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<document id="8424C57FC8FA0E75D32623405DB8B8BB" ID-DOI="10.5281/zenodo.5303719" ID-ISSN="0374-1036" ID-Zenodo-Dep="5303719" ID-ZooBank="69EA2A80-42C1-4041-8255-E9F1189D4BE1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630104705807" checkinUser="marcus" docAuthor="Barclay, Maxwell V. L. &amp; Sw, London" docDate="2015" docId="C878D373FF8E307BFE7A58101028FD73" docLanguage="en" docName="ActaEntMusNatPra.2015.55.2.691-701.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Ivierhipidius Barclay &amp; Sw 2015, gen. nov." docType="treatment" docVersion="4" lastPageNumber="695" masterDocId="3441AB0BFF8F307FFFA75B7E1438FE21" masterDocTitle="Ivierhipidius, an enigmatic new Neotropical genus of Ripiphoridae (Coleoptera: Tenebrionoidea) with four new species" masterLastPageNumber="701" masterPageNumber="691" pageNumber="692" updateTime="1699153561371" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F13FC638656CDA4E7298878568743F16">Ivierhipidius, an enigmatic new Neotropical genus of Ripiphoridae (Coleoptera: Tenebrionoidea) with four new species</mods:title>
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<mods:date id="7A817D64A9B15E047697F3CBFBD9710B">2015</mods:date>
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<paragraph id="406E6265FF8E307EFE7A581016EDFDA7" blockId="1.[477,725,878,902]" box="[477,725,878,902]" pageId="1" pageNumber="692">
<heading id="1B26D509FF8E307EFE7A581016EDFDA7" bold="true" box="[477,725,878,902]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="692" reason="3">
<emphasis id="72A5BE77FF8E307EFE7A581016EDFDA7" bold="true" box="[477,725,878,902]" pageId="1" pageNumber="692">
<taxonomicName id="87D119E6FF8E307EFE7A58101652FDA7" authority="Barclay &amp; Sw, 2015" authorityName="Barclay &amp; Sw" authorityYear="2015" box="[477,618,878,902]" class="Insecta" family="Ripiphoridae" genus="Ivierhipidius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="692" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="72A5BE77FF8E307EFE7A58101652FDA7" bold="true" box="[477,618,878,902]" italics="true" pageId="1" pageNumber="692">Ivierhipidius</emphasis>
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<taxonomicNameLabel id="6996030CFF8E307EFDD6581116EDFDA7" box="[625,725,879,902]" pageId="1" pageNumber="692" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="08CB31EEFF8E307EFD8058F3164FFDEE" pageId="1" pageNumber="692" type="description">
<paragraph id="406E6265FF8E307EFD8058F316B3FD81" blockId="1.[551,651,909,928]" box="[551,651,909,928]" pageId="1" pageNumber="692">
(
<figureCitation id="D8EA7EE0FF8E307EFD8A58F316BCFD81" box="[557,644,909,928]" captionStart-0="Figs 14" captionStart-1="Figures 59" captionStart-2="Figs 1015. 1011" captionStartId-0="2.[151,188,1496,1515]" captionStartId-1="3.[108,172,1340,1359]" captionStartId-2="6.[151,188,1552,1571]" captionTargetBox-0="[267,1018,229,1471]" captionTargetBox-1="[163,1423,402,1314]" captionTargetBox-2="[193,1080,219,1521]" captionTargetId-0="figure-10@2.[239,1040,206,1486]" captionTargetId-1="figure-10@3.[148,1435,391,1336]" captionTargetId-2="figure-10@6.[174,1104,219,1534]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionText-0="Figs 14. Habitus. 12 Ivierhipidius paradoxus sp. nov.: 1 holotype, dorsal view; 2 paratype, lateral view. 34 I. youngi sp. nov.: 3 holotype, dorsal view; 4 holotype, lateral view. Images by Harry Taylor (Natural History Museum London)." captionText-1="Figures 59. Habitus. 56 Ivierhipidius cechorum sp. nov.: 5 holotype, dorsal view; 6 holotype, lateral view. 78 I. monneorum sp. nov.: 7 holotype, dorsal view; 8 paratype, lateral view. 9 Ivierhipidius paradoxus sp. nov., paratype, mounted metathoracic wing. Images by Harry Taylor (Natural History Museum London)." captionText-2="Figs 1015. 1011 detail of heart-shaped, deeply indented metatrochanter (10 I. paradoxus sp. nov., paratype; 11 I. monneorum sp. nov., holotype). 1215 detail of head, frontal view: (12 I. paradoxus sp. nov., paratype; 13 I. youngi sp. nov., holotype; 14 I. cechorum sp. nov., paratype; 15 I. monneorum sp. nov., paratype). Scale bars = 1 mm. Images by Harry Taylor (Natural History Museum, London)." figureDoi-0="http://doi.org/10.5281/zenodo.5303721" figureDoi-1="http://doi.org/10.5281/zenodo.5303723" figureDoi-2="http://doi.org/10.5281/zenodo.5303725" httpUri-0="https://zenodo.org/record/5303721/files/figure.png" httpUri-1="https://zenodo.org/record/5303723/files/figure.png" httpUri-2="https://zenodo.org/record/5303725/files/figure.png" pageId="1" pageNumber="692">Figs 115</figureCitation>
)
</paragraph>
<paragraph id="406E6265FF8E307EFFD658C6164FFDEE" blockId="1.[113,872,952,1007]" box="[113,631,952,975]" pageId="1" pageNumber="692">
<emphasis id="72A5BE77FF8E307EFFD658C6157CFDEE" bold="true" box="[113,324,952,975]" pageId="1" pageNumber="692">Family placement.</emphasis>
<taxonomicName id="87D119E6FF8E307EFEED58C615EDFDEE" authorityName="Barclay &amp; Sw" authorityYear="2015" box="[330,469,952,975]" class="Insecta" family="Ripiphoridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="692" phylum="Arthropoda" rank="family">Ripiphoridae</taxonomicName>
<emphasis id="72A5BE77FF8E307EFE7B58C6164FFDEE" box="[476,631,952,975]" italics="true" pageId="1" pageNumber="692">incertae sedis.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="08CB31EEFF8E307EFFD658A61750FDCE" box="[113,872,984,1007]" pageId="1" pageNumber="692" type="type_taxon">
<paragraph id="406E6265FF8E307EFFD658A61750FDCE" blockId="1.[113,872,952,1007]" box="[113,872,984,1007]" pageId="1" pageNumber="692">
<emphasis id="72A5BE77FF8E307EFFD658A6153DFDCE" bold="true" box="[113,261,984,1007]" pageId="1" pageNumber="692">
<typeStatus id="9F6ADCC7FF8E307EFFD658A61491FDCE" box="[113,169,984,1007]" pageId="1" pageNumber="692">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="87D119E6FF8E307EFEAB58A61633FDCE" authority="Barclay &amp; Sw, 2015" authorityName="Barclay &amp; Sw" authorityYear="2015" box="[268,523,984,1007]" class="Insecta" family="Ripiphoridae" genus="Ivierhipidius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="692" phylum="Arthropoda" rank="species" species="paradoxus" status="sp. nov.">
<emphasis id="72A5BE77FF8E307EFEAB58A61633FDCE" box="[268,523,984,1007]" italics="true" pageId="1" pageNumber="692">Ivierhipidius paradoxus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6996030CFF8E307EFDB658A6165EFDCE" box="[529,614,984,1007]" pageId="1" pageNumber="692" rank="species">sp. nov.</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="08CB31EEFF8E307BFFD65F791661FC13" lastPageId="4" lastPageNumber="695" pageId="1" pageNumber="692" type="description">
<paragraph id="406E6265FF8E307BFFD65F791661FC13" blockId="1.[113,1090,1031,1566]" lastBlockId="4.[151,1127,219,850]" lastPageId="4" lastPageNumber="695" pageId="1" pageNumber="692">
<emphasis id="72A5BE77FF8E307EFFD65F7914C4FA3F" bold="true" box="[113,252,1031,1054]" pageId="1" pageNumber="692">Description.</emphasis>
Only known from males:
<quantity id="8729CF80FF8E307EFDBA5F791693FA3F" box="[541,683,1031,1054]" metricMagnitude="-3" metricUnit="m" metricValue="6.699999999999999" metricValueMax="10.2" metricValueMin="3.2" pageId="1" pageNumber="692" unit="mm" value="6.699999999999999" valueMax="10.2" valueMin="3.2">3.210.2 mm</quantity>
(combined length of head, pronotum and elytra measured individually). Dorsal colour varying from rich orange brown to black, in some species bicolored with elytra contrasting with head and pronotum. Head prognathous. Mandibles elongate, strongly recurved, laterally flattened, untoothed. Maxillary palpi tetramerous, with enlarged palpifer that can resemble an additional palpomere. Apical palpomere enlarged, almost securiform, with distal sensory excavation, and with distinct smooth impunctate impression on dorsal surface. Labrum fused to frontoclypeus to form a distinct process. Labial palps apparently absent. Antennae filiform, with 11 antennomeres, 11
<superScript id="B7A4CF2DFF8E307EFC035F981788FAD5" attach="left" box="[932,944,1254,1268]" fontSize="6" pageId="1" pageNumber="692">th</superScript>
antennomere elongated, 1.82.3× length of 10
<superScript id="B7A4CF2DFF8E307EFE6F5E7815ECFB35" attach="none" box="[456,468,1286,1300]" fontSize="6" pageId="1" pageNumber="692">th</superScript>
. Head broadest across eyes. Eyes large but not contiguous. Front and back of head unite to form a strong transverse carina at vertex, which closes the gap with raised anterior margin of pronotum when head raised. Pronotum wider than head at widest point (at base of apical third), more or less bell-shaped, basal two-thirds of pronotum parallel-sided. Pubescence of pronotum long, semirecumbent, directed towards pronotal disk. Margins of pronotum sinuate at both sides and base either side of hind angle, basal margin more-or-less straight in middle. Scutellum distinct. Elytra with marked shoulders, broader than pronotum, parallel-sided, elytral length shoulderapex 1.72.4× as long as maximum width, rounded at apices, punctured and pubescent, weakly shining. Hind wing (
<figureCitation id="D8EA7EE0FF8E307EFC635D79103BF83F" box="[964,1027,1543,1566]" captionStart="Figures 59" captionStartId="3.[108,172,1340,1359]" captionTargetBox="[163,1423,402,1314]" captionTargetId="figure-10@3.[148,1435,391,1336]" captionTargetPageId="3" captionText="Figures 59. Habitus. 56 Ivierhipidius cechorum sp. nov.: 5 holotype, dorsal view; 6 holotype, lateral view. 78 I. monneorum sp. nov.: 7 holotype, dorsal view; 8 paratype, lateral view. 9 Ivierhipidius paradoxus sp. nov., paratype, mounted metathoracic wing. Images by Harry Taylor (Natural History Museum London)." figureDoi="http://doi.org/10.5281/zenodo.5303723" httpUri="https://zenodo.org/record/5303723/files/figure.png" pageId="1" pageNumber="692">Fig. 9</figureCitation>
) with reduced venation, R3 incomplete, reaching RP only as a weakly indicated line RC absent, R4 not indicated. Apical field with dark area beyond R-R3 junction; apical dark area of characteristic shape with three projections, the much longer middle one extending either side of R1 almost as far as RP (see KUKALOVÁ- PECK &amp; LAWRENCE 1993) (note that
<figureCitation id="D8EA7EE0FF8B307BFC415A45101EFF73" box="[998,1062,315,338]" captionStart="Figures 59" captionStartId="3.[108,172,1340,1359]" captionTargetBox="[163,1423,402,1314]" captionTargetId="figure-10@3.[148,1435,391,1336]" captionTargetPageId="3" captionText="Figures 59. Habitus. 56 Ivierhipidius cechorum sp. nov.: 5 holotype, dorsal view; 6 holotype, lateral view. 78 I. monneorum sp. nov.: 7 holotype, dorsal view; 8 paratype, lateral view. 9 Ivierhipidius paradoxus sp. nov., paratype, mounted metathoracic wing. Images by Harry Taylor (Natural History Museum London)." figureDoi="http://doi.org/10.5281/zenodo.5303723" httpUri="https://zenodo.org/record/5303723/files/figure.png" pageId="4" pageNumber="695">Fig. 9</figureCitation>
has a diagonal fold running across the wing that should not be mistaken for a vein). Abdomen with 5 visible ventrites, in most species with a dense brush of conspicuous dark setae covering the base of the second ventrite, overlapped by the translucent apical margin of the first ventrite. Metatrochanter distinct, deeply indented at apex, so heart-shaped (
<figureCitation id="D8EA7EE0FF8B307BFCC35AC517D8FFF3" box="[868,992,443,466]" captionStart="Figs 1015. 1011" captionStartId="6.[151,188,1552,1571]" captionTargetBox="[193,1080,219,1521]" captionTargetId="figure-10@6.[174,1104,219,1534]" captionTargetPageId="6" captionText="Figs 1015. 1011 detail of heart-shaped, deeply indented metatrochanter (10 I. paradoxus sp. nov., paratype; 11 I. monneorum sp. nov., holotype). 1215 detail of head, frontal view: (12 I. paradoxus sp. nov., paratype; 13 I. youngi sp. nov., holotype; 14 I. cechorum sp. nov., paratype; 15 I. monneorum sp. nov., paratype). Scale bars = 1 mm. Images by Harry Taylor (Natural History Museum, London)." figureDoi="http://doi.org/10.5281/zenodo.5303725" httpUri="https://zenodo.org/record/5303725/files/figure.png" pageId="4" pageNumber="695">Figs 10, 11</figureCitation>
). Legs long, slender, unmodified. Tibiae long, metatibia around 1.5× length of corresponding femur or tarsus. Two tibial spurs, often small, present on all tibiae. Tarsi long, unlobed and unmodified. Tarsal claws simple. Tarsal formula 5-5-4.
</paragraph>
</subSubSection>
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<paragraph id="406E6265FF8D307DFF305EA61545F801" blockId="2.[151,1128,1496,1568]" pageId="2" pageNumber="693">
Figs 14. Habitus. 12
<taxonomicName id="87D119E6FF8D307DFED15EA6167BFBCA" authority="Barclay &amp; Sw, 2015" authorityName="Barclay &amp; Sw" authorityYear="2015" box="[374,579,1496,1515]" class="Insecta" family="Ripiphoridae" genus="Ivierhipidius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="693" phylum="Arthropoda" rank="species" species="paradoxus" status="sp. nov.">
<emphasis id="72A5BE77FF8D307DFED15EA6167BFBCA" box="[374,579,1496,1515]" italics="true" pageId="2" pageNumber="693">Ivierhipidius paradoxus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6996030CFF8D307DFDED5EA616A9FBCA" box="[586,657,1496,1515]" pageId="2" pageNumber="693" rank="species">sp. nov.</taxonomicNameLabel>
: 1 holotype, dorsal view; 2 paratype, lateral view. 34
<taxonomicName id="87D119E6FF8D307DFF685E8C1525F824" authority="Barclay &amp; Sw, 2015" authorityName="Barclay &amp; Sw" authorityYear="2015" box="[207,285,1522,1541]" class="Insecta" family="Ripiphoridae" genus="Ivierhipidius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="693" phylum="Arthropoda" rank="species" species="youngi" status="sp. nov.">
<emphasis id="72A5BE77FF8D307DFF685E8C1525F824" box="[207,285,1522,1541]" italics="true" pageId="2" pageNumber="693">I. youngi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6996030CFF8D307DFE835E8C1553F824" box="[292,363,1522,1541]" pageId="2" pageNumber="693" rank="species">sp. nov.</taxonomicNameLabel>
: 3 holotype, dorsal view; 4 holotype, lateral view. Images by Harry Taylor (Natural History Museum London).
</paragraph>
</caption>
<caption id="14AE32EDFF8C307CFFCB5E421534FBA5" ID-DOI="http://doi.org/10.5281/zenodo.5303723" ID-Zenodo-Dep="5303723" httpUri="https://zenodo.org/record/5303723/files/figure.png" pageId="3" pageNumber="694" startId="3.[108,172,1340,1359]" targetBox="[163,1423,402,1314]" targetPageId="3">
<paragraph id="406E6265FF8C307CFFCB5E421534FBA5" blockId="3.[108,1476,1340,1412]" pageId="3" pageNumber="694">
Figures 59. Habitus. 56
<taxonomicName id="87D119E6FF8C307CFEC45E421611FB6E" authority="Barclay &amp; Sw, 2015" authorityName="Barclay &amp; Sw" authorityYear="2015" box="[355,553,1340,1359]" class="Insecta" family="Ripiphoridae" genus="Ivierhipidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="694" phylum="Arthropoda" rank="species" species="cechorum" status="sp. nov.">
<emphasis id="72A5BE77FF8C307CFEC45E421611FB6E" box="[355,553,1340,1359]" italics="true" pageId="3" pageNumber="694">Ivierhipidius cechorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6996030CFF8C307CFD975E42164FFB6E" box="[560,631,1340,1359]" pageId="3" pageNumber="694" rank="species">sp. nov.</taxonomicNameLabel>
: 5 holotype, dorsal view; 6 holotype, lateral view. 78
<taxonomicName id="87D119E6FF8C307CFB235E4210C5FB6E" authority="Barclay &amp; Sw, 2015" authorityName="Barclay &amp; Sw" authorityYear="2015" box="[1156,1277,1340,1359]" class="Insecta" family="Ripiphoridae" genus="Ivierhipidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="694" phylum="Arthropoda" rank="species" species="monneorum" status="sp. nov.">
<emphasis id="72A5BE77FF8C307CFB235E4210C5FB6E" box="[1156,1277,1340,1359]" italics="true" pageId="3" pageNumber="694">I. monneorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6996030CFF8C307CFAA45E421172FB6E" box="[1283,1354,1340,1359]" pageId="3" pageNumber="694" rank="species">sp. nov.</taxonomicNameLabel>
: 7 holotype, dorsal view; 8 paratype, lateral view. 9
<taxonomicName id="87D119E6FF8C307CFE4E5E29168EFB4B" authority="Barclay &amp; Sw, 2015" authorityName="Barclay &amp; Sw" authorityYear="2015" box="[489,694,1367,1386]" class="Insecta" family="Ripiphoridae" genus="Ivierhipidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="694" phylum="Arthropoda" rank="species" species="paradoxus" status="sp. nov.">
<emphasis id="72A5BE77FF8C307CFE4E5E29168EFB4B" box="[489,694,1367,1386]" italics="true" pageId="3" pageNumber="694">Ivierhipidius paradoxus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6996030CFF8C307CFD1A5E29173BFB4B" box="[701,771,1367,1386]" pageId="3" pageNumber="694" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, mounted metathoracic wing. Images by Harry Taylor (Natural History Museum London).
</paragraph>
</caption>
<subSubSection id="08CB31EEFF8B307BFF3059451749FCF3" pageId="4" pageNumber="695" type="diagnosis">
<paragraph id="406E6265FF8B307BFF3059451749FCF3" blockId="4.[151,1127,219,850]" pageId="4" pageNumber="695">
<emphasis id="72A5BE77FF8B307BFF3059451534FC73" bold="true" box="[151,268,571,594]" pageId="4" pageNumber="695">Diagnosis.</emphasis>
Distinguished from all other genera of
<taxonomicName id="87D119E6FF8B307BFD1159451779FC73" authorityName="Barclay &amp; Sw" authorityYear="2015" box="[694,833,571,594]" class="Insecta" family="Ripiphoridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="695" phylum="Arthropoda" rank="family">Ripiphoridae</taxonomicName>
by the following combination of characters: Antennae simple, symmetrical and unmodified. Elytra complete, simple, unmodified. Legs unmodified, all tibiae with two tibial spurs, tarsi with simple claws. Metatrochanter of characteristic shape, deeply indented at apex, heart-shaped (
<figureCitation id="D8EA7EE0FF8B307BFC0959E51012FC93" box="[942,1066,667,690]" captionStart="Figs 1015. 1011" captionStartId="6.[151,188,1552,1571]" captionTargetBox="[193,1080,219,1521]" captionTargetId="figure-10@6.[174,1104,219,1534]" captionTargetPageId="6" captionText="Figs 1015. 1011 detail of heart-shaped, deeply indented metatrochanter (10 I. paradoxus sp. nov., paratype; 11 I. monneorum sp. nov., holotype). 1215 detail of head, frontal view: (12 I. paradoxus sp. nov., paratype; 13 I. youngi sp. nov., holotype; 14 I. cechorum sp. nov., paratype; 15 I. monneorum sp. nov., paratype). Scale bars = 1 mm. Images by Harry Taylor (Natural History Museum, London)." figureDoi="http://doi.org/10.5281/zenodo.5303725" httpUri="https://zenodo.org/record/5303725/files/figure.png" pageId="4" pageNumber="695">Figs 1011</figureCitation>
). The shape of the metatrochanter is apparently apomorphic for this taxon.
</paragraph>
</subSubSection>
<subSubSection id="08CB31EEFF8B307BFF3059A51028FD73" pageId="4" pageNumber="695" type="etymology">
<paragraph id="406E6265FF8B307BFF3059A51028FD73" blockId="4.[151,1127,219,850]" pageId="4" pageNumber="695">
<emphasis id="72A5BE77FF8B307BFF3059A5152FFCD3" bold="true" box="[151,279,731,754]" pageId="4" pageNumber="695">Etymology.</emphasis>
The name is derived from the suffix -
<emphasis id="72A5BE77FF8B307BFD3E59A516C0FCD3" box="[665,760,731,754]" italics="true" pageId="4" pageNumber="695">rhipidius</emphasis>
, taken from the name of the similar
<taxonomicName id="87D119E6FF8B307BFF3059851605FD33" authority="Iablokoff-Khnzorian, 1986" authorityName="Iablokoff-Khnzorian" authorityYear="1986" box="[151,573,763,786]" class="Insecta" family="Ripiphoridae" genus="Eorhipidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="695" phylum="Arthropoda" rank="genus">
<emphasis id="72A5BE77FF8B307BFF305985152CFD33" box="[151,276,763,786]" italics="true" pageId="4" pageNumber="695">Eorhipidius</emphasis>
Iablokoff-Khnzorian, 1986
</taxonomicName>
(now a synonym of
<taxonomicName id="87D119E6FF8B307BFC8459851014FD33" authority="Reitter, 1895" authorityName="Reitter" authorityYear="1895" box="[803,1068,763,786]" class="Insecta" genus="Pterydrias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="695" phylum="Arthropoda" rank="genus">
<emphasis id="72A5BE77FF8B307BFC84598517ACFD33" box="[803,916,763,786]" italics="true" pageId="4" pageNumber="695">Pterydrias</emphasis>
Reitter, 1895
</taxonomicName>
), and the name Ivie in honour of Michael and Ladonna Ivie of Bozeman,
<collectingRegion id="8215AC87FF8B307BFCCB586517F6FD13" box="[876,974,795,818]" country="United States of America" name="Montana" pageId="4" pageNumber="695">Montana</collectingRegion>
, for their kind hospitality and expert and persistent encouragement. The genus name is masculine.
</paragraph>
</subSubSection>
</treatment>
</document>