212 lines
22 KiB
XML
212 lines
22 KiB
XML
<document id="2EF60E8C604A25A270F9BF7405061074" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. & Marusik, Yuri M." docDate="2021" docId="03F98785035EEF421C43794FFC066F2A" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sosticus montanus Zamani & Chatzaki & Esyunin & Marusik 2021, sp. nov." docType="treatment" docUuid="9C05213D-7ACA-4312-A314-26D6EA50A334" docUuidSource="ZooBank" docVersion="8" lastPageNumber="103" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="101" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3937322105B596CCB69D17F6A96FF3D0">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="42E77F644639898FF0C99ACCB8F68337">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="DF6C4483EA0C4C21C9D1D3A49726CFA6">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:nameIdentifier id="4ADF662DB233C9CED4AC2D72519FBF5A" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FC536923CFC7D85F18C9A95951D79A7F">Chatzaki, Maria</mods:namePart>
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<mods:nameIdentifier id="420F9ADB007D0E8D1B284A75AA898AC8" type="email">maria.chatzaki@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FBF58B805BC66E53CA3386AA72158C3E">Esyunin, Sergei L.</mods:namePart>
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<mods:affiliation id="FC37AF16FA591B505219C73BF784E1F8">Sergei.</mods:affiliation>
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<mods:nameIdentifier id="D15F04E9F18FB412D2003F02C82FEBD3" type="email">Sergei.Esyunin@psu.ru</mods:nameIdentifier>
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<mods:namePart id="8D35901F6AEB0CAD647797FDC57AB8B9">Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier id="A6BD8B0DC2BD3FAC00D692E64AE3EB79" type="email">yurmar@mail.ru</mods:nameIdentifier>
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<mods:title id="5871EE86B8D898DFFAA6090A1685FB23">European Journal of Taxonomy</mods:title>
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<mods:part id="F759AC539890AB4CA03C1AB58F94685C">
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<mods:date id="FABF9245CECC9579EFE6008FE139D38C">2021</mods:date>
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<treatment id="03F98785035EEF421C43794FFC066F2A" ID-DOI="http://doi.org/10.5281/zenodo.4906496" ID-GBIF-Taxon="182597855" ID-Zenodo-Dep="4906496" LSID="urn:lsid:zoobank.org:act:9C05213D-7ACA-4312-A314-26D6EA50A334" httpUri="http://treatment.plazi.org/id/03F98785035EEF421C43794FFC066F2A" lastPageId="35" lastPageNumber="103" pageId="33" pageNumber="101">
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<subSubSection id="C34A6518035EEF401C43794FFC7C6B17" box="[631,957,1596,1622]" pageId="33" pageNumber="101" type="nomenclature">
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<paragraph id="8BEF3693035EEF401C43794FFC7C6B17" blockId="33.[363,1224,1596,1693]" box="[631,957,1596,1622]" pageId="33" pageNumber="101">
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<heading id="D0A781FF035EEF401C43794FFC7C6B17" box="[631,957,1596,1622]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="101" reason="2">
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<taxonomicName id="4C504D10035EEF401C43794FFC9A6B17" ID-CoL="98YVQ" authority="Zamani & Chatzaki & Esyunin & Marusik, 2021" authorityName="Zamani & Chatzaki & Esyunin & Marusik" authorityYear="2021" box="[631,859,1596,1622]" class="Arachnida" family="Gnaphosidae" genus="Sosticus" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="101" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
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<emphasis id="B924EA81035EEF401C43794FFC9A6B17" bold="true" box="[631,859,1596,1622]" italics="true" pageId="33" pageNumber="101">Sosticus montanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21757FA035EEF401D56794FFC7C6B17" box="[866,957,1596,1622]" pageId="33" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C34A6518035EEF401F5F792CFC826BDC" pageId="33" pageNumber="101" type="description">
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<uri id="FFC13A91035EEF401F5F792CFB096B3B" box="[363,1224,1631,1658]" pageId="33" pageNumber="101">
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urn:lsid:zoobank.org:act:
