Early Carboniferous nautiloids from the Central Sahara, southern Algeria Author Korn, Dieter 286CA4F3-7EBC-4AEF-A66A-B2508D001367 Museum für Naturkunde, Leibniz Institut for Research on Evolution and Biodiversity, Invalidenstrasse 43, 10115 Berlin, Germany. dieter.korn@mfn.berlin Author Bockwinkel, Jürgen F7FE7EEA-B678-4FEE-879C-8C429F66BF3A Dechant-Fein-Strasse 22, 51375 Leverkusen, Germany. jbockwinkel@t-online.de text European Journal of Taxonomy 2022 2022-07-20 831 67 108 http://dx.doi.org/10.5852/ejt.2022.831.1871 journal article 105313 10.5852/ejt.2022.831.1871 e697ba5c-5a96-4a28-985c-748890333d6c 2118-9773 6885198 EAA2EAE2-DA8B-4516-B332-CB8423B8EEAA Vestinautilus bicristatus sp. nov. urn:lsid:zoobank.org:act: 69CABBA5-6CDA-4264-B156-0FB2321AEDD5 Figs 20–21 , Table 11 Diagnosis Species of Vestinautilus with strongly depressed, rounded-trapezoidal whorl profile (ww/wh ~ 2.10), venter up to 30 mm conch diameter broadly arched with two shallow submarginal grooves, which disappear in the adult stage. Ventrolateral shoulder defined by a subangular margin; one sharp ridge is located on the umbilical wall near the ventrolateral shoulder. Whorls weakly embracing, coiling very high (WER ~2.55). Ornament with delicate growth lines. Etymology From the Latin ‘ crista ’, meaning ‘crest’ and referring to the two lateral longitudinal ridges. Type material Holotype ALGERIAMouydir , south of Oued Temertasset (locality MOU-C1) ; Argiles de Teguentour ( Helicocyclus-Ouaoufilalites Assemblage; early late Tournaisian ); Wendt et al . Coll.; illustrated in Fig. 20 ; MB.C.30473 . Paratypes ALGERIA3 specimens ; Mouydir , south of Oued Temertasset (locality MOU-C1) ; Argiles de Teguentour ( Helicocyclus-Ouaoufilalites Assemblage; early late Tournaisian ); Korn et al . 2002 Coll.; illustrated in Fig. 21 ; MB.C.30474.1MB.C.30474.3 . Table 11. Conch dimensions (in mm) and ratios of Vestinautilus bicristatus sp. nov.
Specimen dm ww wh uw ah ww/dm ww/wh uw/dm WER IZW
MB .C.30473 38.0 31.9 14.9 14.0 14.3 0.84 2.13 0.37 2.57 0.04
Description Holotype MB.C.30473 is a desert-polished specimen with a diameter of 38 mm , showing the conch shape and small areas of ornament ( Fig. 20 ). The conch is broadly cylindrical and subevolute (ww/dm = 0.84; uw/dm =0.37) with a very high coiling rate (WER =2.57). The whorl profile is strongly depressed (ww/wh =2.13) and shows a flattened venter, which possesses two submarginal external grooves at size stages between 17 and 35 mm diameter. These weaken considerably thereafter and are only visible as shallow longitudinal depressions at the maximum diameter of the specimen. The ventrolateral shoulder is distinguished by three edges, of which the two inner ones are strengthened by longitudinal ridges. Only small areas of shell ornament are visible; these show very fine growth lines on the venter with a deep external sinus. Three paratypes in fragmentary preservation are available. They are larger specimens with whorl widths between 49 and 80 mm , thus belonged to conchs with 60 to 90 mm diameter. Two of them (MB.C.30474.2 and MB.C.30474.3; Fig. 21B–C ) show the imprint of the preceding whorl with sublateral longitudinal grooves in the dorsal region; in paratype MB.C.30474.3 the edges of the ventrolateral shoulder are also still visible. Both have a broadly rounded venter. The largest paratype MB.C.30474.1 shows a shallow, broad depression in the middle of the venter at its largest diameter ( Fig. 21A ). Paratype MB.C.30474.3 shows the suture line with a broad and shallow external lobe and a narrow and shallow lateral lobe, which has a position on the umbilical wall ( Fig. 21C ). The siphuncle has a slightly subcentral position towards the venter. Remarks Vestinautilus bicristatus sp. nov. belongs to the species of the genus that possess a very broad whorl profile (ww/wh> 2.00) and an ornament with only a few spiral ridges. In this respect, V. cariniferus , V. paucicarinatus , V. pinguis and V. semiplicatus are similar, but these four species have a concave venter at least at times in ontogeny ( Sowerby 1825 ; de Koninck 1844 ; Foord 1891 , 1900 ). The most similar V. bicristatus sp. nov. is the species V. padus , which shows the same conch dimensions, but differs in the lack of the submarginal ventral grooves ( Korn et al. 2022 ).