182 lines
20 KiB
XML
182 lines
20 KiB
XML
<document id="512D66BBBE1BD9D59870C3A11F513068" ID-DOI="10.1111/j.1096-3642.2009.00526.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443023" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454731098" checkinUser="felipe" docAuthor="Liew, Thor-Seng, Schilthuizen, Menno & Vermeulen, Jaap Jan" docDate="2009" docId="03F9878AFF96E8190FB98FA5239FFE34" docLanguage="en" docName="j.1096-3642.2009.00526.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Everettia occidentalis Liew & Schilthuizen & Vermeulen 2009, SP. NOV." docType="treatment" docVersion="5" lastPageNumber="541" masterDocId="FFC0FFF2FF8FE8030C238C132365FFCA" masterDocTitle="Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo" masterLastPageNumber="550" masterPageNumber="515" pageNumber="540" updateTime="1699276879878" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B3C2ABF70E61E3F2711DBBDC289BF415">Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo</mods:title>
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<mods:roleTerm id="BF92A5C9BBBED5048DEE8E28C60C4F3C">Author</mods:roleTerm>
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<mods:namePart id="F31570404B04C170B8F1901377734433">Liew, Thor-Seng</mods:namePart>
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<mods:roleTerm id="B64B056B734FD842479EADABE4F2190D">Author</mods:roleTerm>
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<mods:namePart id="EEE36F3E3638345DEA6DE935B9A7B187">Schilthuizen, Menno</mods:namePart>
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<mods:roleTerm id="A1CED092D85093ECEF013C2CBA50D9F6">Author</mods:roleTerm>
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<mods:namePart id="1C3AA0E1D69626C8ED44A8A3880AD875">Vermeulen, Jaap Jan</mods:namePart>
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<mods:typeOfResource id="8A075AAC5BD4FE54888D55BD3F0B07F8">text</mods:typeOfResource>
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<mods:title id="91A4E408B5D44AD7FD6338E7E6B7B9D3">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="4DBACA9E6EDE0514175F9D905BBA00DB">
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<mods:date id="020AA6E292A79B5C822F784D312E43C7">2009</mods:date>
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<mods:number id="E82598653E95B9365A18AB306E0495ED">2009-10-26</mods:number>
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<mods:detail id="720BC60971DAEB466F00E87F567B75D7" type="volume">
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<mods:number id="07BE0D392566C41EC94325079CAD4EEC">157</mods:number>
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<mods:number id="C5422D506B390C7A2E2E19C9D4701498">3</mods:number>
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<mods:url id="8AF697F7F61134B8F4C36DE036CDBC9C">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x</mods:url>
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<mods:identifier id="70DA54601526351626C4261A64664387" type="DOI">10.1111/j.1096-3642.2009.00526.x</mods:identifier>
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<mods:identifier id="C2DF109BB859652A8107B48D9227A9B9" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03F9878AFF96E8190FB98FA5239FFE34" ID-DOI="http://doi.org/10.5281/zenodo.5492201" ID-Zenodo-Dep="5492201" LSID="urn:lsid:plazi:treatment:03F9878AFF96E8190FB98FA5239FFE34" httpUri="http://treatment.plazi.org/id/03F9878AFF96E8190FB98FA5239FFE34" lastPageId="26" lastPageNumber="541" pageId="25" pageNumber="540">
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<subSubSection id="C34A6517FF96E81A0FB98FA52675FB41" pageId="25" pageNumber="540" type="nomenclature">
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<paragraph id="8BEF369CFF96E81A0FB98FA52624FC01" blockId="25.[922,1345,950,971]" box="[922,1345,950,971]" pageId="25" pageNumber="540">
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<heading id="D0A781F0FF96E81A0FB98FA52624FC01" box="[922,1345,950,971]" centered="true" fontSize="9" level="2" pageId="25" pageNumber="540" reason="2">
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<taxonomicName id="4C504D1FFF96E81A0FB98FA527B7FC00" ID-CoL="6HLVS" authority="Liew & Schilthuizen & Vermeulen, 2009" authorityName="Liew & Schilthuizen & Vermeulen" authorityYear="2009" box="[922,1234,950,971]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="540" phylum="Mollusca" rank="species" species="occidentalis" status="sp. nov.">EVERETTIA OCCIDENTALIS</taxonomicName>
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<emphasis id="B924EA8EFF96E81A08FF8FAA2624FC01" bold="true" box="[1244,1345,950,971]" pageId="25" pageNumber="540">
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<taxonomicNameLabel id="A21757F5FF96E81A08FF8FAA2624FC01" box="[1244,1345,950,971]" pageId="25" pageNumber="540" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEF369CFF96E81A0F198FCE2675FB41" blockId="25.[826,1442,988,1163]" pageId="25" pageNumber="540">
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<emphasis id="B924EA8EFF96E81A0F198FCE20E5FC38" box="[826,896,989,1010]" italics="true" pageId="25" pageNumber="540">Types:</emphasis>
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<materialsCitation id="3B383CC1FF96E81A0FAA8FCF2669FB41" collectingDate="2006-06-17" collectionCode="T" collectorName="S. Liew & Meckson. Date" country="Malaysia" county="State" elevation="1200" latitude="4.4" location="Upper Padas" longLatPrecision="1306" longitude="115.73333" municipality="Sipitang District" pageId="25" pageNumber="540" specimenCode="BMNH 20080632" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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<collectingCountry id="F347760CFF96E81A0FAA8FCF2777FC3B" box="[905,1042,988,1009]" name="Malaysia" pageId="25" pageNumber="540">MALAYSIA</collectingCountry>
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:
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<collectingCounty id="628E4E10FF96E81A083C8FCF2739FC3B" box="[1055,1116,988,1009]" pageId="25" pageNumber="540">State</collectingCounty>
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of
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<collectingRegion id="4994F87EFF96E81A08A18FCF27A8FC38" box="[1154,1229,988,1010]" country="Malaysia" name="Sabah" pageId="25" pageNumber="540">Sabah</collectingRegion>
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:
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<collectingMunicipality id="6B8BACE6FF96E81A08FA8FCF26FBFC38" box="[1241,1438,988,1010]" pageId="25" pageNumber="540">Sipitang District</collectingMunicipality>
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||
.
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<location id="8E8F6047FF96E81A0F198FE820B1FBDB" LSID="urn:lsid:plazi:treatment:03F9878AFF96E8190FB98FA5239FFE34:8E8F6047FF96E81A0F198FE820B1FBDB" box="[826,980,1019,1041]" country="Malaysia" county="State" latitude="4.4" longLatPrecision="1306" longitude="115.73333" municipality="Sipitang District" name="Upper Padas" pageId="25" pageNumber="540" stateProvince="Sabah">Upper Padas</location>
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, forest patch near
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<location id="8E8F6047FF96E81A08978FE826F8FBDB" LSID="urn:lsid:plazi:treatment:03F9878AFF96E8190FB98FA5239FFE34:8E8F6047FF96E81A08978FE826F8FBDB" box="[1204,1437,1019,1041]" country="Malaysia" county="State" latitude="4.4" longLatPrecision="1306" longitude="115.73333" municipality="Sipitang District" name="Long Pa Sia village" pageId="25" pageNumber="540" stateProvince="Sabah">Long Pa Sia village</location>
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.
