treatments-xml/data/03/F4/BB/03F4BB77FFD5CE29FF03972EDE7BFDA9.xml

<document id="432B30A283238D8D934681DFDA11DAF9" ID-DOI="10.11646/zootaxa.5507.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13747718" ID-ZooBank="065ECECA-5F0B-47BE-82FC-6C1F68B316FD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726055721707" checkinUser="plazi" docAuthor="Teslenko, Valentina A., Palatov, Dmitry M. &amp; Semenchenko, Alexander A." docDate="2024" docId="03F4BB77FFD5CE29FF03972EDE7BFDA9" docLanguage="en" docName="zootaxa.5507.1.1.pdf" docOrigin="Zootaxa 5507 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Perla palatovi Teslenko, Palatov &amp; Semenchenko, 2024, sp. nov." docType="treatment" docUuid="A700A9D0-CF01-401E-9DF4-F8B42DD8995F" docUuidSource="ZooBank" docVersion="2" lastPageNumber="47" masterDocId="FFCDC30FFFF0CE07FF94901CDA4EFFC8" masterDocTitle="Overview of the Caucasian Perla Geoffroy, 1762 (Plecoptera: Perlidae) based on morphological and molecular data with description of two new species" masterLastPageNumber="56" masterPageNumber="1" pageNumber="38" updateTime="1726056859996" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="109578216002DA0A16DF16C728E3F336">Overview of the Caucasian Perla Geoffroy, 1762 (Plecoptera: Perlidae) based on morphological and molecular data with description of two new species</mods:title>
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<mods:namePart id="6A4C040F4A769AEE3C63B368806CE354">Teslenko, Valentina A.</mods:namePart>
<mods:affiliation id="18BB69856AA98396F04D1C2502211C3D">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences (FSC EATB FEB RAS), 690022, Vladivostok, Russia.</mods:affiliation>
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<mods:namePart id="9E8C4B8F69043E6D75B5158FF3A6C8CF">Palatov, Dmitry M.</mods:namePart>
<mods:affiliation id="7B78E7658528F7079AC33CF9FA52E584">Severtsov A. N. Institute of Ecology and Evolution of RAS, 33 Leninsky prospect, 119071 Moscow, Russia.</mods:affiliation>
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<mods:namePart id="85E7C87A3EF63AA28F52864AD41CCCD6">Semenchenko, Alexander A.</mods:namePart>
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<paragraph id="8BE20A61FFD5CE22FF03972ED8A0F884" blockId="37.[151,750,1842,1905]" box="[151,750,1842,1869]" pageId="37" pageNumber="38">
<heading id="D0AABD0DFFD5CE22FF03972ED8A0F884" bold="true" box="[151,750,1842,1869]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
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<taxonomicName id="4C5D71E2FFD5CE22FF03972EDB0EF885" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[151,320,1842,1869]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFD5CE22FF03972EDB0EF885" bold="true" box="[151,320,1842,1869]" italics="true" pageId="37" pageNumber="38">Perla palatovi</emphasis>
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<taxonomicNameLabel id="A21A6B08FFD5CE22FED3972FDBEBF885" box="[327,421,1843,1869]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
Teslenko &amp; Semenchenko
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</paragraph>
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<subSubSection id="C34759EAFFD5CE22FF03974BDB77F8B9" box="[151,313,1878,1905]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BE20A61FFD5CE22FF03974BDB77F8B9" blockId="37.[151,750,1842,1905]" box="[151,313,1878,1905]" pageId="37" pageNumber="38">
<figureCitation id="136616E4FFD5CE22FF03974BDB77F8B9" box="[151,313,1878,1905]" captionStart-0="FIGURES 128‒131" captionStart-1="FIGURES 132‒136" captionStart-2="FIGURES 137‒140" captionStart-3="FIGURES 141‒147" captionStart-4="FIGURES 148‒149" captionStart-5="FIGURES 150‒156" captionStartId-0="39.[151,264,1854,1878]" captionStartId-1="40.[151,264,1836,1860]" captionStartId-2="41.[151,264,1895,1919]" captionStartId-3="42.[151,264,1742,1766]" captionStartId-4="43.[151,264,1489,1513]" captionStartId-5="44.[151,264,1703,1727]" captionTargetBox-0="[151,1436,191,1830]" captionTargetBox-1="[151,1436,181,1813]" captionTargetBox-2="[158,1430,181,1871]" captionTargetBox-3="[170,1417,181,1718]" captionTargetBox-4="[151,1436,181,1465]" captionTargetBox-5="[151,1435,184,1679]" captionTargetId-0="figure-18@39.[151,1436,181,1830]" captionTargetId-1="figure-24@40.[151,1436,181,1813]" captionTargetId-2="figure-18@41.[158,1430,181,1871]" captionTargetId-3="figure-24@42.[170,1417,181,1718]" captionTargetId-4="figure-18@43.[151,1436,181,1465]" captionTargetId-5="figure-24@44.[151,1435,181,1679]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionTargetPageId-3="42" captionTargetPageId-4="43" captionTargetPageId-5="44" captionText-0="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." captionText-1="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." captionText-2="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." captionText-3="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." captionText-4="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." captionText-5="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi-0="http://doi.org/10.5281/zenodo.13747810" figureDoi-1="http://doi.org/10.5281/zenodo.13747812" figureDoi-2="http://doi.org/10.5281/zenodo.13747814" figureDoi-3="http://doi.org/10.5281/zenodo.13747816" figureDoi-4="http://doi.org/10.5281/zenodo.13747818" figureDoi-5="http://doi.org/10.5281/zenodo.13747820" httpUri-0="https://zenodo.org/record/13747810/files/figure.png" httpUri-1="https://zenodo.org/record/13747812/files/figure.png" httpUri-2="https://zenodo.org/record/13747814/files/figure.png" httpUri-3="https://zenodo.org/record/13747816/files/figure.png" httpUri-4="https://zenodo.org/record/13747818/files/figure.png" httpUri-5="https://zenodo.org/record/13747820/files/figure.png" pageId="37" pageNumber="38">Figs. 128−156</figureCitation>
</paragraph>
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<subSubSection id="C34759EAFFD5CE2AFF039782DEE7F999" lastPageId="45" lastPageNumber="46" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="8BE20A61FFD5CE21FF039782D98AFE19" blockId="37.[151,1436,1950,2013]" lastBlockId="38.[151,1437,151,1977]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<emphasis id="B929D673FFD5CE22FF039782DB5CF870" bold="true" box="[151,274,1950,1976]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
Tergum 9 of males is distinguished by the arrangement of serrated ridges on the mesal field that are relatively short and do not reach the posterior margin (
<figureCitation id="136616E4FFD5CE22FD6497DFD914F815" box="[752,858,1987,2013]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="37" pageNumber="38">Figs. 133</figureCitation>
, 
<figureCitation id="136616E4FFD5CE22FCF197DFD9DDF815" box="[869,915,1987,2013]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="37" pageNumber="38">137</figureCitation>
). The hemitergal hooks are strong, with a weak notch in the apical third along the inner edge; the apex is rectangular and rounded (
<figureCitation id="136616E4FFD6CE21FBB9908BDE9CFF79" box="[1069,1234,151,177]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 132−133</figureCitation>
, 
<figureCitation id="136616E4FFD6CE21FB49908BDF45FF79" box="[1245,1291,151,177]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
). The tube of the penis is 1.8 times longer than the sac. The membranous sac is rough, covered with tufts of tiny setae associated with patches, and arranged in concentric lines following the shape of the lobes (
<figureCitation id="136616E4FFD6CE21FBB090C3DE84FF31" box="[1060,1226,223,249]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 139−140</figureCitation>
, B, D). The apical brush is narrow and pointed ventrally (
<figureCitation id="136616E4FFD6CE21FDC3911FD8B0FED5" box="[599,766,259,285]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 138, 140</figureCitation>
, D). The egg has a short, slightly ribbed collar (
<figureCitation id="136616E4FFD6CE21FABF911FDFDEFED5" box="[1323,1424,259,285]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="38" pageNumber="39">Fig. 144</figureCitation>
). The chorion surface is rough and pitted with follicle hexagonal cell impressions, weakly separated from each other; the cell floor is without a central depression; and the number of central pores ranges from one to four (
<figureCitation id="136616E4FFD6CE21FAB89157DFDEFEAD" box="[1324,1424,331,357]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="38" pageNumber="39">Fig. 146</figureCitation>
). Micropylar orifices lack rims (
<figureCitation id="136616E4FFD6CE21FE669173D818FE41" box="[498,598,367,393]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="38" pageNumber="39">Fig. 147</figureCitation>
). The larva is distinguished by pale bands along the lateral margins of the pronotum; the width of the pale bands is wider than the semi-oval dark band; a paired dark S-shaped spot extends from the anterior margin to the middle of the pronotum (
<figureCitation id="136616E4FFD6CE21FC8491ABD9F6FE19" box="[784,952,439,465]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="38" pageNumber="39">Figs. 148−149</figureCitation>
).
</paragraph>
<paragraph id="8BE20A61FFD6CE21FF5391C6DF52FA9D" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
<emphasis id="B929D673FFD6CE21FF5391C6DB3AFE3C" bold="true" box="[199,372,474,500]" pageId="38" pageNumber="39">Adult habitus.</emphasis>
Body length: males 13.0−14.0 mm (n=6), females 
<quantity id="4CA5A784FFD6CE21FC5691C7DE26FE3D" box="[962,1128,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.27" metricValueMin="1.73" pageId="38" pageNumber="39" unit="mm" value="20.0" valueMax="22.7" valueMin="17.3">17.3−22.7 mm</quantity>
(n=7); length of forewing: males 
<quantity id="4CA5A784FFD6CE21FF7491E3DBC6FDD1" box="[224,392,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.325" metricValueMax="1.3699999999999999" metricValueMin="1.28" pageId="38" pageNumber="39" unit="mm" value="13.25" valueMax="13.7" valueMin="12.8">12.8−13.7 mm</quantity>
; females 
<quantity id="4CA5A784FFD6CE21FE6791E2D8D5FDD1" box="[499,667,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.895" metricValueMax="2.05" metricValueMin="1.7399999999999998" pageId="38" pageNumber="39" unit="mm" value="18.95" valueMax="20.5" valueMin="17.4">17.4−20.5 mm</quantity>
; wingspan: males 
<quantity id="4CA5A784FFD6CE21FCFF91E3DE5DFDD1" box="[875,1043,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="2.835" metricValueMax="2.93" metricValueMin="2.7399999999999998" pageId="38" pageNumber="39" unit="mm" value="28.35" valueMax="29.3" valueMin="27.4">27.4−29.3 mm</quantity>
; females 
<quantity id="4CA5A784FFD6CE21FBEA91E2DF68FDD1" box="[1150,1318,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" metricValueMax="4.35" metricValueMin="3.75" pageId="38" pageNumber="39" unit="mm" value="40.5" valueMax="43.5" valueMin="37.5">37.5−43.5 mm</quantity>
. Head and pronotum are dorsally grayish-yellow with dark brown spots; mesonotum and metanotum are dark brown, especially in females; the three small, pale spots on the prescutum and scutum form a triangle; the abdomen is grayish-yellow (
<figureCitation id="136616E4FFD6CE21FF0B9277DB09FD4D" box="[159,327,619,645]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 128−129</figureCitation>
). The wings are yellowish with dark brown veins (
<figureCitation id="136616E4FFD6CE21FC159277DE67FD4D" box="[897,1065,619,645]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 128−129</figureCitation>
). Antennae and palpi are yellowgrey. The M-line on the head is brownish, connected with a triangular brownish spot on the clypeus; the anterior margin and lateral edges of the clypeus are dark brown, clearly contrasting with a pair of transverse small oval pale spots between the clypeal edges and the triangular spot on the clypeus (
<figureCitation id="136616E4FFD6CE21FC6D92CBDEECFD39" box="[1017,1186,727,753]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 128−131</figureCitation>
). Frontal tentorial pits are brownish (
<figureCitation id="136616E4FFD6CE21FEAC92E7DBD5FCDD" box="[312,411,763,789]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Fig. 130</figureCitation>
). The interocellar area has an X-shaped dark brown spot; the upper branches reach the base of the M-line; the lower branches attain the lateral ocelli; and between the lateral ocelli is a triangular yellowish blurred spot limited basally by a brown stripe of the epicranial suture (
<figureCitation id="136616E4FFD6CE21FC42935FDECEFC95" box="[982,1152,835,861]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 130−131</figureCitation>
). A pair of drop-shaped, bright yellow spots are located between the posterior ocelli and the compound eyes; the tentorial pits are brownish (
<figureCitation id="136616E4FFD6CE21FF0B9397DB4DFC6D" box="[159,259,907,933]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Fig. 131</figureCitation>
). The posterior occipital fold is indicated by a thin light stripe, limited above and below by a narrowdarkened band interrupted in the area of the epicranial stem; the epicranial stem darkened (
<figureCitation id="136616E4FFD6CE21FB0893B3DF0DFC01" box="[1180,1347,943,969]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 130−131</figureCitation>
). A pair of transversely oblique dark bands extends from the base of the occiput to the posterior edge of the compound eyes. Pronotum as wide as head with eyes, slightly narrowing posteriorly, posterolateral margins rounded; anterior margin arcuate, darkened; medial stripe dark brown, widened to anterior and posterior margins, wider at the anterior margin than at the posterior one; pronotal posterior margin darkened; lateral pronotal margins bordered by a dark brown stripe, widening posterolaterally; sometimes the lateral edges are bent down and not visible dorsally (
<figureCitation id="136616E4FFD6CE21FAF0947FDAB3FB69" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="38" pageNumber="39">Figs. 117−118</figureCitation>
). Pronotal rugosities are grayish-yellow, darkly contoured, and form an X-shaped pattern (
<figureCitation id="136616E4FFD6CE21FB7E949BDFDEFB69" box="[1258,1424,1159,1185]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 130‒131</figureCitation>
); in the posterior third, dark rugosities in the form of narrow paired transverse short stripes reach the lateral pronotal margins (
<figureCitation id="136616E4FFD6CE21FE9594D3DBE6FB21" box="[257,424,1231,1257]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 130−131</figureCitation>
). The legs and cerci are grayish-brown. Femur with a dark spot basally and a narrow light stripe dorsally; the distal part darkened; tibia darkened at base with a narrow brownish band in the basal ⅕ (
<figureCitation id="136616E4FFD6CE21FAF094EFDAB3FAF9" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="38" pageNumber="39">Figs. 115−117</figureCitation>
); cowl also darkened.Anal gills are very small, noticeable in newly molted specimens. Cerci are bicolored, each segment with a narrow yellowish ring basally and a thick grayish-yellow band apically (
<figureCitation id="136616E4FFD6CE21FB399527DF41FA9D" box="[1197,1295,1339,1365]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Fig. 128</figureCitation>
).
</paragraph>
<paragraph id="8BE20A61FFD6CE21FF539543DBB4F871" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
<emphasis id="B929D673FFD6CE21FF539543DB45FAB1" bold="true" box="[199,267,1375,1401]" pageId="38" pageNumber="39">Male.</emphasis>
Tergum 8 is medially depressed with a central membranous furrow; posterolateral humps are covered with dense and thick setae; along the posterior margin of the furrow, setae are short and sparse (
<figureCitation id="136616E4FFD6CE21FB44959FDF7DFA55" box="[1232,1331,1411,1437]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Fig. 132</figureCitation>
). Tergum 9 has a sclerotized mesal field; the width is slightly greater than or equal to the length (
<figureCitation id="136616E4FFD6CE21FBEF95BBDF6FFA09" box="[1147,1313,1447,1473]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 132−133</figureCitation>
, 
<figureCitation id="136616E4FFD6CE21FABB95BBDF13FA09" box="[1327,1373,1447,1473]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
). The mesal field bears sharp, heavy sclerotized teeth that merge into long and short longitudinal serrated ridges (8−14), located in the middle of the mesal field, and do not reach the posterior margin (
<figureCitation id="136616E4FFD6CE21FBA695F3DEEEF9C1" box="[1074,1184,1519,1545]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 133</figureCitation>
, 
<figureCitation id="136616E4FFD6CE21FB3B95F3DE93F9C1" box="[1199,1245,1519,1545]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
). Tergum 10 has strong hemitergal hooks (cleared) with a weak notch in the distal third of the inner edge laterally (
<figureCitation id="136616E4FFD6CE21FB7D960FDF1BF9E5" box="[1257,1365,1555,1581]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 134</figureCitation>
, 
<figureCitation id="136616E4FFD6CE21FAF5960FDFC1F9E5" box="[1377,1423,1555,1581]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
); in dorsal view, the notch on the hook corresponds to a shallow depression in the side (
<figureCitation id="136616E4FFD6CE21FB07962BDEB7F999" box="[1171,1273,1591,1617]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Fig. 132</figureCitation>
). The apex of the hook is rectangular and rounded (
<figureCitation id="136616E4FFD6CE21FDD79647D8E6F9BD" box="[579,680,1627,1653]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Fig. 134</figureCitation>
); in its natural state, dorsally, the tip is slightly slanted (
<figureCitation id="136616E4FFD6CE21FAB89647DFDEF9BD" box="[1324,1424,1627,1653]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Fig. 132</figureCitation>
). Sensilla basiconica covers the inner side of the hook from top to outer edge; the base of the hook is rounded and covered with long setae (
<figureCitation id="136616E4FFD6CE21FE2096BFD814F975" box="[436,602,1699,1725]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 132−134</figureCitation>
, 
<figureCitation id="136616E4FFD6CE21FDF396BFD8DBF975" box="[615,661,1699,1725]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
). The artificially everted penis has a tube that is 1.8 times the length of the sac; the dorsal tube is covered with densely arranged longitudinal rows of serrate sclerites, sometimes with small spines; the shape of sclerites in different parts of the tube varies from polygonal to oval elongated; the tube is membranous basally with paired patches of rounded spine bases laterally (
<figureCitation id="136616E4FFD6CE21FC409713DE34F8E1" box="[980,1146,1807,1833]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 137−140</figureCitation>
, A, C). The sac is bulbous and membranous at the base, with two lateral lobes and folds that increase its width (
<figureCitation id="136616E4FFD6CE21FBCC972FDF4EF885" box="[1112,1280,1843,1869]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 139−140</figureCitation>
); thin pointed brown spines are present, forming a relatively narrow apical brush that is pointed ventrally (
<figureCitation id="136616E4FFD6CE21FB03974BDF72F8B9" box="[1175,1340,1879,1905]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 139−140</figureCitation>
, B). The membranous sac is rough and covered with tufts of tiny setae arranged in concentric lines following the shape of the lobes (
<figureCitation id="136616E4FFD6CE21FE9E9783DBE7F871" box="[266,425,1951,1977]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs 139−140</figureCitation>
, B, D).
</paragraph>
<caption id="DF225AE9FFD7CE20FF039722D8AFF856" ID-DOI="http://doi.org/10.5281/zenodo.13747810" ID-Zenodo-Dep="13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,264,1854,1878]" targetBox="[151,1436,191,1830]" targetPageId="39" targetType="figure">
<paragraph id="8BE20A61FFD7CE20FF039722D8AFF856" blockId="39.[151,1437,1854,1951]" pageId="39" pageNumber="40">
<emphasis id="B929D673FFD7CE20FF039722DB22F89E" bold="true" box="[151,364,1854,1878]" pageId="39" pageNumber="40">FIGURES 128‒131.</emphasis>
<taxonomicName id="4C5D71E2FFD7CE20FEE59722D84FF89E" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[369,513,1854,1878]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFD7CE20FEE59722D84FF89E" box="[369,513,1854,1878]" italics="true" pageId="39" pageNumber="40">Perla palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFD7CE20FD929722D818F89E" bold="true" box="[518,598,1854,1878]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="A21A6B08FFD7CE20FD929722D818F89E" box="[518,598,1854,1878]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal.
</paragraph>
</caption>
<caption id="DF225AE9FFD8CE2FFF039730D9B6F844" ID-DOI="http://doi.org/10.5281/zenodo.13747812" ID-Zenodo-Dep="13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,264,1836,1860]" targetBox="[151,1436,181,1813]" targetPageId="40" targetType="figure">
<paragraph id="8BE20A61FFD8CE2FFF039730D9B6F844" blockId="40.[151,1436,1836,1933]" pageId="40" pageNumber="41">
<emphasis id="B929D673FFD8CE2FFF039730DB22F88C" bold="true" box="[151,364,1836,1860]" pageId="40" pageNumber="41">FIGURES 132‒136.</emphasis>
<taxonomicName id="4C5D71E2FFD8CE2FFEE59730D84FF88C" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[369,513,1836,1860]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFD8CE2FFEE59730D84FF88C" box="[369,513,1836,1860]" italics="true" pageId="40" pageNumber="41">Perla palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFD8CE2FFD929731D818F88D" bold="true" box="[518,598,1837,1861]" pageId="40" pageNumber="41">
<taxonomicNameLabel id="A21A6B08FFD8CE2FFD929731D818F88D" box="[518,598,1837,1861]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135 
<emphasis id="B929D673FFD8CE2FFD849769D853F845" bold="true" box="[528,541,1909,1933]" pageId="40" pageNumber="41">‒</emphasis>
136. Female abdominal tip, variations in color.
</paragraph>
</caption>
<caption id="DF225AE9FFD9CE2EFF03977BDF76F823" ID-DOI="http://doi.org/10.5281/zenodo.13747814" ID-Zenodo-Dep="13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,264,1895,1919]" targetBox="[158,1430,181,1871]" targetPageId="41" targetType="figure">
<paragraph id="8BE20A61FFD9CE2EFF03977BDF76F823" blockId="41.[151,1437,1895,2027]" pageId="41" pageNumber="42">
<emphasis id="B929D673FFD9CE2EFF03977BDB3FF8B7" bold="true" box="[151,369,1895,1919]" pageId="41" pageNumber="42">FIGURES 137‒140.</emphasis>
<taxonomicName id="4C5D71E2FFD9CE2EFEEE977BD840F8B7" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[378,526,1895,1919]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFD9CE2EFEEE977BD840F8B7" box="[378,526,1895,1919]" italics="true" pageId="41" pageNumber="42">Perla palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFD9CE2EFD8C977BD822F8B7" bold="true" box="[536,620,1895,1919]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A21A6B08FFD9CE2EFD8C977BD822F8B7" box="[536,620,1895,1919]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines.
</paragraph>
</caption>
<caption id="DF225AE9FFDACE2DFF0396D2D842F852" ID-DOI="http://doi.org/10.5281/zenodo.13747816" ID-Zenodo-Dep="13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,264,1742,1766]" targetBox="[170,1417,181,1718]" targetPageId="42" targetType="figure">
<paragraph id="8BE20A61FFDACE2DFF0396D2D842F852" blockId="42.[151,1437,1742,1946]" pageId="42" pageNumber="43">
<emphasis id="B929D673FFDACE2DFF0396D2DB22F92E" bold="true" box="[151,364,1742,1766]" pageId="42" pageNumber="43">FIGURES 141‒147.</emphasis>
<taxonomicName id="4C5D71E2FFDACE2DFEE596D2D84FF92E" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[369,513,1742,1766]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDACE2DFEE596D2D84FF92E" box="[369,513,1742,1766]" italics="true" pageId="42" pageNumber="43">Perla palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDACE2DFD9296D2D818F92E" bold="true" box="[518,598,1742,1766]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="A21A6B08FFDACE2DFD9296D2D818F92E" box="[518,598,1742,1766]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141. Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142. Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims.
</paragraph>
</caption>
<caption id="DF225AE9FFDBCE2CFF0395CDDE11F9C5" ID-DOI="http://doi.org/10.5281/zenodo.13747818" ID-Zenodo-Dep="13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,264,1489,1513]" targetBox="[151,1436,181,1465]" targetPageId="43" targetType="figure">
<paragraph id="8BE20A61FFDBCE2CFF0395CDDE11F9C5" blockId="43.[151,1436,1489,1549]" pageId="43" pageNumber="44">
<emphasis id="B929D673FFDBCE2CFF0395CDDB22FA21" bold="true" box="[151,364,1489,1513]" pageId="43" pageNumber="44">FIGURES 148‒149.</emphasis>
<taxonomicName id="4C5D71E2FFDBCE2CFEE595CDD84FFA21" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[369,513,1489,1513]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDBCE2CFEE595CDD84FFA21" box="[369,513,1489,1513]" italics="true" pageId="43" pageNumber="44">Perla palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDBCE2CFD9295CDD818FA21" bold="true" box="[518,598,1489,1513]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="A21A6B08FFDBCE2CFD9295CDD818FA21" box="[518,598,1489,1513]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern.
</paragraph>
</caption>
<paragraph id="8BE20A61FFDBCE2CFF539643DBAFF90A" blockId="43.[151,1437,1631,2017]" pageId="43" pageNumber="44">
<emphasis id="B929D673FFDBCE2CFF539643DB6BF9B1" bold="true" box="[199,293,1631,1657]" pageId="43" pageNumber="44">Female.</emphasis>
Sternum 8 has a subgenital plate, occupying almost half of the sternum width, with two small triangular lobes posteromedially. The lobes projecting slightly beyond the anterior margin of sternum 9 are covered with rufous setae (
<figureCitation id="136616E4FFDBCE2CFEB996BBDB9AF90A" box="[301,468,1703,1730]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="43" pageNumber="44">Figs. 135−136</figureCitation>
).
</paragraph>
<paragraph id="8BE20A61FFDBCE2CFF5396D7DA9CF829" blockId="43.[151,1437,1631,2017]" pageId="43" pageNumber="44">
<emphasis id="B929D673FFDBCE2CFF5396D7DAB3F92D" bold="true" box="[199,253,1739,1765]" pageId="43" pageNumber="44">Egg.</emphasis>
Elongate and oval, with mean dimensions of 435 × 272 μm (n = 6), with a short collar with the same diameter at the rim and the base; the rim is weakly ribbed (
<figureCitation id="136616E4FFDBCE2CFC8D96F3D9F1F8C1" box="[793,959,1775,1801]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="43" pageNumber="44">Figs. 141−144</figureCitation>
). The neck is not constricted.A mushroom-shaped anchor plate is present on some eggs and covered with globular bodies concentrated in small groups (
<figureCitation id="136616E4FFDBCE2CFAF0970FDAB0F89A" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="43" pageNumber="44">Figs. 142, 145</figureCitation>
). The chorion surface is rough and pitted; the chorion is covered throughout with follicle hexagonal cell impressions; hexagonal cells with differences in cell widths; cells separated from adjacent ones by poorly expressed boundaries; the cell floor without a central depression with one or four pores (
<figureCitation id="136616E4FFDBCE2CFC6A9763DEEDF851" box="[1022,1187,1919,1945]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="43" pageNumber="44">Figs. 144, 146</figureCitation>
). A ring of micropyles is located ca. 1/3 of the egg length from the anterior pole, the micropylar orifices are without rims (
<figureCitation id="136616E4FFDBCE2CFB6697BFDA8BF829" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="43" pageNumber="44">Figs. 141−142, 147</figureCitation>
).
</paragraph>
<caption id="DF225AE9FFDCCE2BFF0396BBDF54F8E3" ID-DOI="http://doi.org/10.5281/zenodo.13747820" ID-Zenodo-Dep="13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,264,1703,1727]" targetBox="[151,1435,184,1679]" targetPageId="44" targetType="figure">
<paragraph id="8BE20A61FFDCCE2BFF0396BBDF54F8E3" blockId="44.[151,1436,1703,1835]" pageId="44" pageNumber="45">
<emphasis id="B929D673FFDCCE2BFF0396BBDB22F977" bold="true" box="[151,364,1703,1727]" pageId="44" pageNumber="45">FIGURES 150‒156.</emphasis>
<taxonomicName id="4C5D71E2FFDCCE2BFEE596BBD84FF977" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[369,513,1703,1727]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDCCE2BFEE596BBD84FF977" box="[369,513,1703,1727]" italics="true" pageId="44" pageNumber="45">Perla palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDCCE2BFD9296BBD818F977" bold="true" box="[518,598,1703,1727]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="A21A6B08FFDCCE2BFD9296BBD818F977" box="[518,598,1703,1727]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments.
</paragraph>
</caption>
<paragraph id="8BE20A61FFDCCE2AFF539761D9B0FD05" blockId="44.[151,1437,1917,2016]" lastBlockId="45.[151,1437,151,2013]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="B929D673FFDCCE2BFF539761DB59F85F" bold="true" box="[199,279,1917,1943]" pageId="44" pageNumber="45">Larva.</emphasis>
Mature males of medium size, length 
<quantity id="4CA5A784FFDCCE2BFD479761D934F850" box="[723,890,1917,1944]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" metricValueMax="1.24" metricValueMin="1.22" pageId="44" pageNumber="45" unit="mm" value="12.3" valueMax="12.4" valueMin="12.2">12.2−12.4 mm</quantity>
(n=3), antennae 
<quantity id="4CA5A784FFDCCE2BFBA99761DED9F850" box="[1085,1175,1917,1944]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="44" pageNumber="45" unit="mm" value="9.5">9.5 mm</quantity>
, and cerci greater than 
<quantity id="4CA5A784FFDCCE2BFF0397BDDAA3F874" box="[151,237,1953,1980]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="44" pageNumber="45" unit="mm" value="8.8">8.8 mm</quantity>
contain more than 36 segments. The color is generally pale, yellow-brown in dorsal view (
<figureCitation id="136616E4FFDCCE2BFB7D97BDDFDEF873" box="[1257,1424,1953,1979]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="44" pageNumber="45">Figs. 148−149</figureCitation>
), with a distinct dark pattern on the head, pronotum, mesonotum, and metanotum, but with a less distinct pattern on the abdomen. The ventral body side is yellow-brownish. Body covered with short dark clothing hairs, with a dorsomesal band of erect silky white setae more pronounced on the last abdominal terga (
<figureCitation id="136616E4FFDDCE2AFBED90A7DF6EFF1D" box="[1145,1312,187,213]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
). Antennae and cerci are yellow-brown. Clypeus on head basally with paired dark and wide V-shaped spots, outlining branches of yellow M-line from above. Head with a large transverse dark spot, vaguely resembling a pilot's badge with truncated “wings” not extending beyond epicranial arms; the spot limits the pale M-line from below extending from the anterior ocellus to the posterior ocelli; a small triangular spot between the ocelli darkened; epicranial stem and occiput edge also darkened (
<figureCitation id="136616E4FFDDCE2AFE439173D831FE41" box="[471,639,367,393]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
). The occipital ridge is distinct and complete; short spines are arranged in the medial part of the fold; along the postocular crest, spines are noticeably longer and pointed. Mandibles (
<figureCitation id="136616E4FFDDCE2AFAFB918FDA8CFE19" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 153</figureCitation>
) are typical for 
<taxonomicName id="4C5D71E2FFDDCE2AFEEE91ABDBF6FE19" box="[378,440,439,465]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B929D673FFDDCE2AFEEE91ABDBF6FE19" box="[378,440,439,465]" italics="true" pageId="45" pageNumber="46">Perla</emphasis>
</taxonomicName>
: heavily sclerotized, with a rounded outer margin; left mandible with five pointed teeth and two medial rows of setae on the molar area; the marginal row of setae extends from the base of the 4th tooth to the basal third of the mandible; right mandible with six pointed teeth; single medial setal row extends from the base of the first tooth to the base of the mandible; the configuration of the marginal setal row is the same as that of the left mandible. Lacinia bidentate with a large, heavily sclerotized, and strongly curved apical tooth (
<figureCitation id="136616E4FFDDCE2AFB6E925BDF13FDA9" box="[1274,1373,582,609]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 152</figureCitation>
). The subapical tooth is shorter; the length does not exceed 2/3 of the apical tooth length. The marginal fringe of setae along the inner margin is complete; a tuft of long and dense setae of marginal fringe is clearly visible below the subapical tooth. Galea length reaches 1/2 of subapical tooth length (
<figureCitation id="136616E4FFDDCE2AFC1B92AFD9BCFD05" box="[911,1010,690,717]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 152</figureCitation>
).
</paragraph>
<paragraph id="8BE20A61FFDDCE2AFF5392CBDBF9F9E5" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
Pronotum transverse, not wider than the head, with obtusely rounded anterior corners; lateral margins and posterior angles are evenly rounded (
<figureCitation id="136616E4FFDDCE2AFDBA92E7D89AFCDD" box="[558,724,763,789]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
). The pronotal fringe is complete with mixed short and relatively long setae. The pronotum has a thin brown stripe along the edge; paired semi-oval pale bands outline a full (or incomplete, open anterolaterally) oval brown band on the lateral pronotal fields (
<figureCitation id="136616E4FFDDCE2AFBA0935FDE93FC95" box="[1076,1245,835,861]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
). An oval brown band is narrower than semi-oval pale bands, especially in the anterior 1/3. The central pronotal field is yellow with a brown median band narrowed anteriorly and markedly widened posteriorly; a paired vague S-shaped dark spot extends from the anterior margin to the middle of the pronotum (
<figureCitation id="136616E4FFDDCE2AFCFA93B3DE58FC01" box="[878,1046,943,969]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
). Mesonotum and metanotum with an identical Ψ-shaped dark pattern medially, posterior margin darkly margined; two wide dark bands limiting the wing pads; a narrow dark stripe runs along the lateral edge of the wing pads. Legs are yellow and covered with dark closing hairs (
<figureCitation id="136616E4FFDDCE2AFEA29407DB91FBFD" box="[310,479,1051,1077]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
). The femur and tibia bear stout, short, acute bristles along the inner and outer edges and a very dense fringe of long, silky hairs along the outer margins (
<figureCitation id="136616E4FFDDCE2AFC379423DE49FB91" box="[931,1031,1086,1113]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 150</figureCitation>
). The femur is covered with sparse, short, and irregular red bristles; in the distal half closer to the outer edge, with a darkened small and almost oval spot, the distal edge of the femur is also darkened (
<figureCitation id="136616E4FFDDCE2AFD4E949BD973FB69" box="[730,829,1158,1185]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 150</figureCitation>
). Hind femur ca. 3.3−3.6 times as long as wide. Tibia has a diffuse, narrow brownish band basally. The pattern on the abdomen is indistinct; the dark closing hairs are well-marked; the terga are mainly yellowish with a dark median and two lateral longitudinal bands; the lateral dark bands on terga 4−10 are triangular, extending to the anterior margin (
<figureCitation id="136616E4FFDDCE2AFC3C94EFDE1EFAC5" box="[936,1104,1267,1293]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
). The posterior terga margins bear acute bristles that are mostly short, with one or two relatively long setae (
<figureCitation id="136616E4FFDDCE2AFB91950BDE29FAF9" box="[1029,1127,1302,1329]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 151</figureCitation>
). Anal gills are small (
<figureCitation id="136616E4FFDDCE2AFAF0950BDAB1FA9D" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
). Cerci are yellow, and no less than 36 segmented with striking rings of dark spines (
<figureCitation id="136616E4FFDDCE2AFB259527DF16FA9D" box="[1201,1368,1339,1365]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
). Fine pilosity between dark spines is tightly pressed to the segment surface (
<figureCitation id="136616E4FFDDCE2AFC099543DE0DFAB1" box="[925,1091,1374,1401]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Figs. 154−156</figureCitation>
); short fine hairs have the same length as the length of the basal cercal segment; towards the middle of the cerci, fine pilosity length decreases; and on the apical cercal segments, the length of the fine pilosity is slightly&gt;1/3 of the segment length (
<figureCitation id="136616E4FFDDCE2AFB7F95BBDF00FA09" box="[1259,1358,1446,1473]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 156</figureCitation>
). Long silky fine colorless hairs are included in the circlet in tufts (no more than 
<quantity id="4CA5A784FFDDCE2AFC4595D7D9B3FA2D" box="[977,1021,1483,1509]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="45" pageNumber="46" unit="in" value="8.0">8 in</quantity>
each) and form a dorsal longitudinal fringe, especially on the basal segments (
<figureCitation id="136616E4FFDDCE2AFDF795F3D88BF9C1" box="[611,709,1518,1545]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 154</figureCitation>
); the length and density of long silky hairs also decrease towards the apex (
<figureCitation id="136616E4FFDDCE2AFE91960FDBE5F9E5" box="[261,427,1554,1581]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Figs. 155‒156</figureCitation>
).
</paragraph>
<paragraph id="8BE20A61FFDDCE2AFF53962ADEE7F999" blockId="45.[151,1437,151,2013]" box="[199,1193,1590,1617]" pageId="45" pageNumber="46">
<emphasis id="B929D673FFDDCE2AFF53962ADBC7F998" bold="true" box="[199,393,1590,1616]" pageId="45" pageNumber="46">DNA barcoding.</emphasis>
GenBank accession numbers are PP216454‒PP216456 and PP236723.
</paragraph>
</subSubSection>
<subSubSection id="C34759EAFFDDCE2AFF539646DFD3F8CD" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8BE20A61FFDDCE2AFF539646DFD3F8CD" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
<emphasis id="B929D673FFDDCE2AFF539646DBFCF9BC" bold="true" box="[199,434,1626,1652]" pageId="45" pageNumber="46">Material examined.</emphasis>
<materialsCitation id="3B35003CFFDDCE2AFE2E9647D82EF975" collectingDate="2021-08-06" collectionCode="FSC, EATB, FEB" collectorName="D. Palatov" country="Russia" county="Gelendzhik District" latitude="44.569305" location="Pshada River Basin" longLatPrecision="1" longitude="38.42889" municipality="Papay River" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Krasnodar" typeStatus="holotype">
<typeStatus id="54E6B4C3FFDDCE2AFE2E9647D86CF9BD" box="[442,546,1627,1653]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED4C92A4FFDDCE2AFDE49647D8EDF9BD" box="[624,675,1627,1653]" country="Canada" httpUri="http://grbio.org/cool/q309-g0ck" name="Fredericton Stock Culture Collection" pageId="45" pageNumber="46">FSC</collectionCode>
<collectionCode id="ED4C92A4FFDDCE2AFD389647D963F9BD" box="[684,813,1627,1653]" pageId="45" pageNumber="46">EATB FEB</collectionCode>
RAS). Caucasus. 
<collectingCountry id="F34A4AF1FFDDCE2AFC699647DE05F9BD" box="[1021,1099,1627,1653]" name="Russia" pageId="45" pageNumber="46">Russia</collectingCountry>
, 
<collectingRegion id="4999C483FFDDCE2AFBC39647DF41F9BD" box="[1111,1295,1627,1653]" country="Russia" name="Krasnodar" pageId="45" pageNumber="46">Krasnodar Kray</collectingRegion>
: 
<collectingCounty id="628372EDFFDDCE2AFA8F9647DAA0F951" pageId="45" pageNumber="46">Gelendzhik District</collectingCounty>
, 
<collectingMunicipality id="6B86901BFFDDCE2AFF6D9663DBCAF951" box="[249,388,1663,1689]" pageId="45" pageNumber="46">Papay River</collectingMunicipality>
, 
<locationDeviation id="9A501343FFDDCE2AFE1B9662D8F3F951" box="[399,701,1662,1689]" location="Novosadovy" pageId="45" pageNumber="46">
<quantity id="4CA5A784FFDDCE2AFE1B9662DB93F951" box="[399,477,1662,1689]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="45" pageNumber="46" unit="km" value="2.5">2.5 km</quantity>
below Novosadovy
</locationDeviation>
farm, 
<location id="8E825CBAFFDDCE2AFC919663D9AAF951" LSID="urn:lsid:plazi:treatment:03F4BB77FFD5CE29FF03972EDE7BFDA9:8E825CBAFFDDCE2AFC919663D9AAF951" box="[773,996,1663,1689]" country="Russia" county="Gelendzhik District" latitude="44.569305" longLatPrecision="1" longitude="38.42889" municipality="Papay River" name="Pshada River Basin" pageId="45" pageNumber="46" stateProvince="Krasnodar">
<location id="8E825CBAFFDDCE2AFC919663D9D4F951" LSID="urn:lsid:plazi:treatment:03F4BB77FFD5CE29FF03972EDE7BFDA9:8E825CBAFFDDCE2AFC919663D9D4F951" box="[773,922,1663,1689]" country="Russia" county="Gelendzhik District" latitude="44.569305" longLatPrecision="1" longitude="38.42889" municipality="Papay River" name="Pshada River" pageId="45" pageNumber="46" stateProvince="Krasnodar">
Pshada 
<location id="8E825CBAFFDDCE2AFCC89663D9D4F951" LSID="urn:lsid:plazi:treatment:03F4BB77FFD5CE29FF03972EDE7BFDA9:8E825CBAFFDDCE2AFCC89663D9D4F951" box="[860,922,1663,1689]" country="Russia" county="Gelendzhik District" latitude="44.569305" longLatPrecision="1" longitude="38.42889" municipality="Papay River" name="River Basin" pageId="45" pageNumber="46" stateProvince="Krasnodar">River</location>
</location>
Basin
</location>
, collected by a 
<collectingMethod id="521C7276FFDDCE2AFB049663DEB6F951" box="[1168,1272,1663,1689]" pageId="45" pageNumber="46">light trap</collectingMethod>
, 
<geoCoordinate id="EE696CA6FFDDCE2AFA979662DFD6F951" box="[1283,1432,1662,1689]" degrees="44.569305" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="1" value="44.569305">44.569305 N</geoCoordinate>
, 
<geoCoordinate id="EE696CA6FFDDCE2AFF0396BFDB57F975" box="[151,281,1699,1725]" degrees="38.42889" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="1" value="38.42889">38.42889 E</geoCoordinate>
, 
<date id="FFE32CA1FFDDCE2AFEB096BFDBE8F975" box="[292,422,1699,1725]" pageId="45" pageNumber="46" value="2021-08-06">
<collectingDate id="EFA7D549FFDDCE2AFEB096BFDBE8F975" box="[292,422,1699,1725]" pageId="45" pageNumber="46" value="2021-08-06">08.06.2021</collectingDate>
</date>
, coll. 
<collectorName id="26A86FB7FFDDCE2AFE7396BFD82EF975" box="[487,608,1699,1725]" pageId="45" pageNumber="46">D. Palatov</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B35003CFFDDCE2AFDFE96BFDFD6F975" box="[618,1432,1698,1725]" collectionCode="FSC, EATB, FEB" country="Russia" pageId="45" pageNumber="46" specimenCount="10" specimenCount-female="5" specimenCount-larva="2" specimenCount-male="3" stateProvince="Krasnodar" typeStatus="paratype">
<typeStatus id="54E6B4C3FFDDCE2AFDFE96BFD895F975" box="[618,731,1699,1725]" pageId="45" pageNumber="46" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D5BC1E8FFDDCE2AFD7096BFD944F975" box="[740,778,1699,1725]" count="3" pageId="45" pageNumber="46" type="male">3♂</specimenCount>
, 
<specimenCount id="9D5BC1E8FFDDCE2AFC8096BED977F975" box="[788,825,1698,1725]" count="5" pageId="45" pageNumber="46" type="female">5♀</specimenCount>
, 
<specimenCount id="9D5BC1E8FFDDCE2AFCD196BFD9D3F975" box="[837,925,1699,1725]" count="2" pageId="45" pageNumber="46" type="larva">2 larvae</specimenCount>
(
<collectionCode id="ED4C92A4FFDDCE2AFC3F96BFD990F975" box="[939,990,1699,1725]" country="Canada" httpUri="http://grbio.org/cool/q309-g0ck" name="Fredericton Stock Culture Collection" pageId="45" pageNumber="46">FSC</collectionCode>
<collectionCode id="ED4C92A4FFDDCE2AFC7096BFDE2DF975" box="[996,1123,1699,1725]" pageId="45" pageNumber="46">EATB FEB</collectionCode>
RAS), same place and data
</materialsCitation>
; 
<materialsCitation id="3B35003CFFDDCE2AFF0396DBDFD3F8CD" collectingDate="2022-06-22" collectorName="D. Palatov." country="Russia" county="Tenginka village" elevation="22" location="Tuapse District" longitude="38.78621" municipality="Black Sea" pageId="45" pageNumber="46" specimenCount="8" specimenCount-exuvia="3" specimenCount-female="2" specimenCount-larva="1" specimenCount-male="2" stateProvince="Krasnodar" typeStatus="paratype">
<specimenCount id="9D5BC1E8FFDDCE2AFF0396DBDAF3F929" box="[151,189,1735,1761]" count="2" pageId="45" pageNumber="46" type="male">2♂</specimenCount>
, 
<specimenCount id="9D5BC1E8FFDDCE2AFF5396DBDAA2F929" box="[199,236,1735,1761]" count="2" pageId="45" pageNumber="46" type="female">2♀</specimenCount>
, 
<specimenCount id="9D5BC1E8FFDDCE2AFF6C96DBDB09F929" box="[248,327,1735,1761]" count="1" pageId="45" pageNumber="46" type="larva">1 larva</specimenCount>
, 
<specimenCount id="9D5BC1E8FFDDCE2AFEC596DBDBFCF929" box="[337,434,1735,1761]" count="3" pageId="45" pageNumber="46" type="exuvia">3 exuvia</specimenCount>
, 
<location id="8E825CBAFFDDCE2AFE2996DBD824F929" LSID="urn:lsid:plazi:treatment:03F4BB77FFD5CE29FF03972EDE7BFDA9:8E825CBAFFDDCE2AFE2996DBD824F929" box="[445,618,1735,1761]" country="Russia" county="Tenginka village" longitude="38.78621" municipality="Black Sea" name="Tuapse District" pageId="45" pageNumber="46" stateProvince="Krasnodar">Tuapse District</location>
, 
<location id="8E825CBAFFDDCE2AFDE096DBD97BF929" LSID="urn:lsid:plazi:treatment:03F4BB77FFD5CE29FF03972EDE7BFDA9:8E825CBAFFDDCE2AFDE096DBD97BF929" box="[628,821,1735,1761]" country="Russia" county="Tenginka village" longitude="38.78621" municipality="Black Sea" name="Shapsukho River" pageId="45" pageNumber="46" stateProvince="Krasnodar">Shapsukho River</location>
, downstream, 
<quantity id="4CA5A784FFDDCE2AFC4296DBDE40F929" box="[982,1038,1735,1761]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="45" pageNumber="46" unit="km" value="3.0">3 km</quantity>
from the confluence with the 
<collectingMunicipality id="6B86901BFFDDCE2AFACF96DBDA8CF8CD" pageId="45" pageNumber="46">Black Sea</collectingMunicipality>
, near 
<collectingCounty id="628372EDFFDDCE2AFE9096F7DB8CF8CD" box="[260,450,1771,1797]" pageId="45" pageNumber="46">Tenginka village</collectingCounty>
, altitude 
<elevation id="0070ED52FFDDCE2AFDBC96F7D95BF8CD" box="[552,789,1771,1797]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="45" pageNumber="46" unit="m" value="22.0">
<quantity id="4CA5A784FFDDCE2AFDBC96F7D82FF8CD" box="[552,609,1771,1797]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="45" pageNumber="46" unit="m" value="22.0">22 m</quantity>
above sea level
</elevation>
, 44.337647, N 
<geoCoordinate id="EE696CA6FFDDCE2AFC5596F7DE1FF8CD" box="[961,1105,1771,1797]" degrees="38.786211" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="1" value="38.78621">38.786211 E</geoCoordinate>
, 
<date id="FFE32CA1FFDDCE2AFBC996F7DE90F8CD" box="[1117,1246,1771,1797]" pageId="45" pageNumber="46" value="2022-06-22">
<collectingDate id="EFA7D549FFDDCE2AFBC996F7DE90F8CD" box="[1117,1246,1771,1797]" pageId="45" pageNumber="46" value="2022-06-22">22.06.2022</collectingDate>
</date>
, coll. 
<collectorName id="26A86FB7FFDDCE2AFAB596F7DFD3F8CD" box="[1313,1437,1771,1797]" pageId="45" pageNumber="46">D. Palatov.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34759EAFFDDCE2AFF539712DF33F8B9" pageId="45" pageNumber="46" type="distribution">
<paragraph id="8BE20A61FFDDCE2AFF539712DF33F8B9" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
<emphasis id="B929D673FFDDCE2AFF539712DB2FF8E0" bold="true" box="[199,353,1806,1832]" pageId="45" pageNumber="46">Distribution.</emphasis>
North Caucasus. North-eastern coast of the Black Sea. 
<collectingCountry id="F34A4AF1FFDDCE2AFC469713DE6EF8E1" box="[978,1056,1807,1833]" name="Russia" pageId="45" pageNumber="46">Russia</collectingCountry>
. 
<collectingRegion id="4999C483FFDDCE2AFBBF9713DEAFF8E1" box="[1067,1249,1807,1833]" country="Russia" name="Krasnodar" pageId="45" pageNumber="46">Krasnodar Kray</collectingRegion>
. Gelendzhik and Tuapse Districts. The species was found in small foothill rivers along the Black Sea coast (
<figureCitation id="136616E4FFDDCE2AFB1F972FDF7FF885" box="[1163,1329,1842,1869]" captionStart="FIGURES 157‒158" captionStartId="47.[151,264,689,713]" captionTargetBox="[151,1436,181,665]" captionTargetId="figure-18@47.[151,1436,181,665]" captionTargetPageId="47" captionText="FIGURES 157‒158. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray. Type habitat. Gelendzhik District, Papay River, Pshada River Basin." figureDoi="http://doi.org/10.5281/zenodo.13747822" httpUri="https://zenodo.org/record/13747822/files/figure.png" pageId="45" pageNumber="46">Figs. 157−158</figureCitation>
), flowing through low-mountain forest valleys 
<quantity id="4CA5A784FFDDCE2AFDA2974BD8D5F8B9" box="[566,667,1879,1905]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="45" pageNumber="46" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">10−12 m</quantity>
wide with a pebble and rocky bottom and a current up to 
<quantity id="4CA5A784FFDDCE2AFAB7974BDF28F8B9" box="[1315,1382,1879,1905]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="45" pageNumber="46" unit="m" value="0.7">0.7 m</quantity>
/s.
</paragraph>
</subSubSection>
<subSubSection id="C34759EAFFDDCE2AFF539767DB69F815" pageId="45" pageNumber="46" type="etymology">
<paragraph id="8BE20A61FFDDCE2AFF539767DB69F815" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
<emphasis id="B929D673FFDDCE2AFF539767DB00F85D" bold="true" box="[199,334,1915,1941]" pageId="45" pageNumber="46">Etymology.</emphasis>
The species is named in honor of Dr. Dmitry M. Palatov, who collected 
<typeStatus id="54E6B4C3FFDDCE2AFB069767DE8CF85D" box="[1170,1218,1915,1941]" pageId="45" pageNumber="46">type</typeStatus>
material for a new species in the Caucasus as well as other very interesting stonefly material in the Caucasus, Central Asia, and the Middle East.
</paragraph>
</subSubSection>
<subSubSection id="C34759EAFFDECE29FF53908BDE7BFDA9" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8BE20A61FFDECE29FF53908BDE7BFDA9" blockId="46.[151,1437,151,609]" pageId="46" pageNumber="47">
<emphasis id="B929D673FFDECE29FF53908BDB75FF79" bold="true" box="[199,315,151,177]" pageId="46" pageNumber="47">Remarks.</emphasis>
<taxonomicName id="4C5D71E2FFDECE29FED6908BDBACFF79" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[322,482,151,177]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDECE29FED6908BDBACFF79" box="[322,482,151,177]" italics="true" pageId="46" pageNumber="47">Perla palatovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A6B08FFDECE29FE7D908BD80DFF79" box="[489,579,151,177]" pageId="46" pageNumber="47" rank="species">
<emphasis id="B929D673FFDECE29FE7D908BD873FF79" bold="true" box="[489,573,151,177]" pageId="46" pageNumber="47">sp. nov</emphasis>
.
</taxonomicNameLabel>
bears some similarity to the 
<taxonomicName id="4C5D71E2FFDECE29FC10908BD9A0FF79" authorityName="Guerin-Meneville" authorityYear="1843" box="[900,1006,151,177]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B929D673FFDECE29FC10908BD9A0FF79" box="[900,1006,151,177]" italics="true" pageId="46" pageNumber="47">P. pallida</emphasis>
</taxonomicName>
species complex. The most noticeable resemblance between the eggs of 
<taxonomicName id="4C5D71E2FFDECE29FD9890A7D8CCFF1D" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[524,642,187,213]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDECE29FD9890A7D8CCFF1D" box="[524,642,187,213]" italics="true" pageId="46" pageNumber="47">P. palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDECE29FD1C90A7D8AFFF1D" bold="true" box="[648,737,187,213]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A21A6B08FFDECE29FD1C90A7D8AFFF1D" box="[648,737,187,213]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was observed with 
<typeStatus id="54E6B4C3FFDECE29FC2F90A7D9BDFF1D" box="[955,1011,187,213]" pageId="46" pageNumber="47">Type</typeStatus>
2 and 
<typeStatus id="54E6B4C3FFDECE29FBAF90A7DE3DFF1D" box="[1083,1139,187,213]" pageId="46" pageNumber="47">Type</typeStatus>
<quantity id="4CA5A784FFDECE29FBED90A7DEEDFF1D" box="[1145,1187,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="46" pageNumber="47" unit="in" value="3.0">3 in</quantity>
the shape of the collar (
<bibRefCitation id="EFCC7790FFDECE29FF0B90C3DBC8FF31" author="Sivec, I. &amp; Stark, B. P." box="[159,390,223,249]" pageId="46" pageNumber="47" pagination="1 - 33" refId="ref26075" refString="Sivec, I. &amp; Stark, B. P. (2002) The species of Perla (Plecoptera: Perlidae): evidence from egg morphology. Scopolia, 49, 1 - 33." type="journal article" year="2002">Sivec &amp; Stark 2002</bibRefCitation>
). The collar is short, with the same diameter at the rim and base; the neck is not constricted but slightly ribbed. The main differences between 
<taxonomicName id="4C5D71E2FFDECE29FD48911FD918FED5" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[732,854,259,285]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDECE29FD48911FD918FED5" box="[732,854,259,285]" italics="true" pageId="46" pageNumber="47">P. palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDECE29FCCB911FD9F5FED5" bold="true" box="[863,955,259,285]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A21A6B08FFDECE29FCCB911FD9F5FED5" box="[863,955,259,285]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and any 
<typeStatus id="54E6B4C3FFDECE29FBBC911FDE16FED5" box="[1064,1112,259,285]" pageId="46" pageNumber="47">type</typeStatus>
of egg consist of punctured hexagonal chorion cells, the boundaries between which are poorly expressed. The 
<taxonomicName id="4C5D71E2FFDECE29FBD0913BDEF3FE89" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[1092,1213,295,321]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDECE29FBD0913BDEF3FE89" box="[1092,1213,295,321]" italics="true" pageId="46" pageNumber="47">P. palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDECE29FB51913BDF6FFE89" bold="true" box="[1221,1313,295,321]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A21A6B08FFDECE29FB51913BDF6FFE89" box="[1221,1313,295,321]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
hexagonal floors lack depressions and have four pores; the pores and punctuation of the hexagonal floors are uniform. The larvae of 
<taxonomicName id="4C5D71E2FFDECE29FE929173DBCEFE41" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[262,384,367,393]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDECE29FE929173DBCEFE41" box="[262,384,367,393]" italics="true" pageId="46" pageNumber="47">P. palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDECE29FE1D9173DBABFE41" bold="true" box="[393,485,367,393]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A21A6B08FFDECE29FE1D9173DBABFE41" box="[393,485,367,393]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are very similar to the patterning of 
<taxonomicName id="4C5D71E2FFDECE29FC079173DE6EFE40" box="[915,1056,367,392]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="B929D673FFDECE29FC079173DE6EFE40" box="[915,1056,367,392]" italics="true" pageId="46" pageNumber="47">P. caucasica</emphasis>
</taxonomicName>
larvae described by 
<bibRefCitation id="EFCC7790FFDECE29FA869173DB30FE65" author="Zhiltzova, L. A. &amp; Cherchesova, S. K." pageId="46" pageNumber="47" pagination="321 - 326" refId="ref26717" refString="Zhiltzova, L. A. &amp; Cherchesova, S. K. (2003) Description of the larva of genus Perla Geoff. (Plecoptera, Perlidae) from the Caucasus. Entomologicheskoe obozrenie, 82 (2), 321 - 326. [in Russian]" type="journal article" year="2003">Zhiltzova &amp; Cherchesova (2003)</bibRefCitation>
. There are differences in the dark closing hairs, which are denser and more pronounced in 
<taxonomicName id="4C5D71E2FFDECE29FA1C918FDB49FE18" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="B929D673FFDECE29FA1C918FDB49FE18" italics="true" pageId="46" pageNumber="47">P. caucasica</emphasis>
</taxonomicName>
; in 
<taxonomicName id="4C5D71E2FFDECE29FEA491ABDBE8FE19" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[304,422,439,465]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDECE29FEA491ABDBE8FE19" box="[304,422,439,465]" italics="true" pageId="46" pageNumber="47">P. palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDECE29FE3891ABD84AFE19" bold="true" box="[428,516,439,465]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A21A6B08FFDECE29FE3891ABD84AFE19" box="[428,516,439,465]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the pronotal and especially abdominal patterns are lighter and less distinct. For the pronotum of 
<taxonomicName id="4C5D71E2FFDECE29FEBC91C7DBD0FE3D" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[296,414,475,501]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDECE29FEBC91C7DBD0FE3D" box="[296,414,475,501]" italics="true" pageId="46" pageNumber="47">P. palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDECE29FE3091C7DBB3FE3D" bold="true" box="[420,509,475,501]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A21A6B08FFDECE29FE3091C7DBB3FE3D" box="[420,509,475,501]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
larvae, the width of the oval dark bands is narrower than the width of the semi-oval pale bands, and vice versa in 
<taxonomicName id="4C5D71E2FFDECE29FE7091E3D83FFDD0" box="[484,625,511,536]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="B929D673FFDECE29FE7091E3D83FFDD0" box="[484,625,511,536]" italics="true" pageId="46" pageNumber="47">P. caucasica</emphasis>
</taxonomicName>
, where a paired S-shaped dark spot extends from the anterior margin to the middle of the pronotum. On 
<taxonomicName id="4C5D71E2FFDECE29FD97923FD8C0FDF4" box="[515,654,547,572]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="B929D673FFDECE29FD97923FD8C0FDF4" box="[515,654,547,572]" italics="true" pageId="46" pageNumber="47">P. caucasica</emphasis>
</taxonomicName>
, the S-shaped dark spot sometimes continues to the posterior margin. The darkish spot on the 
<taxonomicName id="4C5D71E2FFDECE29FE31925BD853FDA9" authorityName="Teslenko &amp; Palatov &amp; Semenchenko" authorityYear="2024" box="[421,541,583,609]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
<emphasis id="B929D673FFDECE29FE31925BD853FDA9" box="[421,541,583,609]" italics="true" pageId="46" pageNumber="47">P. palatovi</emphasis>
</taxonomicName>
<emphasis id="B929D673FFDECE29FDB7925BD833FDA9" bold="true" box="[547,637,583,609]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A21A6B08FFDECE29FDB7925BD833FDA9" box="[547,637,583,609]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
femur is fuzzy, small, and almost oval.
</paragraph>
</subSubSection>
</treatment>
</document>