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<document id="C7C27B0768C1F68333E62811E3199536" ID-CLB-Dataset="55667" ID-DOI="10.26107/RBZ-2020-0097" ID-GBIF-Dataset="6cb155ce-8b9f-48ce-8e6c-b0dc00f8b1cc" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351295" ID-ZooBank="815E4670-B063-4FD8-B31E-3AD89B3A7942" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630205841501" checkinUser="diego" docAuthor="Schubart, Christoph D. &amp; Ng, Peter K. L." docDate="2020" docId="49152B56FFD9BA18FF1BF96CFA12FB64" docLanguage="en" docName="RafflesBZool.68.891-994.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Pseudosesarma crassimanum" docType="treatment" docVersion="16" lastPageNumber="948" masterDocId="B52C532EFFE1BA21FFDDFFB3FF8DFFB1" masterDocTitle="Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species" masterLastPageNumber="994" masterPageNumber="891" pageNumber="947" updateTime="1701997587259" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="03590900665CE439252900BEAFA953CD">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<mods:namePart id="7CC52FA3F0D9E3D023398C584328918C">Schubart, Christoph D.</mods:namePart>
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<mods:namePart id="F6D5298DBD00A352C6E486A05EAA432A">Ng, Peter K. L.</mods:namePart>
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<emphasis id="F3C84652FFD9BA19FF1BF96CFD80F946" bold="true" box="[198,525,1759,1783]" italics="true" pageId="56" pageNumber="947">Pseudosesarma crassimanum</emphasis>
(
<bibRefCitation id="A52DE7B1FFD9BA19FDC0F96CFD4FF946" author="De Man" box="[541,706,1759,1783]" firstAuthor="De Man" pageId="56" pageNumber="947" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">De Man, 1887</bibRefCitation>
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<subSubSection id="89A6C9CBFFD9BA19FF42F94CFD7DF8A6" box="[159,752,1791,1815]" pageId="56" pageNumber="947" type="description">
<paragraph id="C1039A40FFD9BA19FF42F94CFD7DF8A6" blockId="56.[159,752,1759,1815]" box="[159,752,1791,1815]" pageId="56" pageNumber="947">
(
<figureCitation id="598786C5FFD9BA19FF7AF94CFE9BF8A6" box="[167,278,1791,1815]" captionStart="Fig" captionStartId="41.[113,148,1553,1575]" captionTargetBox="[113,1451,189,1537]" captionTargetId="figure-14@41.[113,1451,189,1537]" captionText="Fig. 22. Overall habitus, Pseudosesarma species. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. edwardsii, male (12.9 × 11.8 mm) (ZRC 1971.9.24.9), Singapore; D, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519005" httpUri="https://zenodo.org/record/6519005/files/figure.png" pageId="56" pageNumber="947">Figs. 22D</figureCitation>
, 
<figureCitation id="598786C5FFD9BA19FEFEF94CFEDFF8A6" box="[291,338,1791,1815]" captionStart="Fig" captionStartId="46.[136,171,1913,1935]" captionTargetBox="[147,1463,189,1899]" captionTargetId="figure-8@46.[147,1463,189,1899]" captionText="Fig. 25. Frontal view of cephalothorax of Pseudosesarma. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; D, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; E, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; F, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; G, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq; H, Contusarma bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; I, Contusarma cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; J, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo." figureDoi="http://doi.org/10.5281/zenodo.6519007" httpUri="https://zenodo.org/record/6519007/files/figure.png" pageId="56" pageNumber="947">25C</figureCitation>
, 
<figureCitation id="598786C5FFD9BA19FE82F94CFE02F8A6" box="[351,399,1791,1815]" captionStart="Fig" captionStartId="49.[113,148,1885,1907]" captionTargetBox="[113,1451,246,1869]" captionTargetId="figure-14@49.[113,1451,246,1869]" captionText="Fig. 27. Outer view of chela of Pseudosesarma. A, P. edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. edwardsii, male (19.4 × 17.3 mm) (ZRC 1971.9.24.8), Singapore; D, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519009" httpUri="https://zenodo.org/record/6519009/files/figure.png" pageId="56" pageNumber="947">27D</figureCitation>
, 
<figureCitation id="598786C5FFD9BA19FE41F94CFE47F8A6" box="[412,458,1791,1815]" captionStart="Fig" captionStartId="52.[136,171,1939,1961]" captionTargetBox="[173,1438,189,1924]" captionTargetId="figure-7@52.[173,1438,189,1924]" captionText="Fig. 30. Male anterior thoracic sternum and pleon of Pseudosesarma. A, P edwardsii, lectotype male (17.5 × 16.1 mm) (RMNH-D17a), Mergui Archipelago; B, P. edwardsii, male (13.4 × 11.9 mm) (ZRC 1965.8.2.81), Pulau Pawai, Singapore; C, P. crassimanum, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore; D, P. glabrum, holotype male (13.8 × 12.0 mm) (CUSAT 2016-1), Kerala, India; E, P. anteactum, holotype male (16.7 × 14.7 mm) (ZRC 2016.0602), Sri Lanka; F, P. anteactum, paratype male (21.2 × 19.7 mm) (ZRC 2016.0603), Sri Lanka; G, P. brehieri, holotype male (17.4 × 15.7 mm) (ZRC 2016.0593), Myanmar; H, P. boulengeri, lectotype male (26.9 × 23.5 mm) (NHM 1919.11.14.1), Basra, Iraq." figureDoi="http://doi.org/10.5281/zenodo.6519011" httpUri="https://zenodo.org/record/6519011/files/figure.png" pageId="56" pageNumber="947">30C</figureCitation>
, 
<figureCitation id="598786C5FFD9BA19FE05F94CFE78F8A6" box="[472,501,1791,1815]" captionStart="Fig" captionStartId="55.[113,148,1344,1366]" captionTargetBox="[113,1451,189,1328]" captionTargetId="figure-15@55.[113,1451,189,1328]" captionTargetPageId="55" captionText="Fig. 32. Male sternopleonal cavity. A, Pseudosesarma edwardsii, male (20.8 × 18.6mm (ZRC 2003.0084), Pulau Tekong, Singapore; B, Pseudosesarma crassimanum, male (15.1 × 13.3 mm) (ZRC 2000.1768), Sungei Benut, Johor, Malaysia; C, Contusarma cheirogonum, male (15.3 × 14.0 mm) (ZRC 2011.0924), Pulau Tioman, Malaysia; D, Miersarma granosimanum, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia. Arrow indicates shallow depression on somite 5 to accept G1 tip when pleon closed (depression absent in Miersarma, D)." figureDoi="http://doi.org/10.5281/zenodo.5351355" httpUri="https://zenodo.org/record/5351355/files/figure.png" pageId="56" pageNumber="947">32</figureCitation>
B, 
<figureCitation id="598786C5FFD9BA19FDC9F94CFDBCF8A6" box="[532,561,1791,1815]" captionStart="Fig" captionStartId="60.[136,171,2006,2028]" captionTargetBox="[204,1406,196,1992]" captionTargetId="figure-8@60.[201,1408,185,1996]" captionTargetPageId="60" captionText="Fig. 35. A–E, Pseudosesarma crassimanum. A–G, lectotype male (16.3 × 14.6 mm) (NHM 1886.52b), Myanmar; H–M, male (15.9 × 14.3 mm) (ZRC 2008.0442), Ranong; N–R, male (15.9 × 14.5 mm) (RMNH-D23313), Singapore. A, N, pleon; B–F, right G1; G, right G2; H–L, O–R, left G1; M, left G2. Scales: A = 2.0 mm; B, G, H, I, M, O, P = 1.0 mm; N = 5.0 mm; C–F, J–L, Q, R = 0.5 mm. After Ng &amp; Schubart (2017: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.5351361" httpUri="https://zenodo.org/record/5351361/files/figure.png" pageId="56" pageNumber="947">35</figureCitation>
, 
<figureCitation id="598786C5FFD9BA19FDE3F94CFDD6F8A6" box="[574,603,1791,1815]" captionStart="Fig" captionStartId="74.[136,171,1898,1920]" captionTargetBox="[136,1474,211,1884]" captionTargetId="figure-7@74.[136,1474,211,1884]" captionTargetPageId="74" captionText="Fig. 44. Vulva. A, Pseudosesarma edwardsii, female (16.5 × 15.3 mm) (ZRC 2016.0608), Malaysia; B, Pseudosesarma crassimanum, female (15.9 × 13.8 mm) (ZRC 2000.1768), Malaysia; C, Pseudosesarma crassimanum, female (18.0 × 15.7 mm) (ZRC 2017.1045), Thailand; D, Pseudosesarma anteactum, paratype female (17.5 × 15.4 mm) (ZRC 2016.0603), Sri Lanka; E, Pseudosesarma brehieri, paratype female (14.9 × 13.4 mm) (ZRC 2016.0594), Myanmar; F, Pseudosesarma boulengeri, female (24.2 × 20.0 mm) (ZRC 2014.0335), Iran; G, Contusarma bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Thailand; H, Contusarma cheirogonum, female (26.6 × 23.0 mm) (ZRC 1964.9.28.127), Sarawak; I, Miersarma granosimanum, female (20.1 × 16.4 mm) (ZRC 1965.7.29.166), Malaysia; J, Manarma moeschii, female (19.3 × 17.3 mm) (ZRC 2017.0170), Thailand." figureDoi="http://doi.org/10.5281/zenodo.5351375" httpUri="https://zenodo.org/record/5351375/files/figure.png" pageId="56" pageNumber="947">44</figureCitation>
B, C, 
<figureCitation id="598786C5FFD9BA19FD47F94CFD3BF8A6" box="[666,694,1791,1815]" captionStart="Fig" captionStartId="88.[136,171,1875,1897]" captionTargetBox="[136,1474,324,1860]" captionTargetId="figure-7@88.[136,1474,324,1860]" captionTargetPageId="88" captionText="Fig. 56. Colour in life. A, B, Pseudosesarma edwardsii, male (20.5 × 19.3 mm) (ZRC 2016.0608), Langkawi [photograph: Paul Ng]; C–E, P. edwardsii, male (19.4 × 18.2 mm) (SMF), Langkawi, Peninsular Malaysia [photograph: CD Schubart]; F, G, P. crassimanum, (12.6 × 10.5 mm) (ZRC 2010.0035), Tioman, Peninsular Malaysia [photograph: PKL Ng]; H, P. boulengeri, specimen not preserved [photograph: Reza Naderloo]." figureDoi="http://doi.org/10.5281/zenodo.5351399" httpUri="https://zenodo.org/record/5351399/files/figure.png" pageId="56" pageNumber="947">56</figureCitation>
F, G)
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<treatmentCitation id="401DBC51FFD9BA19FF55F88DFD22F8E5" author="Man JG De" box="[136,687,1854,1876]" page="649" pageId="56" pageNumber="947" year="1887">
<taxonomicName id="06BCE1C3FFD9BA19FF55F88DFD22F8E5" ID-CoL="84HS6" authority="De Man, 1887: 649" authorityName="De Man" authorityPageNumber="649" authorityYear="1887" box="[136,687,1854,1876]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="947" phylum="Arthropoda" rank="variety" species="edwardsii" variety="crassimana">
<emphasis id="F3C84652FFD9BA19FF55F88DFECFF8E2" box="[136,322,1854,1875]" italics="true" pageId="56" pageNumber="947">Sesarma edwardsii</emphasis>
var. 
<emphasis id="F3C84652FFD9BA19FEA8F88DFE69F8E2" box="[373,484,1854,1875]" italics="true" pageId="56" pageNumber="947">crassimana</emphasis>
<bibRefCitation id="A52DE7B1FFD9BA19FE31F88DFD22F8E5" author="De Man" box="[492,687,1854,1876]" firstAuthor="De Man" page="649" pageId="56" pageNumber="947" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">De Man, 1887: 649</bibRefCitation>
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<taxonomicName id="06BCE1C3FFD9BA19FF55F8E8FE6DF8C1" ID-CoL="84HS6" authority="De Man, 1887" box="[136,480,1882,1904]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="947" phylum="Arthropoda" rank="variety" species="edwardsi" variety="crassimana">
<emphasis id="F3C84652FFD9BA19FF55F8E8FEB6F8DE" box="[136,315,1882,1904]" italics="true" pageId="56" pageNumber="947">Sesarma edwardsi</emphasis>
var. 
<emphasis id="F3C84652FFD9BA19FEACF8E8FE6DF8C1" box="[369,480,1883,1904]" italics="true" pageId="56" pageNumber="947">crassimana</emphasis>
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– 
<bibRefCitation id="A52DE7B1FFD9BA19FE20F8E8FD4CF8C1" author="De Man" box="[509,705,1883,1904]" firstAuthor="De Man" page="188" pageId="56" pageNumber="947" pagination="177 - 240" refId="ref72143" refString="Man JG De (1888) Report on the Podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum. - Part IV. Journal of the Linnean Society of London (Zoology), 22 (139): 177 - 240, pls. 13 - 15." type="journal article" year="1888">
De 
<treatmentCitation id="401DBC51FFD9BA19FDFCF8E8FD4CF8C1" author="Man JG De" box="[545,705,1883,1904]" page="188" pageId="56" pageNumber="947" year="1888">Man, 1888: 188</treatmentCitation>
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, pl. 13 figs. 5, 6; 
<treatmentCitation id="401DBC51FFD9BA19FECEF8C4FE50F83D" author="Zehntner L" box="[275,477,1911,1933]" page="180" pageId="56" pageNumber="947" year="1894">
<bibRefCitation id="A52DE7B1FFD9BA19FECEF8C4FE50F83D" author="Zehntner" box="[275,477,1911,1933]" firstAuthor="Zehntner" page="180" pageId="56" pageNumber="947" pagination="135 - 214" refId="ref78899" refString="Zehntner L (1894) Crustaces de l'Archipel Malais. Voyage de MM. M. Bedot et C. Pictet dans l'Archipel Malais. Revue Suisse de Zoologie et Annales du Musee d'Histoire Naturelle de Geneve, 2: 135 - 214, pls. 7 - 9." type="journal article" year="1894">Zehntner, 1894: 180</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFD9BA19FE35F8C4FD48F83C" author="Lanchester WF" box="[488,709,1911,1933]" page="757" pageId="56" pageNumber="947" year="1900">
<bibRefCitation id="A52DE7B1FFD9BA19FE35F8C4FD48F83C" author="Lanchester" box="[488,709,1911,1933]" firstAuthor="Lanchester" page="757" pageId="56" pageNumber="947" pagination="719 - 770" refId="ref71180" refString="Lanchester WF (1900) On a collection of crustaceans made at Singapore and Malacca. Part I. Crustacea Brachyura. Proceedings of the Zoological Society of London, 1900 (3): 719 - 770, pls. 44 - 47." type="journal article" year="1900">Lanchester, 1900: 757</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD9BA19FF55F827FE95F877" blockId="56.[136,775,1854,2106]" pageId="56" pageNumber="947">
<treatmentCitationGroup id="E1ACBD6EFFD9BA19FF55F827FE95F877" pageId="56" pageNumber="947">
<taxonomicName id="06BCE1C3FFD9BA19FF55F827FD06F81B" ID-CoL="84HS6" authority="(De Man, 1887)" box="[136,651,1940,1962]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="947" phylum="Arthropoda" rank="variety" species="edwardsi" subGenus="Episesarma" variety="crassimana">
<emphasis id="F3C84652FFD9BA19FF55F827FF52F818" box="[136,223,1940,1961]" italics="true" pageId="56" pageNumber="947">Sesarma</emphasis>
(
<emphasis id="F3C84652FFD9BA19FF2FF826FEE7F81B" box="[242,362,1941,1962]" italics="true" pageId="56" pageNumber="947">Episesarma</emphasis>
) 
<emphasis id="F3C84652FFD9BA19FEA3F827FE56F818" box="[382,475,1940,1961]" italics="true" pageId="56" pageNumber="947">edwardsi</emphasis>
var. 
<emphasis id="F3C84652FFD9BA19FDCBF826FD06F81B" box="[534,651,1941,1962]" italics="true" pageId="56" pageNumber="947">crassimana</emphasis>
</taxonomicName>
– 
<bibRefCitation id="A52DE7B1FFD9BA19FD71F826FE99F877" author="De Man" firstAuthor="De Man" page="174" pageId="56" pageNumber="947" pagination="75 - 218" refId="ref72333" refString="Man JG De (1895) Bericht uber die von Herrn Schiffscaptan Storm zu Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der JavaSee gesammelten Decapoden und Stomatopoden. Zweiter Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 9: 75 - 218." type="journal article" year="1895">
De 
<treatmentCitation id="401DBC51FFD9BA19FD0EF826FE99F877" author="Man JG De" page="174" pageId="56" pageNumber="947" year="1895">Man, 1895: 174</treatmentCitation>
</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD9BA19FF55F87DFD6AF852" blockId="56.[136,775,1854,2106]" box="[136,743,1998,2020]" pageId="56" pageNumber="947">
<treatmentCitationGroup id="E1ACBD6EFFD9BA19FF55F87DFD6AF852" box="[136,743,1998,2020]" pageId="56" pageNumber="947">
<taxonomicName id="06BCE1C3FFD9BA19FF55F87DFD91F852" ID-CoL="84HS6" authorityName="(De Man, 1887)" box="[136,540,1998,2019]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="947" phylum="Arthropoda" rank="subSpecies" species="edwardsi" subGenus="Sesarma" subSpecies="crassimana">
<emphasis id="F3C84652FFD9BA19FF55F87DFF56F852" box="[136,219,1998,2019]" italics="true" pageId="56" pageNumber="947">Sesarma</emphasis>
(
<emphasis id="F3C84652FFD9BA19FF36F87DFEB3F852" box="[235,318,1998,2019]" italics="true" pageId="56" pageNumber="947">Sesarma</emphasis>
) 
<emphasis id="F3C84652FFD9BA19FE90F87DFD91F852" box="[333,540,1998,2019]" italics="true" pageId="56" pageNumber="947">edwardsi crassimana</emphasis>
</taxonomicName>
– 
<treatmentCitation id="401DBC51FFD9BA19FDEAF87DFD69F852" author="Tesch JJ" box="[567,740,1998,2020]" page="148" pageId="56" pageNumber="947" year="1917">
<bibRefCitation id="A52DE7B1FFD9BA19FDEAF87DFD69F852" author="Tesch" box="[567,740,1998,2020]" firstAuthor="Tesch" page="148" pageId="56" pageNumber="947" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch, 1917: 148</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD9BA19FF55F858FF79F7AC" blockId="56.[136,775,1854,2106]" pageId="56" pageNumber="947">
<treatmentCitationGroup id="E1ACBD6EFFD9BA19FF55F858FF79F7AC" pageId="56" pageNumber="947">
<taxonomicName id="06BCE1C3FFD9BA19FF55F858FEDEF7B1" ID-CoL="84HS6" authorityName="De Man" authorityYear="1887" box="[136,339,2027,2048]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="947" phylum="Arthropoda" rank="species" species="crassimana">
<emphasis id="F3C84652FFD9BA19FF55F858FEDEF7B1" box="[136,339,2027,2048]" italics="true" pageId="56" pageNumber="947">Sesarma crassimana</emphasis>
</taxonomicName>
– 
<treatmentCitation id="401DBC51FFD9BA19FEACF859FDA3F7B1" author="Tweedie MWF" box="[369,558,2026,2048]" page="92" pageId="56" pageNumber="947" year="1940">
<bibRefCitation id="A52DE7B1FFD9BA19FEACF859FDA3F7B1" author="Tweedie" box="[369,558,2026,2048]" firstAuthor="Tweedie" page="92" pageId="56" pageNumber="947" pagination="88 - 113" refId="ref78385" refString="Tweedie MWF (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea, Brachyura). Bulletin of the Raffles Museum, 16: 88 - 113, figs. 1 - 12, pl. 24." type="journal article" year="1940">Tweedie, 1940: 92</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFD9BA19FDE6F859FC8EF7B1" author="Tweedie MWF" box="[571,771,2026,2048]" page="343" pageId="56" pageNumber="947" year="1950">
<bibRefCitation id="A52DE7B1FFD9BA19FDE6F859FC8EF7B1" author="Tweedie" box="[571,771,2026,2048]" firstAuthor="Tweedie" page="343" pageId="56" pageNumber="947" pagination="338 - 369" refId="ref78455" refString="Tweedie MWF (1950) Grapsoid crabs from Labuan and Sarawak. Sarawak Museum Journal, 5 (2): 338 - 369, figs. 1 - 9." type="journal article" year="1950">Tweedie, 1950: 343</bibRefCitation>
</treatmentCitation>
, fig. 2b.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD9BA19FF55F797FD2CF78B" blockId="56.[136,775,1854,2106]" box="[136,673,2084,2106]" pageId="56" pageNumber="947">
<treatmentCitationGroup id="E1ACBD6EFFD9BA19FF55F797FD2CF78B" box="[136,673,2084,2106]" pageId="56" pageNumber="947">
<taxonomicName id="06BCE1C3FFD9BA19FF55F797FE40F78B" authorityName="De Man" authorityYear="1887" box="[136,461,2084,2106]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="947" phylum="Arthropoda" rank="species" species="crassimanum" subGenus="Sesarma">
<emphasis id="F3C84652FFD9BA19FF55F797FF56F788" box="[136,219,2084,2105]" italics="true" pageId="56" pageNumber="947">Sesarma</emphasis>
(
<emphasis id="F3C84652FFD9BA19FF36F797FEB3F788" box="[235,318,2084,2105]" italics="true" pageId="56" pageNumber="947">Sesarma</emphasis>
) 
<emphasis id="F3C84652FFD9BA19FE90F796FE40F78B" box="[333,461,2085,2106]" italics="true" pageId="56" pageNumber="947">crassimanum</emphasis>
</taxonomicName>
– 
<treatmentCitation id="401DBC51FFD9BA19FE34F797FD10F78B" author="Serene R" box="[489,669,2084,2106]" page="105" pageId="56" pageNumber="947" year="1968">
<bibRefCitation id="A52DE7B1FFD9BA19FE34F797FD10F78B" author="Serene" box="[489,669,2084,2106]" firstAuthor="Serene" page="105" pageId="56" pageNumber="947" pagination="33 - 112" refId="ref76415" refString="Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a check list of the nonplanctonic marine fauna of South East Asia. Singapore National Academy of Science, Special Publication No. 1: 33 - 112." type="book chapter" year="1968">Serène, 1968: 105</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD9BA19FC9EF96CFBA6F8D6" blockId="56.[835,1475,1759,1925]" pageId="56" pageNumber="947">
<treatmentCitationGroup id="E1ACBD6EFFD9BA19FC9EF96CFBA6F8D6" pageId="56" pageNumber="947">
<taxonomicName id="06BCE1C3FFD9BA19FC9EF96CFBEAF944" ID-CoL="78GDP" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[835,1127,1759,1781]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="947" phylum="Arthropoda" rank="species" species="crassimanum">
<emphasis id="F3C84652FFD9BA19FC9EF96CFBEAF944" box="[835,1127,1759,1781]" italics="true" pageId="56" pageNumber="947">Pseudosesarma crassimanum</emphasis>
</taxonomicName>
– 
<treatmentCitation id="401DBC51FFD9BA19FB5BF96CFA07F945" author="Serene R &amp; Soh CL" box="[1158,1418,1759,1781]" page="399" pageId="56" pageNumber="947" year="1970">
<bibRefCitation id="A52DE7B1FFD9BA19FB5BF96CFA07F945" author="Serene &amp; Soh" box="[1158,1418,1759,1781]" firstAuthor="Serene" page="399" pageId="56" pageNumber="947" pagination="387 - 416" refId="ref76461" refString="Serene R &amp; Soh CL (1970) New IndoPacific genera allied to Sesarma Say 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27: 387 - 416." type="journal article" year="1970">Serène &amp; Soh, 1970: 399</bibRefCitation>
</treatmentCitation>
, 406; 
<treatmentCitation id="401DBC51FFD9BA19FCB4F94FFBCCF8A0" author="Tan CGS &amp; Ng PKL" box="[873,1089,1788,1809]" page="82" pageId="56" pageNumber="947" year="1994">
<bibRefCitation id="A52DE7B1FFD9BA19FCB4F94FFBCCF8A0" author="Tan &amp; Ng" box="[873,1089,1788,1809]" firstAuthor="Tan" page="82" pageId="56" pageNumber="947" pagination="75 - 84" refId="ref77891" refString="Tan CGS &amp; Ng PKL (1994) An annotated checklist of mangrove brachyuran crabs in Malaysia and Singapore. Hydrobiologia, 285: 75 - 84." type="journal article" year="1994">Tan &amp; Ng, 1994: 82</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFD9BA19FB8DF94FFAA0F8A0" author="Naiyanetr P" box="[1104,1325,1788,1810]" page="102" pageId="56" pageNumber="947" year="1998">
<bibRefCitation id="A52DE7B1FFD9BA19FB8DF94FFAA0F8A0" author="Naiyanetr" box="[1104,1325,1788,1810]" firstAuthor="Naiyanetr" page="102" pageId="56" pageNumber="947" refId="ref73221" refString="Naiyanetr P (1998) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Environmental Policy and Planning, Bangkok, 161 pp." type="book" year="1998">Naiyanetr, 1998: 102</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFD9BA19FAE7F94FFC42F89E" author="Cuesta JA &amp; Guerao G &amp; Liu H-C &amp; Schubart CD" page="160" pageId="56" pageNumber="947" year="2006">
<bibRefCitation id="A52DE7B1FFD9BA19FAE7F94FFC42F89E" author="Cuesta" etAl="et al." firstAuthor="Cuesta" page="160" pageId="56" pageNumber="947" pagination="151 - 173" refId="ref68440" refString="Cuesta JA, Guerao G, Liu H-C &amp; Schubart CD (2006) Morphology of the first zoeal stages of eleven Sesarmidae (Crustacea, Brachyura, Thoracotremata) from the Indo-West Pacific, with a summary of familial larval characters. Invertebrate Reproduction &amp; Development, 49 (3): 151 - 173." type="journal article" year="2006">Cuesta et al., 2006: 160</bibRefCitation>
</treatmentCitation>
, fig. 9A–E; 
<treatmentCitation id="401DBC51FFD9BA19FB90F8AAFAACF89F" author="Naiyanetr P" box="[1101,1313,1816,1839]" page="115" pageId="56" pageNumber="947" year="2007">
<bibRefCitation id="A52DE7B1FFD9BA19FB90F8AAFAACF89F" author="Naiyanetr" box="[1101,1313,1816,1839]" firstAuthor="Naiyanetr" page="115" pageId="56" pageNumber="947" refId="ref73256" refString="Naiyanetr P (2007) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, 196 pp." type="book" year="2007">Naiyanetr, 2007: 115</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFD9BA19FAF3F8AAFBCFF8FA" author="Rademacher M &amp; Mengedoht O" page="29" pageId="56" pageNumber="947" year="2011">
<bibRefCitation id="A52DE7B1FFD9BA19FAF3F8AAFBCFF8FA" author="Rademacher &amp; Mengedoht" firstAuthor="Rademacher" page="29" pageId="56" pageNumber="947" refId="ref74843" refString="Rademacher M &amp; Mengedoht O (2011) Krabben-Fibel. Die schonsten Krabben fur das Aquaterrarium. Dahne Verlag, Ettlingen, Germany, 95 pp." type="book" year="2011">Rademacher &amp; Mengedoht, 2011: 29</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="401DBC51FFD9BA19FB8DF885FAD2F8FA" author="Ng PKL &amp; Schubart CD" box="[1104,1375,1845,1867]" page="661" pageId="56" pageNumber="947" year="2017">
<bibRefCitation id="A52DE7B1FFD9BA19FB8DF885FAD2F8FA" author="Ng &amp; Schubart" box="[1104,1375,1845,1867]" firstAuthor="Ng" page="661" pageId="56" pageNumber="947" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart, 2017: 661</bibRefCitation>
</treatmentCitation>
, figs. 1D, E, 2C, D, 8–11, 13.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFD9BA19FC9EF8DCFAE7F835" blockId="56.[835,1475,1759,1925]" box="[835,1386,1903,1925]" pageId="56" pageNumber="947">
<treatmentCitationGroup id="E1ACBD6EFFD9BA19FC9EF8DCFAE7F835" box="[835,1386,1903,1925]" pageId="56" pageNumber="947">
<taxonomicName id="06BCE1C3FFD9BA19FC9EF8DCFBFAF834" ID-CoL="78GDP" authorityName=", Serene &amp; Soh" authorityYear="1970" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[835,1143,1903,1925]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="947" phylum="Arthropoda" rank="genus">
“ 
<emphasis id="F3C84652FFD9BA19FC93F8DCFC68F835" box="[846,997,1903,1924]" italics="true" pageId="56" pageNumber="947">Pseudosesarma</emphasis>
” 
<emphasis id="F3C84652FFD9BA19FC2AF8C3FBFAF834" box="[1015,1143,1904,1925]" italics="true" pageId="56" pageNumber="947">crassimanum</emphasis>
</taxonomicName>
– 
<treatmentCitation id="401DBC51FFD9BA19FB4EF8C3FAEBF835" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1171,1382,1903,1925]" page="222" pageId="56" pageNumber="947" year="2008">
<bibRefCitation id="A52DE7B1FFD9BA19FB4EF8C3FAEBF835" author="Ng" box="[1171,1382,1903,1925]" etAl="et al." firstAuthor="Ng" page="222" pageId="56" pageNumber="947" pagination="1 - 286" refId="ref73754" refString="Ng PKL, Guinot D &amp; Davie PJF (2008 a) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008" yearSuffix="a">Ng et al., 2008a: 222</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD9BA18FC9EF81EFD7CF964" lastPageId="57" lastPageNumber="948" pageId="56" pageNumber="947" type="materials_examined">
<paragraph id="C1039A40FFD9BA18FC9EF81EFD7CF964" blockId="56.[835,1475,1965,2085]" lastBlockId="57.[113,753,189,1749]" lastPageId="57" lastPageNumber="948" pageId="56" pageNumber="947">
<emphasis id="F3C84652FFD9BA19FC9EF81EFBA9F874" bold="true" box="[835,1060,1965,1989]" pageId="56" pageNumber="947">Material examined</emphasis>
. 
<materialsCitation id="71D4901DFFD9BA19FBEBF81DFA32F7B4" ID-GBIF-Occurrence="3763237410" collectionCode="NHM" collectorName="J. Anderson" country="Myanmar" location="Myanmar" pageId="56" pageNumber="947" specimenCode="NHM 1886.52" specimenCount="1" typeStatus="lectotype">
<typeStatus id="1E0724E2FFD9BA19FBEBF81DFB25F877" box="[1078,1192,1966,1990]" pageId="56" pageNumber="947" type="lectotype">Lectotype</typeStatus>
: male (16.3 × 
<quantity id="064437A5FFD9BA19FA89F81EFA36F874" box="[1364,1467,1965,1989]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="56" pageNumber="947" unit="mm" value="14.6">14.6 mm</quantity>
) (
<specimenCode id="911A323BFFD9BA19FC96F87EFC7DF854" box="[843,1008,1997,2021]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="56" pageNumber="947" type="Herbarium">NHM 1886.52</specimenCode>
b), mangrove swamps at Zediwon, Mergui Archipelago, 
<collectingCountry id="B9ABDAD0FFD9BA19FC08F85DFBB0F7B7" box="[981,1085,2030,2054]" name="Myanmar" pageId="56" pageNumber="947">Myanmar</collectingCountry>
(= Burma), coll. 
<collectorName id="6C49FF96FFD9BA19FB25F85DFAF7F7B4" box="[1272,1402,2029,2053]" pageId="56" pageNumber="947">J. Anderson</collectorName>
, 1886
</materialsCitation>
. 
<materialsCitation id="71D4901DFFD9BA18FC9EF7BEFF56FF44" ID-GBIF-Occurrence="3763237537" collectionCode="NHM" collectorName="J. Anderson" country="Thailand" lastPageId="57" lastPageNumber="948" location="Myanmar" pageId="56" pageNumber="947" specimenCode="NHM 1886.52" specimenCount="2" specimenCount-female="2" specimenCount-male="1" typeStatus="paralectotype">
<typeStatus id="1E0724E2FFD9BA19FC9EF7BEFC6EF794" box="[835,995,2061,2085]" pageId="56" pageNumber="947" type="paralectotype">Paralectotypes</typeStatus>
: 
<specimenCount id="D7BA51C9FFD9BA19FC30F7BEFBB8F794" box="[1005,1077,2061,2085]" pageId="56" pageNumber="947" type="male">1 male</specimenCount>
(18.8 × 
<quantity id="064437A5FFD9BA19FB4CF7BEFB7EF794" box="[1169,1267,2061,2085]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="56" pageNumber="947" unit="mm" value="16.7">16.7 mm</quantity>
), 
<specimenCount id="D7BA51C9FFD9BA19FADBF7BEFAE3F794" box="[1286,1390,2061,2085]" pageId="56" pageNumber="947" type="female">2 females</specimenCount>
(16.6 × 
<quantity id="064437A5FFD8BA18FFACFF0EFF59FF64" box="[113,212,189,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="57" pageNumber="948" unit="mm" value="14.3">14.3 mm</quantity>
, 12.5 × 11.0 mm) (
<specimenCode id="911A323BFFD8BA18FE78FF0EFDC5FF64" box="[421,584,189,213]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="57" pageNumber="948" type="Herbarium">NHM 1886.52</specimenCode>
c), same data as lectotype.
</materialsCitation>
<materialsCitation id="71D4901DFFD8BA18FF39FF6DFDA2FEE4" ID-GBIF-Occurrence="3763237436" collectingDate="1997-08-12" collectionCode="NHM" collectorName="J. Anderson" country="Thailand" latitude="9.458111" location="Myanmar" longLatPrecision="15" longitude="98.50861" municipality="King Amphoe Suk Sam Lan" pageId="57" pageNumber="948" specimenCode="NHM 1886.52" specimenCount="2" specimenCount-female="2" specimenCount-male="1" stateProvince="Ranong" typeStatus="paralectotype">
<collectingCountry id="B9ABDAD0FFD8BA18FF39FF6DFEE3FF47" box="[228,366,222,246]" name="Thailand" pageId="57" pageNumber="948">THAILAND</collectingCountry>
– 
<specimenCount id="D7BA51C9FFD8BA18FE52FF6EFE57FF44" box="[399,474,221,245]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(15.9 × 
<quantity id="064437A5FFD8BA18FDE3FF6EFD2FFF44" box="[574,674,221,245]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="57" pageNumber="948" unit="mm" value="14.3">14.3 mm</quantity>
) (
<specimenCode id="911A323BFFD8BA18FD66FF6EFF60FEA4" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 2008.0442</specimenCode>
), 
<collectingRegion id="037854A2FFD8BA18FED9FF4DFE40FEA4" box="[260,461,253,278]" country="Thailand" name="Ranong" pageId="57" pageNumber="948">Ranong Province</collectingRegion>
, 
<collectingMunicipality id="2167003AFFD8BA18FE01FF4EFF13FE84" pageId="57" pageNumber="948">King Amphoe Suk Sam Lan</collectingMunicipality>
, 
<location id="C463CC9BFFD8BA18FF74FEADFE8FFE84" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FF74FEADFE8FFE84" box="[169,258,285,309]" country="Thailand" latitude="9.458111" longLatPrecision="15" longitude="98.50861" municipality="King Amphoe Suk Sam Lan" name="Ton Roi" pageId="57" pageNumber="948" stateProvince="Ranong">Ton Roi</location>
waterfall, 
<geoCoordinate id="A488FC87FFD8BA18FEA8FEAFFE76FE84" box="[373,507,284,309]" degrees="9" direction="north" minutes="27" orientation="latitude" pageId="57" pageNumber="948" precision="1" seconds="29.2" value="9.458111">9°27′29.2″N</geoCoordinate>
<geoCoordinate id="A488FC87FFD8BA18FDDFFEAFFD0DFE84" box="[514,640,284,309]" degrees="98" direction="east" minutes="30" orientation="longitude" pageId="57" pageNumber="948" precision="15" seconds="31" value="98.50861">98°30′31″E</geoCoordinate>
, 
<collectingCountry id="B9ABDAD0FFD8BA18FD56FEAEFD60FE84" box="[651,749,285,309]" name="Thailand" pageId="57" pageNumber="948">Thailand</collectingCountry>
, coll. 
<collectorName id="6C49FF96FFD8BA18FF76FE8EFEABFEE4" box="[171,294,317,341]" pageId="57" pageNumber="948">D.C.J. Yeo</collectorName>
et al., 
<date id="B502BC80FFD8BA18FEA4FE8EFDA2FEE4" box="[377,559,317,342]" pageId="57" pageNumber="948" value="1997-08-12">
<collectingDate id="A5464568FFD8BA18FEA4FE8EFDA2FEE4" box="[377,559,317,342]" pageId="57" pageNumber="948" value="1997-08-12">12 August 1997</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FDE3FE8EFEA2FE04" ID-GBIF-Occurrence="3763237503" collectingDate="2017-04" collectionCode="ZRC" country="Thailand" location="Gulf" pageId="57" pageNumber="948" specimenCode="ZRC 2017.0169" specimenCount="2" specimenCount-female="2" specimenCount-male="2" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FDE3FE8EFD19FEE4" box="[574,660,317,341]" pageId="57" pageNumber="948" type="male">2 males</specimenCount>
(21.9 × 
<quantity id="064437A5FFD8BA18FFAFFEEEFF5BFEC4" box="[114,214,349,373]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="57" pageNumber="948" unit="mm" value="19.5">19.5 mm</quantity>
, 18.5 × 
<quantity id="064437A5FFD8BA18FEEEFEEEFE15FEC4" box="[307,408,349,373]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="57" pageNumber="948" unit="mm" value="16.3">16.3 mm</quantity>
), 
<specimenCount id="D7BA51C9FFD8BA18FE73FEEEFD95FEC4" box="[430,536,349,373]" pageId="57" pageNumber="948" type="female">2 females</specimenCount>
(larger 18.0 × 
<quantity id="064437A5FFD8BA18FD1FFEEEFF11FE24" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="57" pageNumber="948" unit="mm" value="15.9">15.9 mm</quantity>
) (
<specimenCode id="911A323BFFD8BA18FF6BFECEFEE0FE24" box="[182,365,381,405]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 2017.0169</specimenCode>
), 
<location id="C463CC9BFFD8BA18FE5EFECEFE39FE24" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FE5EFECEFE39FE24" box="[387,436,381,405]" country="Thailand" name="Gulf" pageId="57" pageNumber="948">Gulf</location>
of 
<collectingCountry id="B9ABDAD0FFD8BA18FE02FECEFDCFFE24" box="[479,578,381,405]" name="Thailand" pageId="57" pageNumber="948">Thailand</collectingCountry>
, coll. aquarium trade, 
<date id="B502BC80FFD8BA18FF68FE2EFEA2FE04" box="[181,303,413,437]" pageId="57" pageNumber="948" value="2017-04">
<collectingDate id="A5464568FFD8BA18FF68FE2EFEA2FE04" box="[181,303,413,437]" pageId="57" pageNumber="948" value="2017-04">April 2017</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="71D4901DFFD8BA18FEE4FE2EFF24FE44" ID-GBIF-Occurrence="3763237419" collectingDate="1959-09-23" collectionCode="ZRC" collectorName="C. L. Soh" country="Singapore" location="Sungei Seletar" pageId="57" pageNumber="948" specimenCode="ZRC 1985.422, 424" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paralectotype">
<collectingCountry id="B9ABDAD0FFD8BA18FEE4FE2EFE42FE04" box="[313,463,413,437]" name="Singapore" pageId="57" pageNumber="948">SINGAPORE</collectingCountry>
– 
<specimenCount id="D7BA51C9FFD8BA18FE31FE2EFDBAFE04" box="[492,567,413,437]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
, 
<specimenCount id="D7BA51C9FFD8BA18FD99FE2EFD21FE04" box="[580,684,413,437]" pageId="57" pageNumber="948" type="female">2 females</specimenCount>
(
<specimenCode id="911A323BFFD8BA18FD66FE2EFF5AFE64" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 1985.422</specimenCode>
– 
<specimenCode id="911A323BFFD8BA18FF38FE0EFE83FE64" box="[229,270,445,469]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">424</specimenCode>
), 
<location id="C463CC9BFFD8BA18FEC3FE0EFE34FE64" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FEC3FE0EFE34FE64" box="[286,441,445,469]" country="Singapore" name="Sungei Seletar" pageId="57" pageNumber="948">Sungei Seletar</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FE2BFE0EFDDBFE64" box="[502,598,445,469]" pageId="57" pageNumber="948">C.L. Soh</collectorName>
, 
<date id="B502BC80FFD8BA18FD82FE0EFF24FE44" pageId="57" pageNumber="948" value="1959-09-23">
<collectingDate id="A5464568FFD8BA18FD82FE0EFF24FE44" pageId="57" pageNumber="948" value="1959-09-23">23 September 1959</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FF6BFE6EFDCEFDA4" ID-GBIF-Occurrence="3763237501" collectingDate="1966-12-31" collectionCode="ZRC" collectorName="C. L. Soh" country="Singapore" location="Sungei Seletar" pageId="57" pageNumber="948" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FF6BFE6EFF72FE44" box="[182,255,477,501]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(17.5 × 15.0 mm) (
<collectionCode id="A7AD0285FFD8BA18FE0EFE6EFD84FE44" box="[467,521,477,501]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
1967.7.21.4), 
<location id="C463CC9BFFD8BA18FD7BFE6EFF33FDA4" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FD7BFE6EFF33FDA4" country="Singapore" name="Sungei Seletar" pageId="57" pageNumber="948">Sungei Seletar</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FEDCFE4EFEE8FDA4" box="[257,357,509,533]" pageId="57" pageNumber="948">C.L. Soh</collectorName>
, 
<date id="B502BC80FFD8BA18FEAFFE4EFDCEFDA4" box="[370,579,509,533]" pageId="57" pageNumber="948" value="1966-12-31">
<collectingDate id="A5464568FFD8BA18FEAFFE4EFDCEFDA4" box="[370,579,509,533]" pageId="57" pageNumber="948" value="1966-12-31">31 December 1966</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FD92FE4EFF53FDE4" ID-GBIF-Occurrence="3763237543" collectionCode="RMNH-D" collectorName="Serene" country="Singapore" location="Singapore" pageId="57" pageNumber="948" specimenCode="RMNH-D23313" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FD92FE4EFD14FDA4" box="[591,665,509,533]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(15.9 × 
<quantity id="064437A5FFD8BA18FFACFDAEFF5CFD84" box="[113,209,541,565]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="57" pageNumber="948" unit="mm" value="14.5">14.5 mm</quantity>
) (
<specimenCode id="911A323BFFD8BA18FF38FDAEFE17FD84" box="[229,410,541,565]" collectionCode="RMNH-D" pageId="57" pageNumber="948">RMNH-D23313</specimenCode>
), probably from 
<collectingCountry id="B9ABDAD0FFD8BA18FD94FDAEFD35FD84" box="[585,696,541,565]" name="Singapore" pageId="57" pageNumber="948">Singapore</collectingCountry>
, don. R. 
<collectorName id="6C49FF96FFD8BA18FF4FFD8EFF53FDE4" box="[146,222,573,597]" pageId="57" pageNumber="948">Serène</collectorName>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FF37FD8EFEEDFDC4" ID-GBIF-Occurrence="3763237460" collectingDate="1966-05-06" collectingDateMax="1967-07-10" collectingDateMin="1966-05-06" collectionCode="ZRC" collectorName="C. L. Soh" country="Singapore" location="Sungei Seletar" pageId="57" pageNumber="948" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FF37FD8EFEC5FDE4" box="[234,328,573,597]" pageId="57" pageNumber="948" type="female">1 female</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FE8AFD8EFE00FDE4" box="[343,397,573,597]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
<date id="B502BC80FFD8BA18FE49FD8EFD8EFDE4" box="[404,515,573,597]" pageId="57" pageNumber="948" value="1967-07-10">
<collectingDate id="A5464568FFD8BA18FE49FD8EFD8EFDE4" box="[404,515,573,597]" pageId="57" pageNumber="948" value="1967-07-10">1967.7.10</collectingDate>
</date>
), 
<location id="C463CC9BFFD8BA18FDCAFD8EFD3BFDE4" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FDCAFD8EFD3BFDE4" box="[535,694,573,597]" country="Singapore" name="Sungei Seletar" pageId="57" pageNumber="948">Sungei Seletar</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FFACFDEEFF5EFDC4" box="[113,211,605,629]" pageId="57" pageNumber="948">C.L. Soh</collectorName>
, 
<date id="B502BC80FFD8BA18FF00FDEEFEEDFDC4" box="[221,352,605,630]" pageId="57" pageNumber="948" value="1966-05-06">
<collectingDate id="A5464568FFD8BA18FF00FDEEFEEDFDC4" box="[221,352,605,630]" pageId="57" pageNumber="948" value="1966-05-06">6 May 1966</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FEB7FDEEFE78FD24" ID-GBIF-Occurrence="3763237397" collectingDate="1966-07-18" collectionCode="ZRC" collectorName="C. L. Soh" country="Singapore" location="Sungei Seletar" pageId="57" pageNumber="948" specimenCount="2" specimenCount-male="2" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FEB7FDEEFE36FDC4" box="[362,443,605,629]" pageId="57" pageNumber="948" type="male">2 males</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FE14FDEEFE72FDC4" box="[457,511,605,629]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
1967.7.10.40), 
<location id="C463CC9BFFD8BA18FD7BFDEEFF33FD24" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FD7BFDEEFF33FD24" country="Singapore" name="Sungei Seletar" pageId="57" pageNumber="948">Sungei Seletar</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FF20FDCEFED3FD24" box="[253,350,637,661]" pageId="57" pageNumber="948">C.L. Soh</collectorName>
, 
<date id="B502BC80FFD8BA18FEB4FDCEFE78FD24" box="[361,501,637,661]" pageId="57" pageNumber="948" value="1966-07-18">
<collectingDate id="A5464568FFD8BA18FEB4FDCEFE78FD24" box="[361,501,637,661]" pageId="57" pageNumber="948" value="1966-07-18">18 July 1966</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FE22FDCEFD61FD04" ID-GBIF-Occurrence="3763237343" collectingDate="1966-02-18" collectionCode="ZRC" collectorName="C. L. Soh" country="Singapore" location="Mak Wai" municipality="Simpang River" pageId="57" pageNumber="948" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FE22FDCEFDCBFD24" box="[511,582,637,661]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FD8EFDCEFD04FD24" box="[595,649,637,661]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
number), 
<collectingMunicipality id="2167003AFFD8BA18FFACFD2EFE99FD04" box="[113,276,669,693]" pageId="57" pageNumber="948">Simpang River</collectingMunicipality>
, 
<location id="C463CC9BFFD8BA18FEC3FD2EFE0EFD04" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FEC3FD2EFE0EFD04" box="[286,387,669,693]" country="Singapore" municipality="Simpang River" name="Mak Wai" pageId="57" pageNumber="948">Mak Wai</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FE1FFD2EFDAEFD04" box="[450,547,669,693]" pageId="57" pageNumber="948">C.L. Soh</collectorName>
, 
<date id="B502BC80FFD8BA18FDF0FD2EFD61FD04" box="[557,748,669,693]" pageId="57" pageNumber="948" value="1966-02-18">
<collectingDate id="A5464568FFD8BA18FDF0FD2EFD61FD04" box="[557,748,669,693]" pageId="57" pageNumber="948" value="1966-02-18">18 February 1966</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FFACFD0EFF76FD44" ID-GBIF-Occurrence="3763237433" collectingDate="1966-03-29" collectionCode="ZRC" collectorName="Sungei Seletar" country="Singapore" pageId="57" pageNumber="948" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FFACFD0EFF53FD64" box="[113,222,701,725]" pageId="57" pageNumber="948" type="female">2 females</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FF2DFD0EFEABFD64" box="[240,294,701,725]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
1973.11.2.493–494), 
<collectorName id="6C49FF96FFD8BA18FDFCFD0EFD4BFD64" box="[545,710,701,725]" pageId="57" pageNumber="948">Sungei Seletar</collectorName>
, 
<date id="B502BC80FFD8BA18FD08FD0EFF76FD44" pageId="57" pageNumber="948" value="1966-03-29">
<collectingDate id="A5464568FFD8BA18FD08FD0EFF76FD44" pageId="57" pageNumber="948" value="1966-03-29">29 March 1966</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FED4FD6EFD60FD44" ID-GBIF-Occurrence="3763237315" box="[265,749,733,757]" collectionCode="ZRC" collectorName="C. L. Soh" country="Singapore" pageId="57" pageNumber="948" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FED4FD6EFED8FD44" box="[265,341,733,757]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FEBBFD6EFE11FD44" box="[358,412,733,757]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
1967.7.10.39), coll. 
<collectorName id="6C49FF96FFD8BA18FD5BFD6EFD60FD44" box="[646,749,733,757]" pageId="57" pageNumber="948">C.L. Soh</collectorName>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FFACFD4EFD61FCA4" ID-GBIF-Occurrence="3763237535" box="[113,748,765,789]" collectionCode="ZRC" collectorName="C. L. Soh" country="Singapore" pageId="57" pageNumber="948" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FFACFD4EFF31FCA4" box="[113,188,765,789]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FF11FD4EFE8FFCA4" box="[204,258,765,789]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
1971.9.22.10), no other data, coll. 
<collectorName id="6C49FF96FFD8BA18FD5AFD4EFD61FCA4" box="[647,748,765,789]" pageId="57" pageNumber="948">C.L. Soh</collectorName>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FFACFCAEFDB0FC84" ID-GBIF-Occurrence="3763237568" box="[113,573,797,821]" collectionCode="ZRC" country="Malaysia" location="MALAYSIA" pageId="57" pageNumber="948" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FFACFCAEFF4CFC84" box="[113,193,797,821]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
, 
<specimenCount id="D7BA51C9FFD8BA18FF0DFCAEFEB0FC84" box="[208,317,797,821]" pageId="57" pageNumber="948" type="female">3 females</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FE92FCAEFE04FC84" box="[335,393,797,821]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
), no other data.
</materialsCitation>
<materialsCitation id="71D4901DFFD8BA18FD9AFCAEFD58FCC4" ID-GBIF-Occurrence="3763237376" collectingDate="2002-08-30" collectionCode="ZRC" collectorName="T. M. Leong" country="Malaysia" location="MALAYSIA" municipality="Mawai" pageId="57" pageNumber="948" specimenCode="ZRC 2003.0054" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Johor" typeStatus="paralectotype">
PENINSULAR 
<collectingCountry id="B9ABDAD0FFD8BA18FFAFFC8EFE8CFCE4" box="[114,257,829,853]" name="Malaysia" pageId="57" pageNumber="948">MALAYSIA</collectingCountry>
– 
<specimenCount id="D7BA51C9FFD8BA18FEFEFC8EFEFDFCE4" box="[291,368,829,853]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(
<specimenCode id="911A323BFFD8BA18FE5CFC8EFDB7FCE4" box="[385,570,829,853]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 2003.0054</specimenCode>
), 
<collectingRegion id="037854A2FFD8BA18FD8CFC8EFD1DFCE4" box="[593,656,829,853]" country="Malaysia" name="Johor" pageId="57" pageNumber="948">Johor</collectingRegion>
, 
<collectingMunicipality id="2167003AFFD8BA18FD42FC8EFD60FCE4" box="[671,749,829,853]" pageId="57" pageNumber="948">Mawai</collectingMunicipality>
, 
<location id="C463CC9BFFD8BA18FFAFFCEEFEB1FCC4" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FFAFFCEEFEB1FCC4" box="[114,316,861,885]" country="Malaysia" municipality="Mawai" name="Sungei Ulu Sedili" pageId="57" pageNumber="948" stateProvince="Johor">Sungei Ulu Sedili</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FE58FCEDFD83FCC7" box="[389,526,862,886]" pageId="57" pageNumber="948">T.M. Leong</collectorName>
, 
<date id="B502BC80FFD8BA18FDC1FCEEFD58FCC4" box="[540,725,861,886]" pageId="57" pageNumber="948" value="2002-08-30">
<collectingDate id="A5464568FFD8BA18FDC1FCEEFD58FCC4" box="[540,725,861,886]" pageId="57" pageNumber="948" value="2002-08-30">30 August 2002</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FD3EFCEEFD58FC04" ID-GBIF-Occurrence="3763237566" collectingDate="1999-09-30" collectionCode="ZRC" collectorName="C. D. Schubart" country="Malaysia" location="Sungei Benut Cintom" pageId="57" pageNumber="948" specimenCode="ZRC 2010.1768" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Johor" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FD3EFCEEFF39FC24" pageId="57" pageNumber="948" type="male">4 males</specimenCount>
, 
<specimenCount id="D7BA51C9FFD8BA18FF1FFCCEFEA3FC24" box="[194,302,893,917]" pageId="57" pageNumber="948" type="female">3 females</specimenCount>
(
<specimenCode id="911A323BFFD8BA18FEE2FCCEFE7AFC24" box="[319,503,893,917]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 2010.1768</specimenCode>
), 
<collectingRegion id="037854A2FFD8BA18FDD0FCCEFDC1FC24" box="[525,588,893,917]" country="Malaysia" name="Johor" pageId="57" pageNumber="948">Johor</collectingRegion>
, 
<location id="C463CC9BFFD8BA18FD87FCCEFF48FC04" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FD87FCCEFF48FC04" country="Malaysia" name="Sungei Benut Cintom" pageId="57" pageNumber="948" stateProvince="Johor">Sungei Benut Cintom</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FED1FC2EFE25FC04" box="[268,424,925,949]" pageId="57" pageNumber="948">C.D. Schubart</collectorName>
et al., 
<date id="B502BC80FFD8BA18FE27FC2EFD58FC04" box="[506,725,925,949]" pageId="57" pageNumber="948" value="1999-09-30">
<collectingDate id="A5464568FFD8BA18FE27FC2EFD58FC04" box="[506,725,925,949]" pageId="57" pageNumber="948" value="1999-09-30">30 September 1999</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FD3EFC2EFE68FC44" ID-GBIF-Occurrence="3763237521" collectionCode="ZRC" collectorName="M. W. F. Tweedie" country="Malaysia" location="Sedili River" pageId="57" pageNumber="948" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Johor" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FD3EFC2EFF3EFC64" pageId="57" pageNumber="948" type="male">2 males</specimenCount>
, 
<specimenCount id="D7BA51C9FFD8BA18FF1FFC0EFEA9FC64" box="[194,292,957,981]" pageId="57" pageNumber="948" type="female">1 female</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FEE8FC0EFEE6FC64" box="[309,363,957,981]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
1964.9.25.210–212), 
<location id="C463CC9BFFD8BA18FDBEFC0EFD61FC64" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FDBEFC0EFD61FC64" box="[611,748,957,981]" country="Malaysia" name="Sedili River" pageId="57" pageNumber="948" stateProvince="Johor">Sedili River</location>
, 
<collectingRegion id="037854A2FFD8BA18FFAFFC6EFF3DFC44" box="[114,176,989,1013]" country="Malaysia" name="Johor" pageId="57" pageNumber="948">Johor</collectingRegion>
, coll. 
<collectorName id="6C49FF96FFD8BA18FF30FC6DFE2EFC44" box="[237,419,989,1013]" pageId="57" pageNumber="948">M.W.F. Tweedie</collectorName>
, 1938
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FE32FC6EFF26FB84" ID-GBIF-Occurrence="3763237449" collectingDate="1999-06-25" collectionCode="ZRC" collectorName="H. H. Tan" country="Malaysia" location="Sungei Keliling" municipality="Pulau Tioman" pageId="57" pageNumber="948" specimenCode="ZRC 1999.990" specimenCount="1" specimenCount-male="1" stateProvince="Johor" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FE32FC6EFDBBFC44" box="[495,566,989,1013]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(
<specimenCode id="911A323BFFD8BA18FD9FFC6EFD68FC44" box="[578,741,989,1013]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 1999.990</specimenCode>
), 
<collectingMunicipality id="2167003AFFD8BA18FFACFC4EFE9FFBA4" box="[113,274,1021,1045]" pageId="57" pageNumber="948">Pulau Tioman</collectingMunicipality>
, 
<location id="C463CC9BFFD8BA18FEFCFC4EFE59FBA4" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FEFCFC4EFE59FBA4" box="[289,468,1021,1045]" country="Malaysia" municipality="Pulau Tioman" name="Sungei Keliling" pageId="57" pageNumber="948" stateProvince="Johor">Sungei Keliling</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FDC0FC4DFD07FBA4" box="[541,650,1022,1045]" pageId="57" pageNumber="948">H.H. Tan</collectorName>
, 
<date id="B502BC80FFD8BA18FD44FC4EFF26FB84" pageId="57" pageNumber="948" value="1999-06-25">
<collectingDate id="A5464568FFD8BA18FD44FC4EFF26FB84" pageId="57" pageNumber="948" value="1999-06-25">25 June 1999</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FF67FBAEFE4AFBC4" ID-GBIF-Occurrence="3763237514" collectingDate="1996-01-27" collectingDateMax="1996-01-28" collectingDateMin="1996-01-27" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Malaysia" location="Sungei Keliling" municipality="Pulau Tioman" pageId="57" pageNumber="948" specimenCode="ZRC 2011.1012" specimenCount="9" specimenCount-female="8" specimenCount-male="1" stateProvince="Johor" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FF67FBAEFE85FB84" box="[186,264,1053,1077]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
, 
<specimenCount id="D7BA51C9FFD8BA18FECAFBAEFE0EFB84" box="[279,387,1053,1077]" pageId="57" pageNumber="948" type="female">8 females</specimenCount>
(largest 20.2 × 17.0 mm) (
<specimenCode id="911A323BFFD8BA18FD66FBAEFF67FBE4" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 2011.1012</specimenCode>
), 
<collectingMunicipality id="2167003AFFD8BA18FEDDFB8EFE12FBE4" box="[256,415,1085,1109]" pageId="57" pageNumber="948">Pulau Tioman</collectingMunicipality>
, 
<location id="C463CC9BFFD8BA18FE73FB8EFDEDFBE4" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FE73FB8EFDEDFBE4" box="[430,608,1085,1109]" country="Malaysia" municipality="Pulau Tioman" name="Sungei Keliling" pageId="57" pageNumber="948" stateProvince="Johor">Sungei Keliling</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FD74FB8DFF1EFBC7" pageId="57" pageNumber="948">P.K.L. Ng</collectorName>
et al., 
<date id="B502BC80FFD8BA18FF39FBEEFE4AFBC4" box="[228,455,1117,1142]" pageId="57" pageNumber="948" value="1996-01-27" valueMax="1996-01-28" valueMin="1996-01-27">
<collectingDate id="A5464568FFD8BA18FF39FBEEFE4AFBC4" box="[228,455,1117,1142]" pageId="57" pageNumber="948" value="1996-01-27" valueMax="1996-01-28" valueMin="1996-01-27">27–28 January 1996</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FE08FBEEFF37FB24" ID-GBIF-Occurrence="3763237326" collectionCode="ZRC" country="Malaysia" pageId="57" pageNumber="948" specimenCount="1" specimenCount-male="1" stateProvince="Johor" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FE08FBEEFDACFBC4" box="[469,545,1117,1141]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(12.6 × 
<quantity id="064437A5FFD8BA18FD58FBEEFD64FBC4" box="[645,745,1117,1141]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="57" pageNumber="948" unit="mm" value="10.5">10.5 mm</quantity>
) (
<collectionCode id="A7AD0285FFD8BA18FFA4FBCEFF3FFB24" box="[121,178,1149,1173]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="71D4901DFFD8BA18FF19FBCEFE54FB64" ID-GBIF-Occurrence="3763237303" collectingDate="2003-08-19" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Malaysia" location="Sungei Keliling" municipality="Pulau Tioman" pageId="57" pageNumber="948" specimenCode="ZRC 2016.0276" specimenCount="1" specimenCount-female="1" stateProvince="Johor" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FF19FBCEFEACFB24" box="[196,289,1149,1173]" pageId="57" pageNumber="948" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFD8BA18FEF3FBCEFE6DFB24" box="[302,480,1149,1173]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 2016.0276</specimenCode>
), small freshwater stream at beginning of mangroves, 
<collectingMunicipality id="2167003AFFD8BA18FE40FB2EFDBAFB04" box="[413,567,1181,1205]" pageId="57" pageNumber="948">Pulau Tioman</collectingMunicipality>
, 
<location id="C463CC9BFFD8BA18FD9CFB2EFD61FB04" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FD9CFB2EFD61FB04" box="[577,748,1181,1205]" country="Malaysia" municipality="Pulau Tioman" name="Sungei Keliling" pageId="57" pageNumber="948" stateProvince="Johor">Sungei Keliling</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FF74FB0DFE97FB67" box="[169,282,1214,1238]" pageId="57" pageNumber="948">P.K.L. Ng</collectorName>
, 
<date id="B502BC80FFD8BA18FEFAFB0EFE54FB64" box="[295,473,1213,1238]" pageId="57" pageNumber="948" value="2003-08-19">
<collectingDate id="A5464568FFD8BA18FEFAFB0EFE54FB64" box="[295,473,1213,1238]" pageId="57" pageNumber="948" value="2003-08-19">19 August 2003</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FE38FB0EFE54FB44" ID-GBIF-Occurrence="3763237487" collectingDate="1959-07-31" collectionCode="ZRC" collectorName="D. S. Johnson" country="Malaysia" pageId="57" pageNumber="948" specimenCode="ZRC 1985.425" specimenCount="1" specimenCount-male="1" stateProvince="Johor" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FE38FB0EFDA2FB64" box="[485,559,1213,1237]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(
<specimenCode id="911A323BFFD8BA18FDE2FB0EFD68FB64" box="[575,741,1213,1237]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 1985.425</specimenCode>
), coll. 
<collectorName id="6C49FF96FFD8BA18FF75FB6EFEB4FB44" box="[168,313,1245,1269]" pageId="57" pageNumber="948">D.S. Johnson</collectorName>
, 
<date id="B502BC80FFD8BA18FE98FB6EFE59FB44" box="[325,468,1245,1269]" pageId="57" pageNumber="948" value="1959-07-31">
<collectingDate id="A5464568FFD8BA18FE98FB6EFE59FB44" box="[325,468,1245,1269]" pageId="57" pageNumber="948" value="1959-07-31">31 July 1959</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="71D4901DFFD8BA18FE3CFB6EFEA2FA84" ID-GBIF-Occurrence="3763237490" collectingDate="1998-06-25" collectionCode="ZRC" collectorName="H. H. Tan" country="Malaysia" location="Sungei Keliling" municipality="Pulau Tioman" pageId="57" pageNumber="948" specimenCode="ZRC 1999.957" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Johor" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FE3CFB6EFDBAFB44" box="[481,567,1245,1269]" pageId="57" pageNumber="948" type="male">5 males</specimenCount>
, 
<specimenCount id="D7BA51C9FFD8BA18FD9EFB6EFD21FB44" box="[579,684,1245,1269]" pageId="57" pageNumber="948" type="female">3 females</specimenCount>
(
<specimenCode id="911A323BFFD8BA18FD66FB6EFF54FAA4" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 1999.957</specimenCode>
), 
<collectingMunicipality id="2167003AFFD8BA18FF36FB4EFE0EFAA4" box="[235,387,1277,1301]" pageId="57" pageNumber="948">Pulau Tioman</collectingMunicipality>
, 
<location id="C463CC9BFFD8BA18FE51FB4EFDBBFAA4" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FE51FB4EFDBBFAA4" box="[396,566,1277,1301]" country="Malaysia" municipality="Pulau Tioman" name="Sungei Keliling" pageId="57" pageNumber="948" stateProvince="Johor">Sungei Keliling</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FDA9FB4DFD5AFAA4" box="[628,727,1278,1301]" pageId="57" pageNumber="948">H.H. Tan</collectorName>
et al., 
<date id="B502BC80FFD8BA18FF47FAAEFEA2FA84" box="[154,303,1309,1333]" pageId="57" pageNumber="948" value="1998-06-25">
<collectingDate id="A5464568FFD8BA18FF47FAAEFEA2FA84" box="[154,303,1309,1333]" pageId="57" pageNumber="948" value="1998-06-25">25 June 1998</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="71D4901DFFD8BA18FEE7FAAEFD81FAE4" ID-GBIF-Occurrence="3763237415" collectionCode="MNHG" collectorName="Bedot &amp; Pichet" country="Malaysia" location="SARAWAK" pageId="57" pageNumber="948" specimenCount="1" specimenCount-male="1">
<collectingCountry id="B9ABDAD0FFD8BA18FEE7FAAEFE4CFA84" box="[314,449,1309,1333]" name="Malaysia" pageId="57" pageNumber="948">SARAWAK</collectingCountry>
– 
<specimenCount id="D7BA51C9FFD8BA18FE02FAAEFDA5FA84" box="[479,552,1309,1333]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(14.0 × 
<quantity id="064437A5FFD8BA18FD5AFAAEFD64FA84" box="[647,745,1309,1333]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="57" pageNumber="948" unit="mm" value="11.9">11.9 mm</quantity>
) (
<collectionCode id="A7AD0285FFD8BA18FFA4FA8EFF5DFAE4" box="[121,208,1341,1365]" pageId="57" pageNumber="948">MNHG</collectionCode>
), coll. 
<collectorName id="6C49FF96FFD8BA18FECBFA8EFEDBFAE4" box="[278,342,1341,1365]" pageId="57" pageNumber="948">Bedot</collectorName>
&amp; 
<collectorName id="6C49FF96FFD8BA18FEABFA8EFE36FAE4" box="[374,443,1341,1365]" pageId="57" pageNumber="948">Pichet</collectorName>
, 1800s;
</materialsCitation>
<materialsCitation id="71D4901DFFD8BA18FDCFFA8EFF27FA24" ID-GBIF-Occurrence="3763237504" collectionCode="ZRC" collectorName="Tweedie MWF" country="Malaysia" location="Kuching" pageId="57" pageNumber="948" specimenCount="24" specimenCount-female="10" specimenCount-male="14" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FDCFFA8EFDFEFAE4" box="[530,627,1341,1365]" pageId="57" pageNumber="948" type="male">14 males</specimenCount>
, 
<specimenCount id="D7BA51C9FFD8BA18FDA0FA8EFD7DFAE4" box="[637,752,1341,1365]" pageId="57" pageNumber="948" type="female">10 females</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FFA4FAEEFF22FAC4" box="[121,175,1373,1397]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
1964.9.25.368–379), 
<location id="C463CC9BFFD8BA18FE45FAEEFE7AFAC4" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FE45FAEEFE7AFAC4" box="[408,503,1373,1397]" country="Malaysia" name="Kuching" pageId="57" pageNumber="948">Kuching</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FDE9FAEDFD05FAC4" box="[564,648,1374,1397]" name="Tweedie MWF" pageId="57" pageNumber="948">M.W.F.</collectorName>
<bibRefCitation id="A52DE7B1FFD8BA18FD53FAEEFF27FA24" author="Tweedie" firstAuthor="Tweedie" pageId="57" pageNumber="948" pagination="338 - 369" refId="ref78455" refString="Tweedie MWF (1950) Grapsoid crabs from Labuan and Sarawak. Sarawak Museum Journal, 5 (2): 338 - 369, figs. 1 - 9." type="journal article" year="1950">Tweedie, 1950</bibRefCitation>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FF6AFACEFEF5FA44" ID-GBIF-Occurrence="3763237378" collectingDate="1996-09-02" collectionCode="ZRC" collectorName="Sungei Gading &amp; H. H. Tan" country="Malaysia" county="Cape Pelandok" location="Landa town" municipality="Kampung Pandan" pageId="57" pageNumber="948" specimenCode="ZRC 1999.851" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FF6AFACEFE8DFA24" box="[183,256,1405,1429]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
(18.9 × 
<quantity id="064437A5FFD8BA18FEBDFACEFE4FFA24" box="[352,450,1405,1429]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="57" pageNumber="948" unit="mm" value="16.7">16.7 mm</quantity>
) (
<specimenCode id="911A323BFFD8BA18FE04FACEFDF2FA24" box="[473,639,1405,1429]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 1999.851</specimenCode>
), stream 
<quantity id="064437A5FFD8BA18FD3EFACEFF1EFA04" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="57" pageNumber="948" unit="km" value="3.0">3 km</quantity>
before turn off to 
<collectingCounty id="2862E2CCFFD8BA18FE8AFA2EFE7AFA04" box="[343,503,1437,1461]" pageId="57" pageNumber="948">Cape Pelandok</collectingCounty>
and 
<collectingMunicipality id="2167003AFFD8BA18FDF6FA2DFD61FA04" box="[555,748,1437,1462]" pageId="57" pageNumber="948">Kampung Pandan</collectingMunicipality>
, after 
<location id="C463CC9BFFD8BA18FF70FA0EFEB9FA64" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FF70FA0EFEB9FA64" box="[173,308,1469,1493]" country="Malaysia" county="Cape Pelandok" municipality="Kampung Pandan" name="Landa town" pageId="57" pageNumber="948">Landa town</location>
, drains from 
<collectorName id="6C49FF96FFD8BA18FE10FA0EFDF9FA64" box="[461,628,1469,1493]" pageId="57" pageNumber="948">Sungei Gading</collectorName>
, coll. 
<collectorName id="6C49FF96FFD8BA18FD61FA0DFF13FA44" pageId="57" pageNumber="948">H.H. Tan</collectorName>
, 
<date id="B502BC80FFD8BA18FF71FA6EFEF5FA44" box="[172,376,1501,1525]" pageId="57" pageNumber="948" value="1996-09-02">
<collectingDate id="A5464568FFD8BA18FF71FA6EFEF5FA44" box="[172,376,1501,1525]" pageId="57" pageNumber="948" value="1996-09-02">2 September 1996</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="71D4901DFFD8BA18FE5BFA6EFDBBF9A4" ID-GBIF-Occurrence="3763237556" collectionCode="ZRC" collectorName="L. K. Charles" country="Indonesia" location="Saribas" municipality="Stambak" pageId="57" pageNumber="948" specimenCount="3" specimenCount-male="3" typeStatus="paralectotype">
<specimenCount id="D7BA51C9FFD8BA18FE5BFA6EFE56FA44" box="[390,475,1501,1525]" pageId="57" pageNumber="948" type="male">3 males</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FE30FA6EFDAEFA44" box="[493,547,1501,1525]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
1964.9.28.14–16), 
<collectingMunicipality id="2167003AFFD8BA18FFACFA4EFF5CF9A4" box="[113,209,1533,1557]" pageId="57" pageNumber="948">Stambak</collectingMunicipality>
, 
<location id="C463CC9BFFD8BA18FF07FA4EFEA6F9A4" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FF07FA4EFEA6F9A4" box="[218,299,1533,1557]" country="Indonesia" municipality="Stambak" name="Saribas" pageId="57" pageNumber="948">Saribas</location>
, coll. 
<collectorName id="6C49FF96FFD8BA18FEBAFA4DFE7CF9A4" box="[359,497,1533,1557]" pageId="57" pageNumber="948">L.K. Charles</collectorName>
, 1950.
</materialsCitation>
<materialsCitation id="71D4901DFFD8BA18FDE6FA4EFE69F9C4" ID-GBIF-Occurrence="3763237442" collectingDate="1995-09-02" collectionCode="ZRC" collectorName="L. K. Charles" country="Indonesia" location="Saribas" municipality="Stambak" pageId="57" pageNumber="948" specimenCode="ZRC 1999.503" specimenCount="3" specimenCount-male="3" stateProvince="Kalimantan Utara" typeStatus="paralectotype">
<collectingCountry id="B9ABDAD0FFD8BA18FDE6FA4EFD41F9A4" box="[571,716,1533,1557]" name="Indonesia" pageId="57" pageNumber="948">INDONESIA</collectingCountry>
– 
<specimenCount id="D7BA51C9FFD8BA18FD3EFA4EFF22F984" pageId="57" pageNumber="948" type="male">2 males</specimenCount>
(17.2 × 15.0 mm, 14.3 × 
<quantity id="064437A5FFD8BA18FE15F9AEFDA4F984" box="[456,553,1565,1589]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="57" pageNumber="948" unit="mm" value="12.3">12.3 mm</quantity>
) (
<specimenCode id="911A323BFFD8BA18FDE2F9AEFD68F984" box="[575,741,1565,1589]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 1999.503</specimenCode>
), 
<collectingMunicipality id="2167003AFFD8BA18FFACF98EFEA1F9E4" box="[113,300,1597,1621]" pageId="57" pageNumber="948">Tanjung Reolep</collectingMunicipality>
, 
<location id="C463CC9BFFD8BA18FEE6F98EFE54F9E4" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FEE6F98EFE54F9E4" box="[315,473,1597,1621]" country="Indonesia" municipality="Tanjung Reolep" name="Sungai Berau" pageId="57" pageNumber="948" stateProvince="Kalimantan Utara">Sungai Berau</location>
, riverbank, 
<collectingRegion id="037854A2FFD8BA18FDBAF98EFD61F9E4" box="[615,748,1597,1621]" country="Indonesia" name="Kalimantan Utara" pageId="57" pageNumber="948">Kalimantan</collectingRegion>
, coll. 
<collectorName id="6C49FF96FFD8BA18FF74F9EDFE9DF9C4" box="[169,272,1629,1653]" pageId="57" pageNumber="948">R. Diesel</collectorName>
, 
<date id="B502BC80FFD8BA18FEC1F9EEFE69F9C4" box="[284,484,1629,1653]" pageId="57" pageNumber="948" value="1995-09-02">
<collectingDate id="A5464568FFD8BA18FEC1F9EEFE69F9C4" box="[284,484,1629,1653]" pageId="57" pageNumber="948" value="1995-09-02">2 September 1995</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="71D4901DFFD8BA18FE32F9EDFEBCF904" ID-GBIF-Occurrence="3763237453" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Cambodia" location="CAMBODIA" pageId="57" pageNumber="948" specimenCode="ZRC 2017.1045" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paralectotype">
<collectingCountry id="B9ABDAD0FFD8BA18FE32F9EDFDF4F9C7" box="[495,633,1630,1654]" name="Thailand" pageId="57" pageNumber="948">THAILAND</collectingCountry>
– 
<specimenCount id="D7BA51C9FFD8BA18FD4AF9EEFD60F9C4" box="[663,749,1629,1653]" pageId="57" pageNumber="948" type="male">2 males</specimenCount>
, 
<specimenCount id="D7BA51C9FFD8BA18FFACF9CEFF5BF924" box="[113,214,1661,1685]" pageId="57" pageNumber="948" type="female">1 female</specimenCount>
(
<specimenCode id="911A323BFFD8BA18FF34F9CEFE2AF924" box="[233,423,1661,1685]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC 2017.1045</specimenCode>
), from aquarium trade, coll. 
<collectorName id="6C49FF96FFD8BA18FFAFF92DFF6BF907" box="[114,230,1694,1718]" pageId="57" pageNumber="948">
P.K.L. 
<bibRefCitation id="A52DE7B1FFD8BA18FF1EF92DFF6BF907" author="Ng" box="[195,230,1694,1718]" firstAuthor="Ng" journalOrPublisher="Zootaxa" pageId="57" pageNumber="948" pagination="263 - 270" part="4311" refId="ref73970" refString="Ng PKL, Rani V &amp; Nandan B (2017 a) A new species of Pseudosesarma Serene &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae) from Cochin in southwest India. Zootaxa, 4311 (2), 263 - 270." title="A new species of Pseudosesarma Serene &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae) from Cochin in southwest India" type="journal article" year="2017">Ng</bibRefCitation>
</collectorName>
, 2017.
</materialsCitation>
<materialsCitation id="71D4901DFFD8BA18FEE6F92EFD60F964" ID-GBIF-Occurrence="3763237491" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Cambodia" location="CAMBODIA" pageId="57" pageNumber="948" specimenCode="ZRC 2017.1045" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paralectotype">
<collectingCountry id="B9ABDAD0FFD8BA18FEE6F92EFE40F904" box="[315,461,1693,1717]" name="Cambodia" pageId="57" pageNumber="948">CAMBODIA</collectingCountry>
– 
<specimenCount id="D7BA51C9FFD8BA18FE33F92EFDB1F904" box="[494,572,1693,1717]" pageId="57" pageNumber="948" type="male">1 male</specimenCount>
, 
<specimenCount id="D7BA51C9FFD8BA18FD97F92EFD27F904" box="[586,682,1693,1717]" pageId="57" pageNumber="948" type="female">1 female</specimenCount>
(
<collectionCode id="A7AD0285FFD8BA18FD66F92EFD7CF904" box="[699,753,1693,1717]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="57" pageNumber="948">ZRC</collectionCode>
1965.7.29.52), 
<location id="C463CC9BFFD8BA18FEC8F90EFE0BF964" LSID="urn:lsid:plazi:treatment:49152B56FFD9BA18FF1BF96CFA12FB64:C463CC9BFFD8BA18FEC8F90EFE0BF964" box="[277,390,1725,1749]" country="Thailand" name="Tonle Sap" pageId="57" pageNumber="948">Tonle Sap</location>
, 
<collectingCountry id="B9ABDAD0FFD8BA18FE4EF90EFE41F964" box="[403,460,1725,1749]" name="Thailand" pageId="57" pageNumber="948">Siam</collectingCountry>
, coll. 
<collectorName id="6C49FF96FFD8BA18FDD2F90DFD25F964" box="[527,680,1725,1749]" pageId="57" pageNumber="948">N. Annandale</collectorName>
, 1918
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD8BA18FFACF94EFA2BFE04" pageId="57" pageNumber="948" type="diagnosis">
<paragraph id="C1039A40FFD8BA18FFACF94EFA2DFF44" blockId="57.[113,752,1789,2101]" lastBlockId="57.[813,1451,189,245]" pageId="57" pageNumber="948">
<emphasis id="F3C84652FFD8BA18FFACF94EFF53F8A4" bold="true" box="[113,222,1789,1813]" pageId="57" pageNumber="948">Diagnosis</emphasis>
. Carapace transversely rectangular; frontal margin relatively wider, median concavity separating lobes more distinct; epibranchial tooth distinct, separated from rest of margin by deep notch; posterolateral margins subparallel; outer surface of chela gently convex, covered with small rounded granules, ventral margin of palm sinuous unarmed; suture between male thoracic sternites 3 and 4 distinct; male pleon broadly triangular; male sternopleonal cavity with press-button of pleonal locking mechanism on sternite 5; distal part of G1 gently swollen with chitinous part forming beak-like structure. In life, chela purplish red to yellow, pigmentation reaching to base of yellowish-white fingers.
</paragraph>
<paragraph id="C1039A40FFD8BA18FCF0FEAEFA2BFE04" blockId="57.[813,1452,285,438]" pageId="57" pageNumber="948">
<emphasis id="F3C84652FFD8BA18FCF0FEAEFCF2FE84" bold="true" box="[813,895,285,309]" pageId="57" pageNumber="948">Colour</emphasis>
. In life, smaller specimens have a darker brown carapace with purple chelae and yellowish fingers (
<figureCitation id="598786C5FFD8BA18FAA3FE8EFCF6FEC4" captionStart="Fig" captionStartId="88.[136,171,1875,1897]" captionTargetBox="[136,1474,324,1860]" captionTargetId="figure-7@88.[136,1474,324,1860]" captionTargetPageId="88" captionText="Fig. 56. Colour in life. A, B, Pseudosesarma edwardsii, male (20.5 × 19.3 mm) (ZRC 2016.0608), Langkawi [photograph: Paul Ng]; C–E, P. edwardsii, male (19.4 × 18.2 mm) (SMF), Langkawi, Peninsular Malaysia [photograph: CD Schubart]; F, G, P. crassimanum, (12.6 × 10.5 mm) (ZRC 2010.0035), Tioman, Peninsular Malaysia [photograph: PKL Ng]; H, P. boulengeri, specimen not preserved [photograph: Reza Naderloo]." figureDoi="http://doi.org/10.5281/zenodo.5351399" httpUri="https://zenodo.org/record/5351399/files/figure.png" pageId="57" pageNumber="948">Fig. 56F, G</figureCitation>
). Larger specimens have carapaces which are lighter brown, with the chelae purplish-red to yellow (
<bibRefCitation id="A52DE7B1FFD8BA18FAF9FECEFBB9FE04" author="Rademacher &amp; Mengedoht" firstAuthor="Rademacher" page="29" pageId="57" pageNumber="948" refId="ref74843" refString="Rademacher M &amp; Mengedoht O (2011) Krabben-Fibel. Die schonsten Krabben fur das Aquaterrarium. Dahne Verlag, Ettlingen, Germany, 95 pp." type="book" year="2011">Rademacher &amp; Mengedoht, 2011: 29</bibRefCitation>
) (see also 
<bibRefCitation id="A52DE7B1FFD8BA18FB71FE2DFA17FE04" author="Ng &amp; Schubart" box="[1196,1434,413,438]" firstAuthor="Ng" pageId="57" pageNumber="948" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart, 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD8BA18FCF0FE6EFB38FCE4" pageId="57" pageNumber="948" type="discussion">
<paragraph id="C1039A40FFD8BA18FCF0FE6EFC6EFD84" blockId="57.[813,1452,477,565]" pageId="57" pageNumber="948">
<emphasis id="F3C84652FFD8BA18FCF0FE6EFC1AFE44" bold="true" box="[813,919,477,501]" pageId="57" pageNumber="948">Remarks</emphasis>
. The differences between 
<taxonomicName id="06BCE1C3FFD8BA18FB14FE6DFADAFE44" authorityName="De Man" authorityYear="1887" box="[1225,1367,477,501]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="948" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="F3C84652FFD8BA18FB14FE6DFADAFE44" box="[1225,1367,477,501]" italics="true" pageId="57" pageNumber="948">P. edwardsii</emphasis>
</taxonomicName>
and 
<taxonomicName id="06BCE1C3FFD8BA18FA49FE6DFC48FDA4" baseAuthorityName="De Man" baseAuthorityYear="1887" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="948" phylum="Arthropoda" rank="species" species="crassimanum">
<emphasis id="F3C84652FFD8BA18FA49FE6DFC48FDA4" italics="true" pageId="57" pageNumber="948">P. crassimanum</emphasis>
</taxonomicName>
have been discussed at length by 
<bibRefCitation id="A52DE7B1FFD8BA18FAB4FE4DFC6DFD84" author="Ng &amp; Schubart" firstAuthor="Ng" pageId="57" pageNumber="948" pagination="655 - 669" refId="ref74064" refString="Ng PKL &amp; Schubart CD (2017) On the taxonomy of Pseudosesarma edwardsii (De Man, 1887) and P. crassimanum (De Man, 1887) (Crustacea: Decapoda: Brachyura: Sesarmidae), with description of a new species from Sri Lanka. Raffles Bulletin of Zoology, 65: 655 - 669." type="journal article" year="2017">Ng &amp; Schubart (2017)</bibRefCitation>
.
</paragraph>
<paragraph id="C1039A40FFD8BA18FCF0FDEEFB38FCE4" blockId="57.[813,1452,605,853]" pageId="57" pageNumber="948">
<bibRefCitation id="A52DE7B1FFD8BA18FCF0FDEEFC56FDC4" author="Li" box="[813,987,605,629]" etAl="et al." firstAuthor="Li" page="3" pageId="57" pageNumber="948" refId="ref71682" refString="Li J-J, Shih H-T &amp; Ng PKL (2020) The Taiwanese and Philippine species of the terrestrial crabs Bresedium Serene and Soh, 1970 and Sesarmops Serene and Soh, 1970 (Crustacea: Decapoda: Brachyura), with descriptions of two new species. Zoological Studies, 59: 16. doi: 10.6620 / ZS. 2020.59 - 16, 32 pp." type="book" year="2020">Li et al. (2020: 3</bibRefCitation>
, 29, 30) list “ 
<taxonomicName id="06BCE1C3FFD8BA18FBBBFDEEFA2DFDC4" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[1126,1440,605,629]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="948" phylum="Arthropoda" rank="species" species="crassimanum">
<emphasis id="F3C84652FFD8BA18FBBBFDEEFA2DFDC4" box="[1126,1440,605,629]" italics="true" pageId="57" pageNumber="948">Pseudosesarma crassimanum</emphasis>
</taxonomicName>
” for the molecular tree they used in a study of 
<taxonomicName id="06BCE1C3FFD8BA18FAEAFDCDFB6CFD04" authority="(H. Milne Edwards, 1837)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="948" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="F3C84652FFD8BA18FAEAFDCDFC13FD04" italics="true" pageId="57" pageNumber="948">Sesarmops impressus</emphasis>
(H. Milne Edwards, 1837)
</taxonomicName>
. Its genetic data, however, aligns with what is here regarded as 
<taxonomicName id="06BCE1C3FFD8BA18FA9CFD0DFBCCFD44" authority="(De Man, 1892)" baseAuthorityName="De Man" baseAuthorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Manarma" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="948" phylum="Arthropoda" rank="species" species="moeschii">
<emphasis id="F3C84652FFD8BA18FA9CFD0DFC01FD44" italics="true" pageId="57" pageNumber="948">Manarma moeschii</emphasis>
(
<bibRefCitation id="A52DE7B1FFD8BA18FC46FD6EFBB4FD44" author="De Man" box="[923,1081,733,757]" firstAuthor="De Man" pageId="57" pageNumber="948" pagination="225 - 264" refId="ref72297" refString="Man JG De (1892) Carcinological studies in the Leyden Museum, No. 6. Notes Leyden Museum, 14 (36): 225 - 264, pls. 7 - 10." type="journal article" year="1892">De Man, 1892</bibRefCitation>
)
</taxonomicName>
(see later). The material that was used for the study by 
<bibRefCitation id="A52DE7B1FFD8BA18FBFEFD4EFB40FCA4" author="Li" box="[1059,1229,765,789]" etAl="et al." firstAuthor="Li" pageId="57" pageNumber="948" refId="ref71682" refString="Li J-J, Shih H-T &amp; Ng PKL (2020) The Taiwanese and Philippine species of the terrestrial crabs Bresedium Serene and Soh, 1970 and Sesarmops Serene and Soh, 1970 (Crustacea: Decapoda: Brachyura), with descriptions of two new species. Zoological Studies, 59: 16. doi: 10.6620 / ZS. 2020.59 - 16, 32 pp." type="book" year="2020">Li et al. (2020)</bibRefCitation>
originated from the aquarium trade and was misidentified, they are actually 
<taxonomicName id="06BCE1C3FFD8BA18FA53FCADFC01FCE4" baseAuthorityName="De Man" baseAuthorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Manarma" kingdom="Animalia" order="Decapoda" pageId="57" pageNumber="948" phylum="Arthropoda" rank="species" species="moeschii">
<emphasis id="F3C84652FFD8BA18FA53FCADFC01FCE4" italics="true" pageId="57" pageNumber="948">M. moeschii</emphasis>
</taxonomicName>
(H.-T. Shih, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD8BA18FCF0FCCEFAA4FBC4" pageId="57" pageNumber="948" type="biology_ecology">
<paragraph id="C1039A40FFD8BA18FCF0FCCEFAA4FBC4" blockId="57.[813,1452,893,1141]" pageId="57" pageNumber="948">
<emphasis id="F3C84652FFD8BA18FCF0FCCEFC0EFC24" bold="true" box="[813,899,893,917]" pageId="57" pageNumber="948">Biology</emphasis>
. This species seems to prefer to hide under small rocks and vegetation along the banks of coastal freshwater streams leading to the sea, but they have also been found in the rearward mangrove zones. They appear to be mainly nocturnal. On Pulau Tioman in 
<collectingCountry id="B9ABDAD0FFD8BA18FB4CFC4EFB7BFBA4" box="[1169,1270,1021,1045]" name="Malaysia" pageId="57" pageNumber="948">Malaysia</collectingCountry>
, they have been observed to climb up shrubs and small trees at night, apparently foraging for young shoots and buds. The larvae have been reported on by 
<bibRefCitation id="A52DE7B1FFD8BA18FB96FBEEFAA8FBC4" author="Cuesta" box="[1099,1317,1117,1141]" etAl="et al." firstAuthor="Cuesta" pageId="57" pageNumber="948" pagination="151 - 173" refId="ref68440" refString="Cuesta JA, Guerao G, Liu H-C &amp; Schubart CD (2006) Morphology of the first zoeal stages of eleven Sesarmidae (Crustacea, Brachyura, Thoracotremata) from the Indo-West Pacific, with a summary of familial larval characters. Invertebrate Reproduction &amp; Development, 49 (3): 151 - 173." type="journal article" year="2006">Cuesta et al. (2006)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFD8BA18FCF0FB2EFA12FB64" pageId="57" pageNumber="948" type="distribution">
<paragraph id="C1039A40FFD8BA18FCF0FB2EFA12FB64" blockId="57.[813,1450,1181,1237]" pageId="57" pageNumber="948">
<emphasis id="F3C84652FFD8BA18FCF0FB2EFC49FB04" bold="true" box="[813,964,1181,1205]" pageId="57" pageNumber="948">Distribution</emphasis>
. Known only from 
<collectingCountry id="B9ABDAD0FFD8BA18FB65FB2EFAAFFB04" box="[1208,1314,1181,1205]" name="Thailand" pageId="57" pageNumber="948">Thailand</collectingCountry>
, Peninsular 
<collectingCountry id="B9ABDAD0FFD8BA18FCF3FB0EFC1FFB64" box="[814,914,1213,1237]" name="Malaysia" pageId="57" pageNumber="948">Malaysia</collectingCountry>
, 
<collectingCountry id="B9ABDAD0FFD8BA18FC42FB0EFB83FB64" box="[927,1038,1213,1237]" name="Singapore" pageId="57" pageNumber="948">Singapore</collectingCountry>
, and Borneo (
<bibRefCitation id="A52DE7B1FFD8BA18FB75FB0EFAC0FB64" author="Tweedie" box="[1192,1357,1213,1237]" firstAuthor="Tweedie" pageId="57" pageNumber="948" pagination="88 - 113" refId="ref78385" refString="Tweedie MWF (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea, Brachyura). Bulletin of the Raffles Museum, 16: 88 - 113, figs. 1 - 12, pl. 24." type="journal article" year="1940">Tweedie, 1940</bibRefCitation>
, 
<bibRefCitation id="A52DE7B1FFD8BA18FA84FB0EFA1EFB64" author="Tweedie" box="[1369,1427,1213,1237]" firstAuthor="Tweedie" pageId="57" pageNumber="948" pagination="338 - 369" refId="ref78455" refString="Tweedie MWF (1950) Grapsoid crabs from Labuan and Sarawak. Sarawak Museum Journal, 5 (2): 338 - 369, figs. 1 - 9." type="journal article" year="1950">1950</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>