307 lines
37 KiB
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307 lines
37 KiB
XML
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<mods:title id="9615263E632C7B724BB11278F2B31D6A">Niuailan, a new genus to accommodate Carneoryctes peterseni (Endrődi, 1967) (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) from New Ireland, Papua New Guinea</mods:title>
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<mods:name id="FF25D6163E416BE9645D42A5D991C9A3" type="personal">
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<mods:namePart id="B891B1B4D52E408374BB03753852D327">Allsopp, Peter G.</mods:namePart>
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<mods:affiliation id="843AFE4CF93B4A6118D429D9AA37EC1E">Queensland Museum, PO Box 3300, South Brisbane 4101, Australia.</mods:affiliation>
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<mods:namePart id="9422F65E68609122DB236D250CF99DAB">Hutchinson, Paul M.</mods:namePart>
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<mods:affiliation id="DC08F5E53144668C4B03B2F394716158">Department of Primary Industries and Regional Development (Quarantine WA), Locked Bag 69, Welshpool DC 6986, Australia. paul. hutchinson @ dpird. wa. gov. au; https: // orcid. org / 0000 - 0001 - 8747 - 1320</mods:affiliation>
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<mods:nameIdentifier id="B42D409BD620814373347691FCAAEB7B" type="email">paul.hutchinson@dpird.wa.gov.au</mods:nameIdentifier>
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<mods:title id="7BAA452840E22C8C558E93F54A5AAD53">Zootaxa</mods:title>
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<mods:date id="322CD0AFCE9DF582EF9CB5C6B609B6F2">2020</mods:date>
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<mods:number id="7069E994EBB416A9B5CB24F2EE9C4DAD">2020-10-09</mods:number>
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<mods:identifier id="BC467BF1142EA497AD91170B4A158D15" type="DOI">10.11646/zootaxa.4860.1.8</mods:identifier>
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<treatment id="C8668C46FFC2FFCF9BEFF4DB851EFDF8" ID-DOI="http://doi.org/10.5281/zenodo.4535829" ID-GBIF-Taxon="177319461" ID-Zenodo-Dep="4535829" LSID="urn:lsid:plazi:treatment:C8668C46FFC2FFCF9BEFF4DB851EFDF8" httpUri="http://treatment.plazi.org/id/C8668C46FFC2FFCF9BEFF4DB851EFDF8" lastPageId="4" lastPageNumber="139" pageId="2" pageNumber="137">
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<subSubSection id="08D56EDBFFC2FFC99BEFF4DB8610FCF4" box="[151,522,763,792]" pageId="2" pageNumber="137" type="nomenclature">
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<paragraph id="40703D50FFC2FFC99BEFF4DB8610FCF4" blockId="2.[151,522,763,792]" box="[151,522,763,792]" pageId="2" pageNumber="137">
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<heading id="1B388A3CFFC2FFC99BEFF4DB8610FCF4" bold="true" box="[151,522,763,792]" fontSize="11" level="1" pageId="2" pageNumber="137" reason="1">
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<taxonomicName id="87CF46D3FFC2FFC99BEFF4DB8610FCF4" authority="Ozdikmen, 2009" authorityName="Ozdikmen" authorityYear="2009" box="[151,522,763,792]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
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<emphasis id="72BBE142FFC2FFC99BEFF4DB8610FCF4" bold="true" box="[151,522,763,792]" pageId="2" pageNumber="137">
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<emphasis id="72BBE142FFC2FFC99BEFF4DB852DFCF4" bold="true" box="[151,311,763,789]" italics="true" pageId="2" pageNumber="137">Carneoryctes</emphasis>
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<bibRefCitation id="245E40A1FFC2FFC99A46F4DC8610FCF4" author="Ozdikmen, H." box="[318,522,763,792]" pageId="2" pageNumber="137" pagination="139 - 147" refId="ref4203" refString="Ozdikmen, H. (2009) Nomenclatural changes for five preoccupied scarab beetle genus group names (Coleoptera: Scarabaeidae). Munis Entomology & Zoology, 4, 139 - 147." type="journal article" year="2009">Özdikmen, 2009</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="08D56EDBFFC2FFC99BEFF56086C0FC97" pageId="2" pageNumber="137" type="reference_group">
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<paragraph id="40703D50FFC2FFC99BEFF560801DFCB9" blockId="2.[151,1031,831,889]" box="[151,1031,831,856]" pageId="2" pageNumber="137">
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<treatmentCitationGroup id="60DF1A7EFFC2FFC99BEFF560801DFCB9" box="[151,1031,831,856]" pageId="2" pageNumber="137">
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<treatmentCitation id="C16E1B41FFC2FFC99BEFF56085CAFCB6" author="Carne, P. B." box="[151,464,831,856]" page="154" pageId="2" pageNumber="137" year="1957">
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<taxonomicName id="87CF46D3FFC2FFC99BEFF56085CAFCB6" ID-CoL="3WSL" authority="Carne, 1957: 154" authorityName="Carne" authorityPageNumber="154" authorityYear="1957" box="[151,464,831,856]" class="Insecta" family="Scarabaeidae" genus="Cryptoryctes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
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<emphasis id="72BBE142FFC2FFC99BEFF5608500FCB6" box="[151,282,832,855]" italics="true" pageId="2" pageNumber="137">Cryptoryctes</emphasis>
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<bibRefCitation id="245E40A1FFC2FFC99A58F56085CAFCB6" author="Carne, P. B." box="[288,464,831,856]" pageId="2" pageNumber="137" refId="ref3747" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957: 154</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(preoccupied by
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<taxonomicName id="87CF46D3FFC2FFC999F8F5608765FCB6" ID-CoL="3WSL" authority="Reed, 1954" authorityName="Reed" authorityYear="1954" box="[640,895,831,856]" class="Insecta" family="Scarabaeidae" genus="Cryptoryctes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
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<emphasis id="72BBE142FFC2FFC999F8F5608719FCB6" box="[640,771,832,855]" italics="true" pageId="2" pageNumber="137">Cryptoryctes</emphasis>
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<bibRefCitation id="245E40A1FFC2FFC99871F5608765FCB6" author="Reed, C. A." box="[777,895,831,856]" pageId="2" pageNumber="137" pagination="102 - 111" refId="ref4237" refString="Reed, C. A. (1954) Some fossorial mammals from the Tertiary of western North America. Journal of Paleontology, 28, 102 - 111." type="journal article" year="1954">Reed, 1954</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="87CF46D3FFC2FFC998F1F56087E6FCB9" authorityName="Linnaeus" authorityYear="1758" box="[905,1020,832,856]" class="Mammalia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="137" phylum="Arthropoda" rank="class">Mammalia</taxonomicName>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="40703D50FFC2FFC99BEFF57F86C0FC97" blockId="2.[151,1031,831,889]" box="[151,730,862,889]" pageId="2" pageNumber="137">
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<treatmentCitationGroup id="60DF1A7EFFC2FFC99BEFF57F86C0FC97" box="[151,730,862,889]" pageId="2" pageNumber="137">
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<treatmentCitation id="C16E1B41FFC2FFC99BEFF57F8618FC97" author="Ozdikmen, H." box="[151,514,862,889]" page="139" pageId="2" pageNumber="137" year="2009">
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<taxonomicName id="87CF46D3FFC2FFC99BEFF57F8618FC97" authority="Ozdikmen, 2009: 139" authorityName="Ozdikmen" authorityPageNumber="139" authorityYear="2009" box="[151,514,862,889]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
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<emphasis id="72BBE142FFC2FFC99BEFF57F8505FC97" box="[151,287,863,886]" italics="true" pageId="2" pageNumber="137">Carneoryctes</emphasis>
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<bibRefCitation id="245E40A1FFC2FFC99A5EF5408618FC97" author="Ozdikmen, H." box="[294,514,862,889]" pageId="2" pageNumber="137" pagination="139 - 147" refId="ref4203" refString="Ozdikmen, H. (2009) Nomenclatural changes for five preoccupied scarab beetle genus group names (Coleoptera: Scarabaeidae). Munis Entomology & Zoology, 4, 139 - 147." type="journal article" year="2009">Özdikmen, 2009: 139</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(replacement name).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="08D56EDBFFC2FFC99BEFF5BC8785FC55" box="[151,927,923,948]" pageId="2" pageNumber="137" type="type_taxon">
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<paragraph id="40703D50FFC2FFC99BEFF5BC8785FC55" blockId="2.[151,927,923,948]" box="[151,927,923,948]" pageId="2" pageNumber="137">
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<emphasis id="72BBE142FFC2FFC99BEFF5BC8539FC52" bold="true" box="[151,291,923,948]" pageId="2" pageNumber="137">
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<typeStatus id="9F7483F2FFC2FFC99BEFF5BC84D6FC55" box="[151,204,924,948]" pageId="2" pageNumber="137">Type</typeStatus>
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species.
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</emphasis>
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||
<taxonomicName id="87CF46D3FFC2FFC99A52F5BC86BEFC55" authority="Blackburn, 1892" authorityName="Blackburn" authorityYear="1892" box="[298,676,924,948]" class="Insecta" family="Dynastidae" genus="Pseudoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="tectus">
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<emphasis id="72BBE142FFC2FFC99A52F5BC85E8FC52" box="[298,498,924,948]" italics="true" pageId="2" pageNumber="137">Pseudoryctes tectus</emphasis>
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Blackburn, 1892
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="08D56EDBFFC2FFCF9BEFF5C1851EFDF8" lastPageId="4" lastPageNumber="139" pageId="2" pageNumber="137" type="description">
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<paragraph id="40703D50FFC2FFC99BEFF5C18634F964" blockId="2.[151,1437,993,1741]" pageId="2" pageNumber="137">
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<emphasis id="72BBE142FFC2FFC99BEFF5C18593FC1D" bold="true" box="[151,393,993,1020]" pageId="2" pageNumber="137">Redescription. Male</emphasis>
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(from
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<bibRefCitation id="245E40A1FFC2FFC99AAEF5C28641FC1C" author="Carne, P. B." box="[470,603,994,1021]" pageId="2" pageNumber="137" refId="ref3747" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne 1957</bibRefCitation>
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;
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<bibRefCitation id="245E40A1FFC2FFC9991FF5C28719FC1C" author="Endrodi, S." box="[615,771,994,1021]" pageId="2" pageNumber="137" refId="ref4044" refString="Endrodi, S. (1985) The Dynastinae of the World. Junk, Dordrecht, Netherlands, 800 pp." type="book" year="1985">Endrődi 1985</bibRefCitation>
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; and P.M.H. specimens) (
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<figureCitation id="D8F421D5FFC2FFC99F5AF5C28080FC1D" box="[1058,1178,994,1020]" captionStart-0="FIGURES 3–7" captionStart-1="FIGURES 8–13" captionStartId-0="3.[151,264,1971,1995]" captionStartId-1="4.[151,264,1492,1516]" captionTargetBox-0="[187,1403,191,1941]" captionTargetBox-1="[168,1424,562,1469]" captionTargetId-0="figure-17@3.[163,1424,181,1947]" captionTargetId-1="figure-125@4.[151,1436,561,1469]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 3–7. Carneoryctes tectus (Blackburn, 1892), male in PMH: 3, dorsal view; 4, ventral view; 5, lateral view; 6, para- meres frontal view; 7, parameres and phallobase lateral view." captionText-1="FIGURES 8–13. Carneoryctes males in PMH: 8, C. brittoni (Carne, 1957), lateral view; 9, C. brittoni, dorsal view; 10, C. psilus (Carne, 1957), lateral view; 11, C. psilus, dorsal view; 12, C. wingarus (Carne, 1957), lateral view; 13, C. wingarus, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.4413597" figureDoi-1="http://doi.org/10.5281/zenodo.4413599" httpUri-0="https://zenodo.org/record/4413597/files/figure.png" httpUri-1="https://zenodo.org/record/4413599/files/figure.png" pageId="2" pageNumber="137">Figs. 3–13</figureCitation>
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). Body reddish brown, bicolorous, or black,
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<quantity id="873790B5FFC2FFC99AFEF226861AFBC1" box="[390,512,1030,1056]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.7" metricValueMin="1.2" pageId="2" pageNumber="137" unit="mm" value="19.5" valueMax="27.0" valueMin="12.0">12–27 mm</quantity>
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long. Mentum strongly compressed and reflexed at ligula; maxillary palp slender, apical palpomere truncate; galea reduced to small, untoothed pieces; labrum usually visible; mandibles visible. Antenna with 10 antennomeres, club longer than shaft. Clypeus at angle to plane of frons, nearly always verti- cal, concave and with truncate anterior face glabrous or setose, disc rarely setose. Clypeofrontal ridge transverse, strongly elevated. Frons frequently setose; ocular canthi large, rounded or subangulate, densely setose. Pronotum with anterior margin corneous and bearing median elevated process or horn, latter tapering to a simple, bifid, trifid, or branched apex; anterolateral angles acute or obtuse; lateral ridges sometimes contracted or interrupted just before basolateral angles, their margins frequently with dorsal setae; basal ridge sometimes continuous; disc often setose, transversely rugulose with deep excavation limited by elevated angles or lateral horns, latter usually simple, disc rarely depressed below level of basal margin. Each elytron with sutural stria linear or linear-punctate, disc with welldefined striae, punctures simple or weakly umbilicate, surface sometimes microreticulate, epipleuron with short lateral setae continuous to apices. Pygidium setose across base, apical ridge glabrous or setose. Postcoxal prosternal process short, setose, often not visible. Abdominal sternites setose across posterior margins. Protibiae tridentate with spur inserted anterior to base of medial denticle; protarsomere 5 usually longer than protarsomere 1, not thick- ened; prothoracic claws simple, slender; metathoracic legs slender or moderately short; metatibiae bicarinate, distal ciliae short, sharp, closely set, spurs unequal, curved; metatarsi elongate, tarsomeres 1 and 5 subequal in length. Parameres symmetrical, external margin broadening pre-apically, in lateral view face arcuate, ratio of phallobase length to paramere length 1.5–2.5:1.
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</paragraph>
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<paragraph id="40703D50FFC2FFC99BBFF0AE869AF92C" blockId="2.[151,1437,993,1741]" pageId="2" pageNumber="137">
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<emphasis id="72BBE142FFC2FFC99BBFF0AE858BF946" bold="true" box="[199,401,1677,1704]" pageId="2" pageNumber="137">Included species.</emphasis>
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General geographic ranges (all in
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<collectingCountry id="38D87DC0FFC2FFC99877F0AE8762F948" box="[783,888,1678,1705]" name="Australia" pageId="2" pageNumber="137">Australia</collectingCountry>
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) were taken from
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<bibRefCitation id="245E40A1FFC2FFC99F3AF0AE80CAF948" author="Carne, P. B." box="[1090,1232,1678,1705]" pageId="2" pageNumber="137" refId="ref3747" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne (1957</bibRefCitation>
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,
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<bibRefCitation id="245E40A1FFC2FFC99FA2F0AE810FF949" author="Carne, P. B." box="[1242,1301,1678,1704]" pageId="2" pageNumber="137" pagination="249 - 252" refId="ref3780" refString="Carne, P. B. (1981) Cryptoryctes minchami sp. n., and notes on other Cryptoryctes spp. (Coleoptera: Scarabaeidae: Dynastinae). Journal of the Australian Entomological Society, 20, 249 - 252. https: // doi. org / 10.1111 / j. 1440 - 6055.1981. tb 01042. x" type="journal article" year="1981">1981</bibRefCitation>
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),
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<bibRefCitation id="245E40A1FFC2FFC99E5FF0AE8534F92D" author="Allsopp, P. G. & Lloyd, R. L." pageId="2" pageNumber="137" pagination="71 - 76" refId="ref3636" refString="Allsopp, P. G. & Lloyd, R. L. (1987) New records of Scarabaeidae (Coleoptera) from central Australia. Australian Entomological Magazine, 13, 71 - 76." type="journal article" year="1987">Allsopp & Lloyd (1987)</bibRefCitation>
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, and
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<bibRefCitation id="245E40A1FFC2FFC99A11F0928666F92C" author="Howden, H. F. & Maly, V." box="[361,636,1714,1741]" pageId="2" pageNumber="137" pagination="274 - 277" refId="ref4097" refString="Howden, H. F. & Maly, V. (2005) Cryptoryctes tricornutus sp. nov. from the Northern Territory, Australia (Scarabaeidae, Dynas- tinae, Oryctini). Canadian Entomologist, 137, 274 - 277. https: // doi. org / 10.4039 / n 04 - 102" type="journal article" year="2005">Howden & Malý (2005)</bibRefCitation>
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.
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</paragraph>
|
||
<paragraph id="40703D50FFC2FFC99BEFF0DA874AF8F5" blockId="2.[151,1054,1786,2029]" box="[151,848,1786,1813]" pageId="2" pageNumber="137">
|
||
<taxonomicName id="87CF46D3FFC2FFC99BEFF0DA85FDF8F4" authority="(Lea, 1917)" baseAuthorityName="Lea" baseAuthorityYear="1917" box="[151,487,1786,1813]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="ater">
|
||
<emphasis id="72BBE142FFC2FFC99BEFF0DA8578F8F5" box="[151,354,1786,1812]" italics="true" pageId="2" pageNumber="137">Carneoryctes ater</emphasis>
|
||
(Lea, 1917)
|
||
</taxonomicName>
|
||
—southwestern
|
||
<collectingRegion id="820BF3B2FFC2FFC999DBF0DA874AF8F5" box="[675,848,1786,1812]" country="Australia" name="South Australia" pageId="2" pageNumber="137">South Australia</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC2FFC99BEFF13E87A6F8D9" blockId="2.[151,1054,1786,2029]" box="[151,956,1822,1849]" pageId="2" pageNumber="137">
|
||
<taxonomicName id="87CF46D3FFC2FFC99BEFF13E863DF8D8" authority="(Carne, 1957)" baseAuthorityName="Carne" baseAuthorityYear="1957" box="[151,551,1822,1849]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="brittoni">
|
||
<emphasis id="72BBE142FFC2FFC99BEFF13E8593F8D9" box="[151,393,1822,1848]" italics="true" pageId="2" pageNumber="137">Carneoryctes brittoni</emphasis>
|
||
(
|
||
<bibRefCitation id="245E40A1FFC2FFC99AE0F13E863AF8D8" author="Carne, P. B." box="[408,544,1822,1849]" pageId="2" pageNumber="137" refId="ref3747" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
—inland-western
|
||
<collectingRegion id="820BF3B2FFC2FFC9998CF13E87A6F8D9" box="[756,956,1822,1848]" country="Australia" name="Western Australia" pageId="2" pageNumber="137">Western Australia</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC2FFC99BEFF16287ABF8BD" blockId="2.[151,1054,1786,2029]" box="[151,945,1858,1885]" pageId="2" pageNumber="137">
|
||
<taxonomicName id="87CF46D3FFC2FFC99BEFF1628655F8BC" authority="(Lea, 1917)" baseAuthorityName="Lea" baseAuthorityYear="1917" box="[151,591,1858,1885]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="griseopilosus">
|
||
<emphasis id="72BBE142FFC2FFC99BEFF16285D0F8BD" box="[151,458,1858,1884]" italics="true" pageId="2" pageNumber="137">Carneoryctes griseopilosus</emphasis>
|
||
(Lea, 1917)
|
||
</taxonomicName>
|
||
—southeastern
|
||
<collectingRegion id="820BF3B2FFC2FFC9987BF16287ABF8BD" box="[771,945,1858,1884]" country="Australia" name="South Australia" pageId="2" pageNumber="137">South Australia</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC2FFC99BEFF14687A6F861" blockId="2.[151,1054,1786,2029]" box="[151,956,1894,1921]" pageId="2" pageNumber="137">
|
||
<taxonomicName id="87CF46D3FFC2FFC99BEFF146865AF860" authority="(Carne, 1981)" baseAuthorityName="Carne" baseAuthorityYear="1981" box="[151,576,1894,1921]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="minchami">
|
||
<emphasis id="72BBE142FFC2FFC99BEFF14685B9F861" box="[151,419,1894,1920]" italics="true" pageId="2" pageNumber="137">Carneoryctes minchami</emphasis>
|
||
(
|
||
<bibRefCitation id="245E40A1FFC2FFC99ACAF1468623F860" author="Carne, P. B." box="[434,569,1894,1921]" pageId="2" pageNumber="137" pagination="249 - 252" refId="ref3780" refString="Carne, P. B. (1981) Cryptoryctes minchami sp. n., and notes on other Cryptoryctes spp. (Coleoptera: Scarabaeidae: Dynastinae). Journal of the Australian Entomological Society, 20, 249 - 252. https: // doi. org / 10.1111 / j. 1440 - 6055.1981. tb 01042. x" type="journal article" year="1981">Carne, 1981</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
—central-eastern
|
||
<collectingRegion id="820BF3B2FFC2FFC99876F14687A6F861" box="[782,956,1894,1920]" country="Australia" name="South Australia" pageId="2" pageNumber="137">South Australia</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC2FFC99BEFF1AA8004F845" blockId="2.[151,1054,1786,2029]" box="[151,1054,1930,1957]" pageId="2" pageNumber="137">
|
||
<taxonomicName id="87CF46D3FFC2FFC99BEFF1AA8699F844" authority="(Blackburn, 1895)" baseAuthorityName="Blackburn" baseAuthorityYear="1895" box="[151,643,1930,1957]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="monstrosus">
|
||
<emphasis id="72BBE142FFC2FFC99BEFF1AA85AEF845" box="[151,436,1930,1956]" italics="true" pageId="2" pageNumber="137">Carneoryctes monstrosus</emphasis>
|
||
(Blackburn, 1895)
|
||
</taxonomicName>
|
||
—central-western
|
||
<collectingRegion id="820BF3B2FFC2FFC9982FF1AA8004F845" box="[855,1054,1930,1956]" country="Australia" name="Western Australia" pageId="2" pageNumber="137">Western Australia</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC2FFC99BEFF18E8789F829" blockId="2.[151,1054,1786,2029]" box="[151,915,1966,1993]" pageId="2" pageNumber="137">
|
||
<taxonomicName id="87CF46D3FFC2FFC99BEFF18E8621F828" authority="(Lea, 1917)" baseAuthorityName="Lea" baseAuthorityYear="1917" box="[151,571,1966,1993]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="nigripennis">
|
||
<emphasis id="72BBE142FFC2FFC99BEFF18E85AFF829" box="[151,437,1966,1992]" italics="true" pageId="2" pageNumber="137">Carneoryctes nigripennis</emphasis>
|
||
(Lea, 1917)
|
||
</taxonomicName>
|
||
—inland southern
|
||
<collectingRegion id="820BF3B2FFC2FFC99877F18E8789F829" box="[783,915,1966,1992]" country="Australia" name="Queensland" pageId="2" pageNumber="137">Queensland</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC2FFC99BEFF1F28760F80D" blockId="2.[151,1054,1786,2029]" box="[151,890,2002,2029]" pageId="2" pageNumber="137">
|
||
<taxonomicName id="87CF46D3FFC2FFC99BEFF1F2863FF80C" authority="(Carne, 1957)" baseAuthorityName="Carne" baseAuthorityYear="1957" box="[151,549,2002,2029]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="pimbus">
|
||
<emphasis id="72BBE142FFC2FFC99BEFF1F2859CF80D" box="[151,390,2002,2028]" italics="true" pageId="2" pageNumber="137">Carneoryctes pimbus</emphasis>
|
||
(
|
||
<bibRefCitation id="245E40A1FFC2FFC99AEDF1F28607F80C" author="Carne, P. B." box="[405,541,2002,2029]" pageId="2" pageNumber="137" refId="ref3747" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
—southern
|
||
<collectingRegion id="820BF3B2FFC2FFC999D4F1F28760F80D" box="[684,890,2002,2028]" country="Australia" name="Northern Territory" pageId="2" pageNumber="137">Northern Territory</collectingRegion>
|
||
</paragraph>
|
||
<caption id="14B06DD8FFC3FFC89BEFF193871EF80E" ID-DOI="http://doi.org/10.5281/zenodo.4413597" ID-Zenodo-Dep="4413597" httpUri="https://zenodo.org/record/4413597/files/figure.png" pageId="3" pageNumber="138" startId="3.[151,264,1971,1995]" targetBox="[187,1403,191,1941]" targetPageId="3">
|
||
<paragraph id="40703D50FFC3FFC89BEFF193871EF80E" blockId="3.[151,1436,1971,2031]" pageId="3" pageNumber="138">
|
||
<emphasis id="72BBE142FFC3FFC89BEFF1938521F82A" bold="true" box="[151,315,1971,1995]" pageId="3" pageNumber="138">FIGURES 3–7.</emphasis>
|
||
<taxonomicName id="87CF46D3FFC3FFC89A39F19386D7F82A" authority="(Blackburn, 1892)" baseAuthorityName="Blackburn" baseAuthorityYear="1892" box="[321,717,1971,1995]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="tectus">
|
||
<emphasis id="72BBE142FFC3FFC89A39F1938611F82A" box="[321,523,1971,1995]" italics="true" pageId="3" pageNumber="138">Carneoryctes tectus</emphasis>
|
||
(Blackburn, 1892)
|
||
</taxonomicName>
|
||
, male in PMH:
|
||
<emphasis id="72BBE142FFC3FFC89817F1938766F82A" bold="true" box="[879,892,1971,1995]" pageId="3" pageNumber="138">3</emphasis>
|
||
, dorsal view;
|
||
<emphasis id="72BBE142FFC3FFC89F74F1938003F82A" bold="true" box="[1036,1049,1971,1995]" pageId="3" pageNumber="138">4</emphasis>
|
||
, ventral view;
|
||
<emphasis id="72BBE142FFC3FFC89FC9F19380A4F82A" bold="true" box="[1201,1214,1971,1995]" pageId="3" pageNumber="138">5</emphasis>
|
||
, lateral view;
|
||
<emphasis id="72BBE142FFC3FFC89E37F1938146F82A" bold="true" box="[1359,1372,1971,1995]" pageId="3" pageNumber="138">6</emphasis>
|
||
, para- meres frontal view;
|
||
<emphasis id="72BBE142FFC3FFC89A1AF1F78575F80E" bold="true" box="[354,367,2007,2031]" pageId="3" pageNumber="138">7</emphasis>
|
||
, parameres and phallobase lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="40703D50FFC4FFCF9BEFF6B7809BFF50" blockId="4.[151,1436,150,537]" box="[151,1153,150,177]" pageId="4" pageNumber="139">
|
||
<taxonomicName id="87CF46D3FFC4FFCF9BEFF6B7860CFF50" authority="(Carne, 1957)" authorityName="" baseAuthorityName="Carne" baseAuthorityYear="1957" box="[151,534,150,177]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="psilus">
|
||
<emphasis id="72BBE142FFC4FFCF9BEFF6B7856CFF50" box="[151,374,151,177]" italics="true" pageId="4" pageNumber="139">Carneoryctes psilus</emphasis>
|
||
(
|
||
<bibRefCitation id="245E40A1FFC4FFCF9AFDF6B78614FF50" author="Carne, P. B." box="[389,526,150,177]" pageId="4" pageNumber="139" refId="ref3747" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
—inland and central-western coastal
|
||
<collectingRegion id="820BF3B2FFC4FFCF98C1F6B7809BFF50" box="[953,1153,151,177]" country="Australia" name="Western Australia" pageId="4" pageNumber="139">Western Australia</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC4FFCF9BEFF69B8771FF34" blockId="4.[151,1436,150,537]" box="[151,875,187,214]" pageId="4" pageNumber="139">
|
||
<taxonomicName id="87CF46D3FFC4FFCF9BEFF69B862CFF37" authority="(Lea, 1917)" baseAuthorityName="Lea" baseAuthorityYear="1917" box="[151,566,187,214]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="semicalvus">
|
||
<emphasis id="72BBE142FFC4FFCF9BEFF69B85AAFF34" box="[151,432,187,213]" italics="true" pageId="4" pageNumber="139">Carneoryctes semicalvus</emphasis>
|
||
(Lea, 1917)
|
||
</taxonomicName>
|
||
—southern
|
||
<collectingRegion id="820BF3B2FFC4FFCF99C5F69B8771FF34" box="[701,875,187,213]" country="Australia" name="South Australia" pageId="4" pageNumber="139">South Australia</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC4FFCF9BEFF6FF8790FF18" blockId="4.[151,1436,150,537]" box="[151,906,223,250]" pageId="4" pageNumber="139">
|
||
<taxonomicName id="87CF46D3FFC4FFCF9BEFF6FF862EFF1B" authority="(Arrow, 1914)" baseAuthorityName="Arrow" baseAuthorityYear="1914" box="[151,564,223,250]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="sulcatus">
|
||
<emphasis id="72BBE142FFC4FFCF9BEFF6FF858BFF18" box="[151,401,223,249]" italics="true" pageId="4" pageNumber="139">Carneoryctes sulcatus</emphasis>
|
||
(Arrow, 1914)
|
||
</taxonomicName>
|
||
—southern
|
||
<collectingRegion id="820BF3B2FFC4FFCF99C3F6FF8790FF18" box="[699,906,223,249]" country="Australia" name="Northern Territory" pageId="4" pageNumber="139">Northern Territory</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC4FFCF9BEFF7238767FEA0" blockId="4.[151,1436,150,537]" pageId="4" pageNumber="139">
|
||
<taxonomicName id="87CF46D3FFC4FFCF9BEFF7238648FEFC" authority="(Blackburn, 1892)" baseAuthorityName="Blackburn" baseAuthorityYear="1892" box="[151,594,259,285]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="tectus">
|
||
<emphasis id="72BBE142FFC4FFCF9BEFF7238560FEFD" box="[151,378,259,285]" italics="true" pageId="4" pageNumber="139">Carneoryctes tectus</emphasis>
|
||
(Blackburn, 1892)
|
||
</taxonomicName>
|
||
—western
|
||
<collectingRegion id="820BF3B2FFC4FFCF99ABF7238024FEFC" box="[723,1086,259,285]" isEnumeration="true" pageId="4" pageNumber="139">Victoria and New South Wales</collectingRegion>
|
||
, southern
|
||
<collectingRegion id="820BF3B2FFC4FFCF9FC0F723858FFEA0" isEnumeration="true" pageId="4" pageNumber="139">South Australia and Northern Territory</collectingRegion>
|
||
, and inland northwestern
|
||
<collectingRegion id="820BF3B2FFC4FFCF99CEF7078767FEA0" box="[694,893,295,321]" country="Australia" name="Western Australia" pageId="4" pageNumber="139">Western Australia</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC4FFCF9BEFF76B803BFE84" blockId="4.[151,1436,150,537]" box="[151,1057,330,357]" pageId="4" pageNumber="139">
|
||
<taxonomicName id="87CF46D3FFC4FFCF9BEFF76B86D6FE84" authority="(Howden & Maly, 2005)" baseAuthorityName="Howden & Maly" baseAuthorityYear="2005" box="[151,716,330,357]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="tricornutus">
|
||
<emphasis id="72BBE142FFC4FFCF9BEFF76B85ABFE85" box="[151,433,331,357]" italics="true" pageId="4" pageNumber="139">Carneoryctes tricornutus</emphasis>
|
||
(
|
||
<bibRefCitation id="245E40A1FFC4FFCF9AB8F76B86DFFE84" author="Howden, H. F. & Maly, V." box="[448,709,330,357]" pageId="4" pageNumber="139" pagination="274 - 277" refId="ref4097" refString="Howden, H. F. & Maly, V. (2005) Cryptoryctes tricornutus sp. nov. from the Northern Territory, Australia (Scarabaeidae, Dynas- tinae, Oryctini). Canadian Entomologist, 137, 274 - 277. https: // doi. org / 10.4039 / n 04 - 102" type="journal article" year="2005">Howden & Malý, 2005</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
—southern
|
||
<collectingRegion id="820BF3B2FFC4FFCF982AF76B803BFE84" box="[850,1057,331,357]" country="Australia" name="Northern Territory" pageId="4" pageNumber="139">Northern Territory</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC4FFCF9BEFF74F85A2FE4C" blockId="4.[151,1436,150,537]" pageId="4" pageNumber="139">
|
||
<taxonomicName id="87CF46D3FFC4FFCF9BEFF74F864CFE68" authority="(Blackburn, 1895)" baseAuthorityName="Blackburn" baseAuthorityYear="1895" box="[151,598,366,393]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="trifidus">
|
||
<emphasis id="72BBE142FFC4FFCF9BEFF74F859CFE68" box="[151,390,367,393]" italics="true" pageId="4" pageNumber="139">Carneoryctes trifidus</emphasis>
|
||
(Blackburn, 1895)
|
||
</taxonomicName>
|
||
—inland southern
|
||
<collectingRegion id="820BF3B2FFC4FFCF9851F74F87AAFE68" box="[809,944,367,393]" country="Australia" name="Queensland" pageId="4" pageNumber="139">Queensland</collectingRegion>
|
||
, northern
|
||
<collectingRegion id="820BF3B2FFC4FFCF9F59F74F80F1FE68" box="[1057,1259,367,393]" country="Australia" name="New South Wales" pageId="4" pageNumber="139">New South Wales</collectingRegion>
|
||
, and northwest- ern
|
||
<collectingRegion id="820BF3B2FFC4FFCF9B88F7B385A2FE4C" box="[240,440,403,429]" country="Australia" name="Western Australia" pageId="4" pageNumber="139">Western Australia</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC4FFCF9BEFF7978734FE30" blockId="4.[151,1436,150,537]" box="[151,814,438,465]" pageId="4" pageNumber="139">
|
||
<taxonomicName id="87CF46D3FFC4FFCF9BEFF7978624FE30" authority="(Carne, 1957)" baseAuthorityName="Carne" baseAuthorityYear="1957" box="[151,574,438,465]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="truncatus">
|
||
<emphasis id="72BBE142FFC4FFCF9BEFF7978585FE31" box="[151,415,439,465]" italics="true" pageId="4" pageNumber="139">Carneoryctes truncatus</emphasis>
|
||
(
|
||
<bibRefCitation id="245E40A1FFC4FFCF9AD6F797862CFE30" author="Carne, P. B." box="[430,566,438,465]" pageId="4" pageNumber="139" refId="ref3747" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
—inland
|
||
<collectingRegion id="820BF3B2FFC4FFCF99D2F7978734FE30" box="[682,814,439,465]" country="Australia" name="Queensland" pageId="4" pageNumber="139">Queensland</collectingRegion>
|
||
</paragraph>
|
||
<paragraph id="40703D50FFC4FFCF9BEFF7FB851EFDF8" blockId="4.[151,1436,150,537]" pageId="4" pageNumber="139">
|
||
<taxonomicName id="87CF46D3FFC4FFCF9BEFF7FB8623FE14" authority="(Carne, 1957)" baseAuthorityName="Carne" baseAuthorityYear="1957" box="[151,569,474,501]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="wingarus">
|
||
<emphasis id="72BBE142FFC4FFCF9BEFF7FB8587FE15" box="[151,413,475,501]" italics="true" pageId="4" pageNumber="139">Carneoryctes wingarus</emphasis>
|
||
(
|
||
<bibRefCitation id="245E40A1FFC4FFCF9AD3F7FB8628FE14" author="Carne, P. B." box="[427,562,474,501]" pageId="4" pageNumber="139" refId="ref3747" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
—
|
||
<materialsCitation id="F0A7370DFFC4FFCF992EF7FA851EFDF8" ID-GBIF-Occurrence="3034030302" location="southwestern Northern" pageId="4" pageNumber="139" specimenCount="1" stateProvince="Western Australia">
|
||
<collectingRegion id="820BF3B2FFC4FFCF992EF7FA873BFE14" box="[598,801,474,501]" country="Australia" name="Western Australia" pageId="4" pageNumber="139">Western Australia</collectingRegion>
|
||
(unknown
|
||
<typeStatus id="9F7483F2FFC4FFCF98D8F7FB87CAFE14" box="[928,976,475,501]" pageId="4" pageNumber="139">type</typeStatus>
|
||
locality) and
|
||
<location id="45106B8BFFC4FFCF9F11F7FB8172FE14" LSID="urn:lsid:plazi:treatment:C8668C46FFC2FFCF9BEFF4DB851EFDF8:45106B8BFFC4FFCF9F11F7FB8172FE14" box="[1129,1384,475,501]" name="southwestern Northern" pageId="4" pageNumber="139" stateProvince="Western Australia">southwestern Northern</location>
|
||
Ter- ritory
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |