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<document id="9CD51FABDBDE74DC1EF6AA4FFA508258" ID-CLB-Dataset="55819" ID-DOI="10.11646/zootaxa.5129.1.4" ID-GBIF-Dataset="87d5d1bc-c72d-4f08-89ae-da6af7187254" ID-ISSN="1175-5326" ID-Zenodo-Dep="6488189" ID-ZooBank="7C85ABDD-B34B-4D1D-A06C-364A8BF8FDCC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650960298758" checkinUser="plazi" docAuthor="Provenzano-Rizzi, Francisco, Argüello, Pablo &amp; Barriga-Salazar, Ramiro" docDate="2022" docId="70502320FFF71E52FF0DFDF1EAD048FD" docLanguage="en" docName="zootaxa.5129.1.4.pdf" docOrigin="Zootaxa 5129 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudohemiodon almendarizi Provenzano-Rizzi &amp; Argüello &amp; Barriga-Salazar 2022, new species" docType="treatment" docVersion="8" lastPageNumber="81" masterDocId="8C695B58FFF51E56FF9AFFB7EA1A4A78" masterDocTitle="The genus Pseudohemiodon (Siluriformes, Loricariidae) in Ecuador, with the description of a new species" masterLastPageNumber="91" masterPageNumber="77" pageNumber="79" updateTime="1699329221450" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AD1F67A2C7AE0DA547962422F724A317">The genus Pseudohemiodon (Siluriformes, Loricariidae) in Ecuador, with the description of a new species</mods:title>
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<mods:namePart id="30D156794FADE6AE64478355D035776F">Provenzano-Rizzi, Francisco</mods:namePart>
<mods:affiliation id="C3F335C495625353BB4FDCF25646E73A">Centro MBUCV, Instituto de Zoología y Ecología Tropical, Facultad de Ciencias, UCV, Caracas, Venezuela.</mods:affiliation>
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<mods:namePart id="DCEC06E047DF45983EE861F9EAD35089">Argüello, Pablo</mods:namePart>
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<mods:affiliation id="B76D4BAC345EB71901C724148D79099C">Departamento de Biología, Facultad de Ciencias, Escuela Politécnica Nacional, Quito, Ecuador. &amp; pablo. arguello @ epn. edu. ec; https: // orcid. org / 0000 - 0001 - 8208 - 441 X</mods:affiliation>
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<mods:namePart id="B260401456F33BEA5130B8F56253CD57">Barriga-Salazar, Ramiro</mods:namePart>
<mods:nameIdentifier id="EC9C9C95893F70540FC4453344C2CA96" type="ORCID">0000-0002-0632-559X</mods:nameIdentifier>
<mods:affiliation id="3216F6F9F7A38807101F4D34A77E96F8">Departamento de Biología, Facultad de Ciencias, Escuela Politécnica Nacional, Quito, Ecuador. &amp; ramiro. barriga @ epn. edu. ec; https: // orcid. org / 0000 - 0002 - 0632 - 559 X</mods:affiliation>
<mods:nameIdentifier id="3D8CB6A4ABA87787C226ECCB6FF0F802" type="email">ramiro.barriga@epn.edu.ec</mods:nameIdentifier>
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<mods:title id="4A48886D6780E83890E3FDA3656A540A">Zootaxa</mods:title>
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<mods:date id="C213F41B08DD2B0BAC6B331D3C6AACD2">2022</mods:date>
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<mods:number id="4F64E6C86F320AB4A399D618B65F8384">5129</mods:number>
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<treatment id="70502320FFF71E52FF0DFDF1EAD048FD" ID-DOI="http://doi.org/10.5281/zenodo.6487816" ID-GBIF-Taxon="194854779" ID-Zenodo-Dep="6487816" LSID="urn:lsid:plazi:treatment:70502320FFF71E52FF0DFDF1EAD048FD" httpUri="http://treatment.plazi.org/id/70502320FFF71E52FF0DFDF1EAD048FD" lastPageId="4" lastPageNumber="81" pageId="2" pageNumber="79">
<subSubSection id="B0E3C1BDFFF71E54FF0DFDF1E88C4818" box="[151,662,582,609]" pageId="2" pageNumber="79" type="nomenclature">
<paragraph id="F8469236FFF71E54FF0DFDF1E88C4818" blockId="2.[151,984,582,681]" box="[151,662,582,609]" pageId="2" pageNumber="79">
<heading id="A30E255AFFF71E54FF0DFDF1E88C4818" bold="true" box="[151,662,582,609]" fontSize="11" level="1" pageId="2" pageNumber="79" reason="1">
<emphasis id="CA8D4E24FFF71E54FF0DFDF1E88C4818" bold="true" box="[151,662,582,609]" pageId="2" pageNumber="79">
<taxonomicName id="3FF9E9B5FFF71E54FF0DFDF1EBE44819" authority="Provenzano-Rizzi &amp; Argüello &amp; Barriga-Salazar, 2022" authorityName="Provenzano-Rizzi &amp; Argüello &amp; Barriga-Salazar" authorityYear="2022" box="[151,510,582,609]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="almendarizi" status="sp. nov.">
<emphasis id="CA8D4E24FFF71E54FF0DFDF1EBE44819" bold="true" box="[151,510,582,609]" italics="true" pageId="2" pageNumber="79">Pseudohemiodon almendarizi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1BEF35FFFF71E54FD9CFDF0E88C4818" box="[518,662,582,609]" pageId="2" pageNumber="79" rank="species">new species</taxonomicNameLabel>
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</paragraph>
</subSubSection>
<subSubSection id="B0E3C1BDFFF71E54FF0DFDDCE9C248D1" pageId="2" pageNumber="79" type="description">
<paragraph id="F8469236FFF71E54FF0DFDDCEB8248FD" blockId="2.[151,984,582,681]" box="[151,408,619,645]" pageId="2" pageNumber="79">
<figureCitation id="60C28EB3FFF71E54FF0DFDDCEB1A48FD" box="[151,256,619,645]" captionStart="FIGURE 1" captionStartId="3.[151,250,1638,1662]" captionTargetBox="[151,1436,309,1615]" captionTargetId="figure-82@3.[151,1436,309,1615]" captionTargetPageId="3" captionText="FIGURE 1. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 1 cm. Photos by P. Argüello." figureDoi="http://doi.org/10.5281/zenodo.6488191" httpUri="https://zenodo.org/record/6488191/files/figure.png" pageId="2" pageNumber="79">Figures 1</figureCitation>
,
<figureCitation id="60C28EB3FFF71E54FE94FDDCEB0448FD" box="[270,286,619,645]" captionStart="FIGURE 2" captionStartId="4.[151,250,1956,1980]" captionTargetBox="[151,1435,669,1933]" captionTargetId="figure-228@4.[151,1435,669,1933]" captionTargetPageId="4" captionText="FIGURE 2. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Ventral view, detail of head contour, mouth and abdomen cover. Scale bar 5 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.6488193" httpUri="https://zenodo.org/record/6488193/files/figure.png" pageId="2" pageNumber="79">2</figureCitation>
,
<figureCitation id="60C28EB3FFF71E54FEB0FDDCEB2348FD" box="[298,313,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1777,1801]" captionTargetBox="[161,1426,813,1754]" captionTargetId="figure-344@6.[161,1426,813,1754]" captionTargetPageId="6" captionText="FIGURE 3. a) P. almendarizi, MEPN 17903, 87.9 mm SL, holotype. b) P. apithanos, MEPN 18374, 97.5 mm SL. c) P. lamina, MEPN 15095, 101.9 mm SL. d) P. laticeps, MCP 9308, 126.1 mm SL. Dorsal views, scale bar 1 cm. Photos a), b) and c) by P. Argüello; d) by B. Calegari." figureDoi="http://doi.org/10.5281/zenodo.6488195" httpUri="https://zenodo.org/record/6488195/files/figure.png" pageId="2" pageNumber="79">3</figureCitation>
,
<tableCitation id="B57BA78DFFF71E54FEDCFDDCEB8248FD" box="[326,408,619,645]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Morphometric data of Pseudohemiodon almendarizi (n=2)." httpUri="http://table.plazi.org/id/AC86C2BEFFF01E53FF0DFF20E9934AD7" pageId="2" pageNumber="79" tableUuid="AC86C2BEFFF01E53FF0DFF20E9934AD7">Table 1</tableCitation>
</paragraph>
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urn:lsid:zoobank.org:act:
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<paragraph id="F8469236FFF71E54FF0DFD60EE5E496D" blockId="2.[151,1437,727,2013]" pageId="2" pageNumber="79">
<materialsCitation id="4891986BFFF71E54FF0DFD60EE5E496D" ID-GBIF-Occurrence="3757616315" collectingDate="1998-07-24" collectionCode="R" collectorName="Barriga &amp; D. J. Stewart." country="Ecuador" county="Aguarico River" latitude="-0.8825" location="River" longLatPrecision="21" longitude="-75.225" municipality="Puerto" pageId="2" pageNumber="79" specimenCode="MEPN 17903" specimenCount="1" stateProvince="Orellana" typeStatus="holotype">
<emphasis id="CA8D4E24FFF71E54FF0DFD60EB114889" bold="true" box="[151,267,727,753]" pageId="2" pageNumber="79">
<typeStatus id="27422C94FFF71E54FF0DFD60EB1C4889" box="[151,262,727,753]" pageId="2" pageNumber="79" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="A85F3A4DFFF71E54FE88FD60EBAA4889" box="[274,432,727,753]" collectionCode="MEPN" pageId="2" pageNumber="79">MEPN 17903</specimenCode>
,
<quantity id="3F013FD3FFF71E54FE21FD60E8054889" box="[443,543,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="8.79" pageId="2" pageNumber="79" unit="mm" value="87.9">87.9 mm</quantity>
SL,
<collectingCountry id="80EED2A6FFF71E54FDCFFD60E8AE4889" box="[597,692,727,753]" name="Ecuador" pageId="2" pageNumber="79">Ecuador</collectingCountry>
,
<collectingRegion id="3A3D5CD4FFF71E54FD25FD60E9974889" box="[703,909,727,753]" country="Ecuador" name="Orellana" pageId="2" pageNumber="79">Orellana Province</collectingRegion>
,
<collectingCounty id="1127EABAFFF71E54FC0DFD60EE5C4889" box="[919,1094,727,753]" pageId="2" pageNumber="79">Aguarico River</collectingCounty>
, near
<collectingMunicipality id="1822084CFFF71E54FB13FD60EEC84889" box="[1161,1234,727,753]" pageId="2" pageNumber="79">Puerto</collectingMunicipality>
<collectingRegion id="3A3D5CD4FFF71E54FB43FD60EF154889" box="[1241,1295,727,753]" country="Ecuador" name="Loja" pageId="2" pageNumber="79">Loja</collectingRegion>
,
<collectingRegion id="3A3D5CD4FFF71E54FA80FD60EF4D4889" box="[1306,1367,727,753]" country="Ecuador" name="Napo" pageId="2" pageNumber="79">Napo</collectingRegion>
<location id="FD26C4EDFFF71E54FAC4FD60EF864889" LSID="urn:lsid:plazi:treatment:70502320FFF71E52FF0DFDF1EAD048FD:FD26C4EDFFF71E54FAC4FD60EF864889" box="[1374,1436,727,753]" country="Ecuador" county="Aguarico River" latitude="-0.8825" longLatPrecision="21" longitude="-75.225" municipality="Puerto" name="River" pageId="2" pageNumber="79" stateProvince="Orellana">River</location>
system, approx.
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<geoCoordinate id="9DCDF4F1FFF71E54FE40FD4DE870496D" box="[474,618,762,789]" degrees="75" direction="west" minutes="13" orientation="longitude" pageId="2" pageNumber="79" precision="15" seconds="30" value="-75.225">75°13’30”W</geoCoordinate>
,
<date id="8C47B4F6FFF71E54FDEFFD4CE913496D" box="[629,777,763,789]" pageId="2" pageNumber="79" value="1998-07-24">
<collectingDate id="9C034D1EFFF71E54FDEFFD4CE913496D" box="[629,777,763,789]" pageId="2" pageNumber="79" value="1998-07-24">24 July 1998</collectingDate>
</date>
,
<collectionCode id="9EE80AF3FFF71E54FC8EFD4CE930496D" box="[788,810,763,789]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="79">R</collectionCode>
.
<collectorName id="550CF7E0FFF71E54FCAEFD4CE992496D" box="[820,904,763,789]" pageId="2" pageNumber="79">Barriga</collectorName>
&amp;
<collectorName id="550CF7E0FFF71E54FC36FD4CEE5E496D" box="[940,1092,763,789]" pageId="2" pageNumber="79">D. J. Stewart.</collectorName>
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</paragraph>
<paragraph id="F8469236FFF71E54FF5DFCA8E9984941" blockId="2.[151,1437,727,2013]" box="[199,898,799,825]" pageId="2" pageNumber="79">
<materialsCitation id="4891986BFFF71E54FF5DFCA8E9654941" ID-GBIF-Occurrence="3757616308" box="[199,895,799,825]" collectingDate="1998-07-24" collectionCode="MEPN" collectorName="Barriga &amp; D. J. Stewart." country="Ecuador" county="Aguarico River" latitude="-0.8825" location="River" longLatPrecision="21" longitude="-75.225" municipality="Puerto" pageId="2" pageNumber="79" specimenCode="MEPN 19491" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
<emphasis id="CA8D4E24FFF71E54FF5DFCA8EB214941" bold="true" box="[199,315,799,825]" pageId="2" pageNumber="79">
<typeStatus id="27422C94FFF71E54FF5DFCA8EB2C4941" box="[199,310,799,825]" pageId="2" pageNumber="79" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCode id="A85F3A4DFFF71E54FED8FCA8EBFA4941" box="[322,480,799,825]" collectionCode="MEPN" pageId="2" pageNumber="79">MEPN 19491</specimenCode>
,
<quantity id="3F013FD3FFF71E54FE71FCA8E8554941" box="[491,591,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="8.03" pageId="2" pageNumber="79" unit="mm" value="80.3">80.3 mm</quantity>
SL, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B0E3C1BDFFF71E54FF5DFCF5EB4D4F75" pageId="2" pageNumber="79" type="diagnosis">
<paragraph id="F8469236FFF71E54FF5DFCF5EB4D4F75" blockId="2.[151,1437,727,2013]" pageId="2" pageNumber="79">
<emphasis id="CA8D4E24FFF71E54FF5DFCF5EB584924" bold="true" box="[199,322,834,860]" pageId="2" pageNumber="79">Diagnosis.</emphasis>
<taxonomicName id="3FF9E9B5FFF71E54FED0FCF4E8804925" authorityName="Provenzano-Rizzi &amp; Argüello &amp; Barriga-Salazar" authorityYear="2022" box="[330,666,835,861]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="almendarizi">
<emphasis id="CA8D4E24FFF71E54FED0FCF4E8804925" box="[330,666,835,861]" italics="true" pageId="2" pageNumber="79">Pseudohemiodon almendarizi</emphasis>
</taxonomicName>
can be distinguished from all its congeners by the combinations of the following characters: abdomen totally covered with small to medium-sized, irregularly shaped plates (
<emphasis id="CA8D4E24FFF71E54FACBFCD0EF6A49F8" box="[1361,1392,871,896]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
abdomen partially covered, in
<taxonomicName id="3FF9E9B5FFF71E54FE43FC3CE88849DD" baseAuthorityName="Kner" baseAuthorityYear="1853" box="[473,658,907,933]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="CA8D4E24FFF71E54FE43FC3CE88849DD" box="[473,658,907,933]" italics="true" pageId="2" pageNumber="79">P. platycephalus</emphasis>
</taxonomicName>
and
<taxonomicName id="3FF9E9B5FFF71E54FD51FC3CE97B49DC" baseAuthorityName="Delsman" baseAuthorityYear="1941" box="[715,865,907,932]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="amazonum">
<emphasis id="CA8D4E24FFF71E54FD51FC3CE97B49DC" box="[715,865,907,932]" italics="true" pageId="2" pageNumber="79">P. amazonum</emphasis>
</taxonomicName>
or covered by one central row of plates wide and rectangular, in
<taxonomicName id="3FF9E9B5FFF71E54FED9FC18EBD849B1" authorityName="Isbrucker" authorityYear="1975" box="[323,450,943,969]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="thorectes">
<emphasis id="CA8D4E24FFF71E54FED9FC18EBD849B1" box="[323,450,943,969]" italics="true" pageId="2" pageNumber="79">P thorectes</emphasis>
</taxonomicName>
); absence of small, bony plates, anterior to gill openings (
<emphasis id="CA8D4E24FFF71E54FBFBFC18EE9A49B0" box="[1121,1152,943,968]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
presence of one to three small plates in front of the gill openings, in
<taxonomicName id="3FF9E9B5FFF71E54FD1DFC64E8EB4995" baseAuthorityName="Gunther" baseAuthorityYear="1868" box="[647,753,979,1005]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="lamina">
<emphasis id="CA8D4E24FFF71E54FD1DFC64E8EB4995" box="[647,753,979,1005]" italics="true" pageId="2" pageNumber="79">P. lamina</emphasis>
</taxonomicName>
); 12–15 coalescing lateral scutes (
<emphasis id="CA8D4E24FFF71E54FBE2FC64EE8D4994" box="[1144,1175,979,1004]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
<quantity id="3F013FD3FFF71E54FB04FC64EF184995" box="[1182,1282,979,1005]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.334" metricValueMin="5.08" pageId="2" pageNumber="79" unit="in" value="20.5" valueMax="21.0" valueMin="20.0">20–21 in</quantity>
<taxonomicName id="3FF9E9B5FFF71E54FA93FC64EF964995" authorityName="Isbrucker" authorityYear="1975" box="[1289,1420,979,1005]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="thorectes">
<emphasis id="CA8D4E24FFF71E54FA93FC64EF964995" box="[1289,1420,979,1005]" italics="true" pageId="2" pageNumber="79">P. thorectes</emphasis>
</taxonomicName>
); thoracic plates (between pectoral and pelvic fins) fold sideways (
<emphasis id="CA8D4E24FFF71E54FCE7FC40E9814E68" box="[893,923,1015,1040]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
thoracic plates do not fold in
<taxonomicName id="3FF9E9B5FFF71E54FB6FFC40EF914E68" baseAuthorityName="Delsman" baseAuthorityYear="1941" box="[1269,1419,1015,1040]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="amazonum">
<emphasis id="CA8D4E24FFF71E54FB6FFC40EF914E68" box="[1269,1419,1015,1040]" italics="true" pageId="2" pageNumber="79">P. amazonum</emphasis>
</taxonomicName>
); eyes relatively smaller, its diameter without notch fits 12.0–12.7 times in HL (
<emphasis id="CA8D4E24FFF71E54FC67FBACEE0E4E4C" box="[1021,1044,1051,1076]" italics="true" pageId="2" pageNumber="79">vs</emphasis>
. 8.3–11.8 times in HL in
<taxonomicName id="3FF9E9B5FFF71E54FAAAFBACEAA44E20" authorityName="Isbrucker &amp; Nijssen" authorityYear="1978" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="apithanos">
<emphasis id="CA8D4E24FFF71E54FAAAFBACEAA44E20" italics="true" pageId="2" pageNumber="79">P. apithanos</emphasis>
</taxonomicName>
, 7.1–9.0 times in HL in
<taxonomicName id="3FF9E9B5FFF71E54FE4DFB88E8584E21" baseAuthorityName="Gunther" baseAuthorityYear="1868" box="[471,578,1087,1113]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="lamina">
<emphasis id="CA8D4E24FFF71E54FE4DFB88E8584E21" box="[471,578,1087,1113]" italics="true" pageId="2" pageNumber="79">P. lamina</emphasis>
</taxonomicName>
, 8.2–8.7 times in HL in
<taxonomicName id="3FF9E9B5FFF71E54FCC1FB88E9D44E21" baseAuthorityName="Regan" baseAuthorityYear="1904" box="[859,974,1087,1113]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="CA8D4E24FFF71E54FCC1FB88E9D44E21" box="[859,974,1087,1113]" italics="true" pageId="2" pageNumber="79">P. laticeps</emphasis>
</taxonomicName>
and 8.1–9.2 times in HL in
<taxonomicName id="3FF9E9B5FFF71E54FA8BFB88EF914E21" authorityName="Rojas, Provenzano &amp; Ramirez" authorityYear="2019" box="[1297,1419,1087,1113]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="unillano">
<emphasis id="CA8D4E24FFF71E54FA8BFB88EF914E21" box="[1297,1419,1087,1113]" italics="true" pageId="2" pageNumber="79">P. unillano</emphasis>
</taxonomicName>
); body less wide at anal-fin origin 10.4%–12.0% SL (
<emphasis id="CA8D4E24FFF71E54FD76FBD4E9104E04" box="[748,778,1123,1148]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
13.4%–17.7% SL in
<taxonomicName id="3FF9E9B5FFF71E54FB98FBD4EE934E05" authorityName="Isbrucker &amp; Nijssen" authorityYear="1978" box="[1026,1161,1123,1149]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="apithanos">
<emphasis id="CA8D4E24FFF71E54FB98FBD4EE934E05" box="[1026,1161,1123,1149]" italics="true" pageId="2" pageNumber="79">P apithanos</emphasis>
</taxonomicName>
, 12.8%–13.9% SL in