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</uri>
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</paragraph>
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<paragraph id="8BEF3693035EEF401CC479F0FC826BDC" blockId="33.[363,1224,1596,1693]" box="[752,835,1667,1693]" pageId="33" pageNumber="101">
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<figureCitation id="136B2A16035EEF401CC479F0FC826BDC" box="[752,835,1667,1693]" captionStart="Fig" captionStartId="34.[189,232,1928,1954]" captionTargetBox="[189,1398,779,1886]" captionTargetId="figure-101@34.[189,1398,779,1890]" captionTargetPageId="34" captionText="Fig. 21. Sosticus montanus sp. nov., holotype, ♀ (MHNG). A, E, G. Epigyne, ventral view. B, D, F, H. Epigyne, dorsal view. C. Habitus, dorsal view. I. Chelicera, posterior view. Scale bars: C = 0.5 mm; G–H = 0.1 mm; I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906065" httpUri="https://zenodo.org/record/4906065/files/figure.png" pageId="33" pageNumber="101">Fig. 21</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6518035EEF401E8979B7FD176A14" pageId="33" pageNumber="101" type="diagnosis">
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<paragraph id="8BEF3693035EEF401E8979B7FEF76B9F" blockId="33.[189,310,1732,1758]" box="[189,310,1732,1758]" pageId="33" pageNumber="101">
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<heading id="D0A781FF035EEF401E8979B7FEF76B9F" bold="true" box="[189,310,1732,1758]" fontSize="11" level="3" pageId="33" pageNumber="101" reason="3">
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<emphasis id="B924EA81035EEF401E8979B7FEF76B9F" bold="true" box="[189,310,1732,1758]" pageId="33" pageNumber="101">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEF3693035EEF401E897980FD176A14" blockId="33.[189,1399,1779,1877]" pageId="33" pageNumber="101">
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Epigyne of the new species is most similar to that of
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<taxonomicName id="4C504D10035EEF401D1B7987FC746A4C" baseAuthorityName="L. Koch" baseAuthorityYear="1866" box="[815,949,1779,1806]" class="Arachnida" family="Gnaphosidae" genus="Sosticus" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="101" phylum="Arthropoda" rank="species" species="loricatus">
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<emphasis id="B924EA81035EEF401D1B7987FC746A4C" box="[815,949,1779,1806]" italics="true" pageId="33" pageNumber="101">S. loricatus</emphasis>
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</taxonomicName>
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but differs by the shape of atrium and the copulatory ducts: atrium triangular, wider posteriorly (vs wider anteriorly) and the ducts converge medially and diverge anteriorly (vs opposite).
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34A6518035EEF401E89780FFE486AA8" pageId="33" pageNumber="101" type="etymology">
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<paragraph id="8BEF3693035EEF401E89780FFE856AD7" blockId="33.[189,324,1916,1942]" box="[189,324,1916,1942]" pageId="33" pageNumber="101">
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<heading id="D0A781FF035EEF401E89780FFE856AD7" bold="true" box="[189,324,1916,1942]" fontSize="11" level="3" pageId="33" pageNumber="101" reason="3">
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<emphasis id="B924EA81035EEF401E89780FFE856AD7" bold="true" box="[189,324,1916,1942]" pageId="33" pageNumber="101">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEF3693035EEF401E8978D8FE486AA8" blockId="33.[189,1398,1963,2025]" pageId="33" pageNumber="101">The specific epithet is an adjective and refers to the mountainous habitat of the new species in the Zagros Mountain Range.</paragraph>
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</subSubSection>
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<subSubSection id="C34A6518035DEF431E897E7AFB3A6C35" pageId="34" pageNumber="102" type="materials_examined">
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<paragraph id="8BEF3693035DEF431E897E7AFEAD6C63" blockId="34.[189,364,264,291]" box="[189,364,264,291]" pageId="34" pageNumber="102">
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<heading id="D0A781FF035DEF431E897E7AFEAD6C63" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="34" pageNumber="102" reason="3">
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<emphasis id="B924EA81035DEF431E897E7AFEAD6C63" bold="true" box="[189,364,264,291]" pageId="34" pageNumber="102">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEF3693035DEF431EDE7E44FB3A6C35" blockId="34.[189,1275,311,373]" pageId="34" pageNumber="102">
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<materialsCitation id="3B383CCE035DEF431EDE7E44FB3A6C35" ID-GBIF-Occurrence="3127434326" collectingDate="2015-07" collectionCode="MHNG" collectorName="A. Zamani" country="Iran" latitude="32.883335" location="Abdanan" longLatPrecision="1208" longitude="47.5" pageId="34" pageNumber="102" specimenCount="1" specimenCount-female="1" stateProvince="Ilam Province" typeStatus="holotype">
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<emphasis id="B924EA81035DEF431EDE7E44FE9D6C10" bold="true" box="[234,348,311,337]" pageId="34" pageNumber="102">
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<typeStatus id="54EB8831035DEF431EDE7E44FE9D6C10" box="[234,348,311,337]" pageId="34" pageNumber="102" type="holotype">Holotype</typeStatus>
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</emphasis>
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<collectingCountry id="F3477603035DEF431E897E29FEC96C35" box="[189,264,346,372]" name="Iran" pageId="34" pageNumber="102">IRAN</collectingCountry>
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•
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<specimenCount id="9D56FD1A035DEF431F167E28FEF86C34" box="[290,313,347,373]" pageId="34" pageNumber="102" type="female">♀</specimenCount>
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;
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<collectingRegion id="4994F871035DEF431F737E29FE2E6C35" box="[327,495,346,372]" country="Iran" name="Ilam" pageId="34" pageNumber="102">Ilam Province</collectingRegion>
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,
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<location id="8E8F6048035DEF431FCE7E29FDA96C35" LSID="urn:lsid:plazi:treatment:03F98785035EEF421C43794FFC066F2A:8E8F6048035DEF431FCE7E29FDA96C35" box="[506,616,346,372]" country="Iran" latitude="32.883335" longLatPrecision="1208" longitude="47.5" name="Abdanan" pageId="34" pageNumber="102" stateProvince="Ilam Province">Abdanan</location>
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;
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<geoCoordinate id="EE645054035DEF431C477E2AFD1E6C34" box="[627,735,345,373]" degrees="32" direction="north" minutes="53" orientation="latitude" pageId="34" pageNumber="102" precision="925" value="32.883335">32°53′ N</geoCoordinate>
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,
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<geoCoordinate id="EE645054035DEF431CDF7E2AFC956C34" box="[747,852,345,373]" degrees="47" direction="east" minutes="30" orientation="longitude" pageId="34" pageNumber="102" precision="925" value="47.5">47°30′ E</geoCoordinate>
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;
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<date id="FFEE1053035DEF431D557E29FC116C34" box="[865,976,346,373]" pageId="34" pageNumber="102" value="2015-07">
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<collectingDate id="EFAAE9BB035DEF431D557E29FC116C34" box="[865,976,346,373]" pageId="34" pageNumber="102" value="2015-07">Jul. 2015</collectingDate>
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</date>
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||
;
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||
<collectorName id="26A55345035DEF431DEE7E29FB996C35" box="[986,1112,346,372]" pageId="34" pageNumber="102">A. Zamani</collectorName>
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leg.;
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||
<collectionCode id="ED41AE56035DEF431AAE7E29FB396C35" box="[1178,1272,346,372]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="34" pageNumber="102" type="Museum">MHNG</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6518035DEF421E897EE8FEF86CB9" lastPageId="35" lastPageNumber="103" pageId="34" pageNumber="102" type="description">
