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<document id="AE9A01CB7C8171A7914E766AF6ACC340" ID-CLB-Dataset="55667" ID-DOI="10.26107/RBZ-2020-0097" ID-GBIF-Dataset="6cb155ce-8b9f-48ce-8e6c-b0dc00f8b1cc" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351295" ID-ZooBank="815E4670-B063-4FD8-B31E-3AD89B3A7942" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630205841501" checkinUser="diego" docAuthor="Schubart, Christoph D. &amp; Ng, Peter K. L." docDate="2020" docId="49152B56FFE6BA2AFCAFF8A9FF75F9A7" docLanguage="en" docName="RafflesBZool.68.891-994.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chiromantes haematocheir" docType="treatment" docVersion="12" lastPageNumber="902" masterDocId="B52C532EFFE1BA21FFDDFFB3FF8DFFB1" masterDocTitle="Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species" masterLastPageNumber="994" masterPageNumber="891" pageNumber="898" updateTime="1699203722016" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FB7C5F3DED417A048E2C48C5A6E9E983">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<mods:namePart id="C8F33DD304FB079C0EA0C3CB84EB2C3E">Schubart, Christoph D.</mods:namePart>
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<mods:namePart id="01BEEB6E0E1C89E416F258817A51C651">Ng, Peter K. L.</mods:namePart>
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<taxonomicName id="06BCE1C3FFE6BA26FCAFF8A9FAEBF883" ID-CoL="69T63" authority="(De Haan, 1833)" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[882,1382,1818,1842]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="898" phylum="Arthropoda" rank="species" species="haematocheir">
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<emphasis id="F3C84652FFE6BA26FCAFF8A9FB2DF883" bold="true" box="[882,1184,1818,1842]" italics="true" pageId="7" pageNumber="898">Chiromantes haematocheir</emphasis>
(
<bibRefCitation id="A52DE7B1FFE6BA26FB6CF8A9FAD3F883" author="De Haan" box="[1201,1374,1818,1842]" firstAuthor="De Haan" pageId="7" pageNumber="898" pagination="1 - 55" refId="ref69759" refString="Haan W De (1833 - 1849) Crustacea. In: Siebold PF (ed.) Fauna Japonica sive Descriptio Animalium, quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, Qui Summum in India Batava Imperium Tenent, Suscepto, Annis 18231830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit, (Crustacea). LugduniBatavorum. Pp. ixvii + ixxxi + iixxvi + 1 - 243, pls. A - J, L - Q, 1 - 55, circ. table 2." type="book chapter" year="1833">De Haan, 1833</bibRefCitation>
)
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<subSubSection id="89A6C9CBFFE6BA26FCB1F889FAE1F8E3" box="[876,1388,1850,1874]" pageId="7" pageNumber="898" type="description">
<paragraph id="C1039A40FFE6BA26FCB1F889FAE1F8E3" blockId="7.[876,1388,1818,1874]" box="[876,1388,1850,1874]" pageId="7" pageNumber="898">
(
<figureCitation id="598786C5FFE6BA26FCA9F889FC59F8E3" box="[884,980,1850,1874]" captionStart="Fig" captionStartId="7.[113,148,1626,1648]" captionTargetBox="[113,1449,189,1611]" captionTargetId="figure-14@7.[113,1449,189,1611]" captionTargetPageId="7" captionText="Fig. 1. Overall habitus. A, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, C. ryukyuanus, holotype male (33.1 × 29.6 mm) (RUMF-ZC-539), Okinawa Island, Japan; C, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; D, O. dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; E, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; F, O. magnum, paratype male (50.6 × 45.6 mm) (ZRC 2013.0173), Ogasawara Island, Japan; G, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong; H, O. patshuni, male (20.4 × 18.2 mm) (ZRC 2012.0032), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351297" httpUri="https://zenodo.org/record/5351297/files/figure.png" pageId="7" pageNumber="898">Figs. 1A</figureCitation>
, 
<figureCitation id="598786C5FFE6BA26FC3FF889FB88F8E3" box="[994,1029,1850,1874]" captionStart="Fig" captionStartId="10.[136,171,1435,1457]" captionTargetBox="[136,1476,189,1419]" captionTargetId="figure-7@10.[136,1476,189,1419]" captionTargetPageId="10" captionText="Fig. 3. Frontal view of cephalothorax. A, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; C, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; D, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; E, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; F, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351301" httpUri="https://zenodo.org/record/5351301/files/figure.png" pageId="7" pageNumber="898">3A</figureCitation>
, 
<figureCitation id="598786C5FFE6BA26FBCFF889FBD9F8E3" box="[1042,1108,1850,1874]" captionStart="Fig" captionStartId="14.[135,170,1987,2009]" captionTargetBox="[136,1476,189,1972]" captionTargetId="figure-7@14.[136,1476,189,1972]" captionTargetPageId="14" captionText="Fig. 5. Chela. A, B, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; C, C. haematocheir, male (18.6 × 20.7 mm) (RUMF-ZC-544), Japan; D–F, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; G, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; H, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong; I, O. patshuni, male (20.4 × 18.2 mm) (ZRC 2012.0032), Hong Kong. A, D, G, H, I, outer view; B, C, F, dorso-lateral view; E, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351305" httpUri="https://zenodo.org/record/5351305/files/figure.png" pageId="7" pageNumber="898">5A–C</figureCitation>
, 
<figureCitation id="598786C5FFE6BA26FBBCF889FB28F8E3" box="[1121,1189,1850,1874]" captionStart="Fig" captionStartId="17.[113,148,1943,1965]" captionTargetBox="[113,1453,223,1927]" captionTargetId="figure-14@17.[113,1453,223,1927]" captionTargetPageId="17" captionText="Fig. 7. A–D, Chiromantes haematocheir, anterior male thoracic sternum; A, male (10.8 × 9.4 mm) (ZRC 2012.1222), Hong Kong; B, male (14.3 × 13.0 mm) (ZRC 2012.1222), Hong Kong; C, male (18.4 × 16.6 mm) (lectotype of Holometopus serenei) (NHM 1978.107), Hong Kong; D, male (27.8 × 23.9 mm) (ZRC 2012.1221), Hong Kong. E–H, left third ambulatory legs; E–G, Orisarma dehaani: E, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; F, male (35.4 × 32.4 mm) (ZRC 2000.2263), Iriomote Island, Japan; G, male (29.6 × 27.6 mm) (ZRC 2002.0223), Hong Kong; H, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China." figureDoi="http://doi.org/10.5281/zenodo.5351309" httpUri="https://zenodo.org/record/5351309/files/figure.png" pageId="7" pageNumber="898">7A–D</figureCitation>
, 
<figureCitation id="598786C5FFE6BA26FB6FF889FB58F8E3" box="[1202,1237,1850,1874]" captionStart="Fig" captionStartId="20.[135,170,1977,1999]" captionTargetBox="[195,1417,189,1962]" captionTargetId="figure-7@20.[195,1417,189,1962]" captionTargetPageId="20" captionText="Fig. 9. Male anterior thoracic sternum and pleon. A, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, C. ryukyuanus, holotype male (33.1 × 29.6 mm) (RUMF-ZC-539), Okinawa Island, Japan; C, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; D, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; E, O. intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; F, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; G, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; H, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351313" httpUri="https://zenodo.org/record/5351313/files/figure.png" pageId="7" pageNumber="898">9A</figureCitation>
, 
<figureCitation id="598786C5FFE6BA26FB3FF889FAA5F8E3" box="[1250,1320,1850,1874]" captionStart="Fig" captionStartId="22.[135,170,1571,1593]" captionTargetBox="[136,1477,191,1554]" captionTargetId="figure-7@22.[120,1496,183,1603]" captionTargetPageId="22" captionText="Fig. 10. A, D, E–I, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, male (26.4 × 22.9 mm) (ZRC 1970.8.27.7), Sagami Bay, Japan; C, C. haematocheir, male (lectotype of Holometopus serenei) (18.4 × 16.6 mm) (NHM 1978.107), Hong Kong; J–L, C. ryukyuanus, holotype male (33.1 × 29.6 mm) (RUMF-ZC-539), Okinawa Island, Japan. A, J, male pleon; B, male pleonal somite 6 and telson; male pleonal somites 4–6, telson; D, anterior thoracic sternites 1–4; E, left G1 (ventral view, denuded); F, left G1 (dorsal view, denuded); G, left distal part of G1 (ventral view, denuded); H, left distal part of G1 (dorsal view, denuded); I, L, left G2 (denuded); K, left G1. Scales: A, B, D = 5.0 mm; C, E, F, I = 2.0 mm; G, H = 1.0 mm. J–L, after Naruse &amp; Ng (2008: fig. 8)." figureDoi="http://doi.org/10.5281/zenodo.5351315" httpUri="https://zenodo.org/record/5351315/files/figure.png" pageId="7" pageNumber="898">10A–I</figureCitation>
, 
<figureCitation id="598786C5FFE6BA26FAE8F889FAE9F8E3" box="[1333,1380,1850,1874]" captionStart="Fig" captionStartId="72.[136,171,1898,1920]" captionTargetBox="[136,1474,211,1882]" captionTargetId="figure-8@72.[136,1474,211,1882]" captionTargetPageId="72" captionText="Fig. 43. Vulva. A, Chiromantes haematocheir, female (27.0 × 23.7 mm) (ZRC 2002.0225), Japan; B, Orisarma dehaani, female (20.4 × 17.8 mm) (ZRC 2012.0057), Japan; C, Orisarma intermedium, female (31.8 × 28.4 mm) (ZRC 2013.0140), Japan; D, Orisarma patshuni, female (20.5 × 18.1 mm) (ZRC 2014.0817), Hong Kong; E, Danarma obtusifrons, female (15.7 × 11.6 mm) (UF-FLMNH 14837), Hawaii; F, Danarma eurymerus, paratype ovigerous female (18.0 × 13.4 mm) (ZRC 2012.0956), Taiwan; G, Cristarma eulimene, female (15.6 × 12.8 mm) (ZRC 1968.1.22.3), Mozambique; H, Cristarma ortmanni, female (13.1 × 10.2 mm) (ZRC 2000.1783), Kenya; I, Trapezarma angolense, female (18.2 × 14.3 mm) (ZRC 2015.0297), Cameroon; J, Platychirarma buettikoferi, ovigerous female (11.3 × 9.1 mm) (ZRC 2015.0298), Cameroon." figureDoi="http://doi.org/10.5281/zenodo.5351373" httpUri="https://zenodo.org/record/5351373/files/figure.png" pageId="7" pageNumber="898">43A</figureCitation>
)
</paragraph>
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<treatmentCitation id="401DBC51FFE6BA26FCF0F8CAFAB2F83E" author="Haan W De" box="[813,1343,1913,1935]" page="62" pageId="7" pageNumber="898" year="1833">
<taxonomicName id="06BCE1C3FFE6BA26FCF0F8CAFAB2F83E" authority="De Haan, 1833: 62" authorityName="De Haan" authorityPageNumber="62" authorityYear="1833" box="[813,1343,1913,1935]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="898" phylum="Arthropoda" rank="species" species="haematocheir" subGenus="Pachysoma">
<emphasis id="F3C84652FFE6BA26FCF0F8CAFCF3F83F" box="[813,894,1913,1934]" italics="true" pageId="7" pageNumber="898">Grapsus</emphasis>
(
<emphasis id="F3C84652FFE6BA26FC57F8CAFC75F83F" box="[906,1016,1913,1934]" italics="true" pageId="7" pageNumber="898">Pachysoma</emphasis>
) 
<emphasis id="F3C84652FFE6BA26FBD8F8CAFB05F83F" box="[1029,1160,1913,1934]" italics="true" pageId="7" pageNumber="898">haematocheir</emphasis>
<bibRefCitation id="A52DE7B1FFE6BA26FB50F8CAFAB2F83E" author="De Haan" box="[1165,1343,1913,1935]" firstAuthor="De Haan" page="62" pageId="7" pageNumber="898" pagination="1 - 55" refId="ref69759" refString="Haan W De (1833 - 1849) Crustacea. In: Siebold PF (ed.) Fauna Japonica sive Descriptio Animalium, quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, Qui Summum in India Batava Imperium Tenent, Suscepto, Annis 18231830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit, (Crustacea). LugduniBatavorum. Pp. ixvii + ixxxi + iixxvi + 1 - 243, pls. A - J, L - Q, 1 - 55, circ. table 2." type="book chapter" year="1833">De Haan, 1833: 62</bibRefCitation>
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, pl. 7 fig. 4.
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<taxonomicName id="06BCE1C3FFE6BA26FCF0F826FBC4F81B" ID-CoL="99432" authority="(De Haan, 1833)" box="[813,1097,1941,1962]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="898" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE6BA26FCF0F826FBC4F81B" box="[813,1097,1941,1962]" italics="true" pageId="7" pageNumber="898">Holometopus haematocheir</emphasis>
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– H. 
<treatmentCitation id="401DBC51FFE6BA26FB51F826FA2AF81A" author="Milne Edwards H" box="[1164,1447,1941,1963]" page="188" pageId="7" pageNumber="898" year="1853">
<bibRefCitation id="A52DE7B1FFE6BA26FB51F826FA2AF81A" author="Milne Edwards" box="[1164,1447,1941,1963]" firstAuthor="Milne Edwards" page="188" pageId="7" pageNumber="898" pagination="163 - 228" refId="ref72839" refString="Milne Edwards H (1853) Memoires sur la famille des Ocypodiens, suite. Annales de Sciences Naturelle, Zoology, Paris, Serie 3 (Zoologie) 20: 163 - 228, pls. 6 - 11. [A continuation of H. Milne Edwards, 1852, and reprinted with it in the undated Melanges Carcinologiques pp. 129 - 196]" type="journal article" year="1853">Milne Edwards, 1853: 188</bibRefCitation>
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; 
<treatmentCitation id="401DBC51FFE6BA26FC8EF801FB96F879" author="Stimpson W" box="[851,1051,1970,1992]" page="106" pageId="7" pageNumber="898" year="1858">
<bibRefCitation id="A52DE7B1FFE6BA26FC8EF801FB96F879" author="Stimpson" box="[851,1051,1970,1992]" firstAuthor="Stimpson" page="106" pageId="7" pageNumber="898" pagination="93 - 110" refId="ref77553" refString="Stimpson W (1858) Crustacea Ocypodoidea. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars V. Proceedings of the Academy of Natural Sciences of Philadelphia, 10: 93 - 110." type="journal article" year="1858">Stimpson, 1858: 106</bibRefCitation>
</treatmentCitation>
(part); 
<treatmentCitation id="401DBC51FFE6BA26FBB9F801FA89F879" author="Heller C" box="[1124,1284,1970,1992]" page="66" pageId="7" pageNumber="898" year="1865">
<bibRefCitation id="A52DE7B1FFE6BA26FBB9F801FA89F879" author="Heller" box="[1124,1284,1970,1992]" firstAuthor="Heller" page="66" pageId="7" pageNumber="898" refId="ref69955" refString="Heller C (1865) Crustaceen. In: Reise der Osterreichischen Fregatte Novara um die Ede in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodores B. von Wullerstorf-Urbair. Zoologischer Theil. Zweiter Band, Volume 3 (1). Pp. 1 - 280, 25 pls." type="journal volume" year="1865">Heller, 1865: 66</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE6BA26FAD3F801FCFBF855" author="Stimpson W" page="137" pageId="7" pageNumber="898" year="1907">
<bibRefCitation id="A52DE7B1FFE6BA26FAD3F801FCFBF855" author="Stimpson" firstAuthor="Stimpson" page="137" pageId="7" pageNumber="898" pagination="1 - 240" refId="ref77609" refString="Stimpson W (1907) Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition, 1853 - 1856. Smithsonian Miscellaneous Collections, 49 (1717): 1 - 240, pls. 1 - 26." type="journal article" year="1907">Stimpson, 1907: 137</bibRefCitation>
</treatmentCitation>
(part); 
<treatmentCitation id="401DBC51FFE6BA26FC61F87CFBFDF854" author="Takeda M" box="[956,1136,1999,2021]" page="146" pageId="7" pageNumber="898" year="1975">
<bibRefCitation id="A52DE7B1FFE6BA26FC61F87CFBFDF854" author="Takeda" box="[956,1136,1999,2021]" firstAuthor="Takeda" page="146" pageId="7" pageNumber="898" pagination="120 - 149" refId="ref77707" refString="Takeda M (1975) Brachyura. In: Utinomi H (ed.) Aquatic Invertebrates. First Edition. Gakken Chukosei zukan, 9: 120 - 149 (fig. with Latin names), 188 - 331 (descriptions)." type="journal article" year="1975">Takeda, 1975: 146</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE6BA26FBA4F87CFB7EF855" author="Soh CL" box="[1145,1267,1999,2021]" page="9" pageId="7" pageNumber="898" year="1978">
<bibRefCitation id="A52DE7B1FFE6BA26FBA4F87CFB7EF855" author="Soh" box="[1145,1267,1999,2021]" firstAuthor="Soh" page="9" pageId="7" pageNumber="898" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh, 1978: 9</bibRefCitation>
</treatmentCitation>
, 10, pl. 2e; 
<treatmentCitation id="401DBC51FFE6BA26FA82F87CFC3AF7B0" author="Takeda M" page="219" pageId="7" pageNumber="898" year="1982">
<bibRefCitation id="A52DE7B1FFE6BA26FA82F87CFC3AF7B0" author="Takeda" firstAuthor="Takeda" page="219" pageId="7" pageNumber="898" refId="ref77752" refString="Takeda M (1982) Keys to the Japanese and foreign crustaceans fully illustrated in colors. Hokuryukan, Tokyo, vi + 285 pp., 58 pp. (keys)." type="book" year="1982">Takeda, 1982: 219</bibRefCitation>
</treatmentCitation>
, fig. 650.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFE6BA28FCF0F7BAFE8CFF5E" blockId="7.[813,1453,1913,2108]" lastBlockId="9.[113,753,189,960]" lastPageId="9" lastPageNumber="900" pageId="7" pageNumber="898">
<treatmentCitationGroup id="E1ACBD6EFFE6BA28FCF0F7BAFE8CFF5E" lastPageId="9" lastPageNumber="900" pageId="7" pageNumber="898">
<taxonomicName id="06BCE1C3FFE6BA26FCF0F7BAFB87F7AF" ID-CoL="7WV46" authorityName="De Haan" authorityYear="1833" box="[813,1034,2057,2078]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="898" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE6BA26FCF0F7BAFB87F7AF" box="[813,1034,2057,2078]" italics="true" pageId="7" pageNumber="898">Sesarma haematocheir</emphasis>
</taxonomicName>
– 
<treatmentCitation id="401DBC51FFE6BA26FBF9F7BAFB56F7AF" author="Herklots JA" box="[1060,1243,2057,2079]" page="17" pageId="7" pageNumber="898" year="1861">
<bibRefCitation id="A52DE7B1FFE6BA26FBF9F7BAFB56F7AF" author="Herklots" box="[1060,1243,2057,2079]" firstAuthor="Herklots" page="17" pageId="7" pageNumber="898" pagination="1 - 43" refId="ref70050" refString="Herklots JA (1861) Symbolae Carcinologicae. I. Catalogue des Crustaces qui ont servi de base au systeme carcinologique de M. W. de Haan, redige d'apres la collection du Musee des Pays- Bas et les Crustaces de la Faune du Japan. Tijdschrift voor Entomologie uitgegeven door de Nederlandische Entomologische Vereeniging, Leyden, 1861: 1 - 43." type="journal article" year="1861">Herklots, 1861: 17</bibRefCitation>
</treatmentCitation>
; 
<bibRefCitation id="A52DE7B1FFE6BA26FB3AF7BAFA25F7AE" author="De Man" box="[1255,1448,2057,2079]" firstAuthor="De Man" page="642" pageId="7" pageNumber="898" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">
De 
<treatmentCitation id="401DBC51FFE6BA26FAD7F7BAFA25F7AE" author="Man JG De" box="[1290,1448,2057,2079]" page="642" pageId="7" pageNumber="898" year="1887">Man, 1887: 642</treatmentCitation>
</bibRefCitation>
; 
<treatmentCitation id="401DBC51FFE6BA26FC8FF795FB87F78A" author="Burger O" box="[850,1034,2085,2107]" page="614" pageId="7" pageNumber="898" year="1893">
<bibRefCitation id="A52DE7B1FFE6BA26FC8FF795FB87F78A" author="Burger" box="[850,1034,2085,2107]" firstAuthor="Burger" page="614" pageId="7" pageNumber="898" pagination="613 - 632" refId="ref68026" refString="Burger O (1893) Beitrage zur Kenntniss der Gattung Sesarma. Zoologischen Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7 (4): 613 - 632, pl. 21." type="journal article" year="1893">Bürger, 1893: 614</bibRefCitation>
</treatmentCitation>
, pl. 21 fig. 3; 
<treatmentCitation id="401DBC51FFE6BA26FB43F795FAEBF78A" author="Ortmann AE" box="[1182,1382,2086,2107]" page="717" pageId="7" pageNumber="898" year="1894">
<bibRefCitation id="A52DE7B1FFE6BA26FB43F795FAEBF78A" author="Ortmann" box="[1182,1382,2086,2107]" firstAuthor="Ortmann" page="717" pageId="7" pageNumber="898" pagination="683 - 772" refId="ref74488" refString="Ortmann AE (1894) Abtheilung: Brachyura (Brachyura genuina Boas), III. Unterabtheilung: Cancroidea. 2. Section: Cancrinea, 2. Gruppe: Catametopa. Die Decapoden-Krebse des Strassburger Museums mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. Theil VIII [= Part 8]. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7 (5): 683 - 772, pl. 23." type="journal article" year="1894">Ortmann, 1894: 717</bibRefCitation>
</treatmentCitation>
(part); 
<treatmentCitation id="401DBC51FFE8BA28FF4AFF0EFE6DFF63" author="Yamaguchi T &amp; Baba K" box="[151,480,189,211]" page="473" pageId="9" pageNumber="900" year="1993">
<bibRefCitation id="A52DE7B1FFE8BA28FF4AFF0EFE6DFF63" author="Yamaguchi &amp; Baba" box="[151,480,189,211]" firstAuthor="Yamaguchi" page="473" pageId="9" pageNumber="900" pagination="145 - 570" refId="ref78696" refString="Yamaguchi T &amp; Baba K (1993) Crustacean specimens collected in Japan by Ph. F. von Siebold and H. Burger and held by the Nationaal Natuurhistorisch Museum in Leiden and other Museums. In: Yamaguchi T (ed.) Ph. F. von Siebold and Natural History of Japan Crustacea. The Carcinological Society of Japan, Tokyo. Pp. 145 - 570." type="book chapter" year="1993">
<collectingRegion id="037854A2FFE8BA28FF4AFF0EFE87FF62" box="[151,266,189,211]" country="Japan" name="Yamaguchi" pageId="9" pageNumber="900">Yamaguchi</collectingRegion>
&amp; 
<collectingRegion id="037854A2FFE8BA28FEECFF0EFEEAFF62" box="[305,359,189,211]" country="Myanmar" name="Chin" pageId="9" pageNumber="900">Baba</collectingRegion>
, 1993: 473
</bibRefCitation>
</treatmentCitation>
, fig. 180c; 
<treatmentCitation id="401DBC51FFE8BA28FD81FF0DFF70FF5E" author="Fransen CHJM &amp; Holthuis LB &amp; Adema JPHM" page="129" pageId="9" pageNumber="900" year="1997">
<bibRefCitation id="A52DE7B1FFE8BA28FD81FF0DFF70FF5E" author="Fransen" etAl="et al." firstAuthor="Fransen" page="129" pageId="9" pageNumber="900" pagination="1 - 344" refId="ref69235" refString="Fransen CHJM, Holthuis LB &amp; Adema JPHM (1997) Typecatalogue of the Decapod Crustacea in the collections of the Nationaal Naturhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Verhandelingen, Leiden, 311: i - xvi, 1 - 344." type="journal article" year="1997">Fransen et al., 1997: 129</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<caption id="95C3CAC8FFE9BA29FF55F835FAD2F79D" ID-DOI="http://doi.org/10.5281/zenodo.5351299" ID-Zenodo-Dep="5351299" httpUri="https://zenodo.org/record/5351299/files/figure.png" pageId="8" pageNumber="899" startId="8.[136,171,1926,1948]" targetBox="[136,1476,212,1910]" targetPageId="8">
<paragraph id="C1039A40FFE9BA29FF55F835FAD2F79D" blockId="8.[136,1475,1926,2092]" pageId="8" pageNumber="899">
Fig. 2. Overall habitus. A, 
<taxonomicName id="06BCE1C3FFE9BA29FE57F835FDEAF82A" ID-CoL="84H2K" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[394,615,1926,1947]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFE9BA29FE57F835FDEAF82A" box="[394,615,1926,1947]" italics="true" pageId="8" pageNumber="899">Orisarma intermedium</emphasis>
</taxonomicName>
, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, 
<taxonomicName id="06BCE1C3FFE9BA29FB65F835FAC2F82A" ID-CoL="84H2K" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[1208,1359,1926,1947]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFE9BA29FB65F835FAC2F82A" box="[1208,1359,1926,1947]" italics="true" pageId="8" pageNumber="899">O. intermedium</emphasis>
</taxonomicName>
, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, 
<taxonomicName id="06BCE1C3FFE9BA29FDFAF810FD33F806" ID-CoL="84H2K" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[551,702,1954,1976]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFE9BA29FDFAF810FD33F806" box="[551,702,1954,1976]" italics="true" pageId="8" pageNumber="899">O. intermedium</emphasis>
</taxonomicName>
, male (27.0 × 25.0 mm) (ZRC 2013.0140), Kyushu, Japan; D, 
<taxonomicName id="06BCE1C3FFE9BA29FAF8F810FA31F806" ID-CoL="84H2K" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[1317,1468,1954,1976]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFE9BA29FAF8F810FA31F806" box="[1317,1468,1954,1976]" italics="true" pageId="8" pageNumber="899">O. intermedium</emphasis>
</taxonomicName>
, male (29.5 × 26.5 mm) (ZRC 2001.0034), Pingtung, Taiwan; E, 
<taxonomicName id="06BCE1C3FFE9BA29FCCDF873FC24F865" ID-CoL="84H2K" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[784,937,1983,2005]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFE9BA29FCCDF873FC24F865" box="[784,937,1983,2005]" italics="true" pageId="8" pageNumber="899">O. intermedium</emphasis>
</taxonomicName>
, male (34.2 × 30.7 mm) (ZRC 2001.0034), Pingtung, Taiwan; F, 
<taxonomicName id="06BCE1C3FFE9BA29FF26F86FFE19F840" ID-CoL="84H2K" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[251,404,2012,2033]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFE9BA29FF26F86FFE19F840" box="[251,404,2012,2033]" italics="true" pageId="8" pageNumber="899">O. intermedium</emphasis>
</taxonomicName>
, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; G, 
<taxonomicName id="06BCE1C3FFE9BA29FBB0F86FFB59F843" ID-CoL="84H2P" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1133,1236,2012,2034]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFE9BA29FBB0F86FFB59F843" box="[1133,1236,2012,2034]" italics="true" pageId="8" pageNumber="899">O. sinense</emphasis>
</taxonomicName>
, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; H, 
<taxonomicName id="06BCE1C3FFE9BA29FDC9F84AFDF4F7BF" ID-CoL="84H2P" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[532,633,2041,2062]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFE9BA29FDC9F84AFDF4F7BF" box="[532,633,2041,2062]" italics="true" pageId="8" pageNumber="899">O. sinense</emphasis>
</taxonomicName>
, paralectotype female (18.8 × 16.9 mm) (MNHN-BP3635b), China; I, 
<taxonomicName id="06BCE1C3FFE9BA29FAFEF84AFA05F7BF" ID-CoL="84H2P" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1315,1416,2041,2062]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFE9BA29FAFEF84AFA05F7BF" box="[1315,1416,2041,2062]" italics="true" pageId="8" pageNumber="899">O. sinense</emphasis>
</taxonomicName>
, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; J, 
<taxonomicName id="06BCE1C3FFE9BA29FD8AF7A5FC8EF79A" ID-CoL="84H2P" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[599,771,2070,2091]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="899" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFE9BA29FD8AF7A5FC8EF79A" box="[599,771,2070,2091]" italics="true" pageId="8" pageNumber="899">Orisarma sinense</emphasis>
</taxonomicName>
, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China.
</paragraph>
</caption>
<paragraph id="C1039A40FFE8BA28FFACFF44FD62FDA1" blockId="9.[113,753,189,960]" pageId="9" pageNumber="900">
<treatmentCitationGroup id="E1ACBD6EFFE8BA28FFACFF44FD62FDA1" pageId="9" pageNumber="900">
<taxonomicName id="06BCE1C3FFE8BA28FFACFF44FE6BFEBD" ID-CoL="7WV46" authorityName="De Haan" authorityYear="1833" box="[113,486,247,268]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir" subGenus="Holometopus">
<emphasis id="F3C84652FFE8BA28FFACFF44FF49FEBD" box="[113,196,247,268]" italics="true" pageId="9" pageNumber="900">Sesarma</emphasis>
(
<emphasis id="F3C84652FFE8BA28FF0EFF44FEDEFEBD" box="[211,339,247,268]" italics="true" pageId="9" pageNumber="900">Holometopus</emphasis>
) 
<emphasis id="F3C84652FFE8BA28FEBFFF44FE6BFEBD" box="[354,486,247,268]" italics="true" pageId="9" pageNumber="900">haematocheir</emphasis>
</taxonomicName>
– 
<treatmentCitation id="401DBC51FFE8BA28FDDDFF44FD26FEBC" author="Tesch JJ" box="[512,683,247,269]" page="156" pageId="9" pageNumber="900" year="1917">
<bibRefCitation id="A52DE7B1FFE8BA28FDDDFF44FD26FEBC" author="Tesch" box="[512,683,247,269]" firstAuthor="Tesch" page="156" pageId="9" pageNumber="900" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch, 1917: 156</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FD6BFF44FF73FE9B" author="Balss H" page="155" pageId="9" pageNumber="900" year="1922">
<bibRefCitation id="A52DE7B1FFE8BA28FD6BFF44FF73FE9B" author="Balss" firstAuthor="Balss" page="155" pageId="9" pageNumber="900" pagination="1" refId="ref67843" refString="Balss H (1922) Ostasiatische Decapoden, IV. Die Brachyrhynchen (Cancridea). Archiv fur Naturgeschichte, 88 A (11): 94 - 165, figs. 1, 2, pls. 1, 2." type="journal article" year="1922">Balss, 1922: 155</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FED6FEA7FE2BFE98" author="Urita T" box="[267,422,276,298]" page="19" pageId="9" pageNumber="900" year="1926">
<bibRefCitation id="A52DE7B1FFE8BA28FED6FEA7FE2BFE98" author="Urita" box="[267,422,276,298]" firstAuthor="Urita" page="19" pageId="9" pageNumber="900" refId="ref78486" refString="Urita T (1926) A check list of Brachyura found in Kagoshima Pref., Japan. The Tsingtao Times, Tsingtao, 40 pp." type="book" year="1926">Urita, 1926: 19</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FE69FEA7FDD7FE98" author="Shen C" box="[436,602,276,298]" page="199" pageId="9" pageNumber="900" year="1932">
<bibRefCitation id="A52DE7B1FFE8BA28FE69FEA7FDD7FE98" author="Shen" box="[436,602,276,298]" firstAuthor="Shen" page="199" pageId="9" pageNumber="900" refId="ref76877" refString="Shen C (1932) The Brachyuran Crustacea of North China. Zoologica Sinica, Series A (Invertebrates of China), Vol. 9, Fasc. 1. The Fan Institute of Biology, Peking [= Beijing], China, x + 321 pp., 171 figs., 10 pls., 1 map." type="book" year="1932">Shen, 1932: 199</bibRefCitation>
</treatmentCitation>
, text-figs. 124, 125, pl. 9 fig. 2; 
<treatmentCitation id="401DBC51FFE8BA28FEE0FE83FE69FEF7" author="Sakai T" box="[317,484,304,326]" page="234" pageId="9" pageNumber="900" year="1936">
<bibRefCitation id="A52DE7B1FFE8BA28FEE0FE83FE69FEF7" author="Sakai" box="[317,484,304,326]" firstAuthor="Sakai" page="234" pageId="9" pageNumber="900" refId="ref75565" refString="Sakai T (1936) Crabs of Japan. 66 plates in life colours with descriptions. Sanseido, Tokyo, 239 pp., 12 pp. [literature cited], 27 pp. [bibliography and index], 66 pls. [dated 1935 but published in 1936]" type="book" year="1936">Sakai, 1936: 234</bibRefCitation>
</treatmentCitation>
, pl. 64, fig. 3; 
<treatmentCitation id="401DBC51FFE8BA28FDAAFE83FF30FED2" author="Sakai T" page="681" pageId="9" pageNumber="900" year="1939">
<bibRefCitation id="A52DE7B1FFE8BA28FDAAFE83FF30FED2" author="Sakai" firstAuthor="Sakai" page="681" pageId="9" pageNumber="900" pagination="1 - 6" refId="ref75618" refString="Sakai T (1939) Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo Co., Ltd., Tokyo. Pp. 365 - 741 + 1 - 23 [= Index], figs. 1 - 129, pls. XLII - CXI, tables 1 - 6." type="book chapter" year="1939">Sakai, 1939: 681</bibRefCitation>
</treatmentCitation>
, pl. 77, fig. 3; 
<treatmentCitation id="401DBC51FFE8BA28FE9AFEFEFE68FED3" author="Shen CJ" box="[327,485,333,355]" page="97" pageId="9" pageNumber="900" year="1940">
<bibRefCitation id="A52DE7B1FFE8BA28FE9AFEFEFE68FED3" author="Shen" box="[327,485,333,355]" firstAuthor="Shen" page="97" pageId="9" pageNumber="900" pagination="211 - 242" refId="ref76939" refString="Shen CJ (1940 a) The brachyuran fauna of Hong Kong. Journal of the Hong Kong Fisheries Research Station, 1: 211 - 242." type="journal article" year="1940" yearSuffix="a">Shen, 1940a: 97</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FE32FEFEFD17FED3" author="Shen C" box="[495,666,333,355]" page="237" pageId="9" pageNumber="900" year="1940">
<bibRefCitation id="A52DE7B1FFE8BA28FE32FEFEFD17FED3" author="Shen" box="[495,666,333,355]" firstAuthor="Shen" page="237" pageId="9" pageNumber="900" pagination="69 - 104" refId="ref76967" refString="Shen C (1940 b) On the collection of crabs of south China. Bulletin of the Fan Memorial Institute of Biology, Zoology Series, Peiping, 10 (2): 69 - 104." type="journal article" year="1940" yearSuffix="b">Shen, 1940b: 237</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FD7EFEFEFF76FECE" author="Kamita T" page="214" pageId="9" pageNumber="900" year="1941">
<bibRefCitation id="A52DE7B1FFE8BA28FD7EFEFEFF76FECE" author="Kamita" firstAuthor="Kamita" page="214" pageId="9" pageNumber="900" refId="ref70869" refString="Kamita T (1941) Studies of the decapod crustaceans of Chosen. Pt. 1. Crabs. The Fisheries Society of Chosen, Keijo, 289 pp., 146 text-figs., 1 map." type="book" year="1941">Kamita, 1941: 214</bibRefCitation>
</treatmentCitation>
, text-fig. 118; 
<treatmentCitation id="401DBC51FFE8BA28FE57FED9FDE0FECE" author="Shen J &amp; Dai A" box="[394,621,362,384]" page="134" pageId="9" pageNumber="900" year="1964">
<bibRefCitation id="A52DE7B1FFE8BA28FE57FED9FDE0FECE" author="Shen &amp; Dai" box="[394,621,362,384]" firstAuthor="Shen" page="134" pageId="9" pageNumber="900" refId="ref77005" refString="Shen J &amp; Dai A (1964) [Illustrations of Animals in China (Crustacea Part II)]. Peking, 142 pp., 277 figs." type="book" year="1964">Shen &amp; Dai, 1964: 134</bibRefCitation>
</treatmentCitation>
, unnumbered fig.; 
<treatmentCitation id="401DBC51FFE8BA28FF18FE34FEE1FE2D" author="Sakai T" box="[197,364,391,413]" page="202" pageId="9" pageNumber="900" year="1965">
<bibRefCitation id="A52DE7B1FFE8BA28FF18FE34FEE1FE2D" author="Sakai" box="[197,364,391,413]" firstAuthor="Sakai" page="202" pageId="9" pageNumber="900" pagination="1 - 92" refId="ref75676" refString="Sakai T (1965) The Crabs of the Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen Co., Tokyo. xvi + 206 pp. [English section], figs. 1 - 27, pls. 1 - 100, pp. 1 - 92 [in Japanese] + 1 - 26 [Bibliography and Index in English] + 27 - 32 [Index in Japanese], 1 map." type="book chapter" year="1965">Sakai, 1965: 202</bibRefCitation>
</treatmentCitation>
, pl. 97, fig. 1; 
<treatmentCitation id="401DBC51FFE8BA28FDDDFE34FD39FE2D" author="Serene R" box="[512,692,391,413]" page="107" pageId="9" pageNumber="900" year="1968">
<bibRefCitation id="A52DE7B1FFE8BA28FDDDFE34FD39FE2D" author="Serene" box="[512,692,391,413]" firstAuthor="Serene" page="107" pageId="9" pageNumber="900" pagination="33 - 112" refId="ref76415" refString="Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a check list of the nonplanctonic marine fauna of South East Asia. Singapore National Academy of Science, Special Publication No. 1: 33 - 112." type="book chapter" year="1968">Serène, 1968: 107</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FD62FE34FF74FE0B" author="Kim HS" page="486" pageId="9" pageNumber="900" year="1973">
<bibRefCitation id="A52DE7B1FFE8BA28FD62FE34FF74FE0B" author="Kim" firstAuthor="Kim" page="486" pageId="9" pageNumber="900" refId="ref70907" refString="Kim HS (1973) Anomura and Brachyura. In: Illustrated Encyclopaedia of Fauna and Flora of Korea. Vol. 14. Ministry of Education, Sam Wha Publishing Co., Republic of Korea, 694 pp., 112 pls." type="book" year="1973">Kim, 1973: 486</bibRefCitation>
</treatmentCitation>
, text-fig. 216, pl. 45 fig. 166; 
<treatmentCitation id="401DBC51FFE8BA28FDCDFE17FD3DFE0B" author="Sakai T" box="[528,688,420,442]" page="655" pageId="9" pageNumber="900" year="1976">
<bibRefCitation id="A52DE7B1FFE8BA28FDCDFE17FD3DFE0B" author="Sakai" box="[528,688,420,442]" firstAuthor="Sakai" page="655" pageId="9" pageNumber="900" pagination="1 - 251" refId="ref75757" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd., Tokyo. In three volumes. xxix + 773 pp. [English Text], 461 pp. [Japanese Text]. Pp. 1 - 16, pls. 1 - 251 [Plates volume]." type="book chapter" year="1976">Sakai, 1976: 655</bibRefCitation>
</treatmentCitation>
(part), pl. 224 fig. 1; 
<bibRefCitation id="A52DE7B1FFE8BA28FEC0FE72FE44FE67" author="Matsuzawa" box="[285,457,449,470]" firstAuthor="Matsuzawa" pageId="9" pageNumber="900" refId="ref72590" refString="Matsuzawa K (1977) Sea Shore Animals of Muroto. Kansai-Insatsu- Kogyi, Japan, unumbered pages, 126 pls." type="book" year="1977">Matsuzawa, 1977</bibRefCitation>
: pl. 110 fig. 1; 
<treatmentCitation id="401DBC51FFE8BA28FD8AFE73FF30FE43" author="Dai AY &amp; Yang SL &amp; Song YZ &amp; Chen GX" page="487" pageId="9" pageNumber="900" year="1986">
<bibRefCitation id="A52DE7B1FFE8BA28FD8AFE73FF30FE43" author="Dai" etAl="et al." firstAuthor="Dai" page="487" pageId="9" pageNumber="900" pagination="1 - 295" refId="ref68573" refString="Dai AY, Yang SL, Song YZ &amp; Chen GX (1986) Crabs of the China Seas. China Ocean Press, Beijing, 17 + 642 pp., figs. 1 - 295, pls. 1 - 74." type="book chapter" year="1986">Dai et al., 1986: 487</bibRefCitation>
</treatmentCitation>
, text-fig. 274(1), pl. 68(6); 
<treatmentCitation id="401DBC51FFE8BA28FE10FE6EFD35FE42" author="Dai AY &amp; Yang SL" box="[461,696,477,499]" page="534" pageId="9" pageNumber="900" year="1991">
<bibRefCitation id="A52DE7B1FFE8BA28FE10FE6EFD35FE42" author="Dai &amp; Yang" box="[461,696,477,499]" firstAuthor="Dai" page="534" pageId="9" pageNumber="900" pagination="1 - 74" refId="ref68539" refString="Dai AY &amp; Yang SL (1991) Crabs of the China Seas. China Ocean Press, Beijing and Berlin, xxi + 681 pp., pls. 1 - 74." type="book chapter" year="1991">Dai &amp; Yang, 1991: 534</bibRefCitation>
</treatmentCitation>
, textfig. 274(1), pl. 68(6); 
<treatmentCitation id="401DBC51FFE8BA28FEB2FE48FDAFFDA1" author="Huang Z-G" box="[367,546,506,528]" page="597" pageId="9" pageNumber="900" year="1994">
<bibRefCitation id="A52DE7B1FFE8BA28FEB2FE48FDAFFDA1" author="Huang" box="[367,546,506,528]" firstAuthor="Huang" page="597" pageId="9" pageNumber="900" pagination="576 - 600" refId="ref70521" refString="Huang Z-G (ed.) (1994) Brachyura. In: Marine species and their distributions in China's Seas. Department of Ocean Management and Monitoring State Oceanic Administration. China Ocean Press, Beijing. Pp. 576 - 600." type="book chapter" year="1994">Huang, 1994: 597</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FDF3FE49FD61FDA1" author="Muraoka K" box="[558,748,506,528]" page="53" pageId="9" pageNumber="900" year="1998">
<bibRefCitation id="A52DE7B1FFE8BA28FDF3FE49FD61FDA1" author="Muraoka" box="[558,748,506,528]" firstAuthor="Muraoka" page="53" pageId="9" pageNumber="900" pagination="1 - 67" refId="ref72999" refString="Muraoka K (1998) Catalogue of the Brachyuran and Anomuran Crabs donated by Prof. Dr. Tune Sakai to the Kanagawa Prefectural Museum. Catalogue of the Collection in the Kanagawa Prefectural Museum of Natural History, Odawara, Kanagawa, Japan, No. 11: 1 - 67." type="book chapter" year="1998">Muraoka, 1998: 53</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFE8BA28FFACFDA4FD19FD9D" blockId="9.[113,753,189,960]" box="[113,660,535,557]" pageId="9" pageNumber="900">
<treatmentCitationGroup id="E1ACBD6EFFE8BA28FFACFDA4FD19FD9D" box="[113,660,535,557]" pageId="9" pageNumber="900">
<taxonomicName id="06BCE1C3FFE8BA28FFACFDA4FEE3FD9D" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[113,366,535,556]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematochir">
<emphasis id="F3C84652FFE8BA28FFACFDA4FEE3FD9D" box="[113,366,535,556]" italics="true" pageId="9" pageNumber="900">Chiromantes haematochir</emphasis>
</taxonomicName>
– 
<treatmentCitation id="401DBC51FFE8BA28FE57FDA4FDD0FD9D" author="Ng PKL &amp; Liu HC" box="[394,605,535,557]" page="229" pageId="9" pageNumber="900" year="1999">
<bibRefCitation id="A52DE7B1FFE8BA28FE57FDA4FDD0FD9D" author="Ng PKL &amp; Liu HC" box="[394,605,535,557]" firstAuthor="Liu" journalOrPublisher="Zoological Studies" pageId="9" pageNumber="900" pagination="228 - 237" part="38" refId="ref73911" refString="Ng PKL &amp; Liu HC (1999) The taxonomy of Sesarma tangi Rathbun, 1931 and S. stormi De Man, 1895 (Crustacea: Decapoda: Brachyura: Grapsidae: Sesarminae), with establishment of a new genus for S. stormi. Zoological Studies, 38 (2): 228 - 237." title="The taxonomy of Sesarma tangi Rathbun, 1931 and S. stormi De Man, 1895 (Crustacea: Decapoda: Brachyura: Grapsidae: Sesarminae), with establishment of a new genus for S. stormi" type="journal article" year="1999">Liu &amp; Ng, 1999: 229</bibRefCitation>
</treatmentCitation>
, 230.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFE8BA28FFACFD87FE3EFCD8" blockId="9.[113,753,189,960]" pageId="9" pageNumber="900">
<treatmentCitationGroup id="E1ACBD6EFFE8BA28FFACFD87FE3EFCD8" pageId="9" pageNumber="900">
<taxonomicName id="06BCE1C3FFE8BA28FFACFD87FE1FFDF8" ID-CoL="69T63" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[113,402,564,585]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE8BA28FFACFD87FE1FFDF8" box="[113,402,564,585]" italics="true" pageId="9" pageNumber="900">Chiromantes haematocheir</emphasis>
</taxonomicName>
– 
<treatmentCitation id="401DBC51FFE8BA28FE68FD87FD09FDF8" author="Miyake S" box="[437,644,564,586]" page="179" pageId="9" pageNumber="900" year="1983">
<bibRefCitation id="A52DE7B1FFE8BA28FE68FD87FD09FDF8" author="Miyake" box="[437,644,564,586]" firstAuthor="Miyake" page="179" pageId="9" pageNumber="900" refId="ref72931" refString="Miyake S (1983) Japanese Crustacean Decapods and Stomatopods in Color, vol. II. Brachyura (Crabs). First Edition. Hoikusha, Osaka, viii + 277 pp., 64 pls., unnumbered figs." type="book" year="1983">Miyake, 1983: 179</bibRefCitation>
</treatmentCitation>
, pl. 60-1; 
<treatmentCitation id="401DBC51FFE8BA28FF45FDE3FEFFFDD7" author="Minemizu R" box="[152,370,592,614]" page="297" pageId="9" pageNumber="900" year="2000">
<bibRefCitation id="A52DE7B1FFE8BA28FF45FDE3FEFFFDD7" author="Minemizu" box="[152,370,592,614]" firstAuthor="Minemizu" page="297" pageId="9" pageNumber="900" refId="ref72906" refString="Minemizu R (2000) [Marine decapod and stomatopod crustaceans mainly from Japan]. Bun-Ichi Sogo Shuppan, Tokyo, 344 pp." type="book" year="2000">Minemizu, 2000: 297</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FEA2FDE3FDD6FDD7" author="Kobayashi S" box="[383,603,592,614]" page="122" pageId="9" pageNumber="900" year="2000">
<bibRefCitation id="A52DE7B1FFE8BA28FEA2FDE3FDD6FDD7" author="Kobayashi" box="[383,603,592,614]" firstAuthor="Kobayashi" page="122" pageId="9" pageNumber="900" pagination="113 - 130" refId="ref70994" refString="Kobayashi S (2000) Distribution pattern and ecology of brachyuran crabs in the riverine environment: their significance in the ecosystem and present condition. Ecological Civil Engineering, 3 (1): 113 - 130." type="journal article" year="2000">Kobayashi, 2000: 122</bibRefCitation>
</treatmentCitation>
, fig. 2p; 
<treatmentCitation id="401DBC51FFE8BA28FD64FDE2FE9DFD33" author="Ng PKL &amp; Wang C-H &amp; Ho P-H &amp; Shih H-T" page="41" pageId="9" pageNumber="900" year="2001">
<bibRefCitation id="A52DE7B1FFE8BA28FD64FDE2FE9DFD33" author="Ng" etAl="et al." firstAuthor="Ng" page="41" pageId="9" pageNumber="900" pagination="1 - 86" refId="ref74237" refString="Ng PKL, Wang C-H, Ho P-H &amp; Shih H-T (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, 11: 1 - 86, 8 colour pls." type="journal article" year="2001">Ng et al., 2001: 41</bibRefCitation>
</treatmentCitation>
[includes references for this species from 
<collectingCountry id="B9ABDAD0FFE8BA28FD75FDDEFD7DFD32" box="[680,752,621,643]" name="Taiwan" pageId="9" pageNumber="900">Taiwan</collectingCountry>
up to 2001]; 
<treatmentCitation id="401DBC51FFE8BA28FEC1FD39FD98FD2E" author="Schubart CD, Cuesta JA &amp; Felder DL" box="[284,533,650,672]" page="30" pageId="9" pageNumber="900" year="2002">
<bibRefCitation id="A52DE7B1FFE8BA28FEC1FD39FD98FD2E" author="Schubart" box="[284,533,650,672]" etAl="et al." firstAuthor="Schubart" journalOrPublisher="Journal of Crustacean Biology" page="30" pageId="9" pageNumber="900" pagination="28 - 44" part="22" refId="ref76061" refString="Schubart CD, Cuesta JA &amp; Felder DL (2002) Glyptograpsidae, a new brachyuran family from Central America: larval and adult morphology, and a molecular phylogeny of the Grapsoidea. Journal of Crustacean Biology, 22 (1): 28 - 44." title="Glyptograpsidae, a new brachyuran family from Central America: larval and adult morphology, and a molecular phylogeny of the Grapsoidea" type="journal article" year="2002">Schubart et al., 2002: 30</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FDFFFD38FD24FD2E" author="Ho P-H" box="[546,681,650,672]" page="38" pageId="9" pageNumber="900" year="2003">
<bibRefCitation id="A52DE7B1FFE8BA28FDFFFD38FD24FD2E" author="Ho" box="[546,681,650,672]" firstAuthor="Ho" page="38" pageId="9" pageNumber="900" refId="ref70247" refString="Ho P-H (2003) Land crabs in the coastal forest of Kenting National Park. Kenting National Park, Pingtung, 47 pp." type="book" year="2003">Ho, 2003: 38</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FD68FD39FEB7FD0D" author="Kwok WPW &amp; Tang W-S" page="3" pageId="9" pageNumber="900" year="2005">
<bibRefCitation id="A52DE7B1FFE8BA28FD68FD39FEB7FD0D" author="Kwok &amp; Tang" firstAuthor="Kwok" page="3" pageId="9" pageNumber="900" pagination="1 - 6" refId="ref71111" refString="Kwok WPW &amp; Tang W-S (2005) An Introduction to Common Sesarmine Crabs of Hong Kong. Hong Kong Biodiversity, Agriculture, Fisheries and Conservation Department Newsletter, 9: 1 - 6." type="journal article" year="2005">Kwok &amp; Tang, 2005: 3</bibRefCitation>
</treatmentCitation>
, fig. 8; 
<treatmentCitation id="401DBC51FFE8BA28FE5AFD14FD08FD0C" author="Schubart CD &amp; Cannicci S &amp; Vannini M &amp; Fratini S" box="[391,645,679,701]" page="195" pageId="9" pageNumber="900" year="2006">
<bibRefCitation id="A52DE7B1FFE8BA28FE5AFD14FD08FD0C" author="Schubart" box="[391,645,679,701]" etAl="et al." firstAuthor="Schubart" page="195" pageId="9" pageNumber="900" pagination="193 - 199" refId="ref75958" refString="Schubart CD, Cannicci S, Vannini M &amp; Fratini S (2006) Molecular phylogeny of grapsoid crabs and allies based on two mitochondrial genes and a proposal for refraining from current superfamily classification. Journal of Zoological Systematics and Evolutionary Research, 44 (3): 193 - 199." type="journal article" year="2006">Schubart et al., 2006: 195</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FD4DFD14FEDEFD68" author="Takeda M &amp; Ueshima R" page="94" pageId="9" pageNumber="900" year="2006">
<bibRefCitation id="A52DE7B1FFE8BA28FD4DFD14FEDEFD68" author="Takeda &amp; Ueshima" firstAuthor="Takeda" page="94" pageId="9" pageNumber="900" pagination="61 - 105" refId="ref77817" refString="Takeda M &amp; Ueshima R (2006) Crabs preserved in the Department of Zoology, the University Museum, the University of Tokyo. In: Ueshima R (ed.) Catalogue of Invertebrate Collection Deposited in the Department of Zoology, The University Museum, the University of Tokyo. The University Museum, the University of Tokyo Materials Reports, 62: 61 - 105. [In Japanese]" type="journal article" year="2006">Takeda &amp; Ueshima, 2006: 94</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FE82FD77FD83FD68" author="Oh SM &amp; Chung DE &amp; Ko HS" box="[351,526,708,730]" page="9" pageId="9" pageNumber="900" year="2007">
<bibRefCitation id="A52DE7B1FFE8BA28FE82FD77FD83FD68" author="Oh" box="[351,526,708,730]" etAl="et al." firstAuthor="Oh" page="9" pageId="9" pageNumber="900" pagination="9 - 20" refId="ref74361" refString="Oh SM, Chung DE &amp; Ko HS (2007) Complete larval development of Chiromantes haematocheir (Crustacea: Decapoda: Brachyura: Grapsidae), with a key to seven sesarminid zoeas in Korea. Korean Journal of Systematics and Zoology, 23 (1): 9 - 20." type="journal article" year="2007">Oh et al., 2007: 9</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FDC7FD77FD61FD68" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[538,748,708,730]" page="220" pageId="9" pageNumber="900" year="2008">
<bibRefCitation id="A52DE7B1FFE8BA28FDC7FD77FD61FD68" author="Ng" box="[538,748,708,730]" etAl="et al." firstAuthor="Ng" page="220" pageId="9" pageNumber="900" pagination="1 - 286" refId="ref73754" refString="Ng PKL, Guinot D &amp; Davie PJF (2008 a) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008" yearSuffix="a">Ng et al., 2008a: 220</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FF4AFD52FE0CFD47" author="Yang S &amp; Chen H &amp; Jiang W" box="[151,385,736,758]" page="801" pageId="9" pageNumber="900" year="2008">
<bibRefCitation id="A52DE7B1FFE8BA28FF4AFD52FE0CFD47" author="Yang" box="[151,385,736,758]" etAl="et al." firstAuthor="Yang" page="801" pageId="9" pageNumber="900" pagination="761 - 810" refId="ref78808" refString="Yang S, Chen H &amp; Jiang W (2008) Crustacea: Decapoda Brachyura. In: Liu R (ed.) Checklist of Marine Biota of China Seas. Institute of Oceanology, Chinese Academy of Sciences. Pp. 761 - 810." type="book chapter" year="2008">Yang et al., 2008: 801</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FE52FD52FDD0FD47" author="Naruse T &amp; Ng PKL" box="[399,605,737,758]" pageId="9" pageNumber="900" year="2008">
<bibRefCitation id="A52DE7B1FFE8BA28FE52FD52FDD0FD47" author="Naruse &amp; Ng" box="[399,605,737,758]" firstAuthor="Naruse" pageId="9" pageNumber="900" pagination="1 - 13" refId="ref73402" refString="Naruse T &amp; Ng PKL (2008) A new species of Chiromantes s. str. (Decapoda: Brachyura: Sesarmidae) from the Ryukyu Islands, Japan, with a note on the identity of Holometopus serenei Soh, 1978. Crustacean Research, 37: 1 - 13." type="journal article" year="2008">Naruse &amp; Ng, 2008</bibRefCitation>
</treatmentCitation>
: figs. 1–4, 5b; 
<treatmentCitation id="401DBC51FFE8BA28FF4AFD4DFEBFFCA3" author="Lee J-H" box="[151,306,765,787]" page="134" pageId="9" pageNumber="900" year="2008">
<bibRefCitation id="A52DE7B1FFE8BA28FF4AFD4DFEBFFCA3" author="Lee" box="[151,306,765,787]" firstAuthor="Lee" page="134" pageId="9" pageNumber="900" refId="ref71440" refString="Lee J-H (2008) A Field Guide to Crabs in Taiwan. Bigtrees Company, Taipei, 174 pp." type="book" year="2008">Lee, 2008: 134</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FEE2FD4EFDF3FCA2" author="Naderloo R &amp; Schubart CD" box="[319,638,765,787]" page="61" pageId="9" pageNumber="900" year="2009">
<bibRefCitation id="A52DE7B1FFE8BA28FEE2FD4EFDF3FCA2" author="Naderloo &amp; Schubart" box="[319,638,765,787]" firstAuthor="Naderloo" page="61" pageId="9" pageNumber="900" pagination="61 - 68" refId="ref73102" refString="Naderloo R &amp; Schubart CD (2009) Redescription and mitochondrial identification of Chiromantes boulengeri (Calman, 1920) (Decapoda: Brachyura: Sesarmidae) based on fresh material from the Persian Gulf, Iran. Zootaxa, 2128: 61 - 68." type="journal article" year="2009">Naderloo &amp; Schubart, 2009: 61</bibRefCitation>
</treatmentCitation>
, 67; 
<treatmentCitation id="401DBC51FFE8BA28FD6EFD4EFEB4FC81" author="Liu H-C &amp; Wang C-H" page="58" pageId="9" pageNumber="900" year="2010">
<bibRefCitation id="A52DE7B1FFE8BA28FD6EFD4EFEB4FC81" author="Liu &amp; Wang" firstAuthor="Liu" page="58" pageId="9" pageNumber="900" refId="ref71886" refString="Liu H-C &amp; Wang C-H (2010) Taiwan Coastal Wetland Crabs. Wild Bird Society of Taipei, Taipei, 79 pp." type="book" year="2010">Liu &amp; Wang, 2010: 58</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FE98FCA9FDAFFC9E" author="Komai T &amp; Ng PKL" box="[325,546,794,816]" page="4" pageId="9" pageNumber="900" year="2013">
<bibRefCitation id="A52DE7B1FFE8BA28FE98FCA9FDAFFC9E" author="Komai &amp; Ng" box="[325,546,794,816]" firstAuthor="Komai" page="4" pageId="9" pageNumber="900" pagination="539 - 551" refId="ref71033" refString="Komai T &amp; Ng PKL (2013) A new species of sesarmid crab of the genus Chiromantes (Crustacea: Decapoda: Brachyura) from the Ogasawara Islands, Japan. Zootaxa, 3681 (5): 539 - 551." type="journal article" year="2013">Komai &amp; Ng, 2013: 4</bibRefCitation>
</treatmentCitation>
, figs. 1–3, 4a, b, 5b; 
<treatmentCitation id="401DBC51FFE8BA28FF4AFC84FE2FFCFD" author="Toyota K &amp; Seki S" box="[151,418,823,845]" page="188" pageId="9" pageNumber="900" year="2014">
<bibRefCitation id="A52DE7B1FFE8BA28FF4AFC84FE2FFCFD" author="Toyota &amp; Seki" box="[151,418,823,845]" firstAuthor="Toyota" page="188" pageId="9" pageNumber="900" refId="ref78208" refString="Toyota K &amp; Seki S (2014) Shrimps and Crabs. Nature Watching Guidebook, Freshwater Shrimps and Crabs in Japan. Seibundo Shinkosha Publishing, 256 pp." type="book" year="2014">Toyota &amp; Seki, 2014: 188</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFE8BA28FE72FC84FD05FCFD" author="Ng PKL &amp; Shih H-T &amp; Ho P-H &amp; Wang C-H" box="[431,648,823,845]" page="103" pageId="9" pageNumber="900" year="2017">
<bibRefCitation id="A52DE7B1FFE8BA28FE72FC84FD05FCFD" author="Ng" box="[431,648,823,845]" etAl="et al." firstAuthor="Ng" page="103" pageId="9" pageNumber="900" pagination="1 - 185" refId="ref74189" refString="Ng PKL, Shih H-T, Ho P-H &amp; Wang C-H (2017 b) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70 (3 - 4): 1 - 185." type="journal article" year="2017" yearSuffix="b">Ng et al., 2017b: 103</bibRefCitation>
</treatmentCitation>
[for other local references for 
<collectingCountry id="B9ABDAD0FFE8BA28FE81FCE0FE24FCD8" box="[348,425,851,873]" name="Taiwan" pageId="9" pageNumber="900">Taiwan</collectingCountry>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFE8BA28FFACFCC3FE97FC13" blockId="9.[113,753,189,960]" pageId="9" pageNumber="900">
<treatmentCitationGroup id="E1ACBD6EFFE8BA28FFACFCC3FE97FC13" pageId="9" pageNumber="900">
<treatmentCitation id="401DBC51FFE8BA28FFACFCC3FE52FC37" author="Soh CL" box="[113,479,880,902]" page="13" pageId="9" pageNumber="900" year="1978">
<taxonomicName id="06BCE1C3FFE8BA28FFACFCC3FE52FC37" ID-CoL="6LXY9" authority="Soh, 1978: 13" authorityName="Soh" authorityPageNumber="13" authorityYear="1978" box="[113,479,880,902]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFE8BA28FFACFCC3FEC8FC37" box="[113,325,880,902]" italics="true" pageId="9" pageNumber="900">Holometopus serenei</emphasis>
<bibRefCitation id="A52DE7B1FFE8BA28FE93FCC3FE52FC37" author="Soh" box="[334,479,880,902]" firstAuthor="Soh" page="13" pageId="9" pageNumber="900" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh, 1978: 13</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. c, d, pl. 1b, e, pl. 2f; 
<treatmentCitation id="401DBC51FFE8BA28FF4AFC3DFE9BFC13" author="Lee SY" box="[151,278,909,931]" page="3" pageId="9" pageNumber="900" year="1995">
<bibRefCitation id="A52DE7B1FFE8BA28FF4AFC3DFE9BFC13" author="Lee" box="[151,278,909,931]" firstAuthor="Lee" page="3" pageId="9" pageNumber="900" pagination="3" refId="ref71500" refString="Lee SY (1995) Endemic sesarmine crabs rediscovered. Porcupine, 12: 3." type="journal article" year="1995">Lee, 1995: 3</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFE8BA28FFACFC19FD0CFC0E" blockId="9.[113,753,189,960]" box="[113,641,938,960]" pageId="9" pageNumber="900">
<treatmentCitationGroup id="E1ACBD6EFFE8BA28FFACFC19FD0CFC0E" box="[113,641,938,960]" pageId="9" pageNumber="900">
<taxonomicName id="06BCE1C3FFE8BA28FFACFC19FEB5FC71" ID-CoL="6LXY9" authority="(Soh, 1978)" box="[113,312,938,960]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">Chiromantes serenei</taxonomicName>
– 
<treatmentCitation id="401DBC51FFE8BA28FE8EFC19FDB6FC0E" author="Kwok WPW &amp; Tang W-S" box="[339,571,938,960]" page="3" pageId="9" pageNumber="900" year="2005">
<bibRefCitation id="A52DE7B1FFE8BA28FE8EFC19FDB6FC0E" author="Kwok &amp; Tang" box="[339,571,938,960]" firstAuthor="Kwok" page="3" pageId="9" pageNumber="900" pagination="1 - 6" refId="ref71111" refString="Kwok WPW &amp; Tang W-S (2005) An Introduction to Common Sesarmine Crabs of Hong Kong. Hong Kong Biodiversity, Agriculture, Fisheries and Conservation Department Newsletter, 9: 1 - 6." type="journal article" year="2005">Kwok &amp; Tang, 2005: 3</bibRefCitation>
</treatmentCitation>
, fig. 9.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFE8BA28FFACFC5BFB78FC84" pageId="9" pageNumber="900" type="materials_examined">
<paragraph id="C1039A40FFE8BA28FFACFC5BFB78FC84" blockId="9.[113,753,1000,2113]" lastBlockId="9.[813,1452,189,822]" pageId="9" pageNumber="900">
<emphasis id="F3C84652FFE8BA28FFACFC5BFEC2FBB1" bold="true" box="[113,335,1000,1024]" pageId="9" pageNumber="900">Material examined</emphasis>
. 
<materialsCitation id="71D4901DFFE8BA28FE82FC5AFDBBFB91" ID-GBIF-Occurrence="3763237516" collectionCode="RMNH" collectorName="Japan &amp; Ph. v. Siebold." pageId="9" pageNumber="900" specimenCode="D160" specimenCount="1" typeStatus="lectotype">
<typeStatus id="1E0724E2FFE8BA28FE82FC5AFE42FBB0" box="[351,463,1001,1025]" pageId="9" pageNumber="900" type="lectotype">Lectotype</typeStatus>
: male (33.7 × 
<quantity id="064437A5FFE8BA28FDAAFC5BFD64FBB1" box="[631,745,1000,1024]" metricMagnitude="-2" metricUnit="m" metricValue="2.965" pageId="9" pageNumber="900" unit="mm" value="29.65">29.65 mm</quantity>
) (
<collectionCode id="A7AD0285FFE8BA28FFA4FBBBFF46FB91" box="[121,203,1032,1056]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="900" type="Museum">RMNH</collectionCode>
<specimenCode id="911A323BFFE8BA28FF0DFBBBFE83FB91" box="[208,270,1032,1056]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="900" type="Museum">D160</specimenCode>
), 
<collectorName id="6C49FF96FFE8BA28FEFDFBBAFED2FB91" box="[288,351,1033,1056]" pageId="9" pageNumber="900">Japan</collectorName>
, coll. 
<collectorName id="6C49FF96FFE8BA28FE41FBBBFDBBFB91" box="[412,566,1032,1056]" pageId="9" pageNumber="900">Ph. v. Siebold.</collectorName>
</materialsCitation>
<materialsCitation id="71D4901DFFE8BA28FDE6FBBBFEF6FBD1" ID-GBIF-Occurrence="3763237424" collectionCode="RMNH" collectorName="Ph. F. von Siebold" location="Japan" pageId="9" pageNumber="900" specimenCode="D 158" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="1E0724E2FFE8BA28FDE6FBBBFD57FB91" box="[571,730,1032,1056]" pageId="9" pageNumber="900" type="paralectotype">Paralectotypes</typeStatus>
: 
<specimenCount id="D7BA51C9FFE8BA28FD3EFBBBFF37FBF1" pageId="9" pageNumber="900" type="female">1 female</specimenCount>
(27.5 × 
<quantity id="064437A5FFE8BA28FECFFB9BFEFFFBF1" box="[274,370,1064,1088]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="9" pageNumber="900" unit="mm" value="23.5">23.5 mm</quantity>
) (
<collectionCode id="A7AD0285FFE8BA28FE5BFB9BFE55FBF1" box="[390,472,1064,1088]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="900" type="Museum">RMNH</collectionCode>
<specimenCode id="911A323BFFE8BA28FE03FB9AFDAFFBF1" box="[478,546,1064,1088]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="900" type="Museum">D 158</specimenCode>
), 
<location id="C463CC9BFFE8BA28FDE9FB9AFDFEFBF1" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FDE9FB9AFDFEFBF1" box="[564,627,1065,1088]" name="Japan" pageId="9" pageNumber="900">Japan</location>
, coll. 
<collectorName id="6C49FF96FFE8BA28FD6FFB9BFF7EFBD1" pageId="9" pageNumber="900">Ph. F. von Siebold</collectorName>
, 1823–1829;
</materialsCitation>
<materialsCitation id="71D4901DFFE8BA28FE5CFBFBFD1BFB11" ID-GBIF-Occurrence="3763237358" collectionCode="RMNH" collectorName="Ph. F. von Siebold" location="Japan" pageId="9" pageNumber="900" specimenCode="D 158" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="1E0724E2FFE8BA28FE5CFBFBFD93FBD1" box="[385,542,1096,1120]" pageId="9" pageNumber="900" type="paralectotype">paralectotypes</typeStatus>
, 
<specimenCount id="D7BA51C9FFE8BA28FDFAFBFBFDFAFBD1" box="[551,631,1096,1120]" pageId="9" pageNumber="900" type="male">4 males</specimenCount>
(16.9–28.9 × 14.8 × 
<quantity id="064437A5FFE8BA28FF0BFBDBFEB7FB31" box="[214,314,1128,1152]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="9" pageNumber="900" unit="mm" value="25.3">25.3 mm</quantity>
), 
<specimenCount id="D7BA51C9FFE8BA28FE93FBDBFE21FB31" box="[334,428,1128,1152]" pageId="9" pageNumber="900" type="female">1 female</specimenCount>
(26.7 × 22.0 mm) (
<collectionCode id="A7AD0285FFE8BA28FD5EFBDBFD58FB31" box="[643,725,1128,1152]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="900" type="Museum">RMNH</collectionCode>
<specimenCode id="911A323BFFE8BA28FD00FBDAFF11FB11" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="900" type="Museum">D 159</specimenCode>
), 
<location id="C463CC9BFFE8BA28FF6EFB3AFF7EFB11" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FF6EFB3AFF7EFB11" box="[179,243,1161,1184]" name="Japan" pageId="9" pageNumber="900">Japan</location>
, coll. 
<collectorName id="6C49FF96FFE8BA28FEE7FB3BFD81FB11" box="[314,524,1160,1184]" pageId="9" pageNumber="900">Ph. F. von Siebold</collectorName>
, 1823–1829
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FD79FB3BFE0CFB51" ID-GBIF-Occurrence="3763237456" collectionCode="MNHN" collectorName="Ph. F. von Siebold &amp; Others" country="Japan" location="Japan" pageId="9" pageNumber="900" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FD79FB3BFD7DFB11" box="[676,752,1160,1184]" pageId="9" pageNumber="900" type="male">1 male</specimenCount>
(29.6 × 
<quantity id="064437A5FFE8BA28FF13FB1BFEB9FB71" box="[206,308,1192,1216]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="9" pageNumber="900" unit="mm" value="25.3">25.3 mm</quantity>
) (
<collectionCode id="A7AD0285FFE8BA28FE91FB1BFE2DFB71" box="[332,416,1192,1216]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="900">MNHN</collectionCode>
B-12475), 
<location id="C463CC9BFFE8BA28FDF9FB1AFDE9FB71" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FDF9FB1AFDE9FB71" box="[548,612,1193,1216]" country="Japan" name="Japan" pageId="9" pageNumber="900">Japan</location>
, coll. 
<collectorName id="6C49FF96FFE8BA28FD70FB1BFF78FB51" pageId="9" pageNumber="900">Ph. F. von Siebold</collectorName>
, 1823–1829.
</materialsCitation>
<materialsCitation id="71D4901DFFE8BA28FE55FB7BFD55FAB1" ID-GBIF-Occurrence="3763237461" collectionCode="MNHN" collectorName="Ph. F. von Siebold &amp; Others" country="Japan" location="Japan" pageId="9" pageNumber="900" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<collectorName id="6C49FF96FFE8BA28FE55FB7BFE5EFB51" box="[392,467,1224,1248]" pageId="9" pageNumber="900">Others</collectorName>
: 
<collectingCountry id="B9ABDAD0FFE8BA28FE03FB7BFDBCFB51" box="[478,561,1224,1248]" name="Japan" pageId="9" pageNumber="900">JAPAN</collectingCountry>
– 
<specimenCount id="D7BA51C9FFE8BA28FD93FB7BFD2CFB51" box="[590,673,1224,1248]" pageId="9" pageNumber="900" type="male">2 males</specimenCount>
(larger 31.7 × 
<quantity id="064437A5FFE8BA28FF63FB5BFE92FAB1" box="[190,287,1256,1280]" metricMagnitude="-2" metricUnit="m" metricValue="2.7399999999999998" pageId="9" pageNumber="900" unit="mm" value="27.4">27.4 mm</quantity>
) (
<collectionCode id="A7AD0285FFE8BA28FEE8FB5BFEE6FAB1" box="[309,363,1256,1280]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC</collectionCode>
1964.9.8.12–13), coll. 
<collectorName id="6C49FF96FFE8BA28FDBBFB5BFD55FAB1" box="[614,728,1256,1280]" pageId="9" pageNumber="900">S. Miyake</collectorName>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FD3EFB5BFD66FAF1" ID-GBIF-Occurrence="3763237570" collectionCode="ZRC" collectorName="T. Sakai" country="Japan" location="Kamakura" municipality="Sagami Bay" pageId="9" pageNumber="900" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FD3EFB5BFF28FA91" pageId="9" pageNumber="900" type="male">1 male</specimenCount>
(26.4 × 
<quantity id="064437A5FFE8BA28FEDAFABBFEE6FA91" box="[263,363,1288,1312]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="9" pageNumber="900" unit="mm" value="22.9">22.9 mm</quantity>
), 
<specimenCount id="D7BA51C9FFE8BA28FE5DFABBFE6DFA91" box="[384,480,1288,1312]" pageId="9" pageNumber="900" type="female">1 female</specimenCount>
(26.6 × 
<quantity id="064437A5FFE8BA28FD9DFABBFD2EFA91" box="[576,675,1288,1312]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="9" pageNumber="900" unit="mm" value="22.4">22.4 mm</quantity>
) (
<collectionCode id="A7AD0285FFE8BA28FD66FABBFD7CFA91" box="[699,753,1288,1312]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC</collectionCode>
1970.8.27.7), 
<collectingMunicipality id="2167003AFFE8BA28FEDAFA9BFE01FAF0" box="[263,396,1320,1345]" pageId="9" pageNumber="900">Sagami Bay</collectingMunicipality>
, 
<location id="C463CC9BFFE8BA28FE45FA9BFD87FAF1" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FE45FA9BFD87FAF1" box="[408,522,1320,1344]" country="Japan" municipality="Sagami Bay" name="Kamakura" pageId="9" pageNumber="900">Kamakura</location>
, coll. 
<collectorName id="6C49FF96FFE8BA28FD96FA9AFD25FAF1" box="[587,680,1320,1344]" pageId="9" pageNumber="900">T. Sakai</collectorName>
, 1968
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FFACFAFAFF26FA71" ID-GBIF-Occurrence="3763237407" collectingDate="2000-08-30" collectionCode="ZRC" collectorName="H. Suzuki" country="Japan" location="Satsuma Peninsula" municipality="Izaku River" pageId="9" pageNumber="900" specimenCode="ZRC 2002.0225" specimenCount="7" specimenCount-female="7" stateProvince="Kagoshima" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FFACFAFAFF50FAD1" box="[113,221,1352,1376]" pageId="9" pageNumber="900" type="female">7 females</specimenCount>
(smallest 18.5 × 
<quantity id="064437A5FFE8BA28FE7AFAFBFD80FAD1" box="[423,525,1352,1376]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="9" pageNumber="900" unit="mm" value="15.6">15.6 mm</quantity>
, largest 28.3 × 
<quantity id="064437A5FFE8BA28FD1CFAFBFF10FA31" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="9" pageNumber="900" unit="mm" value="24.5">24.5 mm</quantity>
) (
<specimenCode id="911A323BFFE8BA28FF6EFADBFEEAFA31" box="[179,359,1384,1408]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC 2002.0225</specimenCode>
), river mouth, 
<collectingMunicipality id="2167003AFFE8BA28FDD5FADBFD07FA31" box="[520,650,1384,1408]" pageId="9" pageNumber="900">Izaku River</collectingMunicipality>
, 
<location id="C463CC9BFFE8BA28FD4BFADBFF56FA11" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FD4BFADBFF56FA11" country="Japan" municipality="Izaku River" name="Satsuma Peninsula" pageId="9" pageNumber="900" stateProvince="Kagoshima">Satsuma Peninsula</location>
, 
<collectingRegion id="037854A2FFE8BA28FF39FA3BFE5EFA11" box="[228,467,1416,1440]" country="Japan" name="Kagoshima" pageId="9" pageNumber="900">Kagoshima Prefecture</collectingRegion>
, coll. 
<collectorName id="6C49FF96FFE8BA28FDD2FA3AFDF7FA11" box="[527,634,1416,1440]" pageId="9" pageNumber="900">H. Suzuki</collectorName>
, 
<date id="B502BC80FFE8BA28FD59FA3BFF26FA71" pageId="9" pageNumber="900" value="2000-08-30">
<collectingDate id="A5464568FFE8BA28FD59FA3BFF26FA71" pageId="9" pageNumber="900" value="2000-08-30">30 August 2000</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FF67FA1BFE5FF991" ID-GBIF-Occurrence="3763237365" collectingDate="2017-10-30" collectionCode="ZRC" collectorName="P.K.L. Ng, T. Naruse, M. Deki et al" country="Japan" county="Kamenoko-jima" elevation="50" latitude="33.125942" location="Kyushu" longLatPrecision="1" longitude="129.67834" municipality="Sasebo" pageId="9" pageNumber="900" specimenCode="ZRC 2017.1004" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kagoshima" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FF67FA1BFE86FA71" box="[186,267,1448,1472]" pageId="9" pageNumber="900" type="male">1 male</specimenCount>
, 
<specimenCount id="D7BA51C9FFE8BA28FEC7FA1BFEF0FA71" box="[282,381,1448,1472]" pageId="9" pageNumber="900" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFE8BA28FE53FA1BFDC4FA71" box="[398,585,1448,1472]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC 2017.1004</specimenCode>
), stream about 
<quantity id="064437A5FFE8BA28FFAFFA7BFF21FA51" box="[114,172,1480,1504]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="900" unit="m" value="50.0">
<elevation id="4A917D73FFE8BA28FFAFFA7BFF21FA51" box="[114,172,1480,1504]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="900" unit="m" value="50.0">50 m</elevation>
</quantity>
before coastline, 
<collectingCounty id="2862E2CCFFE8BA28FEAAFA7BFDA7FA51" box="[375,554,1480,1504]" pageId="9" pageNumber="900">Kamenoko-jima</collectingCounty>
, 
<collectingMunicipality id="2167003AFFE8BA28FDE5FA7BFD0AFA51" box="[568,647,1480,1504]" pageId="9" pageNumber="900">Sasebo</collectingMunicipality>
, 
<location id="C463CC9BFFE8BA28FD4BFA7BFD60FA51" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FD4BFA7BFD60FA51" box="[662,749,1480,1504]" country="Japan" county="Kamenoko-jima" latitude="33.125942" longLatPrecision="1" longitude="129.67834" municipality="Sasebo" name="Kyushu" pageId="9" pageNumber="900" stateProvince="Kagoshima">Kyushu</location>
, 
<geoCoordinate id="A488FC87FFE8BA28FFAFFA5BFE89F9B1" box="[114,260,1512,1536]" degrees="33.125941" direction="north" orientation="latitude" pageId="9" pageNumber="900" precision="1" value="33.125942">33.125941°N</geoCoordinate>
<geoCoordinate id="A488FC87FFE8BA28FED3FA5BFE20F9B1" box="[270,429,1512,1536]" degrees="129.678348" direction="east" orientation="longitude" pageId="9" pageNumber="900" precision="1" value="129.67834">129.678348°E</geoCoordinate>
, coll. 
<collectorName id="6C49FF96FFE8BA28FE2BFA5AFDE0F9B0" box="[502,621,1513,1537]" pageId="9" pageNumber="900">P.K.L. Ng</collectorName>
, 
<collectorName id="6C49FF96FFE8BA28FDA6FA5AFD60F9B1" box="[635,749,1513,1536]" pageId="9" pageNumber="900">T. Naruse</collectorName>
, 
<collectorName id="6C49FF96FFE8BA28FFAFF9BAFF41F991" box="[114,204,1544,1568]" pageId="9" pageNumber="900">M. Deki</collectorName>
et al., 
<date id="B502BC80FFE8BA28FEC5F9BBFE5FF991" box="[280,466,1544,1568]" pageId="9" pageNumber="900" value="2017-10-30">
<collectingDate id="A5464568FFE8BA28FEC5F9BBFE5FF991" box="[280,466,1544,1568]" pageId="9" pageNumber="900" value="2017-10-30">30 October 2017</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="71D4901DFFE8BA28FE03F9BBFD59F9D1" ID-GBIF-Occurrence="3763237450" collectingDate="1985-10-16" collectionCode="ZRC" collectorName="J. - F. Huang" country="Taiwan" location="TAIWAN" pageId="9" pageNumber="900" specimenCount="1" specimenCount-female="2" specimenCount-male="1" typeStatus="paralectotype">
<collectingCountry id="B9ABDAD0FFE8BA28FE03F9BBFDC7F991" box="[478,586,1544,1568]" name="Taiwan" pageId="9" pageNumber="900">TAIWAN</collectingCountry>
– 
<specimenCount id="D7BA51C9FFE8BA28FDBAF9BBFD3CF991" box="[615,689,1544,1568]" pageId="9" pageNumber="900" type="male">1 male</specimenCount>
(26.7 × 
<quantity id="064437A5FFE8BA28FF54F99BFF60F9F1" box="[137,237,1576,1600]" metricMagnitude="-2" metricUnit="m" metricValue="2.42" pageId="9" pageNumber="900" unit="mm" value="24.2">24.2 mm</quantity>
), 
<specimenCount id="D7BA51C9FFE8BA28FEDCF99BFEE7F9F1" box="[257,362,1576,1600]" pageId="9" pageNumber="900" type="female">2 females</specimenCount>
(22.8 × 
<quantity id="064437A5FFE8BA28FE14F99BFDA1F9F1" box="[457,556,1576,1600]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="9" pageNumber="900" unit="mm" value="19.6">19.6 mm</quantity>
, 13.9 × 
<quantity id="064437A5FFE8BA28FD5AF99BFD64F9F1" box="[647,745,1576,1600]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="9" pageNumber="900" unit="mm" value="11.9">11.9 mm</quantity>
) (
<collectionCode id="A7AD0285FFE8BA28FFA4F9FBFF3EF9D1" box="[121,179,1608,1632]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC</collectionCode>
), 
<location id="C463CC9BFFE8BA28FF17F9FAFEBBF9D1" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FF17F9FAFEBBF9D1" box="[202,310,1608,1632]" country="Taiwan" name="Yan Liau" pageId="9" pageNumber="900">Yan Liau</location>
, coll. 
<collectorName id="6C49FF96FFE8BA28FEA3F9FAFD89F9D0" box="[382,516,1609,1633]" pageId="9" pageNumber="900">J.-F. Huang</collectorName>
, 
<date id="B502BC80FFE8BA28FDCEF9FBFD59F9D1" box="[531,724,1608,1632]" pageId="9" pageNumber="900" value="1985-10-16">
<collectingDate id="A5464568FFE8BA28FDCEF9FBFD59F9D1" box="[531,724,1608,1632]" pageId="9" pageNumber="900" value="1985-10-16">16 October 1985</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FD3EF9FBFF27F8B1" ID-GBIF-Occurrence="3763237377" collectingDate="2002-06-21" collectionCode="ZRC" collectorName="H. - C. Liu &amp; P. K. L. Ng" country="Taiwan" county="Lion Museum" location="Toucheng town" municipality="Peikuan" pageId="9" pageNumber="900" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FD3EF9FBFF3DF931" pageId="9" pageNumber="900" type="male">5 males</specimenCount>
(35.6 × 
<quantity id="064437A5FFE8BA28FED2F9DBFEFFF931" box="[271,370,1640,1664]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="9" pageNumber="900" unit="mm" value="31.2">31.2 mm</quantity>
, 29.1 × 
<quantity id="064437A5FFE8BA28FE11F9DBFDA2F931" box="[460,559,1640,1664]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="900" unit="mm" value="25.4">25.4 mm</quantity>
, 25.4 × 
<quantity id="064437A5FFE8BA28FD54F9DBFD61F931" box="[649,748,1640,1664]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="9" pageNumber="900" unit="mm" value="22.9">22.9 mm</quantity>
, 17.7 × 
<quantity id="064437A5FFE8BA28FF1FF93BFEABF911" box="[194,294,1672,1696]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="9" pageNumber="900" unit="mm" value="15.6">15.6 mm</quantity>
, 15.6 × 
<quantity id="064437A5FFE8BA28FE59F93BFE67F911" box="[388,490,1672,1696]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="9" pageNumber="900" unit="mm" value="13.7">13.7 mm</quantity>
), 
<specimenCount id="D7BA51C9FFE8BA28FE22F93BFDD2F911" box="[511,607,1672,1696]" pageId="9" pageNumber="900" type="female">1 female</specimenCount>
(23.4 × 20.0 mm) (
<collectionCode id="A7AD0285FFE8BA28FF69F91BFF60F971" box="[180,237,1704,1728]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC</collectionCode>
), 
<collectingCounty id="2862E2CCFFE8BA28FEDCF91BFE16F971" box="[257,411,1704,1728]" pageId="9" pageNumber="900">Lion Museum</collectingCounty>
, 
<collectingMunicipality id="2167003AFFE8BA28FE75F91BFD8CF971" box="[424,513,1704,1728]" pageId="9" pageNumber="900">Peikuan</collectingMunicipality>
, 
<location id="C463CC9BFFE8BA28FDD0F91BFD39F971" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FDD0F91BFD39F971" box="[525,692,1704,1728]" country="Taiwan" county="Lion Museum" municipality="Peikuan" name="Toucheng town" pageId="9" pageNumber="900">Toucheng town</location>
area, 
<location id="C463CC9BFFE8BA28FFACF97BFEDDF951" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FFACF97BFEDDF951" box="[113,336,1736,1760]" country="Taiwan" county="Lion Museum" municipality="Peikuan" name="northeastern Taiwan" pageId="9" pageNumber="900">northeastern Taiwan</location>
, coll. 
<collectorName id="6C49FF96FFE8BA28FE4FF97BFE77F951" box="[402,506,1736,1760]" pageId="9" pageNumber="900">H.-C. Liu</collectorName>
&amp; 
<collectorName id="6C49FF96FFE8BA28FDC2F97AFD1DF950" box="[543,656,1737,1761]" pageId="9" pageNumber="900">P.K.L. Ng</collectorName>
, 
<date id="B502BC80FFE8BA28FD40F97BFF27F8B1" pageId="9" pageNumber="900" value="2002-06-21">
<collectingDate id="A5464568FFE8BA28FD40F97BFF27F8B1" pageId="9" pageNumber="900" value="2002-06-21">21 June 2002</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FF6AF95BFEC4F8F1" ID-GBIF-Occurrence="3763237466" collectingDate="2000-06-25" collectionCode="ZRC" collectorName="N. K. Ng &amp; P. - H. Ho" country="Taiwan" location="Pei Kuan" municipality="Lion Mountain" pageId="9" pageNumber="900" specimenCode="ZRC 2009.0153" specimenCount="3" specimenCount-male="3" stateProvince="Ilan County" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FF6AF95BFE86F8B1" box="[183,267,1768,1792]" pageId="9" pageNumber="900" type="male">3 males</specimenCount>
(
<specimenCode id="911A323BFFE8BA28FEC1F95BFE5DF8B1" box="[284,464,1768,1792]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC 2009.0153</specimenCode>
), coastal area behind 
<collectingMunicipality id="2167003AFFE8BA28FD63F95BFF51F891" pageId="9" pageNumber="900">Lion Mountain</collectingMunicipality>
, 
<location id="C463CC9BFFE8BA28FF36F8BBFED9F891" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FF36F8BBFED9F891" box="[235,340,1800,1824]" country="Taiwan" municipality="Lion Mountain" name="Pei Kuan" pageId="9" pageNumber="900" stateProvince="Ilan County">Pei Kuan</location>
, 
<collectingRegion id="037854A2FFE8BA28FEBFF8BBFE65F891" box="[354,488,1800,1824]" inferredBySuffix="true" pageId="9" pageNumber="900">Ilan County</collectingRegion>
, coll. 
<collectorName id="6C49FF96FFE8BA28FDF2F8BAFD02F890" box="[559,655,1801,1825]" pageId="9" pageNumber="900">N.K. Ng</collectorName>
&amp; 
<collectorName id="6C49FF96FFE8BA28FD65F8BAFF1BF8F1" pageId="9" pageNumber="900">P.-H. Ho</collectorName>
, 
<date id="B502BC80FFE8BA28FF7BF89BFEC4F8F1" box="[166,329,1832,1856]" pageId="9" pageNumber="900" value="2000-06-25">
<collectingDate id="A5464568FFE8BA28FF7BF89BFEC4F8F1" box="[166,329,1832,1856]" pageId="9" pageNumber="900" value="2000-06-25">25 June 2000</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FE84F89BFDD7F831" ID-GBIF-Occurrence="3763237379" collectingDate="2009-06-06" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Taiwan" location="Hsinchu" pageId="9" pageNumber="900" specimenCode="ZRC 2009.0669" specimenCount="1" specimenCount-juvenile="1" stateProvince="Hsinchu" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FE84F89BFE43F8F1" box="[345,462,1832,1856]" pageId="9" pageNumber="900" type="juvenile">1 juvenile</specimenCount>
male (
<specimenCode id="911A323BFFE8BA28FDFEF89BFD69F8F1" box="[547,740,1832,1856]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC 2009.0669</specimenCode>
), among grass, supralittoral zone, 
<collectingRegion id="037854A2FFE8BA28FE15F8FBFDACF8D1" box="[456,545,1864,1888]" country="Taiwan" name="Hsinchu" pageId="9" pageNumber="900">Hsinchu</collectingRegion>
wetlands, 
<collectingRegion id="037854A2FFE8BA28FD4CF8FBFD60F8D1" box="[657,749,1864,1888]" country="Taiwan" name="Hsinchu" pageId="9" pageNumber="900">Hsinchu</collectingRegion>
, northwest coast, coll. 
<collectorName id="6C49FF96FFE8BA28FE86F8DAFE46F830" box="[347,459,1897,1921]" pageId="9" pageNumber="900">P.K.L. Ng</collectorName>
, 
<date id="B502BC80FFE8BA28FE0BF8DBFDD7F831" box="[470,602,1896,1920]" pageId="9" pageNumber="900" value="2009-06-06">
<collectingDate id="A5464568FFE8BA28FE0BF8DBFDD7F831" box="[470,602,1896,1920]" pageId="9" pageNumber="900" value="2009-06-06">6 June 2009</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FDB8F8DBFDD3F851" ID-GBIF-Occurrence="3763237560" collectingDate="2002-06-22" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; H. - C. Liu" country="Taiwan" latitude="23.981667" location="eastern Taiwan" longLatPrecision="21" longitude="121.610275" municipality="Meilun" pageId="9" pageNumber="900" specimenCode="ZRC 2002.0472" specimenCount="1" specimenCount-male="1" stateProvince="Hualien" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FDB8F8DBFD20F831" box="[613,685,1896,1920]" pageId="9" pageNumber="900" type="male">1 male</specimenCount>
(
<specimenCode id="911A323BFFE8BA28FD66F8DBFF60F811" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC 2002.0472</specimenCode>
), 
<collectingRegion id="037854A2FFE8BA28FED8F83BFE33F811" box="[261,446,1928,1952]" country="Taiwan" name="Hualien" pageId="9" pageNumber="900">Hualien County</collectingRegion>
, mouth of 
<collectingMunicipality id="2167003AFFE8BA28FD9FF83BFD1EF811" box="[578,659,1928,1952]" pageId="9" pageNumber="900">Meilun</collectingMunicipality>
stream, 
<collectingRegion id="037854A2FFE8BA28FFAFF81BFF44F871" box="[114,201,1960,1984]" country="Taiwan" name="Hualien" pageId="9" pageNumber="900">Hualien</collectingRegion>
city, 
<location id="C463CC9BFFE8BA28FECDF81AFE33F871" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FECDF81AFE33F871" box="[272,446,1960,1984]" country="Taiwan" latitude="23.981667" longLatPrecision="21" longitude="121.610275" municipality="Meilun" name="eastern Taiwan" pageId="9" pageNumber="900" stateProvince="Hualien">eastern Taiwan</location>
, 
<geoCoordinate id="A488FC87FFE8BA28FE13F814FDDCF871" box="[462,593,1959,1984]" degrees="23" direction="north" minutes="58" orientation="latitude" pageId="9" pageNumber="900" precision="15" seconds="54" value="23.981667">23°58′54″N</geoCoordinate>
<geoCoordinate id="A488FC87FFE8BA28FD81F814FD61F871" box="[604,748,1959,1984]" degrees="121" direction="east" minutes="36" orientation="longitude" pageId="9" pageNumber="900" precision="15" seconds="37" value="121.610275">121°36′37″E</geoCoordinate>
, coll. 
<collectorName id="6C49FF96FFE8BA28FF71F87AFE93F850" box="[172,286,1993,2017]" pageId="9" pageNumber="900">P.K.L. Ng</collectorName>
&amp; 
<collectorName id="6C49FF96FFE8BA28FE9AF87BFE38F851" box="[327,437,1992,2016]" pageId="9" pageNumber="900">H.-C. Liu</collectorName>
, 
<date id="B502BC80FFE8BA28FE19F87BFDD3F851" box="[452,606,1992,2016]" pageId="9" pageNumber="900" value="2002-06-22">
<collectingDate id="A5464568FFE8BA28FE19F87BFDD3F851" box="[452,606,1992,2016]" pageId="9" pageNumber="900" value="2002-06-22">22 June 2002</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="71D4901DFFE8BA28FDB1F87BFCEBFF64" ID-GBIF-Occurrence="3763237413" collectingDate="2002-09-22" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; S. H. Fang" country="China" county="Chao Pu-Tou" location="Xiamen island" municipality="You Long village" pageId="9" pageNumber="900" specimenCode="ZRC 2002.0558" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="paralectotype">
<collectingCountry id="B9ABDAD0FFE8BA28FDB1F87BFD4CF851" box="[620,705,1992,2016]" name="China" pageId="9" pageNumber="900">CHINA</collectingCountry>
– 
<specimenCount id="D7BA51C9FFE8BA28FD3EF87BFF2BF7B1" pageId="9" pageNumber="900" type="male">1 male</specimenCount>
(
<specimenCode id="911A323BFFE8BA28FF6FF85BFEECF7B1" box="[178,353,2024,2048]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC 2002.0558</specimenCode>
), 
<collectingCounty id="2862E2CCFFE8BA28FEAFF85BFE72F7B1" box="[370,511,2024,2048]" pageId="9" pageNumber="900">Chao Pu-Tou</collectingCounty>
mangroves, 
<collectingMunicipality id="2167003AFFE8BA28FD58F85AFF30F791" pageId="9" pageNumber="900">You Long village</collectingMunicipality>
, mainland, outside 
<location id="C463CC9BFFE8BA28FE4AF7BBFDB4F791" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FE4AF7BBFDB4F791" box="[407,569,2056,2080]" country="China" county="Chao Pu-Tou" municipality="You Long village" name="Xiamen island" pageId="9" pageNumber="900" stateProvince="Fujian">Xiamen island</location>
, 
<location id="C463CC9BFFE8BA28FD9AF7BBFD13F791" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FD9AF7BBFD13F791" box="[583,670,2056,2080]" country="China" county="Chao Pu-Tou" municipality="You Long village" name="Xiamen" pageId="9" pageNumber="900" stateProvince="Fujian">Xiamen</location>
, 
<collectingRegion id="037854A2FFE8BA28FD71F7BBFF5CF7F1" country="China" name="Fujian" pageId="9" pageNumber="900">Fujian province</collectingRegion>
, 
<location id="C463CC9BFFE8BA28FF06F79BFE90F7F1" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FF06F79BFE90F7F1" box="[219,285,2088,2112]" country="China" county="Chao Pu-Tou" municipality="You Long village" name="China" pageId="9" pageNumber="900" stateProvince="Fujian">China</location>
, coll. 
<collectorName id="6C49FF96FFE8BA28FE86F79AFE45F7F0" box="[347,456,2089,2113]" pageId="9" pageNumber="900">P.K.L. Ng</collectorName>
&amp; 
<collectorName id="6C49FF96FFE8BA28FE3AF79BFDD9F7F0" box="[487,596,2088,2113]" pageId="9" pageNumber="900">S.H. Fang</collectorName>
, 
<date id="B502BC80FFE8BA28FD83F79BFCEBFF64" box="[606,870,189,2112]" pageId="9" pageNumber="900" value="2002-09-22">
<collectingDate id="A5464568FFE8BA28FD83F79BFCEBFF64" box="[606,870,189,2112]" pageId="9" pageNumber="900" value="2002-09-22">22 September 2002</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="71D4901DFFE8BA28FCA9FF0EFA99FEA4" ID-GBIF-Occurrence="3763237372" collectingDate="1975-06-08" collectionCode="NHM" collectorName="C. L. Soh" country="Hong Kong" location="Tai Po" pageId="9" pageNumber="900" specimenCode="NHM 1978.107" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<collectingCountry id="B9ABDAD0FFE8BA28FCA9FF0EFB95FF64" box="[884,1048,189,213]" name="Hong Kong" pageId="9" pageNumber="900">HONG KONG</collectingCountry>
– 
<specimenCount id="D7BA51C9FFE8BA28FBE6FF0EFB0AFF64" box="[1083,1159,189,213]" pageId="9" pageNumber="900" type="male">1 male</specimenCount>
(18.2 × 
<quantity id="064437A5FFE8BA28FB30FF0EFADFFF64" box="[1261,1362,189,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="9" pageNumber="900" unit="mm" value="16.6">16.6 mm</quantity>
) (
<specimenCode id="911A323BFFE8BA28FAB6FF0EFC1EFF44" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="9" pageNumber="900" type="Herbarium">NHM 1978.107</specimenCode>
) (
<typeStatus id="1E0724E2FFE8BA28FC71FF6EFB9DFF44" box="[940,1040,221,245]" pageId="9" pageNumber="900" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="06BCE1C3FFE8BA28FBE6FF6EFA12FF44" authority="Soh, 1978" authorityName="Soh" authorityYear="1978" box="[1083,1439,221,245]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFE8BA28FBE6FF6EFAACFF44" box="[1083,1313,221,245]" italics="true" pageId="9" pageNumber="900">Holometopus serenei</emphasis>
<bibRefCitation id="A52DE7B1FFE8BA28FAF6FF6EFA12FF44" author="Soh" box="[1323,1439,221,245]" firstAuthor="Soh" pageId="9" pageNumber="900" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh, 1978</bibRefCitation>
</taxonomicName>
), 
<location id="C463CC9BFFE8BA28FCF0FF4EFCF8FEA4" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FCF0FF4EFCF8FEA4" box="[813,885,253,277]" country="Hong Kong" name="Tai Po" pageId="9" pageNumber="900">Tai Po</location>
ricefield, coll. 
<collectorName id="6C49FF96FFE8BA28FBC1FF4EFB0DFEA4" box="[1052,1152,253,277]" pageId="9" pageNumber="900">C.L. Soh</collectorName>
, 
<date id="B502BC80FFE8BA28FB50FF4EFA99FEA4" box="[1165,1300,253,277]" pageId="9" pageNumber="900" value="1975-06-08">
<collectingDate id="A5464568FFE8BA28FB50FF4EFA99FEA4" box="[1165,1300,253,277]" pageId="9" pageNumber="900" value="1975-06-08">8 June 1975</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FAFCFF4EFA25FE84" ID-GBIF-Occurrence="3763237448" collectingDate="2000-07" collectionCode="ZRC" collectorName="H. H. Tan" country="Hong Kong" pageId="9" pageNumber="900" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FAFCFF4EFAE6FEA4" box="[1313,1387,253,277]" pageId="9" pageNumber="900" type="male">1 male</specimenCount>
(35.3 × 
<quantity id="064437A5FFE8BA28FC9EFEAEFC2EFE84" box="[835,931,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="3.06" pageId="9" pageNumber="900" unit="mm" value="30.6">30.6 mm</quantity>
) (
<collectionCode id="A7AD0285FFE8BA28FC64FEAEFC7FFE84" box="[953,1010,285,309]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC</collectionCode>
), in aquarium, coll. 
<collectorName id="6C49FF96FFE8BA28FB14FEADFABDFE84" box="[1225,1328,286,309]" pageId="9" pageNumber="900">H.H. Tan</collectorName>
, 
<date id="B502BC80FFE8BA28FAE6FEAEFA25FE84" box="[1339,1448,285,309]" pageId="9" pageNumber="900" value="2000-07">
<collectingDate id="A5464568FFE8BA28FAE6FEAEFA25FE84" box="[1339,1448,285,309]" pageId="9" pageNumber="900" value="2000-07">July 2000</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FCF0FE8DFADDFE04" ID-GBIF-Occurrence="3763237422" collectingDate="1996-06-06" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; S. Y. Lee" country="Hong Kong" county="Tai Tam" location="Tai Tam Dam" municipality="Hong Kong Island" pageId="9" pageNumber="900" specimenCode="ZRC 1997.761" specimenCount="7" specimenCount-female="7" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FCF0FE8DFC15FEE4" box="[813,920,317,341]" pageId="9" pageNumber="900" type="female">7 females</specimenCount>
(largest 12.0 × 
<quantity id="064437A5FFE8BA28FB8CFE8EFB35FEE4" box="[1105,1208,317,341]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="9" pageNumber="900" unit="mm" value="10.7">10.7 mm</quantity>
, smallest 24.5 × 
<quantity id="064437A5FFE8BA28FAA1FE8EFCD5FEC4" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="9" pageNumber="900" unit="mm" value="21.1">21.1 mm</quantity>
) (
<specimenCode id="911A323BFFE8BA28FCB2FEEEFB98FEC4" box="[879,1045,349,373]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC 1997.761</specimenCode>
), salt marsh, 
<collectingCounty id="2862E2CCFFE8BA28FB7AFEEEFA8AFEC4" box="[1191,1287,349,373]" pageId="9" pageNumber="900">Tai Tam</collectingCounty>
, south coast of 
<collectingMunicipality id="2167003AFFE8BA28FCF0FECDFC73FE24" box="[813,1022,381,406]" pageId="9" pageNumber="900">Hong Kong Island</collectingMunicipality>
, immediately downstream of 
<location id="C463CC9BFFE8BA28FA91FECEFCE9FE04" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FA91FECEFCE9FE04" country="Hong Kong" county="Tai Tam" municipality="Hong Kong Island" name="Tai Tam Dam" pageId="9" pageNumber="900">Tai Tam Dam</location>
, coll. 
<collectorName id="6C49FF96FFE8BA28FC73FE2DFBAFFE07" box="[942,1058,414,438]" pageId="9" pageNumber="900">P.K.L. Ng</collectorName>
&amp; 
<collectorName id="6C49FF96FFE8BA28FB91FE2EFB3EFE04" box="[1100,1203,413,437]" pageId="9" pageNumber="900">S.Y. Lee</collectorName>
, 
<date id="B502BC80FFE8BA28FB1CFE2EFADDFE04" box="[1217,1360,413,437]" pageId="9" pageNumber="900" value="1996-06-06">
<collectingDate id="A5464568FFE8BA28FB1CFE2EFADDFE04" box="[1217,1360,413,437]" pageId="9" pageNumber="900" value="1996-06-06">6 June 1996</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FA82FE2EFB3BFDA4" ID-GBIF-Occurrence="3763237381" collectingDate="2010-10-03" collectionCode="ZRC" collectorName="Tai Tam Dam &amp; K. Wong" country="Hong Kong" location="Hong Kong Island" municipality="Tai Tam" pageId="9" pageNumber="900" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FA82FE2EFA21FE04" box="[1375,1452,413,437]" pageId="9" pageNumber="900" type="male">1 male</specimenCount>
(30.8 × 
<quantity id="064437A5FFE8BA28FC59FE0EFC6BFE64" box="[900,998,445,469]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="9" pageNumber="900" unit="mm" value="26.9">26.9 mm</quantity>
) (
<collectionCode id="A7AD0285FFE8BA28FC20FE0EFBBBFE64" box="[1021,1078,445,469]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC</collectionCode>
), salt marsh, 
<collectingMunicipality id="2167003AFFE8BA28FB1AFE0EFAAAFE64" box="[1223,1319,445,469]" pageId="9" pageNumber="900">Tai Tam</collectingMunicipality>
, south coast of 
<location id="C463CC9BFFE8BA28FC97FE6DFB9FFE44" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FC97FE6DFB9FFE44" box="[842,1042,477,502]" country="Hong Kong" municipality="Tai Tam" name="Hong Kong Island" pageId="9" pageNumber="900">Hong Kong Island</location>
, immediately downstream of 
<collectorName id="6C49FF96FFE8BA28FA8DFE6EFCE9FDA4" pageId="9" pageNumber="900">Tai Tam Dam</collectorName>
, coll. 
<collectorName id="6C49FF96FFE8BA28FC7CFE4DFB88FDA7" box="[929,1029,510,534]" pageId="9" pageNumber="900">K. Wong</collectorName>
, 
<date id="B502BC80FFE8BA28FBD3FE4EFB3BFDA4" box="[1038,1206,509,533]" pageId="9" pageNumber="900" value="2010-10-03">
<collectingDate id="A5464568FFE8BA28FBD3FE4EFB3BFDA4" box="[1038,1206,509,533]" pageId="9" pageNumber="900" value="2010-10-03">3 October 2010</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FB1DFE4EFC55FD04" ID-GBIF-Occurrence="3763237551" collectingDate="2012-03-27" collectionCode="ZRC" collectorName="Tai Tam Dam &amp; K. Wong" country="Hong Kong" location="Hong Kong Island" municipality="Tai Tam" pageId="9" pageNumber="900" specimenCode="ZRC 2012.1221" specimenCount="6" specimenCount-female="6" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FB1DFE4EFA93FDA4" box="[1216,1310,509,533]" pageId="9" pageNumber="900" type="male">15 males</specimenCount>
(smallest 9.8 × 
<quantity id="064437A5FFE8BA28FC9BFDAEFC27FD84" box="[838,938,541,565]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="9" pageNumber="900" unit="mm" value="11.2">11.2 mm</quantity>
, largest 27.8 × 
<quantity id="064437A5FFE8BA28FB86FDAEFB4CFD84" box="[1115,1217,541,565]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="9" pageNumber="900" unit="mm" value="23.9">23.9 mm</quantity>
), 
<specimenCount id="D7BA51C9FFE8BA28FB0AFDAEFACCFD84" box="[1239,1345,541,565]" pageId="9" pageNumber="900" type="female">6 females</specimenCount>
(smallest 11.1 × 
<quantity id="064437A5FFE8BA28FCA7FD8EFC43FDE4" box="[890,974,573,597]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="900" unit="mm" value="9.5">9.5 mm</quantity>
, largest 21.1 × 
<quantity id="064437A5FFE8BA28FBABFD8EFB5AFDE4" box="[1142,1239,573,597]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="9" pageNumber="900" unit="mm" value="18.4">18.4 mm</quantity>
) (
<specimenCode id="911A323BFFE8BA28FB30FD8EFA2DFDE4" box="[1261,1440,573,597]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC 2012.1221</specimenCode>
), salt marsh, 
<collectingMunicipality id="2167003AFFE8BA28FC6EFDEEFB9AFDC4" box="[947,1047,605,629]" pageId="9" pageNumber="900">Tai Tam</collectingMunicipality>
, south coast of 
<location id="C463CC9BFFE8BA28FB0FFDEDFA2AFDC4" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FB0FFDEDFA2AFDC4" box="[1234,1447,605,630]" country="Hong Kong" municipality="Tai Tam" name="Hong Kong Island" pageId="9" pageNumber="900">Hong Kong Island</location>
, immediately downstream of 
<collectorName id="6C49FF96FFE8BA28FBBEFDCEFB72FD24" box="[1123,1279,637,661]" pageId="9" pageNumber="900">Tai Tam Dam</collectorName>
, coll. 
<collectorName id="6C49FF96FFE8BA28FA9FFDCDFA25FD27" box="[1346,1448,638,662]" pageId="9" pageNumber="900">K. Wong</collectorName>
, 
<date id="B502BC80FFE8BA28FCF0FD2EFC55FD04" box="[813,984,669,693]" pageId="9" pageNumber="900" value="2012-03-27">
<collectingDate id="A5464568FFE8BA28FCF0FD2EFC55FD04" box="[813,984,669,693]" pageId="9" pageNumber="900" value="2012-03-27">27 March 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFE8BA28FC38FD2EFB7CFC84" ID-GBIF-Occurrence="3763237333" collectingDate="2012-09-17" collectionCode="ZRC" collectorName="Tai Tam Dam &amp; K. Wong" country="Hong Kong" location="Hong Kong Island" municipality="Tai Tam" pageId="9" pageNumber="900" specimenCode="ZRC 2012.1222" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFE8BA28FC38FD2EFBB7FD04" box="[997,1082,669,693]" pageId="9" pageNumber="900" type="male">4 males</specimenCount>
(10.2 × 
<quantity id="064437A5FFE8BA28FB41FD2EFB7EFD04" box="[1180,1267,669,693]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="9" pageNumber="900" unit="mm" value="8.7">8.7 mm</quantity>
, 10.8 × 
<quantity id="064437A5FFE8BA28FA8CFD2EFA2AFD04" box="[1361,1447,669,693]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="9" pageNumber="900" unit="mm" value="9.4">9.4 mm</quantity>
, 11.6 × 
<quantity id="064437A5FFE8BA28FCA2FD0EFC5AFD64" box="[895,983,701,725]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="9" pageNumber="900" unit="mm" value="9.6">9.6 mm</quantity>
, 14.3 × 13.0 mm), 
<specimenCount id="D7BA51C9FFE8BA28FB6BFD0EFA9BFD64" box="[1206,1302,701,725]" pageId="9" pageNumber="900" type="female">1 female</specimenCount>
(17.5 × 
<quantity id="064437A5FFE8BA28FAA1FD0EFCD5FD44" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="9" pageNumber="900" unit="mm" value="15.1">15.1 mm</quantity>
) (
<specimenCode id="911A323BFFE8BA28FCACFD6EFBA5FD44" box="[881,1064,733,757]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="900">ZRC 2012.1222</specimenCode>
), salt marsh, 
<collectingMunicipality id="2167003AFFE8BA28FB1DFD6EFAAFFD44" box="[1216,1314,733,757]" pageId="9" pageNumber="900">Tai Tam</collectingMunicipality>
, south coast of 
<location id="C463CC9BFFE8BA28FC97FD4DFB9FFCA4" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFE8BA28FC97FD4DFB9FFCA4" box="[842,1042,765,790]" country="Hong Kong" municipality="Tai Tam" name="Hong Kong Island" pageId="9" pageNumber="900">Hong Kong Island</location>
, immediately downstream of 
<collectorName id="6C49FF96FFE8BA28FA8DFD4EFCE9FC84" pageId="9" pageNumber="900">Tai Tam Dam</collectorName>
, coll. 
<collectorName id="6C49FF96FFE8BA28FC7AFCADFB83FC87" box="[935,1038,798,822]" pageId="9" pageNumber="900">K. Wong</collectorName>
, 
<date id="B502BC80FFE8BA28FBC7FCAEFB7CFC84" box="[1050,1265,797,821]" pageId="9" pageNumber="900" value="2012-09-17">
<collectingDate id="A5464568FFE8BA28FBC7FCAEFB7CFC84" box="[1050,1265,797,821]" pageId="9" pageNumber="900" value="2012-09-17">17 September 2012</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFE8BA28FCF0FCEEFAE8FB04" pageId="9" pageNumber="900" type="diagnosis">
<paragraph id="C1039A40FFE8BA28FCF0FCEEFB91FBA4" blockId="9.[813,1452,861,1045]" pageId="9" pageNumber="900">
<emphasis id="F3C84652FFE8BA28FCF0FCEEFC2CFCC4" bold="true" box="[813,929,861,885]" pageId="9" pageNumber="900">Diagnosis</emphasis>
. Ambulatory legs relatively stouter, shorter, distal end of second ambulatory merus just reaches frontal margin when folded; row of granules on dorsal margin of cheliped dactylus only distinct in smaller specimens, very low or undiscernible in large males; male pleonal somite 6 is relatively narrower.
</paragraph>
<paragraph id="C1039A40FFE8BA28FCF0FB8EFAE8FB04" blockId="9.[813,1452,1085,1206]" pageId="9" pageNumber="900">
<emphasis id="F3C84652FFE8BA28FCF0FB8EFCF0FBE4" bold="true" box="[813,893,1085,1109]" pageId="9" pageNumber="900">Colour</emphasis>
. The carapace of adult 
<taxonomicName id="06BCE1C3FFE8BA28FB58FB8DFAB7FBE4" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[1157,1338,1085,1109]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE8BA28FB58FB8DFAB7FBE4" box="[1157,1338,1085,1109]" italics="true" pageId="9" pageNumber="900">C. haematocheir</emphasis>
</taxonomicName>
is usually dark green with the anterior parts and margins yellowish to red (see also 
<bibRefCitation id="A52DE7B1FFE8BA28FC19FBCEFBEDFB24" author="Miyake" box="[964,1120,1149,1173]" firstAuthor="Miyake" pageId="9" pageNumber="900" refId="ref72931" refString="Miyake S (1983) Japanese Crustacean Decapods and Stomatopods in Color, vol. II. Brachyura (Crabs). First Edition. Hoikusha, Osaka, viii + 277 pp., 64 pls., unnumbered figs." type="book" year="1983">Miyake, 1983</bibRefCitation>
; 
<bibRefCitation id="A52DE7B1FFE8BA28FBB3FBCEFAAAFB24" author="Minemizu" box="[1134,1319,1149,1173]" firstAuthor="Minemizu" pageId="9" pageNumber="900" refId="ref72906" refString="Minemizu R (2000) [Marine decapod and stomatopod crustaceans mainly from Japan]. Bun-Ichi Sogo Shuppan, Tokyo, 344 pp." type="book" year="2000">Minemizu, 2000</bibRefCitation>
; 
<bibRefCitation id="A52DE7B1FFE8BA28FAE8FBCDFCD5FB04" author="Lee" firstAuthor="Lee" page="134" pageId="9" pageNumber="900" refId="ref71440" refString="Lee J-H (2008) A Field Guide to Crabs in Taiwan. Bigtrees Company, Taipei, 174 pp." type="book" year="2008">Lee, 2008: 134</bibRefCitation>
; 
<bibRefCitation id="A52DE7B1FFE8BA28FCB8FB2EFBEFFB04" author="Liu &amp; Wang" box="[869,1122,1181,1206]" firstAuthor="Liu" page="58" pageId="9" pageNumber="900" refId="ref71886" refString="Liu H-C &amp; Wang C-H (2010) Taiwan Coastal Wetland Crabs. Wild Bird Society of Taipei, Taipei, 79 pp." type="book" year="2010">Liu &amp; Wang, 2010: 58</bibRefCitation>
; 
<bibRefCitation id="A52DE7B1FFE8BA28FBB2FB2DFAD4FB04" author="Toyota &amp; Seki" box="[1135,1369,1181,1206]" firstAuthor="Toyota" pageId="9" pageNumber="900" refId="ref78208" refString="Toyota K &amp; Seki S (2014) Shrimps and Crabs. Nature Watching Guidebook, Freshwater Shrimps and Crabs in Japan. Seibundo Shinkosha Publishing, 256 pp." type="book" year="2014">Toyota &amp; Seki, 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFE8BA2AFCF0FB6EFD97FB24" lastPageId="11" lastPageNumber="902" pageId="9" pageNumber="900" type="discussion">
<paragraph id="C1039A40FFE8BA28FCF0FB6EFC42FAC7" blockId="9.[813,1452,1245,1398]" pageId="9" pageNumber="900">
<emphasis id="F3C84652FFE8BA28FCF0FB6EFC18FB44" bold="true" box="[813,917,1245,1269]" pageId="9" pageNumber="900">Remarks</emphasis>
. There are 
<specimenCount id="D7BA51C9FFE8BA28FBCEFB6EFBE9FB47" box="[1043,1124,1245,1270]" pageId="9" pageNumber="900" type="generic">25 type</specimenCount>
specimens (and a set of dried mouthparts) of this species in RMNH (
<bibRefCitation id="A52DE7B1FFE8BA28FB14FB4EFA12FAA4" author="Fransen" box="[1225,1439,1277,1301]" etAl="et al." firstAuthor="Fransen" pageId="9" pageNumber="900" pagination="1 - 344" refId="ref69235" refString="Fransen CHJM, Holthuis LB &amp; Adema JPHM (1997) Typecatalogue of the Decapod Crustacea in the collections of the Nationaal Naturhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Verhandelingen, Leiden, 311: i - xvi, 1 - 344." type="journal article" year="1997">Fransen et al., 1997</bibRefCitation>
), with the 
<typeStatus id="1E0724E2FFE8BA28FC55FAAEFC67FA84" box="[904,1002,1309,1333]" pageId="9" pageNumber="900" type="lectotype">lectotype</typeStatus>
designated by 
<bibRefCitation id="A52DE7B1FFE8BA28FB5AFAAEFA25FA84" author="Yamaguchi &amp; Baba" box="[1159,1448,1309,1333]" firstAuthor="Yamaguchi" pageId="9" pageNumber="900" pagination="145 - 570" refId="ref78696" refString="Yamaguchi T &amp; Baba K (1993) Crustacean specimens collected in Japan by Ph. F. von Siebold and H. Burger and held by the Nationaal Natuurhistorisch Museum in Leiden and other Museums. In: Yamaguchi T (ed.) Ph. F. von Siebold and Natural History of Japan Crustacea. The Carcinological Society of Japan, Tokyo. Pp. 145 - 570." type="book chapter" year="1993">
<collectingRegion id="037854A2FFE8BA28FB5AFAAEFA8FFA84" box="[1159,1282,1309,1333]" country="Japan" name="Yamaguchi" pageId="9" pageNumber="900">Yamaguchi</collectingRegion>
&amp; 
<collectingRegion id="037854A2FFE8BA28FAFCFAAEFAD5FA84" box="[1313,1368,1309,1333]" country="Myanmar" name="Chin" pageId="9" pageNumber="900">Baba</collectingRegion>
(1993)
</bibRefCitation>
. The 
<typeStatus id="1E0724E2FFE8BA28FCBCFA8EFC48FAE4" box="[865,965,1341,1365]" pageId="9" pageNumber="900" type="lectotype">lectotype</typeStatus>
male is a specimen in excellent condition, despite its age.
</paragraph>
<paragraph id="C1039A40FFE8BA2BFCF0FA2EFC85F947" blockId="9.[812,1452,1437,2102]" lastBlockId="10.[136,776,1598,1783]" lastPageId="10" lastPageNumber="901" pageId="9" pageNumber="900">
<bibRefCitation id="A52DE7B1FFE8BA28FCF0FA2EFC2CFA04" author="Soh" box="[813,929,1437,1461]" firstAuthor="Soh" pageId="9" pageNumber="900" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh (1978)</bibRefCitation>
described 
<taxonomicName id="06BCE1C3FFE8BA28FBCDFA2EFB60FA04" authorityName="Soh" authorityYear="1978" box="[1040,1261,1437,1461]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFE8BA28FBCDFA2EFB60FA04" box="[1040,1261,1437,1461]" italics="true" pageId="9" pageNumber="900">Holometopus serenei</emphasis>
</taxonomicName>
from 
<collectingCountry id="B9ABDAD0FFE8BA28FAF7FA2DFA25FA07" box="[1322,1448,1438,1462]" name="Hong Kong" pageId="9" pageNumber="900">Hong Kong</collectingCountry>
. 
<bibRefCitation id="A52DE7B1FFE8BA28FCF1FA0EFC70FA64" author="Shen" box="[812,1021,1469,1493]" firstAuthor="Shen" page="237" pageId="9" pageNumber="900" pagination="211 - 242" refId="ref76939" refString="Shen CJ (1940 a) The brachyuran fauna of Hong Kong. Journal of the Hong Kong Fisheries Research Station, 1: 211 - 242." type="journal article" year="1940" yearSuffix="a">Shen (1940a: 237)</bibRefCitation>
had earlier recorded 
<taxonomicName id="06BCE1C3FFE8BA28FB2EFA0DFA21FA64" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[1267,1452,1469,1493]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE8BA28FB2EFA0DFA21FA64" box="[1267,1452,1469,1493]" italics="true" pageId="9" pageNumber="900">C. haematocheir</emphasis>
</taxonomicName>
from 
<collectingCountry id="B9ABDAD0FFE8BA28FCB6FA6DFC60FA47" box="[875,1005,1502,1526]" name="Hong Kong" pageId="9" pageNumber="900">Hong Kong</collectingCountry>
from a site called Wong Chuk Hang, a small village near Aberdeen, but 
<bibRefCitation id="A52DE7B1FFE8BA28FB48FA4EFAB5F9A4" author="Soh" box="[1173,1336,1533,1557]" firstAuthor="Soh" page="10" pageId="9" pageNumber="900" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh (1978: 10)</bibRefCitation>
noted that he could not find the species there as the area had since been developed. Examination of the 
<typeStatus id="1E0724E2FFE8BA28FB50F98EFB7DF9E4" box="[1165,1264,1597,1621]" pageId="9" pageNumber="900" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="06BCE1C3FFE8BA28FAC1F98EFC76F9C4" authority="Soh, 1978" authorityName="Soh" authorityYear="1978" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFE8BA28FAC1F98EFCF1F9C4" italics="true" pageId="9" pageNumber="900">Holometopus serenei</emphasis>
<bibRefCitation id="A52DE7B1FFE8BA28FC5BF9EEFC76F9C4" author="Soh" box="[902,1019,1629,1653]" firstAuthor="Soh" pageId="9" pageNumber="900" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh, 1978</bibRefCitation>
</taxonomicName>
, as well as numerous specimens from various locations around 
<collectingCountry id="B9ABDAD0FFE8BA28FB91F9CDFB5FF927" box="[1100,1234,1662,1686]" name="Hong Kong" pageId="9" pageNumber="900">Hong Kong</collectingCountry>
, confirms that it is only a junior synonym of 
<taxonomicName id="06BCE1C3FFE8BA28FB90F92DFA8FF904" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[1101,1282,1693,1717]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE8BA28FB90F92DFA8FF904" box="[1101,1282,1693,1717]" italics="true" pageId="9" pageNumber="900">C. haematocheir</emphasis>
</taxonomicName>
. All specimens reported or here examined are only juveniles or subadults of 
<taxonomicName id="06BCE1C3FFE8BA28FC90F96DFB89F944" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[845,1028,1757,1781]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE8BA28FC90F96DFB89F944" box="[845,1028,1757,1781]" italics="true" pageId="9" pageNumber="900">C. haematocheir</emphasis>
</taxonomicName>
(see also 
<bibRefCitation id="A52DE7B1FFE8BA28FBA4F96DFABFF944" author="Ng" box="[1145,1330,1757,1782]" etAl="et al." firstAuthor="Ng" pageId="9" pageNumber="900" pagination="1 - 286" refId="ref73754" refString="Ng PKL, Guinot D &amp; Davie PJF (2008 a) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008" yearSuffix="a">Ng et al., 2008a</bibRefCitation>
; 
<bibRefCitation id="A52DE7B1FFE8BA28FA9DF96DFC17F8A4" author="Naruse &amp; Ng" firstAuthor="Naruse" pageId="9" pageNumber="900" pagination="1 - 13" refId="ref73402" refString="Naruse T &amp; Ng PKL (2008) A new species of Chiromantes s. str. (Decapoda: Brachyura: Sesarmidae) from the Ryukyu Islands, Japan, with a note on the identity of Holometopus serenei Soh, 1978. Crustacean Research, 37: 1 - 13." type="journal article" year="2008">Naruse &amp; Ng, 2008</bibRefCitation>
). The 
<typeStatus id="1E0724E2FFE8BA28FC38F94EFBC8F8A4" box="[997,1093,1789,1813]" pageId="9" pageNumber="900" type="holotype">holotype</typeStatus>
male of 
<taxonomicName id="06BCE1C3FFE8BA28FB6DF94DFAA9F8A4" authorityName="Soh" authorityYear="1978" box="[1200,1316,1790,1813]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFE8BA28FB6DF94DFAA9F8A4" box="[1200,1316,1790,1813]" italics="true" pageId="9" pageNumber="900">H. serenei</emphasis>
</taxonomicName>
is not fully mature, as evidenced by its poorly developed chelae. The G1 of the type of 
<taxonomicName id="06BCE1C3FFE8BA28FC23F88DFBFDF8E4" authorityName="Soh" authorityYear="1978" box="[1022,1136,1854,1877]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFE8BA28FC23F88DFBFDF8E4" box="[1022,1136,1854,1877]" italics="true" pageId="9" pageNumber="900">H. serenei</emphasis>
</taxonomicName>
, however, is relatively well developed and is almost identical to that of 
<taxonomicName id="06BCE1C3FFE8BA28FB29F8EDFA28F8C4" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[1268,1445,1885,1909]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE8BA28FB29F8EDFA28F8C4" box="[1268,1445,1885,1909]" italics="true" pageId="9" pageNumber="900">C. haematocheir</emphasis>
</taxonomicName>
. The male pleon of 
<taxonomicName id="06BCE1C3FFE8BA28FBDFF8CDFBFEF824" authorityName="Soh" authorityYear="1978" box="[1026,1139,1918,1941]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFE8BA28FBDFF8CDFBFEF824" box="[1026,1139,1918,1941]" italics="true" pageId="9" pageNumber="900">H. serenei</emphasis>
</taxonomicName>
has the telson and somite 6 relatively more elongate than adult 
<taxonomicName id="06BCE1C3FFE8BA28FB7CF82DFADFF804" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[1185,1362,1949,1973]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE8BA28FB7CF82DFADFF804" box="[1185,1362,1949,1973]" italics="true" pageId="9" pageNumber="900">C. haematocheir</emphasis>
</taxonomicName>
, but this is also clearly associated with size and age. Medium-sized specimens of 
<taxonomicName id="06BCE1C3FFE8BA28FC63F86DFBFDF844" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[958,1136,2013,2037]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE8BA28FC63F86DFBFDF844" box="[958,1136,2013,2037]" italics="true" pageId="9" pageNumber="900">C. haematocheir</emphasis>
</taxonomicName>
from 
<collectingCountry id="B9ABDAD0FFE8BA28FB6DF86DFB60F844" box="[1200,1261,2014,2037]" name="Japan" pageId="9" pageNumber="900">Japan</collectingCountry>
have male pleons intermediate in form between the 
<typeStatus id="1E0724E2FFE8BA28FB4FF84EFB62F7A4" box="[1170,1263,2045,2069]" pageId="9" pageNumber="900" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="06BCE1C3FFE8BA28FACFF84DFAF2F7A4" authorityName="Soh" authorityYear="1978" box="[1298,1407,2046,2069]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFE8BA28FACFF84DFAF2F7A4" box="[1298,1407,2046,2069]" italics="true" pageId="9" pageNumber="900">H. serenei</emphasis>
</taxonomicName>
and adult 
<taxonomicName id="06BCE1C3FFE8BA28FCB7F7ADFB93F784" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[874,1054,2077,2101]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFE8BA28FCB7F7ADFB93F784" box="[874,1054,2077,2101]" italics="true" pageId="9" pageNumber="900">C. haematocheir</emphasis>
</taxonomicName>
. The 
<typeStatus id="1E0724E2FFE8BA28FB83F7ADFB37F787" box="[1118,1210,2078,2102]" pageId="9" pageNumber="900" type="paratype">paratype</typeStatus>
females of 
<taxonomicName id="06BCE1C3FFE8BA28FAE1F7ADFA26F784" authorityName="Soh" authorityYear="1978" box="[1340,1451,2078,2101]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="900" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFE8BA28FAE1F7ADFA26F784" box="[1340,1451,2078,2101]" italics="true" pageId="9" pageNumber="900">H. serenei</emphasis>
</taxonomicName>
are already mature despite being relatively small, but such early maturity is not abnormal—we similarly have a good series of small adult females of 
<taxonomicName id="06BCE1C3FFEBBA2BFDDAF9CCFD4BF927" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[519,710,1662,1686]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFEBBA2BFDDAF9CCFD4BF927" box="[519,710,1662,1686]" italics="true" pageId="10" pageNumber="901">C. haematocheir</emphasis>
</taxonomicName>
from 
<collectingCountry id="B9ABDAD0FFEBBA2BFF54F92DFF54F907" box="[137,217,1694,1718]" name="Taiwan" pageId="10" pageNumber="901">Taiwan</collectingCountry>
and 
<collectingCountry id="B9ABDAD0FFEBBA2BFECCF92CFEDCF907" box="[273,337,1695,1718]" name="Japan" pageId="10" pageNumber="901">Japan</collectingCountry>
. Direct comparisons of specimens of 
<taxonomicName id="06BCE1C3FFEBBA2BFD33F92CFE96F967" baseAuthorityName="De Haan" baseAuthorityYear="1833" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFEBBA2BFD33F92CFE96F967" italics="true" pageId="10" pageNumber="901">C. haematocheir</emphasis>
</taxonomicName>
comparable in size from 
<collectingCountry id="B9ABDAD0FFEBBA2BFDE9F90DFD09F967" box="[564,644,1726,1750]" name="Taiwan" pageId="10" pageNumber="901">Taiwan</collectingCountry>
to the male 
<typeStatus id="1E0724E2FFEBBA2BFF55F96DFF68F947" box="[136,229,1758,1782]" pageId="10" pageNumber="901" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="06BCE1C3FFEBBA2BFED2F96CFEF3F947" authorityName="Soh" authorityYear="1978" box="[271,382,1759,1782]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFEBBA2BFED2F96CFEF3F947" box="[271,382,1759,1782]" italics="true" pageId="10" pageNumber="901">H. serenei</emphasis>
</taxonomicName>
do not show any major differences.
</paragraph>
<caption id="95C3CAC8FFEBBA2BFF55FA28FBEFF9B6" ID-DOI="http://doi.org/10.5281/zenodo.5351301" ID-Zenodo-Dep="5351301" httpUri="https://zenodo.org/record/5351301/files/figure.png" pageId="10" pageNumber="901" startId="10.[136,171,1435,1457]" targetBox="[136,1476,189,1419]" targetPageId="10">
<paragraph id="C1039A40FFEBBA2BFF55FA28FBEFF9B6" blockId="10.[136,1476,1435,1543]" pageId="10" pageNumber="901">
Fig. 3. Frontal view of cephalothorax. A, 
<taxonomicName id="06BCE1C3FFEBBA2BFDC8FA28FC96FA01" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[533,795,1435,1456]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFEBBA2BFDC8FA28FC96FA01" box="[533,795,1435,1456]" italics="true" pageId="10" pageNumber="901">Chiromantes haematocheir</emphasis>
</taxonomicName>
, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, 
<taxonomicName id="06BCE1C3FFEBBA2BFABAFA28FF58FA7D" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="dehaani">
<emphasis id="F3C84652FFEBBA2BFABAFA28FF58FA7D" italics="true" pageId="10" pageNumber="901">Orisarma dehaani</emphasis>
</taxonomicName>
, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; C, 
<taxonomicName id="06BCE1C3FFEBBA2BFC9FFA0BFC4BFA7D" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[834,966,1463,1485]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="neglectum">
<emphasis id="F3C84652FFEBBA2BFC9FFA0BFC4BFA7D" box="[834,966,1463,1485]" italics="true" pageId="10" pageNumber="901">O. neglectum</emphasis>
</taxonomicName>
, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; D, 
<taxonomicName id="06BCE1C3FFEBBA2BFE87FA66FE78FA58" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[346,501,1492,1514]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="intermedium">
<emphasis id="F3C84652FFEBBA2BFE87FA66FE78FA58" box="[346,501,1492,1514]" italics="true" pageId="10" pageNumber="901">O. intermedium</emphasis>
</taxonomicName>
, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; E, 
<taxonomicName id="06BCE1C3FFEBBA2BFBF9FA66FB06FA5B" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1060,1163,1493,1514]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="sinense">
<emphasis id="F3C84652FFEBBA2BFBF9FA66FB06FA5B" box="[1060,1163,1493,1514]" italics="true" pageId="10" pageNumber="901">O. sinense</emphasis>
</taxonomicName>
, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; F, 
<taxonomicName id="06BCE1C3FFEBBA2BFE13FA42FDCDF9B7" authorityName="Soh" authorityYear="1978" box="[462,576,1521,1542]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="patshuni">
<emphasis id="F3C84652FFEBBA2BFE13FA42FDCDF9B7" box="[462,576,1521,1542]" italics="true" pageId="10" pageNumber="901">O. patshuni</emphasis>
</taxonomicName>
, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong.
</paragraph>
</caption>
<paragraph id="C1039A40FFEBBA2BFF55F8ADFB8EF866" blockId="10.[136,775,1822,2102]" lastBlockId="10.[835,1475,1598,2007]" pageId="10" pageNumber="901">
The good series of specimens from 
<collectingCountry id="B9ABDAD0FFEBBA2BFDD9F8ACFD0CF886" box="[516,641,1823,1847]" name="Hong Kong" pageId="10" pageNumber="901">Hong Kong</collectingCountry>
has allowed us to estimate the size at which males and females of this species mature. Two small female specimens (11.1 × 
<quantity id="064437A5FFEBBA2BFD38F8EDFF39F827" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="10" pageNumber="901" unit="mm" value="9.5">9.5 mm</quantity>
, 14.1 × 
<quantity id="064437A5FFEBBA2BFEDAF8CDFEEAF827" box="[263,359,1918,1942]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="10" pageNumber="901" unit="mm" value="11.8">11.8 mm</quantity>
) (
<specimenCode id="911A323BFFEBBA2BFEA6F8CDFDA4F827" box="[379,553,1918,1942]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="901">ZRC 2012.1221</specimenCode>
) are not fully mature with the pleon not completely covering the thoracic sternum, although a slightly larger specimen (14.5 × 
<quantity id="064437A5FFEBBA2BFDBFF80DFD49F867" box="[610,708,1982,2006]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="10" pageNumber="901" unit="mm" value="12.1">12.1 mm</quantity>
) from the same locality is already mature. The species therefore can mature at a relatively small size. The males also appear to be able to reach maturity at smaller sizes. As noted above, the small 
<typeStatus id="1E0724E2FFEBBA2BFC73F98DFB9DF9E7" box="[942,1040,1598,1622]" pageId="10" pageNumber="901" type="lectotype">lectotype</typeStatus>
male of 
<taxonomicName id="06BCE1C3FFEBBA2BFBAFF98CFB6DF9E7" authorityName="Soh" authorityYear="1978" box="[1138,1248,1599,1622]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFEBBA2BFBAFF98CFB6DF9E7" box="[1138,1248,1599,1622]" italics="true" pageId="10" pageNumber="901">H. serenei</emphasis>
</taxonomicName>
(18.2 × 
<quantity id="064437A5FFEBBA2BFAE2F98DFA2CF9E7" box="[1343,1441,1598,1622]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="10" pageNumber="901" unit="mm" value="16.6">16.6 mm</quantity>
) is already mature, with the G1 fully developed. 
<location id="C463CC9BFFEBBA2BFAEAF9EDFA33F9C7" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFEBBA2BFAEAF9EDFA33F9C7" box="[1335,1470,1630,1654]" country="Hong Kong" county="We" municipality="Taiwan" name="Surprisingly" pageId="10" pageNumber="901">Surprisingly</location>
, an even smaller male (9.8 × 
<quantity id="064437A5FFEBBA2BFB54F9CDFB63F927" box="[1161,1262,1662,1686]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="10" pageNumber="901" unit="mm" value="11.2">11.2 mm</quantity>
) (
<specimenCode id="911A323BFFEBBA2BFADAF9CDFA36F927" box="[1287,1467,1662,1686]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="901">ZRC 2012.1222</specimenCode>
) is also mature. 
<location id="C463CC9BFFEBBA2BFC30F92DFB9BF907" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFEBBA2BFC30F92DFB9BF907" box="[1005,1046,1694,1718]" country="Hong Kong" county="We" municipality="Taiwan" name="The" pageId="10" pageNumber="901">The</location>
structure of the male thoracic sternum also varies with size. 
<location id="C463CC9BFFEBBA2BFBE3F90DFBF3F967" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFEBBA2BFBE3F90DFBF3F967" box="[1086,1150,1726,1750]" country="Hong Kong" county="We" municipality="Taiwan" name="Small" pageId="10" pageNumber="901">Small</location>
specimens (10.8 × 
<quantity id="064437A5FFEBBA2BFABFF90DFA37F967" box="[1378,1466,1726,1750]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="10" pageNumber="901" unit="mm" value="9.4">9.4 mm</quantity>
) (
<specimenCode id="911A323BFFEBBA2BFC96F96DFC71F947" box="[843,1020,1758,1782]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="901">ZRC 2012.1222</specimenCode>
) have a sternite 4 that is relatively wider, becoming gradually longer as they increase in size (
<figureCitation id="598786C5FFEBBA2BFA4BF94DFC04F887" captionStart="Fig" captionStartId="17.[113,148,1943,1965]" captionTargetBox="[113,1453,223,1927]" captionTargetId="figure-14@17.[113,1453,223,1927]" captionTargetPageId="17" captionText="Fig. 7. A–D, Chiromantes haematocheir, anterior male thoracic sternum; A, male (10.8 × 9.4 mm) (ZRC 2012.1222), Hong Kong; B, male (14.3 × 13.0 mm) (ZRC 2012.1222), Hong Kong; C, male (18.4 × 16.6 mm) (lectotype of Holometopus serenei) (NHM 1978.107), Hong Kong; D, male (27.8 × 23.9 mm) (ZRC 2012.1221), Hong Kong. E–H, left third ambulatory legs; E–G, Orisarma dehaani: E, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; F, male (35.4 × 32.4 mm) (ZRC 2000.2263), Iriomote Island, Japan; G, male (29.6 × 27.6 mm) (ZRC 2002.0223), Hong Kong; H, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China." figureDoi="http://doi.org/10.5281/zenodo.5351309" httpUri="https://zenodo.org/record/5351309/files/figure.png" pageId="10" pageNumber="901">Fig. 7A–D</figureCitation>
). 
<location id="C463CC9BFFEBBA2BFC46F8ADFC42F887" LSID="urn:lsid:plazi:treatment:49152B56FFE6BA2AFCAFF8A9FF75F9A7:C463CC9BFFEBBA2BFC46F8ADFC42F887" box="[923,975,1822,1846]" country="Hong Kong" county="We" municipality="Taiwan" name="Why" pageId="10" pageNumber="901">Why</location>
some specimens from 
<collectingCountry id="B9ABDAD0FFEBBA2BFB1AF8ACFAC8F886" box="[1223,1349,1823,1847]" name="Hong Kong" pageId="10" pageNumber="901">Hong Kong</collectingCountry>
can mature at these smaller sizes is not known, some of the specimens we have from there are larger and agree with what we know from other parts of its range (e.g., in 
<collectingCountry id="B9ABDAD0FFEBBA2BFB37F8CCFAAAF827" box="[1258,1319,1919,1942]" name="Japan" pageId="10" pageNumber="901">Japan</collectingCountry>
and 
<collectingMunicipality id="2167003AFFEBBA2BFABFF8CDFA3AF827" box="[1378,1463,1918,1942]" pageId="10" pageNumber="901">Taiwan</collectingMunicipality>
). 
<collectingCounty id="2862E2CCFFEBBA2BFC9EF82CFCE5F807" box="[835,872,1951,1974]" pageId="10" pageNumber="901">We</collectingCounty>
are also uncertain if this phenomenon occurs in other parts of its range.
</paragraph>
<paragraph id="C1039A40FFEBBA2AFC9EF84DFD5DFEA4" blockId="10.[835,1474,2046,2102]" lastBlockId="11.[113,753,189,277]" lastPageId="11" lastPageNumber="902" pageId="10" pageNumber="901">
The lateral carapace margin of 
<taxonomicName id="06BCE1C3FFEBBA2BFB41F84CFADCF7A7" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[1180,1361,2046,2070]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="901" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFEBBA2BFB41F84CFADCF7A7" box="[1180,1361,2046,2070]" italics="true" pageId="10" pageNumber="901">C. haematocheir</emphasis>
</taxonomicName>
is usually entire, especially in adults and males. In smaller female specimens, however, the margin has two very broad and low lobiform teeth separated by very short and very narrow fissures, which are just discernible under magnification.
</paragraph>
<paragraph id="C1039A40FFEABA2AFFACFE8EFECEFD04" blockId="11.[113,754,317,693]" pageId="11" pageNumber="902">
Although most of the literature describes the dactylar finger of adult males of 
<taxonomicName id="06BCE1C3FFEABA2AFEF9FEEDFE5FFEC4" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[292,466,349,373]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="902" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFEABA2AFEF9FEEDFE5FFEC4" box="[292,466,349,373]" italics="true" pageId="11" pageNumber="902">C. haematocheir</emphasis>
</taxonomicName>
as unarmed, this is actually not the case. As discussed at length by 
<bibRefCitation id="A52DE7B1FFEABA2AFDD2FECDFD61FE24" author="Naruse &amp; Ng" box="[527,748,381,406]" firstAuthor="Naruse" pageId="11" pageNumber="902" pagination="1 - 13" refId="ref73402" refString="Naruse T &amp; Ng PKL (2008) A new species of Chiromantes s. str. (Decapoda: Brachyura: Sesarmidae) from the Ryukyu Islands, Japan, with a note on the identity of Holometopus serenei Soh, 1978. Crustacean Research, 37: 1 - 13." type="journal article" year="2008">Naruse &amp; Ng (2008)</bibRefCitation>
, the dorsal margin of the dactylar finger is actually lined with a row of regularly shaped low subrectangular granules that are especially prominent in smaller specimens (
<figureCitation id="598786C5FFEABA2AFD51FE6EFD69FE44" box="[652,740,477,501]" captionStart="Fig" captionStartId="14.[135,170,1987,2009]" captionTargetBox="[136,1476,189,1972]" captionTargetId="figure-7@14.[136,1476,189,1972]" captionTargetPageId="14" captionText="Fig. 5. Chela. A, B, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; C, C. haematocheir, male (18.6 × 20.7 mm) (RUMF-ZC-544), Japan; D–F, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; G, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; H, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong; I, O. patshuni, male (20.4 × 18.2 mm) (ZRC 2012.0032), Hong Kong. A, D, G, H, I, outer view; B, C, F, dorso-lateral view; E, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351305" httpUri="https://zenodo.org/record/5351305/files/figure.png" pageId="11" pageNumber="902">Fig. 5C</figureCitation>
), becoming very low or undiscernible in large individuals (
<figureCitation id="598786C5FFEABA2AFD18FE4EFF1EFD84" captionStart="Fig" captionStartId="14.[135,170,1987,2009]" captionTargetBox="[136,1476,189,1972]" captionTargetId="figure-7@14.[136,1476,189,1972]" captionTargetPageId="14" captionText="Fig. 5. Chela. A, B, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; C, C. haematocheir, male (18.6 × 20.7 mm) (RUMF-ZC-544), Japan; D–F, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; G, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; H, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong; I, O. patshuni, male (20.4 × 18.2 mm) (ZRC 2012.0032), Hong Kong. A, D, G, H, I, outer view; B, C, F, dorso-lateral view; E, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351305" httpUri="https://zenodo.org/record/5351305/files/figure.png" pageId="11" pageNumber="902">Fig. 5B</figureCitation>
). In fact, 
<taxonomicName id="06BCE1C3FFEABA2AFF20FDAEFDD4FD84" authority="Soh, 1978" authorityName="Soh" authorityYear="1978" box="[253,601,541,565]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="902" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFEABA2AFF20FDAEFE6CFD84" box="[253,481,541,565]" italics="true" pageId="11" pageNumber="902">Holometopus serenei</emphasis>
<bibRefCitation id="A52DE7B1FFEABA2AFE34FDAEFDD4FD84" author="Soh" box="[489,601,541,565]" firstAuthor="Soh" pageId="11" pageNumber="902" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh, 1978</bibRefCitation>
</taxonomicName>
, was partially diagnosed on this basis. Specimens of the closely related 
<taxonomicName id="06BCE1C3FFEABA2AFFAFFDEDFE9EFDC4" authorityName="Naruse &amp; Ng" authorityYear="2008" box="[114,275,605,629]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="902" phylum="Arthropoda" rank="species" species="ryukyuanus">
<emphasis id="F3C84652FFEABA2AFFAFFDEDFE9EFDC4" box="[114,275,605,629]" italics="true" pageId="11" pageNumber="902">C. ryukyuanus</emphasis>
</taxonomicName>
of a comparable size to 
<taxonomicName id="06BCE1C3FFEABA2AFDEBFDEDFD7CFDC4" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[566,753,605,629]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="902" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFEABA2AFDEBFDEDFD7CFDC4" box="[566,753,605,629]" italics="true" pageId="11" pageNumber="902">C. haematocheir</emphasis>
</taxonomicName>
have proportionately larger granules, that are distinct even in large specimens.
</paragraph>
<paragraph id="C1039A40FFEABA2AFFACFD6EFD8DFBA4" blockId="11.[113,754,733,1045]" pageId="11" pageNumber="902">
Although we do not have morphological support, our phylogenetic reconstruction (
<figureCitation id="598786C5FFEABA2AFE1BFD4EFD90FCA4" box="[454,541,765,789]" captionStart="Fig" captionStartId="91.[113,149,1757,1779]" captionTargetBox="[116,1448,352,1737]" captionTargetId="figure-15@91.[86,1475,347,1749]" captionTargetPageId="91" captionText="Fig. 59. Phylogenetic relationships among selected taxa of Sesarmidae (91 OTUs), including all representatives of Chiromantes and Pseudosesarma previous to current revision of these genera. In addition, several species of Bresedium and Sesarmops are considered, as the revision also involves some of their former members. Topology and confidence values were determined by means of a phylogenetic reconstruction based on 1,096 basepairs of mtDNA (16S-tRNALeu-ND1), conducted with MrBayes, GTR+I+G as substitution model, and Parasesarma plicatum as outgroup (midpoint rooting). Posterior probabilities were determined after 4 million generations with a burn-in of 25%. All species so far classified within Chiromantes (without obvious anterolateral teeth) are shaded in blue, whereas those species that used to be classified in Pseudosesarma (carrying at least one pair of anterolateral teeth) are shaded in yellow." figureDoi="http://doi.org/10.5281/zenodo.5351405" httpUri="https://zenodo.org/record/5351405/files/figure.png" pageId="11" pageNumber="902">Fig. 59</figureCitation>
) does suggest the possibility that there could still be more than one species within 
<taxonomicName id="06BCE1C3FFEABA2AFF1DFC8EFE67FCE4" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[192,490,829,853]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="902" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFEABA2AFF1DFC8EFE67FCE4" box="[192,490,829,853]" italics="true" pageId="11" pageNumber="902">Chiromantes haematocheir</emphasis>
</taxonomicName>
as defined here. If this is the case, it remains possible that 
<taxonomicName id="06BCE1C3FFEABA2AFE25FCEDFDEAFCC4" authorityName="Soh" authorityYear="1978" box="[504,615,862,885]" class="Malacostraca" family="Sesarmidae" genus="Holometopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="902" phylum="Arthropoda" rank="species" species="serenei">
<emphasis id="F3C84652FFEABA2AFE25FCEDFDEAFCC4" box="[504,615,862,885]" italics="true" pageId="11" pageNumber="902">C. serenei</emphasis>
</taxonomicName>
may still be recognised in the future as a valid cryptic or pseudocryptic species. A more detailed study with a larger series of samples (both morphological and molecular) across the known range of 
<taxonomicName id="06BCE1C3FFEABA2AFF4FFC6EFE36FC44" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[146,443,989,1013]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="902" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFEABA2AFF4FFC6EFE36FC44" box="[146,443,989,1013]" italics="true" pageId="11" pageNumber="902">Chiromantes haematocheir</emphasis>
</taxonomicName>
will need to be undertaken before this problem can be resolved.
</paragraph>
<paragraph id="C1039A40FFEABA2AFFACFB8EFD97FB24" blockId="11.[113,753,1085,1174]" pageId="11" pageNumber="902">
The record of “ 
<taxonomicName id="06BCE1C3FFEABA2AFEFDFB8EFDC5FBE4" baseAuthorityName="De Haan" baseAuthorityYear="1833" box="[288,584,1085,1109]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="902" phylum="Arthropoda" rank="species" species="haematocheir">
<emphasis id="F3C84652FFEABA2AFEFDFB8EFDC5FBE4" box="[288,584,1085,1109]" italics="true" pageId="11" pageNumber="902">Chiromantes haematocheir</emphasis>
</taxonomicName>
” by 
<bibRefCitation id="A52DE7B1FFEABA2AFD5EFB8EFF7DFBC4" author="Liu &amp; He" firstAuthor="Liu" page="162" pageId="11" pageNumber="902" refId="ref71910" refString="Liu W &amp; He W (2007) The benthic macro-invertebrates in Yangtze Estuary. Shanghai Scientific &amp; Technical Publisher, Shanghai, 203 pp." type="book" year="2007">Liu &amp; He (2007: 162)</bibRefCitation>
from 
<collectingCountry id="B9ABDAD0FFEABA2AFEEEFBEEFEFEFBC4" box="[307,371,1117,1141]" name="China" pageId="11" pageNumber="902">China</collectingCountry>
is actually 
<taxonomicName id="06BCE1C3FFEABA2AFE32FBEDFF75FB24" authority="(De Haan, 1837)" baseAuthorityName="De Haan" baseAuthorityYear="1837" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="902" phylum="Arthropoda" rank="species" species="affine">
<emphasis id="F3C84652FFEABA2AFE32FBEDFD4DFBC4" box="[495,704,1117,1141]" italics="true" pageId="11" pageNumber="902">Parasesarma affine</emphasis>
(De Haan, 1837)
</taxonomicName>
(cf. 
<bibRefCitation id="A52DE7B1FFEABA2AFEF0FBCEFD83FB24" author="Rahayu &amp; Ng" box="[301,526,1149,1174]" firstAuthor="Rahayu" pageId="11" pageNumber="902" pagination="1 - 22" refId="ref74919" refString="Rahayu DL &amp; Ng PKL (2010) Revision of the Parasesarma plicatum (Latreille, 1803) species-group (Crustacea: Decapoda: Brachyura: Sesarmidae). Zootaxa, 2327: 1 - 22." type="journal article" year="2010">Rahayu &amp; Ng, 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFEABA2AFFACFB0EFD4EFA24" pageId="11" pageNumber="902" type="biology_ecology">
<paragraph id="C1039A40FFEABA2AFFACFB0EFD4EFA24" blockId="11.[113,753,1213,1429]" pageId="11" pageNumber="902">
<emphasis id="F3C84652FFEABA2AFFACFB0EFF47FB64" bold="true" box="[113,202,1213,1237]" pageId="11" pageNumber="902">Biology</emphasis>
. This species is found in supralittoral habitats, usually some distance from the sea, often adjacent to or near freshwater habitats. They may be found several kilometres inland if the area is flat and still subjected to some tidal influence. The larval development has been reported by 
<bibRefCitation id="A52DE7B1FFEABA2AFFAFFAEEFEF7FAC4" author="Fukuda &amp; Baba" box="[114,378,1373,1397]" firstAuthor="Fukuda" pageId="11" pageNumber="902" pagination="61 - 75" refId="ref69321" refString="Fukuda Y &amp; Baba K (1976) Complete larval development of the sesarminid crabs, Chiromantes bidens, Holometopus haematocheir, Parasesarma plicatum, and Sesarmops intermedius, reared in the laboratory. Memoirs of the Faculty of Education, Kumamoto University, Natural Science, 25: 61 - 75." type="journal article" year="1976">
Fukuda &amp; 
<collectingRegion id="037854A2FFEABA2AFF33FAEEFEABFAC4" box="[238,294,1373,1397]" country="Myanmar" name="Chin" pageId="11" pageNumber="902">Baba</collectingRegion>
(1976)
</bibRefCitation>
and 
<bibRefCitation id="A52DE7B1FFEABA2AFE6BFAEEFDFFFAC4" author="Oh" box="[438,626,1373,1397]" etAl="et al." firstAuthor="Oh" pageId="11" pageNumber="902" pagination="9 - 20" refId="ref74361" refString="Oh SM, Chung DE &amp; Ko HS (2007) Complete larval development of Chiromantes haematocheir (Crustacea: Decapoda: Brachyura: Grapsidae), with a key to seven sesarminid zoeas in Korea. Korean Journal of Systematics and Zoology, 23 (1): 9 - 20." type="journal article" year="2007">Oh et al. (2007)</bibRefCitation>
. Nematode parasites have been reported on by 
<bibRefCitation id="A52DE7B1FFEABA2AFE2BFACEFD32FA24" author="Yoshimura" box="[502,703,1405,1429]" firstAuthor="Yoshimura" pageId="11" pageNumber="902" pagination="755 - 761" refId="ref78855" refString="Yoshimura K (1990) Two new species of the genus Monhystrium Cobb, 1920 (Monhysteridae: Nematoda) from terrestrial crabs of subfamily Sesarminae (Brachyura) in Japan. Zoological Science, 7 (4): 755 - 761." type="journal article" year="1990">Yoshimura (1990)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFEABA2AFFACFA0EFF75F9A7" pageId="11" pageNumber="902" type="distribution">
<paragraph id="C1039A40FFEABA2AFFACFA0EFF75F9A7" blockId="11.[113,752,1469,1558]" pageId="11" pageNumber="902">
<emphasis id="F3C84652FFEABA2AFFACFA0EFF71FA64" bold="true" box="[113,252,1469,1493]" pageId="11" pageNumber="902">Distribution</emphasis>
. Widely distributed in 
<collectingCountry id="B9ABDAD0FFEABA2AFE21FA0DFDEBFA64" box="[508,614,1469,1493]" name="China" pageId="11" pageNumber="902">East Asia</collectingCountry>
, from 
<collectingCountry id="B9ABDAD0FFEABA2AFD72FA0DFD7DFA64" box="[687,752,1470,1493]" name="South Korea" pageId="11" pageNumber="902">Korea</collectingCountry>
to mainland 
<collectingCountry id="B9ABDAD0FFEABA2AFF24FA6DFEB5FA44" box="[249,312,1502,1525]" name="Japan" pageId="11" pageNumber="902">Japan</collectingCountry>
, 
<collectingCountry id="B9ABDAD0FFEABA2AFE98FA6EFE15FA44" box="[325,408,1501,1525]" name="Taiwan" pageId="11" pageNumber="902">Taiwan</collectingCountry>
, and mainland 
<collectingCountry id="B9ABDAD0FFEABA2AFDE0FA6EFDF2FA44" box="[573,639,1501,1525]" name="China" pageId="11" pageNumber="902">China</collectingCountry>
, including 
<collectingCountry id="B9ABDAD0FFEABA2AFFACFA4DFF79F9A7" box="[113,244,1534,1558]" name="Hong Kong" pageId="11" pageNumber="902">Hong Kong</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>