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<document id="B27D0A63C0A03CC89ABF41145AFC1103" ID-DOI="10.5252/geodiversitas2024v46a2" ID-ISSN="1638-9395" ID-ZooBank="urn:lsid:zoobank.org:pub:A141C894-DEC7-42FD-BC7C-2A8A692B0A31" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706171845289" checkinUser="plazi" docAuthor="Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni &amp; Chény, Cédric" docDate="2024" docId="704E030AFFA8FF9BDD08FD59FC6E2AEC" docLanguage="en" docName="geodiversitas.2024.46.2.pdf" docOrigin="Geodiversitas 46 (2)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a2.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Parahyas Charbonnier &amp; Garassino &amp; Pasini &amp; Chény 2024, n. gen." docType="treatment" docUuid="B38E2FCA-0FC0-43E8-99A4-3F94395BEA5E" docUuidSource="ZooBank" docVersion="1" lastPageNumber="19" masterDocId="8C777B72FFA0FF93D910FFAFFF9D2C21" masterDocTitle="Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria" masterLastPageNumber="29" masterPageNumber="13" pageNumber="19" updateTime="1706171949445" updateUser="GgImagineBatch" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="F0C548CFF3F0E307F4561B8CA48BB7F1">
<mods:title id="A89041A1B38C48BFE1C2AE99BBCFDE3D">Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria</mods:title>
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<mods:namePart id="3D29ADCE53A99D2A11381A948ACF8382">Charbonnier, Sylvain</mods:namePart>
<mods:affiliation id="9F34F977600E2CCE9A65E2CE7E00BCA5">Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), MNHN, CNRS, Sorbonne Université, Muséum national d’Histoire naturelle, 57 rue Cuvier, F- 75231 Paris, cedex 05 (France) sylvain. charbonnier @ mnhn. fr (corresponding author)</mods:affiliation>
<mods:nameIdentifier id="A6D8040F158084E1925E517F9B666EE8" type="email">sylvain.charbonnier@mnhn.fr</mods:nameIdentifier>
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<mods:roleTerm id="3D24AC4E34A1389477DB73B3CD0B619B">Author</mods:roleTerm>
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<mods:namePart id="BCAF45A18465EAFE7F6754277FB187E8">Garassino, Alessandro</mods:namePart>
<mods:affiliation id="265C31D26463F50430E3013DE7142792">Department of Earth and Biological Sciences, Loma Linda University, Loma Linda, 92350 CA (United States) alegarassino @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="D766956B42E045F9BF9B51AEA0225FC7" type="email">alegarassino@gmail.com</mods:nameIdentifier>
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<mods:namePart id="1C271AC36BF4CC403EE0C2ECB002F44A">Pasini, Giovanni</mods:namePart>
<mods:affiliation id="B6FB6C6722F67D3D8A252E9EAB1A4202">Via Alessandro Volta, 16 22070 Appiano Gentile (Italy) giovannialdopasini @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="5B9CA3C57002FA7978E26519942F10A2" type="email">giovannialdopasini@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0E3216684D013D4D173DC83B65143B16">Chény, Cédric</mods:namePart>
<mods:affiliation id="7D5BE77BD5A1782CEDB149A87478200E">Géosciences Rennes, UMR 6118, CNRS Université de Rennes, F- 35000 Rennes (France)</mods:affiliation>
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<mods:title id="295A0F6DBAA118F9E9D50FC6DE9250D7">Geodiversitas</mods:title>
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<mods:date id="099E9A65D2E1FADCF50267BE65DF18CA">2024</mods:date>
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<mods:number id="D57632EA2500F58F150AE48F0D847CD1">2024-01-25</mods:number>
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<mods:number id="CEA9BFBD353C3DBB314910511DEFDB28">46</mods:number>
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<mods:url id="143F2D99ED1C3E3E75024A99CC11874B">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a2.pdf</mods:url>
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<treatment id="704E030AFFA8FF9BDD08FD59FC6E2AEC" LSID="urn:lsid:zoobank.org:act:B38E2FCA-0FC0-43E8-99A4-3F94395BEA5E" httpUri="http://treatment.plazi.org/id/704E030AFFA8FF9BDD08FD59FC6E2AEC" lastPageNumber="19" pageId="8" pageNumber="19">
<subSubSection id="B0FDE197FFA8FF9BDD08FD59FA322F68" pageId="8" pageNumber="19" type="nomenclature">
<paragraph id="F858B21CFFA8FF9BDD08FD59FB592F31" blockId="8.[1048,1220,758,784]" box="[1048,1220,758,784]" pageId="8" pageNumber="19">
<heading id="A3100570FFA8FF9BDD08FD59FB592F31" box="[1048,1220,758,784]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="19" reason="2">
<taxonomicName id="3FE7C99FFFA8FF9BDD08FD59FBEE2F31" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[1048,1139,758,784]" class="Malacostraca" family="Epialtidae" genus="Parahyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CA936E0EFFA8FF9BDD08FD59FBEE2F31" box="[1048,1139,758,784]" italics="true" pageId="8" pageNumber="19">Parahyas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1A0D375FFA8FF9BDD6AFD59FB592F31" box="[1146,1220,758,784]" pageId="8" pageNumber="19" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="F858B21CFFA8FF9BDA3DFC97FA322F68" blockId="8.[813,1455,824,841]" box="[813,1455,824,841]" pageId="8" pageNumber="19">
<uri id="8C76BE1EFFA8FF9BDA3DFC97FA322F68" box="[813,1455,824,841]" pageId="8" pageNumber="19">
urn:lsid:zoobank.org:act:
<uuid id="8C4188C9FFA8FF9BDD17FC97FA322F68" box="[1031,1455,824,841]" pageId="8" pageNumber="19">B38E2FCA-0FC0-43E8-99A4-3F94395BEA5E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA8FF9BDA3DFCD5FC312FE7" pageId="8" pageNumber="19" type="etymology">
<paragraph id="F858B21CFFA8FF9BDA3DFCD5FC312FE7" blockId="8.[813,1455,890,966]" pageId="8" pageNumber="19">
<smallCapsWord id="FEBE24C0FFA8FF9BDA3DFCD5FC3B2FB1" baselines="907,908" box="[813,934,890,913]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="8" pageNumber="19">ETYMOLOGY</smallCapsWord>
. — A combination between
<emphasis id="CA936E0EFFA8FF9BDDDDFCD5FB652FB0" box="[1229,1272,890,913]" italics="true" pageId="8" pageNumber="19">Para</emphasis>
(Latin: similar to) and the extant genus
<taxonomicName id="3FE7C99FFFA8FF9BDD13FC3AFB282F8D" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[1027,1205,916,940]" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA8FF9BDD13FC3AFBAC2F8D" box="[1027,1073,917,940]" italics="true" pageId="8" pageNumber="19">Hyas</emphasis>
Leach, 1814
</taxonomicName>
. The gender of the genus is masculine.
</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA8FF9BDA3DFC70FABB2FD7" box="[813,1318,990,1014]" pageId="8" pageNumber="19" type="materials_examined">
<paragraph id="F858B21CFFA8FF9BDA3DFC70FABB2FD7" blockId="8.[813,1318,990,1014]" box="[813,1318,990,1014]" pageId="8" pageNumber="19">
<smallCapsWord id="FEBE24C0FFA8FF9BDA3DFC70FCC02FD5" baselines="1008,1008" box="[813,861,991,1014]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="8" pageNumber="19">
<typeStatus id="275C0CBEFFA8FF9BDA3DFC70FCC02FD5" box="[813,861,991,1014]" pageId="8" pageNumber="19">TYPE</typeStatus>
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<smallCapsWord id="FEBE24C0FFA8FF9BDA74FC4BFC362FD4" baselines="1009" box="[868,939,996,1013]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="8" pageNumber="19">SPECIES</smallCapsWord>
. —
<taxonomicName id="3FE7C99FFFA8FF9BDAC7FC70FABE2FD7" authority="Van Straelen, 1937" authorityName="Van Straelen" authorityYear="1937" box="[983,1315,990,1014]" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="oranensis">
<emphasis id="CA936E0EFFA8FF9BDAC7FC70FBC22FD7" box="[983,1119,991,1014]" italics="true" pageId="8" pageNumber="19">Hyas oranensis</emphasis>
<bibRefCitation id="9C76CFEDFFA8FF9BDD75FC70FABE2FD7" author="VAN STRAELEN V." box="[1125,1315,990,1014]" pageId="8" pageNumber="19" pagination="473 - 480" refId="ref12191" refString="VAN STRAELEN V. 1937. - Sur quelques crabes du Sahelien moyen des environs d'Oran (Algerie). Bulletin de la Societe geologique de France, serie 5, 6: 473 - 480." type="journal article" year="1937">Van Straelen, 1937</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA8FF9BDA3DFBA1FBD7288A" pageId="8" pageNumber="19" type="diagnosis">
<paragraph id="F858B21CFFA8FF9BDA3DFBA1FBD7288A" blockId="8.[811,1457,1038,1195]" pageId="8" pageNumber="19">
<smallCapsWord id="FEBE24C0FFA8FF9BDA3DFBA1FC0A2805" baselines="1055,1056" box="[813,919,1038,1061]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="8" pageNumber="19">DIAGNOSIS</smallCapsWord>
. — Carapace pyriform; orbit composed of supraorbital rim and widely cup-shaped postorbital spine bearing marginal spines; inflated branchial regions with posterolateral margin bearing row of tubercles; intestinal region protruding distally beyond and under branchial regions; urogastric and cardiac regions limited by deep branchiocardiac groove.
</paragraph>
</subSubSection>
<subSubSection id="B0FDE197FFA8FF9BDA3DFB7AFC6E2AEC" pageId="8" pageNumber="19" type="discussion">
<paragraph id="F858B21CFFA8FF9BDA3DFB7AFC3328CC" blockId="8.[811,1458,1237,1741]" box="[813,942,1237,1263]" pageId="8" pageNumber="19">
<smallCapsWord id="FEBE24C0FFA8FF9BDA3DFB7AFC3328CC" baselines="1256,1256" box="[813,942,1237,1263]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Discussion" pageId="8" pageNumber="19">DISCUSSION</smallCapsWord>
</paragraph>
<paragraph id="F858B21CFFA8FF9BDA3DFB5BFC6E2AEC" blockId="8.[811,1458,1237,1741]" pageId="8" pageNumber="19">
<taxonomicName id="3FE7C99FFFA8FF9BDA3DFB5BFC15292F" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[813,904,1268,1294]" class="Malacostraca" family="Epialtidae" genus="Parahyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CA936E0EFFA8FF9BDA3DFB5BFC15292F" box="[813,904,1268,1294]" italics="true" pageId="8" pageNumber="19">Parahyas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1A0D375FFA8FF9BDA9FFB5AFC46292E" box="[911,987,1269,1295]" pageId="8" pageNumber="19" rank="genus">n. gen.</taxonomicNameLabel>
is assigned to the
<taxonomicName id="3FE7C99FFFA8FF9BDDB5FB5BFAB6292F" authorityName="Garth" authorityYear="1958" box="[1189,1323,1268,1294]" class="Malacostraca" family="Oregoniidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="family">Oregoniidae</taxonomicName>
based upon the widely cup-shaped postorbital spine, which is well visible in several extant representatives such as
<taxonomicName id="3FE7C99FFFA8FF9BDDD7FA9AFCFE294E" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="coarctatus">
<emphasis id="CA936E0EFFA8FF9BDDD7FA9AFAFF296F" box="[1223,1378,1333,1358]" italics="true" pageId="8" pageNumber="19">Hyas coarctatus</emphasis>
Leach, 1815
</taxonomicName>
and
<taxonomicName id="3FE7C99FFFA8FF9BDA88FAFAFB2E294F" authority="Rathbun, 1924" authorityName="Rathbun" authorityYear="1924" box="[920,1203,1364,1390]" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="ursinus">
<emphasis id="CA936E0EFFA8FF9BDA88FAFAFB99294F" box="[920,1028,1365,1390]" italics="true" pageId="8" pageNumber="19">H. ursinus</emphasis>
Rathbun, 1924
</taxonomicName>
.