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<paragraph id="8BEF3693035DEF431E897EE8FE8F6CF4" blockId="34.[189,334,411,437]" box="[189,334,411,437]" pageId="34" pageNumber="102">
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<heading id="D0A781FF035DEF431E897EE8FE8F6CF4" bold="true" box="[189,334,411,437]" fontSize="11" level="3" pageId="34" pageNumber="102" reason="3">
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<emphasis id="B924EA81035DEF431E897EE8FE8F6CF4" bold="true" box="[189,334,411,437]" pageId="34" pageNumber="102">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEF3693035DEF431EDE7EB9FE846CA5" blockId="34.[189,1401,458,660]" box="[234,325,458,484]" pageId="34" pageNumber="102">
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<emphasis id="B924EA81035DEF431EDE7EB9FE846CA5" bold="true" box="[234,325,458,484]" pageId="34" pageNumber="102">Female</emphasis>
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</paragraph>
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<paragraph id="8BEF3693035DEF431E897E9EFE2A6FD5" blockId="34.[189,1401,458,660]" pageId="34" pageNumber="102">
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Habitus as in
|
||
<figureCitation id="136B2A16035DEF431F697E9EFE066F49" box="[349,455,493,520]" captionStart="Fig" captionStartId="34.[189,232,1928,1954]" captionTargetBox="[189,1398,779,1886]" captionTargetId="figure-101@34.[189,1398,779,1890]" captionTargetPageId="34" captionText="Fig. 21. Sosticus montanus sp. nov., holotype, ♀ (MHNG). A, E, G. Epigyne, ventral view. B, D, F, H. Epigyne, dorsal view. C. Habitus, dorsal view. I. Chelicera, posterior view. Scale bars: C = 0.5 mm; G–H = 0.1 mm; I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906065" httpUri="https://zenodo.org/record/4906065/files/figure.png" pageId="34" pageNumber="102">Fig. 21C</figureCitation>
|
||
. Total length 2.68. Carapace 1.67 long, 1.32 wide. Eye sizes and interdistance of PMEs: AME = 0.06, ALE = 0.08, PME = 0.11, PLE = 0.07, PME–PME = 0.04. Carapace, chelicerae, sternum, maxillae and labium yellow. Chelicera (
|
||
<figureCitation id="136B2A16035DEF431D277D47FCB56F0F" box="[787,884,564,590]" captionStart="Fig" captionStartId="34.[189,232,1928,1954]" captionTargetBox="[189,1398,779,1886]" captionTargetId="figure-101@34.[189,1398,779,1890]" captionTargetPageId="34" captionText="Fig. 21. Sosticus montanus sp. nov., holotype, ♀ (MHNG). A, E, G. Epigyne, ventral view. B, D, F, H. Epigyne, dorsal view. C. Habitus, dorsal view. I. Chelicera, posterior view. Scale bars: C = 0.5 mm; G–H = 0.1 mm; I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906065" httpUri="https://zenodo.org/record/4906065/files/figure.png" pageId="34" pageNumber="102">Fig. 21I</figureCitation>
|
||
) with 3 promarginal teeth. Abdomen pale, yellowish, with four dorsal sigilla, ventrally with distinct tracheal marks. Spinnerets pale and uniform in color. All legs missing.
|
||
</paragraph>
|
||
<caption id="DF2F661B035DEF431E8978FBFDC86AA8" ID-DOI="http://doi.org/10.5281/zenodo.4906065" ID-Zenodo-Dep="4906065" httpUri="https://zenodo.org/record/4906065/files/figure.png" pageId="34" pageNumber="102" startId="34.[189,232,1928,1954]" targetBox="[189,1398,779,1886]" targetPageId="34">
|
||
<paragraph id="8BEF3693035DEF431E8978FBFDC86AA8" blockId="34.[189,1400,1928,2025]" pageId="34" pageNumber="102">
|
||
<emphasis id="B924EA81035DEF431E8978FBFEDD6AE3" bold="true" box="[189,284,1928,1954]" pageId="34" pageNumber="102">Fig. 21.</emphasis>
|
||
<taxonomicName id="4C504D10035DEF431F1278FBFDC26AE3" authority="Zamani & Chatzaki & Esyunin & Marusik, 2021" authorityName="Zamani & Chatzaki & Esyunin & Marusik" authorityYear="2021" box="[294,515,1928,1954]" class="Arachnida" family="Gnaphosidae" genus="Sosticus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="102" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
|
||
<emphasis id="B924EA81035DEF431F1278FBFDC26AE3" box="[294,515,1928,1954]" italics="true" pageId="34" pageNumber="102">Sosticus montanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21757FA035DEF431C3978FAFDAA6AE2" box="[525,619,1929,1955]" pageId="34" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♀ (MHNG).
|
||
<emphasis id="B924EA81035DEF431DBA78FBFC2E6AE3" bold="true" box="[910,1007,1928,1954]" pageId="34" pageNumber="102">A, E, G</emphasis>
|
||
. Epigyne, ventral view.
|
||
<emphasis id="B924EA81035DEF431B2378FBFF156A84" bold="true" pageId="34" pageNumber="102">B, D, F, H</emphasis>
|
||
. Epigyne, dorsal view.
|
||
<emphasis id="B924EA81035DEF431FDC78D8FE3C6A84" bold="true" box="[488,509,1963,1989]" pageId="34" pageNumber="102">C</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="B924EA81035DEF431D3F78D8FCD76A84" bold="true" box="[779,790,1963,1989]" pageId="34" pageNumber="102">I</emphasis>
|
||
. Chelicera, posterior view. Scale bars: C = 0.5 mm; G–H = 0.1 mm; I = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEF3693035CEF421E897E7AFDDA6CD0" blockId="35.[189,1399,265,401]" pageId="35" pageNumber="103">
|
||
Epigyne as in
|
||
<figureCitation id="136B2A16035CEF421F517E7AFDFD6C62" box="[357,572,265,291]" captionStart="Fig" captionStartId="34.[189,232,1928,1954]" captionTargetBox="[189,1398,779,1886]" captionTargetId="figure-101@34.[189,1398,779,1890]" captionTargetPageId="34" captionText="Fig. 21. Sosticus montanus sp. nov., holotype, ♀ (MHNG). A, E, G. Epigyne, ventral view. B, D, F, H. Epigyne, dorsal view. C. Habitus, dorsal view. I. Chelicera, posterior view. Scale bars: C = 0.5 mm; G–H = 0.1 mm; I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906065" httpUri="https://zenodo.org/record/4906065/files/figure.png" pageId="35" pageNumber="103">Fig. 21A–B, D–H</figureCitation>
|
||
. Atrium triangular; scape large, longer than atrium; receptacles round, touching each other; copulatory ducts semicircular, converging anteriorly; receptacles and ducts form 2 vertical arcs in opposite directions; accessorial duct long, directed anteriorly and terminating at the level of mid part of the scape.
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035CEF421EDE7ECAFEEB6C92" blockId="35.[189,313,441,504]" box="[234,298,441,467]" pageId="35" pageNumber="103">
|
||
<emphasis id="B924EA81035CEF421EDE7ECAFEEB6C92" bold="true" box="[234,298,441,467]" pageId="35" pageNumber="103">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035CEF421E897EADFEF86CB9" blockId="35.[189,313,441,504]" box="[189,313,478,504]" pageId="35" pageNumber="103">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A6518035CEF421E897D53FC066F2A" box="[189,967,544,619]" pageId="35" pageNumber="103" type="distribution">
|
||
<paragraph id="8BEF3693035CEF421E897D53FE976F7B" blockId="35.[189,342,544,570]" box="[189,342,544,570]" pageId="35" pageNumber="103">
|
||
<heading id="D0A781FF035CEF421E897D53FE976F7B" bold="true" box="[189,342,544,570]" fontSize="11" level="3" pageId="35" pageNumber="103" reason="3">
|
||
<emphasis id="B924EA81035CEF421E897D53FE976F7B" bold="true" box="[189,342,544,570]" pageId="35" pageNumber="103">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEF3693035CEF421E897D22FC066F2A" blockId="35.[189,967,592,619]" box="[189,967,592,619]" pageId="35" pageNumber="103">
|
||
Known only from the
|
||
<typeStatus id="54EB8831035CEF421FF67D22FE346F2A" box="[450,501,593,619]" pageId="35" pageNumber="103">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="4994F871035CEF421C4D7D23FCE06F2B" box="[633,801,592,618]" country="Iran" name="Ilam" pageId="35" pageNumber="103">Ilam Province</collectingRegion>
|
||
, western
|
||
<collectingCountry id="F3477603035CEF421DA47D22FC026F2A" box="[912,963,593,619]" name="Iran" pageId="35" pageNumber="103">Iran</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |