Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo
Author
Liew, Thor-Seng
Author
Schilthuizen, Menno
Author
Vermeulen, Jaap Jan
text
Zoological Journal of the Linnean Society
2009
2009-10-26
157
3
515
550
https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x
journal article
10.1111/j.1096-3642.2009.00526.x
0024-4082
5443023
EVERETTIA OCCIDENTALIS
SP. NOV.
Types:
MALAYSIA
:
State
of
Sabah
:
Sipitang District
.
Upper Padas
, forest patch near
Long Pa Sia village
.
At
1200 m
alt. (
4°24
′
N
,
115°44
′
E
). Collected by T.-
S. Liew
and
Meckson. Date
:
17.vi.2006
,
holotype
, BOR/ MOL 4345,
Figure 2E
;
eight paratypes
, four in
BMNH 20080632
and four in SP 13064
.
Etymology:
This species is named
occidentalis
for its distribution range in the western mountain ranges of
Sabah
.
Material examined:
SABAH
– Crocker Range: BOR/ MOL 946, Lian cave,
iv.2000
. JJ 1087, Lian cave,
vii.1986
. Tenom: BOR/MOL 1373, Tenom Rafflesia Garden,
400 m
alt.,
vi.2003
; BOR/MOL
4244, 350 m
alt.,
i.2006
. Sipitang: BOR/MOL 2993, Muaya waterfall, Lumaku,
x.2003
.
Diagnostic characteristics:
Regularly spaced radial grooves above shell which are different from the radial corrugation of
E. dominiki
.
Description:
Shell (
Figs 2E
,
11D
): medium-sized, rather thin, brownish, moderately elevated, outer whorls slightly shouldered below the suture. Periphery round to slightly angular. Above the periphery, shell shiny with rather regularly placed radial grooves,
c.
2–4 per mm. These radial grooves make the shell seem sculptured with radial ridges under the naked eye. Below the periphery, shell has very fine, shallow, densely placed spiral grooves. Height up to 11.0 mm; width up to 13.0 mm; diameter of the first three whorls 0.9, 0.5, and
1.2 mm
, respectively; number of whorls up to four and three-quarters; height aperture up to
4.9 mm
; width aperture up to
6.8 mm
. Genitalia: unknown. Animal: unknown.
Distribution and habitat:
Primary forest.
Sabah
: Long Pa Sia, Tenom, and Sipitang, interior of
Sabah
(
Fig. 5A
).