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<document id="651B8BD6D4D70F6EC5FD83451F9BA05F" ID-CLB-Dataset="105313" ID-DOI="10.5852/ejt.2022.831.1871" ID-GBIF-Dataset="e697ba5c-5a96-4a28-985c-748890333d6c" ID-ISSN="2118-9773" ID-Zenodo-Dep="6885198" ID-ZooBank="EAA2EAE2-DA8B-4516-B332-CB8423B8EEAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658507519427" checkinUser="felipe" docAuthor="Korn, Dieter &amp; Bockwinkel, Jürgen" docDate="2022" docId="9327592FF4321948FD9F1AAEFC0250DD" docLanguage="en" docName="ejt-831_67-108.pdf" docOrigin="European Journal of Taxonomy 831" docSource="http://dx.doi.org/10.5852/ejt.2022.831.1871" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Vestinautilus bicristatus Korn &amp; Bockwinkel 2022, sp. nov." docType="treatment" docUuid="69CABBA5-6CDA-4264-B156-0FB2321AEDD5" docUuidSource="ZooBank" docVersion="7" lastPageNumber="92" masterDocId="6F1E2157F42A1952FFE11E6BFF97516E" masterDocTitle="Early Carboniferous nautiloids from the Central Sahara, southern Algeria" masterLastPageNumber="108" masterPageNumber="67" pageNumber="91" updateTime="1699361693651" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F31204501813B3C396AA88301FB573BA">Early Carboniferous nautiloids from the Central Sahara, southern Algeria</mods:title>
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<mods:namePart id="3FB0FED954EB8D3A695BBB4670AD5623">Korn, Dieter</mods:namePart>
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<mods:namePart id="50A0927391F97804F42D8C0080581B8D">Bockwinkel, Jürgen</mods:namePart>
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<mods:title id="12C6F9764086B52EB02BB16310195F7F">European Journal of Taxonomy</mods:title>
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<mods:date id="7E76D73FD96123D43C6DECFC6A1F9E75">2022</mods:date>
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<treatment id="9327592FF4321948FD9F1AAEFC0250DD" ID-DOI="http://doi.org/10.5281/zenodo.6903234" ID-GBIF-Taxon="197852054" ID-Zenodo-Dep="6903234" LSID="urn:lsid:zoobank.org:act:69CABBA5-6CDA-4264-B156-0FB2321AEDD5" httpUri="http://treatment.plazi.org/id/9327592FF4321948FD9F1AAEFC0250DD" lastPageId="26" lastPageNumber="92" pageId="24" pageNumber="91">
<subSubSection id="5394BBB2F432194AFD9F1AAEFB96558E" box="[638,1025,1221,1248]" pageId="24" pageNumber="91" type="nomenclature">
<paragraph id="1B31E839F432194AFD9F1AAEFB96558E" blockId="24.[391,1272,1221,1317]" box="[638,1025,1221,1248]" pageId="24" pageNumber="91">
<heading id="40795F55F432194AFD9F1AAEFB96558E" box="[638,1025,1221,1248]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="91" reason="2">
<taxonomicName id="DC8E93BAF432194AFD9F1AAEFC37558E" authority="Korn &amp; Bockwinkel, 2022" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" box="[638,928,1221,1248]" class="Cephalopoda" family="Trigonoceratidae" genus="Vestinautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="24" pageNumber="91" phylum="Mollusca" rank="species" species="bicristatus" status="sp. nov.">
<emphasis id="29FA342BF432194AFD9F1AAEFC37558E" bold="true" box="[638,928,1221,1248]" italics="true" pageId="24" pageNumber="91">Vestinautilus bicristatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C98950F432194AFC461AADFB96558E" box="[935,1025,1222,1248]" pageId="24" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="5394BBB2F432194AFE661A83FC2D544B" pageId="24" pageNumber="91" type="description">
<paragraph id="1B31E839F432194AFE661A83FB6F546D" blockId="24.[391,1272,1221,1317]" box="[391,1272,1256,1283]" pageId="24" pageNumber="91">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="1B31E839F432194AFD241B60FC2D544B" blockId="24.[391,1272,1221,1317]" box="[709,954,1291,1317]" pageId="24" pageNumber="91">
<figureCitation id="83B5F4BCF432194AFD241B60FCDD544B" box="[709,842,1291,1317]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="25.[227,270,1999,2025]" captionStartId-1="26.[227,270,1963,1989]" captionTargetBox-0="[281,1386,1515,1950]" captionTargetBox-1="[232,1430,552,1899]" captionTargetId-0="figure-338@25.[269,1394,1504,1958]" captionTargetId-1="figure-117@26.[227,1437,550,1911]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="Fig. 20. Vestinautilus bicristatus sp. nov., holotype MB.C.30473 (Wendt &amp; Kaufmann Coll.) from Oued Temertasset. Scale bar units = 1 mm." captionText-1="Fig. 21. Vestinautilus bicristatus sp. nov. from Oued Temertasset (all Korn et al. 2002 Coll.). A. Paratype MB.C.30474.1. B. Paratype MB.C.30474.2. C. Paratype MB.C.30474.3. Scale bar units =1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6902636" figureDoi-1="http://doi.org/10.5281/zenodo.6885248" httpUri-0="https://zenodo.org/record/6902636/files/figure.png" httpUri-1="https://zenodo.org/record/6885248/files/figure.png" pageId="24" pageNumber="91">Figs 2021</figureCitation>
,
<tableCitation id="560CDD82F432194AFCB71B60FC2D544B" box="[854,954,1291,1317]" captionStart="Table 11" captionStartId="25.[227,296,302,328]" captionTargetBox="[234,1415,375,448]" captionTargetPageId="25" captionText="Table 11. Conch dimensions (in mm) and ratios of Vestinautilus bicristatus sp. nov." httpUri="http://table.plazi.org/id/4FF1B8B1F433194BFF021F45FB215027" pageId="24" pageNumber="91" tableUuid="4FF1B8B1F433194BFF021F45FB215027">Table 11</tableCitation>
</paragraph>
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<subSubSection id="5394BBB2F432194AFF021B27FC135771" pageId="24" pageNumber="91" type="diagnosis">
<paragraph id="1B31E839F432194AFF021B27FECB5408" blockId="24.[227,348,1356,1382]" box="[227,348,1356,1382]" pageId="24" pageNumber="91">
<heading id="40795F55F432194AFF021B27FECB5408" bold="true" box="[227,348,1356,1382]" fontSize="11" level="3" pageId="24" pageNumber="91" reason="3">
<emphasis id="29FA342BF432194AFF021B27FECB5408" bold="true" box="[227,348,1356,1382]" pageId="24" pageNumber="91">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="1B31E839F432194AFF021B11FC135771" blockId="24.[227,1437,1402,1568]" pageId="24" pageNumber="91">
Species of
<taxonomicName id="DC8E93BAF432194AFE8A1B11FD9654FA" authorityName="Ryckholt" authorityYear="1852" box="[363,513,1402,1428]" class="Cephalopoda" family="Trigonoceratidae" genus="Vestinautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="24" pageNumber="91" phylum="Mollusca" rank="genus">
<emphasis id="29FA342BF432194AFE8A1B11FD9654FA" box="[363,513,1402,1428]" italics="true" pageId="24" pageNumber="91">Vestinautilus</emphasis>
</taxonomicName>
with strongly depressed, rounded-trapezoidal whorl profile (ww/wh ~ 2.10), venter up to
<quantity id="DC7645DCF432194AFE651BF6FE4C54D9" box="[388,475,1437,1463]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="91" unit="mm" value="30.0">30 mm</quantity>
conch diameter broadly arched with two shallow submarginal grooves, which disappear in the adult stage. Ventrolateral shoulder defined by a subangular margin; one sharp ridge is located on the umbilical wall near the ventrolateral shoulder. Whorls weakly embracing, coiling very high (WER ~2.55). Ornament with delicate growth lines.
</paragraph>
</subSubSection>
<subSubSection id="5394BBB2F432194AFF02182DFA9A57E0" box="[227,1293,1606,1678]" pageId="24" pageNumber="91" type="etymology">
<paragraph id="1B31E839F432194AFF02182DFEFD570E" blockId="24.[227,362,1606,1632]" box="[227,362,1606,1632]" pageId="24" pageNumber="91">
<heading id="40795F55F432194AFF02182DFEFD570E" bold="true" box="[227,362,1606,1632]" fontSize="11" level="3" pageId="24" pageNumber="91" reason="3">
<emphasis id="29FA342BF432194AFF02182DFEFD570E" bold="true" box="[227,362,1606,1632]" pageId="24" pageNumber="91">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="1B31E839F432194AFF02181FFA9A57E0" blockId="24.[227,1293,1652,1678]" box="[227,1293,1652,1678]" pageId="24" pageNumber="91">
From the Latin
<emphasis id="29FA342BF432194AFE45181EFE7057E0" box="[420,487,1653,1678]" italics="true" pageId="24" pageNumber="91">crista</emphasis>
, meaning crest and referring to the two lateral longitudinal ridges.
</paragraph>
</subSubSection>
<subSubSection id="5394BBB2F432194AFF0218DEFCC05960" pageId="24" pageNumber="91" type="materials_examined">
<paragraph id="1B31E839F432194AFF0218DEFE0657A0" blockId="24.[227,401,1716,1743]" box="[227,401,1716,1743]" pageId="24" pageNumber="91">
<heading id="40795F55F432194AFF0218DEFE0657A0" bold="true" box="[227,401,1716,1743]" fontSize="11" level="3" pageId="24" pageNumber="91" reason="3">
<emphasis id="29FA342BF432194AFF0218DEFE0657A0" bold="true" box="[227,401,1716,1743]" pageId="24" pageNumber="91">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="1B31E839F432194AFEF11888FE71560B" blockId="24.[227,1436,1763,1893]" pageId="24" pageNumber="91">
<materialsCitation id="ABE6E264F432194AFEF11888FE71560B" ID-GBIF-Occurrence="3865861326" collectionCode="MB" collectorName="Wendt et al." country="Algeria" location="Mouydir" municipality="Argiles de Teguentour" pageId="24" pageNumber="91" specimenCode="MB.C.30473" specimenCount="1" typeStatus="holotype">
<emphasis id="29FA342BF432194AFEF11888FE155793" bold="true" box="[272,386,1763,1789]" pageId="24" pageNumber="91">
<typeStatus id="C435569BF432194AFEF11888FE155793" box="[272,386,1763,1789]" pageId="24" pageNumber="91" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="6399A8A9F432194AFF02196EFEF35671" box="[227,356,1797,1823]" name="Algeria" pageId="24" pageNumber="91">ALGERIA</collectingCountry>
<location id="1E51BEE2F432194AFE70196EFE6D5671" LSID="urn:lsid:plazi:treatment:9327592FF4321948FD9F1AAEFC0250DD:1E51BEE2F432194AFE70196EFE6D5671" box="[401,506,1797,1823]" country="Algeria" municipality="Argiles de Teguentour" name="Mouydir" pageId="24" pageNumber="91">Mouydir</location>
,
<locationDeviation id="0A83F11BF432194AFDF1196EFBFB5671" box="[528,1132,1797,1824]" pageId="24" pageNumber="91">south of Oued Temertasset (locality MOU-C1)</locationDeviation>
;
<location id="1E51BEE2F432194AFB63196EFA0B564E" LSID="urn:lsid:plazi:treatment:9327592FF4321948FD9F1AAEFC0250DD:1E51BEE2F432194AFB63196EFA0B564E" box="[1154,1436,1797,1824]" country="Algeria" municipality="Argiles de Teguentour" name="Argiles de Teguentour" pageId="24" pageNumber="91">Argiles de Teguentour</location>
(
<geologicalTimeScale id="69AE6892F432194AFF0C1943FC63562C" box="[237,1012,1832,1858]" pageId="24" pageNumber="91">
<emphasis id="29FA342BF432194AFF0C1943FDA2562C" box="[237,565,1832,1858]" italics="true" pageId="24" pageNumber="91">Helicocyclus-Ouaoufilalites</emphasis>
Assemblage; early late Tournaisian
</geologicalTimeScale>
);
<collectorName id="B67B8DEFF432194AFBED1943FB31562D" box="[1036,1190,1832,1859]" pageId="24" pageNumber="91">
Wendt
<emphasis id="29FA342BF432194AFB871942FB08562C" box="[1126,1183,1832,1858]" italics="true" pageId="24" pageNumber="91">et al</emphasis>
.
</collectorName>
Coll.; illustrated in
<figureCitation id="83B5F4BCF432194AFF021920FEAD560B" box="[227,314,1867,1893]" captionStart="Fig" captionStartId="25.[227,270,1999,2025]" captionTargetBox="[281,1386,1515,1950]" captionTargetId="figure-338@25.[269,1394,1504,1958]" captionTargetPageId="25" captionText="Fig. 20. Vestinautilus bicristatus sp. nov., holotype MB.C.30473 (Wendt &amp; Kaufmann Coll.) from Oued Temertasset. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902636" httpUri="https://zenodo.org/record/6902636/files/figure.png" pageId="24" pageNumber="91">Fig. 20</figureCitation>
;
<specimenCode id="4B284042F432194AFEA41920FE75560B" box="[325,482,1867,1893]" pageId="24" pageNumber="91">MB.C.30473</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="1B31E839F432194AFEF119E7FCC05960" blockId="24.[227,1437,1932,2062]" pageId="24" pageNumber="91">
<materialsCitation id="ABE6E264F432194AFEF119E7FCC05960" ID-GBIF-Occurrence="3865861320" collectionCode="MB" collectorName="Korn et al." country="Algeria" location="Mouydir" municipality="Argiles de Teguentour" pageId="24" pageNumber="91" specimenCode="MB.C.30474.1, MB.C.30474.3" specimenCount="3" typeStatus="paratype">
<emphasis id="29FA342BF432194AFEF119E7FE1A56C8" bold="true" box="[272,397,1932,1958]" pageId="24" pageNumber="91">
<typeStatus id="C435569BF432194AFEF119E7FE1A56C8" box="[272,397,1932,1958]" pageId="24" pageNumber="91" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="6399A8A9F432194AFF0219C5FEF356A6" box="[227,356,1966,1992]" name="Algeria" pageId="24" pageNumber="91">ALGERIA</collectingCountry>
<specimenCount id="0D8823B0F432194AFE6A19C5FDBF56A6" box="[395,552,1966,1992]" pageId="24" pageNumber="91" type="generic">3 specimens</specimenCount>
;
<location id="1E51BEE2F432194AFDDD19C5FD3256A6" LSID="urn:lsid:plazi:treatment:9327592FF4321948FD9F1AAEFC0250DD:1E51BEE2F432194AFDDD19C5FD3256A6" box="[572,677,1966,1992]" country="Algeria" municipality="Argiles de Teguentour" name="Mouydir" pageId="24" pageNumber="91">Mouydir</location>
,
<locationDeviation id="0A83F11BF432194AFD5919C5FA9F56A6" box="[696,1288,1966,1993]" pageId="24" pageNumber="91">south of Oued Temertasset (locality MOU-C1)</locationDeviation>
;
<location id="1E51BEE2F432194AFAFA19C5FEFF5685" LSID="urn:lsid:plazi:treatment:9327592FF4321948FD9F1AAEFC0250DD:1E51BEE2F432194AFAFA19C5FEFF5685" country="Algeria" municipality="Argiles de Teguentour" name="Argiles de Teguentour" pageId="24" pageNumber="91">Argiles de Teguentour</location>
(
<geologicalTimeScale id="69AE6892F432194AFE9A19BAFBE35685" box="[379,1140,2001,2027]" pageId="24" pageNumber="91">
<emphasis id="29FA342BF432194AFE9A19BAFD545685" box="[379,707,2001,2027]" italics="true" pageId="24" pageNumber="91">Helicocyclus-Ouaoufilalites</emphasis>
Assemblage; early late Tournaisian
</geologicalTimeScale>
);
<collectorName id="B67B8DEFF432194AFB6819BAFA9B5685" box="[1161,1292,2001,2027]" pageId="24" pageNumber="91">
Korn
<emphasis id="29FA342BF432194AFB3319B9FA925685" box="[1234,1285,2001,2027]" italics="true" pageId="24" pageNumber="91">et al</emphasis>
.
</collectorName>
2002 Coll.; illustrated in
<figureCitation id="83B5F4BCF432194AFE61199FFE415960" box="[384,470,2036,2062]" captionStart="Fig" captionStartId="26.[227,270,1963,1989]" captionTargetBox="[232,1430,552,1899]" captionTargetId="figure-117@26.[227,1437,550,1911]" captionTargetPageId="26" captionText="Fig. 21. Vestinautilus bicristatus sp. nov. from Oued Temertasset (all Korn et al. 2002 Coll.). A. Paratype MB.C.30474.1. B. Paratype MB.C.30474.2. C. Paratype MB.C.30474.3. Scale bar units =1 mm." figureDoi="http://doi.org/10.5281/zenodo.6885248" httpUri="https://zenodo.org/record/6885248/files/figure.png" pageId="24" pageNumber="91">Fig. 21</figureCitation>
;
<specimenCode id="4B284042F432194AFE02199FFD055960" box="[483,658,2036,2062]" pageId="24" pageNumber="91">MB.C.30474.1</specimenCode>
<specimenCode id="4B284042F432194AFD43199FFCC45960" box="[674,851,2036,2062]" pageId="24" pageNumber="91">MB.C.30474.3</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="4FF1B8B1F433194BFF021F45FB215027" ID-Table-UUID="4FF1B8B1F433194BFF021F45FB215027" box="[227,1206,302,329]" httpUri="http://table.plazi.org/id/4FF1B8B1F433194BFF021F45FB215027" pageId="25" pageNumber="92" startId="25.[227,296,302,328]" targetBox="[234,1415,375,448]" targetIsTable="true" targetPageId="25">
<paragraph id="1B31E839F433194BFF021F45FB215027" blockId="25.[227,1206,302,329]" box="[227,1206,302,329]" pageId="25" pageNumber="92">
<emphasis id="29FA342BF433194BFF021F45FEC45026" bold="true" box="[227,339,302,328]" pageId="25" pageNumber="92">Table 11.</emphasis>
Conch dimensions (in mm) and ratios of
<taxonomicName id="DC8E93BAF433194BFCDA1F45FBC35026" authority="Korn &amp; Bockwinkel, 2022" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" box="[827,1108,302,328]" class="Cephalopoda" family="Trigonoceratidae" genus="Vestinautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="25" pageNumber="92" phylum="Mollusca" rank="species" species="bicristatus" status="sp. nov.">
<emphasis id="29FA342BF433194BFCDA1F45FBC35026" box="[827,1108,302,328]" italics="true" pageId="25" pageNumber="92">Vestinautilus bicristatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C98950F433194BFBBA1F44FB215027" box="[1115,1206,303,329]" pageId="25" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="1B31E839F433194BFEED1F1CFA1450AE" pageId="25" pageNumber="92">
<table id="698E1A99F433E6ADFF0B1F1CFA1050AE" box="[234,1415,375,448]" gridcols="11" gridrows="2" pageId="25" pageNumber="92">
<tr id="A5BEEA7BF433E6ADFF0B1F1CFA1050E1" box="[234,1415,375,399]" gridrow="0" pageId="25" pageNumber="92">
<th id="E66F8307F433E6ADFF0B1F1CFEED50E1" box="[234,378,375,399]" gridcol="0" gridrow="0" pageId="25" pageNumber="92">Specimen</th>
<td id="E66F8307F433E6ADFE571F1CFE7250E1" box="[438,485,375,399]" gridcol="1" gridrow="0" pageId="25" pageNumber="92">dm</td>
<td id="E66F8307F433E6ADFDEE1F1CFDA950E1" box="[527,574,375,399]" gridcol="2" gridrow="0" pageId="25" pageNumber="92">ww</td>
<td id="E66F8307F433E6ADFD8C1F1CFD0B50E1" box="[621,668,375,399]" gridcol="3" gridrow="0" pageId="25" pageNumber="92">wh</td>
<td id="E66F8307F433E6ADFD311F1CFD6850E1" box="[720,767,375,399]" gridcol="4" gridrow="0" pageId="25" pageNumber="92">uw</td>
<td id="E66F8307F433E6ADFCCC1F1CFCCB50E1" box="[813,860,375,399]" gridcol="5" gridrow="0" pageId="25" pageNumber="92">ah</td>
<td id="E66F8307F433E6ADFC631F1CFC4250E1" box="[898,981,375,399]" gridcol="6" gridrow="0" pageId="25" pageNumber="92">ww/dm</td>
<td id="E66F8307F433E6ADFC1B1F1CFBDD50E1" box="[1018,1098,375,399]" gridcol="7" gridrow="0" pageId="25" pageNumber="92">ww/wh</td>
<td id="E66F8307F433E6ADFB8E1F1CFB2A50E1" box="[1135,1213,375,399]" gridcol="8" gridrow="0" pageId="25" pageNumber="92">uw/dm</td>
<td id="E66F8307F433E6ADFB041F1CFAB250E1" box="[1253,1317,375,399]" gridcol="9" gridrow="0" pageId="25" pageNumber="92">
<collectionCode id="7D9F70FCF433194BFB041F1CFAB250E1" box="[1253,1317,375,399]" pageId="25" pageNumber="92">WER</collectionCode>
</td>
<td id="E66F8307F433E6ADFAB11F1CFA1050E1" box="[1360,1415,375,399]" gridcol="10" gridrow="0" pageId="25" pageNumber="92">
<emphasis id="29FA342BF433194BFAB11F1CFA1050E1" bold="true" box="[1360,1415,375,399]" pageId="25" pageNumber="92">
<collectionCode id="7D9F70FCF433194BFAB11F1CFA1050E1" box="[1360,1415,375,399]" country="Poland" httpUri="http://grbio.org/cool/jvnn-vr63" name="Institut Zoologii" pageId="25" pageNumber="92">IZW</collectionCode>
</emphasis>
</td>
</tr>
<tr id="A5BEEA7BF433E6ADFF0B1FC3FA1050AE" box="[234,1415,424,448]" gridrow="1" pageId="25" pageNumber="92">
<th id="E66F8307F433E6ADFF0B1FC3FEED50AE" box="[234,378,424,448]" gridcol="0" gridrow="1" pageId="25" pageNumber="92">
<collectionCode id="7D9F70FCF433194BFF0B1FC3FE8250AE" box="[234,277,424,448]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="25" pageNumber="92">MB</collectionCode>
.C.30473
</th>
<td id="E66F8307F433E6ADFE571FC3FE7250AE" box="[438,485,424,448]" gridcol="1" gridrow="1" pageId="25" pageNumber="92">38.0</td>
<td id="E66F8307F433E6ADFDEE1FC3FDA950AE" box="[527,574,424,448]" gridcol="2" gridrow="1" pageId="25" pageNumber="92">31.9</td>
<td id="E66F8307F433E6ADFD8C1FC3FD0B50AE" box="[621,668,424,448]" gridcol="3" gridrow="1" pageId="25" pageNumber="92">14.9</td>
<td id="E66F8307F433E6ADFD311FC3FD6850AE" box="[720,767,424,448]" gridcol="4" gridrow="1" pageId="25" pageNumber="92">14.0</td>
<td id="E66F8307F433E6ADFCCC1FC3FCCB50AE" box="[813,860,424,448]" gridcol="5" gridrow="1" pageId="25" pageNumber="92">14.3</td>
<td id="E66F8307F433E6ADFC631FC3FC4250AE" box="[898,981,424,448]" gridcol="6" gridrow="1" pageId="25" pageNumber="92">0.84</td>
<td id="E66F8307F433E6ADFC1B1FC3FBDD50AE" box="[1018,1098,424,448]" gridcol="7" gridrow="1" pageId="25" pageNumber="92">2.13</td>
<td id="E66F8307F433E6ADFB8E1FC3FB2A50AE" box="[1135,1213,424,448]" gridcol="8" gridrow="1" pageId="25" pageNumber="92">0.37</td>
<td id="E66F8307F433E6ADFB041FC3FAB250AE" box="[1253,1317,424,448]" gridcol="9" gridrow="1" pageId="25" pageNumber="92">2.57</td>
<td id="E66F8307F433E6ADFAB11FC3FA1050AE" box="[1360,1415,424,448]" gridcol="10" gridrow="1" pageId="25" pageNumber="92">0.04</td>
</tr>
</table>
</paragraph>
<subSubSection id="5394BBB2F433194BFF021C67FD305460" pageId="25" pageNumber="92" type="description">
<paragraph id="1B31E839F433194BFF021C67FEE35348" blockId="25.[227,372,524,550]" box="[227,372,524,550]" pageId="25" pageNumber="92">
<heading id="40795F55F433194BFF021C67FEE35348" bold="true" box="[227,372,524,550]" fontSize="11" level="3" pageId="25" pageNumber="92" reason="3">
<emphasis id="29FA342BF433194BFF021C67FEE35348" bold="true" box="[227,372,524,550]" pageId="25" pageNumber="92">Description</emphasis>
</heading>
</paragraph>
<paragraph id="1B31E839F433194BFF021C56FE5C52EF" blockId="25.[226,1437,573,897]" pageId="25" pageNumber="92">
<typeStatus id="C435569BF433194BFF021C56FEC75339" box="[227,336,573,599]" pageId="25" pageNumber="92" type="holotype">Holotype</typeStatus>
MB.C.30473 is a desert-polished specimen with a diameter of
<quantity id="DC7645DCF433194BFBB21C56FB3B5339" box="[1107,1196,573,599]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="92" unit="mm" value="38.0">38 mm</quantity>
, showing the conch shape and small areas of ornament (
<figureCitation id="83B5F4BCF433194BFD621C09FD4D5312" box="[643,730,610,636]" captionStart="Fig" captionStartId="25.[227,270,1999,2025]" captionTargetBox="[281,1386,1515,1950]" captionTargetId="figure-338@25.[269,1394,1504,1958]" captionTargetPageId="25" captionText="Fig. 20. Vestinautilus bicristatus sp. nov., holotype MB.C.30473 (Wendt &amp; Kaufmann Coll.) from Oued Temertasset. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902636" httpUri="https://zenodo.org/record/6902636/files/figure.png" pageId="25" pageNumber="92">Fig. 20</figureCitation>
). The conch is broadly cylindrical and subevolute (ww/dm = 0.84; uw/dm =0.37) with a very high coiling rate (WER =2.57). The whorl profile is strongly depressed (ww/wh =2.13) and shows a flattened venter, which possesses two submarginal external grooves at size stages between 17 and
<quantity id="DC7645DCF433194BFE161CB9FDDC5383" box="[503,587,722,749]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="92" unit="mm" value="35.0">35 mm</quantity>
diameter. These weaken considerably thereafter and are only visible as shallow longitudinal depressions at the maximum diameter of the specimen. The ventrolateral shoulder is distinguished by three edges, of which the two inner ones are strengthened by longitudinal ridges. Only small areas of shell ornament are visible; these show very fine growth lines on the venter with a deep external sinus.
</paragraph>
<paragraph id="1B31E839F433194BFF021DC0FBE155EE" blockId="25.[224,1437,939,1152]" pageId="25" pageNumber="92">
Three
<typeStatus id="C435569BF433194BFECF1DC0FE0952AB" box="[302,414,939,965]" pageId="25" pageNumber="92" type="paratype">paratypes</typeStatus>
in fragmentary preservation are available. They are larger specimens with whorl widths between 49 and
<quantity id="DC7645DCF433194BFE7B1DBBFE795285" box="[410,494,976,1003]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="92" unit="mm" value="80.0">80 mm</quantity>
, thus belonged to conchs with
<quantity id="DC7645DCF433194BFCA81DBBFC405285" box="[841,983,976,1003]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="25" pageNumber="92" unit="mm" value="75.0" valueMax="90.0" valueMin="60.0">60 to 90 mm</quantity>
diameter. Two of them (MB.C.30474.2 and MB.C.30474.3;
<figureCitation id="83B5F4BCF433194BFE321D9EFDC9557E" box="[467,606,1013,1040]" captionStart="Fig" captionStartId="26.[227,270,1963,1989]" captionTargetBox="[232,1430,552,1899]" captionTargetId="figure-117@26.[227,1437,550,1911]" captionTargetPageId="26" captionText="Fig. 21. Vestinautilus bicristatus sp. nov. from Oued Temertasset (all Korn et al. 2002 Coll.). A. Paratype MB.C.30474.1. B. Paratype MB.C.30474.2. C. Paratype MB.C.30474.3. Scale bar units =1 mm." figureDoi="http://doi.org/10.5281/zenodo.6885248" httpUri="https://zenodo.org/record/6885248/files/figure.png" pageId="25" pageNumber="92">Fig. 21BC</figureCitation>
) show the imprint of the preceding whorl with sublateral longitudinal grooves in the dorsal region; in
<typeStatus id="C435569BF433194BFDB91A70FD2A555B" box="[600,701,1051,1077]" pageId="25" pageNumber="92" type="paratype">paratype</typeStatus>
MB.C.30474.3 the edges of the ventrolateral shoulder are also still visible. Both have a broadly rounded venter. The largest
<typeStatus id="C435569BF433194BFC531A2AFB805535" box="[946,1047,1089,1115]" pageId="25" pageNumber="92" type="paratype">paratype</typeStatus>
MB.C.30474.1 shows a shallow, broad depression in the middle of the venter at its largest diameter (
<figureCitation id="83B5F4BCF433194BFC1A1A0EFBFF55EE" box="[1019,1128,1125,1152]" captionStart="Fig" captionStartId="26.[227,270,1963,1989]" captionTargetBox="[232,1430,552,1899]" captionTargetId="figure-117@26.[227,1437,550,1911]" captionTargetPageId="26" captionText="Fig. 21. Vestinautilus bicristatus sp. nov. from Oued Temertasset (all Korn et al. 2002 Coll.). A. Paratype MB.C.30474.1. B. Paratype MB.C.30474.2. C. Paratype MB.C.30474.3. Scale bar units =1 mm." figureDoi="http://doi.org/10.5281/zenodo.6885248" httpUri="https://zenodo.org/record/6885248/files/figure.png" pageId="25" pageNumber="92">Fig. 21A</figureCitation>
).
</paragraph>
<paragraph id="1B31E839F433194BFF021AC1FD305460" blockId="25.[227,1436,1193,1294]" pageId="25" pageNumber="92">
<typeStatus id="C435569BF433194BFF021AC1FEDD55AA" box="[227,330,1194,1220]" pageId="25" pageNumber="92" type="paratype">Paratype</typeStatus>
MB.C.30474.3 shows the suture line with a broad and shallow external lobe and a narrow and shallow lateral lobe, which has a position on the umbilical wall (
<figureCitation id="83B5F4BCF433194BFC371AA5FBD55587" box="[982,1090,1230,1257]" captionStart="Fig" captionStartId="26.[227,270,1963,1989]" captionTargetBox="[232,1430,552,1899]" captionTargetId="figure-117@26.[227,1437,550,1911]" captionTargetPageId="26" captionText="Fig. 21. Vestinautilus bicristatus sp. nov. from Oued Temertasset (all Korn et al. 2002 Coll.). A. Paratype MB.C.30474.1. B. Paratype MB.C.30474.2. C. Paratype MB.C.30474.3. Scale bar units =1 mm." figureDoi="http://doi.org/10.5281/zenodo.6885248" httpUri="https://zenodo.org/record/6885248/files/figure.png" pageId="25" pageNumber="92">Fig. 21C</figureCitation>
). The siphuncle has a slightly subcentral position towards the venter.
</paragraph>
</subSubSection>
<subSubSection id="5394BBB2F4331948FF021B53FC0250DD" lastPageId="26" lastPageNumber="93" pageId="25" pageNumber="92" type="discussion">
<paragraph id="1B31E839F433194BFF021B53FEC2543C" blockId="25.[227,341,1336,1362]" box="[227,341,1336,1362]" pageId="25" pageNumber="92">
<heading id="40795F55F433194BFF021B53FEC2543C" bold="true" box="[227,341,1336,1362]" fontSize="11" level="3" pageId="25" pageNumber="92" reason="3">
<emphasis id="29FA342BF433194BFF021B53FEC2543C" bold="true" box="[227,341,1336,1362]" pageId="25" pageNumber="92">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="1B31E839F4331948FF021B03FC0250DD" blockId="25.[227,1436,1384,1448]" lastBlockId="26.[225,1436,303,435]" lastPageId="26" lastPageNumber="93" pageId="25" pageNumber="92">
<taxonomicName id="DC8E93BAF433194BFF021B03FE6854EC" authority="Korn &amp; Bockwinkel, 2022" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" box="[227,511,1384,1410]" class="Cephalopoda" family="Trigonoceratidae" genus="Vestinautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="25" pageNumber="92" phylum="Mollusca" rank="species" species="bicristatus" status="sp. nov.">
<emphasis id="29FA342BF433194BFF021B03FE6854EC" box="[227,511,1384,1410]" italics="true" pageId="25" pageNumber="92">Vestinautilus bicristatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C98950F433194BFDE91B02FDF254ED" box="[520,613,1385,1411]" pageId="25" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the species of the genus that possess a very broad whorl profile (ww/wh&gt; 2.00) and an ornament with only a few spiral ridges. In this respect,
<taxonomicName id="DC8E93BAF433194BFB141BE5FA0254C9" authorityName="Sowerby" authorityYear="1825" box="[1269,1429,1421,1447]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="25" pageNumber="92" phylum="Mollusca" rank="species" species="cariniferus">
<emphasis id="29FA342BF433194BFB141BE5FA0254C9" box="[1269,1429,1421,1447]" italics="true" pageId="25" pageNumber="92">V. cariniferus</emphasis>
</taxonomicName>
,
<taxonomicName id="DC8E93BAF4301948FF021F5BFE395027" authorityName="Foord" authorityYear="1891" box="[227,430,304,329]" class="Cephalopoda" family="Goniatitidae" genus="Coelonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="26" pageNumber="93" phylum="Mollusca" rank="species" species="paucicarinatus">
<emphasis id="29FA342BF4301948FF021F5BFE395027" box="[227,430,304,329]" italics="true" pageId="26" pageNumber="93">V. paucicarinatus</emphasis>
</taxonomicName>
,
<taxonomicName id="DC8E93BAF4301948FE5A1F5BFDBA5027" authorityName="de Koninck" authorityYear="1844" box="[443,557,304,329]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="26" pageNumber="93" phylum="Mollusca" rank="species" species="pinguis">
<emphasis id="29FA342BF4301948FE5A1F5BFDBA5027" box="[443,557,304,329]" italics="true" pageId="26" pageNumber="93">V. pinguis</emphasis>
</taxonomicName>
and
<taxonomicName id="DC8E93BAF4301948FD831F5BFC865027" authorityName="Foord" authorityYear="1900" box="[610,785,303,329]" class="Cephalopoda" family="Trigonoceratidae" genus="Vestinautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="26" pageNumber="93" phylum="Mollusca" rank="species" species="semiplicatus">
<emphasis id="29FA342BF4301948FD831F5BFC865027" box="[610,785,303,329]" italics="true" pageId="26" pageNumber="93">V. semiplicatus</emphasis>
</taxonomicName>
are similar, but these four species have a concave venter at least at times in ontogeny (
<bibRefCitation id="7F1F95C8F4301948FDA61F39FD6F5003" author="Sowerby J. D. C." box="[583,760,338,365]" pageId="26" pageNumber="93" refId="ref18499" refString="Sowerby J. D. C. 1825. The Mineral Conchology of Great Britain; or Coloured Figures and Descriptions of those Remains of Testaceous Animals or Shells, which have been preserved at various Times and Depths in the Earth: Vol. 5. Richard Taylor, London." type="book" year="1825">Sowerby 1825</bibRefCitation>
;
<bibRefCitation id="7F1F95C8F4301948FCE71F39FC405002" author="de Koninck L. G." box="[774,983,338,364]" pageId="26" pageNumber="93" refId="ref17367" refString="de Koninck L. G. 1844. Description des animaux fossiles qui se trouvent dans le terrain carbonifere de la Belgique. H. Dessain, Brussels." type="book" year="1844">de Koninck 1844</bibRefCitation>
;
<bibRefCitation id="7F1F95C8F4301948FC041F39FBE65002" author="Foord A. H." box="[997,1137,338,364]" pageId="26" pageNumber="93" refId="ref17610" refString="Foord A. H. 1891. Catalogue of the Fossil Cephalopoda in the British Museum, part II. Containing the Remainder of the Suborder Nautiloidea, Consisting of the Families Lituitidae, Trochoceratidae, and Nautilidae, with a Supplement. Order of the Trustees, London." type="book" year="1891">Foord 1891</bibRefCitation>
,
<bibRefCitation id="7F1F95C8F4301948FB9E1F39FB2D5002" author="Foord A. H." box="[1151,1210,338,364]" pageId="26" pageNumber="93" pagination="49 - 126" refId="ref17660" refString="Foord A. H. 1900. Monograph on the Carboniferous Cephalopoda of Ireland. Part III. Containing the Families Tainoceratidae, Trigonoceratidae, Triboloceratidae, Rineceratidae, Coloceratidae, and Solenocheilidae (in Part). Monographs of the Palaeontographical Society 54: 49 - 126. https: // doi. org / 10.1080 / 02693445.1900.12035492" type="journal article" year="1900">1900</bibRefCitation>
). The most similar
<taxonomicName id="DC8E93BAF4301948FF041F1DFEEC50E1" authority="Korn &amp; Bockwinkel, 2022" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" box="[229,379,373,399]" class="Cephalopoda" family="Trigonoceratidae" genus="Vestinautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="26" pageNumber="93" phylum="Mollusca" rank="species" species="bicristatus" status="sp. nov.">
<emphasis id="29FA342BF4301948FF041F1DFEEC50E1" box="[229,379,373,399]" italics="true" pageId="26" pageNumber="93">V. bicristatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C98950F4301948FE601F1DFE4C50FE" box="[385,475,374,400]" pageId="26" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
is the species
<taxonomicName id="DC8E93BAF4301948FD611F1DFD7550E1" authorityName="Korn, Miao &amp; Bockwinkel" authorityYear="2022" box="[640,738,373,399]" class="Cephalopoda" family="Trigonoceratidae" genus="Vestinautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="26" pageNumber="93" phylum="Mollusca" rank="species" species="padus">
<emphasis id="29FA342BF4301948FD611F1DFD7550E1" box="[640,738,373,399]" italics="true" pageId="26" pageNumber="93">V. padus</emphasis>
</taxonomicName>
, which shows the same conch dimensions, but differs in the lack of the submarginal ventral grooves (
<bibRefCitation id="7F1F95C8F4301948FD221FF3FC1150DC" author="Korn D. &amp; Miao L. &amp; Bockwinkel J." box="[707,902,408,434]" pageId="26" pageNumber="93" pagination="104 - 129" refId="ref18166" refString="Korn D., Miao L. &amp; Bockwinkel J. 2022. The nautiloids from the Dalle a Merocanites of Timimoun (Tournaisian - Visean boundary, Algeria). European Journal of Taxonomy 789: 104 - 129. https: // doi. org / 10.5852 / ejt. 2022.789.1635" type="journal article" year="2022">
Korn
<emphasis id="29FA342BF4301948FCE91FF2FCD450DC" box="[776,835,408,434]" italics="true" pageId="26" pageNumber="93">et al.</emphasis>
2022
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>