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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.912.47719" ID-GBIF-Dataset="26396469-500e-42b8-acba-f6a22f652923" ID-PMC="PMC7040049" ID-Pensoft-Pub="1313-2970-912-65" ID-Pensoft-UUID="D1FDF644344C5B5B83E6024904E40A05" ID-PubMed="32123500" ID-ZooBank="4F361E38B2684B70944DD6B6FCC66542" ModsDocID="1313-2970-912-65" checkinTime="1581958640118" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2020" docId="D3A047A92396570EA75DE9C3F536B26A" docLanguage="en" docName="ZooKeys 912: 65-124" docOrigin="ZooKeys 912" docSource="http://dx.doi.org/10.3897/zookeys.912.47719" docTitle="Pseudotheopea kimotoi Lee &amp; Bezděk 2020, sp. nov." docType="treatment" docUuid="5A493FF3-7DD0-4DCB-81FF-893F6FCA9420" docUuidSource="ZooBank" docVersion="4" id="D1FDF644344C5B5B83E6024904E40A05" lastPageNumber="65" masterDocId="D1FDF644344C5B5B83E6024904E40A05" masterDocTitle="Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species" masterLastPageNumber="124" masterPageNumber="65" pageNumber="65" updateTime="1668168482312" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of the Theopea genus group (Coleoptera, Chrysomelidae, Galerucinae), part III: Descriptions of two new genera and nine new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Jan</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>912</mods:number>
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<mods:start>65</mods:start>
<mods:end>124</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.912.47719</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.912.47719</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-912-65</mods:identifier>
<mods:identifier type="ZooBank">4F361E38B2684B70944DD6B6FCC66542</mods:identifier>
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<treatment ID-GBIF-Taxon="162147249" LSID="urn:lsid:zoobank.org:act:5A493FF3-7DD0-4DCB-81FF-893F6FCA9420" httpUri="http://treatment.plazi.org/id/D3A047A92396570EA75DE9C3F536B26A" lastPageNumber="65" pageId="0" pageNumber="65">
<subSubSection pageId="0" pageNumber="65" type="nomenclature">
<paragraph pageId="0" pageNumber="65">
<taxonomicName LSID="D3A047A9-2396-570E-A75D-E9C3F536B26A" authority="Lee &amp; Bezděk, 2020" authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea kimotoi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="kimotoi" status="sp. nov.">Pseudotheopea kimotoi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="65">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Pseudotheopea kimotoi sp. nov., P. nigrita, and P. leehsuehae sp. nov. A P. kimotoi sp. nov., male, dorsal view B Same, ventral view C P. kimotoi sp. nov., female, dorsal view D P. kimotoi sp. nov., male, from Dalat E P. nigrita, holotype, dorsal view F P. nigrita, male, dorsal view G P. leehsuehae sp. nov., male, dorsal view H Same, ventral view I P. leehsuehae sp. nov., female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure18" httpUri="https://binary.pensoft.net/fig/380673" pageId="0" pageNumber="65">Figs 18A-D</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Heads of Pseudotheopea kimotoi sp. nov., P. sufangae sp. nov., and P. sauteri. A P. kimotoi sp. nov., male, dorsofrontal view B Same, front view C P. sufangae sp. nov., male, dorsofrontal view D Same, front view E P. sufangae sp. nov., female, dorsofrontal view F P. sauteri, female, dorsofrontal view." figureDoi="10.3897/zookeys.912.47719.figure19" httpUri="https://binary.pensoft.net/fig/380674" pageId="0" pageNumber="65">; 19A, B</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">; 20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="reference_group">
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Chujo.Solid" authorityYear="1935" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea sauteri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="sauteri">Theopea sauteri</taxonomicName>
: Kimoto, 1989: 200 (part); Medvedev, 2000: 178 (part).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="types">
<paragraph pageId="0" pageNumber="65">Types.</paragraph>
<paragraph pageId="0" pageNumber="65">
Holotype ♂ (BPBM): &quot;VIET NAM: 7km SE / of Dilinh (Djiring) / 990m, 2.V.1960 [p, w] // ♂ [p, w] // R. E. Leech / Collector / BISHOP [p, w] // Theopea / sauteri / Chujo [h] / Det. S. Kimoto, 19 [p] 87 [h, w]&quot;. This specimen was misidentified as
<emphasis italics="true" pageId="0" pageNumber="65">Theopea sauteri</emphasis>
by
<bibRefCitation author="Kimoto, S" journalOrPublisher="Esakia" pageId="0" pageNumber="65" pagination="1 - 241" refId="B10" refString="Kimoto, S, 1989. Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos, and Vietnam. IV. Gallerucinaea. Esakia 27: 1 - 241" title="Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos, and Vietnam. IV. Gallerucinaea." volume="27" year="1989">Kimoto (1989)</bibRefCitation>
. Paratypes.
<emphasis bold="true" pageId="0" pageNumber="65">LAOS</emphasis>
. Boli Kham Xai: 1♀ (JBCB), Ban Nape (8km NE), ~600 m,
<geoCoordinate degrees="18" direction="north" minutes="21" orientation="latitude" precision="925" value="18.35">18°21'N</geoCoordinate>
<geoCoordinate degrees="105" direction="east" minutes="08" orientation="longitude" precision="925" value="105.13333">105°08'E</geoCoordinate>
, 1-18.V.2001, leg. C. L.
<normalizedToken originalValue="Peša">Pesa</normalizedToken>
; Champassak: 1♂, 5♀♀ (HNHM), Dong Hua Xao NBCA, 2 km S of Ban Nong Luang, bank of Touay-Guai Stream,
<geoCoordinate degrees="15" direction="north" minutes="04" orientation="latitude" precision="925" value="15.066667">15°04'N</geoCoordinate>
<geoCoordinate degrees="106" direction="east" minutes="13" orientation="longitude" precision="925" value="106.21667">106°13'E</geoCoordinate>
, 800 m, 1-5.IV.1998, leg. O. Merkl and G. Csorba (identified as
<taxonomicName authorityName="Chujo.Solid" authorityYear="1935" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea sauteri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea sauteri</emphasis>
</taxonomicName>
by
<bibRefCitation author="Medvedev, LN" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="65" pagination="161 - 182" refId="B13" refString="Medvedev, LN, 2000. Chrysomelidae (Coleoptera) of Laos from the collection of the Hungarian Natural History Museum. Annales Historico-Naturales Musei Nationalis Hungarici 92: 161 - 182" title="Chrysomelidae (Coleoptera) of Laos from the collection of the Hungarian Natural History Museum." volume="92" year="2000">Medvedev (2000)</bibRefCitation>
); Hua Pan: 3♀♀ (JBCB), Ban Kangpabong env., 25km SE Vieng Xai (by road),
<geoCoordinate degrees="20" direction="north" minutes="19" orientation="latitude" precision="925" value="20.316668">20°19'N</geoCoordinate>
<geoCoordinate degrees="104" direction="east" minutes="25" orientation="longitude" precision="925" value="104.416664">104°25'E</geoCoordinate>
, 14-18.V.2001, leg. J.
<normalizedToken originalValue="Bezděk">Bezdek</normalizedToken>
; Khammouane: 2♀♀ (RBCN), Nakai env., Rout no 8,
<geoCoordinate degrees="17" direction="north" minutes="42.8" orientation="latitude" precision="92" value="17.713333">17°42.8'N</geoCoordinate>
<geoCoordinate degrees="105" direction="east" minutes="09.1" orientation="longitude" precision="92" value="105.151665">105°09.1'E</geoCoordinate>
, 560 m, 4-8.V.1998, leg. E. Jendek and O.
<normalizedToken originalValue="Šauša">Sausa</normalizedToken>
; Louangphrabang: 4♀♀ (NHMB), Thong Khan,
<geoCoordinate degrees="19" direction="north" minutes="33" orientation="latitude" precision="925" value="19.55">19°33'N</geoCoordinate>
<geoCoordinate degrees="101" direction="east" minutes="58" orientation="longitude" precision="925" value="101.96667">101°58'E</geoCoordinate>
, 750 m, 11-21.V.2002, leg. V.
<normalizedToken originalValue="Kubáň">Kuban</normalizedToken>
;
<emphasis bold="true" pageId="0" pageNumber="65">THAILAND</emphasis>
. Loei: 1♀ (NMPC), Phu Kradung N.P., 16-17.V.1999, leg. D. Hauck;
<emphasis bold="true" pageId="0" pageNumber="65">VIETNAM</emphasis>
. 1♀ (ZSM), Tam Dao, 1982, leg. L. Medvedev; Cao Bang: 1♂ (NMPC), Bao-Lac; Lam Dong: 1♂, 1♀ (BPBM), 6 km S Dalat, 1400-1500 m, 6.VI.-7.VII.1961, leg. N. R. Spencer, identified as
<taxonomicName authorityName="Chujo.Solid" authorityYear="1935" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theopea sauteri" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis italics="true" pageId="0" pageNumber="65">Theopea sauteri</emphasis>
</taxonomicName>
by
<bibRefCitation author="Kimoto, S" journalOrPublisher="Esakia" pageId="0" pageNumber="65" pagination="1 - 241" refId="B10" refString="Kimoto, S, 1989. Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos, and Vietnam. IV. Gallerucinaea. Esakia 27: 1 - 241" title="Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos, and Vietnam. IV. Gallerucinaea." volume="27" year="1989">Kimoto (1989)</bibRefCitation>
; Ninh Binh: 1♀ (NHMB), Cuc Phuong N.P., 21-27.V.1996, leg.
<normalizedToken originalValue="Pacholátko">Pacholatko</normalizedToken>
and
<normalizedToken originalValue="Dembický">Dembicky</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="description">
<paragraph pageId="0" pageNumber="65">Description.</paragraph>
<paragraph pageId="0" pageNumber="65">
Length 6.6-7.5 mm, width 2.6-3.2 mm. Body color (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Pseudotheopea kimotoi sp. nov., P. nigrita, and P. leehsuehae sp. nov. A P. kimotoi sp. nov., male, dorsal view B Same, ventral view C P. kimotoi sp. nov., female, dorsal view D P. kimotoi sp. nov., male, from Dalat E P. nigrita, holotype, dorsal view F P. nigrita, male, dorsal view G P. leehsuehae sp. nov., male, dorsal view H Same, ventral view I P. leehsuehae sp. nov., female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure18" httpUri="https://binary.pensoft.net/fig/380673" pageId="0" pageNumber="65">18A-D</figureCitation>
) metallic blue or purple, but antennae and legs yellowish brown, mouth parts dark brown. Frontoclypeus (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Heads of Pseudotheopea kimotoi sp. nov., P. sufangae sp. nov., and P. sauteri. A P. kimotoi sp. nov., male, dorsofrontal view B Same, front view C P. sufangae sp. nov., male, dorsofrontal view D Same, front view E P. sufangae sp. nov., female, dorsofrontal view F P. sauteri, female, dorsofrontal view." figureDoi="10.3897/zookeys.912.47719.figure19" httpUri="https://binary.pensoft.net/fig/380674" pageId="0" pageNumber="65">19A, B</figureCitation>
) transverse and weakly excavated between eyes in males, semi-circular, the annular concavity 0.8
<normalizedToken originalValue="×">x</normalizedToken>
as wide as interspace between eyes, with cluster of long setae near middle of anterior margin, some shorter setae scattered along anterior margin. Antennae filiform in males (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20A</figureCitation>
), but relatively broader than those of females (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus of Pseudotheopea hsingtzungi sp. nov. and P. smaragdina. A P. hsingtzungi sp. nov., male, dorsal view B P. hsingtzungi sp. nov., female, dorsal view C Same, ventral view D P. smaragdina, male, dorsal view E P. smaragdina, female, dorsal view F Same, ventral view." figureDoi="10.3897/zookeys.912.47719.figure15" httpUri="https://binary.pensoft.net/fig/380670" pageId="0" pageNumber="65">15A</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.3: 0.9: 1.1: 1.2: 1.0: 1.1: 1.0: 1.0: 0.9: 1.1, length to width ratios of antennomeres I-XI 3.3: 1.6: 3.4: 4.4: 4.9: 4.2: 4.8: 4.5: 4.7: 4.8: 4.7; filiform in females (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20B</figureCitation>
), length ratios of antennomeres I-XI 1.0: 0.3: 0.7: 0.9: 1.0: 0.9: 0.9: 0.8: 0.8: 0.8: 0.9, length to width ratios of antennomeres I-XI 4.2: 2.2: 4.1: 5.1: 5.6: 5.4: 5.7: 5.1: 5.5: 5.2: 6.2. Elytra elongate, parallel-sided, 1.7-1.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; disc with dense, coarse punctures, arranged into longitudinal rows, with one distinct and convex longitudinal ridge between two longitudinal rows of punctures, with convex area behind scutellum, ridges reduced at convex area. Tarsomeres I of front legs strongly swollen in males; subparallel in females. Aedeagus (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20D-F</figureCitation>
) slender, 8.4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex with shallow notch; tectum elongate, from apical 1/13 to basal 1/3; almost straight in lateral view, angular at apical 2/5, moderately curved near base; triangular sclerites elongate; internal sac covered with stout teeth, with elongate endophallic sclerite, 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus, some small, stout teeth at apical 1/8 to 2/5. Gonocoxae (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20H</figureCitation>
) elongate, both gonocoxae fused from basal 1/4 to apical 1/5; apices convergent and narrowly rounded, each gonocoxa with eight setae along lateral margin from apex to apical 1/6, four much longer than others; lateral processes reduced. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20G</figureCitation>
) elongate and well sclerotized; disc with several long setae at sides and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20I</figureCitation>
) strongly swollen; pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, narrow and short.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="variation">
<paragraph pageId="0" pageNumber="65">Variation.</paragraph>
<paragraph pageId="0" pageNumber="65">
One male collected from Dalat has a smaller body (5.3 mm long, 2.2 mm wide) and the convex area on the elytra is indistinct and with longitudinal ridges (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20D</figureCitation>
), antennomeres IV-VI are curved (VII-XI lost, Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20C</figureCitation>
), and the frontoclypeus lacks a concavity.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="diagnosis">
<paragraph pageId="0" pageNumber="65">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea kimotoi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea kimotoi</emphasis>
</taxonomicName>
sp. nov. is similar to
<taxonomicName lsidName="P. clypealis" pageId="0" pageNumber="65" rank="species" species="clypealis">
<emphasis italics="true" pageId="0" pageNumber="65">P. clypealis</emphasis>
</taxonomicName>
(Medvedev) and
<taxonomicName lsidName="P. leehsuehae" pageId="0" pageNumber="65" rank="species" species="leehsuehae">
<emphasis italics="true" pageId="0" pageNumber="65">P. leehsuehae</emphasis>
</taxonomicName>
sp. nov. based on the convex and distinct longitudinal ridges on the elytra but differs in having all longitudinal ridges convex and distinct (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Pseudotheopea kimotoi sp. nov., P. nigrita, and P. leehsuehae sp. nov. A P. kimotoi sp. nov., male, dorsal view B Same, ventral view C P. kimotoi sp. nov., female, dorsal view D P. kimotoi sp. nov., male, from Dalat E P. nigrita, holotype, dorsal view F P. nigrita, male, dorsal view G P. leehsuehae sp. nov., male, dorsal view H Same, ventral view I P. leehsuehae sp. nov., female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure18" httpUri="https://binary.pensoft.net/fig/380673" pageId="0" pageNumber="65">18A, C, D</figureCitation>
) (intertwined with convex distinct ridges and weak indistinct ones in others (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D, F</figureCitation>
;
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Pseudotheopea kimotoi sp. nov., P. nigrita, and P. leehsuehae sp. nov. A P. kimotoi sp. nov., male, dorsal view B Same, ventral view C P. kimotoi sp. nov., female, dorsal view D P. kimotoi sp. nov., male, from Dalat E P. nigrita, holotype, dorsal view F P. nigrita, male, dorsal view G P. leehsuehae sp. nov., male, dorsal view H Same, ventral view I P. leehsuehae sp. nov., female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure18" httpUri="https://binary.pensoft.net/fig/380673" pageId="0" pageNumber="65">18F, H</figureCitation>
), with a convex area surrounding the scutellum and longitudinal ridges reduced on the convex area in males (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Pseudotheopea kimotoi sp. nov., P. nigrita, and P. leehsuehae sp. nov. A P. kimotoi sp. nov., male, dorsal view B Same, ventral view C P. kimotoi sp. nov., female, dorsal view D P. kimotoi sp. nov., male, from Dalat E P. nigrita, holotype, dorsal view F P. nigrita, male, dorsal view G P. leehsuehae sp. nov., male, dorsal view H Same, ventral view I P. leehsuehae sp. nov., female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure18" httpUri="https://binary.pensoft.net/fig/380673" pageId="0" pageNumber="65">18A</figureCitation>
) (without convex area surrounding scutellum in those of others (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., male, dorsal view B Same, ventral view C P. boreri, female, dorsal view D P. clypealis, male, dorsal view E Same, ventral view F P. clypealis, female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure8" httpUri="https://binary.pensoft.net/fig/380663" pageId="0" pageNumber="65">8D</figureCitation>
,
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Pseudotheopea kimotoi sp. nov., P. nigrita, and P. leehsuehae sp. nov. A P. kimotoi sp. nov., male, dorsal view B Same, ventral view C P. kimotoi sp. nov., female, dorsal view D P. kimotoi sp. nov., male, from Dalat E P. nigrita, holotype, dorsal view F P. nigrita, male, dorsal view G P. leehsuehae sp. nov., male, dorsal view H Same, ventral view I P. leehsuehae sp. nov., female, dorsal view." figureDoi="10.3897/zookeys.912.47719.figure18" httpUri="https://binary.pensoft.net/fig/380673" pageId="0" pageNumber="65">18F</figureCitation>
)) and the shallow concavity between it in males (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Heads of Pseudotheopea kimotoi sp. nov., P. sufangae sp. nov., and P. sauteri. A P. kimotoi sp. nov., male, dorsofrontal view B Same, front view C P. sufangae sp. nov., male, dorsofrontal view D Same, front view E P. sufangae sp. nov., female, dorsofrontal view F P. sauteri, female, dorsofrontal view." figureDoi="10.3897/zookeys.912.47719.figure19" httpUri="https://binary.pensoft.net/fig/380674" pageId="0" pageNumber="65">19A, B</figureCitation>
) (deep concavity in others (Figs
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Heads of males of Pseudotheopea boreri sp. nov. and P. clypealis. A P. boreri sp. nov., dorsofrontal view B Same, front view C P. clypealis, dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure9" httpUri="https://binary.pensoft.net/fig/380664" pageId="0" pageNumber="65">9C, D</figureCitation>
;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of males of Pseudotheopea gressitti sp. nov. and P. leehsuehae sp. nov. A P. gressitti sp. nov., dorsofrontal view B Same, front view C P. leehsuehae sp. nov., dorsofrontal view D Same, front view." figureDoi="10.3897/zookeys.912.47719.figure13" httpUri="https://binary.pensoft.net/fig/380668" pageId="0" pageNumber="65">13C, D</figureCitation>
)). Males of
<taxonomicName lsidName="P. kimotoi" pageId="0" pageNumber="65" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">P. kimotoi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20E, F</figureCitation>
) are similar to those of
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17D, E</figureCitation>
) in possessing elongate triangular aedeagal sclerites but differ in the absence of lateral sclerites attached to the median elongate sclerite, with small spines near apex of median elongate sclerite, which is undivided (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Diagnostic characters of Pseudotheopea kimotoi sp. nov. A Antenna, male B Antenna, female C Antenna, male, from Dalat D Aedeagus, dorsal view E Aedeagus, lateral view F Aedeagus, ventral view G Abdominal ventrite VIII H Gonocoxae I Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65">20D, E</figureCitation>
) (with one pair of hook-like sclerites at sides of median elongate sclerite, without small spines near apex of median elongate sclerite, and divided at middle in those of
<taxonomicName lsidName="P. hsingtzungi" pageId="0" pageNumber="65" rank="species" species="hsingtzungi">
<emphasis italics="true" pageId="0" pageNumber="65">P. hsingtzungi</emphasis>
</taxonomicName>
sp. nov. Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Diagnostic characters of Pseudotheopea hsingtzungi sp. nov. A Antenna, male B Antenna, female C Aedeagus, dorsal view D Aedeagus, lateral view E Aedeagus, ventral view F Abdominal ventrite VIII G Gonocoxae H Spermatheca." figureDoi="10.3897/zookeys.912.47719.figure17" httpUri="https://binary.pensoft.net/fig/380672" pageId="0" pageNumber="65">17C, D</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="etymology">
<paragraph pageId="0" pageNumber="65">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="65">
This new species is dedicated to late Dr. Shinsaku Kimoto for his great contribution to taxonomy of oriental and Palaearctic
<taxonomicName authorityName="Latreille" authorityYear="1802" family="Chrysomelidae" lsidName="" pageId="0" pageNumber="65" rank="family">Chrysomelidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="distribution">
<paragraph pageId="0" pageNumber="65">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="65">Laos, Thailand, Vietnam.</paragraph>
<caption doi="10.3897/zookeys.912.47719.figure18" httpUri="https://binary.pensoft.net/fig/380673" pageId="0" pageNumber="65" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 18.</emphasis>
Habitus of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea kimotoi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea kimotoi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="P. nigrita" pageId="0" pageNumber="65" rank="species" species="nigrita">
<emphasis italics="true" pageId="0" pageNumber="65">P. nigrita</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="P. leehsuehae" pageId="0" pageNumber="65" rank="species" species="leehsuehae">
<emphasis italics="true" pageId="0" pageNumber="65">P. leehsuehae</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
<taxonomicName lsidName="P. kimotoi" pageId="0" pageNumber="65" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">P. kimotoi</emphasis>
</taxonomicName>
sp. nov., male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Same, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
<taxonomicName lsidName="P. kimotoi" pageId="0" pageNumber="65" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">P. kimotoi</emphasis>
</taxonomicName>
sp. nov., female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
<taxonomicName lsidName="P. kimotoi" pageId="0" pageNumber="65" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">P. kimotoi</emphasis>
</taxonomicName>
sp. nov., male, from Dalat
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
<taxonomicName lsidName="P. nigrita" pageId="0" pageNumber="65" rank="species" species="nigrita">
<emphasis italics="true" pageId="0" pageNumber="65">P. nigrita</emphasis>
</taxonomicName>
, holotype, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
<taxonomicName lsidName="P. nigrita" pageId="0" pageNumber="65" rank="species" species="nigrita">
<emphasis italics="true" pageId="0" pageNumber="65">P. nigrita</emphasis>
</taxonomicName>
, male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
<taxonomicName lsidName="P. leehsuehae" pageId="0" pageNumber="65" rank="species" species="leehsuehae">
<emphasis italics="true" pageId="0" pageNumber="65">P. leehsuehae</emphasis>
</taxonomicName>
sp. nov., male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
Same, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">I</emphasis>
<taxonomicName lsidName="P. leehsuehae" pageId="0" pageNumber="65" rank="species" species="leehsuehae">
<emphasis italics="true" pageId="0" pageNumber="65">P. leehsuehae</emphasis>
</taxonomicName>
sp. nov., female, dorsal view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.912.47719.figure19" httpUri="https://binary.pensoft.net/fig/380674" pageId="0" pageNumber="65" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 19.</emphasis>
Heads of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea kimotoi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea kimotoi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="P. sufangae" pageId="0" pageNumber="65" rank="species" species="sufangae">
<emphasis italics="true" pageId="0" pageNumber="65">P. sufangae</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName lsidName="P. sauteri" pageId="0" pageNumber="65" rank="species" species="sauteri">
<emphasis italics="true" pageId="0" pageNumber="65">P. sauteri</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
<taxonomicName lsidName="P. kimotoi" pageId="0" pageNumber="65" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">P. kimotoi</emphasis>
</taxonomicName>
sp. nov., male, dorsofrontal view
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Same, front view
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
<taxonomicName lsidName="P. sufangae" pageId="0" pageNumber="65" rank="species" species="sufangae">
<emphasis italics="true" pageId="0" pageNumber="65">P. sufangae</emphasis>
</taxonomicName>
sp. nov., male, dorsofrontal view
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
Same, front view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
<taxonomicName lsidName="P. sufangae" pageId="0" pageNumber="65" rank="species" species="sufangae">
<emphasis italics="true" pageId="0" pageNumber="65">P. sufangae</emphasis>
</taxonomicName>
sp. nov., female, dorsofrontal view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
<taxonomicName lsidName="P. sauteri" pageId="0" pageNumber="65" rank="species" species="sauteri">
<emphasis italics="true" pageId="0" pageNumber="65">P. sauteri</emphasis>
</taxonomicName>
, female, dorsofrontal view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.912.47719.figure20" httpUri="https://binary.pensoft.net/fig/380675" pageId="0" pageNumber="65" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="65">
<emphasis bold="true" pageId="0" pageNumber="65">Figure 20.</emphasis>
Diagnostic characters of
<taxonomicName authorityName="Lee &amp; Bezděk" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Pseudotheopea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudotheopea kimotoi" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="kimotoi">
<emphasis italics="true" pageId="0" pageNumber="65">Pseudotheopea kimotoi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
Antenna, male
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
Antenna, female
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
Antenna, male, from Dalat
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
Aedeagus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
Aedeagus, lateral view
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
Aedeagus, ventral view
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
Abdominal ventrite VIII
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
Gonocoxae
<emphasis bold="true" pageId="0" pageNumber="65">I</emphasis>
Spermatheca.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>