treatments-xml/data/00/3B/BA/003BBAE10183519EB9CCCD9CF06CCBBE.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.96.52860" ID-GBIF-Dataset="e27c18de-98bd-41e1-9354-11d49edd5d33" ID-GBIF-Taxon="167407637" ID-Pensoft-Pub="1860-0743-2-577" ID-Pensoft-UUID="D4882E78D81B5FA394A6A156886835E4" ModsDocID="1860-0743-2-577" checkinTime="1599019521532" checkinUser="pensoft" docAuthor="Vinarski, Maxim V., Aksenova, Olga V. &amp; Bolotov, Ivan N." docDate="2020" docId="003BBAE10183519EB9CCCD9CF06CCBBE" docLanguage="en" docName="ZoosystEvol 96(2): 577-608" docOrigin="Zoosystematics and Evolution 96 (2)" docSource="http://dx.doi.org/10.3897/zse.96.52860" docTitle="Radix (Radix) alticola" docType="treatment" docVersion="4" id="D4882E78D81B5FA394A6A156886835E4" lastPageNumber="577" masterDocId="D4882E78D81B5FA394A6A156886835E4" masterDocTitle="Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae)" masterLastPageNumber="608" masterPageNumber="577" pageId="0" pageNumber="577" updateTime="1634717030393" updateUser="ExternalLinkService">
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<mods:title>Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vinarski, Maxim V.</mods:namePart>
<mods:affiliation>Laboratory of Macroecology and Biogeography of Invertebrates, Saint-Petersburg State University, 7 / 9 Universitetskaya Emb., 199034, Saint-Petersburg, Russia &amp; Omsk State University, 28 Adrianova Str., 644077, Omsk, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">radix.vinarski@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Aksenova, Olga V.</mods:namePart>
<mods:affiliation>Laboratory of Macroecology and Biogeography of Invertebrates, Saint-Petersburg State University, 7 / 9 Universitetskaya Emb., 199034, Saint-Petersburg, Russia &amp; N. Laverov Federal Center for Integrated Arctic Research, Ural Branch of the Russian Academy of Sciences, 23 Severnaya Dvina Emb., 163000, Arkhangelsk, Russia</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bolotov, Ivan N.</mods:namePart>
<mods:affiliation>N. Laverov Federal Center for Integrated Arctic Research, Ural Branch of the Russian Academy of Sciences, 23 Severnaya Dvina Emb., 163000, Arkhangelsk, Russia &amp; Northern (Arctic) Federal University, 17 Severnaya Dvina Emb., 163002, Arkhangelsk, Russia</mods:affiliation>
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<mods:date>2020</mods:date>
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<mods:number>96</mods:number>
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<treatment ID-GBIF-Taxon="167407637" LSID="urn:lsid:plazi:treatment:003BBAE10183519EB9CCCD9CF06CCBBE" httpUri="http://treatment.plazi.org/id/003BBAE10183519EB9CCCD9CF06CCBBE" lastPageNumber="577" pageId="0" pageNumber="577">
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2. 
<taxonomicName LSID="003BBAE1-0183-519E-B9CC-CD9CF06CCBBE" authority="(Izzatullaev, Kruglov &amp; Starobogatov, 1983)" baseAuthorityName="Izzatullaev, Kruglov &amp; Starobogatov" baseAuthorityYear="1983" class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radix (Radix) alticola" order="Hygrophila" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="alticola" subGenus="Radix">Radix (Radix) alticola (Izzatullaev, Kruglov &amp; Starobogatov, 1983)</taxonomicName>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of species in the genus Radix. A. Radix auricularia (28.08.2013, Kazakhstan, Karaganda Region, Suresai River; LMBI); B. R. alticola (20.06.2012, Tajikistan, a hot spring near Djelandy village; LMBI); C. R. alticola (01.07.2016, Tajikistan, a warm brook near Dzhaushangoz village; LMBI); D. R. brevicauda (Kashmir, a syntype; NHMUK); E. R. brevicauda (05.08.1948, Tajikistan, Pamir Mts., Shaimak village, in a warm spring; ZISP); F. R. euphratica (26.06.2016, Tajikistan, Dushanbe, a fountain near the President's palace; LMBI); G. R. euphratica (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI); H. R. euphratica (without date, Iraq, Bagdad; NHMUK); I. R. makhrovi, the holotype (China, Tibet, a roadside ditch west of the Lhasa River mouth; ZISP); J. R. plicatula, a probable syntype (China, Chusan, NHMUK); K. R. plicatula (26.07.2017, China, Uyghuria, Bagrash-kol' Lake; LMBI); L. R. plicatula (26.11.2014, China, Bejing, an artificial pond in the former Emperor's summer palace; LMBI). Scale bars: 2 mm (B-C, F-L), 5 mm (A, D-E, K)." figureDoi="10.3897/zse.96.52860.figure3" httpUri="https://binary.pensoft.net/fig/448267" pageId="0" pageNumber="577">Figs 3B, C</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory apparatuses of the species of the genus Radix. A. Radix auricularia (19.07.2007, Russia, Tyumen' Region, Vylposl channel near Labytnangi Town); B. R. alticola (01.07.2016, Tajikistan, a warm brook near Dzhaushangoz village); C. R. euphratica (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village); D. R. makhrovi, a paratype (after Aksenova et al. 2018 a, modified). E. R. plicatula (26.11.2014, China, Bejing, an artificial pond in the former Emperor's summer palace). F. R. rubiginosa (Malaysia, Kuala-Lumpur); G. R. rufescens (24.11.2016, Myanmar, Yetho River, near the dam). H. R. natalensis (05.08.2018, Uganda, crater lake Kyamwiga). Scale bars: 2 mm." figureDoi="10.3897/zse.96.52860.figure4" httpUri="https://binary.pensoft.net/fig/448268" pageId="0" pageNumber="577">; 4B</figureCitation>
; 
<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characterisation of shells and copulative apparatuses of several species in the genus Radix." pageId="0" pageNumber="577">Table 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="577" type="reference_group">
<paragraph pageId="0" pageNumber="577">
<taxonomicName authorityName="Izzatullaev, Kruglov &amp; Starobogatov" authorityYear="1983" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lymnaea (Radix) alticola" order="Hygrophila" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="alticola" subGenus="Radix">Lymnaea (Radix) alticola</taxonomicName>
<bibRefCitation author="Izzatullaev, ZI" journalOrPublisher="Izvestiya Akademii Nauk Tajikskoy SSR, otdelenie biologicheskikh nauk" pageId="0" pageNumber="577" pagination="53 - 57" refId="B52" refString="Izzatullaev, ZI, Kruglov, ND, Starobogatov, YI, 1983. New and ill-known species of mollusks of the subgenus Radix of the genus Lymnaea of the USSR fauna from the Central Asia (Gastropoda, Pulmonata). Izvestiya Akademii Nauk Tajikskoy SSR, otdelenie biologicheskikh nauk 4: 53 - 57" title="New and ill-known species of mollusks of the subgenus Radix of the genus Lymnaea of the USSR fauna from the Central Asia (Gastropoda, Pulmonata)." volume="4" year="1983">Izzatullaev et al. 1983</bibRefCitation>
: 53, figs 1, 2; 
<bibRefCitation author="Kruglov, ND" journalOrPublisher="Ruthenica: The Russian Malacological Journal" pageId="0" pageNumber="577" pagination="65 - 92" refId="B69" refString="Kruglov, ND, Starobogatov, YI, 1993a. Annotated and illustrated catalogue of species of the family Lymnaeidae (Gastropoda Pulmonata Lymnaeiformes) of Palaearctic and adjacent river drainage areas. Part I. Ruthenica: The Russian Malacological Journal 3: 65 - 92" title="Annotated and illustrated catalogue of species of the family Lymnaeidae (Gastropoda Pulmonata Lymnaeiformes) of Palaearctic and adjacent river drainage areas. Part I." volume="3" year="1993 a">Kruglov and Starobogatov 1993a</bibRefCitation>
: 85, fig. 11C; 
<bibRefCitation author="Kruglov, ND" journalOrPublisher="SGPU Publishing, Smolensk" pageId="0" pageNumber="577" publicationUrl="http://herba.msu.ru/shipunov/school/books/kruglov2005_lymnaeidae.djvu" refId="B62" refString="Kruglov, ND, 2005. Mollusks of the Family Lymnaeidae (GastropodaPulmonata) in Europe and Northern Asia. SGPU Publishing, Smolensk, http://herba.msu.ru/shipunov/school/books/kruglov2005_lymnaeidae.djvu" title="Mollusks of the Family Lymnaeidae (GastropodaPulmonata) in Europe and Northern Asia." url="http://herba.msu.ru/shipunov/school/books/kruglov2005_lymnaeidae.djvu" year="2005">Kruglov 2005</bibRefCitation>
: 261, figs 146(1)-148; 
<bibRefCitation author="Sitnikova, TY" journalOrPublisher="Zoologicheskie Issledovania" pageId="0" pageNumber="577" pagination="7 - 37" publicationUrl="http://zmmu.msu.ru/files/books/zool_issl_16.pdf" refId="B116" refString="Sitnikova, TY, Sysoev, AV, Prozorova, LA, 2014. Types of freshwater gastropods described by Ya.I. Starobogatov, with additional data on the species: family Lymnaeidae. Zoologicheskie Issledovania 16: 7 - 37, http://zmmu.msu.ru/files/books/zool_issl_16.pdf" title="Types of freshwater gastropods described by Ya. I. Starobogatov, with additional data on the species: family Lymnaeidae." url="http://zmmu.msu.ru/files/books/zool_issl_16.pdf" volume="16" year="2014">Sitnikova et al. 2014</bibRefCitation>
: 25, fig. 8D.
</paragraph>
<paragraph pageId="0" pageNumber="577">
<taxonomicName baseAuthorityName="Izzatullaev, Kruglov &amp; Starobogatov" baseAuthorityYear="1983" class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radix (Radix) alticola" order="Hygrophila" pageId="0" pageNumber="577" phylum="Mollusca" rank="species" species="alticola" subGenus="Radix">Radix (Radix) alticola</taxonomicName>
- 
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="577" refId="B142" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
: 320.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="577" type="tl">
<paragraph pageId="0" pageNumber="577">TL.</paragraph>
<paragraph pageId="0" pageNumber="577">
Tajikistan, a hot spring near the 
<normalizedToken originalValue="Yashilkul’">Yashilkul'</normalizedToken>
Lake (approximately 
<geoCoordinate degrees="37" direction="north" minutes="47" orientation="latitude" precision="15" seconds="00" value="37.783333">37°47'00&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="72" direction="east" minutes="51" orientation="longitude" precision="15" seconds="00" value="72.85">72°51'00&quot;E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="577" type="types">
<paragraph pageId="0" pageNumber="577">Types.</paragraph>
<paragraph pageId="0" pageNumber="577">
ZISP (
<bibRefCitation author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="577" refId="B142" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
). The holotype is illustrated by 
<bibRefCitation author="Sitnikova, TY" journalOrPublisher="Zoologicheskie Issledovania" pageId="0" pageNumber="577" pagination="7 - 37" publicationUrl="http://zmmu.msu.ru/files/books/zool_issl_16.pdf" refId="B116" refString="Sitnikova, TY, Sysoev, AV, Prozorova, LA, 2014. Types of freshwater gastropods described by Ya.I. Starobogatov, with additional data on the species: family Lymnaeidae. Zoologicheskie Issledovania 16: 7 - 37, http://zmmu.msu.ru/files/books/zool_issl_16.pdf" title="Types of freshwater gastropods described by Ya. I. Starobogatov, with additional data on the species: family Lymnaeidae." url="http://zmmu.msu.ru/files/books/zool_issl_16.pdf" volume="16" year="2014">Sitnikova et al. (2014)</bibRefCitation>
. We studied both the holotype and the paratypes of 
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="577">
According to our data (
<bibRefCitation DOI="https://doi.org/10.1038/s41598-018-29451-1" author="Aksenova, OV" journalOrPublisher="Scientific Reports" pageId="0" pageNumber="577" pagination="1 - 17" refId="B1" refString="Aksenova, OV, Bolotov, IN, Gofarov, MY, Kondakov, AV, Vinarski, MV, Bespalaya, YV, Kolosova, YS, Palatov, DM, Sokolova, SE, Spitsyn, VM, Tomilova, AA, Travina, OV, Vikhrev, IV, 2018a. Species richness, molecular taxonomy and biogeography of the radicine pond snails (Gastropoda: Lymnaeidae) in the Old World. Scientific Reports 8 (11199): 1 - 17, DOI: https://doi.org/10.1038/s41598-018-29451-1" title="Species richness, molecular taxonomy and biogeography of the radicine pond snails (Gastropoda: Lymnaeidae) in the Old World." url="https://doi.org/10.1038/s41598-018-29451-1" volume="8" year="2018 a">Aksenova et al. 2018a</bibRefCitation>
), 
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
is endemic to the High Asia mountains; found in Tajikistan (from several sites, including the type locality) and Nepal. In mountain Tajikistan (Pamir), this species inhabits hot springs and satellite streams with relatively warm water.
</paragraph>
<paragraph pageId="0" pageNumber="577">
The shell shape of 
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
is similar to that of 
<taxonomicName genus="R." lsidName="R. auricularia" pageId="0" pageNumber="577" rank="species" species="auricularia">
<emphasis italics="true" pageId="0" pageNumber="577">R. auricularia</emphasis>
</taxonomicName>
, but the former species is of much smaller size; its shell height does not exceed 16 mm (see Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characterisation of shells and copulative apparatuses of several species in the genus Radix." httpUri="http://table.plazi.org/id/72A87B6AE3E005FDC64F6F3D96361C95" pageId="0" pageNumber="577" tableUuid="72A87B6AE3E005FDC64F6F3D96361C95">1</tableCitation>
), whereas the shells of 
<taxonomicName genus="R." lsidName="R. auricularia" pageId="0" pageNumber="577" rank="species" species="auricularia">
<emphasis italics="true" pageId="0" pageNumber="577">R. auricularia</emphasis>
</taxonomicName>
may reach 35-40 mm in height. The proportions of the copulatory organ of 
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
are very similar to 
<taxonomicName genus="R." lsidName="R. auricularia" pageId="0" pageNumber="577" rank="species" species="auricularia">
<emphasis italics="true" pageId="0" pageNumber="577">R. auricularia</emphasis>
</taxonomicName>
(compare Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of species in the genus Radix. A. Radix auricularia (28.08.2013, Kazakhstan, Karaganda Region, Suresai River; LMBI); B. R. alticola (20.06.2012, Tajikistan, a hot spring near Djelandy village; LMBI); C. R. alticola (01.07.2016, Tajikistan, a warm brook near Dzhaushangoz village; LMBI); D. R. brevicauda (Kashmir, a syntype; NHMUK); E. R. brevicauda (05.08.1948, Tajikistan, Pamir Mts., Shaimak village, in a warm spring; ZISP); F. R. euphratica (26.06.2016, Tajikistan, Dushanbe, a fountain near the President's palace; LMBI); G. R. euphratica (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI); H. R. euphratica (without date, Iraq, Bagdad; NHMUK); I. R. makhrovi, the holotype (China, Tibet, a roadside ditch west of the Lhasa River mouth; ZISP); J. R. plicatula, a probable syntype (China, Chusan, NHMUK); K. R. plicatula (26.07.2017, China, Uyghuria, Bagrash-kol' Lake; LMBI); L. R. plicatula (26.11.2014, China, Bejing, an artificial pond in the former Emperor's summer palace; LMBI). Scale bars: 2 mm (B-C, F-L), 5 mm (A, D-E, K)." figureDoi="10.3897/zse.96.52860.figure3" httpUri="https://binary.pensoft.net/fig/448267" pageId="0" pageNumber="577">3A</figureCitation>
and 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of species in the genus Radix. A. Radix auricularia (28.08.2013, Kazakhstan, Karaganda Region, Suresai River; LMBI); B. R. alticola (20.06.2012, Tajikistan, a hot spring near Djelandy village; LMBI); C. R. alticola (01.07.2016, Tajikistan, a warm brook near Dzhaushangoz village; LMBI); D. R. brevicauda (Kashmir, a syntype; NHMUK); E. R. brevicauda (05.08.1948, Tajikistan, Pamir Mts., Shaimak village, in a warm spring; ZISP); F. R. euphratica (26.06.2016, Tajikistan, Dushanbe, a fountain near the President's palace; LMBI); G. R. euphratica (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI); H. R. euphratica (without date, Iraq, Bagdad; NHMUK); I. R. makhrovi, the holotype (China, Tibet, a roadside ditch west of the Lhasa River mouth; ZISP); J. R. plicatula, a probable syntype (China, Chusan, NHMUK); K. R. plicatula (26.07.2017, China, Uyghuria, Bagrash-kol' Lake; LMBI); L. R. plicatula (26.11.2014, China, Bejing, an artificial pond in the former Emperor's summer palace; LMBI). Scale bars: 2 mm (B-C, F-L), 5 mm (A, D-E, K)." figureDoi="10.3897/zse.96.52860.figure3" httpUri="https://binary.pensoft.net/fig/448267" pageId="0" pageNumber="577">3B</figureCitation>
).
</paragraph>
<caption ID-Table-UUID="72A87B6AE3E005FDC64F6F3D96361C95" httpUri="http://table.plazi.org/id/72A87B6AE3E005FDC64F6F3D96361C95" pageId="0" pageNumber="577" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="577">
<emphasis bold="true" pageId="0" pageNumber="577">Table 1.</emphasis>
Morphological characterisation of shells and copulative apparatuses of several species in the genus 
<taxonomicName class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radix" pageId="0" pageNumber="577" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="577">Radix</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="577">
<table pageId="0" pageNumber="577">
<tr pageId="0" pageNumber="577">
<th colspan="1" pageId="0" pageNumber="577" rowspan="3">Character / index</th>
<th colspan="10" pageId="0" pageNumber="577" rowspan="1">Species / locality</th>
</tr>
<tr pageId="0" pageNumber="577">
<th colspan="2" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. alticola" pageId="0" pageNumber="577" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="577">R. alticola</emphasis>
</taxonomicName>
</th>
<th colspan="2" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. euphratica" pageId="0" pageNumber="577" rank="species" species="euphratica">
<emphasis italics="true" pageId="0" pageNumber="577">R. euphratica</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. brevicauda" pageId="0" pageNumber="577" rank="species" species="brevicauda">
<emphasis italics="true" pageId="0" pageNumber="577">R. brevicauda</emphasis>
</taxonomicName>
</th>
<th colspan="2" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. plicatula" pageId="0" pageNumber="577" rank="species" species="plicatula">
<emphasis italics="true" pageId="0" pageNumber="577">R. plicatula</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. natalensis" pageId="0" pageNumber="577" rank="species" species="natalensis">
<emphasis italics="true" pageId="0" pageNumber="577">R. natalensis</emphasis>
</taxonomicName>
</th>
<th colspan="2" pageId="0" pageNumber="577" rowspan="1">
<taxonomicName genus="R." lsidName="R. rufescens" pageId="0" pageNumber="577" rank="species" species="rufescens">
<emphasis italics="true" pageId="0" pageNumber="577">R. rufescens</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="577">
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Tajikistan, Zor-Kul Lake (paratypes)</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Tajikistan, a warm brook near Dzhaushangoz village*</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
Tajikistan, Dushanbe, a fountain near 
<normalizedToken originalValue="President’s">President's</normalizedToken>
palace*
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Tajikistan, a roadside ditch near Kurban-Shakhid village*</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Tajikistan, a warm spring near Shaimak village</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
China, Bejing, a pond near the former 
<normalizedToken originalValue="Emperor’s">Emperor's</normalizedToken>
palace*
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">
China, Uyghuria, 
<normalizedToken originalValue="Bagrashkol’">Bagrashkol'</normalizedToken>
Lake*
</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Uganda, the Kyamwiga crater lake*, **</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">India, &quot;Khipru, Sind&quot;</th>
<th colspan="1" pageId="0" pageNumber="577" rowspan="1">Myanmar, Yetho River near the dam*</th>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">Repository</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">ZISP</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">ZISP</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">NHMUK</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">LMBI</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">Number of specimens measured (dissected)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">45 (0)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">44 (16)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">4(3)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">21(15)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">21(0)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">29(11)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">16(9)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">26(20)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">40(0)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12(6)</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Whorls number</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.00-3.75</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.00-3.75</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.87-4.25</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.25-4.12</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.25-4.00</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.75-4.00</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.50-4.12</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.25-3.75</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.75-4.75</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.87-4.50</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.40
<normalizedToken originalValue="±">+/-</normalizedToken>
0.18
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.30
<normalizedToken originalValue="±">+/-</normalizedToken>
0.17
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.06
<normalizedToken originalValue="±">+/-</normalizedToken>
0.16
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.71
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.48
<normalizedToken originalValue="±">+/-</normalizedToken>
0.19
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.08
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.76
<normalizedToken originalValue="±">+/-</normalizedToken>
0.23
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.51
<normalizedToken originalValue="±">+/-</normalizedToken>
0.15
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.17
<normalizedToken originalValue="±">+/-</normalizedToken>
0.21
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.22
<normalizedToken originalValue="±">+/-</normalizedToken>
0.22
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Shell height, mm (SH)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.0-13.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.4-15.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.6-15.1</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.5-20.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.8-15.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">13.6-20.7</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.9-22.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">13.6-18.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">37.9-43.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">17.5-28.9</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
13.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.8
<normalizedToken originalValue="±">+/-</normalizedToken>
2.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
12.1
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
16.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
15.8
<normalizedToken originalValue="±">+/-</normalizedToken>
3.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
16.2
<normalizedToken originalValue="±">+/-</normalizedToken>
1.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
39.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
23.2
<normalizedToken originalValue="±">+/-</normalizedToken>
3.8
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Shell width, mm (SW)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.4-10.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.8-13.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.4-9.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.6-13.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.4-13.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.9-14.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.2-14.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.0-13.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">17.8-20.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.1-13.6</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
8.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
8.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.3
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.1
<normalizedToken originalValue="±">+/-</normalizedToken>
2.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
18.8
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.0
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Spire height, mm (SpH)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">1.4-2.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.0-4.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.8-4.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.7-5.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.2-4.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.4-6.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.7-7.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.8-4.7</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">10.6-14.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">4.5-10.8</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
2.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
2.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
2.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
5.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
12.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.8
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Body whorl height, mm (BWH)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.4-12.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.7-14.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">10.1-12.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">11.0-17.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.9-13.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">11.7-18.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">11.5-20.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.3-17.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">32.8-37.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">15.6-25.2</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
13.1
<normalizedToken originalValue="±">+/-</normalizedToken>
1.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.1
<normalizedToken originalValue="±">+/-</normalizedToken>
2.8
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.8
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
34.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
20.1
<normalizedToken originalValue="±">+/-</normalizedToken>
3.3
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Aperture height, mm (AH)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.6-11.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.7-12.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.1-12.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.4-15.1</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">7.7-11.7</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">8.8-16.0</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">9.4-16.1</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">10.2-14.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">27.0-31.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.7-19.1</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.6
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
10.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
9.6
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.8
<normalizedToken originalValue="±">+/-</normalizedToken>
1.7
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
11.6
<normalizedToken originalValue="±">+/-</normalizedToken>
2.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
12.6
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
28.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
15.5
<normalizedToken originalValue="±">+/-</normalizedToken>
2.2
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Aperture width, mm (AW)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.0-8.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.2-10.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.8-7.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.8-11.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">5.5-8.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.3-12.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.1-10.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.0-9.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">12.3-16.3</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">6.5-11.3</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
6.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.7
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
6.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
6.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.0
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
8.2
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
7.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.7
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
14.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
8.6
<normalizedToken originalValue="±">+/-</normalizedToken>
1.4
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Praeputium length, mm (PP)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.2-5.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.8-3.6</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.2-4.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.4-6.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.0-5.4</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.1-4.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.8-4.2</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">Penis sheath length, mm (PS)</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.2-5.1</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.9-3.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.2-3.9</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.6-9.2</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.6-7.8</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">3.0-5.5</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">2.2-3.6</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
3.1
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
5.8
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
5.5
<normalizedToken originalValue="±">+/-</normalizedToken>
1.3
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
4.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
2.9
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">SW/SH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.65-0.81</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.72-0.87</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.62-0.67</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.61-0.73</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.79-0.94</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.53-0.76</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.55-0.72</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.57-0.71</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.44-0.50</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.41-0.52</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.74
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.80
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.64
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.66
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.85
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.63
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.64
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.65
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.47
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.48
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">SpH/SH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.13-0.24</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.18-0.27</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.28-0.32</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.20-0.30</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.18-0.28</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.20-0.42</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.20-0.33</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.19-0.27</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.27-0.38</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.26-0.37</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.19
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.22
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.30
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.23
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.31
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.28
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.23
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.32
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.33
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">BWH/SH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.87-0.93</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.88-0.98</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.80-0.86</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.83-0.94</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.87-0.92</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.83-0.95</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.86-0.92</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.86-0.93</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.84-0.89</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.80-0.90</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.90
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.91
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.84
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.89
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.90
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.87
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.89
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.91
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.87
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.86
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">AH/SH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.65-0.81</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.75-0.85</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.70-0.74</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.70-0.86</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.72-0.86</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.59-0.86</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.69-0.81</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.74-0.82</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.69-0.77</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.63-0.75</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.74
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.81
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.73
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.77
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.80
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.71
<normalizedToken originalValue="±">+/-</normalizedToken>
0.06
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.74
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.78
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.73
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.67
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">AW/AH</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.59-0.83</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.66-0.91</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.63-0.67</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.57-0.78</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.67-0.81</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.63-0.78</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.56-0.73</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.57-0.67</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.45-0.55</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.50-0.61</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.69
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.72
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.65
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.65
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.73
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.70
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.64
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.62
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.50
<normalizedToken originalValue="±">+/-</normalizedToken>
0.03
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.55
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04
</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">ICA</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.81-1.18</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.93-1.16</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">1.10-1.69</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.54-0.96</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.64-0.90</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">0.84-1.13</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">1.09-1.27</td>
</tr>
<tr pageId="0" pageNumber="577">
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.97
<normalizedToken originalValue="±">+/-</normalizedToken>
0.11
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
1.01
<normalizedToken originalValue="±">+/-</normalizedToken>
0.12
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
1.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.18
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.76
<normalizedToken originalValue="±">+/-</normalizedToken>
0.14
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.79
<normalizedToken originalValue="±">+/-</normalizedToken>
0.08
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
0.96
<normalizedToken originalValue="±">+/-</normalizedToken>
0.07
</td>
<td colspan="1" pageId="0" pageNumber="577" rowspan="1">
1.19
<normalizedToken originalValue="±">+/-</normalizedToken>
0.06
</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="577">
<tableNote pageId="0" pageNumber="577">
*Sample studied genetically. ** The short-spired morph of 
<taxonomicName genus="R." lsidName="R. natalensis" pageId="0" pageNumber="577" rank="species" species="natalensis">
<emphasis italics="true" pageId="0" pageNumber="577">R. natalensis</emphasis>
</taxonomicName>
.
</tableNote>
</paragraph>
</subSubSection>
</treatment>
</document>