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<location id="8E8F6047FF96E81A0F19880A2030FBE4" LSID="urn:lsid:plazi:treatment:03F9878AFF96E8190FB98FA5239FFE34:8E8F6047FF96E81A0F19880A2030FBE4" box="[826,853,1049,1070]" country="Malaysia" county="State" latitude="4.4" longLatPrecision="1306" longitude="115.73333" municipality="Sipitang District" name="At" pageId="25" pageNumber="540" stateProvince="Sabah">At</location>
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<quantity id="4CA89B79FF96E81A0F43880A20D1FBE5" box="[864,948,1049,1071]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="540" unit="m" value="1200.0">
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<elevation id="007DD1AFFF96E81A0F43880A20D1FBE5" box="[864,948,1049,1071]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="540" unit="m" value="1200.0">1200 m</elevation>
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</quantity>
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alt. (
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<geoCoordinate id="EE64505BFF96E81A0FD4880A2723FBE5" box="[1015,1094,1048,1071]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="25" pageNumber="540" precision="925" value="4.4">
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4°24
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<emphasis id="B924EA8EFF96E81A0809880B2755FBE5" box="[1066,1072,1048,1071]" italics="true" pageId="25" pageNumber="540">′</emphasis>
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N
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</geoCoordinate>
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,
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<geoCoordinate id="EE64505BFF96E81A0875880A27D8FBE5" box="[1110,1213,1048,1071]" degrees="115" direction="east" minutes="44" orientation="longitude" pageId="25" pageNumber="540" precision="925" value="115.73333">
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115°44
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<emphasis id="B924EA8EFF96E81A0880880B27CCFBE5" box="[1187,1193,1048,1071]" italics="true" pageId="25" pageNumber="540">′</emphasis>
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E
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</geoCoordinate>
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). Collected by T.-
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<collectorName id="26A5534AFF96E81A09AD880A2017FB84" pageId="25" pageNumber="540">S. Liew</collectorName>
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and
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<collectorName id="26A5534AFF96E81A0F93882B273AFB87" box="[944,1119,1080,1102]" pageId="25" pageNumber="540">Meckson. Date</collectorName>
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:
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<date id="FFEE105CFF96E81A084F882B2786FB84" box="[1132,1251,1080,1102]" pageId="25" pageNumber="540" value="2006-06-17">
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<collectingDate id="EFAAE9B4FF96E81A084F882B2786FB84" box="[1132,1251,1080,1102]" pageId="25" pageNumber="540" value="2006-06-17">17.vi.2006</collectingDate>
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</date>
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,
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<typeStatus id="54EB883EFF96E81A08D1882B2630FB84" box="[1266,1365,1080,1102]" pageId="25" pageNumber="540" type="holotype">holotype</typeStatus>
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, BOR/ MOL 4345,
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<figureCitation id="136B2A19FF96E81A0FFF88452734FBA6" box="[988,1105,1110,1132]" captionStart="Figure 2" captionStartId="7.[164,243,1411,1430]" captionTargetBox="[326,1281,196,1382]" captionTargetId="figure-180@7.[323,1283,194,1382]" captionTargetPageId="7" captionText="Figure 2. Shells. A, Everettia corrugata corrugata; B, Everettia corrugata williamsi ssp. nov.; C, Everettia interior sp. nov.; D, Everettia jucunda; E, Everettia occidentalis sp. nov.; F, Everettia jucundior sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443027" httpUri="https://zenodo.org/record/5443027/files/figure.png" pageId="25" pageNumber="540">Figure 2E</figureCitation>
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;
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<specimenCount id="9D56FD15FF96E81A084988452648FBA6" box="[1130,1325,1110,1132]" pageId="25" pageNumber="540" type="generic" typeStatus="paratypes">eight paratypes</specimenCount>
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, four in
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<specimenCode id="DBF69EE7FF96E81A0F1988652766FB41" box="[826,1027,1141,1163]" collectionCode="BMNH" pageId="25" pageNumber="540">BMNH 20080632</specimenCode>
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and four in SP 13064
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6517FF96E8190F1988A7239FFE34" lastPageId="26" lastPageNumber="541" pageId="25" pageNumber="540" type="etymology">
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<paragraph id="8BEF369CFF96E81A0F1988A720ECFACD" blockId="25.[826,1441,1204,1287]" pageId="25" pageNumber="540">
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<emphasis id="B924EA8EFF96E81A0F1988A720DDFB03" box="[826,952,1204,1225]" italics="true" pageId="25" pageNumber="540">Etymology:</emphasis>
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This species is named
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<taxonomicName id="4C504D1FFF96E81A08EF88A72634FB03" authorityName="Liew & Schilthuizen & Vermeulen" authorityYear="2009" box="[1228,1361,1204,1225]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="540" phylum="Mollusca" rank="species" species="occidentalis">
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<emphasis id="B924EA8EFF96E81A08EF88A72634FB03" box="[1228,1361,1204,1225]" italics="true" pageId="25" pageNumber="540">occidentalis</emphasis>
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</taxonomicName>
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for its distribution range in the western mountain ranges of
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<collectingRegion id="4994F87EFF96E81A0F1988E220E0FACD" box="[826,901,1265,1287]" country="Malaysia" name="Sabah" pageId="25" pageNumber="540">Sabah</collectingRegion>
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.
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</paragraph>
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<paragraph id="8BEF369CFF96E81A0F19893C2751FA17" blockId="25.[826,1441,1327,1502]" pageId="25" pageNumber="540">
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<emphasis id="B924EA8EFF96E81A0F19893C277EFA8E" box="[826,1051,1327,1348]" italics="true" pageId="25" pageNumber="540">Material examined:</emphasis>
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<collectingRegion id="4994F87EFF96E81A0804893C27E5FA8E" box="[1063,1152,1327,1348]" country="Malaysia" name="Sabah" pageId="25" pageNumber="540">SABAH</collectingRegion>
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– Crocker Range: BOR/ MOL 946, Lian cave,
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<date id="FFEE105CFF96E81A0868895D27FBFAAE" box="[1099,1182,1358,1380]" pageId="25" pageNumber="540" value="2000-04">iv.2000</date>
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. JJ 1087, Lian cave,
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<date id="FFEE105CFF96E81A0F19897F20FDFA48" box="[826,920,1388,1410]" pageId="25" pageNumber="540" value="1986-07">vii.1986</date>
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. Tenom: BOR/MOL 1373, Tenom Rafflesia Garden,
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<quantity id="4CA89B79FF96E81A0F8189982082FA6B" box="[930,999,1419,1441]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="540" unit="m" value="400.0">400 m</quantity>
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||
alt.,
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<date id="FFEE105CFF96E81A08088998271AFA6B" box="[1067,1151,1419,1441]" pageId="25" pageNumber="540" value="2003-06">vi.2003</date>
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||
; BOR/MOL
|
||
<quantity id="4CA89B79FF96E81A0931899826C4FA6B" box="[1298,1441,1419,1441]" metricMagnitude="6" metricUnit="m" metricValue="4.24435" pageId="25" pageNumber="540" unit="m" value="4244350.0">4244, 350 m</quantity>
|
||
alt.,
|
||
<date id="FFEE105CFF96E81A0F5389B920DCFA0A" box="[880,953,1450,1472]" pageId="25" pageNumber="540" value="2006-01">i.2006</date>
|
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. Sipitang: BOR/MOL 2993, Muaya waterfall, Lumaku,
|
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<date id="FFEE105CFF96E81A0FC189DB2755FA17" box="[994,1072,1480,1501]" pageId="25" pageNumber="540" value="2003-10">x.2003</date>
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||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF96E81A0F198A1427A1F993" blockId="25.[826,1441,1543,1626]" pageId="25" pageNumber="540">
|
||
<emphasis id="B924EA8EFF96E81A0F198A14270AF9D6" box="[826,1135,1543,1564]" italics="true" pageId="25" pageNumber="540">Diagnostic characteristics:</emphasis>
|
||
Regularly spaced radial grooves above shell which are different from the radial corrugation of
|
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<emphasis id="B924EA8EFF96E81A08178A5627A1F993" box="[1076,1220,1604,1626]" italics="true" pageId="25" pageNumber="540">
|
||
<taxonomicName id="4C504D1FFF96E81A08178A5627A5F993" authorityName="Liew & Schilthuizen & Vermeulen" authorityYear="2009" box="[1076,1216,1604,1626]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="540" phylum="Mollusca" rank="species" species="dominiki">E. dominiki</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF96E8190F198A9021A5FEBF" blockId="25.[826,1441,1667,1903]" lastBlockId="26.[144,759,198,373]" lastPageId="26" lastPageNumber="541" pageId="25" pageNumber="540">
|
||
<emphasis id="B924EA8EFF96E81A0F198A9020A7F952" box="[826,962,1667,1688]" italics="true" pageId="25" pageNumber="540">Description:</emphasis>
|
||
Shell (
|
||
<figureCitation id="136B2A19FF96E81A08108A9027E9F952" box="[1075,1164,1667,1689]" captionStart="Figure 2" captionStartId="7.[164,243,1411,1430]" captionTargetBox="[326,1281,196,1382]" captionTargetId="figure-180@7.[323,1283,194,1382]" captionTargetPageId="7" captionText="Figure 2. Shells. A, Everettia corrugata corrugata; B, Everettia corrugata williamsi ssp. nov.; C, Everettia interior sp. nov.; D, Everettia jucunda; E, Everettia occidentalis sp. nov.; F, Everettia jucundior sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443027" httpUri="https://zenodo.org/record/5443027/files/figure.png" pageId="25" pageNumber="540">Figs 2E</figureCitation>
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,
|
||
<figureCitation id="136B2A19FF96E81A088B8A9027BDF953" box="[1192,1240,1667,1689]" captionStart="Figure 11" captionStartId="18.[144,223,1209,1228]" captionTargetBox="[305,1263,196,1180]" captionTargetId="figure-241@18.[303,1263,196,1180]" captionTargetPageId="18" captionText="Figure 11. Shell surface sculptures (scanning electron micrographs). A, upper surface of Everettia corrugata williamsi ssp. nov.; B, underside of E. corrugata williamsi ssp. nov.; C, underside of Everettia lapidini; D, upper surface of Everettia occidentalis sp. nov.; E, corrugation at the suture of Everettia corrugata corrugata; F, upper surface of Everettia jucundior sp. nov. Scale bar = 500 Mm." figureDoi="http://doi.org/10.5281/zenodo.5443045" httpUri="https://zenodo.org/record/5443045/files/figure.png" pageId="25" pageNumber="540">11D</figureCitation>
|
||
): medium-sized, rather thin, brownish, moderately elevated, outer whorls slightly shouldered below the suture. Periphery round to slightly angular. Above the periphery, shell shiny with rather regularly placed radial grooves,
|
||
<emphasis id="B924EA8EFF96E81A0F818B0F20D1F8FB" box="[930,948,1820,1841]" italics="true" pageId="25" pageNumber="540">c.</emphasis>
|
||
2–4 per mm. These radial grooves make the shell seem sculptured with radial ridges under the naked eye. Below the periphery, shell has very fine, shallow, densely placed spiral grooves. Height up to 11.0 mm; width up to 13.0 mm; diameter of the first three whorls 0.9, 0.5, and
|
||
<quantity id="4CA89B79FF95E8190DDB8D10212AFED3" box="[504,591,259,281]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="541" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, respectively; number of whorls up to four and three-quarters; height aperture up to
|
||
<quantity id="4CA89B79FF95E8190DBA8D532295FE9C" box="[409,496,320,342]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="26" pageNumber="541" unit="mm" value="4.9">4.9 mm</quantity>
|
||
; width aperture up to
|
||
<quantity id="4CA89B79FF95E8190CB38D4C238DFEBE" box="[144,232,351,373]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="26" pageNumber="541" unit="mm" value="6.8">6.8 mm</quantity>
|
||
. Genitalia: unknown. Animal: unknown.
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF95E8190CB38DB9239FFE34" blockId="26.[144,759,426,510]" pageId="26" pageNumber="541">
|
||
<emphasis id="B924EA8EFF95E8190CB38DB922C8FE75" box="[144,429,426,447]" italics="true" pageId="26" pageNumber="541">Distribution and habitat:</emphasis>
|
||
Primary forest.
|
||
<collectingRegion id="4994F87EFF95E8190E4B8DB921D6FE0A" box="[616,691,426,448]" country="Malaysia" name="Sabah" pageId="26" pageNumber="541">Sabah</collectingRegion>
|
||
: Long Pa Sia, Tenom, and Sipitang, interior of
|
||
<collectingRegion id="4994F87EFF95E8190E8C8DDA2192FE15" box="[687,759,457,479]" country="Malaysia" name="Sabah" pageId="26" pageNumber="541">Sabah</collectingRegion>
|
||
(
|
||
<figureCitation id="136B2A19FF95E8190CBB8DFB238BFE34" box="[152,238,488,510]" captionStart="Figure 5" captionStartId="10.[144,223,1342,1361]" captionTargetBox="[305,1261,197,1311]" captionTargetId="figure-168@10.[303,1263,195,1313]" captionTargetPageId="10" captionText="Figure 5. Distribution maps (Sabah). A, Everettia subconsul in the east and west coast lowland, Everettia jucunda in the west coast, and Everettia occidentalis sp. nov. in the western mountain range; B, Everettia klemmantanica in the central mountain ranges, Everettia interior sp. nov. in the interior area, Everettia paulbasintali sp. nov. in the east coast lowland, Everettia themis in the central mountain ranges; C, Everettia lapidini sp. nov. in the central mountain ranges, Everettia jucundior sp. nov. in the east coast lowland; D, Everettia monticola sp. nov. in the central mountain ranges, Everettia planispira sp. nov. in the eastern and northern parts of Sabah." figureDoi="http://doi.org/10.5281/zenodo.5443033" httpUri="https://zenodo.org/record/5443033/files/figure.png" pageId="26" pageNumber="541">Fig. 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |