treatments-xml/data/00/3B/BA/003BBAE10183519EB9CCCD9CF06CCBBE.xml
2024-06-21 12:18:41 +02:00

1193 lines
57 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zse.96.52860" ID-GBIF-Dataset="e27c18de-98bd-41e1-9354-11d49edd5d33" ID-GBIF-Taxon="167407637" ID-Pensoft-Pub="1860-0743-2-577" ID-Pensoft-UUID="D4882E78D81B5FA394A6A156886835E4" ModsDocID="1860-0743-2-577" checkinTime="1599019521532" checkinUser="pensoft" docAuthor="Vinarski, Maxim V., Aksenova, Olga V. &amp; Bolotov, Ivan N." docDate="2020" docId="003BBAE10183519EB9CCCD9CF06CCBBE" docLanguage="en" docName="ZoosystEvol 96(2): 577-608" docOrigin="Zoosystematics and Evolution 96 (2)" docSource="http://dx.doi.org/10.3897/zse.96.52860" docTitle="Radix (Radix) alticola" docType="treatment" docVersion="4" id="D4882E78D81B5FA394A6A156886835E4" lastPageNumber="577" masterDocId="D4882E78D81B5FA394A6A156886835E4" masterDocTitle="Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae)" masterLastPageNumber="608" masterPageNumber="577" pageId="0" pageNumber="577" updateTime="1634717030393" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vinarski, Maxim V.</mods:namePart>
<mods:affiliation>Laboratory of Macroecology and Biogeography of Invertebrates, Saint-Petersburg State University, 7 / 9 Universitetskaya Emb., 199034, Saint-Petersburg, Russia &amp; Omsk State University, 28 Adrianova Str., 644077, Omsk, Russia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7644-4164</mods:nameIdentifier>
<mods:nameIdentifier type="email">radix.vinarski@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aksenova, Olga V.</mods:namePart>
<mods:affiliation>Laboratory of Macroecology and Biogeography of Invertebrates, Saint-Petersburg State University, 7 / 9 Universitetskaya Emb., 199034, Saint-Petersburg, Russia &amp; N. Laverov Federal Center for Integrated Arctic Research, Ural Branch of the Russian Academy of Sciences, 23 Severnaya Dvina Emb., 163000, Arkhangelsk, Russia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-0817-7105</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bolotov, Ivan N.</mods:namePart>
<mods:affiliation>N. Laverov Federal Center for Integrated Arctic Research, Ural Branch of the Russian Academy of Sciences, 23 Severnaya Dvina Emb., 163000, Arkhangelsk, Russia &amp; Northern (Arctic) Federal University, 17 Severnaya Dvina Emb., 163002, Arkhangelsk, Russia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3878-4192</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>96</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>577</mods:start>
<mods:end>608</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zse.96.52860</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.52860</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-2-577</mods:identifier>
<mods:identifier type="Pensoft-UUID">D4882E78D81B5FA394A6A156886835E4</mods:identifier>
<mods:identifier type="ZooBank">B8CF4E84-1FEE-46F3-B275-BAFA6824902B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="167407637" LSID="urn:lsid:plazi:treatment:003BBAE10183519EB9CCCD9CF06CCBBE" httpUri="http://treatment.plazi.org/id/003BBAE10183519EB9CCCD9CF06CCBBE" lastPageNumber="577" pageId="0" pageNumber="577">
<subSubSection pageId="0" pageNumber="577" type="nomenclature">
<paragraph pageId="0" pageNumber="577">
2.
<taxonomicName LSID="003BBAE1-0183-519E-B9CC-CD9CF06CCBBE" authority="(Izzatullaev, Kruglov &amp; Starobogatov, 1983)" baseAuthorityName="Izzatullaev, Kruglov &amp; Starobogatov" baseAuthorityYear="1983" class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radix (Radix) alticola" order="Hygrophila" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="alticola" subGenus="Radix">Radix (Radix) alticola (Izzatullaev, Kruglov &amp; Starobogatov, 1983)</taxonomicName>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of species in the genus Radix. A. Radix auricularia (28.08.2013, Kazakhstan, Karaganda Region, Suresai River; LMBI); B. R. alticola (20.06.2012, Tajikistan, a hot spring near Djelandy village; LMBI); C. R. alticola (01.07.2016, Tajikistan, a warm brook near Dzhaushangoz village; LMBI); D. R. brevicauda (Kashmir, a syntype; NHMUK); E. R. brevicauda (05.08.1948, Tajikistan, Pamir Mts., Shaimak village, in a warm spring; ZISP); F. R. euphratica (26.06.2016, Tajikistan, Dushanbe, a fountain near the President's palace; LMBI); G. R. euphratica (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI); H. R. euphratica (without date, Iraq, Bagdad; NHMUK); I. R. makhrovi, the holotype (China, Tibet, a roadside ditch west of the Lhasa River mouth; ZISP); J. R. plicatula, a probable syntype (China, Chusan, NHMUK); K. R. plicatula (26.07.2017, China, Uyghuria, Bagrash-kol' Lake; LMBI); L. R. plicatula (26.11.2014, China, Bejing, an artificial pond in the former Emperor's summer palace; LMBI). Scale bars: 2 mm (B-C, F-L), 5 mm (A, D-E, K)." figureDoi="10.3897/zse.96.52860.figure3" httpUri="https://binary.pensoft.net/fig/448267" pageId="0" pageNumber="577">Figs 3B, C</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory apparatuses of the species of the genus Radix. A. Radix auricularia (19.07.2007, Russia, Tyumen' Region, Vylposl channel near Labytnangi Town); B. R. alticola (01.07.2016, Tajikistan, a warm brook near Dzhaushangoz village); C. R. euphratica (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village); D. R. makhrovi, a paratype (after Aksenova et al. 2018 a, modified). E. R. plicatula (26.11.2014, China, Bejing, an artificial pond in the former Emperor's summer palace). F. R. rubiginosa (Malaysia, Kuala-Lumpur); G. R. rufescens (24.11.2016, Myanmar, Yetho River, near the dam). H. R. natalensis (05.08.2018, Uganda, crater lake Kyamwiga). Scale bars: 2 mm." figureDoi="10.3897/zse.96.52860.figure4" httpUri="https://binary.pensoft.net/fig/448268" pageId="0" pageNumber="577">; 4B</figureCitation>
;
<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characterisation of shells and copulative apparatuses of several species in the genus Radix." pageId="0" pageNumber="577">Table 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="577" type="reference_group">
<paragraph pageId="0" pageNumber="577">
<taxonomicName authorityName="Izzatullaev, Kruglov &amp; Starobogatov" authorityYear="1983" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lymnaea (Radix) alticola" order="Hygrophila" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="alticola" subGenus="Radix">Lymnaea (Radix) alticola</taxonomicName>
<bibRefCitation author="Izzatullaev, ZI" journalOrPublisher="Izvestiya Akademii Nauk Tajikskoy SSR, otdelenie biologicheskikh nauk" pageId="0" pageNumber="577" pagination="53 - 57" refId="B52" refString="Izzatullaev, ZI, Kruglov, ND, Starobogatov, YI, 1983. New and ill-known species of mollusks of the subgenus Radix of the genus Lymnaea of the USSR fauna from the Central Asia (Gastropoda, Pulmonata). Izvestiya Akademii Nauk Tajikskoy SSR, otdelenie biologicheskikh nauk 4: 53 - 57" title="New and ill-known species of mollusks of the subgenus Radix of the genus Lymnaea of the USSR fauna from the Central Asia (Gastropoda, Pulmonata)." volume="4" year="1983">Izzatullaev et al. 1983</bibRefCitation>
: 53, figs 1, 2;
<bibRefCitation author="Kruglov, ND" journalOrPublisher="Ruthenica: The Russian Malacological Journal" pageId="0" pageNumber="577" pagination="65 - 92" refId="B69" refString="Kruglov, ND, Starobogatov, YI, 1993a. Annotated and illustrated catalogue of species of the family Lymnaeidae (Gastropoda Pulmonata Lymnaeiformes) of Palaearctic and adjacent river drainage areas. Part I. Ruthenica: The Russian Malacological Journal 3: 65 - 92" title="Annotated and illustrated catalogue of species of the family Lymnaeidae (Gastropoda Pulmonata Lymnaeiformes) of Palaearctic and adjacent river drainage areas. Part I." volume="3" year="1993 a">Kruglov and Starobogatov 1993a</bibRefCitation>
: 85, fig. 11C;
<bibRefCitation author="Kruglov, ND" journalOrPublisher="SGPU Publishing, Smolensk" pageId="0" pageNumber="577" publicationUrl="http://herba.msu.ru/shipunov/school/books/kruglov2005_lymnaeidae.djvu" refId="B62" refString="Kruglov, ND, 2005. Mollusks of the Family Lymnaeidae (GastropodaPulmonata) in Europe and Northern Asia. SGPU Publishing, Smolensk, http://herba.msu.ru/shipunov/school/books/kruglov2005_lymnaeidae.djvu" title="Mollusks of the Family Lymnaeidae (GastropodaPulmonata) in Europe and Northern Asia." url="http://herba.msu.ru/shipunov/school/books/kruglov2005_lymnaeidae.djvu" year="2005">Kruglov 2005</bibRefCitation>
: 261, figs 146(1)-148;
<bibRefCitation author="Sitnikova, TY" journalOrPublisher="Zoologicheskie Issledovania" pageId="0" pageNumber="577" pagination="7 - 37" publicationUrl="http://zmmu.msu.ru/files/books/zool_issl_16.pdf" refId="B116" refString="Sitnikova, TY, Sysoev, AV, Prozorova, LA, 2014. Types of freshwater gastropods described by Ya.I. Starobogatov, with additional data on the species: family Lymnaeidae. Zoologicheskie Issledovania 16: 7 - 37, http://zmmu.msu.ru/files/books/zool_issl_16.pdf" title="Types of freshwater gastropods described by Ya. I. Starobogatov, with additional data on the species: family Lymnaeidae." url="http://zmmu.msu.ru/files/books/zool_issl_16.pdf" volume="16" year="2014">Sitnikova et al. 2014</bibRefCitation>
: 25, fig. 8D.
</paragraph>
<paragraph pageId="0" pageNumber="577">
<taxonomicName baseAuthorityName="Izzatullaev, Kruglov &amp; Starobogatov" baseAuthorityYear="1983" class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radix (Radix) alticola" order="Hygrophila" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="alticola" subGenus="Radix">Radix (Radix) alticola</taxonomicName>
-
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="577" refId="B142" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
: 320.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="577" type="tl">
<paragraph pageId="0" pageNumber="577">TL.</paragraph>
<paragraph pageId="0" pageNumber="577">
Tajikistan, a hot spring near the
<normalizedToken originalValue="Yashilkul">Yashilkul'</normalizedToken>
Lake (approximately
<geoCoordinate degrees="37" direction="north" minutes="47" orientation="latitude" precision="15" seconds="00" value="37.783333">37°47'00&quot;N</geoCoordinate>
,
<geoCoordinate degrees="72" direction="east" minutes="51" orientation="longitude" precision="15" seconds="00" value="72.85">72°51'00&quot;E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="577" type="types">
<paragraph pageId="0" pageNumber="577">Types.</paragraph>
<paragraph pageId="0" pageNumber="577">
ZISP (
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="577" refId="B142" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
). The holotype is illustrated by
<bibRefCitation author="Sitnikova, TY" journalOrPublisher="Zoologicheskie Issledovania" pageId="0" pageNumber="577" pagination="7 - 37" publicationUrl="http://zmmu.msu.ru/files/books/zool_issl_16.pdf" refId="B116" refString="Sitnikova, TY, Sysoev, AV, Prozorova, LA, 2014. Types of freshwater gastropods described by Ya.I. Starobogatov, with additional data on the species: family Lymnaeidae. Zoologicheskie Issledovania 16: 7 - 37, http://zmmu.msu.ru/files/books/zool_issl_16.pdf" title="Types of freshwater gastropods described by Ya. I. Starobogatov, with additional data on the species: family Lymnaeidae." url="http://zmmu.msu.ru/files/books/zool_issl_16.pdf" volume="16" year="2014">Sitnikova et al. (2014)</bibRefCitation>
. We studied both the holotype and the paratypes of
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="577">
According to our data (
<bibRefCitation DOI="https://doi.org/10.1038/s41598-018-29451-1" author="Aksenova, OV" journalOrPublisher="Scientific Reports" pageId="0" pageNumber="577" pagination="1 - 17" refId="B1" refString="Aksenova, OV, Bolotov, IN, Gofarov, MY, Kondakov, AV, Vinarski, MV, Bespalaya, YV, Kolosova, YS, Palatov, DM, Sokolova, SE, Spitsyn, VM, Tomilova, AA, Travina, OV, Vikhrev, IV, 2018a. Species richness, molecular taxonomy and biogeography of the radicine pond snails (Gastropoda: Lymnaeidae) in the Old World. Scientific Reports 8 (11199): 1 - 17, DOI: https://doi.org/10.1038/s41598-018-29451-1" title="Species richness, molecular taxonomy and biogeography of the radicine pond snails (Gastropoda: Lymnaeidae) in the Old World." url="https://doi.org/10.1038/s41598-018-29451-1" volume="8" year="2018 a">Aksenova et al. 2018a</bibRefCitation>
),
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
is endemic to the High Asia mountains; found in Tajikistan (from several sites, including the type locality) and Nepal. In mountain Tajikistan (Pamir), this species inhabits hot springs and satellite streams with relatively warm water.
</paragraph>
<paragraph pageId="0" pageNumber="577">
The shell shape of
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName genus="R." lsidName="R. auricularia" pageId="0" pageNumber="577" rank="species" species="auricularia">
<emphasis italics="true" pageId="0" pageNumber="577">R. auricularia</emphasis>
</taxonomicName>
, but the former species is of much smaller size; its shell height does not exceed 16 mm (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characterisation of shells and copulative apparatuses of several species in the genus Radix." httpUri="http://table.plazi.org/id/72A87B6AE3E005FDC64F6F3D96361C95" pageId="0" pageNumber="577" tableUuid="72A87B6AE3E005FDC64F6F3D96361C95">1</tableCitation>
), whereas the shells of
<taxonomicName genus="R." lsidName="R. auricularia" pageId="0" pageNumber="577" rank="species" species="auricularia">
<emphasis italics="true" pageId="0" pageNumber="577">R. auricularia</emphasis>
</taxonomicName>
may reach 35-40 mm in height. The proportions of the copulatory organ of
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
are very similar to
<taxonomicName genus="R." lsidName="R. auricularia" pageId="0" pageNumber="577" rank="species" species="auricularia">
<emphasis italics="true" pageId="0" pageNumber="577">R. auricularia</emphasis>
</taxonomicName>
(compare Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of species in the genus Radix. A. Radix auricularia (28.08.2013, Kazakhstan, Karaganda Region, Suresai River; LMBI); B. R. alticola (20.06.2012, Tajikistan, a hot spring near Djelandy village; LMBI); C. R. alticola (01.07.2016, Tajikistan, a warm brook near Dzhaushangoz village; LMBI); D. R. brevicauda (Kashmir, a syntype; NHMUK); E. R. brevicauda (05.08.1948, Tajikistan, Pamir Mts., Shaimak village, in a warm spring; ZISP); F. R. euphratica (26.06.2016, Tajikistan, Dushanbe, a fountain near the President's palace; LMBI); G. R. euphratica (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI); H. R. euphratica (without date, Iraq, Bagdad; NHMUK); I. R. makhrovi, the holotype (China, Tibet, a roadside ditch west of the Lhasa River mouth; ZISP); J. R. plicatula, a probable syntype (China, Chusan, NHMUK); K. R. plicatula (26.07.2017, China, Uyghuria, Bagrash-kol' Lake; LMBI); L. R. plicatula (26.11.2014, China, Bejing, an artificial pond in the former Emperor's summer palace; LMBI). Scale bars: 2 mm (B-C, F-L), 5 mm (A, D-E, K)." figureDoi="10.3897/zse.96.52860.figure3" httpUri="https://binary.pensoft.net/fig/448267" pageId="0" pageNumber="577">3A</figureCitation>
and
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of species in the genus Radix. A. Radix auricularia (28.08.2013, Kazakhstan, Karaganda Region, Suresai River; LMBI); B. R. alticola (20.06.2012, Tajikistan, a hot spring near Djelandy village; LMBI); C. R. alticola (01.07.2016, Tajikistan, a warm brook near Dzhaushangoz village; LMBI); D. R. brevicauda (Kashmir, a syntype; NHMUK); E. R. brevicauda (05.08.1948, Tajikistan, Pamir Mts., Shaimak village, in a warm spring; ZISP); F. R. euphratica (26.06.2016, Tajikistan, Dushanbe, a fountain near the President's palace; LMBI); G. R. euphratica (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI); H. R. euphratica (without date, Iraq, Bagdad; NHMUK); I. R. makhrovi, the holotype (China, Tibet, a roadside ditch west of the Lhasa River mouth; ZISP); J. R. plicatula, a probable syntype (China, Chusan, NHMUK); K. R. plicatula (26.07.2017, China, Uyghuria, Bagrash-kol' Lake; LMBI); L. R. plicatula (26.11.2014, China, Bejing, an artificial pond in the former Emperor's summer palace; LMBI). Scale bars: 2 mm (B-C, F-L), 5 mm (A, D-E, K)." figureDoi="10.3897/zse.96.52860.figure3" httpUri="https://binary.pensoft.net/fig/448267" pageId="0" pageNumber="577">3B</figureCitation>
).
</paragraph>
<caption ID-Table-UUID="72A87B6AE3E005FDC64F6F3D96361C95" httpUri="http://table.plazi.org/id/72A87B6AE3E005FDC64F6F3D96361C95" pageId="0" pageNumber="577" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="577">
<emphasis bold="true" pageId="0" pageNumber="577">Table 1.</emphasis>
Morphological characterisation of shells and copulative apparatuses of several species in the genus
<taxonomicName class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radix" pageId="0" pageNumber="577" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="577">Radix</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="577">
<table pageId="0" pageNumber="577">
<tr pageId="0" pageNumber="577">
<th colspan="1" pageId="0" pageNumber="577" rowspan="3">Character / index</th>
<th colspan="10" pageId="0" pageNumber="577" rowspan="1">Species / locality</th>
</tr>
<tr pageId="0" pageNumber="577">
<th colspan="2" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
</th>
<th colspan="2" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. euphratica" pageId="0" pageNumber="577" rank="species" species="euphratica">
<emphasis italics="true" pageId="0" pageNumber="577">R. euphratica</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. brevicauda" pageId="0" pageNumber="577" rank="species" species="brevicauda">
<emphasis italics="true" pageId="0" pageNumber="577">R. brevicauda</emphasis>
</taxonomicName>
</th>
<th colspan="2" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. plicatula" pageId="0" pageNumber="577" rank="species" species="plicatula">
<emphasis italics="true" pageId="0" pageNumber="577">R. plicatula</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. natalensis" pageId="0" pageNumber="577" rank="species" species="natalensis">
<emphasis italics="true" pageId="0" pageNumber="577">R. natalensis</emphasis>
</taxonomicName>
</th>
<th colspan="2" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. rufescens" pageId="0" pageNumber="577" rank="species" species="rufescens">
<emphasis italics="true" pageId="0" pageNumber="577">R. rufescens</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="577">
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Tajikistan, Zor-Kul Lake (paratypes)</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Tajikistan, a warm brook near Dzhaushangoz village*</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
Tajikistan, Dushanbe, a fountain near
<normalizedToken originalValue="Presidents">President's</normalizedToken>
palace*
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Tajikistan, a roadside ditch near Kurban-Shakhid village*</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Tajikistan, a warm spring near Shaimak village</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
China, Bejing, a pond near the former
<normalizedToken originalValue="Emperors">Emperor's</normalizedToken>
palace*
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
China, Uyghuria,
<normalizedToken originalValue="Bagrashkol">Bagrashkol'</normalizedToken>
Lake*
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Uganda, the Kyamwiga crater lake*, **</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">India, &quot;Khipru, Sind&quot;</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Myanmar, Yetho River near the dam*</th>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">Repository</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">ZISP</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">ZISP</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">NHMUK</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">Number of specimens measured (dissected)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">45 (0)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">44 (16)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">4(3)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">21(15)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">21(0)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">29(11)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">16(9)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">26(20)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">40(0)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12(6)</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Whorls number</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.00-3.75</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.00-3.75</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.87-4.25</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.25-4.12</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.25-4.00</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.75-4.00</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.50-4.12</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.25-3.75</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.75-4.75</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.87-4.50</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.40
<normalizedToken originalValue="±">+/-</normalizedToken>
0.18
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.30
<normalizedToken originalValue="±">+/-</normalizedToken>
0.17
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.06
<normalizedToken originalValue="±">+/-</normalizedToken>
0.16
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.71
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.48
<normalizedToken originalValue="±">+/-</normalizedToken>
0.19
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.08
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.76
<normalizedToken originalValue="±">+/-</normalizedToken>
0.23
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.51
<normalizedToken originalValue="±">+/-</normalizedToken>
0.15
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.17
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.22
<normalizedToken originalValue="±">+/-</normalizedToken>
0.22
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Shell height, mm (SH)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.0-13.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.4-15.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.6-15.1</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.5-20.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.8-15.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">13.6-20.7</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.9-22.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">13.6-18.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">37.9-43.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">17.5-28.9</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
13.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.8
<normalizedToken originalValue="±">+/-</normalizedToken>
2.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
12.1
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
16.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
15.8
<normalizedToken originalValue="±">+/-</normalizedToken>
3.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
16.2
<normalizedToken originalValue="±">+/-</normalizedToken>
1.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
39.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
23.2
<normalizedToken originalValue="±">+/-</normalizedToken>
3.8
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Shell width, mm (SW)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.4-10.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.8-13.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.4-9.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.6-13.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.4-13.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.9-14.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.2-14.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.0-13.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">17.8-20.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.1-13.6</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
8.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
8.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.3
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.1
<normalizedToken originalValue="±">+/-</normalizedToken>
2.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
18.8
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.0
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Spire height, mm (SpH)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">1.4-2.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.0-4.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.8-4.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.7-5.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.2-4.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.4-6.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.7-7.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.8-4.7</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">10.6-14.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">4.5-10.8</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
2.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
2.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
2.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
5.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
12.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.8
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Body whorl height, mm (BWH)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.4-12.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.7-14.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">10.1-12.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">11.0-17.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.9-13.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">11.7-18.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">11.5-20.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.3-17.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">32.8-37.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">15.6-25.2</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
13.1
<normalizedToken originalValue="±">+/-</normalizedToken>
1.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.1
<normalizedToken originalValue="±">+/-</normalizedToken>
2.8
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.8
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
34.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
20.1
<normalizedToken originalValue="±">+/-</normalizedToken>
3.3
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Aperture height, mm (AH)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.6-11.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.7-12.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.1-12.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.4-15.1</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.7-11.7</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.8-16.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.4-16.1</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">10.2-14.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">27.0-31.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.7-19.1</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.6
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.6
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.8
<normalizedToken originalValue="±">+/-</normalizedToken>
1.7
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.6
<normalizedToken originalValue="±">+/-</normalizedToken>
2.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
12.6
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
28.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
15.5
<normalizedToken originalValue="±">+/-</normalizedToken>
2.2
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Aperture width, mm (AW)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.0-8.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.2-10.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.8-7.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.8-11.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.5-8.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.3-12.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.1-10.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.0-9.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.3-16.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.5-11.3</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
6.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.7
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
6.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
6.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.0
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
8.2
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.7
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
8.6
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Praeputium length, mm (PP)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.2-5.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.8-3.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.2-4.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.4-6.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.0-5.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.1-4.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.8-4.2</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Penis sheath length, mm (PS)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.2-5.1</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.9-3.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.2-3.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.6-9.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.6-7.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.0-5.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.2-3.6</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.1
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
5.8
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
5.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
2.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">SW/SH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.65-0.81</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.72-0.87</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.62-0.67</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.61-0.73</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.79-0.94</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.53-0.76</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.55-0.72</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.57-0.71</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.44-0.50</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.41-0.52</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.74
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.80
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.64
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.66
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.85
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.63
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.64
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.65
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.47
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.48
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">SpH/SH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.13-0.24</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.18-0.27</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.28-0.32</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.20-0.30</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.18-0.28</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.20-0.42</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.20-0.33</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.19-0.27</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.27-0.38</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.26-0.37</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.19
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.22
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.30
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.23
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.31
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.28
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.23
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.32
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.33
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">BWH/SH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.87-0.93</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.88-0.98</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.80-0.86</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.83-0.94</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.87-0.92</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.83-0.95</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.86-0.92</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.86-0.93</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.84-0.89</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.80-0.90</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.90
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.91
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.84
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.89
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.90
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.87
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.89
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.91
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.87
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.86
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">AH/SH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.65-0.81</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.75-0.85</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.70-0.74</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.70-0.86</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.72-0.86</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.59-0.86</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.69-0.81</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.74-0.82</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.69-0.77</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.63-0.75</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.74
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.81
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.73
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.77
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.80
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.71
<normalizedToken originalValue="±">+/-</normalizedToken>
0.06
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.74
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.78
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.73
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.67
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">AW/AH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.59-0.83</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.66-0.91</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.63-0.67</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.57-0.78</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.67-0.81</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.63-0.78</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.56-0.73</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.57-0.67</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.45-0.55</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.50-0.61</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.69
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.72
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.65
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.65
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.73
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.70
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.64
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.62
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.50
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.55
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">ICA</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.81-1.18</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.93-1.16</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">1.10-1.69</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.54-0.96</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.64-0.90</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.84-1.13</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">1.09-1.27</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.97
<normalizedToken originalValue="±">+/-</normalizedToken>
0.11
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
1.01
<normalizedToken originalValue="±">+/-</normalizedToken>
0.12
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
1.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.18
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.76
<normalizedToken originalValue="±">+/-</normalizedToken>
0.14
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.79
<normalizedToken originalValue="±">+/-</normalizedToken>
0.08
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.96
<normalizedToken originalValue="±">+/-</normalizedToken>
0.07
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
1.19
<normalizedToken originalValue="±">+/-</normalizedToken>
0.06
</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="577">
<tableNote pageId="0" pageNumber="577">
*Sample studied genetically. ** The short-spired morph of
<taxonomicName genus="R." lsidName="R. natalensis" pageId="0" pageNumber="577" rank="species" species="natalensis">
<emphasis italics="true" pageId="0" pageNumber="577">R. natalensis</emphasis>
</taxonomicName>
.
</tableNote>
</paragraph>
</subSubSection>
</treatment>
</document>