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<mods:namePart id="D962407BAAC37B86C636B37632A59007">Teslenko, Valentina A.</mods:namePart>
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<mods:namePart id="31889D37DC7957A7D508858BD2A73AD0">Palatov, Dmitry M.</mods:namePart>
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<mods:affiliation id="0D51F8AFD32875E5BAB64ECB2306FD63">Severtsov A. N. Institute of Ecology and Evolution of RAS, 33 Leninsky prospect, 119071 Moscow, Russia.</mods:affiliation>
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<mods:namePart id="D5D2AF1B146C6A428E4983D807168297">Semenchenko, Alexander A.</mods:namePart>
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<mods:affiliation id="22670507972F57FCB528AE62C44D0D74">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences (FSC EATB FEB RAS), 690022, Vladivostok, Russia.</mods:affiliation>
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<mods:nameIdentifier id="A0A68422DAD8CAB494A53B0B2825485B" type="email">semenchenko_alexander@mail.ru</mods:nameIdentifier>
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@ -72,32 +73,32 @@ Teslenko & Semenchenko
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<figureCitation id="136616E4FFD5CE22FF03974BDB77F8B9" box="[151,313,1878,1905]" captionStart-0="FIGURES 128‒131" captionStart-1="FIGURES 132‒136" captionStart-2="FIGURES 137‒140" captionStart-3="FIGURES 141‒147" captionStart-4="FIGURES 148‒149" captionStart-5="FIGURES 150‒156" captionStartId-0="39.[151,264,1854,1878]" captionStartId-1="40.[151,264,1836,1860]" captionStartId-2="41.[151,264,1895,1919]" captionStartId-3="42.[151,264,1742,1766]" captionStartId-4="43.[151,264,1489,1513]" captionStartId-5="44.[151,264,1703,1727]" captionTargetBox-0="[151,1436,191,1830]" captionTargetBox-1="[151,1436,181,1813]" captionTargetBox-2="[158,1430,181,1871]" captionTargetBox-3="[170,1417,181,1718]" captionTargetBox-4="[151,1436,181,1465]" captionTargetBox-5="[151,1435,184,1679]" captionTargetId-0="figure-18@39.[151,1436,181,1830]" captionTargetId-1="figure-24@40.[151,1436,181,1813]" captionTargetId-2="figure-18@41.[158,1430,181,1871]" captionTargetId-3="figure-24@42.[170,1417,181,1718]" captionTargetId-4="figure-18@43.[151,1436,181,1465]" captionTargetId-5="figure-24@44.[151,1435,181,1679]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionTargetPageId-3="42" captionTargetPageId-4="43" captionTargetPageId-5="44" captionText-0="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." captionText-1="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." captionText-2="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." captionText-3="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." captionText-4="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." captionText-5="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="37" pageNumber="38">Figs. 128−156</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FF03974BDB77F8B9" box="[151,313,1878,1905]" captionStart-0="FIGURES 128‒131" captionStart-1="FIGURES 132‒136" captionStart-2="FIGURES 137‒140" captionStart-3="FIGURES 141‒147" captionStart-4="FIGURES 148‒149" captionStart-5="FIGURES 150‒156" captionStartId-0="39.[151,264,1854,1878]" captionStartId-1="40.[151,264,1836,1860]" captionStartId-2="41.[151,264,1895,1919]" captionStartId-3="42.[151,264,1742,1766]" captionStartId-4="43.[151,264,1489,1513]" captionStartId-5="44.[151,264,1703,1727]" captionTargetBox-0="[151,1436,191,1830]" captionTargetBox-1="[151,1436,181,1813]" captionTargetBox-2="[158,1430,181,1871]" captionTargetBox-3="[170,1417,181,1718]" captionTargetBox-4="[151,1436,181,1465]" captionTargetBox-5="[151,1435,184,1679]" captionTargetId-0="figure-18@39.[151,1436,181,1830]" captionTargetId-1="figure-24@40.[151,1436,181,1813]" captionTargetId-2="figure-18@41.[158,1430,181,1871]" captionTargetId-3="figure-24@42.[170,1417,181,1718]" captionTargetId-4="figure-18@43.[151,1436,181,1465]" captionTargetId-5="figure-24@44.[151,1435,181,1679]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionTargetPageId-3="42" captionTargetPageId-4="43" captionTargetPageId-5="44" captionText-0="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." captionText-1="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." captionText-2="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." captionText-3="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." captionText-4="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." captionText-5="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi-0="http://doi.org/10.5281/zenodo.13747810" figureDoi-1="http://doi.org/10.5281/zenodo.13747812" figureDoi-2="http://doi.org/10.5281/zenodo.13747814" figureDoi-3="http://doi.org/10.5281/zenodo.13747816" figureDoi-4="http://doi.org/10.5281/zenodo.13747818" figureDoi-5="http://doi.org/10.5281/zenodo.13747820" httpUri-0="https://zenodo.org/record/13747810/files/figure.png" httpUri-1="https://zenodo.org/record/13747812/files/figure.png" httpUri-2="https://zenodo.org/record/13747814/files/figure.png" httpUri-3="https://zenodo.org/record/13747816/files/figure.png" httpUri-4="https://zenodo.org/record/13747818/files/figure.png" httpUri-5="https://zenodo.org/record/13747820/files/figure.png" pageId="37" pageNumber="38">Figs. 128−156</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C34759EAFFD5CE2AFF039782DEE7F999" lastPageId="45" lastPageNumber="46" pageId="37" pageNumber="38" type="diagnosis">
|
||||
<paragraph id="8BE20A61FFD5CE21FF039782D98AFE19" blockId="37.[151,1436,1950,2013]" lastBlockId="38.[151,1437,151,1977]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
|
||||
<emphasis id="B929D673FFD5CE22FF039782DB5CF870" bold="true" box="[151,274,1950,1976]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
|
||||
Tergum 9 of males is distinguished by the arrangement of serrated ridges on the mesal field that are relatively short and do not reach the posterior margin (
|
||||
<figureCitation id="136616E4FFD5CE22FD6497DFD914F815" box="[752,858,1987,2013]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="37" pageNumber="38">Figs. 133</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FD6497DFD914F815" box="[752,858,1987,2013]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="37" pageNumber="38">Figs. 133</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD5CE22FCF197DFD9DDF815" box="[869,915,1987,2013]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="37" pageNumber="38">137</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FCF197DFD9DDF815" box="[869,915,1987,2013]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="37" pageNumber="38">137</figureCitation>
|
||||
). The hemitergal hooks are strong, with a weak notch in the apical third along the inner edge; the apex is rectangular and rounded (
|
||||
<figureCitation id="136616E4FFD6CE21FBB9908BDE9CFF79" box="[1069,1234,151,177]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="38" pageNumber="39">Figs. 132−133</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FBB9908BDE9CFF79" box="[1069,1234,151,177]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 132−133</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD6CE21FB49908BDF45FF79" box="[1245,1291,151,177]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">137</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FB49908BDF45FF79" box="[1245,1291,151,177]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
|
||||
). The tube of the penis is 1.8 times longer than the sac. The membranous sac is rough, covered with tufts of tiny setae associated with patches, and arranged in concentric lines following the shape of the lobes (
|
||||
<figureCitation id="136616E4FFD6CE21FBB090C3DE84FF31" box="[1060,1226,223,249]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">Figs. 139−140</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FBB090C3DE84FF31" box="[1060,1226,223,249]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 139−140</figureCitation>
|
||||
, B, D). The apical brush is narrow and pointed ventrally (
|
||||
<figureCitation id="136616E4FFD6CE21FDC3911FD8B0FED5" box="[599,766,259,285]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">Figs. 138, 140</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FDC3911FD8B0FED5" box="[599,766,259,285]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 138, 140</figureCitation>
|
||||
, D). The egg has a short, slightly ribbed collar (
|
||||
<figureCitation id="136616E4FFD6CE21FABF911FDFDEFED5" box="[1323,1424,259,285]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." pageId="38" pageNumber="39">Fig. 144</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FABF911FDFDEFED5" box="[1323,1424,259,285]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="38" pageNumber="39">Fig. 144</figureCitation>
|
||||
). The chorion surface is rough and pitted with follicle hexagonal cell impressions, weakly separated from each other; the cell floor is without a central depression; and the number of central pores ranges from one to four (
|
||||
<figureCitation id="136616E4FFD6CE21FAB89157DFDEFEAD" box="[1324,1424,331,357]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." pageId="38" pageNumber="39">Fig. 146</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FAB89157DFDEFEAD" box="[1324,1424,331,357]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="38" pageNumber="39">Fig. 146</figureCitation>
|
||||
). Micropylar orifices lack rims (
|
||||
<figureCitation id="136616E4FFD6CE21FE669173D818FE41" box="[498,598,367,393]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." pageId="38" pageNumber="39">Fig. 147</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FE669173D818FE41" box="[498,598,367,393]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="38" pageNumber="39">Fig. 147</figureCitation>
|
||||
). The larva is distinguished by pale bands along the lateral margins of the pronotum; the width of the pale bands is wider than the semi-oval dark band; a paired dark S-shaped spot extends from the anterior margin to the middle of the pronotum (
|
||||
<figureCitation id="136616E4FFD6CE21FC8491ABD9F6FE19" box="[784,952,439,465]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="38" pageNumber="39">Figs. 148−149</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FC8491ABD9F6FE19" box="[784,952,439,465]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="38" pageNumber="39">Figs. 148−149</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFD6CE21FF5391C6DF52FA9D" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||||
|
|
@ -113,68 +114,68 @@ Body length: males 13.0−14.0 mm (n=6), females
|
|||
; females
|
||||
<quantity id="4CA5A784FFD6CE21FBEA91E2DF68FDD1" box="[1150,1318,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" metricValueMax="4.35" metricValueMin="3.75" pageId="38" pageNumber="39" unit="mm" value="40.5" valueMax="43.5" valueMin="37.5">37.5−43.5 mm</quantity>
|
||||
. Head and pronotum are dorsally grayish-yellow with dark brown spots; mesonotum and metanotum are dark brown, especially in females; the three small, pale spots on the prescutum and scutum form a triangle; the abdomen is grayish-yellow (
|
||||
<figureCitation id="136616E4FFD6CE21FF0B9277DB09FD4D" box="[159,327,619,645]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Figs. 128−129</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FF0B9277DB09FD4D" box="[159,327,619,645]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 128−129</figureCitation>
|
||||
). The wings are yellowish with dark brown veins (
|
||||
<figureCitation id="136616E4FFD6CE21FC159277DE67FD4D" box="[897,1065,619,645]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Figs. 128−129</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FC159277DE67FD4D" box="[897,1065,619,645]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 128−129</figureCitation>
|
||||
). Antennae and palpi are yellowgrey. The M-line on the head is brownish, connected with a triangular brownish spot on the clypeus; the anterior margin and lateral edges of the clypeus are dark brown, clearly contrasting with a pair of transverse small oval pale spots between the clypeal edges and the triangular spot on the clypeus (
|
||||
<figureCitation id="136616E4FFD6CE21FC6D92CBDEECFD39" box="[1017,1186,727,753]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Figs. 128−131</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FC6D92CBDEECFD39" box="[1017,1186,727,753]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 128−131</figureCitation>
|
||||
). Frontal tentorial pits are brownish (
|
||||
<figureCitation id="136616E4FFD6CE21FEAC92E7DBD5FCDD" box="[312,411,763,789]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Fig. 130</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FEAC92E7DBD5FCDD" box="[312,411,763,789]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Fig. 130</figureCitation>
|
||||
). The interocellar area has an X-shaped dark brown spot; the upper branches reach the base of the M-line; the lower branches attain the lateral ocelli; and between the lateral ocelli is a triangular yellowish blurred spot limited basally by a brown stripe of the epicranial suture (
|
||||
<figureCitation id="136616E4FFD6CE21FC42935FDECEFC95" box="[982,1152,835,861]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Figs. 130−131</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FC42935FDECEFC95" box="[982,1152,835,861]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 130−131</figureCitation>
|
||||
). A pair of drop-shaped, bright yellow spots are located between the posterior ocelli and the compound eyes; the tentorial pits are brownish (
|
||||
<figureCitation id="136616E4FFD6CE21FF0B9397DB4DFC6D" box="[159,259,907,933]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Fig. 131</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FF0B9397DB4DFC6D" box="[159,259,907,933]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Fig. 131</figureCitation>
|
||||
). The posterior occipital fold is indicated by a thin light stripe, limited above and below by a narrowdarkened band interrupted in the area of the epicranial stem; the epicranial stem darkened (
|
||||
<figureCitation id="136616E4FFD6CE21FB0893B3DF0DFC01" box="[1180,1347,943,969]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Figs. 130−131</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FB0893B3DF0DFC01" box="[1180,1347,943,969]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 130−131</figureCitation>
|
||||
). A pair of transversely oblique dark bands extends from the base of the occiput to the posterior edge of the compound eyes. Pronotum as wide as head with eyes, slightly narrowing posteriorly, posterolateral margins rounded; anterior margin arcuate, darkened; medial stripe dark brown, widened to anterior and posterior margins, wider at the anterior margin than at the posterior one; pronotal posterior margin darkened; lateral pronotal margins bordered by a dark brown stripe, widening posterolaterally; sometimes the lateral edges are bent down and not visible dorsally (
|
||||
<figureCitation id="136616E4FFD6CE21FAF0947FDAB3FB69" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="38" pageNumber="39">Figs. 117−118</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FAF0947FDAB3FB69" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="38" pageNumber="39">Figs. 117−118</figureCitation>
|
||||
). Pronotal rugosities are grayish-yellow, darkly contoured, and form an X-shaped pattern (
|
||||
<figureCitation id="136616E4FFD6CE21FB7E949BDFDEFB69" box="[1258,1424,1159,1185]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Figs. 130‒131</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FB7E949BDFDEFB69" box="[1258,1424,1159,1185]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 130‒131</figureCitation>
|
||||
); in the posterior third, dark rugosities in the form of narrow paired transverse short stripes reach the lateral pronotal margins (
|
||||
<figureCitation id="136616E4FFD6CE21FE9594D3DBE6FB21" box="[257,424,1231,1257]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Figs. 130−131</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FE9594D3DBE6FB21" box="[257,424,1231,1257]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Figs. 130−131</figureCitation>
|
||||
). The legs and cerci are grayish-brown. Femur with a dark spot basally and a narrow light stripe dorsally; the distal part darkened; tibia darkened at base with a narrow brownish band in the basal ⅕ (
|
||||
<figureCitation id="136616E4FFD6CE21FAF094EFDAB3FAF9" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="38" pageNumber="39">Figs. 115−117</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FAF094EFDAB3FAF9" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="38" pageNumber="39">Figs. 115−117</figureCitation>
|
||||
); cowl also darkened.Anal gills are very small, noticeable in newly molted specimens. Cerci are bicolored, each segment with a narrow yellowish ring basally and a thick grayish-yellow band apically (
|
||||
<figureCitation id="136616E4FFD6CE21FB399527DF41FA9D" box="[1197,1295,1339,1365]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." pageId="38" pageNumber="39">Fig. 128</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FB399527DF41FA9D" box="[1197,1295,1339,1365]" captionStart="FIGURES 128‒131" captionStartId="39.[151,264,1854,1878]" captionTargetBox="[151,1436,191,1830]" captionTargetId="figure-18@39.[151,1436,181,1830]" captionTargetPageId="39" captionText="FIGURES 128‒131. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="38" pageNumber="39">Fig. 128</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFD6CE21FF539543DBB4F871" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||||
<emphasis id="B929D673FFD6CE21FF539543DB45FAB1" bold="true" box="[199,267,1375,1401]" pageId="38" pageNumber="39">Male.</emphasis>
|
||||
Tergum 8 is medially depressed with a central membranous furrow; posterolateral humps are covered with dense and thick setae; along the posterior margin of the furrow, setae are short and sparse (
|
||||
<figureCitation id="136616E4FFD6CE21FB44959FDF7DFA55" box="[1232,1331,1411,1437]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="38" pageNumber="39">Fig. 132</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FB44959FDF7DFA55" box="[1232,1331,1411,1437]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Fig. 132</figureCitation>
|
||||
). Tergum 9 has a sclerotized mesal field; the width is slightly greater than or equal to the length (
|
||||
<figureCitation id="136616E4FFD6CE21FBEF95BBDF6FFA09" box="[1147,1313,1447,1473]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="38" pageNumber="39">Figs. 132−133</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FBEF95BBDF6FFA09" box="[1147,1313,1447,1473]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 132−133</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD6CE21FABB95BBDF13FA09" box="[1327,1373,1447,1473]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">137</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FABB95BBDF13FA09" box="[1327,1373,1447,1473]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
|
||||
). The mesal field bears sharp, heavy sclerotized teeth that merge into long and short longitudinal serrated ridges (8−14), located in the middle of the mesal field, and do not reach the posterior margin (
|
||||
<figureCitation id="136616E4FFD6CE21FBA695F3DEEEF9C1" box="[1074,1184,1519,1545]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="38" pageNumber="39">Figs. 133</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FBA695F3DEEEF9C1" box="[1074,1184,1519,1545]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 133</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD6CE21FB3B95F3DE93F9C1" box="[1199,1245,1519,1545]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">137</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FB3B95F3DE93F9C1" box="[1199,1245,1519,1545]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
|
||||
). Tergum 10 has strong hemitergal hooks (cleared) with a weak notch in the distal third of the inner edge laterally (
|
||||
<figureCitation id="136616E4FFD6CE21FB7D960FDF1BF9E5" box="[1257,1365,1555,1581]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="38" pageNumber="39">Figs. 134</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FB7D960FDF1BF9E5" box="[1257,1365,1555,1581]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 134</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD6CE21FAF5960FDFC1F9E5" box="[1377,1423,1555,1581]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">137</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FAF5960FDFC1F9E5" box="[1377,1423,1555,1581]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
|
||||
); in dorsal view, the notch on the hook corresponds to a shallow depression in the side (
|
||||
<figureCitation id="136616E4FFD6CE21FB07962BDEB7F999" box="[1171,1273,1591,1617]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="38" pageNumber="39">Fig. 132</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FB07962BDEB7F999" box="[1171,1273,1591,1617]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Fig. 132</figureCitation>
|
||||
). The apex of the hook is rectangular and rounded (
|
||||
<figureCitation id="136616E4FFD6CE21FDD79647D8E6F9BD" box="[579,680,1627,1653]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="38" pageNumber="39">Fig. 134</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FDD79647D8E6F9BD" box="[579,680,1627,1653]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Fig. 134</figureCitation>
|
||||
); in its natural state, dorsally, the tip is slightly slanted (
|
||||
<figureCitation id="136616E4FFD6CE21FAB89647DFDEF9BD" box="[1324,1424,1627,1653]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="38" pageNumber="39">Fig. 132</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FAB89647DFDEF9BD" box="[1324,1424,1627,1653]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Fig. 132</figureCitation>
|
||||
). Sensilla basiconica covers the inner side of the hook from top to outer edge; the base of the hook is rounded and covered with long setae (
|
||||
<figureCitation id="136616E4FFD6CE21FE2096BFD814F975" box="[436,602,1699,1725]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="38" pageNumber="39">Figs. 132−134</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FE2096BFD814F975" box="[436,602,1699,1725]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="38" pageNumber="39">Figs. 132−134</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD6CE21FDF396BFD8DBF975" box="[615,661,1699,1725]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">137</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FDF396BFD8DBF975" box="[615,661,1699,1725]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">137</figureCitation>
|
||||
). The artificially everted penis has a tube that is 1.8 times the length of the sac; the dorsal tube is covered with densely arranged longitudinal rows of serrate sclerites, sometimes with small spines; the shape of sclerites in different parts of the tube varies from polygonal to oval elongated; the tube is membranous basally with paired patches of rounded spine bases laterally (
|
||||
<figureCitation id="136616E4FFD6CE21FC409713DE34F8E1" box="[980,1146,1807,1833]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">Figs. 137−140</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FC409713DE34F8E1" box="[980,1146,1807,1833]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 137−140</figureCitation>
|
||||
, A, C). The sac is bulbous and membranous at the base, with two lateral lobes and folds that increase its width (
|
||||
<figureCitation id="136616E4FFD6CE21FBCC972FDF4EF885" box="[1112,1280,1843,1869]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">Figs. 139−140</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FBCC972FDF4EF885" box="[1112,1280,1843,1869]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 139−140</figureCitation>
|
||||
); thin pointed brown spines are present, forming a relatively narrow apical brush that is pointed ventrally (
|
||||
<figureCitation id="136616E4FFD6CE21FB03974BDF72F8B9" box="[1175,1340,1879,1905]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">Figs. 139−140</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FB03974BDF72F8B9" box="[1175,1340,1879,1905]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs. 139−140</figureCitation>
|
||||
, B). The membranous sac is rough and covered with tufts of tiny setae arranged in concentric lines following the shape of the lobes (
|
||||
<figureCitation id="136616E4FFD6CE21FE9E9783DBE7F871" box="[266,425,1951,1977]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." pageId="38" pageNumber="39">Figs 139−140</figureCitation>
|
||||
<figureCitation id="136616E4FFD6CE21FE9E9783DBE7F871" box="[266,425,1951,1977]" captionStart="FIGURES 137‒140" captionStartId="41.[151,264,1895,1919]" captionTargetBox="[158,1430,181,1871]" captionTargetId="figure-18@41.[158,1430,181,1871]" captionTargetPageId="41" captionText="FIGURES 137‒140. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines." figureDoi="http://doi.org/10.5281/zenodo.13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="38" pageNumber="39">Figs 139−140</figureCitation>
|
||||
, B, D).
|
||||
</paragraph>
|
||||
<caption id="DF225AE9FFD7CE20FF039722D8AFF856" pageId="39" pageNumber="40" startId="39.[151,264,1854,1878]" targetBox="[151,1436,191,1830]" targetPageId="39" targetType="figure">
|
||||
<caption id="DF225AE9FFD7CE20FF039722D8AFF856" ID-DOI="http://doi.org/10.5281/zenodo.13747810" ID-Zenodo-Dep="13747810" httpUri="https://zenodo.org/record/13747810/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,264,1854,1878]" targetBox="[151,1436,191,1830]" targetPageId="39" targetType="figure">
|
||||
<paragraph id="8BE20A61FFD7CE20FF039722D8AFF856" blockId="39.[151,1437,1854,1951]" pageId="39" pageNumber="40">
|
||||
<emphasis id="B929D673FFD7CE20FF039722DB22F89E" bold="true" box="[151,364,1854,1878]" pageId="39" pageNumber="40">FIGURES 128‒131.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFD7CE20FEE59722D84FF89E" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[369,513,1854,1878]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
|
||||
|
|
@ -186,7 +187,7 @@ Tergum 8 is medially depressed with a central membranous furrow; posterolateral
|
|||
Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. 128. Paratype. Female habitus, dorsal. 129. Holotype. Male, habitus, dorsal. 130. Holotype, male, head and pronotum, dorsal. 131. Paratype, female, head and pronotum, dorsal.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFD8CE2FFF039730D9B6F844" pageId="40" pageNumber="41" startId="40.[151,264,1836,1860]" targetBox="[151,1436,181,1813]" targetPageId="40" targetType="figure">
|
||||
<caption id="DF225AE9FFD8CE2FFF039730D9B6F844" ID-DOI="http://doi.org/10.5281/zenodo.13747812" ID-Zenodo-Dep="13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,264,1836,1860]" targetBox="[151,1436,181,1813]" targetPageId="40" targetType="figure">
|
||||
<paragraph id="8BE20A61FFD8CE2FFF039730D9B6F844" blockId="40.[151,1436,1836,1933]" pageId="40" pageNumber="41">
|
||||
<emphasis id="B929D673FFD8CE2FFF039730DB22F88C" bold="true" box="[151,364,1836,1860]" pageId="40" pageNumber="41">FIGURES 132‒136.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFD8CE2FFEE59730D84FF88C" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[369,513,1836,1860]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
|
||||
|
|
@ -200,7 +201,7 @@ Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River
|
|||
136. Female abdominal tip, variations in color.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFD9CE2EFF03977BDF76F823" pageId="41" pageNumber="42" startId="41.[151,264,1895,1919]" targetBox="[158,1430,181,1871]" targetPageId="41" targetType="figure">
|
||||
<caption id="DF225AE9FFD9CE2EFF03977BDF76F823" ID-DOI="http://doi.org/10.5281/zenodo.13747814" ID-Zenodo-Dep="13747814" httpUri="https://zenodo.org/record/13747814/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,264,1895,1919]" targetBox="[158,1430,181,1871]" targetPageId="41" targetType="figure">
|
||||
<paragraph id="8BE20A61FFD9CE2EFF03977BDF76F823" blockId="41.[151,1437,1895,2027]" pageId="41" pageNumber="42">
|
||||
<emphasis id="B929D673FFD9CE2EFF03977BDB3FF8B7" bold="true" box="[151,369,1895,1919]" pageId="41" pageNumber="42">FIGURES 137‒140.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFD9CE2EFEEE977BD840F8B7" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[378,526,1895,1919]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
|
||||
|
|
@ -212,7 +213,7 @@ Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River
|
|||
Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Male. 137. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 138. Penis, ventral. 139. Chaetotaxy of the penis, dorsal. 140. Chaetotaxy of the penis, ventral. A. Leaf-shaped scales with small spines. B, D. Tufts of tiny setae arranged in concentric lines, following the shape of the lobes. C. Oval scales with small spines.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFDACE2DFF0396D2D842F852" pageId="42" pageNumber="43" startId="42.[151,264,1742,1766]" targetBox="[170,1417,181,1718]" targetPageId="42" targetType="figure">
|
||||
<caption id="DF225AE9FFDACE2DFF0396D2D842F852" ID-DOI="http://doi.org/10.5281/zenodo.13747816" ID-Zenodo-Dep="13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,264,1742,1766]" targetBox="[170,1417,181,1718]" targetPageId="42" targetType="figure">
|
||||
<paragraph id="8BE20A61FFDACE2DFF0396D2D842F852" blockId="42.[151,1437,1742,1946]" pageId="42" pageNumber="43">
|
||||
<emphasis id="B929D673FFDACE2DFF0396D2DB22F92E" bold="true" box="[151,364,1742,1766]" pageId="42" pageNumber="43">FIGURES 141‒147.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFDACE2DFEE596D2D84FF92E" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[369,513,1742,1766]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
|
||||
|
|
@ -224,7 +225,7 @@ Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River
|
|||
Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141. Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142. Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFDBCE2CFF0395CDDE11F9C5" pageId="43" pageNumber="44" startId="43.[151,264,1489,1513]" targetBox="[151,1436,181,1465]" targetPageId="43" targetType="figure">
|
||||
<caption id="DF225AE9FFDBCE2CFF0395CDDE11F9C5" ID-DOI="http://doi.org/10.5281/zenodo.13747818" ID-Zenodo-Dep="13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,264,1489,1513]" targetBox="[151,1436,181,1465]" targetPageId="43" targetType="figure">
|
||||
<paragraph id="8BE20A61FFDBCE2CFF0395CDDE11F9C5" blockId="43.[151,1436,1489,1549]" pageId="43" pageNumber="44">
|
||||
<emphasis id="B929D673FFDBCE2CFF0395CDDB22FA21" bold="true" box="[151,364,1489,1513]" pageId="43" pageNumber="44">FIGURES 148‒149.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFDBCE2CFEE595CDD84FFA21" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[369,513,1489,1513]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
|
||||
|
|
@ -239,22 +240,22 @@ Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River
|
|||
<paragraph id="8BE20A61FFDBCE2CFF539643DBAFF90A" blockId="43.[151,1437,1631,2017]" pageId="43" pageNumber="44">
|
||||
<emphasis id="B929D673FFDBCE2CFF539643DB6BF9B1" bold="true" box="[199,293,1631,1657]" pageId="43" pageNumber="44">Female.</emphasis>
|
||||
Sternum 8 has a subgenital plate, occupying almost half of the sternum width, with two small triangular lobes posteromedially. The lobes projecting slightly beyond the anterior margin of sternum 9 are covered with rufous setae (
|
||||
<figureCitation id="136616E4FFDBCE2CFEB996BBDB9AF90A" box="[301,468,1703,1730]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." pageId="43" pageNumber="44">Figs. 135−136</figureCitation>
|
||||
<figureCitation id="136616E4FFDBCE2CFEB996BBDB9AF90A" box="[301,468,1703,1730]" captionStart="FIGURES 132‒136" captionStartId="40.[151,264,1836,1860]" captionTargetBox="[151,1436,181,1813]" captionTargetId="figure-24@40.[151,1436,181,1813]" captionTargetPageId="40" captionText="FIGURES 132‒136. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. 132. Male, abdominal tip, dorsal. 133. Male, abdominal tip, dorsal, cleared. 134. Male, hemitergal hook, lateral, cleared. 135‒136. Female abdominal tip, variations in color." figureDoi="http://doi.org/10.5281/zenodo.13747812" httpUri="https://zenodo.org/record/13747812/files/figure.png" pageId="43" pageNumber="44">Figs. 135−136</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFDBCE2CFF5396D7DA9CF829" blockId="43.[151,1437,1631,2017]" pageId="43" pageNumber="44">
|
||||
<emphasis id="B929D673FFDBCE2CFF5396D7DAB3F92D" bold="true" box="[199,253,1739,1765]" pageId="43" pageNumber="44">Egg.</emphasis>
|
||||
Elongate and oval, with mean dimensions of 435 × 272 μm (n = 6), with a short collar with the same diameter at the rim and the base; the rim is weakly ribbed (
|
||||
<figureCitation id="136616E4FFDBCE2CFC8D96F3D9F1F8C1" box="[793,959,1775,1801]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." pageId="43" pageNumber="44">Figs. 141−144</figureCitation>
|
||||
<figureCitation id="136616E4FFDBCE2CFC8D96F3D9F1F8C1" box="[793,959,1775,1801]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="43" pageNumber="44">Figs. 141−144</figureCitation>
|
||||
). The neck is not constricted.A mushroom-shaped anchor plate is present on some eggs and covered with globular bodies concentrated in small groups (
|
||||
<figureCitation id="136616E4FFDBCE2CFAF0970FDAB0F89A" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." pageId="43" pageNumber="44">Figs. 142, 145</figureCitation>
|
||||
<figureCitation id="136616E4FFDBCE2CFAF0970FDAB0F89A" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="43" pageNumber="44">Figs. 142, 145</figureCitation>
|
||||
). The chorion surface is rough and pitted; the chorion is covered throughout with follicle hexagonal cell impressions; hexagonal cells with differences in cell widths; cells separated from adjacent ones by poorly expressed boundaries; the cell floor without a central depression with one or four pores (
|
||||
<figureCitation id="136616E4FFDBCE2CFC6A9763DEEDF851" box="[1022,1187,1919,1945]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." pageId="43" pageNumber="44">Figs. 144, 146</figureCitation>
|
||||
<figureCitation id="136616E4FFDBCE2CFC6A9763DEEDF851" box="[1022,1187,1919,1945]" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="43" pageNumber="44">Figs. 144, 146</figureCitation>
|
||||
). A ring of micropyles is located ca. 1/3 of the egg length from the anterior pole, the micropylar orifices are without rims (
|
||||
<figureCitation id="136616E4FFDBCE2CFB6697BFDA8BF829" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." pageId="43" pageNumber="44">Figs. 141−142, 147</figureCitation>
|
||||
<figureCitation id="136616E4FFDBCE2CFB6697BFDA8BF829" captionStart="FIGURES 141‒147" captionStartId="42.[151,264,1742,1766]" captionTargetBox="[170,1417,181,1718]" captionTargetId="figure-24@42.[170,1417,181,1718]" captionTargetPageId="42" captionText="FIGURES 141‒147. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin. Egg. 141.Appearance without mushroom-shaped anchor plate, micropylar row, lateral. 142.Appearance with mushroomshaped anchor plate, micropylar row, lateral. 143. Appearance, dorsolateral. 144. Collar with weakly ribbed rim, lateral. 145. Mushroom-shaped anchor with globular bodies concentrated in groups, lateral. 146. Chorion structure with follicle hexagonal cell impressions, the cell floor without a central depression with one or four pores. 147. Chorion structure with micropylar row, the micropylar orifices without rims." figureDoi="http://doi.org/10.5281/zenodo.13747816" httpUri="https://zenodo.org/record/13747816/files/figure.png" pageId="43" pageNumber="44">Figs. 141−142, 147</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF225AE9FFDCCE2BFF0396BBDF54F8E3" pageId="44" pageNumber="45" startId="44.[151,264,1703,1727]" targetBox="[151,1435,184,1679]" targetPageId="44" targetType="figure">
|
||||
<caption id="DF225AE9FFDCCE2BFF0396BBDF54F8E3" ID-DOI="http://doi.org/10.5281/zenodo.13747820" ID-Zenodo-Dep="13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,264,1703,1727]" targetBox="[151,1435,184,1679]" targetPageId="44" targetType="figure">
|
||||
<paragraph id="8BE20A61FFDCCE2BFF0396BBDF54F8E3" blockId="44.[151,1436,1703,1835]" pageId="44" pageNumber="45">
|
||||
<emphasis id="B929D673FFDCCE2BFF0396BBDB22F977" bold="true" box="[151,364,1703,1727]" pageId="44" pageNumber="45">FIGURES 150‒156.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFDCCE2BFEE596BBD84FF977" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[369,513,1703,1727]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="palatovi" status="sp. nov.">
|
||||
|
|
@ -275,54 +276,54 @@ Mature males of medium size, length
|
|||
, and cerci greater than
|
||||
<quantity id="4CA5A784FFDCCE2BFF0397BDDAA3F874" box="[151,237,1953,1980]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="44" pageNumber="45" unit="mm" value="8.8">8.8 mm</quantity>
|
||||
contain more than 36 segments. The color is generally pale, yellow-brown in dorsal view (
|
||||
<figureCitation id="136616E4FFDCCE2BFB7D97BDDFDEF873" box="[1257,1424,1953,1979]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="44" pageNumber="45">Figs. 148−149</figureCitation>
|
||||
<figureCitation id="136616E4FFDCCE2BFB7D97BDDFDEF873" box="[1257,1424,1953,1979]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="44" pageNumber="45">Figs. 148−149</figureCitation>
|
||||
), with a distinct dark pattern on the head, pronotum, mesonotum, and metanotum, but with a less distinct pattern on the abdomen. The ventral body side is yellow-brownish. Body covered with short dark clothing hairs, with a dorsomesal band of erect silky white setae more pronounced on the last abdominal terga (
|
||||
<figureCitation id="136616E4FFDDCE2AFBED90A7DF6EFF1D" box="[1145,1312,187,213]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFBED90A7DF6EFF1D" box="[1145,1312,187,213]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
). Antennae and cerci are yellow-brown. Clypeus on head basally with paired dark and wide V-shaped spots, outlining branches of yellow M-line from above. Head with a large transverse dark spot, vaguely resembling a pilot's badge with truncated “wings” not extending beyond epicranial arms; the spot limits the pale M-line from below extending from the anterior ocellus to the posterior ocelli; a small triangular spot between the ocelli darkened; epicranial stem and occiput edge also darkened (
|
||||
<figureCitation id="136616E4FFDDCE2AFE439173D831FE41" box="[471,639,367,393]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFE439173D831FE41" box="[471,639,367,393]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
). The occipital ridge is distinct and complete; short spines are arranged in the medial part of the fold; along the postocular crest, spines are noticeably longer and pointed. Mandibles (
|
||||
<figureCitation id="136616E4FFDDCE2AFAFB918FDA8CFE19" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Fig. 153</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFAFB918FDA8CFE19" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 153</figureCitation>
|
||||
) are typical for
|
||||
<taxonomicName id="4C5D71E2FFDDCE2AFEEE91ABDBF6FE19" box="[378,440,439,465]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B929D673FFDDCE2AFEEE91ABDBF6FE19" box="[378,440,439,465]" italics="true" pageId="45" pageNumber="46">Perla</emphasis>
|
||||
</taxonomicName>
|
||||
: heavily sclerotized, with a rounded outer margin; left mandible with five pointed teeth and two medial rows of setae on the molar area; the marginal row of setae extends from the base of the 4th tooth to the basal third of the mandible; right mandible with six pointed teeth; single medial setal row extends from the base of the first tooth to the base of the mandible; the configuration of the marginal setal row is the same as that of the left mandible. Lacinia bidentate with a large, heavily sclerotized, and strongly curved apical tooth (
|
||||
<figureCitation id="136616E4FFDDCE2AFB6E925BDF13FDA9" box="[1274,1373,582,609]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Fig. 152</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFB6E925BDF13FDA9" box="[1274,1373,582,609]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 152</figureCitation>
|
||||
). The subapical tooth is shorter; the length does not exceed 2/3 of the apical tooth length. The marginal fringe of setae along the inner margin is complete; a tuft of long and dense setae of marginal fringe is clearly visible below the subapical tooth. Galea length reaches 1/2 of subapical tooth length (
|
||||
<figureCitation id="136616E4FFDDCE2AFC1B92AFD9BCFD05" box="[911,1010,690,717]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Fig. 152</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFC1B92AFD9BCFD05" box="[911,1010,690,717]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 152</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFDDCE2AFF5392CBDBF9F9E5" blockId="45.[151,1437,151,2013]" pageId="45" pageNumber="46">
|
||||
Pronotum transverse, not wider than the head, with obtusely rounded anterior corners; lateral margins and posterior angles are evenly rounded (
|
||||
<figureCitation id="136616E4FFDDCE2AFDBA92E7D89AFCDD" box="[558,724,763,789]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFDBA92E7D89AFCDD" box="[558,724,763,789]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
). The pronotal fringe is complete with mixed short and relatively long setae. The pronotum has a thin brown stripe along the edge; paired semi-oval pale bands outline a full (or incomplete, open anterolaterally) oval brown band on the lateral pronotal fields (
|
||||
<figureCitation id="136616E4FFDDCE2AFBA0935FDE93FC95" box="[1076,1245,835,861]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFBA0935FDE93FC95" box="[1076,1245,835,861]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
). An oval brown band is narrower than semi-oval pale bands, especially in the anterior 1/3. The central pronotal field is yellow with a brown median band narrowed anteriorly and markedly widened posteriorly; a paired vague S-shaped dark spot extends from the anterior margin to the middle of the pronotum (
|
||||
<figureCitation id="136616E4FFDDCE2AFCFA93B3DE58FC01" box="[878,1046,943,969]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFCFA93B3DE58FC01" box="[878,1046,943,969]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
). Mesonotum and metanotum with an identical Ψ-shaped dark pattern medially, posterior margin darkly margined; two wide dark bands limiting the wing pads; a narrow dark stripe runs along the lateral edge of the wing pads. Legs are yellow and covered with dark closing hairs (
|
||||
<figureCitation id="136616E4FFDDCE2AFEA29407DB91FBFD" box="[310,479,1051,1077]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFEA29407DB91FBFD" box="[310,479,1051,1077]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
). The femur and tibia bear stout, short, acute bristles along the inner and outer edges and a very dense fringe of long, silky hairs along the outer margins (
|
||||
<figureCitation id="136616E4FFDDCE2AFC379423DE49FB91" box="[931,1031,1086,1113]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Fig. 150</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFC379423DE49FB91" box="[931,1031,1086,1113]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 150</figureCitation>
|
||||
). The femur is covered with sparse, short, and irregular red bristles; in the distal half closer to the outer edge, with a darkened small and almost oval spot, the distal edge of the femur is also darkened (
|
||||
<figureCitation id="136616E4FFDDCE2AFD4E949BD973FB69" box="[730,829,1158,1185]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Fig. 150</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFD4E949BD973FB69" box="[730,829,1158,1185]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 150</figureCitation>
|
||||
). Hind femur ca. 3.3−3.6 times as long as wide. Tibia has a diffuse, narrow brownish band basally. The pattern on the abdomen is indistinct; the dark closing hairs are well-marked; the terga are mainly yellowish with a dark median and two lateral longitudinal bands; the lateral dark bands on terga 4−10 are triangular, extending to the anterior margin (
|
||||
<figureCitation id="136616E4FFDDCE2AFC3C94EFDE1EFAC5" box="[936,1104,1267,1293]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFC3C94EFDE1EFAC5" box="[936,1104,1267,1293]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
). The posterior terga margins bear acute bristles that are mostly short, with one or two relatively long setae (
|
||||
<figureCitation id="136616E4FFDDCE2AFB91950BDE29FAF9" box="[1029,1127,1302,1329]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Fig. 151</figureCitation>
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||||
<figureCitation id="136616E4FFDDCE2AFB91950BDE29FAF9" box="[1029,1127,1302,1329]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 151</figureCitation>
|
||||
). Anal gills are small (
|
||||
<figureCitation id="136616E4FFDDCE2AFAF0950BDAB1FA9D" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
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<figureCitation id="136616E4FFDDCE2AFAF0950BDAB1FA9D" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
). Cerci are yellow, and no less than 36 segmented with striking rings of dark spines (
|
||||
<figureCitation id="136616E4FFDDCE2AFB259527DF16FA9D" box="[1201,1368,1339,1365]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
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<figureCitation id="136616E4FFDDCE2AFB259527DF16FA9D" box="[1201,1368,1339,1365]" captionStart="FIGURES 148‒149" captionStartId="43.[151,264,1489,1513]" captionTargetBox="[151,1436,181,1465]" captionTargetId="figure-18@43.[151,1436,181,1465]" captionTargetPageId="43" captionText="FIGURES 148‒149. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva, males, habitus, dorsal. Variation of the pattern." figureDoi="http://doi.org/10.5281/zenodo.13747818" httpUri="https://zenodo.org/record/13747818/files/figure.png" pageId="45" pageNumber="46">Figs. 148−149</figureCitation>
|
||||
). Fine pilosity between dark spines is tightly pressed to the segment surface (
|
||||
<figureCitation id="136616E4FFDDCE2AFC099543DE0DFAB1" box="[925,1091,1374,1401]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Figs. 154−156</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFC099543DE0DFAB1" box="[925,1091,1374,1401]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Figs. 154−156</figureCitation>
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||||
); short fine hairs have the same length as the length of the basal cercal segment; towards the middle of the cerci, fine pilosity length decreases; and on the apical cercal segments, the length of the fine pilosity is slightly>1/3 of the segment length (
|
||||
<figureCitation id="136616E4FFDDCE2AFB7F95BBDF00FA09" box="[1259,1358,1446,1473]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Fig. 156</figureCitation>
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||||
<figureCitation id="136616E4FFDDCE2AFB7F95BBDF00FA09" box="[1259,1358,1446,1473]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 156</figureCitation>
|
||||
). Long silky fine colorless hairs are included in the circlet in tufts (no more than
|
||||
<quantity id="4CA5A784FFDDCE2AFC4595D7D9B3FA2D" box="[977,1021,1483,1509]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="45" pageNumber="46" unit="in" value="8.0">8 in</quantity>
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each) and form a dorsal longitudinal fringe, especially on the basal segments (
|
||||
<figureCitation id="136616E4FFDDCE2AFDF795F3D88BF9C1" box="[611,709,1518,1545]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Fig. 154</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFDF795F3D88BF9C1" box="[611,709,1518,1545]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Fig. 154</figureCitation>
|
||||
); the length and density of long silky hairs also decrease towards the apex (
|
||||
<figureCitation id="136616E4FFDDCE2AFE91960FDBE5F9E5" box="[261,427,1554,1581]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." pageId="45" pageNumber="46">Figs. 155‒156</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFE91960FDBE5F9E5" box="[261,427,1554,1581]" captionStart="FIGURES 150‒156" captionStartId="44.[151,264,1703,1727]" captionTargetBox="[151,1435,184,1679]" captionTargetId="figure-24@44.[151,1435,181,1679]" captionTargetPageId="44" captionText="FIGURES 150‒156. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray, Gelendzhik District, Papay River, Pshada River Basin; Tuapse District, Shapsukho River. Larva. 150. Right middle leg, dorsal. 151. Chaetotaxy of abdominal terga 7 and 8, dorsal, cleared. 152. Left lacinia, galea, maxillary palpus, ventral. 153. Left and right mandibles, ventral. 154−156. Chaetotaxy of the right cercus, lateral. 154. Basal cercal segments. 155. Middle cercal segments. 156. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747820" httpUri="https://zenodo.org/record/13747820/files/figure.png" pageId="45" pageNumber="46">Figs. 155‒156</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFDDCE2AFF53962ADEE7F999" blockId="45.[151,1437,151,2013]" box="[199,1193,1590,1617]" pageId="45" pageNumber="46">
|
||||
|
|
@ -429,7 +430,7 @@ North Caucasus. North-eastern coast of the Black Sea.
|
|||
.
|
||||
<collectingRegion id="4999C483FFDDCE2AFBBF9713DEAFF8E1" box="[1067,1249,1807,1833]" country="Russia" name="Krasnodar" pageId="45" pageNumber="46">Krasnodar Kray</collectingRegion>
|
||||
. Gelendzhik and Tuapse Districts. The species was found in small foothill rivers along the Black Sea coast (
|
||||
<figureCitation id="136616E4FFDDCE2AFB1F972FDF7FF885" box="[1163,1329,1842,1869]" captionStart="FIGURES 157‒158" captionStartId="47.[151,264,689,713]" captionTargetBox="[151,1436,181,665]" captionTargetId="figure-18@47.[151,1436,181,665]" captionTargetPageId="47" captionText="FIGURES 157‒158. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray. Type habitat. Gelendzhik District, Papay River, Pshada River Basin." pageId="45" pageNumber="46">Figs. 157−158</figureCitation>
|
||||
<figureCitation id="136616E4FFDDCE2AFB1F972FDF7FF885" box="[1163,1329,1842,1869]" captionStart="FIGURES 157‒158" captionStartId="47.[151,264,689,713]" captionTargetBox="[151,1436,181,665]" captionTargetId="figure-18@47.[151,1436,181,665]" captionTargetPageId="47" captionText="FIGURES 157‒158. Perla palatovi sp. nov. Caucasus, Russia, Krasnodar Kray. Type habitat. Gelendzhik District, Papay River, Pshada River Basin." figureDoi="http://doi.org/10.5281/zenodo.13747822" httpUri="https://zenodo.org/record/13747822/files/figure.png" pageId="45" pageNumber="46">Figs. 157−158</figureCitation>
|
||||
), flowing through low-mountain forest valleys
|
||||
<quantity id="4CA5A784FFDDCE2AFDA2974BD8D5F8B9" box="[566,667,1879,1905]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="45" pageNumber="46" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">10−12 m</quantity>
|
||||
wide with a pebble and rocky bottom and a current up to
|
||||
|
|
|
|||
|
|
@ -1,60 +1,61 @@
|
|||
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<mods:title id="8C98BDF754202137D9F6647104A37F22">Overview of the Caucasian Perla Geoffroy, 1762 (Plecoptera: Perlidae) based on morphological and molecular data with description of two new species</mods:title>
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<mods:namePart id="E135DC918A6E4E4C225BC20E6D419D1F">Palatov, Dmitry M.</mods:namePart>
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<treatment id="03F4BB77FFE0CE12FF03918EDF70FDF5" LSID="urn:lsid:plazi:treatment:03F4BB77FFE0CE12FF03918EDF70FDF5" httpUri="http://treatment.plazi.org/id/03F4BB77FFE0CE12FF03918EDF70FDF5" lastPageId="21" lastPageNumber="22" pageId="16" pageNumber="17">
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<paragraph id="8BE20A61FFE0CE17FF0391ABDB55FE19" blockId="16.[151,569,402,465]" box="[151,283,438,465]" pageId="16" pageNumber="17">
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<figureCitation id="136616E4FFE0CE17FF0391ABDB55FE19" box="[151,283,438,465]" captionStart-0="FIGURES 50‒53" captionStart-1="FIGURES 54‒58" captionStart-2="FIGURES 59‒61" captionStart-3="FIGURES 62‒68" captionStartId-0="17.[151,264,1918,1942]" captionStartId-1="18.[151,264,1861,1885]" captionStartId-2="19.[151,264,1414,1438]" captionStartId-3="20.[151,264,1845,1869]" captionTargetBox-0="[151,1436,181,1886]" captionTargetBox-1="[158,1430,273,1831]" captionTargetBox-2="[151,1436,188,1390]" captionTargetBox-3="[151,1436,181,1822]" captionTargetId-0="figure-18@17.[151,1436,181,1894]" captionTargetId-1="figure-79@18.[158,1430,273,1838]" captionTargetId-2="figure-18@19.[151,1436,181,1390]" captionTargetId-3="figure-24@20.[151,1436,181,1822]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." captionText-1="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." captionText-2="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." captionText-3="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="16" pageNumber="17">Figs. 50‒68</figureCitation>
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||||
<figureCitation id="136616E4FFE0CE17FF0391ABDB55FE19" box="[151,283,438,465]" captionStart-0="FIGURES 50‒53" captionStart-1="FIGURES 54‒58" captionStart-2="FIGURES 59‒61" captionStart-3="FIGURES 62‒68" captionStartId-0="17.[151,264,1918,1942]" captionStartId-1="18.[151,264,1861,1885]" captionStartId-2="19.[151,264,1414,1438]" captionStartId-3="20.[151,264,1845,1869]" captionTargetBox-0="[151,1436,181,1886]" captionTargetBox-1="[158,1430,273,1831]" captionTargetBox-2="[151,1436,188,1390]" captionTargetBox-3="[151,1436,181,1822]" captionTargetId-0="figure-18@17.[151,1436,181,1894]" captionTargetId-1="figure-79@18.[158,1430,273,1838]" captionTargetId-2="figure-18@19.[151,1436,181,1390]" captionTargetId-3="figure-24@20.[151,1436,181,1822]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." captionText-1="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." captionText-2="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." captionText-3="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi-0="http://doi.org/10.5281/zenodo.13747764" figureDoi-1="http://doi.org/10.5281/zenodo.13747770" figureDoi-2="http://doi.org/10.5281/zenodo.13747772" figureDoi-3="http://doi.org/10.5281/zenodo.13747774" httpUri-0="https://zenodo.org/record/13747764/files/figure.png" httpUri-1="https://zenodo.org/record/13747770/files/figure.png" httpUri-2="https://zenodo.org/record/13747772/files/figure.png" httpUri-3="https://zenodo.org/record/13747774/files/figure.png" pageId="16" pageNumber="17">Figs. 50‒68</figureCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFE0CE17FF0391E7D8BBFDDB" blockId="16.[151,1160,507,715]" box="[151,757,507,531]" pageId="16" pageNumber="17">
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||||
<bibRefCitation id="EFCC7790FFE0CE17FF0391E7DBD7FDDB" author="Zhiltzova, L. A." box="[151,409,507,531]" pageId="16" pageNumber="17" pagination="872 - 880" refId="ref26643" refString="Zhiltzova, L. A. (1961) On the study of the fauna of Plecoptera of the Caucasus V. Plecoptera of Armenia. Entomologicheskoe obozrenie, 40 (4), 872 - 880. [in Russian]" type="journal article" year="1961">Zhiltzova, 1961:874‒876</bibRefCitation>
|
||||
|
|
@ -130,70 +131,70 @@ is one of the darkest-colored species of Caucasian
|
|||
.
|
||||
</emphasis>
|
||||
The male is distinguished by the long mesal field, which is widened and rounded posteriorly; the length does not exceed its width; and the longitudinal serrated ridges (7−8) are concentrated in the middle (
|
||||
<figureCitation id="136616E4FFE0CE17FCE9935DDE49FC94" box="[893,1031,833,860]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." pageId="16" pageNumber="17">Figs. 55, 57</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FCE9935DDE49FC94" box="[893,1031,833,860]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." figureDoi="http://doi.org/10.5281/zenodo.13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="16" pageNumber="17">Figs. 55, 57</figureCitation>
|
||||
). The hemitergal hooks are strongly swollen in the place of the bend, forming a noticeable rounded protrusion, and the apexes are bluntly rounded (
|
||||
<figureCitation id="136616E4FFE0CE17FF0B9395DB62FC6C" box="[159,300,905,932]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." pageId="16" pageNumber="17">Figs. 54−57</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FF0B9395DB62FC6C" box="[159,300,905,932]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." figureDoi="http://doi.org/10.5281/zenodo.13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="16" pageNumber="17">Figs. 54−57</figureCitation>
|
||||
). An artificially everted penis bears the bulbous, membranous sac with a narrow apical brush ending in triangles of spines ventrally and dorsally; the membranous sac is rough, covered with dense tiny setae, clearly visible near the folds of the lobes (
|
||||
<figureCitation id="136616E4FFE0CE17FDA793CDD8DAFC24" box="[563,660,977,1004]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." pageId="16" pageNumber="17">Figs. 58</figureCitation>
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<figureCitation id="136616E4FFE0CE17FDA793CDD8DAFC24" box="[563,660,977,1004]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." figureDoi="http://doi.org/10.5281/zenodo.13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="16" pageNumber="17">Figs. 58</figureCitation>
|
||||
, D). The egg is spindle-shaped, tapering evenly to each pole (
|
||||
<figureCitation id="136616E4FFE0CE17FAFB93CDDAF9FBC7" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="16" pageNumber="17">Fig. 62</figureCitation>
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||||
<figureCitation id="136616E4FFE0CE17FAFB93CDDAF9FBC7" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi="http://doi.org/10.5281/zenodo.13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="16" pageNumber="17">Fig. 62</figureCitation>
|
||||
). The collar is short, but the rim is thick and rough (
|
||||
<figureCitation id="136616E4FFE0CE17FC8593E9D929FBC7" box="[785,871,1013,1039]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="16" pageNumber="17">Fig. 64</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FC8593E9D929FBC7" box="[785,871,1013,1039]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi="http://doi.org/10.5281/zenodo.13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="16" pageNumber="17">Fig. 64</figureCitation>
|
||||
). The chorion is rough and punctuate throughout; punctations are mostly uniform, but small punctations are also present near the micropylar row located near the equator; micropylar orifices with a raised thin edge are placed laterally on the pit wall; inside the pit there may be additional 1−2 small orifices (
|
||||
<figureCitation id="136616E4FFE0CE17FE7C947DD8D3FBB4" box="[488,669,1121,1148]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="16" pageNumber="17">Figs. 62, 65−68</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FE7C947DD8D3FBB4" box="[488,669,1121,1148]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi="http://doi.org/10.5281/zenodo.13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="16" pageNumber="17">Figs. 62, 65−68</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFE0CE17FF539499DBF3F987" blockId="16.[151,1437,761,2012]" pageId="16" pageNumber="17">
|
||||
<emphasis id="B929D673FFE0CE17FF539499D89CFB57" bold="true" box="[199,722,1157,1183]" pageId="16" pageNumber="17">Complimentary description—adult habitus.</emphasis>
|
||||
Males and females are macropterous (
|
||||
<figureCitation id="136616E4FFE0CE17FB169499DE91FB68" box="[1154,1247,1157,1184]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." pageId="16" pageNumber="17">Figs. 50</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FB169499DE91FB68" box="[1154,1247,1157,1184]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747764" httpUri="https://zenodo.org/record/13747764/files/figure.png" pageId="16" pageNumber="17">Figs. 50</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFE0CE17FB7F9499DF45FB68" box="[1259,1291,1157,1184]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="16" pageNumber="17">59</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FB7F9499DF45FB68" box="[1259,1291,1157,1184]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="16" pageNumber="17">59</figureCitation>
|
||||
). The M-line on the head is rufous, smooth, and clearly defined (
|
||||
<figureCitation id="136616E4FFE0CE17FD4C94B5D962FB0C" box="[728,812,1193,1220]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." pageId="16" pageNumber="17">Fig. 53</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FD4C94B5D962FB0C" box="[728,812,1193,1220]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747764" httpUri="https://zenodo.org/record/13747764/files/figure.png" pageId="16" pageNumber="17">Fig. 53</figureCitation>
|
||||
). The interocellar area has a dark brown spot extending upward to the M-line and down to the lateral ocelli, with the lateral branches of this spot reaching the lateral edges of the head above tentorial pits; the lateral edges of the clypeus are also dark brown; and paired rufous spots are located behind the compound eyes (
|
||||
<figureCitation id="136616E4FFE0CE17FE469509D87EFAF8" box="[466,560,1301,1328]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." pageId="16" pageNumber="17">Figs. 53</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FE469509D87EFAF8" box="[466,560,1301,1328]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747764" httpUri="https://zenodo.org/record/13747764/files/figure.png" pageId="16" pageNumber="17">Figs. 53</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFE0CE17FDA89509D8C9FAF8" box="[572,647,1301,1328]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="16" pageNumber="17">59–60</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FDA89509D8C9FAF8" box="[572,647,1301,1328]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="16" pageNumber="17">59–60</figureCitation>
|
||||
). Along the anterior margin of the clypeus, paired patches between the lateral ocelli and compound eyes, as well as paired patches on the occiput, are yellow (
|
||||
<figureCitation id="136616E4FFE0CE17FBF99525DE85FA9C" box="[1133,1227,1337,1364]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." pageId="16" pageNumber="17">Figs. 53</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FBF99525DE85FA9C" box="[1133,1227,1337,1364]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747764" httpUri="https://zenodo.org/record/13747764/files/figure.png" pageId="16" pageNumber="17">Figs. 53</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFE0CE17FB4C9525DF6AFA9C" box="[1240,1316,1337,1364]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="16" pageNumber="17">59−60</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FB4C9525DF6AFA9C" box="[1240,1316,1337,1364]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="16" pageNumber="17">59−60</figureCitation>
|
||||
). Tentorial pits above lateral ocelli are clearly defined, bean-shaped, yellow, and have a small, dark spot inside (
|
||||
<figureCitation id="136616E4FFE0CE17FA929541DF2DFAB0" box="[1286,1379,1373,1400]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." pageId="16" pageNumber="17">Figs. 53</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FA929541DF2DFAB0" box="[1286,1379,1373,1400]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747764" httpUri="https://zenodo.org/record/13747764/files/figure.png" pageId="16" pageNumber="17">Figs. 53</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFE0CE17FAE49541DFDEFABF" box="[1392,1424,1373,1399]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="16" pageNumber="17">60</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FAE49541DFDEFABF" box="[1392,1424,1373,1399]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="16" pageNumber="17">60</figureCitation>
|
||||
). Pronotum with a dark medial band noticeably widening towards the anterior margin; band surrounded by pale rugosities of an X-shaped pattern; lateral fields the same color as the medial band (
|
||||
<figureCitation id="136616E4FFE0CE17FBBF95B9DEC6FA08" box="[1067,1160,1445,1472]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." pageId="16" pageNumber="17">Figs. 53</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FBBF95B9DEC6FA08" box="[1067,1160,1445,1472]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747764" httpUri="https://zenodo.org/record/13747764/files/figure.png" pageId="16" pageNumber="17">Figs. 53</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFE0CE17FB0795B9DE90FA08" box="[1171,1246,1445,1472]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="16" pageNumber="17">59–60</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FB0795B9DE90FA08" box="[1171,1246,1445,1472]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="16" pageNumber="17">59–60</figureCitation>
|
||||
). Legs are brown (
|
||||
<figureCitation id="136616E4FFE0CE17FF0B95D5DB65FA2C" box="[159,299,1481,1508]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." pageId="16" pageNumber="17">Figs. 50, 52</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FF0B95D5DB65FA2C" box="[159,299,1481,1508]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747764" httpUri="https://zenodo.org/record/13747764/files/figure.png" pageId="16" pageNumber="17">Figs. 50, 52</figureCitation>
|
||||
). Femur darkened distally with a thin yellow band along the outer edge (
|
||||
<figureCitation id="136616E4FFE0CE17FBFC95D5DEF3FA2C" box="[1128,1213,1481,1508]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="16" pageNumber="17">Fig. 59</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FBFC95D5DEF3FA2C" box="[1128,1213,1481,1508]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="16" pageNumber="17">Fig. 59</figureCitation>
|
||||
); dark closing hairs and scattered brown bristles along the inner and outer edges; a medial band without hairs and bristles with a small oval pale spot distally. Tibia has a thin yellow band along the outer edge; the base darkened (
|
||||
<figureCitation id="136616E4FFE0CE17FB38960DDF47F9E4" box="[1196,1289,1553,1580]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." pageId="16" pageNumber="17">Figs. 50</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FB38960DDF47F9E4" box="[1196,1289,1553,1580]" captionStart="FIGURES 50‒53" captionStartId="17.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1886]" captionTargetId="figure-18@17.[151,1436,181,1894]" captionTargetPageId="17" captionText="FIGURES 50‒53. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan,Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747764" httpUri="https://zenodo.org/record/13747764/files/figure.png" pageId="16" pageNumber="17">Figs. 50</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFE0CE17FA82960DDF78F9E4" box="[1302,1334,1553,1580]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="16" pageNumber="17">59</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FA82960DDF78F9E4" box="[1302,1334,1553,1580]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="16" pageNumber="17">59</figureCitation>
|
||||
); there is a small dark band distally.
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFE0CE15FF539645DEA0FF31" blockId="16.[151,1437,761,2012]" lastBlockId="18.[151,1437,150,249]" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
|
||||
<emphasis id="B929D673FFE0CE17FF539645DB45F9BB" bold="true" box="[199,267,1625,1651]" pageId="16" pageNumber="17">Male.</emphasis>
|
||||
Tergum 8 medially has a membranous furrow and posterolateral humps covered with dense, thick, and long setae. Tergum 9 has a depressed, sclerotized mesal field slightly widened posteriorly, and the length does not exceed its width. On the mesal plate, the longitudinal serrated ridges (7–8) are heavily sclerotized and concentrated mesally and do not reach the posterior edge, which is smooth and rounded with a weak notch in the middle (
|
||||
<figureCitation id="136616E4FFE0CE17FAF096D9DAAFF8CC" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." pageId="16" pageNumber="17">Figs. 55, 57</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FAF096D9DAAFF8CC" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." figureDoi="http://doi.org/10.5281/zenodo.13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="16" pageNumber="17">Figs. 55, 57</figureCitation>
|
||||
). Tergum 10 has hemitergites that are curved at an obtuse angle and directed obliquely upward; each hook is strongly swollen in place of the bend on the outer and dorsal sides, forming a noticeable rounded protrusion covered with long setae; the same setae cover the outer edge of the hook (
|
||||
<figureCitation id="136616E4FFE0CE17FCF9972DD9B8F884" box="[877,1014,1841,1868]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." pageId="16" pageNumber="17">Figs. 54‒57</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FCF9972DD9B8F884" box="[877,1014,1841,1868]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." figureDoi="http://doi.org/10.5281/zenodo.13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="16" pageNumber="17">Figs. 54‒57</figureCitation>
|
||||
). The apex of the hook is rounded (on a cleared specimen, the hook apex is rectangular-rounded) and covered with sensilla basiconica, which is extended at the tip to the front and outer edges (
|
||||
<figureCitation id="136616E4FFE0CE17FDD89765D899F85C" box="[588,727,1913,1940]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." pageId="16" pageNumber="17">Figs. 56‒57</figureCitation>
|
||||
<figureCitation id="136616E4FFE0CE17FDD89765D899F85C" box="[588,727,1913,1940]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." figureDoi="http://doi.org/10.5281/zenodo.13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="16" pageNumber="17">Figs. 56‒57</figureCitation>
|
||||
). An artificially everted penis has a tube covered dorsally with densely arranged longitudinal rows of serrate sclerites, turning into scales sometimes with small spines; the shape of sclerites and rows in different parts of the tube is different; the tube is membranous basally with paired patches of rounded spine bases laterally (
|
||||
<figureCitation id="136616E4FFE2CE15FE7D908BD80BFF79" box="[489,581,150,177]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." pageId="18" pageNumber="19">Figs. 58</figureCitation>
|
||||
<figureCitation id="136616E4FFE2CE15FE7D908BD80BFF79" box="[489,581,150,177]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." figureDoi="http://doi.org/10.5281/zenodo.13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="18" pageNumber="19">Figs. 58</figureCitation>
|
||||
, A‒B). The sac is bulbous and membranous, with thin pointed brown spines, forming a narrow apical brush ending in triangles of spines ventrally and dorsally (
|
||||
<figureCitation id="136616E4FFE2CE15FBD790A7DEEFFF1D" box="[1091,1185,186,213]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." pageId="18" pageNumber="19">Figs. 58</figureCitation>
|
||||
<figureCitation id="136616E4FFE2CE15FBD790A7DEEFFF1D" box="[1091,1185,186,213]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." figureDoi="http://doi.org/10.5281/zenodo.13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="18" pageNumber="19">Figs. 58</figureCitation>
|
||||
, C). The membranous sac is rough, covered with dense, tiny setae, clearly visible near the folds of the lobes (
|
||||
<figureCitation id="136616E4FFE2CE15FBF490C3DEF3FF31" box="[1120,1213,222,249]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." pageId="18" pageNumber="19">Figs. 58</figureCitation>
|
||||
<figureCitation id="136616E4FFE2CE15FBF490C3DEF3FF31" box="[1120,1213,222,249]" captionStart="FIGURES 54‒58" captionStartId="18.[151,264,1861,1885]" captionTargetBox="[158,1430,273,1831]" captionTargetId="figure-79@18.[158,1430,273,1838]" captionTargetPageId="18" captionText="FIGURES 54‒58. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes." figureDoi="http://doi.org/10.5281/zenodo.13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="18" pageNumber="19">Figs. 58</figureCitation>
|
||||
, D).
|
||||
</paragraph>
|
||||
<caption id="DF225AE9FFE1CE16FF039762DF77F872" pageId="17" pageNumber="18" startId="17.[151,264,1918,1942]" targetBox="[151,1436,181,1886]" targetPageId="17" targetType="figure">
|
||||
<caption id="DF225AE9FFE1CE16FF039762DF77F872" ID-DOI="http://doi.org/10.5281/zenodo.13747764" ID-Zenodo-Dep="13747764" httpUri="https://zenodo.org/record/13747764/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,264,1918,1942]" targetBox="[151,1436,181,1886]" targetPageId="17" targetType="figure">
|
||||
<paragraph id="8BE20A61FFE1CE16FF039762DF77F872" blockId="17.[151,1437,1918,1979]" pageId="17" pageNumber="18">
|
||||
<emphasis id="B929D673FFE1CE16FF039762DB1CF85E" bold="true" box="[151,338,1918,1942]" pageId="17" pageNumber="18">FIGURES 50‒53.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFE1CE16FEC29762D8E1F85E" authority="Zhiltzova, 1961" authorityName="Zhiltzova" authorityYear="1961" box="[342,687,1918,1942]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kiritshenkoi">
|
||||
|
|
@ -203,7 +204,7 @@ Tergum 8 medially has a membranous furrow and posterolateral humps covered with
|
|||
. Caucasus:Armenia, Stepanavan, Kamenka River;Azerbaijan, Lelyakoran. Paratypes. Males. 50. Habitus, dorsal. 51. Abdominal tip, dorsal. 52. Habitus, dorsal. 53. Head and pronotum, dorsal.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFE2CE15FF039759D95EF825" pageId="18" pageNumber="19" startId="18.[151,264,1861,1885]" targetBox="[158,1430,273,1831]" targetPageId="18" targetType="figure">
|
||||
<caption id="DF225AE9FFE2CE15FF039759D95EF825" ID-DOI="http://doi.org/10.5281/zenodo.13747770" ID-Zenodo-Dep="13747770" httpUri="https://zenodo.org/record/13747770/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,264,1861,1885]" targetBox="[158,1430,273,1831]" targetPageId="18" targetType="figure">
|
||||
<paragraph id="8BE20A61FFE2CE15FF039759D95EF825" blockId="18.[151,1437,1861,2029]" pageId="18" pageNumber="19">
|
||||
<emphasis id="B929D673FFE2CE15FF039759DB1AF895" bold="true" box="[151,340,1861,1885]" pageId="18" pageNumber="19">FIGURES 54‒58.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFE2CE15FECE9759D8F5F895" authority="Zhiltzova, 1961" authorityName="Zhiltzova" authorityYear="1961" box="[346,699,1861,1885]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kiritshenkoi">
|
||||
|
|
@ -213,7 +214,7 @@ Tergum 8 medially has a membranous furrow and posterolateral humps covered with
|
|||
. Caucasus, Armenia. Male. 54. Paratype male, abdominal tip, dorsal. 55. Mesal plate and hemitergites, dorsal, alcohol. 56. Hemitergal hook swollen in the place of the bend, dorsocaudally, alcohol. 57. Mesal plate, dorsal, and hemitergites, lateral, cleared. 58. Chaetotaxy of artificially everted penis, dorsal, cleared. A. Leafshaped scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. C. Pointed brown spines in the apical brush. D. Tiny setae covered the sac lobes.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFE3CE14FF03959AD974FA0A" pageId="19" pageNumber="20" startId="19.[151,264,1414,1438]" targetBox="[151,1436,188,1390]" targetPageId="19" targetType="figure">
|
||||
<caption id="DF225AE9FFE3CE14FF03959AD974FA0A" ID-DOI="http://doi.org/10.5281/zenodo.13747772" ID-Zenodo-Dep="13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,264,1414,1438]" targetBox="[151,1436,188,1390]" targetPageId="19" targetType="figure">
|
||||
<paragraph id="8BE20A61FFE3CE14FF03959AD974FA0A" blockId="19.[151,1436,1414,1474]" pageId="19" pageNumber="20">
|
||||
<emphasis id="B929D673FFE3CE14FF03959ADB1BFA56" bold="true" box="[151,341,1414,1438]" pageId="19" pageNumber="20">FIGURES 59‒61.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFE3CE14FEC8959AD8F0FA56" authority="Zhiltzova, 1961" authorityName="Zhiltzova" authorityYear="1961" box="[348,702,1414,1438]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kiritshenkoi">
|
||||
|
|
@ -226,11 +227,11 @@ Tergum 8 medially has a membranous furrow and posterolateral humps covered with
|
|||
<paragraph id="8BE20A61FFE3CE14FF539608D887F953" blockId="19.[151,1437,1556,1978]" pageId="19" pageNumber="20">
|
||||
<emphasis id="B929D673FFE3CE14FF539608DB6BF9E6" bold="true" box="[199,293,1556,1582]" pageId="19" pageNumber="20">Female.</emphasis>
|
||||
Subgenital plate occupies ⅔ of the width of sternum 8 with bilobed projection posteromedially (
|
||||
<figureCitation id="136616E4FFE3CE14FAFB9608DAF8F99A" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="19" pageNumber="20">Fig. 61</figureCitation>
|
||||
<figureCitation id="136616E4FFE3CE14FAFB9608DAF8F99A" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="19" pageNumber="20">Fig. 61</figureCitation>
|
||||
); lobes are small, narrow, pointed, and covered with small, reddish setae (
|
||||
<figureCitation id="136616E4FFE3CE14FC799624DE0EF99A" box="[1005,1088,1592,1618]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="19" pageNumber="20">Fig. 61</figureCitation>
|
||||
<figureCitation id="136616E4FFE3CE14FC799624DE0EF99A" box="[1005,1088,1592,1618]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="19" pageNumber="20">Fig. 61</figureCitation>
|
||||
). The cerci are long, projecting far beyond the end of the wings; the basal cercal segments are brown; and the distal cercal segments are bicolored, yellowish at the base and brown distally (
|
||||
<figureCitation id="136616E4FFE3CE14FDFC969CD8F2F953" box="[616,700,1664,1691]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." pageId="19" pageNumber="20">Fig. 59</figureCitation>
|
||||
<figureCitation id="136616E4FFE3CE14FDFC969CD8F2F953" box="[616,700,1664,1691]" captionStart="FIGURES 59‒61" captionStartId="19.[151,264,1414,1438]" captionTargetBox="[151,1436,188,1390]" captionTargetId="figure-18@19.[151,1436,181,1390]" captionTargetPageId="19" captionText="FIGURES 59‒61. Perla kiritshenkoi Zhiltzova, 1961. Caucasus, Armenia, Stepanavan, Kamenka River. Female. 59. Habitus, dorsal. 60. Head and pronotum, dorsal. 61. Abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747772" httpUri="https://zenodo.org/record/13747772/files/figure.png" pageId="19" pageNumber="20">Fig. 59</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFE3CE14FF5396B8DA8AF85E" blockId="19.[151,1437,1556,1978]" pageId="19" pageNumber="20">
|
||||
|
|
@ -238,17 +239,17 @@ Subgenital plate occupies ⅔ of the width of sternum 8 with bilobed projection
|
|||
The
|
||||
<typeStatus id="54E6B4C3FFE3CE14FEA096B9DB2AF977" box="[308,356,1701,1727]" pageId="19" pageNumber="20">type</typeStatus>
|
||||
material egg is spindle-shaped, tapering evenly to each pole (
|
||||
<figureCitation id="136616E4FFE3CE14FB9D96B8DEDFF976" box="[1033,1169,1700,1726]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="19" pageNumber="20">Figs. 62–63</figureCitation>
|
||||
<figureCitation id="136616E4FFE3CE14FB9D96B8DEDFF976" box="[1033,1169,1700,1726]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi="http://doi.org/10.5281/zenodo.13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="19" pageNumber="20">Figs. 62–63</figureCitation>
|
||||
), with mean dimensions of 428 x 242 µm (n = 3; length without anchor). The collar is short, and the rim is thick and rough, with a series of small pits around the base (
|
||||
<figureCitation id="136616E4FFE3CE14FE5C96F0D81CF8CE" box="[456,594,1772,1798]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="19" pageNumber="20">Figs. 63–64</figureCitation>
|
||||
<figureCitation id="136616E4FFE3CE14FE5C96F0D81CF8CE" box="[456,594,1772,1798]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi="http://doi.org/10.5281/zenodo.13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="19" pageNumber="20">Figs. 63–64</figureCitation>
|
||||
). Chorion is rough and punctuate throughout (
|
||||
<figureCitation id="136616E4FFE3CE14FBC196F0DF49F8CF" box="[1109,1287,1772,1799]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="19" pageNumber="20">Figs. 62, 65–68</figureCitation>
|
||||
<figureCitation id="136616E4FFE3CE14FBC196F0DF49F8CF" box="[1109,1287,1772,1799]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi="http://doi.org/10.5281/zenodo.13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="19" pageNumber="20">Figs. 62, 65–68</figureCitation>
|
||||
); punctations are large, deep, and mostly uniform, but small punctations are also present, especially near the micropylar row, which is located almost at the equator (
|
||||
<figureCitation id="136616E4FFE3CE14FDDB9728D8EAF886" box="[591,676,1844,1870]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="19" pageNumber="20">Fig. 62</figureCitation>
|
||||
<figureCitation id="136616E4FFE3CE14FDDB9728D8EAF886" box="[591,676,1844,1870]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi="http://doi.org/10.5281/zenodo.13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="19" pageNumber="20">Fig. 62</figureCitation>
|
||||
); micropylar orifices with a slightly raised thin edge are located not on the bottom but laterally on the pit wall (
|
||||
<figureCitation id="136616E4FFE3CE14FDE59744D88AF8BA" box="[625,708,1880,1906]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="19" pageNumber="20">Fig. 67</figureCitation>
|
||||
<figureCitation id="136616E4FFE3CE14FDE59744D88AF8BA" box="[625,708,1880,1906]" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi="http://doi.org/10.5281/zenodo.13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="19" pageNumber="20">Fig. 67</figureCitation>
|
||||
); inside the pit there may be an additional 1–2 small orifices (
|
||||
<figureCitation id="136616E4FFE3CE14FAFB9744DAF9F85E" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." pageId="19" pageNumber="20">Fig. 68</figureCitation>
|
||||
<figureCitation id="136616E4FFE3CE14FAFB9744DAF9F85E" captionStart="FIGURES 62‒68" captionStartId="20.[151,264,1845,1869]" captionTargetBox="[151,1436,181,1822]" captionTargetId="figure-24@20.[151,1436,181,1822]" captionTargetPageId="20" captionText="FIGURES 62‒68. Perla kiritshenkoi Zhiltzova, 1961.Caucasus:Armenia, Kamenka River.Egg. 62.Appearance with micropylar row, lateral. 63. Same, dorsolateral. 64. Collar rim, and small pits around the base, lateral. 65−66. Chorion structure with deep punctations. 67. Micropyle with a slightly raised, thin edge. 68. Micropyle on the pit wall with the addition small orifices." figureDoi="http://doi.org/10.5281/zenodo.13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="19" pageNumber="20">Fig. 68</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFE3CE14FF5397BCDBDAF872" blockId="19.[151,1437,1556,1978]" box="[199,404,1952,1978]" pageId="19" pageNumber="20">
|
||||
|
|
@ -256,7 +257,7 @@ material egg is spindle-shaped, tapering evenly to each pole (
|
|||
Unknown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF225AE9FFE4CE13FF039729DF24F85E" pageId="20" pageNumber="21" startId="20.[151,264,1845,1869]" targetBox="[151,1436,181,1822]" targetPageId="20" targetType="figure">
|
||||
<caption id="DF225AE9FFE4CE13FF039729DF24F85E" ID-DOI="http://doi.org/10.5281/zenodo.13747774" ID-Zenodo-Dep="13747774" httpUri="https://zenodo.org/record/13747774/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,264,1845,1869]" targetBox="[151,1436,181,1822]" targetPageId="20" targetType="figure">
|
||||
<paragraph id="8BE20A61FFE4CE13FF039729DF24F85E" blockId="20.[151,1436,1845,1942]" pageId="20" pageNumber="21">
|
||||
<emphasis id="B929D673FFE4CE13FF039729DB1CF886" bold="true" box="[151,338,1845,1870]" pageId="20" pageNumber="21">FIGURES 62‒68.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFE4CE13FEC2972AD8E1F886" authority="Zhiltzova, 1961" authorityName="Zhiltzova" authorityYear="1961" box="[342,687,1846,1870]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="kiritshenkoi">
|
||||
|
|
@ -412,7 +413,7 @@ Darilmaz
|
|||
2016
|
||||
</bibRefCitation>
|
||||
) (
|
||||
<figureCitation id="136616E4FFE5CE12FB83918FDE35FE65" box="[1047,1147,402,429]" captionStart="FIGURE 159" captionStartId="47.[151,250,1780,1804]" captionTargetBox="[158,1430,846,1757]" captionTargetId="figure-40@47.[158,1430,846,1757]" captionTargetPageId="47" captionText="FIGURE 159. The map of the Perla species distribution in Caucasus according to our data. Perla caicasica—pink, P. persica— green, P. kiritshenkoi—blue, P. schapsugica sp. nov.—white, P. pallida—yellow, and P. palatovi sp. nov.—red." pageId="21" pageNumber="22">Fig. 159</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FB83918FDE35FE65" box="[1047,1147,402,429]" captionStart="FIGURE 159" captionStartId="47.[151,250,1780,1804]" captionTargetBox="[158,1430,846,1757]" captionTargetId="figure-40@47.[158,1430,846,1757]" captionTargetPageId="47" captionText="FIGURE 159. The map of the Perla species distribution in Caucasus according to our data. Perla caicasica—pink, P. persica— green, P. kiritshenkoi—blue, P. schapsugica sp. nov.—white, P. pallida—yellow, and P. palatovi sp. nov.—red." figureDoi="http://doi.org/10.5281/zenodo.13747824" httpUri="https://zenodo.org/record/13747824/files/figure.png" pageId="21" pageNumber="22">Fig. 159</figureCitation>
|
||||
). Adults emerge from the second half of May through the first half of July. Adults were found in
|
||||
<collectingCountry id="F34A4AF1FFE5CE12FC2791ABDE5BFE19" box="[947,1045,439,465]" name="Armenia" pageId="21" pageNumber="22">Armenia</collectingCountry>
|
||||
at altitudes of
|
||||
|
|
|
|||
|
|
@ -1,60 +1,61 @@
|
|||
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<mods:title id="5EE2333B1272B9F6362E9DF2AA4DF1A0">Overview of the Caucasian Perla Geoffroy, 1762 (Plecoptera: Perlidae) based on morphological and molecular data with description of two new species</mods:title>
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<mods:namePart id="4AB992A91C10611320288161A0DBCB3C">Teslenko, Valentina A.</mods:namePart>
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<mods:affiliation id="B94488F08DBE6339E62616B09DEE3694">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences (FSC EATB FEB RAS), 690022, Vladivostok, Russia.</mods:affiliation>
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<mods:namePart id="1A61F28EF2B779C1E03BF16234875393">Teslenko, Valentina A.</mods:namePart>
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<mods:affiliation id="BC826676F0308A16D51C702085A751DA">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences (FSC EATB FEB RAS), 690022, Vladivostok, Russia.</mods:affiliation>
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<mods:roleTerm id="B886797027048DA8DDFDEBC00B645FE7">Author</mods:roleTerm>
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||||
<mods:namePart id="E3F6A1924E997F30E47BA6D42814B5C1">Palatov, Dmitry M.</mods:namePart>
|
||||
<mods:affiliation id="FC9C7384F1AC8220B0AA98591B28EC29">Severtsov A. N. Institute of Ecology and Evolution of RAS, 33 Leninsky prospect, 119071 Moscow, Russia.</mods:affiliation>
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||||
<mods:namePart id="B6EFF0C39E090E5285A2744E6EF65FEB">Palatov, Dmitry M.</mods:namePart>
|
||||
<mods:affiliation id="C58972E05E139B35ED46098192DC3B78">Severtsov A. N. Institute of Ecology and Evolution of RAS, 33 Leninsky prospect, 119071 Moscow, Russia.</mods:affiliation>
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|
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<mods:role id="A4759916E98103126485353274316CD1">
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</mods:role>
|
||||
<mods:namePart id="83E747262BC9AD4B6EBCDECEEBA1B62D">Semenchenko, Alexander A.</mods:namePart>
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<mods:affiliation id="40764C43FA94F44E75DE8A3EEC2E0934">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences (FSC EATB FEB RAS), 690022, Vladivostok, Russia.</mods:affiliation>
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<mods:affiliation id="6A20B763828052B7030A2076F8B505E4">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences (FSC EATB FEB RAS), 690022, Vladivostok, Russia.</mods:affiliation>
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@ -72,36 +73,36 @@ Teslenko & Palatov
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<subSubSection id="C34759EAFFE5CE12FF0392AFDB55FD05" box="[151,283,691,717]" pageId="21" pageNumber="22" type="description">
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<paragraph id="8BE20A61FFE5CE12FF0392AFDB55FD05" blockId="21.[151,724,654,717]" box="[151,283,691,717]" pageId="21" pageNumber="22">
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<figureCitation id="136616E4FFE5CE12FF0392AFDB55FD05" box="[151,283,691,717]" captionStart-0="FIGURES 69‒72" captionStart-1="FIGURES 73‒76" captionStart-2="FIGURES 77‒81" captionStart-3="FIGURES 82‒89" captionStart-4="FIGURES 90‒92" captionStart-5="FIGURES 93‒98" captionStartId-0="22.[151,264,1890,1914]" captionStartId-1="23.[151,264,1636,1660]" captionStartId-2="24.[151,264,1826,1850]" captionStartId-3="25.[151,264,1901,1925]" captionStartId-4="26.[151,264,1856,1880]" captionStartId-5="27.[151,264,1775,1799]" captionTargetBox-0="[164,1423,181,1866]" captionTargetBox-1="[151,1436,181,1612]" captionTargetBox-2="[183,1405,181,1793]" captionTargetBox-3="[158,1430,181,1877]" captionTargetBox-4="[151,1436,181,1833]" captionTargetBox-5="[151,1436,187,1752]" captionTargetId-0="figure-24@22.[164,1423,181,1866]" captionTargetId-1="figure-18@23.[151,1436,181,1612]" captionTargetId-2="figure-24@24.[183,1405,181,1802]" captionTargetId-3="figure-18@25.[158,1430,181,1877]" captionTargetId-4="figure-24@26.[151,1436,181,1833]" captionTargetId-5="figure-18@27.[151,1436,181,1752]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionTargetPageId-3="25" captionTargetPageId-4="26" captionTargetPageId-5="27" captionText-0="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." captionText-1="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." captionText-2="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." captionText-3="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." captionText-4="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." captionText-5="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="21" pageNumber="22">Figs. 69‒98</figureCitation>
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<figureCitation id="136616E4FFE5CE12FF0392AFDB55FD05" box="[151,283,691,717]" captionStart-0="FIGURES 69‒72" captionStart-1="FIGURES 73‒76" captionStart-2="FIGURES 77‒81" captionStart-3="FIGURES 82‒89" captionStart-4="FIGURES 90‒92" captionStart-5="FIGURES 93‒98" captionStartId-0="22.[151,264,1890,1914]" captionStartId-1="23.[151,264,1636,1660]" captionStartId-2="24.[151,264,1826,1850]" captionStartId-3="25.[151,264,1901,1925]" captionStartId-4="26.[151,264,1856,1880]" captionStartId-5="27.[151,264,1775,1799]" captionTargetBox-0="[164,1423,181,1866]" captionTargetBox-1="[151,1436,181,1612]" captionTargetBox-2="[183,1405,181,1793]" captionTargetBox-3="[158,1430,181,1877]" captionTargetBox-4="[151,1436,181,1833]" captionTargetBox-5="[151,1436,187,1752]" captionTargetId-0="figure-24@22.[164,1423,181,1866]" captionTargetId-1="figure-18@23.[151,1436,181,1612]" captionTargetId-2="figure-24@24.[183,1405,181,1802]" captionTargetId-3="figure-18@25.[158,1430,181,1877]" captionTargetId-4="figure-24@26.[151,1436,181,1833]" captionTargetId-5="figure-18@27.[151,1436,181,1752]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionTargetPageId-3="25" captionTargetPageId-4="26" captionTargetPageId-5="27" captionText-0="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." captionText-1="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." captionText-2="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." captionText-3="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." captionText-4="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." captionText-5="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi-0="http://doi.org/10.5281/zenodo.13747780" figureDoi-1="http://doi.org/10.5281/zenodo.13747784" figureDoi-2="http://doi.org/10.5281/zenodo.13747788" figureDoi-3="http://doi.org/10.5281/zenodo.13747790" figureDoi-4="http://doi.org/10.5281/zenodo.13747792" figureDoi-5="http://doi.org/10.5281/zenodo.13747794" httpUri-0="https://zenodo.org/record/13747780/files/figure.png" httpUri-1="https://zenodo.org/record/13747784/files/figure.png" httpUri-2="https://zenodo.org/record/13747788/files/figure.png" httpUri-3="https://zenodo.org/record/13747790/files/figure.png" httpUri-4="https://zenodo.org/record/13747792/files/figure.png" httpUri-5="https://zenodo.org/record/13747794/files/figure.png" pageId="21" pageNumber="22">Figs. 69‒98</figureCitation>
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C34759EAFFE5CE1AFF0392E6DEC4F82A" lastPageId="29" lastPageNumber="30" pageId="21" pageNumber="22" type="diagnosis">
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<paragraph id="8BE20A61FFE5CE12FF0392E6D88AFB21" blockId="21.[151,1437,762,2013]" pageId="21" pageNumber="22">
|
||||
<emphasis id="B929D673FFE5CE12FF0392E6DB5CFCDC" bold="true" box="[151,274,762,788]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
|
||||
Tergum 9 of males is distinguished by a butterfly-shaped mesal field; the width is greater than the length (
|
||||
<figureCitation id="136616E4FFE5CE12FF0B9303DABAFCF1" box="[159,244,799,825]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="21" pageNumber="22">Fig. 73</figureCitation>
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||||
<figureCitation id="136616E4FFE5CE12FF0B9303DABAFCF1" box="[159,244,799,825]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="21" pageNumber="22">Fig. 73</figureCitation>
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); the longitudinal serrated ridges are short at the lateral margins, the long serrated rides are located in the middle of the mesal field (
|
||||
<figureCitation id="136616E4FFE5CE12FE29935FD85FFC95" box="[445,529,835,861]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="21" pageNumber="22">Fig. 73</figureCitation>
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<figureCitation id="136616E4FFE5CE12FE29935FD85FFC95" box="[445,529,835,861]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="21" pageNumber="22">Fig. 73</figureCitation>
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||||
). The hemitergal hook is rounded at the apex (
|
||||
<figureCitation id="136616E4FFE5CE12FB82935FDE3DFC95" box="[1046,1139,835,861]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="21" pageNumber="22">Figs. 73</figureCitation>
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<figureCitation id="136616E4FFE5CE12FB82935FDE3DFC95" box="[1046,1139,835,861]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="21" pageNumber="22">Figs. 73</figureCitation>
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||||
,
|
||||
<figureCitation id="136616E4FFE5CE12FBEA935FDED0FC95" box="[1150,1182,835,861]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." pageId="21" pageNumber="22">77</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FBEA935FDED0FC95" box="[1150,1182,835,861]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="21" pageNumber="22">77</figureCitation>
|
||||
). The tube of the penis is ca. 2.3 times longer than the sac. The membranous sac is smooth, with a pair of rounded lateral lobes dorsally and ventrally and a tongue-shaped medial lobe dorsally; ventrally, the sac is deeply depressed, and the lobes are almost indistinguishable except for a pair of lateral lobes (
|
||||
<figureCitation id="136616E4FFE5CE12FD7493B3D925FC01" box="[736,875,943,969]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." pageId="21" pageNumber="22">Figs. 77‒81</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FD7493B3D925FC01" box="[736,875,943,969]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="21" pageNumber="22">Figs. 77‒81</figureCitation>
|
||||
). The apical brush is long, widened dorsally, and slightly pointed ventrally (
|
||||
<figureCitation id="136616E4FFE5CE12FE5793CFD803FC25" box="[451,589,979,1005]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." pageId="21" pageNumber="22">Figs. 79‒81</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FE5793CFD803FC25" box="[451,589,979,1005]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="21" pageNumber="22">Figs. 79‒81</figureCitation>
|
||||
). The egg is round-oval, anchor small, and covers only the collar rim (
|
||||
<figureCitation id="136616E4FFE5CE12FAFB93CFDB45FBD9" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="21" pageNumber="22">Fig. 82, 84‒85</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FAFB93CFDB45FBD9" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="21" pageNumber="22">Fig. 82, 84‒85</figureCitation>
|
||||
). Collar wide, with barely noticeable discernable ribs; rim smooth, with weak incisions (
|
||||
<figureCitation id="136616E4FFE5CE12FB4A93EBDFDFFBD9" box="[1246,1425,1014,1041]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="21" pageNumber="22">Figs. 83, 85‒86</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FB4A93EBDFDFFBD9" box="[1246,1425,1014,1041]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="21" pageNumber="22">Figs. 83, 85‒86</figureCitation>
|
||||
). Chorion is coarsely pitted; pits are round, large, and deep; the chorion surface between pits is rough (
|
||||
<figureCitation id="136616E4FFE5CE12FA929407DFC1FBFD" box="[1286,1423,1051,1077]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="21" pageNumber="22">Figs. 84, 88</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FA929407DFC1FBFD" box="[1286,1423,1051,1077]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="21" pageNumber="22">Figs. 84, 88</figureCitation>
|
||||
). Micropyles in the pits are set not on flows but on the walls of the pits; orifices are raised with thin rims (
|
||||
<figureCitation id="136616E4FFE5CE12FA989423DF11FB91" box="[1292,1375,1087,1113]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="21" pageNumber="22">Fig. 89</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FA989423DF11FB91" box="[1292,1375,1087,1113]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="21" pageNumber="22">Fig. 89</figureCitation>
|
||||
). The larvae are distinctive by the color pattern of the head, pronotum, and cercal segment pilosity (
|
||||
<figureCitation id="136616E4FFE5CE12FB3B947FDF79FBB5" box="[1199,1335,1123,1149]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 90‒91</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FB3B947FDF79FBB5" box="[1199,1335,1123,1149]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="21" pageNumber="22">Figs. 90‒91</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFE5CE12FAD6947FDFC3FBB5" box="[1346,1421,1123,1149]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="21" pageNumber="22">96‒98</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FAD6947FDFC3FBB5" box="[1346,1421,1123,1149]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="21" pageNumber="22">96‒98</figureCitation>
|
||||
).
|
||||
<emphasis id="B929D673FFE5CE12FF03949ADB0AFB68" bold="true" box="[151,324,1158,1184]" pageId="21" pageNumber="22">Adult habitus.</emphasis>
|
||||
Body length: males
|
||||
|
|
@ -111,45 +112,45 @@ Body length: males
|
|||
(n=5); macropterous forewings are long, equal to body length, reaching males 21.3−23.0 mm and females
|
||||
<quantity id="4CA5A784FFE5CE12FC4D94B7DE30FB0D" box="[985,1150,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="2.495" metricValueMax="2.87" metricValueMin="2.12" pageId="21" pageNumber="22" unit="mm" value="24.95" valueMax="28.7" valueMin="21.2">21.2−28.7 mm</quantity>
|
||||
(
|
||||
<figureCitation id="136616E4FFE5CE12FB1894B7DF58FB0D" box="[1164,1302,1195,1221]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 69‒70</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FB1894B7DF58FB0D" box="[1164,1302,1195,1221]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Figs. 69‒70</figureCitation>
|
||||
); wingspan: males 44.9−49.0 mm and females
|
||||
<quantity id="4CA5A784FFE5CE12FD8C94D2D88EFB21" box="[536,704,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="4.745" metricValueMax="4.97" metricValueMin="4.52" pageId="21" pageNumber="22" unit="mm" value="47.45" valueMax="49.7" valueMin="45.2">45.2−49.7 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFE5CE12FF5394EFDE38F8E1" blockId="21.[151,1437,762,2013]" pageId="21" pageNumber="22">
|
||||
Head and pronotum are dorsally yellow-brown with dark brown and pale spots (
|
||||
<figureCitation id="136616E4FFE5CE12FBCE94EFDEA9FAC5" box="[1114,1255,1267,1293]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 71−72</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FBCE94EFDEA9FAC5" box="[1114,1255,1267,1293]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Figs. 71−72</figureCitation>
|
||||
). The wings are yellowish with dark brown veins (
|
||||
<figureCitation id="136616E4FFE5CE12FD8D950BD8EAFAF9" box="[537,676,1303,1329]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 69–70</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FD8D950BD8EAFAF9" box="[537,676,1303,1329]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Figs. 69–70</figureCitation>
|
||||
). Antennae are dark brown, palpi yellow. The M-line is dark brown with sharp projections, between which there is a small yellow spot connected to a brownish spot on the clypeus; the lateral edges of the clypeus are light brown (
|
||||
<figureCitation id="136616E4FFE5CE12FD289543D909FAB1" box="[700,839,1375,1401]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 71–72</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FD289543D909FAB1" box="[700,839,1375,1401]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Figs. 71–72</figureCitation>
|
||||
). Frontal tentorial pits are darkened. The interocellar area is yellow and the tentorial pits are dark brown (
|
||||
<figureCitation id="136616E4FFE5CE12FD4E959FD92DFA55" box="[730,867,1411,1437]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 71‒72</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FD4E959FD92DFA55" box="[730,867,1411,1437]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Figs. 71‒72</figureCitation>
|
||||
). The posterior occipital fold looks like a thin, dark brown stripe (
|
||||
<figureCitation id="136616E4FFE5CE12FEA595BBDBCAFA09" box="[305,388,1447,1473]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Fig. 72</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FEA595BBDBCAFA09" box="[305,388,1447,1473]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Fig. 72</figureCitation>
|
||||
).A pair of transversely oblique dark bands extends from the base of the occiput to the posterior occipital fold. Pronotum brown, as wide as the head with eyes, slightly narrowing posteriorly; anterolateral margins obtuse, posterolateral margins rounded; medial stripe dark brown, widened to anterior and posterior margins; anterior pronotal margin darkened medially; lateral pronotal margins bordered by a dark brown stripe, widening posterolaterally; sometimes the lateral edges are bent down and not visible dorsally (
|
||||
<figureCitation id="136616E4FFE5CE12FBA5962BDEF4F999" box="[1073,1210,1591,1617]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 71−72</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FBA5962BDEF4F999" box="[1073,1210,1591,1617]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Figs. 71−72</figureCitation>
|
||||
). Pronotal rugosities are grayish-yellow, darkly contoured, and form an X-shaped pattern; in the posterior 1/3, pale rugosities in the form of narrow paired transverse short stripes almost reach the lateral pronotal margins (
|
||||
<figureCitation id="136616E4FFE5CE12FBA09663DEF1F951" box="[1076,1215,1663,1689]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 71−72</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FBA09663DEF1F951" box="[1076,1215,1663,1689]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Figs. 71−72</figureCitation>
|
||||
). The legs and cerci are yellow-brown. Femur is yellow-brown with a narrow, dark brown stripe along the distal edge. Tibia is darker than the femur, with a narrow dark brown stripe along the inner edge and a short oblique dark stripe along the outer edge at the base; the distal edge of the tibia has a thin dark brown stripe (
|
||||
<figureCitation id="136616E4FFE5CE12FC4F96F7DE29F8CD" box="[987,1127,1771,1797]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 69‒70</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FC4F96F7DE29F8CD" box="[987,1127,1771,1797]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Figs. 69‒70</figureCitation>
|
||||
). Anal gills are absent. The cerci are yellowish in the basal third of their length and brownish apically (
|
||||
<figureCitation id="136616E4FFE5CE12FC749713DE24F8E1" box="[992,1130,1807,1833]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." pageId="21" pageNumber="22">Figs. 69‒70</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FC749713DE24F8E1" box="[992,1130,1807,1833]" captionStart="FIGURES 69‒72" captionStartId="22.[151,264,1890,1914]" captionTargetBox="[164,1423,181,1866]" captionTargetId="figure-24@22.[164,1423,181,1866]" captionTargetPageId="22" captionText="FIGURES 69‒72. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="21" pageNumber="22">Figs. 69‒70</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFE5CE12FF53972FDFD3F815" blockId="21.[151,1437,762,2013]" pageId="21" pageNumber="22">
|
||||
<emphasis id="B929D673FFE5CE12FF53972FDB45F885" bold="true" box="[199,267,1843,1869]" pageId="21" pageNumber="22">Male.</emphasis>
|
||||
Tergum 8 is medially depressed with a central membranous furrow; posterolateral humps are covered with dense and thick setae (
|
||||
<figureCitation id="136616E4FFE5CE12FE41974BD865F8B9" box="[469,555,1879,1905]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="21" pageNumber="22">Fig. 73</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FE41974BD865F8B9" box="[469,555,1879,1905]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="21" pageNumber="22">Fig. 73</figureCitation>
|
||||
). Tergum 9 has a sclerotized mesal field that is butterfly-shaped; the width is greater than the length (
|
||||
<figureCitation id="136616E4FFE5CE12FE349767DBBAF85D" box="[416,500,1915,1941]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="21" pageNumber="22">Fig. 73</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FE349767DBBAF85D" box="[416,500,1915,1941]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="21" pageNumber="22">Fig. 73</figureCitation>
|
||||
). The mesal field bears sharp, heavy sclerotized teeth that merge into long and short longitudinal serrated ridges; the long serrated ridges are located in the middle of the mesal field (
|
||||
<figureCitation id="136616E4FFE5CE12FB4A9783DF7DF871" box="[1246,1331,1951,1977]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="21" pageNumber="22">Fig. 73</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FB4A9783DF7DF871" box="[1246,1331,1951,1977]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="21" pageNumber="22">Fig. 73</figureCitation>
|
||||
). Tergum 10 has finger-shaped hemitergal hooks (cleared) with a weak notch in the distal third (
|
||||
<figureCitation id="136616E4FFE5CE12FBE097DFDE87F815" box="[1140,1225,1987,2013]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="21" pageNumber="22">Fig. 73</figureCitation>
|
||||
<figureCitation id="136616E4FFE5CE12FBE097DFDE87F815" box="[1140,1225,1987,2013]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="21" pageNumber="22">Fig. 73</figureCitation>
|
||||
); apex is rounded.
|
||||
</paragraph>
|
||||
<caption id="DF225AE9FFE6CE11FF03977EDB02F82E" pageId="22" pageNumber="23" startId="22.[151,264,1890,1914]" targetBox="[164,1423,181,1866]" targetPageId="22" targetType="figure">
|
||||
<caption id="DF225AE9FFE6CE11FF03977EDB02F82E" ID-DOI="http://doi.org/10.5281/zenodo.13747780" ID-Zenodo-Dep="13747780" httpUri="https://zenodo.org/record/13747780/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,264,1890,1914]" targetBox="[164,1423,181,1866]" targetPageId="22" targetType="figure">
|
||||
<paragraph id="8BE20A61FFE6CE11FF03977EDB02F82E" blockId="22.[151,1437,1890,2022]" pageId="22" pageNumber="23">
|
||||
<emphasis id="B929D673FFE6CE11FF03977EDB1AF8B2" bold="true" box="[151,340,1890,1914]" pageId="22" pageNumber="23">FIGURES 69‒72.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFE6CE11FECE977ED85BF8B2" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[346,533,1890,1914]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="schapsugica" status="sp. nov.">
|
||||
|
|
@ -161,7 +162,7 @@ Tergum 8 is medially depressed with a central membranous furrow; posterolateral
|
|||
Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 69. Paratype. Female habitus, dorsal. 70. Holotype. Male, habitus, dorsal. 71. Holotype, male, head and pronotum, dorsal. 72. Paratype, female, head and pronotum, dorsal.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFE7CE10FF039678D8F0F920" pageId="23" pageNumber="24" startId="23.[151,264,1636,1660]" targetBox="[151,1436,181,1612]" targetPageId="23" targetType="figure">
|
||||
<caption id="DF225AE9FFE7CE10FF039678D8F0F920" ID-DOI="http://doi.org/10.5281/zenodo.13747784" ID-Zenodo-Dep="13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,264,1636,1660]" targetBox="[151,1436,181,1612]" targetPageId="23" targetType="figure">
|
||||
<paragraph id="8BE20A61FFE7CE10FF039678D8F0F920" blockId="23.[151,1437,1636,1768]" pageId="23" pageNumber="24">
|
||||
<emphasis id="B929D673FFE7CE10FF039678DB1AF9B4" bold="true" box="[151,340,1636,1660]" pageId="23" pageNumber="24">FIGURES 73‒76.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFE7CE10FECE9678D85BF9B4" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[346,533,1636,1660]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="schapsugica" status="sp. nov.">
|
||||
|
|
@ -175,7 +176,7 @@ Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District,
|
|||
76. Female abdominal tip, the subgenital plate, variations.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFE8CE1FFF03973EDBD4F826" pageId="24" pageNumber="25" startId="24.[151,264,1826,1850]" targetBox="[183,1405,181,1793]" targetPageId="24" targetType="figure">
|
||||
<caption id="DF225AE9FFE8CE1FFF03973EDBD4F826" ID-DOI="http://doi.org/10.5281/zenodo.13747788" ID-Zenodo-Dep="13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,264,1826,1850]" targetBox="[183,1405,181,1793]" targetPageId="24" targetType="figure">
|
||||
<paragraph id="8BE20A61FFE8CE1FFF03973EDBD4F826" blockId="24.[151,1437,1826,2030]" pageId="24" pageNumber="25">
|
||||
<emphasis id="B929D673FFE8CE1FFF03973EDB1AF8F2" bold="true" box="[151,340,1826,1850]" pageId="24" pageNumber="25">FIGURES 77‒81.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFE8CE1FFECE973ED85BF8F2" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[346,533,1826,1850]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="schapsugica" status="sp. nov.">
|
||||
|
|
@ -187,7 +188,7 @@ Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District,
|
|||
Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFE9CE1EFF039771DFCDF839" pageId="25" pageNumber="26" startId="25.[151,264,1901,1925]" targetBox="[158,1430,181,1877]" targetPageId="25" targetType="figure">
|
||||
<caption id="DF225AE9FFE9CE1EFF039771DFCDF839" ID-DOI="http://doi.org/10.5281/zenodo.13747790" ID-Zenodo-Dep="13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,264,1901,1925]" targetBox="[158,1430,181,1877]" targetPageId="25" targetType="figure">
|
||||
<paragraph id="8BE20A61FFE9CE1EFF039771DFCDF839" blockId="25.[151,1437,1901,2033]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B929D673FFE9CE1EFF039771DB1AF84D" bold="true" box="[151,340,1901,1925]" pageId="25" pageNumber="26">FIGURES 82‒89.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFE9CE1EFECE9771D85BF84D" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[346,533,1901,1925]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="schapsugica" status="sp. nov.">
|
||||
|
|
@ -199,7 +200,7 @@ Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District,
|
|||
Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFEACE1DFF03975CDB38F869" pageId="26" pageNumber="27" startId="26.[151,264,1856,1880]" targetBox="[151,1436,181,1833]" targetPageId="26" targetType="figure">
|
||||
<caption id="DF225AE9FFEACE1DFF03975CDB38F869" ID-DOI="http://doi.org/10.5281/zenodo.13747792" ID-Zenodo-Dep="13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,264,1856,1880]" targetBox="[151,1436,181,1833]" targetPageId="26" targetType="figure">
|
||||
<paragraph id="8BE20A61FFEACE1DFF03975CDB38F869" blockId="26.[151,1437,1856,1953]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B929D673FFEACE1DFF03975CDB1AF890" bold="true" box="[151,340,1856,1880]" pageId="26" pageNumber="27">FIGURES 90‒92.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFEACE1DFECE975DD85BF891" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[346,533,1857,1881]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="schapsugica" status="sp. nov.">
|
||||
|
|
@ -211,7 +212,7 @@ Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District,
|
|||
Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFEBCE1CFF0396F3D89FF8BC" pageId="27" pageNumber="28" startId="27.[151,264,1775,1799]" targetBox="[151,1436,187,1752]" targetPageId="27" targetType="figure">
|
||||
<caption id="DF225AE9FFEBCE1CFF0396F3D89FF8BC" ID-DOI="http://doi.org/10.5281/zenodo.13747794" ID-Zenodo-Dep="13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,264,1775,1799]" targetBox="[151,1436,187,1752]" targetPageId="27" targetType="figure">
|
||||
<paragraph id="8BE20A61FFEBCE1CFF0396F3D89FF8BC" blockId="27.[151,1437,1775,1908]" pageId="27" pageNumber="28">
|
||||
<emphasis id="B929D673FFEBCE1CFF0396F3DB1AF8C0" bold="true" box="[151,340,1775,1800]" pageId="27" pageNumber="28">FIGURES 93‒98.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFEBCE1CFECE96ECD85BF8C0" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[346,533,1776,1800]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="schapsugica" status="sp. nov.">
|
||||
|
|
@ -225,43 +226,43 @@ Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District,
|
|||
</caption>
|
||||
<paragraph id="8BE20A61FFECCE1BFF53908BDB7BFDF5" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||||
Sensilla basiconica are concentrated into an oval patch covering the inner side and top of the hook; the base of the hook is slightly rounded and covered with long setae (
|
||||
<figureCitation id="136616E4FFECCE1BFCCA90A7D9F0FF1D" box="[862,958,187,213]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="28" pageNumber="29">Figs. 73</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFCCA90A7D9F0FF1D" box="[862,958,187,213]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="28" pageNumber="29">Figs. 73</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFECCE1BFC5990A7D9A3FF1D" box="[973,1005,187,213]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." pageId="28" pageNumber="29">77</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFC5990A7D9A3FF1D" box="[973,1005,187,213]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="28" pageNumber="29">77</figureCitation>
|
||||
). In lateral view, the hemitergites are bent almost at a right angle and directed obliquely forward and downward towards the mesal field (
|
||||
<figureCitation id="136616E4FFECCE1BFA9390C3DF12FF31" box="[1287,1372,223,249]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="28" pageNumber="29">Fig. 74</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFA9390C3DF12FF31" box="[1287,1372,223,249]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="28" pageNumber="29">Fig. 74</figureCitation>
|
||||
). The artificially everted penis has a tube that is ca. 2.3 times the length of the sac. The tube is widened in basal half; dorsally and ventrally covered with densely arranged longitudinal rows of serrate sclerites, sometimes with small spines; the shape of sclerites changes from serrated and oval elongated to polygonal pattern in different parts of the tube (
|
||||
<figureCitation id="136616E4FFECCE1BFF429173DBD3FE41" box="[214,413,367,393]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." pageId="28" pageNumber="29">Figs. 79, A‒C, 81</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFF429173DBD3FE41" box="[214,413,367,393]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="28" pageNumber="29">Figs. 79, A‒C, 81</figureCitation>
|
||||
); the tube basally with paired lateral patches of rounded spine bases (
|
||||
<figureCitation id="136616E4FFECCE1BFB0F9173DF68FE41" box="[1179,1318,367,393]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." pageId="28" pageNumber="29">Figs. 79−81</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFB0F9173DF68FE41" box="[1179,1318,367,393]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="28" pageNumber="29">Figs. 79−81</figureCitation>
|
||||
); ventrally tube membranous in basal half. The sac is membranous and smooth, and at the base there is a pair of rounded lateral lobes dorsally and ventrally (
|
||||
<figureCitation id="136616E4FFECCE1BFE7091ABD83EFE19" box="[484,624,439,465]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." pageId="28" pageNumber="29">Figs. 77, 79</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFE7091ABD83EFE19" box="[484,624,439,465]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="28" pageNumber="29">Figs. 77, 79</figureCitation>
|
||||
). On the dorsal side, the sac has a tongue-shaped medial lobe, tapering towards the base of the sac; ventrally, the sac is deeply depressed, and the lobes are almost indistinguishable (
|
||||
<figureCitation id="136616E4FFECCE1BFAF091C7DAAEFDD1" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." pageId="28" pageNumber="29">Figs. 77, 79</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFAF091C7DAAEFDD1" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="28" pageNumber="29">Figs. 77, 79</figureCitation>
|
||||
). The apical brush of pointed brown spines is relatively long, widened dorsally, and slightly pointed ventrally (
|
||||
<figureCitation id="136616E4FFECCE1BFF0B923FDB67FDF5" box="[159,297,547,573]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." pageId="28" pageNumber="29">Figs. 79‒81</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFF0B923FDB67FDF5" box="[159,297,547,573]" captionStart="FIGURES 77‒81" captionStartId="24.[151,264,1826,1850]" captionTargetBox="[183,1405,181,1793]" captionTargetId="figure-24@24.[183,1405,181,1802]" captionTargetPageId="24" captionText="FIGURES 77‒81. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 77. Male. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 78. Abdominal tip, artificially everted penis, ventral, cleared. 79. Chaetotaxy of the penis, dorsal. A. Polygonal scales with small spines. B. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. D. Pointed brown spines in the apical brush. 80. Penis, ventral. 81. Penis, lateral." figureDoi="http://doi.org/10.5281/zenodo.13747788" httpUri="https://zenodo.org/record/13747788/files/figure.png" pageId="28" pageNumber="29">Figs. 79‒81</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFECCE1BFF53925BDFCFFD61" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B929D673FFECCE1BFF53925BDB6BFDA9" bold="true" box="[199,293,583,609]" pageId="28" pageNumber="29">Female.</emphasis>
|
||||
Sternum 8 has a subgenital plate occupying>1/2 of the sternum width and is slightly convergent in the medial plane. The subgenital plate has a small bilobed projection posteromedially; the lobes are small, rounded or slightly pointed, covered with rufous setae, and project onto sternum 9, covering 1/4 of its length. (
|
||||
<figureCitation id="136616E4FFECCE1BFB7F9293DF3BFD61" box="[1259,1397,654,681]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." pageId="28" pageNumber="29">Figs. 75‒76</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFB7F9293DF3BFD61" box="[1259,1397,654,681]" captionStart="FIGURES 73‒76" captionStartId="23.[151,264,1636,1660]" captionTargetBox="[151,1436,181,1612]" captionTargetId="figure-18@23.[151,1436,181,1612]" captionTargetPageId="23" captionText="FIGURES 73‒76. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, nameless small stream, the right tributary of the Kuapse River opposite Verkhnyaya Mamedka village. 73. Paratype. Male. Abdominal tip, mesal plate, hemitergal hooks, dorsal, cleared. 74. Abdominal tip, hemitergal hooks, lateral, cleared. 75‒76. Female abdominal tip, the subgenital plate, variations." figureDoi="http://doi.org/10.5281/zenodo.13747784" httpUri="https://zenodo.org/record/13747784/files/figure.png" pageId="28" pageNumber="29">Figs. 75‒76</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFECCE1BFF5392AFDE73FC49" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B929D673FFECCE1BFF5392AFDAB8FD05" bold="true" box="[199,246,691,717]" pageId="28" pageNumber="29">Egg</emphasis>
|
||||
. Round-oval, 488 x 322 µm (n=7), length with anchor (
|
||||
<figureCitation id="136616E4FFECCE1BFCFE92AFD9BAFD05" box="[874,1012,691,717]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="28" pageNumber="29">Figs. 82, 84</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFCFE92AFD9BAFD05" box="[874,1012,691,717]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="28" pageNumber="29">Figs. 82, 84</figureCitation>
|
||||
). Anchor is small, mushroom-shaped, and covers only the collar rim (
|
||||
<figureCitation id="136616E4FFECCE1BFE6B92CBD8C5FD39" box="[511,651,726,753]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="28" pageNumber="29">Figs. 82, 85</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFE6B92CBD8C5FD39" box="[511,651,726,753]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="28" pageNumber="29">Figs. 82, 85</figureCitation>
|
||||
). Collar wide, circumference with barely noticeable discernable ribs; rim smooth, with weak incisions (
|
||||
<figureCitation id="136616E4FFECCE1BFD8792E7D8D2FCDD" box="[531,668,763,789]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="28" pageNumber="29">Figs. 82‒86</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFD8792E7D8D2FCDD" box="[531,668,763,789]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="28" pageNumber="29">Figs. 82‒86</figureCitation>
|
||||
). Chorion is coarsely pitted; pits are regularly round in shape or oval (in micropylar row), relatively large and deep; and the chorion surface between pits is rough (
|
||||
<figureCitation id="136616E4FFECCE1BFB439303DFDEFCF1" box="[1239,1424,799,825]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="28" pageNumber="29">Figs. 84, 87‒88</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFB439303DFDEFCF1" box="[1239,1424,799,825]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="28" pageNumber="29">Figs. 84, 87‒88</figureCitation>
|
||||
). Micropylar row in the form of a broken line located close to the equator (
|
||||
<figureCitation id="136616E4FFECCE1BFC55935FDE06FC95" box="[961,1096,835,861]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="28" pageNumber="29">Figs. 82, 87</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFC55935FDE06FC95" box="[961,1096,835,861]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="28" pageNumber="29">Figs. 82, 87</figureCitation>
|
||||
). Micropyles in the pits are set not on flows but on the walls of the pits; orifices are raised with thin rims (
|
||||
<figureCitation id="136616E4FFECCE1BFC49937BDE7FFC49" box="[989,1073,871,897]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." pageId="28" pageNumber="29">Fig. 89</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFC49937BDE7FFC49" box="[989,1073,871,897]" captionStart="FIGURES 82‒89" captionStartId="25.[151,264,1901,1925]" captionTargetBox="[158,1430,181,1877]" captionTargetId="figure-18@25.[158,1430,181,1877]" captionTargetPageId="25" captionText="FIGURES 82‒89. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, the unnamed stream in the Krasnoaleksandrovskaya cave. Egg. 82. Appearance with mushroom anchor and micropylar row, lateral. 83. Collar, dorsal. 84. Appearance without anchor, lateral. 85. Posterior pole with anchor and collar, lateral. 86. Collar, dorsolateral. 87. Micropylar row. 88. Chorion structure. 89. Micropyles set on the walls of the pits; orifices raised with rims." figureDoi="http://doi.org/10.5281/zenodo.13747790" httpUri="https://zenodo.org/record/13747790/files/figure.png" pageId="28" pageNumber="29">Fig. 89</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFECCE1BFF539397DEE3F975" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||||
|
|
@ -269,45 +270,45 @@ Sternum 8 has a subgenital plate occupying>1/2 of the sternum width and is sl
|
|||
Female larva length 28.0−37.0 mm, length of antennae 11.5‒13.0 mm, and cerci 14.5–17.0 mm (n=9); male larva length 17.0 mm, antennae
|
||||
<quantity id="4CA5A784FFECCE1BFDDB93B2D8F9FC01" box="[591,695,942,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="28" pageNumber="29" unit="mm" value="11.5">11.5 mm</quantity>
|
||||
, and cerci 13.0 mm (n=1). The main color is olive-brown, with contrasting lighter patterns on the head, pronotum, mesonotum, and metanotum, and a less distinct pattern on the abdomen (
|
||||
<figureCitation id="136616E4FFECCE1BFE9B93EBDBD5FBD9" box="[271,411,1015,1041]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="28" pageNumber="29">Figs. 90‒91</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFE9B93EBDBD5FBD9" box="[271,411,1015,1041]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="28" pageNumber="29">Figs. 90‒91</figureCitation>
|
||||
). Body covered with dark clothing hairs; a dorsomesal band of erect silky setae is faintly visible (
|
||||
<figureCitation id="136616E4FFECCE1BFF659407DB35FBFD" box="[241,379,1051,1077]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="28" pageNumber="29">Figs. 90, 92</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFF659407DB35FBFD" box="[241,379,1051,1077]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="28" pageNumber="29">Figs. 90, 92</figureCitation>
|
||||
). Antennae, cerci, and sterna are pale. The head is large, and its width along the posterior edge does not exceed the width of the pronotum (
|
||||
<figureCitation id="136616E4FFECCE1BFD139423D892FB91" box="[647,732,1087,1113]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFD139423D892FB91" box="[647,732,1087,1113]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
). Clypeus olive-brownish until M-line, with pale tentorial pits; lateral clypeal margins darkened. M-line pale, a large transverse brown spot below M-line extending from the anterior ocellus to the posterior ocelli and lateral margins of the head; a triangular spot between the posterior ocelli and small triangular spots near the posterior ocelli laterally are pale; the posterior tentorial pits are internally light brown (
|
||||
<figureCitation id="136616E4FFECCE1BFF7994D3DB0EFB21" box="[237,320,1231,1257]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFF7994D3DB0EFB21" box="[237,320,1231,1257]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
). A short, relatively wide brown stripe with rectangular lateral edges below the epicranial suture does not reach the lateral edges of the head; the epicranial stem and occiput edge are brownish (
|
||||
<figureCitation id="136616E4FFECCE1BFB0494EFDEABFAC5" box="[1168,1253,1267,1293]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFB0494EFDEABFAC5" box="[1168,1253,1267,1293]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
). The transverse occipital fold and the fold running around the eye merge into a regularly curved line. Setation along the occipital fold extends only to behind the eye but not forward along the edge of the flat, expanded side of the head. No wavy section of the occipital fold is near the inner corner of the eye. The labrum is brownish with a small pale spot medially. Mandibles are heavily sclerotized (
|
||||
<figureCitation id="136616E4FFECCE1BFD07959FD8A6FA55" box="[659,744,1411,1437]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="28" pageNumber="29">Fig. 93</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFD07959FD8A6FA55" box="[659,744,1411,1437]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="28" pageNumber="29">Fig. 93</figureCitation>
|
||||
), with a rounded, dark outer margin. Left mandible with five sclerotized, bluntly pointed teeth; the apical tooth is elongated; the next three teeth are distinctly shorter; the last tooth is small and obtuse; the molar area bears two rows of short setae and a marginal row of long setae; the first row of short setae extends from the base of the first tooth, the second from the base of the third (
|
||||
<figureCitation id="136616E4FFECCE1BFB1E95F3DE93F9C1" box="[1162,1245,1519,1545]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="28" pageNumber="29">Fig. 93</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFB1E95F3DE93F9C1" box="[1162,1245,1519,1545]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="28" pageNumber="29">Fig. 93</figureCitation>
|
||||
). Right mandible with six bluntly pointed teeth; the last tooth is small, sometimes hidden by a marginal row of setae; the molar area bears a marginal row of long setae and one medial row of small setae stretching from the base of the first tooth to the mandibular base (
|
||||
<figureCitation id="136616E4FFECCE1BFE119647DB96F9BD" box="[389,472,1627,1653]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="28" pageNumber="29">Fig. 93</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFE119647DB96F9BD" box="[389,472,1627,1653]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="28" pageNumber="29">Fig. 93</figureCitation>
|
||||
). Lacinia bidentate has a large, darkly curved apical tooth and a much shorter subapical tooth (
|
||||
<figureCitation id="136616E4FFECCE1BFF759663DB78F951" box="[225,310,1663,1689]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="28" pageNumber="29">Fig. 94</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFF759663DB78F951" box="[225,310,1663,1689]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="28" pageNumber="29">Fig. 94</figureCitation>
|
||||
). The marginal fringe of setae along the inner lacinial margin is complete; one strong seta is visible near the subapical tooth. Galea length reaches almost half of the apical tooth length (
|
||||
<figureCitation id="136616E4FFECCE1BFBD896BFDEEEF975" box="[1100,1184,1699,1725]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="28" pageNumber="29">Fig. 94</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFBD896BFDEEEF975" box="[1100,1184,1699,1725]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="28" pageNumber="29">Fig. 94</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFECCE1AFF5396DBDBAAFE41" blockId="28.[151,1437,151,2013]" lastBlockId="29.[151,1437,151,393]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||||
Pronotum transverse, not wider than the head posteriorly, with obtusely rounded anterior corners; lateral margins and posterior angles are evenly rounded (
|
||||
<figureCitation id="136616E4FFECCE1BFDF996F7D8B7F8CD" box="[621,761,1771,1797]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="28" pageNumber="29">Figs. 90‒92</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFDF996F7D8B7F8CD" box="[621,761,1771,1797]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="28" pageNumber="29">Figs. 90‒92</figureCitation>
|
||||
). The pronotal fringe is complete with mostly short setae; a few relatively long setae are present on the posterolateral angles. Pronotum with thin brown stripes along lateral margins; paired semi-oval pale bands outline semi-oval dark brown bands within the lateral pronotal fields; brown bands are wider than pale bands (
|
||||
<figureCitation id="136616E4FFECCE1BFD92974BD817F8B9" box="[518,601,1879,1905]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFD92974BD817F8B9" box="[518,601,1879,1905]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
). The central pronotal field is pale with a light brown median band widened anteriorly and posteriorly; the band is surrounded by pale rugosities of an X-shaped pattern; paired vague S-shaped brown bands extend from the anterior to posterior pronotal margin; and the lateral fields are the same pale as the X-shaped medial pattern (
|
||||
<figureCitation id="136616E4FFECCE1BFE5097DFD852F815" box="[452,540,1987,2013]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
<figureCitation id="136616E4FFECCE1BFE5097DFD852F815" box="[452,540,1987,2013]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="28" pageNumber="29">Fig. 92</figureCitation>
|
||||
). Mesonotum and metanotum with similar Ψ-shaped brown pattern medially; paired wide dark brown bands limiting the wing pad bases; a narrow dark stripe runs along the lateral edge of the wing pads (
|
||||
<figureCitation id="136616E4FFEDCE1AFE8E90A7DBEAFF1D" box="[282,420,187,213]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="29" pageNumber="30">Figs. 90‒91</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFE8E90A7DBEAFF1D" box="[282,420,187,213]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="29" pageNumber="30">Figs. 90‒91</figureCitation>
|
||||
). The legs are light yellow-brown. The femur has a diffuse brownish spot in the distal half closer to the outer edge; the basal and distal edges are also brownish; the stout, hind femur is 3.2−3.5 times as long as wide (
|
||||
<figureCitation id="136616E4FFEDCE1AFF68911FDB14FED5" box="[252,346,259,285]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="29" pageNumber="30">Figs. 91</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFF68911FDB14FED5" box="[252,346,259,285]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="29" pageNumber="30">Figs. 91</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFEDCE1AFEF2911EDBC8FED5" box="[358,390,258,285]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="29" pageNumber="30">95</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFEF2911EDBC8FED5" box="[358,390,258,285]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="29" pageNumber="30">95</figureCitation>
|
||||
). The femur and tibia are densely covered with dark clothing hairs bearing stout, short, acute red bristles along the inner and outer edges and a fringe of very dense, long, silky hairs along the outer margins (
|
||||
<figureCitation id="136616E4FFEDCE1AFAFB913BDAF9FEAD" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="29" pageNumber="30">Fig. 95</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFAFB913BDAF9FEAD" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="29" pageNumber="30">Fig. 95</figureCitation>
|
||||
). Sparse red bristles also covered their surface irregularly. Tibia have diffuse, narrow, dark brown bands on the inner and outer edges basally.
|
||||
</paragraph>
|
||||
<caption id="DF225AE9FFEDCE1AFF039596D9F4FA22" pageId="29" pageNumber="30" startId="29.[151,264,1418,1442]" targetBox="[151,1436,417,1394]" targetPageId="29" targetType="figure">
|
||||
<caption id="DF225AE9FFEDCE1AFF039596D9F4FA22" ID-DOI="http://doi.org/10.5281/zenodo.13747796" ID-Zenodo-Dep="13747796" httpUri="https://zenodo.org/record/13747796/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,264,1418,1442]" targetBox="[151,1436,417,1394]" targetPageId="29" targetType="figure">
|
||||
<paragraph id="8BE20A61FFEDCE1AFF039596D9F4FA22" blockId="29.[151,1436,1418,1514]" pageId="29" pageNumber="30">
|
||||
<emphasis id="B929D673FFEDCE1AFF039596DB2EFA6A" bold="true" box="[151,352,1418,1442]" pageId="29" pageNumber="30">FIGURES 99‒101.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFEDCE1AFEF19596D86EFA6A" authorityName="Teslenko & Palatov & Semenchenko" authorityYear="2024" box="[357,544,1418,1442]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="schapsugica" status="sp. nov.">
|
||||
|
|
@ -321,23 +322,23 @@ Habitat. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky D
|
|||
</caption>
|
||||
<paragraph id="8BE20A61FFEDCE1AFF539620DEC4F82A" blockId="29.[151,1436,1596,2019]" pageId="29" pageNumber="30">
|
||||
The abdomen is dark brown, with variation in banding between female and male larvae (
|
||||
<figureCitation id="136616E4FFEDCE1AFB309620DF60F99E" box="[1188,1326,1596,1622]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="29" pageNumber="30">Figs. 90‒91</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFB309620DF60F99E" box="[1188,1326,1596,1622]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="29" pageNumber="30">Figs. 90‒91</figureCitation>
|
||||
). On each abdominal tergum of the male larva are two pale transverse spots on the sides of the medial brown line; along the lateral tergal margins, there is another paired pale spot, not always clear (
|
||||
<figureCitation id="136616E4FFEDCE1AFC4C9698DE62F956" box="[984,1068,1668,1694]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="29" pageNumber="30">Fig. 91</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFC4C9698DE62F956" box="[984,1068,1668,1694]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="29" pageNumber="30">Fig. 91</figureCitation>
|
||||
). On the last tergum of the male larva, the pale spots merge with each other (
|
||||
<figureCitation id="136616E4FFEDCE1AFD1796B4D899F90A" box="[643,727,1704,1730]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="29" pageNumber="30">Fig. 91</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFD1796B4D899F90A" box="[643,727,1704,1730]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="29" pageNumber="30">Fig. 91</figureCitation>
|
||||
). The posterior terga margins bear acute bristles that are mostly short, with one or two relatively long, thin setae. Anal gills are very small (
|
||||
<figureCitation id="136616E4FFEDCE1AFC4C96D0DE62F92E" box="[984,1068,1740,1766]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="29" pageNumber="30">Fig. 90</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFC4C96D0DE62F92E" box="[984,1068,1740,1766]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="29" pageNumber="30">Fig. 90</figureCitation>
|
||||
). Cerci are uncolored, yellowish, and have about 39 segments with striking rings of dark spines (
|
||||
<figureCitation id="136616E4FFEDCE1AFCDA96ECD9EFF8C2" box="[846,929,1776,1802]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." pageId="29" pageNumber="30">Fig. 90</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFCDA96ECD9EFF8C2" box="[846,929,1776,1802]" captionStart="FIGURES 90‒92" captionStartId="26.[151,264,1856,1880]" captionTargetBox="[151,1436,181,1833]" captionTargetId="figure-24@26.[151,1436,181,1833]" captionTargetPageId="26" captionText="FIGURES 90‒92. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 90. Female, habitus, dorsal. 91. Male, habitus, dorsal. 92. Head and pronotum, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747792" httpUri="https://zenodo.org/record/13747792/files/figure.png" pageId="29" pageNumber="30">Fig. 90</figureCitation>
|
||||
). Fine pilosity is present between dark spines, tightly pressed to the segment surface; their length is 1/2 the length of the basal and middle cercal segments; on the apical cercal segments, the length of the thin hairs is slightly>1/3 of the length of the segment (
|
||||
<figureCitation id="136616E4FFEDCE1AFB519724DF01F89A" box="[1221,1359,1848,1874]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="29" pageNumber="30">Figs. 96‒98</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFB519724DF01F89A" box="[1221,1359,1848,1874]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="29" pageNumber="30">Figs. 96‒98</figureCitation>
|
||||
). Long silky colorless hairs are present in the circlet in tufts (at least
|
||||
<quantity id="4CA5A784FFEDCE1AFCDF9740D9CAF8BE" box="[843,900,1884,1911]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="29" pageNumber="30" unit="in" value="15.0">15 in</quantity>
|
||||
each on the basal and middle cercal segments), forming a dorsal longitudinal fringe, tightly pressed to the segments; the length and density of long silky hairs decrease towards the cercal apex (
|
||||
<figureCitation id="136616E4FFEDCE1AFD8597B8D8D4F876" box="[529,666,1956,1982]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="29" pageNumber="30">Figs. 96–98</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFD8597B8D8D4F876" box="[529,666,1956,1982]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="29" pageNumber="30">Figs. 96–98</figureCitation>
|
||||
). On the middle and apical cercal segments, short and thin intercalary setae are visible; their length does not exceed the length of fine circlet hairs (
|
||||
<figureCitation id="136616E4FFEDCE1AFC6097D4DE33F82A" box="[1012,1149,1992,2018]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." pageId="29" pageNumber="30">Figs. 97‒98</figureCitation>
|
||||
<figureCitation id="136616E4FFEDCE1AFC6097D4DE33F82A" box="[1012,1149,1992,2018]" captionStart="FIGURES 93‒98" captionStartId="27.[151,264,1775,1799]" captionTargetBox="[151,1436,187,1752]" captionTargetId="figure-18@27.[151,1436,181,1752]" captionTargetPageId="27" captionText="FIGURES 93‒98. Perla schapsugica sp. nov. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. Larva. 93. Left and right mandibles, ventral. 94. Left lacinia, galea, maxillary palpus, ventral. 95. Right hind leg, dorsal. 96‒98. Chaetotaxy of the right cercus, lateral: 96. Basal cercal segments. 97. Middle cercal segments. 98. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747794" httpUri="https://zenodo.org/record/13747794/files/figure.png" pageId="29" pageNumber="30">Figs. 97‒98</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -561,7 +562,7 @@ and
|
|||
<emphasis id="B929D673FFEECE19FBFB92E7DE92FCDC" box="[1135,1244,763,788]" italics="true" pageId="30" pageNumber="31">P. persica</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136616E4FFEECE19FB7892E7DF00FCDD" box="[1260,1358,763,789]" captionStart="FIGURE 160" captionStartId="48.[151,250,1826,1850]" captionTargetBox="[152,1430,566,1798]" captionTargetId="figure-206@48.[151,1435,561,1803]" captionTargetPageId="48" captionText="FIGURE 160. Ultrametric Bayesian inference (BI) tree inferred from the cytochrome c oxidase I (COI) gene fragment and the results of species delineation of the genus Perla Geoffroy, 1762, and out-group Kamimuria coreana. Bayesian posterior probabilities (> 0.7) are noted above the tree nodes. Vertical bars correspond to molecular operational taxonomic units by ASAP, mPTP, GMYC, and BINs of BOLD. The color of the specimens indicates the development stage of stoneflies in accordance with the legend, while the sequences obtained in this study are in bold." pageId="30" pageNumber="31">Fig. 160</figureCitation>
|
||||
<figureCitation id="136616E4FFEECE19FB7892E7DF00FCDD" box="[1260,1358,763,789]" captionStart="FIGURE 160" captionStartId="48.[151,250,1826,1850]" captionTargetBox="[152,1430,566,1798]" captionTargetId="figure-206@48.[151,1435,561,1803]" captionTargetPageId="48" captionText="FIGURE 160. Ultrametric Bayesian inference (BI) tree inferred from the cytochrome c oxidase I (COI) gene fragment and the results of species delineation of the genus Perla Geoffroy, 1762, and out-group Kamimuria coreana. Bayesian posterior probabilities (> 0.7) are noted above the tree nodes. Vertical bars correspond to molecular operational taxonomic units by ASAP, mPTP, GMYC, and BINs of BOLD. The color of the specimens indicates the development stage of stoneflies in accordance with the legend, while the sequences obtained in this study are in bold." figureDoi="http://doi.org/10.5281/zenodo.13747826" httpUri="https://zenodo.org/record/13747826/files/figure.png" pageId="30" pageNumber="31">Fig. 160</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -590,7 +591,7 @@ occurs in the small karst streams of the Black Sea coast of the Caucasus from Tu
|
|||
,
|
||||
<collectingCountry id="F34A4AF1FFEECE19FD6A93B3D907FC01" box="[766,841,943,969]" name="Russia" pageId="30" pageNumber="31">Russia</collectingCountry>
|
||||
(
|
||||
<figureCitation id="136616E4FFEECE19FCCC93B3D9F7FC01" box="[856,953,942,969]" captionStart="FIGURE 159" captionStartId="47.[151,250,1780,1804]" captionTargetBox="[158,1430,846,1757]" captionTargetId="figure-40@47.[158,1430,846,1757]" captionTargetPageId="47" captionText="FIGURE 159. The map of the Perla species distribution in Caucasus according to our data. Perla caicasica—pink, P. persica— green, P. kiritshenkoi—blue, P. schapsugica sp. nov.—white, P. pallida—yellow, and P. palatovi sp. nov.—red." pageId="30" pageNumber="31">Fig. 159</figureCitation>
|
||||
<figureCitation id="136616E4FFEECE19FCCC93B3D9F7FC01" box="[856,953,942,969]" captionStart="FIGURE 159" captionStartId="47.[151,250,1780,1804]" captionTargetBox="[158,1430,846,1757]" captionTargetId="figure-40@47.[158,1430,846,1757]" captionTargetPageId="47" captionText="FIGURE 159. The map of the Perla species distribution in Caucasus according to our data. Perla caicasica—pink, P. persica— green, P. kiritshenkoi—blue, P. schapsugica sp. nov.—white, P. pallida—yellow, and P. palatovi sp. nov.—red." figureDoi="http://doi.org/10.5281/zenodo.13747824" httpUri="https://zenodo.org/record/13747824/files/figure.png" pageId="30" pageNumber="31">Fig. 159</figureCitation>
|
||||
). The material was collected at the outlet of a small stream from a cave, quickly falling into a karst,
|
||||
<quantity id="4CA5A784FFEECE19FC9B93CED9C1FC25" box="[783,911,978,1005]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="30" pageNumber="31" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">100–150 m</quantity>
|
||||
long and 2.0–
|
||||
|
|
@ -598,7 +599,7 @@ long and 2.0–
|
|||
wide, with a current ≤
|
||||
<quantity id="4CA5A784FFEECE19FAED93CFDAFEFBD9" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="31" unit="m" value="0.6">0.6 m</quantity>
|
||||
/s. The bottom was enclosed with large stones covered with moss (
|
||||
<figureCitation id="136616E4FFEECE19FC0D93EBDE7CFBD9" box="[921,1074,1015,1041]" captionStart="FIGURES 99‒101" captionStartId="29.[151,264,1418,1442]" captionTargetBox="[151,1436,417,1394]" captionTargetId="figure-146@29.[151,1436,417,1394]" captionTargetPageId="29" captionText="FIGURES 99‒101. Perla schapsugica sp. nov. Habitat. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. 99. The Krasnoaleksandrovskaya cave, exit. 100. Outlet of a small stream from Krasnoaleksandrovskaya cave. 101. A shallow beech ravine." pageId="30" pageNumber="31">Figs. 99−101</figureCitation>
|
||||
<figureCitation id="136616E4FFEECE19FC0D93EBDE7CFBD9" box="[921,1074,1015,1041]" captionStart="FIGURES 99‒101" captionStartId="29.[151,264,1418,1442]" captionTargetBox="[151,1436,417,1394]" captionTargetId="figure-146@29.[151,1436,417,1394]" captionTargetPageId="29" captionText="FIGURES 99‒101. Perla schapsugica sp. nov. Habitat. Caucasus, Russia, Krasnodar Kray, urban District of Sochi, Lazarevsky District, unnamed stream in the Krasnoaleksandrovskaya cave. 99. The Krasnoaleksandrovskaya cave, exit. 100. Outlet of a small stream from Krasnoaleksandrovskaya cave. 101. A shallow beech ravine." figureDoi="http://doi.org/10.5281/zenodo.13747796" httpUri="https://zenodo.org/record/13747796/files/figure.png" pageId="30" pageNumber="31">Figs. 99−101</figureCitation>
|
||||
). This species was also found in small rivers and streams on the Black Sea coast of the Tuapse District, flowing in low-mountain beech forest valleys up to
|
||||
<quantity id="4CA5A784FFEECE19FF429423DB53FB91" box="[214,285,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="31" unit="m" value="200.0">200 m</quantity>
|
||||
above sea level. The stream has a width of 1.5−3.0 m, a depth varying from
|
||||
|
|
|
|||
|
|
@ -1,60 +1,61 @@
|
|||
<document id="55E95CC1B24D57B60506752DDE37A5DD" ID-DOI="10.11646/zootaxa.5507.1.1" ID-ISSN="1175-5326" ID-ZooBank="065ECECA-5F0B-47BE-82FC-6C1F68B316FD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726055721707" checkinUser="plazi" docAuthor="Teslenko, Valentina A., Palatov, Dmitry M. & Semenchenko, Alexander A." docDate="2024" docId="03F4BB77FFEECE22FF0394D2DB51F929" docLanguage="en" docName="zootaxa.5507.1.1.pdf" docOrigin="Zootaxa 5507 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Perla pallida Guerin-Meneville 1843" docType="treatment" docVersion="1" lastPageNumber="38" masterDocId="FFCDC30FFFF0CE07FF94901CDA4EFFC8" masterDocTitle="Overview of the Caucasian Perla Geoffroy, 1762 (Plecoptera: Perlidae) based on morphological and molecular data with description of two new species" masterLastPageNumber="56" masterPageNumber="1" pageNumber="31" updateTime="1726055957189" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:mods id="DB9280CA4DD1F6082C4DE568B26CAF7A" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="8756016F93E029144D7D9E75F4CDFDCF">
|
||||
<mods:title id="E4C4FFEBD3222DB6793D0E5B16C5E84D">Overview of the Caucasian Perla Geoffroy, 1762 (Plecoptera: Perlidae) based on morphological and molecular data with description of two new species</mods:title>
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<mods:title id="D4A97D4537E3123A5F6DE11EC92CE8A2">Overview of the Caucasian Perla Geoffroy, 1762 (Plecoptera: Perlidae) based on morphological and molecular data with description of two new species</mods:title>
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<figureCitation id="136616E4FFEECE19FF0394EFDB79FAC5" box="[151,311,1267,1293]" captionStart-0="FIGURES 102‒107" captionStart-1="FIGURES 108‒111" captionStart-2="FIGURES 112‒119" captionStart-3="FIGURES 120‒121" captionStart-4="FIGURES 122‒127" captionStartId-0="31.[151,264,1888,1912]" captionStartId-1="32.[151,264,1864,1888]" captionStartId-2="33.[151,264,1837,1861]" captionStartId-3="34.[151,264,1516,1540]" captionStartId-4="35.[151,264,1788,1812]" captionTargetBox-0="[151,1436,181,1864]" captionTargetBox-1="[170,1416,181,1840]" captionTargetBox-2="[183,1404,181,1813]" captionTargetBox-3="[151,1436,181,1493]" captionTargetBox-4="[151,1436,181,1764]" captionTargetId-0="figure-18@31.[151,1436,181,1864]" captionTargetId-1="figure-24@32.[170,1416,181,1840]" captionTargetId-2="figure-18@33.[183,1404,181,1813]" captionTargetId-3="figure-24@34.[151,1436,181,1493]" captionTargetId-4="figure-18@35.[151,1436,181,1764]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="34" captionTargetPageId-4="35" captionText-0="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." captionText-1="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." captionText-2="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." captionText-3="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." captionText-4="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." pageId="30" pageNumber="31">Figs. 102‒127</figureCitation>
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<figureCitation id="136616E4FFEECE19FF0394EFDB79FAC5" box="[151,311,1267,1293]" captionStart-0="FIGURES 102‒107" captionStart-1="FIGURES 108‒111" captionStart-2="FIGURES 112‒119" captionStart-3="FIGURES 120‒121" captionStart-4="FIGURES 122‒127" captionStartId-0="31.[151,264,1888,1912]" captionStartId-1="32.[151,264,1864,1888]" captionStartId-2="33.[151,264,1837,1861]" captionStartId-3="34.[151,264,1516,1540]" captionStartId-4="35.[151,264,1788,1812]" captionTargetBox-0="[151,1436,181,1864]" captionTargetBox-1="[170,1416,181,1840]" captionTargetBox-2="[183,1404,181,1813]" captionTargetBox-3="[151,1436,181,1493]" captionTargetBox-4="[151,1436,181,1764]" captionTargetId-0="figure-18@31.[151,1436,181,1864]" captionTargetId-1="figure-24@32.[170,1416,181,1840]" captionTargetId-2="figure-18@33.[183,1404,181,1813]" captionTargetId-3="figure-24@34.[151,1436,181,1493]" captionTargetId-4="figure-18@35.[151,1436,181,1764]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="34" captionTargetPageId-4="35" captionText-0="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." captionText-1="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." captionText-2="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." captionText-3="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." captionText-4="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." figureDoi-0="http://doi.org/10.5281/zenodo.13747798" figureDoi-1="http://doi.org/10.5281/zenodo.13747800" figureDoi-2="http://doi.org/10.5281/zenodo.13747802" figureDoi-3="http://doi.org/10.5281/zenodo.13747804" figureDoi-4="http://doi.org/10.5281/zenodo.13747806" httpUri-0="https://zenodo.org/record/13747798/files/figure.png" httpUri-1="https://zenodo.org/record/13747800/files/figure.png" httpUri-2="https://zenodo.org/record/13747802/files/figure.png" httpUri-3="https://zenodo.org/record/13747804/files/figure.png" httpUri-4="https://zenodo.org/record/13747806/files/figure.png" pageId="30" pageNumber="31">Figs. 102‒127</figureCitation>
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<bibRefCitation id="EFCC7790FFEECE19FF03952BDBB9FA87" author="Guerin-Meneville, F. E." box="[151,503,1335,1359]" pageId="30" pageNumber="31" pagination="393 - 395" refId="ref24390" refString="Guerin-Meneville, F. E. (1843) Genre Perle. In: Guerin-Meneville, F. E. (Ed.), Iconographie du Regne Animal de G. Cuvier. Insectes, 1843, pp. 393 - 395." type="journal article" year="1843">Guérin-Méneville, 1843: 393−394</bibRefCitation>
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@ -148,24 +149,24 @@ to be a species complex of three or more species or subspecies distributed in th
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|
||||
<emphasis id="B929D673FFEECE19FF53972ADB0CF898" bold="true" box="[199,322,1846,1872]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
|
||||
Mesal field on tergum 9 of male slightly expanded anteriorly and posteriorly, width slightly greater than length; longitudinal serrated ridges (10−12) closer to anterior and lateral margins; posterior margin avoids serrated ridge rows (
|
||||
<figureCitation id="136616E4FFEECE19FEEF9763DBABF851" box="[379,485,1919,1945]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="30" pageNumber="31">Figs. 106</figureCitation>
|
||||
<figureCitation id="136616E4FFEECE19FEEF9763DBABF851" box="[379,485,1919,1945]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="30" pageNumber="31">Figs. 106</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFEECE19FE659763D851F851" box="[497,543,1919,1945]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="30" pageNumber="31">108</figureCitation>
|
||||
<figureCitation id="136616E4FFEECE19FE659763D851F851" box="[497,543,1919,1945]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="30" pageNumber="31">108</figureCitation>
|
||||
). Hemitergites are strong, narrowed towards the rounded apex, and hook wide at the base with a weak notch distally (
|
||||
<figureCitation id="136616E4FFEECE19FDBE97BFD882F875" box="[554,716,1955,1981]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="30" pageNumber="31">Figs. 104, 106</figureCitation>
|
||||
<figureCitation id="136616E4FFEECE19FDBE97BFD882F875" box="[554,716,1955,1981]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="30" pageNumber="31">Figs. 104, 106</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFEECE19FD4C97BFD948F875" box="[728,774,1955,1981]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="30" pageNumber="31">108</figureCitation>
|
||||
<figureCitation id="136616E4FFEECE19FD4C97BFD948F875" box="[728,774,1955,1981]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="30" pageNumber="31">108</figureCitation>
|
||||
). The tube of the penis is 1.6 times the length of the sac and has a constriction before the sac; the sac is large, wider than the tube, and covered with a patch of coarse triangular spines, forming a wide, loose triangular apical brush; the spines of the brush along the lateral edges of the sac are absent; ventrally, the brush end is triangular; dorsally, the edge of the apical brush is semi-rounded (
|
||||
<figureCitation id="136616E4FFD2CE25FF0B96C0DB06F93E" box="[159,328,1756,1782]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="34" pageNumber="35">Figs. 108−111</figureCitation>
|
||||
<figureCitation id="136616E4FFD2CE25FF0B96C0DB06F93E" box="[159,328,1756,1782]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="34" pageNumber="35">Figs. 108−111</figureCitation>
|
||||
). Egg oval; collar with flanged rim that is incised and distinctly wider than the neck; the collar sides covered ribs; chorion smooth, covered with follicle hexagonal cell impressions, limiting at the corners by six tiny round punctuations and by six small, slit-shaped punctuations located between them; cells have boundaries, a central depression, and one pore on the floor; pore and hexagonal punctuation are different in size (
|
||||
<figureCitation id="136616E4FFD2CE25FB799754DFDEF8AA" box="[1261,1424,1864,1890]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="34" pageNumber="35">Figs. 112‒119</figureCitation>
|
||||
<figureCitation id="136616E4FFD2CE25FB799754DFDEF8AA" box="[1261,1424,1864,1890]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="34" pageNumber="35">Figs. 112‒119</figureCitation>
|
||||
). The micropylar orifices lack rims (
|
||||
<figureCitation id="136616E4FFD2CE25FDBF9770D89AF84E" box="[555,724,1900,1926]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="34" pageNumber="35">Figs. 117−118</figureCitation>
|
||||
<figureCitation id="136616E4FFD2CE25FDBF9770D89AF84E" box="[555,724,1900,1926]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="34" pageNumber="35">Figs. 117−118</figureCitation>
|
||||
). Head of larvae with a large trapezoidal pale spot expanding from the anterior ocellus to the labrum base; the lateral edges of the spot with two pair pointed projections; paired projections near the labrum wide; a thin pale M-line delimiting the spot from below; the epicranial sutures brown (
|
||||
<figureCitation id="136616E4FFD2CE25FF0B97C4DB08F83A" box="[159,326,2008,2034]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." pageId="34" pageNumber="35">Figs. 120−121</figureCitation>
|
||||
<figureCitation id="136616E4FFD2CE25FF0B97C4DB08F83A" box="[159,326,2008,2034]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747804" httpUri="https://zenodo.org/record/13747804/files/figure.png" pageId="34" pageNumber="35">Figs. 120−121</figureCitation>
|
||||
). Anal gills are absent.
|
||||
</paragraph>
|
||||
<caption id="DF225AE9FFEFCE18FF03977CD895F82C" pageId="31" pageNumber="32" startId="31.[151,264,1888,1912]" targetBox="[151,1436,181,1864]" targetPageId="31" targetType="figure">
|
||||
<caption id="DF225AE9FFEFCE18FF03977CD895F82C" ID-DOI="http://doi.org/10.5281/zenodo.13747798" ID-Zenodo-Dep="13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,264,1888,1912]" targetBox="[151,1436,181,1864]" targetPageId="31" targetType="figure">
|
||||
<paragraph id="8BE20A61FFEFCE18FF03977CD895F82C" blockId="31.[151,1437,1888,2020]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B929D673FFEFCE18FF03977CDB3EF8B0" bold="true" box="[151,368,1888,1912]" pageId="31" pageNumber="32">FIGURES 102‒107.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFEFCE18FEED977CD94BF8B0" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[377,773,1888,1912]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pallida">
|
||||
|
|
@ -175,7 +176,7 @@ Mesal field on tergum 9 of male slightly expanded anteriorly and posteriorly, wi
|
|||
. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFD0CE27FF039754DBA8F838" pageId="32" pageNumber="33" startId="32.[151,264,1864,1888]" targetBox="[170,1416,181,1840]" targetPageId="32" targetType="figure">
|
||||
<caption id="DF225AE9FFD0CE27FF039754DBA8F838" ID-DOI="http://doi.org/10.5281/zenodo.13747800" ID-Zenodo-Dep="13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,264,1864,1888]" targetBox="[170,1416,181,1840]" targetPageId="32" targetType="figure">
|
||||
<paragraph id="8BE20A61FFD0CE27FF039754DBA8F838" blockId="32.[151,1437,1864,2032]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B929D673FFD0CE27FF039754DB23F8A8" bold="true" box="[151,365,1864,1888]" pageId="32" pageNumber="33">FIGURES 108‒111.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFD0CE27FEE09754D8B1F8A8" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[372,767,1864,1888]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pallida">
|
||||
|
|
@ -185,7 +186,7 @@ Mesal field on tergum 9 of male slightly expanded anteriorly and posteriorly, wi
|
|||
. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFD1CE26FF039731D8A5F831" pageId="33" pageNumber="34" startId="33.[151,264,1837,1861]" targetBox="[183,1404,181,1813]" targetPageId="33" targetType="figure">
|
||||
<caption id="DF225AE9FFD1CE26FF039731D8A5F831" ID-DOI="http://doi.org/10.5281/zenodo.13747802" ID-Zenodo-Dep="13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,264,1837,1861]" targetBox="[183,1404,181,1813]" targetPageId="33" targetType="figure">
|
||||
<paragraph id="8BE20A61FFD1CE26FF039731D8A5F831" blockId="33.[151,1437,1837,2041]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B929D673FFD1CE26FF039731DB22F88D" bold="true" box="[151,364,1837,1861]" pageId="33" pageNumber="34">FIGURES 112‒119.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFD1CE26FEE09731D8B1F88D" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[372,767,1837,1861]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pallida">
|
||||
|
|
@ -197,7 +198,7 @@ Mesal field on tergum 9 of male slightly expanded anteriorly and posteriorly, wi
|
|||
118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFD2CE25FF0395F0D8EEF985" pageId="34" pageNumber="35" startId="34.[151,264,1516,1540]" targetBox="[151,1436,181,1493]" targetPageId="34" targetType="figure">
|
||||
<caption id="DF225AE9FFD2CE25FF0395F0D8EEF985" ID-DOI="http://doi.org/10.5281/zenodo.13747804" ID-Zenodo-Dep="13747804" httpUri="https://zenodo.org/record/13747804/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,264,1516,1540]" targetBox="[151,1436,181,1493]" targetPageId="34" targetType="figure">
|
||||
<paragraph id="8BE20A61FFD2CE25FF0395F0D8EEF985" blockId="34.[151,1437,1516,1613]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B929D673FFD2CE25FF0395F0DB21F9CD" bold="true" box="[151,367,1516,1541]" pageId="34" pageNumber="35">FIGURES 120‒121.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFD2CE25FEE395F1D94FF9CD" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[375,769,1517,1541]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pallida">
|
||||
|
|
@ -207,7 +208,7 @@ Mesal field on tergum 9 of male slightly expanded anteriorly and posteriorly, wi
|
|||
. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFD3CE24FF0396E0DFCBF848" pageId="35" pageNumber="36" startId="35.[151,264,1788,1812]" targetBox="[151,1436,181,1764]" targetPageId="35" targetType="figure">
|
||||
<caption id="DF225AE9FFD3CE24FF0396E0DFCBF848" ID-DOI="http://doi.org/10.5281/zenodo.13747806" ID-Zenodo-Dep="13747806" httpUri="https://zenodo.org/record/13747806/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,264,1788,1812]" targetBox="[151,1436,181,1764]" targetPageId="35" targetType="figure">
|
||||
<paragraph id="8BE20A61FFD3CE24FF0396E0DFCBF848" blockId="35.[151,1437,1788,1920]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B929D673FFD3CE24FF0396E0DB21F8DC" bold="true" box="[151,367,1788,1812]" pageId="35" pageNumber="36">FIGURES 122‒127.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFD3CE24FEE396E0D94FF8DC" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[375,769,1788,1812]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pallida">
|
||||
|
|
@ -242,66 +243,66 @@ The coloration, external structures of the male and female genitalia, eggs, and
|
|||
<paragraph id="8BE20A61FFD4CE23FF539156DFC5FDA9" blockId="36.[151,1437,150,2013]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B929D673FFD4CE23FF539156DB3BFEAC" bold="true" box="[199,373,330,356]" pageId="36" pageNumber="37">Adult habitus.</emphasis>
|
||||
The main body color is yellow-brown (
|
||||
<figureCitation id="136616E4FFD4CE23FCD39156D9E3FEAC" box="[839,941,330,356]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Fig. 102</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FCD39156D9E3FEAC" box="[839,941,330,356]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Fig. 102</figureCitation>
|
||||
). The head is mostly yellow; the M-line is yellow and moderately defined (
|
||||
<figureCitation id="136616E4FFD4CE23FD979172D8E7FE41" box="[515,681,366,393]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 102, 105</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FD979172D8E7FE41" box="[515,681,366,393]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 102, 105</figureCitation>
|
||||
). The interocellar area bears a dark brown spot reaching the M-line and directed to compound eyes above yellow posterior tentorial pits; the posterior edge between the lateral ocelli with a solid pale line (
|
||||
<figureCitation id="136616E4FFD4CE23FE1B91AADBB2FE19" box="[399,508,438,465]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 105</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FE1B91AADBB2FE19" box="[399,508,438,465]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 105</figureCitation>
|
||||
). The female head is darker due to a larger dark brown spot in the interocellar area, with more distinct lateral edges approaching the M-line and the posterior tentorial pits closely but not going beyond them (
|
||||
<figureCitation id="136616E4FFD4CE23FF4991E2DB71FDD1" box="[221,319,510,537]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Fig. 105</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FF4991E2DB71FDD1" box="[221,319,510,537]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Fig. 105</figureCitation>
|
||||
). A triangular light brown spot, widening towards the anterior margin of the clypeus, is in front of the M-line and anterior ocellus; the antenna is brown (
|
||||
<figureCitation id="136616E4FFD4CE23FD5E923ED9E8FDF5" box="[714,934,546,573]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 102–103, 105</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FD5E923ED9E8FDF5" box="[714,934,546,573]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 102–103, 105</figureCitation>
|
||||
). Occiput bears a wide, V-shaped brown spot in the area of the epicranial suture and stem, the spot is darker and more pronounced on females (
|
||||
<figureCitation id="136616E4FFD4CE23FB4D925ADF30FDA9" box="[1241,1406,582,609]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 103, 105</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FB4D925ADF30FDA9" box="[1241,1406,582,609]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 103, 105</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFD4CE23FF539277D82AFC00" blockId="36.[151,1437,150,2013]" pageId="36" pageNumber="37">
|
||||
Pronotum is trapezoidal, narrowed, and rounded posteriorly (
|
||||
<figureCitation id="136616E4FFD4CE23FCEB9276DE11FD4D" box="[895,1119,618,645]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 102‒103, 105</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FCEB9276DE11FD4D" box="[895,1119,618,645]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 102‒103, 105</figureCitation>
|
||||
). The anterior and posterior margins arched, the lateral margins straight, the anterior corners almost straight; yellow, with a narrow dark brown stripe along the anterior and posterior margins; the lateral margins with a relatively wide dark sinuous band sometimes the lateral edges bent down, and the dark band is not visible dorsally (
|
||||
<figureCitation id="136616E4FFD4CE23FBBF92CADF44FD39" box="[1067,1290,726,753]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 102‒103, 105</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FBBF92CADF44FD39" box="[1067,1290,726,753]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 102‒103, 105</figureCitation>
|
||||
). The medial pronotal band is brown and relatively narrow, widening more anteriorly than posteriorly; lateral pronotal fields have dark, rounded, patches on which light rugosities are visible (
|
||||
<figureCitation id="136616E4FFD4CE23FCC69302D9B2FCF1" box="[850,1020,798,825]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 103, 105</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FCC69302D9B2FCF1" box="[850,1020,798,825]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 103, 105</figureCitation>
|
||||
). Legs yellow; femur with a narrow longitudinal grayish band close to the outer edge and a narrow dark brown band distally; tibia with a short, narrow transverse brownish band in the basal third; tarsi brown apically (
|
||||
<figureCitation id="136616E4FFD4CE23FCF3937AD986FC48" box="[871,968,870,896]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Fig. 102</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FCF3937AD986FC48" box="[871,968,870,896]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Fig. 102</figureCitation>
|
||||
). The cerci are long, projecting far beyond the end of the wings; they are bicolored, with each segment yellowish at the base and brown distally. The wings are yellow with dark brown veins (
|
||||
<figureCitation id="136616E4FFD4CE23FE6193B2D819FC00" box="[501,599,942,968]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Fig. 102</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FE6193B2D819FC00" box="[501,599,942,968]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Fig. 102</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFD4CE23FF5393CED930F974" blockId="36.[151,1437,150,2013]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B929D673FFD4CE23FF5393CEDB45FC24" bold="true" box="[199,267,978,1004]" pageId="36" pageNumber="37">Male.</emphasis>
|
||||
Tergum 8 medially with a membranous furrow narrowed posteriorly; posterolateral humps and posterior tergal margins covered with dense, relatively long setae (
|
||||
<figureCitation id="136616E4FFD4CE23FC9A93EAD921FBD8" box="[782,879,1014,1040]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Fig. 106</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FC9A93EAD921FBD8" box="[782,879,1014,1040]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Fig. 106</figureCitation>
|
||||
). Tergum 9 with a sclerotized mesal field; width is slightly greater than length; slightly expanded anteriorly and posteriorly; posterior margin with a small membranous notch in the middle (
|
||||
<figureCitation id="136616E4FFD4CE23FEEE9422DBAAFB90" box="[378,484,1086,1112]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 106</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FEEE9422DBAAFB90" box="[378,484,1086,1112]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 106</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD4CE23FE7B9422D853FB90" box="[495,541,1086,1112]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">108</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FE7B9422D853FB90" box="[495,541,1086,1112]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">108</figureCitation>
|
||||
). Longitudinal serrated ridges (10−12) cover the mesal field closer to the anterior and lateral margins; the posterior margin avoids serrated ridge rows (
|
||||
<figureCitation id="136616E4FFD4CE23FC33947EDE5DFBB4" box="[935,1043,1122,1148]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 106</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FC33947EDE5DFBB4" box="[935,1043,1122,1148]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 106</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD4CE23FBB5947EDE01FBB4" box="[1057,1103,1122,1148]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">108</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FBB5947EDE01FBB4" box="[1057,1103,1122,1148]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">108</figureCitation>
|
||||
). Hemitergites are strong and slightly narrowed towards the rounded apex (
|
||||
<figureCitation id="136616E4FFD4CE23FD04949AD97CFB68" box="[656,818,1158,1184]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 104, 106</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FD04949AD97CFB68" box="[656,818,1158,1184]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 104, 106</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD4CE23FCAA949AD922FB68" box="[830,876,1158,1184]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">108</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FCAA949AD922FB68" box="[830,876,1158,1184]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">108</figureCitation>
|
||||
). In dorsal view, the hemitergal hook (not cleared) is wide at the base, with a weak notch on the inner edge in the distal 1/3 (
|
||||
<figureCitation id="136616E4FFD4CE23FC4F94B6DE70FB0C" box="[987,1086,1194,1220]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Fig. 104</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FC4F94B6DE70FB0C" box="[987,1086,1194,1220]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Fig. 104</figureCitation>
|
||||
); the top of the hook and inner edge are covered with sensilla basiconica (
|
||||
<figureCitation id="136616E4FFD4CE23FDEE94D2D950FB20" box="[634,798,1230,1256]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Figs. 104, 106</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FDEE94D2D950FB20" box="[634,798,1230,1256]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Figs. 104, 106</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD4CE23FCBF94D2D917FB20" box="[811,857,1230,1256]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">108</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FCBF94D2D917FB20" box="[811,857,1230,1256]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">108</figureCitation>
|
||||
). The artificially everted penis carries a tube that is 1.6 times the length of the sac, a tube with a noticeable constriction in the distal third before the sac, covered dorsally with densely arranged longitudinal rows of serrate sclerites, changing into scales with small spines or without them (
|
||||
<figureCitation id="136616E4FFD4CE23FF0B9526DB0CFA9C" box="[159,322,1338,1364]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">Figs. 108‒111</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FF0B9526DB0CFA9C" box="[159,322,1338,1364]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">Figs. 108‒111</figureCitation>
|
||||
). The base of the tube is membranous, with paired patches of round spine bases laterally (
|
||||
<figureCitation id="136616E4FFD4CE23FAB29526DA8DFAB0" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">Figs. 110– 111</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FAB29526DA8DFAB0" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">Figs. 110– 111</figureCitation>
|
||||
). Ventrally, the tube is partly membranous and covered with rows of oval-elongated or polygonal sclerites with or without small spines, the same as on the dorsal surface (
|
||||
<figureCitation id="136616E4FFD4CE23FCA6959ED9D2FA54" box="[818,924,1410,1436]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">Figs. 111</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FCA6959ED9D2FA54" box="[818,924,1410,1436]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">Figs. 111</figureCitation>
|
||||
, A−C). The membranous sac is spherical and large; the sac width exceeds the tube width by 1/3; in dorsal view, the sac bears two rounded lobes at the base and a central tongue-shaped lobe narrowed to the sac base; ventrally, the sac bears two rounded lobes (
|
||||
<figureCitation id="136616E4FFD4CE23FB7895D6DFDEFA2C" box="[1260,1424,1482,1508]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">Figs. 108‒111</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FB7895D6DFDEFA2C" box="[1260,1424,1482,1508]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">Figs. 108‒111</figureCitation>
|
||||
). The apex of the sac is covered with a patch of rather coarse triangular brown spines, forming a relatively wide, loose apical brush, reaching almost to the middle of the tongue-shaped lobe (
|
||||
<figureCitation id="136616E4FFD4CE23FC23960EDEDFF9E4" box="[951,1169,1554,1580]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">Figs. 108, 110‒111</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FC23960EDEDFF9E4" box="[951,1169,1554,1580]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">Figs. 108, 110‒111</figureCitation>
|
||||
, D); there are no spines along the lateral edges of the sac. Ventrally, the apical brush ends with a triangular protrusion; dorsally, the edge of the brush is semi-rounded (
|
||||
<figureCitation id="136616E4FFD4CE23FE5C9646D8EDF9BC" box="[456,675,1626,1652]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">Figs. 108, 110–111</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FE5C9646D8EDF9BC" box="[456,675,1626,1652]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">Figs. 108, 110–111</figureCitation>
|
||||
). The configuration of the apical brush of the penis in the Caucasian population differs from the pattern of
|
||||
<taxonomicName id="4C5D71E2FFD4CE23FDD09663D8FEF950" authorityName="Guerin-Meneville" authorityYear="1843" box="[580,688,1662,1688]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pallida">
|
||||
<emphasis id="B929D673FFD4CE23FDD09663D8FEF950" box="[580,688,1662,1688]" italics="true" pageId="36" pageNumber="37">P. pallida</emphasis>
|
||||
|
|
@ -320,25 +321,25 @@ The subgenital plate is typical of several
|
|||
species
|
||||
</taxonomicName>
|
||||
, occupying>1/2 of sternum 8 width and has a small bilobed projection posteromedially. The lobes are small, pointed, and covered with reddish setae; the lateral edges of the lobes are slightly beveled; and the lobes slightly protrude beyond the anterior margin of the sternum 9 (
|
||||
<figureCitation id="136616E4FFD4CE23FF0B972EDB4CF884" box="[159,258,1842,1868]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." pageId="36" pageNumber="37">Fig. 107</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FF0B972EDB4CF884" box="[159,258,1842,1868]" captionStart="FIGURES 102‒107" captionStartId="31.[151,264,1888,1912]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure-18@31.[151,1436,181,1864]" captionTargetPageId="31" captionText="FIGURES 102‒107. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, Papay River, Pshada River Basin; 3rd tributary Kutarka River, Sochi River Basin. 102. Male habitus, dorsal. 103. Male, head and pronotum, dorsal. 104. Male, hemitergal hook, lateral. 105. Female, head and pronotum, dorsolateral. 106. Male, abdominal tip, mesal plate and hemitergites, dorsal. 107. Female, abdominal tip, ventral." figureDoi="http://doi.org/10.5281/zenodo.13747798" httpUri="https://zenodo.org/record/13747798/files/figure.png" pageId="36" pageNumber="37">Fig. 107</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFD4CE22FF53974AD9BBFEAD" blockId="36.[151,1437,150,2013]" lastBlockId="37.[151,1437,151,1761]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
|
||||
<emphasis id="B929D673FFD4CE23FF53974ADAB3F8B8" bold="true" box="[199,253,1878,1904]" pageId="36" pageNumber="37">Egg.</emphasis>
|
||||
Oval of medium size, with mean dimensions of 452×300 μm (n=8) (
|
||||
<figureCitation id="136616E4FFD4CE23FBAF974ADEEEF8B8" box="[1083,1184,1878,1904]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." pageId="36" pageNumber="37">Fig. 111</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FBAF974ADEEEF8B8" box="[1083,1184,1878,1904]" captionStart="FIGURES 108‒111" captionStartId="32.[151,264,1864,1888]" captionTargetBox="[170,1416,181,1840]" captionTargetId="figure-24@32.[170,1416,181,1840]" captionTargetPageId="32" captionText="FIGURES 108‒111. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin. Male. 108. Abdominal tip, mesal plate, hemitergal hooks, and artificially everted penis, dorsal, cleared. 109. Penis, ventral. 110. Chaetotaxy of the penis, dorsal. 111. Chaetotaxy of the penis, ventral. A. Oval scales with small spines. B. Leaf-shaped scales with small spines. C. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747800" httpUri="https://zenodo.org/record/13747800/files/figure.png" pageId="36" pageNumber="37">Fig. 111</figureCitation>
|
||||
). The anchor plate is mushroom-shaped and covered with globular bodies gathered in small groups (
|
||||
<figureCitation id="136616E4FFD4CE23FBA09766DE91F85C" box="[1076,1247,1914,1940]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="36" pageNumber="37">Figs. 113, 116</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FBA09766DE91F85C" box="[1076,1247,1914,1940]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="36" pageNumber="37">Figs. 113, 116</figureCitation>
|
||||
). The collar has a flanged rim, which is incised and distinctly wider than the neck (
|
||||
<figureCitation id="136616E4FFD4CE23FC0B9782DE0BF870" box="[927,1093,1950,1976]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="36" pageNumber="37">Figs. 114‒116</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FC0B9782DE0BF870" box="[927,1093,1950,1976]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="36" pageNumber="37">Figs. 114‒116</figureCitation>
|
||||
). The sides of the collar have prominent, continuous ribs extending from the shoulders to the rim (
|
||||
<figureCitation id="136616E4FFD4CE23FC0097DEDE79F814" box="[916,1079,1986,2012]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="36" pageNumber="37">Figs. 114, 116</figureCitation>
|
||||
<figureCitation id="136616E4FFD4CE23FC0097DEDE79F814" box="[916,1079,1986,2012]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="36" pageNumber="37">Figs. 114, 116</figureCitation>
|
||||
). The chorion surface is smooth and pitted; the chorion is covered throughout with follicle hexagonal cell impressions, limiting at the corners by six tiny round punctuations and six small, slit-shaped punctuations located between them; cells are separated from adjacent ones by expressed boundaries; the cell floor has one pore, and the pore and hexagonal punctuations are different in size (
|
||||
<figureCitation id="136616E4FFD5CE22FEC7911FDBB8FED5" box="[339,502,259,285]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="37" pageNumber="38">Figs. 118‒119</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FEC7911FDBB8FED5" box="[339,502,259,285]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="37" pageNumber="38">Figs. 118‒119</figureCitation>
|
||||
). A central depression in the cell flow is well pronounced in eggs from the Armenian population (
|
||||
<figureCitation id="136616E4FFD5CE22FEB5913BD84CFE89" box="[289,514,295,321]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="37" pageNumber="38">Figs. 113, 116‒117</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FEB5913BD84CFE89" box="[289,514,295,321]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="37" pageNumber="38">Figs. 113, 116‒117</figureCitation>
|
||||
). The ring of micropyles is located ca. 1/3 of the egg length removed from the anterior pole, the micropylar orifices without rims (
|
||||
<figureCitation id="136616E4FFD5CE22FD439157D9A7FEAD" box="[727,1001,331,357]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." pageId="37" pageNumber="38">Figs. 112–113, 117‒118</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FD439157D9A7FEAD" box="[727,1001,331,357]" captionStart="FIGURES 112‒119" captionStartId="33.[151,264,1837,1861]" captionTargetBox="[183,1404,181,1813]" captionTargetId="figure-18@33.[183,1404,181,1813]" captionTargetPageId="33" captionText="FIGURES 112‒119. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Armenia, Syunik Marz, Tzav River. Egg. 112. Appearance with micropylar row, lateral. 113. Appearance with mushroom-shaped anchor plate and micropylar row, lateral. 114. Collar with flanged rim and raised ribs, lateral. 115. Collar, dorsal. 116. Collar with anchor, covered with globular bodies, flanged rim and raised ribs, lateral. 117‒118. Chorion structure with micropylar row. 119. Chorion structure with follicle hexagonal cell impressions." figureDoi="http://doi.org/10.5281/zenodo.13747802" httpUri="https://zenodo.org/record/13747802/files/figure.png" pageId="37" pageNumber="38">Figs. 112–113, 117‒118</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFD5CE22FF539173DF62FBFD" blockId="37.[151,1437,151,1761]" pageId="37" pageNumber="38">
|
||||
|
|
@ -346,32 +347,32 @@ Oval of medium size, with mean dimensions of 452×300 μm (n=8) (
|
|||
The appearance of the larva agrees well with the description by
|
||||
<bibRefCitation id="EFCC7790FFD5CE22FC649173DF28FE41" author="Zhiltzova, L. A. & Cherchesova, S. K." box="[1008,1382,367,393]" pageId="37" pageNumber="38" pagination="321 - 326" refId="ref26717" refString="Zhiltzova, L. A. & Cherchesova, S. K. (2003) Description of the larva of genus Perla Geoff. (Plecoptera, Perlidae) from the Caucasus. Entomologicheskoe obozrenie, 82 (2), 321 - 326. [in Russian]" type="journal article" year="2003">Zhiltzova & Cherchesova (2003)</bibRefCitation>
|
||||
. The body is brown with contrasting pale patterning, covered with short dark clothing hairs, and has a dorsomesal band of erect silky white setae that is more pronounced on the last abdominal terga (
|
||||
<figureCitation id="136616E4FFD5CE22FB9891ABDEFCFE19" box="[1036,1202,439,465]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." pageId="37" pageNumber="38">Figs. 120–121</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FB9891ABDEFCFE19" box="[1036,1202,439,465]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747804" httpUri="https://zenodo.org/record/13747804/files/figure.png" pageId="37" pageNumber="38">Figs. 120–121</figureCitation>
|
||||
). The head is brown, bearing a large trapezoidal pale spot expanding from the anterior ocellus to the labrum base; the lateral edges of the spot with two pair pointed projections; paired projections near the labrum are wide; a thin pale M-line medially delimiting the spot from below; an interocellar area with a short slit-shaped pale spot between lateral ocelli; and the epicranial sutures are brown (
|
||||
<figureCitation id="136616E4FFD5CE22FE71925BD8C5FDA9" box="[485,651,583,609]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." pageId="37" pageNumber="38">Figs. 120–121</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FE71925BD8C5FDA9" box="[485,651,583,609]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747804" httpUri="https://zenodo.org/record/13747804/files/figure.png" pageId="37" pageNumber="38">Figs. 120–121</figureCitation>
|
||||
). Mandible strongly sclerotized, outer edge rounded, base with tuft of thin setae at outer edge; mandibular teeth strongly pointed; left mandible with 5 teeth; right mandible with 6 teeth; the number and arrangement of rows of short setae on the molar areas of the left and right mandibles is the same as in other species (
|
||||
<figureCitation id="136616E4FFD5CE22FEC092AFDBF8FD05" box="[340,438,691,717]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." pageId="37" pageNumber="38">Fig. 122</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FEC092AFDBF8FD05" box="[340,438,691,717]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747806" httpUri="https://zenodo.org/record/13747806/files/figure.png" pageId="37" pageNumber="38">Fig. 122</figureCitation>
|
||||
). Lacinia bidentate, teeth are pointed, terminal tooth strongly curved, subapical tooth less curved, and extending ca. 2/3 of terminal tooth length. Below the subapical tooth, there is a small downward-sloping projection with 3–4 submarginal setae (
|
||||
<figureCitation id="136616E4FFD5CE22FDCF92E7D88EFCDD" box="[603,704,763,789]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." pageId="37" pageNumber="38">Fig. 123</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FDCF92E7D88EFCDD" box="[603,704,763,789]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747806" httpUri="https://zenodo.org/record/13747806/files/figure.png" pageId="37" pageNumber="38">Fig. 123</figureCitation>
|
||||
). The marginal fringe of setae along the inner lacinial margin is complete (
|
||||
<figureCitation id="136616E4FFD5CE22FE9A9303DB3CFCF1" box="[270,370,799,825]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." pageId="37" pageNumber="38">Fig. 123</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FE9A9303DB3CFCF1" box="[270,370,799,825]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747806" httpUri="https://zenodo.org/record/13747806/files/figure.png" pageId="37" pageNumber="38">Fig. 123</figureCitation>
|
||||
). Abdomen brown, each tergum with two transverse pale spots on the sides of the medial line and paired small pale spots laterally; in the female larva, the spots on the last abdominal terga are faintly visible (
|
||||
<figureCitation id="136616E4FFD5CE22FF0B937BDB08FC49" box="[159,326,871,897]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." pageId="37" pageNumber="38">Figs. 120–121</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FF0B937BDB08FC49" box="[159,326,871,897]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747804" httpUri="https://zenodo.org/record/13747804/files/figure.png" pageId="37" pageNumber="38">Figs. 120–121</figureCitation>
|
||||
). The anal gills are absent. The legs pale, the femur darkened basally and distally, with a dark spot in the distal half close to the outer edge; the tibia basally has a short brown band on the outer edge (
|
||||
<figureCitation id="136616E4FFD5CE22FB609397DFD6FC6D" box="[1268,1432,907,933]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." pageId="37" pageNumber="38">Figs. 120–121</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FB609397DFD6FC6D" box="[1268,1432,907,933]" captionStart="FIGURES 120‒121" captionStartId="34.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure-24@34.[151,1436,181,1493]" captionTargetPageId="34" captionText="FIGURES 120‒121. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 120. Habitus, female, dorsal. 121. Habitus, male, dorsal." figureDoi="http://doi.org/10.5281/zenodo.13747804" httpUri="https://zenodo.org/record/13747804/files/figure.png" pageId="37" pageNumber="38">Figs. 120–121</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFD5CE22FF0393B3DA8BFC01" box="[151,197,943,969]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." pageId="37" pageNumber="38">124</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FF0393B3DA8BFC01" box="[151,197,943,969]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747806" httpUri="https://zenodo.org/record/13747806/files/figure.png" pageId="37" pageNumber="38">124</figureCitation>
|
||||
). The fine pilosity between dark short spines in the cercal circlet is the same as that of other
|
||||
<taxonomicName id="4C5D71E2FFD5CE22FB5493B3DF18FC01" box="[1216,1366,943,969]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B929D673FFD5CE22FB5493B3DEB0FC01" box="[1216,1278,943,969]" italics="true" pageId="37" pageNumber="38">Perla</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="136616E4FFD5CE22FAF093B3DAB1FC25" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." pageId="37" pageNumber="38">Figs. 125−127</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FAF093B3DAB1FC25" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747806" httpUri="https://zenodo.org/record/13747806/files/figure.png" pageId="37" pageNumber="38">Figs. 125−127</figureCitation>
|
||||
). A dorsal longitudinal fringe of swimming hairs is tightly pressed to the segments; the length and density of long silky hairs decrease towards the cercal apex (
|
||||
<figureCitation id="136616E4FFD5CE22FD4893EBD9CFFBD9" box="[732,897,1014,1041]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." pageId="37" pageNumber="38">Figs. 125–127</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FD4893EBD9CFFBD9" box="[732,897,1014,1041]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747806" httpUri="https://zenodo.org/record/13747806/files/figure.png" pageId="37" pageNumber="38">Figs. 125–127</figureCitation>
|
||||
). On the middle and apical cercal segments, long and thin intercalary setae are visible; their length exceeds the length of fine circlet hairs (
|
||||
<figureCitation id="136616E4FFD5CE22FBEE9407DF6EFBFD" box="[1146,1312,1051,1077]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." pageId="37" pageNumber="38">Figs. 126‒127</figureCitation>
|
||||
<figureCitation id="136616E4FFD5CE22FBEE9407DF6EFBFD" box="[1146,1312,1051,1077]" captionStart="FIGURES 122‒127" captionStartId="35.[151,264,1788,1812]" captionTargetBox="[151,1436,181,1764]" captionTargetId="figure-18@35.[151,1436,181,1764]" captionTargetPageId="35" captionText="FIGURES 122‒127. Perla pallida Guérin-Méneville, 1843. Caucasus, Russia, Krasnodar Kray, 3rd tributary Kutarka River, Sochi River Basin; Republic of North Ossetia-Alania, Prigorodny District, Mairamadag River, Terek River Basin. Larva. 122. Left and right mandibles, ventral. 123. Left lacinia, galea, maxillary palpus, ventral. 124. Right hind leg, dorsal. 125−127. Chaetotaxy of the left cercus, lateral. 125. Basal cercal segments. 126. Middle cercal segments. 127. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747806" httpUri="https://zenodo.org/record/13747806/files/figure.png" pageId="37" pageNumber="38">Figs. 126‒127</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFD5CE22FF539422D8ECFAF9" blockId="37.[151,1437,151,1761]" pageId="37" pageNumber="38">
|
||||
|
|
@ -393,7 +394,7 @@ from the Caucasus showed deep COI divergence between populations from
|
|||
(AEB9929), whose minimum p-distances were 8.1%, 4.8%, 4.7%, and 2.8%, respectively (
|
||||
<tableCitation id="C6DF3FDAFFD5CE22FEE794B7DB87FB0D" box="[371,457,1195,1221]" captionStart="TABLE 2" captionStartId="51.[151,239,509,535]" captionText="TABLE 2. Maximum intraspecific and minimum interspecific sequence divergence (%) based on p-distances for the cytochrome c oxidase subunit 1 (COI) mitochondrial DNA gene of 16 mOTUs of Perla according to BINs in BOLD systems." pageId="37" pageNumber="38">Table 2</tableCitation>
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). However, the phylogenetic analysis showed that this species is polyphyletic (
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<figureCitation id="136616E4FFD5CE22FABB94B7DFDEFB0D" box="[1327,1424,1195,1221]" captionStart="FIGURE 160" captionStartId="48.[151,250,1826,1850]" captionTargetBox="[152,1430,566,1798]" captionTargetId="figure-206@48.[151,1435,561,1803]" captionTargetPageId="48" captionText="FIGURE 160. Ultrametric Bayesian inference (BI) tree inferred from the cytochrome c oxidase I (COI) gene fragment and the results of species delineation of the genus Perla Geoffroy, 1762, and out-group Kamimuria coreana. Bayesian posterior probabilities (> 0.7) are noted above the tree nodes. Vertical bars correspond to molecular operational taxonomic units by ASAP, mPTP, GMYC, and BINs of BOLD. The color of the specimens indicates the development stage of stoneflies in accordance with the legend, while the sequences obtained in this study are in bold." pageId="37" pageNumber="38">Fig. 160</figureCitation>
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<figureCitation id="136616E4FFD5CE22FABB94B7DFDEFB0D" box="[1327,1424,1195,1221]" captionStart="FIGURE 160" captionStartId="48.[151,250,1826,1850]" captionTargetBox="[152,1430,566,1798]" captionTargetId="figure-206@48.[151,1435,561,1803]" captionTargetPageId="48" captionText="FIGURE 160. Ultrametric Bayesian inference (BI) tree inferred from the cytochrome c oxidase I (COI) gene fragment and the results of species delineation of the genus Perla Geoffroy, 1762, and out-group Kamimuria coreana. Bayesian posterior probabilities (> 0.7) are noted above the tree nodes. Vertical bars correspond to molecular operational taxonomic units by ASAP, mPTP, GMYC, and BINs of BOLD. The color of the specimens indicates the development stage of stoneflies in accordance with the legend, while the sequences obtained in this study are in bold." figureDoi="http://doi.org/10.5281/zenodo.13747826" httpUri="https://zenodo.org/record/13747826/files/figure.png" pageId="37" pageNumber="38">Fig. 160</figureCitation>
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). In addition,
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||||
<taxonomicName id="4C5D71E2FFD5CE22FE8994D3DBC8FB21" authorityName="Guerin-Meneville" authorityYear="1843" box="[285,390,1231,1257]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pallida">
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<emphasis id="B929D673FFD5CE22FE8994D3DBC8FB21" box="[285,390,1231,1257]" italics="true" pageId="37" pageNumber="38">P. pallida</emphasis>
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|
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|
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@ -1,60 +1,61 @@
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|
||||
<figureCitation id="136616E4FFF2CE05FF039303DB43FCF1" box="[151,269,799,825]" captionStart-0="FIGURES 1‒6" captionStart-1="FIGURES 7–10" captionStart-2="FIGURES 11–15" captionStart-3="FIGURES 16–17" captionStart-4="FIGURES 18–23" captionStartId-0="4.[151,264,1867,1891]" captionStartId-1="5.[151,264,1894,1918]" captionStartId-2="6.[151,264,1437,1461]" captionStartId-3="7.[151,264,1941,1965]" captionStartId-4="8.[151,264,1859,1883]" captionTargetBox-0="[170,1417,179,1841]" captionTargetBox-1="[151,1436,273,1871]" captionTargetBox-2="[170,1417,181,1414]" captionTargetBox-3="[170,1436,561,1918]" captionTargetBox-4="[170,1417,190,1835]" captionTargetId-0="figure-24@4.[170,1417,179,1841]" captionTargetId-1="figure-64@5.[151,1436,273,1871]" captionTargetId-2="figure-24@6.[170,1417,181,1414]" captionTargetId-3="figure-222@7.[151,1436,561,1918]" captionTargetId-4="figure-24@8.[170,1417,181,1835]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." captionText-1="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." captionText-2="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." captionText-3="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." captionText-4="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi-0="http://doi.org/10.5281/zenodo.13747723" figureDoi-1="http://doi.org/10.5281/zenodo.13747725" figureDoi-2="http://doi.org/10.5281/zenodo.13747733" figureDoi-3="http://doi.org/10.5281/zenodo.13747735" figureDoi-4="http://doi.org/10.5281/zenodo.13747739" httpUri-0="https://zenodo.org/record/13747723/files/figure.png" httpUri-1="https://zenodo.org/record/13747725/files/figure.png" httpUri-2="https://zenodo.org/record/13747733/files/figure.png" httpUri-3="https://zenodo.org/record/13747735/files/figure.png" httpUri-4="https://zenodo.org/record/13747739/files/figure.png" pageId="2" pageNumber="3">Figs. 1‒23</figureCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFF2CE05FF03937FDB8EFCB3" blockId="2.[151,1437,867,1351]" box="[151,448,867,891]" pageId="2" pageNumber="3">
|
||||
<bibRefCitation id="EFCC7790FFF2CE05FF03937FDBF2FCB3" author="Guerin-Meneville, F. E." box="[151,444,867,891]" pageId="2" pageNumber="3" pagination="393 - 395" refId="ref24390" refString="Guerin-Meneville, F. E. (1843) Genre Perle. In: Guerin-Meneville, F. E. (Ed.), Iconographie du Regne Animal de G. Cuvier. Insectes, 1843, pp. 393 - 395." type="journal article" year="1843">Guérin-Méneville, 1843:394</bibRefCitation>
|
||||
|
|
@ -174,23 +175,23 @@ Cherchesova
|
|||
<paragraph id="8BE20A61FFF2CE05FF039569DB0CF8D3" blockId="2.[151,1437,1397,2036]" pageId="2" pageNumber="3">
|
||||
<emphasis id="B929D673FFF2CE05FF039569DB5CFA47" bold="true" box="[151,274,1397,1423]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
|
||||
The male is distinguished by a long mesal field on tergum 9 (
|
||||
<figureCitation id="136616E4FFF2CE05FC5C9569DE43FA47" box="[968,1037,1397,1423]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FC5C9569DE43FA47" box="[968,1037,1397,1423]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||||
). The field expands anteriorly, with lateral longitudinal serrated ridges that are longer than the central ridges of teeth and an anterior part and posterior margin of the mesal field devoid of serrated ridges (
|
||||
<figureCitation id="136616E4FFF2CE05FD5E95A1D97CFA10" box="[714,818,1469,1496]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="2" pageNumber="3">Figs. 2, 5</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FD5E95A1D97CFA10" box="[714,818,1469,1496]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="2" pageNumber="3">Figs. 2, 5</figureCitation>
|
||||
). The hemitergal hook is triangular, without projections, narrowed towards the top, apex rounded (
|
||||
<figureCitation id="136616E4FFF2CE05FDFF95FDD8BBFA34" box="[619,757,1505,1532]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="2" pageNumber="3">Figs. 2, 5‒6</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FDFF95FDD8BBFA34" box="[619,757,1505,1532]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="2" pageNumber="3">Figs. 2, 5‒6</figureCitation>
|
||||
). The sac of the penis bears a narrow apical brush, ending in triangles ventrally and dorsally (
|
||||
<figureCitation id="136616E4FFF2CE05FE6B9619D836F9D7" box="[511,632,1541,1567]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." pageId="2" pageNumber="3">Figs. 9‒10</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FE6B9619D836F9D7" box="[511,632,1541,1567]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747725" httpUri="https://zenodo.org/record/13747725/files/figure.png" pageId="2" pageNumber="3">Figs. 9‒10</figureCitation>
|
||||
, C). Ventrally, on the sac, there is an additional barely noticeable ventral brush in the form of a short and narrow stripe, consisting of tiny spines (
|
||||
<figureCitation id="136616E4FFF2CE05FC5C9635DE6BF98B" box="[968,1061,1577,1603]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." pageId="2" pageNumber="3">Figs. 10</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FC5C9635DE6BF98B" box="[968,1061,1577,1603]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747725" httpUri="https://zenodo.org/record/13747725/files/figure.png" pageId="2" pageNumber="3">Figs. 10</figureCitation>
|
||||
, D). The female is recognized by the distinctive subgenital plate, with a small rectangular projection with rounded corners posteriorly (
|
||||
<figureCitation id="136616E4FFF2CE05FA819651DF12F9AF" box="[1301,1372,1613,1639]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FA819651DF12F9AF" box="[1301,1372,1613,1639]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||||
). The egg is oval-elongated; collar short, with irregular ribs; rim apex smooth with very slight irregular incisions; chorion is rough with numerous funnel-shaped punctations throughout; micropylar orifices with thin raised rims set in pits (
|
||||
<figureCitation id="136616E4FFF2CE05FF0B96A5DB67F91C" box="[159,297,1721,1748]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." pageId="2" pageNumber="3">Figs. 11‒15</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FF0B96A5DB67F91C" box="[159,297,1721,1748]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." figureDoi="http://doi.org/10.5281/zenodo.13747733" httpUri="https://zenodo.org/record/13747733/files/figure.png" pageId="2" pageNumber="3">Figs. 11‒15</figureCitation>
|
||||
). The larva is distinguished by a wide, V-shaped pale spot on the head without serrations laterally, reaching the clypeus; by the narrow semi-oval pale bands on the pronotum; and by the pilosity of the cerci (
|
||||
<figureCitation id="136616E4FFF2CE05FAF096C1DAAEF8D3" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="2" pageNumber="3">Figs. 16‒17</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FAF096C1DAAEF8D3" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="2" pageNumber="3">Figs. 16‒17</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFF2CE05FF7F971DDB78F8D3" box="[235,310,1793,1819]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="2" pageNumber="3">21‒23</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FF7F971DDB78F8D3" box="[235,310,1793,1819]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="2" pageNumber="3">21‒23</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFF2CE05FF539739DF7AF8AB" blockId="2.[151,1437,1397,2036]" pageId="2" pageNumber="3">
|
||||
|
|
@ -206,61 +207,61 @@ and
|
|||
<paragraph id="8BE20A61FFF2CE04FF539771DBF7FE65" blockId="2.[151,1437,1397,2036]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||||
<emphasis id="B929D673FFF2CE05FF539771DB3FF84F" bold="true" box="[199,369,1901,1927]" pageId="2" pageNumber="3">Adult habitus.</emphasis>
|
||||
The general body color is brownish-yellow (
|
||||
<figureCitation id="136616E4FFF2CE05FCF79771D9EDF84F" box="[867,931,1901,1927]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FCF79771D9EDF84F" box="[867,931,1901,1927]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||||
). The M-line on the head is slightly darkened and clearly defined (
|
||||
<figureCitation id="136616E4FFF2CE05FEEF978DDBF1F863" box="[379,447,1937,1963]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||||
<figureCitation id="136616E4FFF2CE05FEEF978DDBF1F863" box="[379,447,1937,1963]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||||
). A pair of thin, longitudinal, narrow light spots parallel to the lateral clypeal edges reach the anterior edge of the clypeus; the clypeus has dark lateral margins. The anterior and lateral ocelli are outlined by a black band; lateral margins of the head darkened. The interocellar area is yellow-orange, extending to M-line and compound eyes; the tentorial pits dorsal to the lateral ocelli are light brown; the occiput is darkened (
|
||||
<figureCitation id="136616E4FFF3CE04FAF0908BDA89FF1D" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Figs. 1, 3</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FAF0908BDA89FF1D" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Figs. 1, 3</figureCitation>
|
||||
). Pronotum brown; rugosities are pale-grayish, forming an X-shaped pattern (
|
||||
<figureCitation id="136616E4FFF3CE04FBC690A7DE8DFF1D" box="[1106,1219,187,213]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Figs. 1, 3</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FBC690A7DE8DFF1D" box="[1106,1219,187,213]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Figs. 1, 3</figureCitation>
|
||||
); anterolateral and posterolateral margins darkened; medial stripe brownish, widened to anterior and posterior margins; lateral margins of pronotum bordered by brown stripe, widening posterolaterally; sometimes the lateral edges are bent down and not visible dorsally. The legs and cerci are yellow-brown (
|
||||
<figureCitation id="136616E4FFF3CE04FCB1913BD929FE89" box="[805,871,295,321]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FCB1913BD929FE89" box="[805,871,295,321]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||||
). The femur bears a narrow, darkish band close to the outer edge and a black band distally. Tibia darkened basally with an additional narrow brownish band at the base (
|
||||
<figureCitation id="136616E4FFF3CE04FF0B9173DAAEFE41" box="[159,224,367,393]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FF0B9173DAAEFE41" box="[159,224,367,393]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||||
); tarsi also darkened.Anal gills small and visible only in newly molted females. Each cercal segment slightly darkened distally (
|
||||
<figureCitation id="136616E4FFF3CE04FEF3918FDBE6FE65" box="[359,424,403,429]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FEF3918FDBE6FE65" box="[359,424,403,429]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFF3CE04FF5391ABDE44FC25" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B929D673FFF3CE04FF5391ABDB45FE19" bold="true" box="[199,267,439,465]" pageId="3" pageNumber="4">Male.</emphasis>
|
||||
Tergum 8 medially has a membranous furrow and posterolateral humps covered with dense and thick setae (
|
||||
<figureCitation id="136616E4FFF3CE04FF4A91C7DB6CFE3D" box="[222,290,475,501]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FF4A91C7DB6CFE3D" box="[222,290,475,501]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||||
). Tergum 9 with a depressed sclerotized mesal field reaching the end of the posterior margin; the length of the field exceeds its width, expanded anteriorly, covered with longitudinal ridges (10–11) of sclerotized pointed teeth; lateral longitudinal serrated ridges longer than central rows of teeth; devoid of serrated ridges in the anterior part and posterior margin (
|
||||
<figureCitation id="136616E4FFF3CE04FE56925BD860FDA9" box="[450,558,582,609]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Figs. 2, 5</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FE56925BD860FDA9" box="[450,558,582,609]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Figs. 2, 5</figureCitation>
|
||||
). Tergum 10 bears hemitergites; the paired triangular hooks narrowed towards the tip, apices rounded, the base of the hook distended, and covered with long setae (
|
||||
<figureCitation id="136616E4FFF3CE04FBFE9277DEB9FD4D" box="[1130,1271,618,645]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Figs. 2, 5‒6</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FBFE9277DEB9FD4D" box="[1130,1271,618,645]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Figs. 2, 5‒6</figureCitation>
|
||||
). Hemitergites without projections, with an inner edge covered with sensilla basiconica, which is extended at the tip to the front and outer edges (
|
||||
<figureCitation id="136616E4FFF3CE04FECD92AFDBAAFD05" box="[345,484,690,717]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Figs. 2, 5‒6</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FECD92AFDBAAFD05" box="[345,484,690,717]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Figs. 2, 5‒6</figureCitation>
|
||||
). The artificially everted penis has a tube that is 1.8 times the length of the sac (
|
||||
<figureCitation id="136616E4FFF3CE04FAFB92AFDAE9FD39" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." pageId="3" pageNumber="4">Fig. 7</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FAFB92AFDAE9FD39" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747725" httpUri="https://zenodo.org/record/13747725/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
|
||||
); the base of the tube is membranous with patches of rounded spine bases laterally; the dorsal tube is covered with densely arranged longitudinal rows of heavy sclerotized serrate sclerites, sometimes changed into scales with small spines (
|
||||
<figureCitation id="136616E4FFF3CE04FEA69303DBD1FCF1" box="[306,415,799,825]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." pageId="3" pageNumber="4">Figs. 7, 9</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FEA69303DBD1FCF1" box="[306,415,799,825]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747725" httpUri="https://zenodo.org/record/13747725/files/figure.png" pageId="3" pageNumber="4">Figs. 7, 9</figureCitation>
|
||||
, A‒B); ventrally, the tube at the basal half is weakly sclerotized; in the distal 1/3 there are longitudinal rows of leaf-shaped scales with small spines (
|
||||
<figureCitation id="136616E4FFF3CE04FCB5935FD9D4FC95" box="[801,922,835,861]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." pageId="3" pageNumber="4">Figs. 8, 10</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FCB5935FD9D4FC95" box="[801,922,835,861]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747725" httpUri="https://zenodo.org/record/13747725/files/figure.png" pageId="3" pageNumber="4">Figs. 8, 10</figureCitation>
|
||||
). The sac is bulbous, membranous, at the base with lobes (
|
||||
<figureCitation id="136616E4FFF3CE04FE8C937BDBCDFC49" box="[280,387,871,897]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." pageId="3" pageNumber="4">Figs. 7–8</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FE8C937BDBCDFC49" box="[280,387,871,897]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747725" httpUri="https://zenodo.org/record/13747725/files/figure.png" pageId="3" pageNumber="4">Figs. 7–8</figureCitation>
|
||||
); thin pointed brown spines, forming a relatively narrow apical brush, noticeable in the form of a triangle ventrally and dorsally (
|
||||
<figureCitation id="136616E4FFF3CE04FD9E9397D8CDFC6D" box="[522,643,907,933]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." pageId="3" pageNumber="4">Figs. 9‒10</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FD9E9397D8CDFC6D" box="[522,643,907,933]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747725" httpUri="https://zenodo.org/record/13747725/files/figure.png" pageId="3" pageNumber="4">Figs. 9‒10</figureCitation>
|
||||
, D). Ventrally between membranous lobes, closer to the sac base, there is also a barely noticeable ventral brush in the form of a short and narrow stripe consisting of tiny spines; the spine size in the ventral brush is half the size of the apical brush spines (
|
||||
<figureCitation id="136616E4FFF3CE04FCEA93CFD995FC25" box="[894,987,979,1005]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." pageId="3" pageNumber="4">Figs. 10</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FCEA93CFD995FC25" box="[894,987,979,1005]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1894,1918]" captionTargetBox="[151,1436,273,1871]" captionTargetId="figure-64@5.[151,1436,273,1871]" captionTargetPageId="5" captionText="FIGURES 7–10. Perla caucasica Guérin-Méneville, 1843. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush." figureDoi="http://doi.org/10.5281/zenodo.13747725" httpUri="https://zenodo.org/record/13747725/files/figure.png" pageId="3" pageNumber="4">Figs. 10</figureCitation>
|
||||
, C).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFF3CE04FF5393EBDF03FBFD" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B929D673FFF3CE04FF5393EBDB6BFBD9" bold="true" box="[199,293,1015,1041]" pageId="3" pageNumber="4">Female.</emphasis>
|
||||
Color and pigmentation are similar for males. The lateral margins of the subgenital plate are weakly pronounced. The subgenital plate has medially a small rectangular projection with rounded corners (
|
||||
<figureCitation id="136616E4FFF3CE04FB6E9407DF71FBFD" box="[1274,1343,1051,1077]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." pageId="3" pageNumber="4">Fig. 4</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FB6E9407DF71FBFD" box="[1274,1343,1051,1077]" captionStart="FIGURES 1‒6" captionStartId="4.[151,264,1867,1891]" captionTargetBox="[170,1417,179,1841]" captionTargetId="figure-24@4.[170,1417,179,1841]" captionTargetPageId="4" captionText="FIGURES 1‒6. Perla caucasica Guérin-Méneville, 1843. Caucasus: Russia, Adygea, Belaya River; Republic of North Ossetia- Alania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared." figureDoi="http://doi.org/10.5281/zenodo.13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE20A61FFF3CE04FF539423DEADFB21" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B929D673FFF3CE04FF539423DAB3FB91" bold="true" box="[199,253,1087,1113]" pageId="3" pageNumber="4">Egg.</emphasis>
|
||||
Oval-elongate, with mean dimensions of 509 x 270 µm (n = 3), and a mushroom-shaped anchor (
|
||||
<figureCitation id="136616E4FFF3CE04FAA99423DFC0FB91" box="[1341,1422,1087,1113]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." pageId="3" pageNumber="4">Fig. 11</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FAA99423DFC0FB91" box="[1341,1422,1087,1113]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." figureDoi="http://doi.org/10.5281/zenodo.13747733" httpUri="https://zenodo.org/record/13747733/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
|
||||
). Collar short, with irregular ribs extending from rim to shoulders; rim apex smooth with very slight irregular incisions (
|
||||
<figureCitation id="136616E4FFF3CE04FF0B949BDB65FB69" box="[159,299,1159,1185]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." pageId="3" pageNumber="4">Figs. 12–13</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FF0B949BDB65FB69" box="[159,299,1159,1185]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." figureDoi="http://doi.org/10.5281/zenodo.13747733" httpUri="https://zenodo.org/record/13747733/files/figure.png" pageId="3" pageNumber="4">Figs. 12–13</figureCitation>
|
||||
). The chorion is pitted, with numerous funnel-shaped punctations throughout (
|
||||
<figureCitation id="136616E4FFF3CE04FB26949BDF46FB69" box="[1202,1288,1159,1185]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." pageId="3" pageNumber="4">Fig. 14</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FB26949BDF46FB69" box="[1202,1288,1159,1185]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." figureDoi="http://doi.org/10.5281/zenodo.13747733" httpUri="https://zenodo.org/record/13747733/files/figure.png" pageId="3" pageNumber="4">Fig. 14</figureCitation>
|
||||
). Micropylar row is approximately 1/3 length from the anterior pole; orifices with thin raised rims are set in pits (
|
||||
<figureCitation id="136616E4FFF3CE04FA9194B7DFC1FB0D" box="[1285,1423,1194,1221]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." pageId="3" pageNumber="4">Figs. 12, 15</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FA9194B7DFC1FB0D" box="[1285,1423,1194,1221]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1437,1461]" captionTargetBox="[170,1417,181,1414]" captionTargetId="figure-24@6.[170,1417,181,1414]" captionTargetPageId="6" captionText="FIGURES 11–15. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River. Egg. 11. Appearance with a mushroom-shaped anchor plate, lateral. 12. Same, without the anchor plate. 13. Collar, lateral. 14. Chorion structure with funnel-shaped punctations. 15. Micropyles, orifices with thin raised rims." figureDoi="http://doi.org/10.5281/zenodo.13747733" httpUri="https://zenodo.org/record/13747733/files/figure.png" pageId="3" pageNumber="4">Figs. 12, 15</figureCitation>
|
||||
). The shape and the chorion structure agree well with the original description (
|
||||
<bibRefCitation id="EFCC7790FFF3CE04FC6194D3DE99FB21" author="Sivec, I. & Stark, B. P." box="[1013,1239,1231,1257]" pageId="3" pageNumber="4" pagination="1 - 33" refId="ref26075" refString="Sivec, I. & Stark, B. P. (2002) The species of Perla (Plecoptera: Perlidae): evidence from egg morphology. Scopolia, 49, 1 - 33." type="journal article" year="2002">Sivec & Stark 2002</bibRefCitation>
|
||||
).
|
||||
|
|
@ -276,26 +277,26 @@ Mature larvae were associated with adults and corroborated the results of DNA ba
|
|||
larva by
|
||||
<bibRefCitation id="EFCC7790FFF3CE04FC159527DEB8FA9D" author="Zhiltzova, L. A. & Cherchesova, S. K." box="[897,1270,1339,1365]" pageId="3" pageNumber="4" pagination="321 - 326" refId="ref26717" refString="Zhiltzova, L. A. & Cherchesova, S. K. (2003) Description of the larva of genus Perla Geoff. (Plecoptera, Perlidae) from the Caucasus. Entomologicheskoe obozrenie, 82 (2), 321 - 326. [in Russian]" type="journal article" year="2003">Zhiltzova & Cherchesova (2003)</bibRefCitation>
|
||||
, except for the predominance of brown color and the pattern of the abdomen. Color yellow-brown (
|
||||
<figureCitation id="136616E4FFF3CE04FBA29543DEF5FAB1" box="[1078,1211,1375,1401]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="3" pageNumber="4">Figs. 16–17</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FBA29543DEF5FAB1" box="[1078,1211,1375,1401]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="3" pageNumber="4">Figs. 16–17</figureCitation>
|
||||
) with a distinct dark pattern on the head, pronotum, mesonotum, metanotum, and abdomen. Body covered with numerous short dark clothing hairs; bands of erect silky white hairs upward of the epicranial suture on the head and along the median body line, sometimes well pronounced (
|
||||
<figureCitation id="136616E4FFF3CE04FD8E95D7D820FA2D" box="[538,622,1483,1509]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="3" pageNumber="4">Fig. 16</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FD8E95D7D820FA2D" box="[538,622,1483,1509]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="3" pageNumber="4">Fig. 16</figureCitation>
|
||||
). Antennae and cerci are yellow-brown. M-line is pale, a wide V-shaped spot expanding to the clypeus is also pale, and the lateral clypeal margins are brown (
|
||||
<figureCitation id="136616E4FFF3CE04FBD495F3DE86F9C1" box="[1088,1224,1519,1545]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="3" pageNumber="4">Figs. 16–17</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FBD495F3DE86F9C1" box="[1088,1224,1519,1545]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="3" pageNumber="4">Figs. 16–17</figureCitation>
|
||||
). Head with a large transverse brown band, bounded above by the M-line and below by pale posterior tentorial pits and branches of the epicranial suture; a small triangular spot between the ocelli is pale; epicranial stem and base of the occiput brownish (
|
||||
<figureCitation id="136616E4FFF3CE04FF0B9647DB64F9BD" box="[159,298,1627,1653]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="3" pageNumber="4">Figs. 16–17</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FF0B9647DB64F9BD" box="[159,298,1627,1653]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="3" pageNumber="4">Figs. 16–17</figureCitation>
|
||||
). The transverse occipital fold and the fold running around the eye merge in a regularly curved line. Setation along the occipital fold extends only to behind the eye but not forward along the edge of the flat, expanded side of the head. Mesothoracic supracoxal gills with double trunks. Mandibles (
|
||||
<figureCitation id="136616E4FFF3CE04FBA796BFDEC6F975" box="[1075,1160,1699,1725]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="3" pageNumber="4">Fig. 18</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FBA796BFDEC6F975" box="[1075,1160,1699,1725]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="3" pageNumber="4">Fig. 18</figureCitation>
|
||||
) are heavily sclerotized along a rounded outer margin. The number of teeth and the form of the molar region are asymmetrical between the right and left mandibles. Left mandible ventrally with five pointed heavy sclerotized teeth; bearing three rows of setae; the marginal brush extends from the base of the 4th tooth to the basal third of the mandible; the submarginal setal row stretches from the base of the 3rd tooth to the mandible basal third; the last submarginal setal row prolongs from the base of the first tooth to the mandible base (
|
||||
<figureCitation id="136616E4FFF3CE04FD70974BD979F8B9" box="[740,823,1879,1905]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="3" pageNumber="4">Fig. 18</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FD70974BD979F8B9" box="[740,823,1879,1905]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="3" pageNumber="4">Fig. 18</figureCitation>
|
||||
). Right mandible ventrally with six teeth, the last tooth being small, obtuse-angled, sometimes not developed or poorly visible; marginal brush extending from the base of the 4th tooth to the basal 1/3 of the mandible; the submarginal setal row prolongs from the base of the first tooth to the mandible base (
|
||||
<figureCitation id="136616E4FFF3CE04FEE097DFDB87F815" box="[372,457,1987,2013]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="3" pageNumber="4">Fig. 18</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FEE097DFDB87F815" box="[372,457,1987,2013]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="3" pageNumber="4">Fig. 18</figureCitation>
|
||||
). Lacinia bidentate (
|
||||
<figureCitation id="136616E4FFF3CE04FD2697DFD94BF815" box="[690,773,1987,2013]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="3" pageNumber="4">Fig. 19</figureCitation>
|
||||
<figureCitation id="136616E4FFF3CE04FD2697DFD94BF815" box="[690,773,1987,2013]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="3" pageNumber="4">Fig. 19</figureCitation>
|
||||
) has a large, curved, and strongly sclerotized apical tooth; the subapical tooth is shorter, extending 2/3 of the apical tooth length. The marginal fringe of setae along the inner margin is complete; short median setae are present in a sparse patch; basal setae are sporadic. Galea length reaches half of apical tooth length (
|
||||
<figureCitation id="136616E4FFF5CE02FE5C90C3D855FF31" box="[456,539,223,249]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="5" pageNumber="6">Fig. 19</figureCitation>
|
||||
<figureCitation id="136616E4FFF5CE02FE5C90C3D855FF31" box="[456,539,223,249]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="5" pageNumber="6">Fig. 19</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF225AE9FFF4CE03FF039757DBE0F83B" pageId="4" pageNumber="5" startId="4.[151,264,1867,1891]" targetBox="[170,1417,179,1841]" targetPageId="4" targetType="figure">
|
||||
<caption id="DF225AE9FFF4CE03FF039757DBE0F83B" ID-DOI="http://doi.org/10.5281/zenodo.13747723" ID-Zenodo-Dep="13747723" httpUri="https://zenodo.org/record/13747723/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,264,1867,1891]" targetBox="[170,1417,179,1841]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="8BE20A61FFF4CE03FF039757DBE0F83B" blockId="4.[151,1437,1867,2035]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B929D673FFF4CE03FF039757DB74F8AB" bold="true" box="[151,314,1867,1891]" pageId="4" pageNumber="5">FIGURES 1‒6.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFF4CE03FEAB9757D891F8AB" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[319,735,1867,1891]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caucasica">
|
||||
|
|
@ -305,7 +306,7 @@ larva by
|
|||
. Caucasus: Russia, Adygea, Belaya River; Republic of North OssetiaAlania, Mairamadag River. Georgia, Sukhum District, Kelasuri River. 1. Habitus, male, dorsal. 2. Male abdominal tip, dorsal. 3. Head and pronotum, male, dorsal. 4. Female subgenital plate, ventral. 5. Mesal field with longitudinal serrated ridges, dorsolateral, and hemitergal hooks covered with sensilla basiconica, lateral, cleared. 6. Hemitergal hooks covered with sensilla basiconica, caudal, cleared.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFF5CE02FF03977AD98FF822" pageId="5" pageNumber="6" startId="5.[151,264,1894,1918]" targetBox="[151,1436,273,1871]" targetPageId="5" targetType="figure">
|
||||
<caption id="DF225AE9FFF5CE02FF03977AD98FF822" ID-DOI="http://doi.org/10.5281/zenodo.13747725" ID-Zenodo-Dep="13747725" httpUri="https://zenodo.org/record/13747725/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,264,1894,1918]" targetBox="[151,1436,273,1871]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="8BE20A61FFF5CE02FF03977AD98FF822" blockId="5.[151,1437,1894,2026]" pageId="5" pageNumber="6">
|
||||
<emphasis id="B929D673FFF5CE02FF03977ADB05F8B6" bold="true" box="[151,331,1894,1918]" pageId="5" pageNumber="6">FIGURES 7–10.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFF5CE02FEC1977AD94CF8B6" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[341,770,1894,1918]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="caucasica">
|
||||
|
|
@ -315,7 +316,7 @@ larva by
|
|||
. Caucasus, Russia, Adygea, Belaya River. 7. Hemitergal hooks and artificially everted penis, dorsal. 8. Penis, ventral. 9. Chaetotaxy of artificially everted penis, dorsal. A. Densely arranged longitudinal rows of heavy sclerotized serrate sclerites on the tube of the penis. B. Scales with small spines. 10. Same, ventral. C. Tiny spines in the ventral brush. D. Pointed brown spines in the apical brush.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFF6CE01FF039581DE5FFA36" pageId="6" pageNumber="7" startId="6.[151,264,1437,1461]" targetBox="[170,1417,181,1414]" targetPageId="6" targetType="figure">
|
||||
<caption id="DF225AE9FFF6CE01FF039581DE5FFA36" ID-DOI="http://doi.org/10.5281/zenodo.13747733" ID-Zenodo-Dep="13747733" httpUri="https://zenodo.org/record/13747733/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,264,1437,1461]" targetBox="[170,1417,181,1414]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="8BE20A61FFF6CE01FF039581DE5FFA36" blockId="6.[151,1436,1437,1534]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B929D673FFF6CE01FF039581DB18FA7D" bold="true" box="[151,342,1437,1461]" pageId="6" pageNumber="7">FIGURES 11–15.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFF6CE01FECB9582D944FA7E" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[351,778,1438,1462]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caucasica">
|
||||
|
|
@ -327,41 +328,41 @@ larva by
|
|||
</caption>
|
||||
<paragraph id="8BE20A61FFF6CE00FF53964CD995FDD1" blockId="6.[151,1437,1616,2038]" lastBlockId="7.[151,1437,151,537]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||||
Pronotum transverse, approximately the same width as the head, with obtusely rounded anterior corners; lateral margins and posterior angles are evenly rounded (
|
||||
<figureCitation id="136616E4FFF6CE01FD2F9668D90CF946" box="[699,834,1652,1678]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
<figureCitation id="136616E4FFF6CE01FD2F9668D90CF946" box="[699,834,1652,1678]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
). The pronotal fringe is complete with mixed short and relatively long setae. Pronotum with a thin brown stripe along the edges; paired narrow semi-oval pale bands from the anterior corners to the middle of the posterior margin outline an oval brown band; lateral pronotal fields pale (
|
||||
<figureCitation id="136616E4FFF6CE01FF0B96FCDB66F932" box="[159,296,1760,1786]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
<figureCitation id="136616E4FFF6CE01FF0B96FCDB66F932" box="[159,296,1760,1786]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
). Medial pronotal band brown, widening anteriorly and posteriorly; band surrounded by pale rugosites of an X-shaped pattern; a paired vague S-shaped dark spot extends from the anterior margin to the middle of the pronotum (
|
||||
<figureCitation id="136616E4FFF6CE01FE809734DBD0F88A" box="[276,414,1832,1858]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
<figureCitation id="136616E4FFF6CE01FE809734DBD0F88A" box="[276,414,1832,1858]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
). Mesonotum and metanotum with identical Ψ-shaped brown pattern medially; posteriorly darkly margined; two wide brown bands limiting the wing pad bases; a brown stripe runs along the lateral edge of the wing pads (
|
||||
<figureCitation id="136616E4FFF6CE01FEAB976CDB89F842" box="[319,455,1904,1930]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
<figureCitation id="136616E4FFF6CE01FEAB976CDB89F842" box="[319,455,1904,1930]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
). Legs are yellow; the femur and tibia bear stout, short, acute bristles along the inner and outer edges, and a very dense fringe of long, silky hairs along the outer margins. Hind femur ca. 3.5–3.9 times as long as wide. The femur surface is covered with sparse, short, and irregular red bristles; there is a diffuse brownish spot in the distal half closer to the outer edge, and the distal edge of the femur is also darkened (
|
||||
<figureCitation id="136616E4FFF6CE01FB4F97C0DF2AF83E" box="[1243,1380,2012,2038]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
<figureCitation id="136616E4FFF6CE01FB4F97C0DF2AF83E" box="[1243,1380,2012,2038]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
|
||||
,
|
||||
<figureCitation id="136616E4FFF6CE01FAE497C0DFDEF83E" box="[1392,1424,2012,2038]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="6" pageNumber="7">20</figureCitation>
|
||||
<figureCitation id="136616E4FFF6CE01FAE497C0DFDEF83E" box="[1392,1424,2012,2038]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="6" pageNumber="7">20</figureCitation>
|
||||
). Tibia with a diffuse, narrow brownish band basally (
|
||||
<figureCitation id="136616E4FFF7CE00FD77908BD90CFF79" box="[739,834,151,177]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="7" pageNumber="8">Figs. 20</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FD77908BD90CFF79" box="[739,834,151,177]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="7" pageNumber="8">Figs. 20</figureCitation>
|
||||
). Abdomen brown, each tergum with two median and two lateral oval pale spots forming longitudinal rows (
|
||||
<figureCitation id="136616E4FFF7CE00FC9990A7D9D7FF1D" box="[781,921,187,213]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="7" pageNumber="8">Figs. 16–17</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FC9990A7D9D7FF1D" box="[781,921,187,213]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="7" pageNumber="8">Figs. 16–17</figureCitation>
|
||||
). The two last terga of the female larvae are somewhat darker than the previous ones (
|
||||
<figureCitation id="136616E4FFF7CE00FDF690C3D8FDFF31" box="[610,691,223,249]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." pageId="7" pageNumber="8">Fig. 16</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FDF690C3D8FDFF31" box="[610,691,223,249]" captionStart="FIGURES 16–17" captionStartId="7.[151,264,1941,1965]" captionTargetBox="[170,1436,561,1918]" captionTargetId="figure-222@7.[151,1436,561,1918]" captionTargetPageId="7" captionText="FIGURES 16–17. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva." figureDoi="http://doi.org/10.5281/zenodo.13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="7" pageNumber="8">Fig. 16</figureCitation>
|
||||
), the posterior margin of sternum 7 is incomplete for female larvae. Anal gills are small. Cerci brown; each cercal circlet with rings of stout brown spines and fine pilosity between spines, tightly pressed to the segment surface (
|
||||
<figureCitation id="136616E4FFF7CE00FD36913BD962FE89" box="[674,812,295,321]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="7" pageNumber="8">Figs. 21‒23</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FD36913BD962FE89" box="[674,812,295,321]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="7" pageNumber="8">Figs. 21‒23</figureCitation>
|
||||
); short fine hairs are the same length as the basal cercal segment length (
|
||||
<figureCitation id="136616E4FFF7CE00FEC59157DBEAFEAD" box="[337,420,331,357]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="7" pageNumber="8">Fig. 21</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FEC59157DBEAFEAD" box="[337,420,331,357]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="7" pageNumber="8">Fig. 21</figureCitation>
|
||||
); towards the middle of the cerci, fine pilosity length decreases (
|
||||
<figureCitation id="136616E4FFF7CE00FBE29157DE84FEAD" box="[1142,1226,331,357]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="7" pageNumber="8">Fig. 22</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FBE29157DE84FEAD" box="[1142,1226,331,357]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="7" pageNumber="8">Fig. 22</figureCitation>
|
||||
), and on the apical cercal segments, the length of the fine pilosity is about a quarter of the apical segment length (
|
||||
<figureCitation id="136616E4FFF7CE00FB2F9173DF5EFE41" box="[1211,1296,367,393]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="7" pageNumber="8">Fig. 23</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FB2F9173DF5EFE41" box="[1211,1296,367,393]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="7" pageNumber="8">Fig. 23</figureCitation>
|
||||
). Long silky colorless hairs are included in the circlet in tufts and form a dorsal longitudinal swimming fringe, well visible especially on the basal cercal segments (
|
||||
<figureCitation id="136616E4FFF7CE00FDF791ABD8F9FE19" box="[611,695,439,465]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="7" pageNumber="8">Fig. 21</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FDF791ABD8F9FE19" box="[611,695,439,465]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="7" pageNumber="8">Fig. 21</figureCitation>
|
||||
); the length and density of long silky hairs also decrease towards the apex (
|
||||
<figureCitation id="136616E4FFF7CE00FE9691C7DBCEFE3D" box="[258,384,475,501]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="7" pageNumber="8">Fig. 21–23</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FE9691C7DBCEFE3D" box="[258,384,475,501]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="7" pageNumber="8">Fig. 21–23</figureCitation>
|
||||
). Additionally, a few fine intercalary setae are present on each cercal segment, and their density and length increase from the basal to apical cercal segments (
|
||||
<figureCitation id="136616E4FFF7CE00FCD291E3D99EFDD1" box="[838,976,511,537]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." pageId="7" pageNumber="8">Figs. 21‒23</figureCitation>
|
||||
<figureCitation id="136616E4FFF7CE00FCD291E3D99EFDD1" box="[838,976,511,537]" captionStart="FIGURES 18–23" captionStartId="8.[151,264,1859,1883]" captionTargetBox="[170,1417,190,1835]" captionTargetId="figure-24@8.[170,1417,181,1835]" captionTargetPageId="8" captionText="FIGURES 18–23. Perla caucasica Guérin-Méneville, 1843. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. 18. Left and right mandibles, ventral. 19. Left lacinia, galea, maxillary palpus, ventral. 20. Right middle leg, dorsal. 21‒23. Chaetotaxy of the right cercus, lateral. 21. Basal cercal segments. 22. Middle cercal segments. 23. Apical cercal segments." figureDoi="http://doi.org/10.5281/zenodo.13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="7" pageNumber="8">Figs. 21‒23</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF225AE9FFF7CE00FF039789D989F819" pageId="7" pageNumber="8" startId="7.[151,264,1941,1965]" targetBox="[170,1436,561,1918]" targetPageId="7" targetType="figure">
|
||||
<caption id="DF225AE9FFF7CE00FF039789D989F819" ID-DOI="http://doi.org/10.5281/zenodo.13747735" ID-Zenodo-Dep="13747735" httpUri="https://zenodo.org/record/13747735/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,264,1941,1965]" targetBox="[170,1436,561,1918]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8BE20A61FFF7CE00FF039789D989F819" blockId="7.[151,1436,1941,2001]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B929D673FFF7CE00FF039789DB16F865" bold="true" box="[151,344,1941,1965]" pageId="7" pageNumber="8">FIGURES 16–17.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFF7CE00FEF69789D941F865" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[354,783,1941,1965]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="caucasica">
|
||||
|
|
@ -371,7 +372,7 @@ Pronotum transverse, approximately the same width as the head, with obtusely rou
|
|||
. Caucasus, Georgia, Sukhum District, Kelasuri River; Russia, Adygea, Belaya River. Larvae, habitus, dorsal. 16. Female larva. 17. Male larva.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF225AE9FFF8CE0FFF03975FDB0EF800" pageId="8" pageNumber="9" startId="8.[151,264,1859,1883]" targetBox="[170,1417,190,1835]" targetPageId="8" targetType="figure">
|
||||
<caption id="DF225AE9FFF8CE0FFF03975FDB0EF800" ID-DOI="http://doi.org/10.5281/zenodo.13747739" ID-Zenodo-Dep="13747739" httpUri="https://zenodo.org/record/13747739/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,264,1859,1883]" targetBox="[170,1417,190,1835]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8BE20A61FFF8CE0FFF03975FDB0EF800" blockId="8.[151,1437,1859,1992]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B929D673FFF8CE0FFF03975FDB16F893" bold="true" box="[151,344,1859,1883]" pageId="8" pageNumber="9">FIGURES 18–23.</emphasis>
|
||||
<taxonomicName id="4C5D71E2FFF8CE0FFEF6975FD941F894" authority="Guerin-Meneville, 1843" authorityName="Guerin-Meneville" authorityYear="1843" box="[354,783,1859,1884]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="caucasica">
|
||||
|
|
@ -518,7 +519,7 @@ Northern slopes of the Greater Caucasus: Adygea, Caucasus Nature Reserve, Karach
|
|||
, Lagodekhi River (
|
||||
<bibRefCitation id="EFCC7790FFF9CE0EFF70923FDBDFFDF5" author="Zhiltzova, L. A." box="[228,401,547,573]" pageId="9" pageNumber="10" pagination="101 - 114" refId="ref26685" refString="Zhiltzova, L. A. (1964) Die Plecopteren des europaischen Teils der Sowjetunion und des Kaukasus. Gewasser und Abwasser, 34 / 35, 101 - 114." type="journal article" year="1964">Zhiltzova 1964</bibRefCitation>
|
||||
) (
|
||||
<figureCitation id="136616E4FFF9CE0EFE3C923FD844FDF5" box="[424,522,546,573]" captionStart="FIGURE 159" captionStartId="47.[151,250,1780,1804]" captionTargetBox="[158,1430,846,1757]" captionTargetId="figure-40@47.[158,1430,846,1757]" captionTargetPageId="47" captionText="FIGURE 159. The map of the Perla species distribution in Caucasus according to our data. Perla caicasica—pink, P. persica— green, P. kiritshenkoi—blue, P. schapsugica sp. nov.—white, P. pallida—yellow, and P. palatovi sp. nov.—red." pageId="9" pageNumber="10">Fig. 159</figureCitation>
|
||||
<figureCitation id="136616E4FFF9CE0EFE3C923FD844FDF5" box="[424,522,546,573]" captionStart="FIGURE 159" captionStartId="47.[151,250,1780,1804]" captionTargetBox="[158,1430,846,1757]" captionTargetId="figure-40@47.[158,1430,846,1757]" captionTargetPageId="47" captionText="FIGURE 159. The map of the Perla species distribution in Caucasus according to our data. Perla caicasica—pink, P. persica— green, P. kiritshenkoi—blue, P. schapsugica sp. nov.—white, P. pallida—yellow, and P. palatovi sp. nov.—red." figureDoi="http://doi.org/10.5281/zenodo.13747824" httpUri="https://zenodo.org/record/13747824/files/figure.png" pageId="9" pageNumber="10">Fig. 159</figureCitation>
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).
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<collectingRegion id="4999C483FFF9CE0EFD88923FD8C4FDF5" box="[540,650,547,573]" country="Georgia" name="Abkhazia" pageId="9" pageNumber="10">Abkhazia</collectingRegion>
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, Kalasuri River. The question of whether
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