<taxonomicName id="3FE7C99FFFA8FF9BDDACFAFBFA8A294F" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[1212,1303,1364,1390]" class="Malacostraca" family="Epialtidae" genus="Parahyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CA936E0EFFA8FF9BDDACFAFBFA8A294F" box="[1212,1303,1364,1390]" italics="true" pageId="8" pageNumber="19">Parahyas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1A0D375FFA8FF9BDC0EFAFAFAFA294E" box="[1310,1383,1365,1391]" pageId="8" pageNumber="19" rank="genus">n. gen.</taxonomicNameLabel>
differs from
<taxonomicName id="3FE7C99FFFA8FF9BDA7BFADAFC5629AE" authorityName=", Dana" authorityYear="1851" box="[875,971,1397,1423]" class="Malacostraca" family="Oregoniidae" genus="Oregonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA8FF9BDA7BFADAFC5629AE" box="[875,971,1397,1423]" italics="true" pageId="8" pageNumber="19">Oregonia</emphasis>
</taxonomicName>
,
<taxonomicName id="3FE7C99FFFA8FF9BDACAFADBFBC029AF" authorityName="KrOyer" authorityYear="1838" box="[986,1117,1396,1422]" class="Malacostraca" family="Oregoniidae" genus="Chionoecetes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA8FF9BDACAFADBFBC029AF" box="[986,1117,1396,1422]" italics="true" pageId="8" pageNumber="19">Chionoecetes</emphasis>
</taxonomicName>
,
<taxonomicName id="3FE7C99FFFA8FF9BDD7DFADAFB6129AF" authorityName="Sakai" authorityYear="1978" box="[1133,1276,1397,1422]" class="Malacostraca" family="Oregoniidae" genus="Macroregonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA8FF9BDD7DFADAFB6129AF" box="[1133,1276,1397,1422]" italics="true" pageId="8" pageNumber="19">Macroregonia</emphasis>
</taxonomicName>
by the presence of widely cup-shaped postorbital spine (absent in all these genera).
<taxonomicName id="3FE7C99FFFA8FF9BDA9BFA1BFC7B29EF" authority="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény, 2024" authorityName="Charbonnier &amp; Garassino &amp; Pasini &amp; Chény" authorityYear="2024" box="[907,998,1460,1486]" class="Malacostraca" family="Epialtidae" genus="Parahyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CA936E0EFFA8FF9BDA9BFA1BFC7B29EF" box="[907,998,1460,1486]" italics="true" pageId="8" pageNumber="19">Parahyas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1A0D375FFA8FF9BDAFEFA1BFBA629EF" box="[1006,1083,1460,1486]" pageId="8" pageNumber="19" rank="genus">n. gen.</taxonomicNameLabel>
differs from
<taxonomicName id="3FE7C99FFFA8FF9BDDDDFA1AFA9D29EF" authorityName="Leach" authorityYear="1814" box="[1229,1280,1461,1486]" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA8FF9BDDDDFA1AFA9D29EF" box="[1229,1280,1461,1486]" italics="true" pageId="8" pageNumber="19">Hyas</emphasis>
</taxonomicName>
by the presence of a row of large tubercles extending along the entire length of the posterolateral margin (
<emphasis id="CA936E0EFFA8FF9BDD7FFA5BFB362A2C" box="[1135,1195,1524,1549]" italics="true" pageId="8" pageNumber="19">versus</emphasis>
sometimes scarce small tubercles along the margin in
<taxonomicName id="3FE7C99FFFA8FF9BDD67F9BBFB352A0C" authorityName="Leach" authorityYear="1814" box="[1143,1192,1556,1581]" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA8FF9BDD67F9BBFB352A0C" box="[1143,1192,1556,1581]" italics="true" pageId="8" pageNumber="19">Hyas</emphasis>
</taxonomicName>
), dorsal surface of carapace covered by small tubercles arranged uniformly (
<emphasis id="CA936E0EFFA8FF9BDC63F99BFA322A6C" box="[1395,1455,1588,1613]" italics="true" pageId="8" pageNumber="19">versus</emphasis>
scattered granules and tubercles of varying size in
<taxonomicName id="3FE7C99FFFA8FF9BDC2DF9FBFAED2A4C" authorityName="Leach" authorityYear="1814" box="[1341,1392,1620,1645]" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA8FF9BDC2DF9FBFAED2A4C" box="[1341,1392,1620,1645]" italics="true" pageId="8" pageNumber="19">Hyas</emphasis>
</taxonomicName>
), and gastric, urogastric, and cardiac regions without median raised tubercles (
<emphasis id="CA936E0EFFA8FF9BDA89F93BFC492A8C" box="[921,980,1684,1709]" italics="true" pageId="8" pageNumber="19">versus</emphasis>
above-mentioned regions with raised median tubercles in
<taxonomicName id="3FE7C99FFFA8FF9BDAA0F91BFC7E2AEC" authorityName="Leach" authorityYear="1814" box="[944,995,1716,1741]" class="Malacostraca" family="Oregoniidae" genus="Hyas" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="CA936E0EFFA8FF9BDAA0F91BFC7E2AEC" box="[944,995,1716,1741]" italics="true" pageId="8" pageNumber="19">Hyas</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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