71" box="[836,956,873,898]" gridcol="3" gridrow="6" pageId="14" pageNumber="517">0</td>
<td id="CB605DBCFFD10063F795FCF1862CFC71" box="[985,1055,873,898]" gridcol="4" gridrow="6" pageId="14" pageNumber="517">12</td>
<td id="CB605DBCFFD10063F07BFCF1868EFC71" box="[1079,1213,873,898]" gridcol="5" gridrow="6" pageId="14" pageNumber="517">0</td>
<td id="CB605DBCFFD10063F092FCF1877CFC71" box="[1246,1359,873,898]" gridcol="6" gridrow="6" pageId="14" pageNumber="517">0</td>
<td id="CB605DBCFFD10063F121FCF187EAFC71" box="[1389,1497,873,898]" gridcol="7" gridrow="6" pageId="14" pageNumber="517">1222</td>
<td id="CB605DBCFFD10063F1B5FCF1846CFC71" box="[1529,1631,873,898]" gridcol="8" gridrow="6" pageId="14" pageNumber="517">016</td>
<td id="CB605DBCFFD10063F2CDFCF1853CFC71" box="[1665,1807,873,898]" gridcol="9" gridrow="6" pageId="14" pageNumber="517">0</td>
<td id="CB605DBCFFD10063F360FCF1855FFC71" box="[1836,1900,873,898]" gridcol="10" gridrow="6" pageId="14" pageNumber="517">36</td>
<td id="CB605DBCFFD10063F3C6FCF185EDFC71" box="[1930,2014,873,898]" gridcol="11" gridrow="6" pageId="14" pageNumber="517">0</td>
</tr>
<tr id="88B134C0FFD10063F4E9FC1785EDFC5B" box="[165,2014,911,936]" gridrow="7" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FC1783D8FC5B" box="[165,491,911,936]" gridcol="0" gridrow="7" pageId="14" pageNumber="517">Supralabials</th>
<td id="CB605DBCFFD10063F644FC1780B1FC5B" box="[520,642,911,936]" gridcol="1" gridrow="7" pageId="14" pageNumber="517">6</td>
<td id="CB605DBCFFD10063F6EDFC178117FC5B" box="[673,804,911,936]" gridcol="2" gridrow="7" pageId="14" pageNumber="517">6</td>
<td id="CB605DBCFFD10063F708FC17818FFC5B" box="[836,956,911,936]" gridcol="3" gridrow="7" pageId="14" pageNumber="517">45</td>
<td id="CB605DBCFFD10063F795FC17862CFC5B" box="[985,1055,911,936]" gridcol="4" gridrow="7" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F07BFC17868EFC5B" box="[1079,1213,911,936]" gridcol="5" gridrow="7" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F092FC17877CFC5B" box="[1246,1359,911,936]" gridcol="6" gridrow="7" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F121FC1787EAFC5B" box="[1389,1497,911,936]" gridcol="7" gridrow="7" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F1B5FC17846CFC5B" box="[1529,1631,911,936]" gridcol="8" gridrow="7" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F2CDFC17853CFC5B" box="[1665,1807,911,936]" gridcol="9" gridrow="7" pageId="14" pageNumber="517">45</td>
<td id="CB605DBCFFD10063F360FC17855FFC5B" box="[1836,1900,911,936]" gridcol="10" gridrow="7" pageId="14" pageNumber="517">45</td>
<td id="CB605DBCFFD10063F3C6FC1785EDFC5B" box="[1930,2014,911,936]" gridcol="11" gridrow="7" pageId="14" pageNumber="517">4</td>
</tr>
<tr id="88B134C0FFD10063F4E9FC2C85EDFC3E" box="[165,2014,948,973]" gridrow="8" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FC2C83D8FC3E" box="[165,491,948,973]" gridcol="0" gridrow="8" pageId="14" pageNumber="517">Infralabials</th>
<td id="CB605DBCFFD10063F644FC2C80B1FC3E" box="[520,642,948,973]" gridcol="1" gridrow="8" pageId="14" pageNumber="517">6</td>
<td id="CB605DBCFFD10063F6EDFC2C8117FC3E" box="[673,804,948,973]" gridcol="2" gridrow="8" pageId="14" pageNumber="517">6</td>
<td id="CB605DBCFFD10063F708FC2C818FFC3E" box="[836,956,948,973]" gridcol="3" gridrow="8" pageId="14" pageNumber="517">56</td>
<td id="CB605DBCFFD10063F795FC2C862CFC3E" box="[985,1055,948,973]" gridcol="4" gridrow="8" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F07BFC2C868EFC3E" box="[1079,1213,948,973]" gridcol="5" gridrow="8" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F092FC2C877CFC3E" box="[1246,1359,948,973]" gridcol="6" gridrow="8" pageId="14" pageNumber="517">6</td>
<td id="CB605DBCFFD10063F121FC2C87EAFC3E" box="[1389,1497,948,973]" gridcol="7" gridrow="8" pageId="14" pageNumber="517">6</td>
<td id="CB605DBCFFD10063F1B5FC2C846CFC3E" box="[1529,1631,948,973]" gridcol="8" gridrow="8" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F2CDFC2C853CFC3E" box="[1665,1807,948,973]" gridcol="9" gridrow="8" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F360FC2C855FFC3E" box="[1836,1900,948,973]" gridcol="10" gridrow="8" pageId="14" pageNumber="517">5</td>
<td id="CB605DBCFFD10063F3C6FC2C85EDFC3E" box="[1930,2014,948,973]" gridcol="11" gridrow="8" pageId="14" pageNumber="517">5</td>
</tr>
<tr id="88B134C0FFD10063F4E9FC4285EDFC00" box="[165,2014,986,1011]" gridrow="9" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FC4283D8FC00" box="[165,491,986,1011]" gridcol="0" gridrow="9" pageId="14" pageNumber="517">Supraoculars</th>
<td id="CB605DBCFFD10063F644FC4280B1FC00" box="[520,642,986,1011]" gridcol="1" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F6EDFC428117FC00" box="[673,804,986,1011]" gridcol="2" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F708FC42818FFC00" box="[836,956,986,1011]" gridcol="3" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F795FC42862CFC00" box="[985,1055,986,1011]" gridcol="4" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F07BFC42868EFC00" box="[1079,1213,986,1011]" gridcol="5" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F092FC42877CFC00" box="[1246,1359,986,1011]" gridcol="6" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F121FC4287EAFC00" box="[1389,1497,986,1011]" gridcol="7" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F1B5FC42846CFC00" box="[1529,1631,986,1011]" gridcol="8" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F2CDFC42853CFC00" box="[1665,1807,986,1011]" gridcol="9" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F360FC42855FFC00" box="[1836,1900,986,1011]" gridcol="10" gridrow="9" pageId="14" pageNumber="517">4</td>
<td id="CB605DBCFFD10063F3C6FC4285EDFC00" box="[1930,2014,986,1011]" gridcol="11" gridrow="9" pageId="14" pageNumber="517">4</td>
</tr>
<tr id="88B134C0FFD10063F4E9FB9885EDFBEA" box="[165,2014,1024,1049]" gridrow="10" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FB9883D8FBEA" box="[165,491,1024,1049]" gridcol="0" gridrow="10" pageId="14" pageNumber="517">Supraciliars</th>
<td id="CB605DBCFFD10063F644FB9880B1FBEA" box="[520,642,1024,1049]" gridcol="1" gridrow="10" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F6EDFB988117FBEA" box="[673,804,1024,1049]" gridcol="2" gridrow="10" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F708FB98818FFBEA" box="[836,956,1024,1049]" gridcol="3" gridrow="10" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F795FB98862CFBEA" box="[985,1055,1024,1049]" gridcol="4" gridrow="10" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F07BFB98868EFBEA" box="[1079,1213,1024,1049]" gridcol="5" gridrow="10" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F092FB98877CFBEA" box="[1246,1359,1024,1049]" gridcol="6" gridrow="10" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F121FB9887EAFBEA" box="[1389,1497,1024,1049]" gridcol="7" gridrow="10" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F1B5FB98846CFBEA" box="[1529,1631,1024,1049]" gridcol="8" gridrow="10" pageId="14" pageNumber="517">34</td>
<td id="CB605DBCFFD10063F2CDFB98853CFBEA" box="[1665,1807,1024,1049]" gridcol="9" gridrow="10" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F360FB98855FFBEA" box="[1836,1900,1024,1049]" gridcol="10" gridrow="10" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F3C6FB9885EDFBEA" box="[1930,2014,1024,1049]" gridcol="11" gridrow="10" pageId="14" pageNumber="517">3</td>
</tr>
<tr id="88B134C0FFD10063F4E9FBBE85EDFBCC" box="[165,2014,1062,1087]" gridrow="11" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FBBE83D8FBCC" box="[165,491,1062,1087]" gridcol="0" gridrow="11" pageId="14" pageNumber="517">Loreal</th>
<td id="CB605DBCFFD10063F644FBBE80B1FBCC" box="[520,642,1062,1087]" gridcol="1" gridrow="11" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F6EDFBBE8117FBCC" box="[673,804,1062,1087]" gridcol="2" gridrow="11" pageId="14" pageNumber="517">FUSED</td>
<td id="CB605DBCFFD10063F708FBBE818FFBCC" box="[836,956,1062,1087]" gridcol="3" gridrow="11" pageId="14" pageNumber="517">01</td>
<td id="CB605DBCFFD10063F795FBBE862CFBCC" box="[985,1055,1062,1087]" gridcol="4" gridrow="11" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F07BFBBE868EFBCC" box="[1079,1213,1062,1087]" gridcol="5" gridrow="11" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F092FBBE877CFBCC" box="[1246,1359,1062,1087]" gridcol="6" gridrow="11" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F121FBBE87EAFBCC" box="[1389,1497,1062,1087]" gridcol="7" gridrow="11" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F1B5FBBE846CFBCC" box="[1529,1631,1062,1087]" gridcol="8" gridrow="11" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F2CDFBBE853CFBCC" box="[1665,1807,1062,1087]" gridcol="9" gridrow="11" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F360FBBE855FFBCC" box="[1836,1900,1062,1087]" gridcol="10" gridrow="11" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F3C6FBBE85EDFBCC" box="[1930,2014,1062,1087]" gridcol="11" gridrow="11" pageId="14" pageNumber="517">1</td>
</tr>
<tr id="88B134C0FFD10063F4E9FBD385EDFB97" box="[165,2014,1099,1124]" gridrow="12" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FBD383D8FB97" box="[165,491,1099,1124]" gridcol="0" gridrow="12" pageId="14" pageNumber="517">Suboculars</th>
<td id="CB605DBCFFD10063F644FBD380B1FB97" box="[520,642,1099,1124]" gridcol="1" gridrow="12" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F6EDFBD38117FB97" box="[673,804,1099,1124]" gridcol="2" gridrow="12" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F708FBD3818FFB97" box="[836,956,1099,1124]" gridcol="3" gridrow="12" pageId="14" pageNumber="517">23</td>
<td id="CB605DBCFFD10063F795FBD3862CFB97" box="[985,1055,1099,1124]" gridcol="4" gridrow="12" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F07BFBD3868EFB97" box="[1079,1213,1099,1124]" gridcol="5" gridrow="12" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F092FBD3877CFB97" box="[1246,1359,1099,1124]" gridcol="6" gridrow="12" pageId="14" pageNumber="517">34</td>
<td id="CB605DBCFFD10063F121FBD387EAFB97" box="[1389,1497,1099,1124]" gridcol="7" gridrow="12" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F1B5FBD3846CFB97" box="[1529,1631,1099,1124]" gridcol="8" gridrow="12" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F2CDFBD3853CFB97" box="[1665,1807,1099,1124]" gridcol="9" gridrow="12" pageId="14" pageNumber="517">23</td>
<td id="CB605DBCFFD10063F360FBD3855FFB97" box="[1836,1900,1099,1124]" gridcol="10" gridrow="12" pageId="14" pageNumber="517">3</td>
<td id="CB605DBCFFD10063F3C6FBD385EDFB97" box="[1930,2014,1099,1124]" gridcol="11" gridrow="12" pageId="14" pageNumber="517">23</td>
</tr>
<tr id="88B134C0FFD10063F4E9FBE985EDFB79" box="[165,2014,1137,1162]" gridrow="13" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FBE983D8FB79" box="[165,491,1137,1162]" gridcol="0" gridrow="13" pageId="14" pageNumber="517">Preoculars</th>
<td id="CB605DBCFFD10063F644FBE980B1FB79" box="[520,642,1137,1162]" gridcol="1" gridrow="13" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F6EDFBE98117FB79" box="[673,804,1137,1162]" gridcol="2" gridrow="13" pageId="14" pageNumber="517">FUSED</td>
<td id="CB605DBCFFD10063F708FBE9818FFB79" box="[836,956,1137,1162]" gridcol="3" gridrow="13" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F795FBE9862CFB79" box="[985,1055,1137,1162]" gridcol="4" gridrow="13" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F07BFBE9868EFB79" box="[1079,1213,1137,1162]" gridcol="5" gridrow="13" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F092FBE9877CFB79" box="[1246,1359,1137,1162]" gridcol="6" gridrow="13" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F121FBE987EAFB79" box="[1389,1497,1137,1162]" gridcol="7" gridrow="13" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F1B5FBE9846CFB79" box="[1529,1631,1137,1162]" gridcol="8" gridrow="13" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F2CDFBE9853CFB79" box="[1665,1807,1137,1162]" gridcol="9" gridrow="13" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F360FBE9855FFB79" box="[1836,1900,1137,1162]" gridcol="10" gridrow="13" pageId="14" pageNumber="517">1</td>
<td id="CB605DBCFFD10063F3C6FBE985EDFB79" box="[1930,2014,1137,1162]" gridcol="11" gridrow="13" pageId="14" pageNumber="517">1</td>
</tr>
<tr id="88B134C0FFD10063F4E9FB0F85EDFB43" box="[165,2014,1175,1200]" gridrow="14" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FB0F83D8FB43" box="[165,491,1175,1200]" gridcol="0" gridrow="14" pageId="14" pageNumber="517">Transverse Gular Scales</th>
<td id="CB605DBCFFD10063F644FB0F80B1FB43" box="[520,642,1175,1200]" gridcol="1" gridrow="14" pageId="14" pageNumber="517">17</td>
<td id="CB605DBCFFD10063F6EDFB0F8117FB43" box="[673,804,1175,1200]" gridcol="2" gridrow="14" pageId="14" pageNumber="517">17</td>
<td id="CB605DBCFFD10063F708FB0F818FFB43" box="[836,956,1175,1200]" gridcol="3" gridrow="14" pageId="14" pageNumber="517">1417</td>
<td id="CB605DBCFFD10063F795FB0F862CFB43" box="[985,1055,1175,1200]" gridcol="4" gridrow="14" pageId="14" pageNumber="517">15</td>
<td id="CB605DBCFFD10063F07BFB0F868EFB43" box="[1079,1213,1175,1200]" gridcol="5" gridrow="14" pageId="14" pageNumber="517">1516</td>
<td id="CB605DBCFFD10063F092FB0F877CFB43" box="[1246,1359,1175,1200]" gridcol="6" gridrow="14" pageId="14" pageNumber="517">1721</td>
<td id="CB605DBCFFD10063F121FB0F87EAFB43" box="[1389,1497,1175,1200]" gridcol="7" gridrow="14" pageId="14" pageNumber="517">1820</td>
<td id="CB605DBCFFD10063F1B5FB0F846CFB43" box="[1529,1631,1175,1200]" gridcol="8" gridrow="14" pageId="14" pageNumber="517">1824</td>
<td id="CB605DBCFFD10063F2CDFB0F853CFB43" box="[1665,1807,1175,1200]" gridcol="9" gridrow="14" pageId="14" pageNumber="517">18</td>
<td id="CB605DBCFFD10063F360FB0F855FFB43" box="[1836,1900,1175,1200]" gridcol="10" gridrow="14" pageId="14" pageNumber="517">18</td>
<td id="CB605DBCFFD10063F3C6FB0F85EDFB43" box="[1930,2014,1175,1200]" gridcol="11" gridrow="14" pageId="14" pageNumber="517">18</td>
</tr>
<tr id="88B134C0FFD10063F4E9FB2585EDFB25" box="[165,2014,1213,1238]" gridrow="15" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FB2583D8FB25" box="[165,491,1213,1238]" gridcol="0" gridrow="15" pageId="14" pageNumber="517">Dorsal Transverse Rows</th>
<td id="CB605DBCFFD10063F644FB2580B1FB25" box="[520,642,1213,1238]" gridcol="1" gridrow="15" pageId="14" pageNumber="517">25</td>
<td id="CB605DBCFFD10063F6EDFB258117FB25" box="[673,804,1213,1238]" gridcol="2" gridrow="15" pageId="14" pageNumber="517">25</td>
<td id="CB605DBCFFD10063F708FB25818FFB25" box="[836,956,1213,1238]" gridcol="3" gridrow="15" pageId="14" pageNumber="517">2426</td>
<td id="CB605DBCFFD10063F795FB25862CFB25" box="[985,1055,1213,1238]" gridcol="4" gridrow="15" pageId="14" pageNumber="517">24</td>
<td id="CB605DBCFFD10063F07BFB25868EFB25" box="[1079,1213,1213,1238]" gridcol="5" gridrow="15" pageId="14" pageNumber="517">2325</td>
<td id="CB605DBCFFD10063F092FB25877CFB25" box="[1246,1359,1213,1238]" gridcol="6" gridrow="15" pageId="14" pageNumber="517">2428</td>
<td id="CB605DBCFFD10063F121FB2587EAFB25" box="[1389,1497,1213,1238]" gridcol="7" gridrow="15" pageId="14" pageNumber="517">2326</td>
<td id="CB605DBCFFD10063F1B5FB25846CFB25" box="[1529,1631,1213,1238]" gridcol="8" gridrow="15" pageId="14" pageNumber="517">2528</td>
<td id="CB605DBCFFD10063F2CDFB25853CFB25" box="[1665,1807,1213,1238]" gridcol="9" gridrow="15" pageId="14" pageNumber="517">22</td>
<td id="CB605DBCFFD10063F360FB25855FFB25" box="[1836,1900,1213,1238]" gridcol="10" gridrow="15" pageId="14" pageNumber="517">21</td>
<td id="CB605DBCFFD10063F3C6FB2585EDFB25" box="[1930,2014,1213,1238]" gridcol="11" gridrow="15" pageId="14" pageNumber="517">21</td>
</tr>
<tr id="88B134C0FFD10063F4E9FB7B85EDFB0F" box="[165,2014,1251,1276]" gridrow="16" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FB7B83D8FB0F" box="[165,491,1251,1276]" gridcol="0" gridrow="16" pageId="14" pageNumber="517">Dosolateral Longitudinal Rows</th>
<td id="CB605DBCFFD10063F644FB7B80B1FB0F" box="[520,642,1251,1276]" gridcol="1" gridrow="16" pageId="14" pageNumber="517">23</td>
<td id="CB605DBCFFD10063F6EDFB7B8117FB0F" box="[673,804,1251,1276]" gridcol="2" gridrow="16" pageId="14" pageNumber="517">23</td>
<td id="CB605DBCFFD10063F708FB7B818FFB0F" box="[836,956,1251,1276]" gridcol="3" gridrow="16" pageId="14" pageNumber="517">24</td>
<td id="CB605DBCFFD10063F795FB7B862CFB0F" box="[985,1055,1251,1276]" gridcol="4" gridrow="16" pageId="14" pageNumber="517">23</td>
<td id="CB605DBCFFD10063F07BFB7B868EFB0F" box="[1079,1213,1251,1276]" gridcol="5" gridrow="16" pageId="14" pageNumber="517">2324</td>
<td id="CB605DBCFFD10063F092FB7B877CFB0F" box="[1246,1359,1251,1276]" gridcol="6" gridrow="16" pageId="14" pageNumber="517">2227</td>
<td id="CB605DBCFFD10063F121FB7B87EAFB0F" box="[1389,1497,1251,1276]" gridcol="7" gridrow="16" pageId="14" pageNumber="517">23</td>
<td id="CB605DBCFFD10063F1B5FB7B846CFB0F" box="[1529,1631,1251,1276]" gridcol="8" gridrow="16" pageId="14" pageNumber="517">2324</td>
<td id="CB605DBCFFD10063F2CDFB7B853CFB0F" box="[1665,1807,1251,1276]" gridcol="9" gridrow="16" pageId="14" pageNumber="517">1819</td>
<td id="CB605DBCFFD10063F360FB7B855FFB0F" box="[1836,1900,1251,1276]" gridcol="10" gridrow="16" pageId="14" pageNumber="517">1819</td>
<td id="CB605DBCFFD10063F3C6FB7B85EDFB0F" box="[1930,2014,1251,1276]" gridcol="11" gridrow="16" pageId="14" pageNumber="517">1819</td>
</tr>
<tr id="88B134C0FFD10063F4E9FA9085EDFAD2" box="[165,2014,1288,1313]" gridrow="17" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FA9083D8FAD2" box="[165,491,1288,1313]" gridcol="0" gridrow="17" pageId="14" pageNumber="517">Ventral Transverse Rows</th>
<td id="CB605DBCFFD10063F644FA9080B1FAD2" box="[520,642,1288,1313]" gridcol="1" gridrow="17" pageId="14" pageNumber="517">23</td>
<td id="CB605DBCFFD10063F6EDFA908117FAD2" box="[673,804,1288,1313]" gridcol="2" gridrow="17" pageId="14" pageNumber="517">23</td>
<td id="CB605DBCFFD10063F708FA90818FFAD2" box="[836,956,1288,1313]" gridcol="3" gridrow="17" pageId="14" pageNumber="517">2124</td>
<td id="CB605DBCFFD10063F795FA90862CFAD2" box="[985,1055,1288,1313]" gridcol="4" gridrow="17" pageId="14" pageNumber="517">22</td>
<td id="CB605DBCFFD10063F07BFA90868EFAD2" box="[1079,1213,1288,1313]" gridcol="5" gridrow="17" pageId="14" pageNumber="517">2122</td>
<td id="CB605DBCFFD10063F092FA90877CFAD2" box="[1246,1359,1288,1313]" gridcol="6" gridrow="17" pageId="14" pageNumber="517">2125</td>
<td id="CB605DBCFFD10063F121FA9087EAFAD2" box="[1389,1497,1288,1313]" gridcol="7" gridrow="17" pageId="14" pageNumber="517">2126</td>
<td id="CB605DBCFFD10063F1B5FA90846CFAD2" box="[1529,1631,1288,1313]" gridcol="8" gridrow="17" pageId="14" pageNumber="517">2022</td>
<td id="CB605DBCFFD10063F2CDFA90853CFAD2" box="[1665,1807,1288,1313]" gridcol="9" gridrow="17" pageId="14" pageNumber="517">2526</td>
<td id="CB605DBCFFD10063F360FA90855FFAD2" box="[1836,1900,1288,1313]" gridcol="10" gridrow="17" pageId="14" pageNumber="517">25</td>
<td id="CB605DBCFFD10063F3C6FA9085EDFAD2" box="[1930,2014,1288,1313]" gridcol="11" gridrow="17" pageId="14" pageNumber="517">24</td>
</tr>
<tr id="88B134C0FFD10063F4E9FAB685EDFAB4" box="[165,2014,1326,1351]" gridrow="18" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FAB683D8FAB4" box="[165,491,1326,1351]" gridcol="0" gridrow="18" pageId="14" pageNumber="517">Ventral Longitudinal Rows</th>
<td id="CB605DBCFFD10063F644FAB680B1FAB4" box="[520,642,1326,1351]" gridcol="1" gridrow="18" pageId="14" pageNumber="517">17</td>
<td id="CB605DBCFFD10063F6EDFAB68117FAB4" box="[673,804,1326,1351]" gridcol="2" gridrow="18" pageId="14" pageNumber="517">17</td>
<td id="CB605DBCFFD10063F708FAB6818FFAB4" box="[836,956,1326,1351]" gridcol="3" gridrow="18" pageId="14" pageNumber="517">1516</td>
<td id="CB605DBCFFD10063F795FAB6862CFAB4" box="[985,1055,1326,1351]" gridcol="4" gridrow="18" pageId="14" pageNumber="517">17</td>
<td id="CB605DBCFFD10063F07BFAB6868EFAB4" box="[1079,1213,1326,1351]" gridcol="5" gridrow="18" pageId="14" pageNumber="517">1416</td>
<td id="CB605DBCFFD10063F092FAB6877CFAB4" box="[1246,1359,1326,1351]" gridcol="6" gridrow="18" pageId="14" pageNumber="517">1418</td>
<td id="CB605DBCFFD10063F121FAB687EAFAB4" box="[1389,1497,1326,1351]" gridcol="7" gridrow="18" pageId="14" pageNumber="517">1316</td>
<td id="CB605DBCFFD10063F1B5FAB6846CFAB4" box="[1529,1631,1326,1351]" gridcol="8" gridrow="18" pageId="14" pageNumber="517">1314</td>
<td id="CB605DBCFFD10063F2CDFAB6853CFAB4" box="[1665,1807,1326,1351]" gridcol="9" gridrow="18" pageId="14" pageNumber="517">12</td>
<td id="CB605DBCFFD10063F360FAB6855FFAB4" box="[1836,1900,1326,1351]" gridcol="10" gridrow="18" pageId="14" pageNumber="517">12</td>
<td id="CB605DBCFFD10063F3C6FAB685EDFAB4" box="[1930,2014,1326,1351]" gridcol="11" gridrow="18" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F3C6FAB68591FAB4" box="[1930,1954,1326,1351]" pageId="14" pageNumber="517" value="2011-12-12">11</date>
</td>
</tr>
<tr id="88B134C0FFD10063F4E9FACB85EDFA9E" box="[165,2014,1363,1389]" gridrow="19" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FACB83D8FA9E" box="[165,491,1363,1389]" gridcol="0" gridrow="19" pageId="14" pageNumber="517">
Caudal Scales 11
<superScript id="C1F49BCAFFD1FF92F51FFACB836DFA92" attach="left" box="[339,350,1363,1377]" fontSize="6" pageId="14" pageNumber="517">th</superScript>
Whorl
</th>
<td id="CB605DBCFFD10063F644FACB80B1FA9E" box="[520,642,1363,1389]" gridcol="1" gridrow="19" pageId="14" pageNumber="517">10</td>
<td id="CB605DBCFFD10063F6EDFACB8117FA9E" box="[673,804,1363,1389]" gridcol="2" gridrow="19" pageId="14" pageNumber="517">10</td>
<td id="CB605DBCFFD10063F708FACB818FFA9E" box="[836,956,1363,1389]" gridcol="3" gridrow="19" pageId="14" pageNumber="517">1012</td>
<td id="CB605DBCFFD10063F795FACB862CFA9E" box="[985,1055,1363,1389]" gridcol="4" gridrow="19" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F795FACC81C1FA9E" box="[985,1010,1364,1389]" pageId="14" pageNumber="517" value="2010-10-10" valueMax="2010-12-10" valueMin="2010-10-10">10</date>
</td>
<td id="CB605DBCFFD10063F07BFACB868EFA9E" box="[1079,1213,1363,1389]" gridcol="5" gridrow="19" pageId="14" pageNumber="517">10</td>
<td id="CB605DBCFFD10063F092FACB877CFA9E" box="[1246,1359,1363,1389]" gridcol="6" gridrow="19" pageId="14" pageNumber="517">1011</td>
<td id="CB605DBCFFD10063F121FACB87EAFA9E" box="[1389,1497,1363,1389]" gridcol="7" gridrow="19" pageId="14" pageNumber="517">10</td>
<td id="CB605DBCFFD10063F1B5FACB846CFA9E" box="[1529,1631,1363,1389]" gridcol="8" gridrow="19" pageId="14" pageNumber="517">10</td>
<td id="CB605DBCFFD10063F2CDFACB853CFA9E" box="[1665,1807,1363,1389]" gridcol="9" gridrow="19" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F2CDFACC84A9FA9E" box="[1665,1690,1364,1389]" pageId="14" pageNumber="517" value="2010-10-10">10</date>
</td>
<td id="CB605DBCFFD10063F360FACB855FFA9E" box="[1836,1900,1363,1389]" gridcol="10" gridrow="19" pageId="14" pageNumber="517">11</td>
<td id="CB605DBCFFD10063F3C6FACB85EDFA9E" box="[1930,2014,1363,1389]" gridcol="11" gridrow="19" pageId="14" pageNumber="517">11</td>
</tr>
<tr id="88B134C0FFD10063F4E9FAE085EDFA60" box="[165,2014,1400,1427]" gridrow="20" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FAE083D8FA60" box="[165,491,1400,1427]" gridcol="0" gridrow="20" pageId="14" pageNumber="517">
Caudal Scales 15
<superScript id="C1F49BCAFFD1FF92F518FAE0836CFA75" attach="left" box="[340,351,1400,1414]" fontSize="6" pageId="14" pageNumber="517">th</superScript>
Whorl
</th>
<td id="CB605DBCFFD10063F644FAE080B1FA60" box="[520,642,1400,1427]" gridcol="1" gridrow="20" pageId="14" pageNumber="517">8</td>
<td id="CB605DBCFFD10063F6EDFAE08117FA60" box="[673,804,1400,1427]" gridcol="2" gridrow="20" pageId="14" pageNumber="517">8</td>
<td id="CB605DBCFFD10063F708FAE0818FFA60" box="[836,956,1400,1427]" gridcol="3" gridrow="20" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F708FAE2816EFA60" box="[836,861,1402,1427]" pageId="14" pageNumber="517" value="2010-08-08">10</date>
</td>
<td id="CB605DBCFFD10063F795FAE0862CFA60" box="[985,1055,1400,1427]" gridcol="4" gridrow="20" pageId="14" pageNumber="517">10</td>
<td id="CB605DBCFFD10063F07BFAE0868EFA60" box="[1079,1213,1400,1427]" gridcol="5" gridrow="20" pageId="14" pageNumber="517">10</td>
<td id="CB605DBCFFD10063F092FAE0877CFA60" box="[1246,1359,1400,1427]" gridcol="6" gridrow="20" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F092FAE28723FA60" box="[1246,1296,1402,1427]" pageId="14" pageNumber="517" value="2010-08-10" valueMax="2010-10-10" valueMin="2010-08-10">810</date>
</td>
<td id="CB605DBCFFD10063F121FAE087EAFA60" box="[1389,1497,1400,1427]" gridcol="7" gridrow="20" pageId="14" pageNumber="517">1011</td>
<td id="CB605DBCFFD10063F1B5FAE0846CFA60" box="[1529,1631,1400,1427]" gridcol="8" gridrow="20" pageId="14" pageNumber="517">9</td>
<td id="CB605DBCFFD10063F2CDFAE0853CFA60" box="[1665,1807,1400,1427]" gridcol="9" gridrow="20" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F2CDFAE284BDFA60" box="[1665,1678,1402,1427]" pageId="14" pageNumber="517" value="2011-08-09">8</date>
</td>
<td id="CB605DBCFFD10063F360FAE0855FFA60" box="[1836,1900,1400,1427]" gridcol="10" gridrow="20" pageId="14" pageNumber="517">0</td>
<td id="CB605DBCFFD10063F3C6FAE085EDFA60" box="[1930,2014,1400,1427]" gridcol="11" gridrow="20" pageId="14" pageNumber="517">0</td>
</tr>
<tr id="88B134C0FFD10063F4E9FA3885EDFA4A" box="[165,2014,1440,1465]" gridrow="21" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FA3883D8FA4A" box="[165,491,1440,1465]" gridcol="0" gridrow="21" pageId="14" pageNumber="517">Subdigital Lamellae F1: F2: F3:</th>
<td id="CB605DBCFFD10063F644FA3880B1FA4A" box="[520,642,1440,1465]" gridcol="1" gridrow="21" pageId="14" pageNumber="517">4: 8: 10:</td>
<td id="CB605DBCFFD10063F6EDFA388117FA4A" box="[673,804,1440,1465]" gridcol="2" gridrow="21" pageId="14" pageNumber="517"></td>
<td id="CB605DBCFFD10063F708FA38818FFA4A" box="[836,956,1440,1465]" gridcol="3" gridrow="21" pageId="14" pageNumber="517">56: 79:</td>
<td id="CB605DBCFFD10063F795FA38862CFA4A" box="[985,1055,1440,1465]" gridcol="4" gridrow="21" pageId="14" pageNumber="517">6: 8:</td>
<td id="CB605DBCFFD10063F07BFA38868EFA4A" box="[1079,1213,1440,1465]" gridcol="5" gridrow="21" pageId="14" pageNumber="517">56:</td>
<td id="CB605DBCFFD10063F092FA38877CFA4A" box="[1246,1359,1440,1465]" gridcol="6" gridrow="21" pageId="14" pageNumber="517">5: 89:</td>
<td id="CB605DBCFFD10063F121FA3887EAFA4A" box="[1389,1497,1440,1465]" gridcol="7" gridrow="21" pageId="14" pageNumber="517">46:</td>
<td id="CB605DBCFFD10063F1B5FA38846CFA4A" box="[1529,1631,1440,1465]" gridcol="8" gridrow="21" pageId="14" pageNumber="517">56: 89:</td>
<td id="CB605DBCFFD10063F2CDFA38853CFA4A" box="[1665,1807,1440,1465]" gridcol="9" gridrow="21" pageId="14" pageNumber="517">4: 8: 11: 12: 5</td>
<td id="CB605DBCFFD10063F360FA38855FFA4A" box="[1836,1900,1440,1465]" gridcol="10" gridrow="21" pageId="14" pageNumber="517">4: 9:</td>
<td id="CB605DBCFFD10063F3C6FA3885EDFA4A" box="[1930,2014,1440,1465]" gridcol="11" gridrow="21" pageId="14" pageNumber="517">4: 8: 11:</td>
</tr>
<tr id="88B134C0FFD10063F4E9FA5C85EDFA2E" box="[165,2014,1476,1501]" gridrow="22" pageId="14" pageNumber="517" rowspan-2="1" rowspan-9="1">
<th id="CB605DBCFFD10063F4E9FA5C83D8FA2E" box="[165,491,1476,1501]" gridcol="0" gridrow="22" pageId="14" pageNumber="517">F4: F5</th>
<td id="CB605DBCFFD10063F644FA5C80B1FA2E" box="[520,642,1476,1501]" gridcol="1" gridrow="22" pageId="14" pageNumber="517">12: 8</td>
<td id="CB605DBCFFD10063F708FA5C818FFA2E" box="[836,956,1476,1501]" gridcol="3" gridrow="22" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F708FA5C81B4FA2E" box="[836,903,1476,1501]" pageId="14" pageNumber="517" value="2012-10-08">1012</date>
:
</td>
<td id="CB605DBCFFD10063F795FA5C862CFA2E" box="[985,1055,1476,1501]" gridcol="4" gridrow="22" pageId="14" pageNumber="517">11:</td>
<td id="CB605DBCFFD10063F07BFA5C868EFA2E" box="[1079,1213,1476,1501]" gridcol="5" gridrow="22" pageId="14" pageNumber="517">910: 1112:</td>
<td id="CB605DBCFFD10063F092FA5C877CFA2E" box="[1246,1359,1476,1501]" gridcol="6" gridrow="22" pageId="14" pageNumber="517">1112:</td>
<td id="CB605DBCFFD10063F121FA5C87EAFA2E" box="[1389,1497,1476,1501]" gridcol="7" gridrow="22" pageId="14" pageNumber="517">79: 911:</td>
<td id="CB605DBCFFD10063F1B5FA5C846CFA2E" box="[1529,1631,1476,1501]" gridcol="8" gridrow="22" pageId="14" pageNumber="517">11: 11: 8</td>
<td id="CB605DBCFFD10063F360FA5C855FFA2E" box="[1836,1900,1476,1501]" gridcol="10" gridrow="22" pageId="14" pageNumber="517">11:</td>
<td id="CB605DBCFFD10063F3C6FA5C85EDFA2E" box="[1930,2014,1476,1501]" gridcol="11" gridrow="22" pageId="14" pageNumber="517">12: 0</td>
</tr>
<tr id="88B134C0FFD10063F4E9FA7085EDF9D2" box="[165,2014,1512,1569]" gridrow="23" pageId="14" pageNumber="517">
<th id="CB605DBCFFD10063F4E9FA7083D8F9D2" box="[165,491,1512,1569]" gridcol="0" gridrow="23" pageId="14" pageNumber="517">Subdigital Lamellae T1: T2:</th>
<td id="CB605DBCFFD10063F644FA7080B1F9D2" box="[520,642,1512,1569]" gridcol="1" gridrow="23" pageId="14" pageNumber="517">6: 9: 11: 14:</td>
<td id="CB605DBCFFD10063F6EDFA708117F9D2" box="[673,804,1512,1569]" gridcol="2" gridrow="23" pageId="14" pageNumber="517"></td>
<td id="CB605DBCFFD10063F708FA70818FF9D2" box="[836,956,1512,1569]" gridcol="3" gridrow="23" pageId="14" pageNumber="517">1113: 79 57: 810:</td>
<td id="CB605DBCFFD10063F795FA70862CF9D2" box="[985,1055,1512,1569]" gridcol="4" gridrow="23" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F795FA7081C5F9F2" box="[985,1014,1512,1537]" pageId="14" pageNumber="517" value="2012-09-07">12</date>
: 8 6: 9:
</td>
<td id="CB605DBCFFD10063F07BFA70868EF9D2" box="[1079,1213,1512,1569]" gridcol="5" gridrow="23" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F07BFA708649F9F2" box="[1079,1146,1512,1537]" pageId="14" pageNumber="517" value="2013-12-08">1213</date>
: 89 57: 810:
</td>
<td id="CB605DBCFFD10063F092FA70877CF9D2" box="[1246,1359,1512,1569]" gridcol="6" gridrow="23" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F092FA708713F9F2" box="[1246,1312,1512,1537]" pageId="14" pageNumber="517" value="2013-11-08" valueMax="2013-11-09" valueMin="2013-11-08">1113</date>
: 89 6: 810:
</td>
<td id="CB605DBCFFD10063F121FA7087EAF9D2" box="[1389,1497,1512,1569]" gridcol="7" gridrow="23" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F121FA708792F9F2" box="[1389,1441,1512,1537]" pageId="14" pageNumber="517" value="2011-09-08" valueMax="2011-09-09" valueMin="2011-09-08">911</date>
: 7.9 5: 89:
</td>
<td id="CB605DBCFFD10063F1B5FA70846CF9D2" box="[1529,1631,1512,1569]" gridcol="8" gridrow="23" pageId="14" pageNumber="517">6: 910:</td>
<td id="CB605DBCFFD10063F2CDFA70853CF9D2" box="[1665,1807,1512,1569]" gridcol="9" gridrow="23" pageId="14" pageNumber="517">19: 10: 12:13:</td>
<td id="CB605DBCFFD10063F360FA70855FF9D2" box="[1836,1900,1512,1569]" gridcol="10" gridrow="23" pageId="14" pageNumber="517">12: 4 9: 11:</td>
<td id="CB605DBCFFD10063F3C6FA7085EDF9D2" box="[1930,2014,1512,1569]" gridcol="11" gridrow="23" pageId="14" pageNumber="517"></td>
</tr>
<tr id="88B134C0FFD10063F4E9F9B485EDF9B6" box="[165,2014,1580,1605]" gridrow="24" pageId="14" pageNumber="517" rowspan-11="1" rowspan-2="1">
<th id="CB605DBCFFD10063F4E9F9B483D8F9B6" box="[165,491,1580,1605]" gridcol="0" gridrow="24" pageId="14" pageNumber="517">T3: T4: T5</th>
<td id="CB605DBCFFD10063F644F9B480B1F9B6" box="[520,642,1580,1605]" gridcol="1" gridrow="24" pageId="14" pageNumber="517">10</td>
<td id="CB605DBCFFD10063F708F9B4818FF9B6" box="[836,956,1580,1605]" gridcol="3" gridrow="24" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F708F9B4816FF9B6" box="[836,860,1580,1605]" pageId="14" pageNumber="517" value="2011-10-05">11</date>
12:
</td>
<td id="CB605DBCFFD10063F795F9B4862CF9B6" box="[985,1055,1580,1605]" gridcol="4" gridrow="24" pageId="14" pageNumber="517">11:</td>
<td id="CB605DBCFFD10063F07BF9B4868EF9B6" box="[1079,1213,1580,1605]" gridcol="5" gridrow="24" pageId="14" pageNumber="517">1114: 14</td>
<td id="CB605DBCFFD10063F092F9B4877CF9B6" box="[1246,1359,1580,1605]" gridcol="6" gridrow="24" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F092F9B48712F9B6" box="[1246,1313,1580,1605]" pageId="14" pageNumber="517" value="2013-12-14">1213</date>
:
</td>
<td id="CB605DBCFFD10063F121F9B487EAF9B6" box="[1389,1497,1580,1605]" gridcol="7" gridrow="24" pageId="14" pageNumber="517">1012:</td>
<td id="CB605DBCFFD10063F1B5F9B4846CF9B6" box="[1529,1631,1580,1605]" gridcol="8" gridrow="24" pageId="14" pageNumber="517">1112:</td>
<td id="CB605DBCFFD10063F2CDF9B4853CF9B6" box="[1665,1807,1580,1605]" gridcol="9" gridrow="24" pageId="14" pageNumber="517">10</td>
<td id="CB605DBCFFD10063F360F9B4855FF9B6" box="[1836,1900,1580,1605]" gridcol="10" gridrow="24" pageId="14" pageNumber="517">11:</td>
</tr>
<tr id="88B134C0FFD10063F4E9F9C885EDF99A" box="[165,2014,1616,1641]" gridrow="25" pageId="14" pageNumber="517" rowspan-0="1" rowspan-1="1" rowspan-11="1" rowspan-2="1" rowspan-9="1">
<td id="CB605DBCFFD10063F708F9C8818FF99A" box="[836,956,1616,1641]" gridcol="3" gridrow="25" pageId="14" pageNumber="517">1314:</td>
<td id="CB605DBCFFD10063F795F9C8862CF99A" box="[985,1055,1616,1641]" gridcol="4" gridrow="25" pageId="14" pageNumber="517">13:</td>
<td id="CB605DBCFFD10063F07BF9C8868EF99A" box="[1079,1213,1616,1641]" gridcol="5" gridrow="25" pageId="14" pageNumber="517">17: 1012</td>
<td id="CB605DBCFFD10063F092F9C8877CF99A" box="[1246,1359,1616,1641]" gridcol="6" gridrow="25" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F092F9C88713F99A" box="[1246,1312,1616,1641]" pageId="14" pageNumber="517" value="2010-12-12" valueMax="2010-12-15" valueMin="2010-12-12">1215</date>
:
</td>
<td id="CB605DBCFFD10063F121F9C887EAF99A" box="[1389,1497,1616,1641]" gridcol="7" gridrow="25" pageId="14" pageNumber="517">1114:</td>
<td id="CB605DBCFFD10063F1B5F9C8846CF99A" box="[1529,1631,1616,1641]" gridcol="8" gridrow="25" pageId="14" pageNumber="517">14: 1011</td>
<td id="CB605DBCFFD10063F360F9C8855FF99A" box="[1836,1900,1616,1641]" gridcol="10" gridrow="25" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F360F9C8857AF99A" box="[1836,1865,1616,1641]" pageId="14" pageNumber="517" value="2012-11-10">12</date>
: 9
</td>
</tr>
<tr id="88B134C0FFD10063F4E9F9EC85EDF97E" box="[165,2014,1652,1677]" gridrow="26" pageId="14" pageNumber="517" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
<td id="CB605DBCFFD10063F708F9EC818FF97E" box="[836,956,1652,1677]" gridcol="3" gridrow="26" pageId="14" pageNumber="517">1011</td>
<td id="CB605DBCFFD10063F795F9EC862CF97E" box="[985,1055,1652,1677]" gridcol="4" gridrow="26" pageId="14" pageNumber="517">11</td>
<td id="CB605DBCFFD10063F092F9EC877CF97E" box="[1246,1359,1652,1677]" gridcol="6" gridrow="26" pageId="14" pageNumber="517">
<date id="423F1042FFD1FF92F092F9EC86C4F97E" box="[1246,1271,1652,1677]" pageId="14" pageNumber="517" value="2010-11-10" valueMax="2010-11-11" valueMin="2010-11-10">10</date>
12
</td>
<td id="CB605DBCFFD10063F121F9EC87EAF97E" box="[1389,1497,1652,1677]" gridcol="7" gridrow="26" pageId="14" pageNumber="517">911</td>
</tr>
</table>
</paragraph>
<paragraph id="363E3682FFD0FF93F48BFF008764FEEC" blockId="15.[151,1437,152,827]" pageId="15" pageNumber="518">
<emphasis id="04F5EA90FFD0FF93F48BFF00830FFF40" bold="true" box="[199,316,152,179]" pageId="15" pageNumber="518">Variation.</emphasis>
Variation in meristic counts of the type series is reported in
<tableCitation id="7B030339FFD0FF93F78DFF018620FF40" box="[961,1043,153,179]" captionStart="TABLE 2" captionStartId="14.[151,239,513,539]" captionTargetPageId="14" captionText="TABLE 2. Variation in meristic counts for specimens of Cordylus phonolithos sp. nov., C. namakuiyus, C. machadoi and C. angolensis. All measurements are presented in millimeters (mm). Data of C. namakuiyus retrieved from Stanley et al. (2016); data for C. machadoi retrieved from Stanley et al. (2016) and two additional specimens from this study; data for C. phonolithos sp. nov. and C. angolensis collected in this study" httpUri="http://table.plazi.org/id/62FE660AFFD1FF92F4DBFD998196FD90" pageId="15" pageNumber="518" tableUuid="62FE660AFFD1FF92F4DBFD998196FD90">Table 2</tableCitation>
. The single juvenile
<typeStatus id="E93A8820FFD0FF93F0BEFF018763FF40" box="[1266,1360,153,179]" pageId="15" pageNumber="518" type="paratype">paratype</typeStatus>
agrees almost entirely with the
<typeStatus id="E93A8820FFD0FF93F5D2FF258033FF24" box="[414,512,189,215]" pageId="15" pageNumber="518" type="holotype">holotype</typeStatus>
, although, the loreal and the preocular scales are fused and the right posterior parietal is entire. Measurements of the
<typeStatus id="E93A8820FFD0FF93F5A1FF798078FF08" box="[493,587,225,251]" pageId="15" pageNumber="518" type="paratype">paratype</typeStatus>
are the following: SVL
<quantity id="F1799B67FFD0FF93F716FF79818CFF08" box="[858,959,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="4.99" pageId="15" pageNumber="518" unit="mm" value="49.9">49.9 mm</quantity>
; HL
<quantity id="F1799B67FFD0FF93F7BBFF78866EFF08" box="[1015,1117,224,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="15" pageNumber="518" unit="mm" value="15.2">15.2 mm</quantity>
; HW
<quantity id="F1799B67FFD0FF93F0D2FF798737FF08" box="[1182,1284,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="15" pageNumber="518" unit="mm" value="10.7">10.7 mm</quantity>
; HH
<quantity id="F1799B67FFD0FF93F10CFF7987AAFF08" box="[1344,1433,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="15" pageNumber="518" unit="mm" value="6.4">6.4 mm</quantity>
; SEL
<quantity id="F1799B67FFD0FF93F482FE9C8316FEEC" box="[206,293,260,287]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="15" pageNumber="518" unit="mm" value="5.8">5.8 mm</quantity>
; SAL
<quantity id="F1799B67FFD0FF93F526FE9D83FDFEEC" box="[362,462,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="15" pageNumber="518" unit="mm" value="20.2">20.2 mm</quantity>
; AGD
<quantity id="F1799B67FFD0FF93F656FE9D804CFEEC" box="[538,639,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="15" pageNumber="518" unit="mm" value="23.1">23.1 mm</quantity>
; HML
<quantity id="F1799B67FFD0FF93F681FE9D8123FEEC" box="[717,784,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="518" unit="mm" value="6.0">6 mm</quantity>
; RUL
<quantity id="F1799B67FFD0FF93F714FE9C819CFEEC" box="[856,943,260,287]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="15" pageNumber="518" unit="mm" value="5.4">5.4 mm</quantity>
; FL
<quantity id="F1799B67FFD0FF93F7ACFE9D8604FEEC" box="[992,1079,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="15" pageNumber="518" unit="mm" value="7.9">7.9 mm</quantity>
; TFL
<quantity id="F1799B67FFD0FF93F034FE9D86FCFEEC" box="[1144,1231,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="15" pageNumber="518" unit="mm" value="7.8">7.8 mm</quantity>
; LTL
<quantity id="F1799B67FFD0FF93F15CFE9D8760FEEC" box="[1296,1363,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="518" unit="mm" value="8.0">8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="7E9B6509FFD0FF93F48BFEB08626FDCC" pageId="15" pageNumber="518" type="distribution">
<paragraph id="363E3682FFD0FF93F48BFEB086CBFE78" blockId="15.[151,1437,152,827]" pageId="15" pageNumber="518">
<emphasis id="04F5EA90FFD0FF93F48BFEB08352FEB0" bold="true" box="[199,353,296,323]" pageId="15" pageNumber="518">Distribution.</emphasis>
The new species is known only from NDolondolo, at the base of the Serra de Neve Inselberg in
<collectingRegion id="F445F860FFD0FF93F4DBFED58368FE94" box="[151,347,333,359]" country="Angola" name="Namibe" pageId="15" pageNumber="518">Namibe Province</collectingRegion>
,
<collectingCountry id="4E967612FFD0FF93F528FED58389FE94" box="[356,442,333,359]" name="Angola" pageId="15" pageNumber="518">Angola</collectingCountry>
. The observational record cited by
<bibRefCitation id="52104B73FFD0FF93F777FED5862DFE9B" author="Stanley, E. L. &amp; Ceriaco, L. M. P. &amp; Bandeira, S. &amp; Valerio, H. &amp; Bates, M. F. &amp; Branch, W. R." box="[827,1054,333,360]" pageId="15" pageNumber="518" pagination="201 - 226" refId="ref12195" refString="Stanley, E. L., Ceriaco, L. M. P., Bandeira, S., Valerio, H., Bates, M. F. &amp; Branch, W. R. (2016) A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola with the description of a new species from the Pro-Namib desert. Zootaxa, 4061 (3), 201 - 226. https: // doi. org / 10.11646 / zootaxa. 4061.3.1" type="journal article" year="2016">
Stanley
<emphasis id="04F5EA90FFD0FF93F7D9FED681FFFE94" box="[917,972,333,359]" italics="true" pageId="15" pageNumber="518">et al.</emphasis>
(2016)
</bibRefCitation>
, originally identified as
<taxonomicName id="F1814D01FFD0FF93F165FED582E1FE78" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="518" phylum="Chordata" rank="species" species="namakuyius">
<emphasis id="04F5EA90FFD0FF93F165FED582E1FE78" italics="true" pageId="15" pageNumber="518">C. namakuyius</emphasis>
</taxonomicName>
from “Sera [sic] de Neve” (= Serra da Neve) is likely to correspond to
<taxonomicName id="F1814D01FFD0FF93F7BDFEE986A5FE78" authority="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[1009,1174,369,395]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="518" phylum="Chordata" rank="species" species="phonolithos" status="sp. nov.">
<emphasis id="04F5EA90FFD0FF93F7BDFEE986A5FE78" box="[1009,1174,369,395]" italics="true" pageId="15" pageNumber="518">C. phonolithos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FC657EBFFD0FF93F0D1FEE986CBFE78" box="[1181,1272,369,395]" pageId="15" pageNumber="518" rank="species">
<emphasis id="04F5EA90FFD0FF93F0D1FEE986C2FE78" bold="true" box="[1181,1265,369,395]" pageId="15" pageNumber="518">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
<paragraph id="363E3682FFD0FF93F48BFE0C8626FDCC" blockId="15.[151,1437,152,827]" pageId="15" pageNumber="518">
<emphasis id="04F5EA90FFD0FF93F48BFE0C805FFE5C" bold="true" box="[199,620,404,431]" pageId="15" pageNumber="518">Habitat and Natural history notes.</emphasis>
This species was found in granite outcrops in “sparse Miombo” forest (
<figureCitation id="AEBA2A07FFD0FF93F4D3FE2182C2FE20" box="[159,241,441,467]" captionStart="FIGURE 10" captionStartId="12.[151,250,1899,1924]" captionTargetBox="[157,1432,919,1875]" captionTargetId="figure@12.[151,1436,913,1879]" captionTargetPageId="12" captionText="FIGURE 10. Habitat view of the type locality of Cordylus phonolithos sp. nov. (Photo by Ishan Agarwal)." figureDoi="http://doi.org/10.5281/zenodo.3449928" httpUri="https://zenodo.org/record/3449928/files/figure.png" pageId="15" pageNumber="518">Fig. 10</figureCitation>
), dominated by trees of the genera
<taxonomicName id="F1814D01FFD0FF93F634FE21813FFE20" authorityName="Bentham" authorityYear="1865" box="[632,780,441,467]" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="518" phylum="Tracheophyta" rank="genus">
<emphasis id="04F5EA90FFD0FF93F634FE21813FFE20" box="[632,780,441,467]" italics="true" pageId="15" pageNumber="518">Brachystegia</emphasis>
</taxonomicName>
and
<emphasis id="04F5EA90FFD0FF93F70DFE2181F9FE20" box="[833,970,441,467]" italics="true" pageId="15" pageNumber="518">Jubernardia</emphasis>
(
<bibRefCitation id="52104B73FFD0FF93F794FE2186CDFE20" author="Grandvaux-Barbosa L. A." box="[984,1278,441,467]" pageId="15" pageNumber="518" refId="ref10577" refString="Grandvaux-Barbosa L. A. (1970) Carta fitogeorafica de Angola. Instituto de Investigacao Cientifica de Angola, Luanda, 323 pp." type="book" year="1970">Grandvaux-Barbosa 1970</bibRefCitation>
). The juvenile
<typeStatus id="E93A8820FFD0FF93F4DBFE4582C4FE04" box="[151,247,477,503]" pageId="15" pageNumber="518" type="paratype">paratype</typeStatus>
was found under a small granite rock on a larger rock, while the
<typeStatus id="E93A8820FFD0FF93F78EFE458617FE04" box="[962,1060,477,503]" pageId="15" pageNumber="518" type="holotype">holotype</typeStatus>
was collected from within a crevice in a granite boulder. The latter was exposed outside of the crevice, quickly sheltering in response to our presence. CT scanning of the
<typeStatus id="E93A8820FFD0FF93F538FDBD83E5FDCC" box="[372,470,549,575]" pageId="15" pageNumber="518" type="holotype">holotype</typeStatus>
revealed a myriapod carapace in its digestive tract.
</paragraph>
</subSubSection>
<subSubSection id="7E9B6509FFD0FF93F48BFDD181E2FCC8" pageId="15" pageNumber="518" type="etymology">
<paragraph id="363E3682FFD0FF93F48BFDD181E2FCC8" blockId="15.[151,1437,152,827]" pageId="15" pageNumber="518">
<emphasis id="04F5EA90FFD0FF93F48BFDD1837DFD90" bold="true" box="[199,334,585,611]" pageId="15" pageNumber="518">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="F1814D01FFD0FF93F61DFDD180E6FD90" authorityName="Marques &amp; Ceríaco &amp; Stanley &amp; Bandeira &amp; Agarwal &amp; Bauer" authorityYear="2019" box="[593,725,585,611]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="518" phylum="Chordata" rank="species" species="phonolithos">
<emphasis id="04F5EA90FFD0FF93F61DFDD180E6FD90" box="[593,725,585,611]" italics="true" pageId="15" pageNumber="518">phonolithos</emphasis>
</taxonomicName>
” is a noun in apposition from the Greek “
<emphasis id="04F5EA90FFD0FF93F08AFDD1872FFD90" box="[1222,1308,585,611]" italics="true" pageId="15" pageNumber="518">phono”</emphasis>
= sound + “
<emphasis id="04F5EA90FFD0FF93F4E8FDF582C1FD74" box="[164,242,621,647]" italics="true" pageId="15" pageNumber="518">lithos”</emphasis>
= rock, which means “sound stone”. In the local Mucobal language the
<typeStatus id="E93A8820FFD0FF93F05FFDF58670FD74" box="[1043,1091,621,647]" pageId="15" pageNumber="518">type</typeStatus>
locality name, “NDolondolo” means literally “rock that sounds like a bell” or “bell” and stems from the presence of a large and famous phonolite stone at the locality. Phonolites are rare igneous volcanic stones of intermediate composition between felsic and maphic, with aphanitic to porphyritic texture that produce a very distinctive metallic sound when hit, similar to the ringing of a metallic bell. We suggest “NDolondolo Girdled Lizard” and “Lagarto Espinhoso de NDolondolo” as the English and Portuguese common names, respectively, for this species.
</paragraph>
</subSubSection>
</treatment>
</document>