<taxonomicName id="3FF9E9B5FFF71E54FA12FBD4EAF44ED9" baseAuthorityName="Regan" baseAuthorityYear="1904" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="CA8D4E24FFF71E54FA12FBD4EAF44ED9" italics="true" pageId="2" pageNumber="79">P. laticeps</emphasis>
</taxonomicName>
, 11.9%–17.1% SL in
<taxonomicName id="3FF9E9B5FFF71E54FE7DFB30E87A4ED9" authorityName="Rojas, Provenzano &amp; Ramirez" authorityYear="2019" box="[487,608,1159,1185]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="unillano">
<emphasis id="CA8D4E24FFF71E54FE7DFB30E87A4ED9" box="[487,608,1159,1185]" italics="true" pageId="2" pageNumber="79">P. unillano</emphasis>
</taxonomicName>
); six or seven wide and dark transverse bands, posterior to dorsal-fin (
<emphasis id="CA8D4E24FFF71E54FAE4FB30EF864ED8" box="[1406,1436,1159,1184]" italics="true" pageId="2" pageNumber="79">vs.</emphasis>
bands absent in
<taxonomicName id="3FF9E9B5FFF71E54FED3FB1CEBA14EBD" baseAuthorityName="Regan" baseAuthorityYear="1904" box="[329,443,1195,1221]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="CA8D4E24FFF71E54FED3FB1CEBA14EBD" box="[329,443,1195,1221]" italics="true" pageId="2" pageNumber="79">P. laticeps</emphasis>
</taxonomicName>
,
<taxonomicName id="3FF9E9B5FFF71E54FE52FB1CE85A4EBD" authorityName="Rojas, Provenzano &amp; Ramirez" authorityYear="2019" box="[456,576,1195,1221]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="unillano">
<emphasis id="CA8D4E24FFF71E54FE52FB1CE85A4EBD" box="[456,576,1195,1221]" italics="true" pageId="2" pageNumber="79">P. unillano</emphasis>
</taxonomicName>
, and medium to large sized
<taxonomicName id="3FF9E9B5FFF71E54FCE3FB1CE9FB4EBD" baseAuthorityName="Gunther" baseAuthorityYear="1868" box="[889,993,1195,1221]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="lamina">
<emphasis id="CA8D4E24FFF71E54FCE3FB1CE9FB4EBD" box="[889,993,1195,1221]" italics="true" pageId="2" pageNumber="79">P. lamina</emphasis>
</taxonomicName>
(over
<quantity id="3F013FD3FFF71E54FBB3FB1CEE6D4EBD" box="[1065,1143,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="79" unit="mm" value="60.0">60 mm</quantity>
SL) or no more than three or four transverse dark bands posterior to dorsal-fin in
<taxonomicName id="3FF9E9B5FFF71E54FD6FFB78E9674E91" authorityName="Isbrucker &amp; Nijssen" authorityYear="1978" box="[757,893,1231,1257]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="apithanos">
<emphasis id="CA8D4E24FFF71E54FD6FFB78E9674E91" box="[757,893,1231,1257]" italics="true" pageId="2" pageNumber="79">P. apithanos</emphasis>
</taxonomicName>
, or up to eight in very small sized (below
<quantity id="3F013FD3FFF71E54FAD4FB79EF874E91" box="[1358,1437,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="79" unit="mm" value="55.0">55 mm</quantity>
SL) of
<taxonomicName id="3FF9E9B5FFF71E54FF7DFB44EB4A4F75" baseAuthorityName="Gunther" baseAuthorityYear="1868" box="[231,336,1267,1293]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="79" phylum="Chordata" rank="species" species="lamina">
<emphasis id="CA8D4E24FFF71E54FF7DFB44EB4A4F75" box="[231,336,1267,1293]" italics="true" pageId="2" pageNumber="79">P. lamina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B0E3C1BDFFF71E52FF5DFAA1E9D84862" lastPageId="4" lastPageNumber="81" pageId="2" pageNumber="79" type="description">
<paragraph id="F8469236FFF71E54FF5DFAA1E9B64C71" blockId="2.[151,1437,727,2013]" pageId="2" pageNumber="79">
<emphasis id="CA8D4E24FFF71E54FF5DFAA1EB414F48" bold="true" box="[199,347,1302,1328]" pageId="2" pageNumber="79">Description:</emphasis>
Morphometric data presented in table 1. Head and body very depressed, caudal peduncle long, narrow and very depressed, without adipose-fin (
<figureCitation id="60C28EB3FFF71E54FD2EFA8CE8EF4F2D" box="[692,757,1339,1365]" captionStart="FIGURE 1" captionStartId="3.[151,250,1638,1662]" captionTargetBox="[151,1436,309,1615]" captionTargetId="figure-82@3.[151,1436,309,1615]" captionTargetPageId="3" captionText="FIGURE 1. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 1 cm. Photos by P. Argüello." figureDoi="http://doi.org/10.5281/zenodo.6488191" httpUri="https://zenodo.org/record/6488191/files/figure.png" pageId="2" pageNumber="79">Fig. 1</figureCitation>
). Maximum body depth at dorsal-fin origin, or slightly ahead and maximum width at cleithrum, becoming narrower posteriorly, gradually, to caudal-fin origin. Dorsal profile of body from tip of snout through anterior border of eye, straight and scarcely sloping up, from this point to dorsal-fin origin, straight and gently inclined or gently convex, then descending straight to caudal-fin origin. Ventral profile of body flat and straight. Pectoral-fin insertions at vertical through posterior margin of orbit. Dorsal-fin origin opposite pelvic-fin insertions. Anal-fin origin at lateral plate number 11 (
<figureCitation id="60C28EB3FFF71E54FCFAFA58E9BB4C71" box="[864,929,1519,1545]" captionStart="FIGURE 1" captionStartId="3.[151,250,1638,1662]" captionTargetBox="[151,1436,309,1615]" captionTargetId="figure-82@3.[151,1436,309,1615]" captionTargetPageId="3" captionText="FIGURE 1. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 1 cm. Photos by P. Argüello." figureDoi="http://doi.org/10.5281/zenodo.6488191" httpUri="https://zenodo.org/record/6488191/files/figure.png" pageId="2" pageNumber="79">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="F8469236FFF71E54FF5DF9A4EE054CC5" blockId="2.[151,1437,727,2013]" pageId="2" pageNumber="79">
Head shape in dorsal view an acute triangle with straight edges. Snout little projected, with rounded tip (
<figureCitation id="60C28EB3FFF71E54FAF1F9A4EABF4C29" captionStart="FIGURE 1" captionStartId="3.[151,250,1638,1662]" captionTargetBox="[151,1436,309,1615]" captionTargetId="figure-82@3.[151,1436,309,1615]" captionTargetPageId="3" captionText="FIGURE 1. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 1 cm. Photos by P. Argüello." figureDoi="http://doi.org/10.5281/zenodo.6488191" httpUri="https://zenodo.org/record/6488191/files/figure.png" pageId="2" pageNumber="79">Figs 1</figureCitation>
,
<figureCitation id="60C28EB3FFF71E54FF2FF980EADF4C29" box="[181,197,1591,1617]" captionStart="FIGURE 2" captionStartId="4.[151,250,1956,1980]" captionTargetBox="[151,1435,669,1933]" captionTargetId="figure-228@4.[151,1435,669,1933]" captionTargetPageId="4" captionText="FIGURE 2. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Ventral view, detail of head contour, mouth and abdomen cover. Scale bar 5 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.6488193" httpUri="https://zenodo.org/record/6488193/files/figure.png" pageId="2" pageNumber="79">2</figureCitation>
,
<figureCitation id="60C28EB3FFF71E54FF49F980EAF94C29" box="[211,227,1591,1617]" captionStart="FIGURE 3" captionStartId="6.[151,250,1777,1801]" captionTargetBox="[161,1426,813,1754]" captionTargetId="figure-344@6.[161,1426,813,1754]" captionTargetPageId="6" captionText="FIGURE 3. a) P. almendarizi, MEPN 17903, 87.9 mm SL, holotype. b) P. apithanos, MEPN 18374, 97.5 mm SL. c) P. lamina, MEPN 15095, 101.9 mm SL. d) P. laticeps, MCP 9308, 126.1 mm SL. Dorsal views, scale bar 1 cm. Photos a), b) and c) by P. Argüello; d) by B. Calegari." figureDoi="http://doi.org/10.5281/zenodo.6488195" httpUri="https://zenodo.org/record/6488195/files/figure.png" pageId="2" pageNumber="79">3</figureCitation>
). Eyes located dorsally, orbits with shallow, ventral and posterior notches. Keels weak with parallel path, from nostrils, through eyes to anterior tip of supraoccipital. Over the supraoccipital, keels convergent posteriorly, becoming closer and again parallel to posterior tip of supraoccipital. Pre-dorsal area with three single plates, first and second with low parallel keels, third with single low keel at midline (
<figureCitation id="60C28EB3FFF71E54FC57F914EE144CC5" box="[973,1038,1699,1725]" captionStart="FIGURE 1" captionStartId="3.[151,250,1638,1662]" captionTargetBox="[151,1436,309,1615]" captionTargetId="figure-82@3.[151,1436,309,1615]" captionTargetPageId="3" captionText="FIGURE 1. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 1 cm. Photos by P. Argüello." figureDoi="http://doi.org/10.5281/zenodo.6488191" httpUri="https://zenodo.org/record/6488191/files/figure.png" pageId="2" pageNumber="79">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="F8469236FFF71E55FF5DF970EF234B65" blockId="2.[151,1437,727,2013]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="80" pageId="2" pageNumber="79">
Ventral surface of head naked except for plates along border and snout, no plates anterior to the gill openings. Branchiostegal membrane smooth and uniform, without wrinkled flap on anterior margin or any protuberance or fold (
<figureCitation id="60C28EB3FFF71E54FF4EF8B8EB004D51" box="[212,282,1807,1833]" captionStart="FIGURE 2" captionStartId="4.[151,250,1956,1980]" captionTargetBox="[151,1435,669,1933]" captionTargetId="figure-228@4.[151,1435,669,1933]" captionTargetPageId="4" captionText="FIGURE 2. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Ventral view, detail of head contour, mouth and abdomen cover. Scale bar 5 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.6488193" httpUri="https://zenodo.org/record/6488193/files/figure.png" pageId="2" pageNumber="79">Fig. 2</figureCitation>
). Mouth ventral with lips laminar and thin. Upper lip very narrow, imperceptible, its border with conical, small, elongated, unbranched barblets, decreasing in size toward middle. Surface of upper lip with sparse small papillae laterally. Border of upper lip continuous with maxillary barbels that extend to gill opening. Maxillary barbel with small, conical, unbranched barblets. Lower lip wide, its edge with elongated, branched, and conical barblets; barblets slightly shorter at middle. Lower lip surface covered with short, fleshy, thick papillae, sometimes slightly elongated (
<figureCitation id="60C28EB3FFF71E54FE89F874EB4D4DA5" box="[275,343,1987,2013]" captionStart="FIGURE 2" captionStartId="4.[151,250,1956,1980]" captionTargetBox="[151,1435,669,1933]" captionTargetId="figure-228@4.[151,1435,669,1933]" captionTargetPageId="4" captionText="FIGURE 2. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Ventral view, detail of head contour, mouth and abdomen cover. Scale bar 5 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.6488193" httpUri="https://zenodo.org/record/6488193/files/figure.png" pageId="2" pageNumber="79">Fig. 2</figureCitation>
). Teeth present in both jaws, minute but evident, same length, asymmetrical bicuspids with medial cusp more developed and spoon-shaped, lateral cusp very small, sometimes not visible, pointed, apex yellow or golden, stalk white. Premaxillary teeth 2–5, dentary teeth 5–7 (
<figureCitation id="60C28EB3FFF61E55FCFAFF0CE9BC4AAD" box="[864,934,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1956,1980]" captionTargetBox="[151,1435,669,1933]" captionTargetId="figure-228@4.[151,1435,669,1933]" captionTargetPageId="4" captionText="FIGURE 2. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Ventral view, detail of head contour, mouth and abdomen cover. Scale bar 5 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.6488193" httpUri="https://zenodo.org/record/6488193/files/figure.png" pageId="3" pageNumber="80">Fig. 2</figureCitation>
). Buccal ornamentation composed of two or three small, fleshy, cylindrical, elongate and unbranched barblets, at distal side of each premaxillae. Inside mouth, behind premaxillaries, just at middle, with single long, fleshy, cylindrical and unbranched barblet (
<figureCitation id="60C28EB3FFF61E55FB7DFEB4EF364B65" box="[1255,1324,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1956,1980]" captionTargetBox="[151,1435,669,1933]" captionTargetId="figure-228@4.[151,1435,669,1933]" captionTargetPageId="4" captionText="FIGURE 2. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Ventral view, detail of head contour, mouth and abdomen cover. Scale bar 5 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.6488193" httpUri="https://zenodo.org/record/6488193/files/figure.png" pageId="3" pageNumber="80">Fig. 2</figureCitation>
).
</paragraph>
<caption id="AC86C2BEFFF61E55FF0DF9D1EBA24CDA" ID-DOI="http://doi.org/10.5281/zenodo.6488191" ID-Zenodo-Dep="6488191" httpUri="https://zenodo.org/record/6488191/files/figure.png" pageId="3" pageNumber="80" startId="3.[151,250,1638,1662]" targetBox="[151,1436,309,1615]" targetPageId="3">
<paragraph id="F8469236FFF61E55FF0DF9D1EBA24CDA" blockId="3.[151,1436,1638,1698]" pageId="3" pageNumber="80">
<emphasis id="CA8D4E24FFF61E55FF0DF9D1EB0E4C06" bold="true" box="[151,276,1638,1662]" pageId="3" pageNumber="80">FIGURE 1.</emphasis>
<taxonomicName id="3FF9E9B5FFF61E55FE81F9D1E8864C05" authority=", MEPN" authorityName="MEPN" box="[283,668,1638,1662]" class="Actinopterygii" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="80" phylum="Chordata" rank="species" species="almendarizi">
<emphasis id="CA8D4E24FFF61E55FE81F9D1E8534C06" box="[283,585,1638,1662]" italics="true" pageId="3" pageNumber="80">Pseudohemiodon almendarizi</emphasis>
, MEPN
</taxonomicName>
17903, 87.9 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 1 cm. Photos by P. Argüello.
</paragraph>
</caption>
<paragraph id="F8469236FFF61E55FF5DF967EEDA4D4A" blockId="3.[151,1437,1744,2023]" pageId="3" pageNumber="80">
Abdomen completely covered with irregularly polygonal-shaped plates, plates small to medium-sized, smaller near pectoral girdle. Anus projected as very small tube, urogenital papilla not visible, apparently attached to posterior surface of anal tube. Anus delimited by a narrow naked area, surrounded by plates (
<figureCitation id="60C28EB3FFF61E55FBF4F8AFEEA94D4A" box="[1134,1203,1816,1842]" captionStart="FIGURE 2" captionStartId="4.[151,250,1956,1980]" captionTargetBox="[151,1435,669,1933]" captionTargetId="figure-228@4.[151,1435,669,1933]" captionTargetPageId="4" captionText="FIGURE 2. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Ventral view, detail of head contour, mouth and abdomen cover. Scale bar 5 mm. Photo by F. Provenzano." figureDoi="http://doi.org/10.5281/zenodo.6488193" httpUri="https://zenodo.org/record/6488193/files/figure.png" pageId="3" pageNumber="80">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="F8469236FFF61E52FF5DF88BEF344AAD" blockId="3.[151,1437,1744,2023]" lastBlockId="4.[151,1437,151,645]" lastPageId="4" lastPageNumber="81" pageId="3" pageNumber="80">
Body with 33 plates in median lateral series, 13 coalescent plates (double keel) and 20 posterior plates (one keel). Seven to eight thoracic plates (between posterior end of pectoral-fin base and origin of pelvic-fin base). Postdorsal plates 20–22 and post-anal plates 17–20. Four plates border dorsal-fin base; and two or three border anal-fin base. Dorsal-fin rays i,7; pectoral-fin rays i,6; pelvic-fin rays i,5; anal-fin rays i,5; and caudal-fin rays i,10,i. Tip of pelvic-fin surpasses anal-fin origin. Caudal-fin slightly bifurcated, with unbranched rays longer than branched. Upper unbranched caudal-fin rays projecting as long filaments (but broken in both specimens). In available material, first unbranched rays (spines) of dorsal, pectoral, pelvic and anal-fins not elongated as filaments (
<figureCitation id="60C28EB3FFF11E52FB41FF0CEF074AAD" box="[1243,1309,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1638,1662]" captionTargetBox="[151,1436,309,1615]" captionTargetId="figure-82@3.[151,1436,309,1615]" captionTargetPageId="3" captionText="FIGURE 1. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 1 cm. Photos by P. Argüello." figureDoi="http://doi.org/10.5281/zenodo.6488191" httpUri="https://zenodo.org/record/6488191/files/figure.png" pageId="4" pageNumber="81">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="F8469236FFF11E52FF5DFF69EF654BA9" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="81">
<emphasis id="CA8D4E24FFF11E52FF5DFF69EB954A81" bold="true" box="[199,399,222,249]" pageId="4" pageNumber="81">Color in alcohol:</emphasis>
In specimens preserved in 70% alcohol, dorsal surface of head and body brown or yellowish brown, uniform. On head to end of dorsal-fin base, light and dark areas randomly positioned (
<figureCitation id="60C28EB3FFF11E52FB2DFEB4EEE24B65" box="[1207,1272,259,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1638,1662]" captionTargetBox="[151,1436,309,1615]" captionTargetId="figure-82@3.[151,1436,309,1615]" captionTargetPageId="3" captionText="FIGURE 1. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 1 cm. Photos by P. Argüello." figureDoi="http://doi.org/10.5281/zenodo.6488191" httpUri="https://zenodo.org/record/6488191/files/figure.png" pageId="4" pageNumber="81">Fig. 1</figureCitation>
). Six or seven dark, transverse bands, decreasing in width and intensity posteriorly located from posterior end of dorsal-fin base to caudal peduncle. Ventral surface of head and body, whitish or yellowish, uniform (
<figureCitation id="60C28EB3FFF11E52FBAEFEFCEE6F4B1D" box="[1076,1141,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1638,1662]" captionTargetBox="[151,1436,309,1615]" captionTargetId="figure-82@3.[151,1436,309,1615]" captionTargetPageId="3" captionText="FIGURE 1. Pseudohemiodon almendarizi, MEPN 17903, 87.9 mm SL, holotype. Lateral, dorsal and ventral views. Scale bar 1 cm. Photos by P. Argüello." figureDoi="http://doi.org/10.5281/zenodo.6488191" httpUri="https://zenodo.org/record/6488191/files/figure.png" pageId="4" pageNumber="81">Fig. 1</figureCitation>
). Dorsal and pectoral-fins rays and interradial membrane dark blackish. Pectoral-fin spine paler brown or yellow with four diffuse (faint) dark bands. Pelvic-fin rays and interradial membranes light brown or yellow with dark area at middle. Anal-fin whitish or yellowish, uniform. Caudal-fin rays dark brown becoming pale brown, posterior as the interradial membrane.
</paragraph>
<paragraph id="F8469236FFF11E52FF5DFE6DE9D84862" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="81">
<emphasis id="CA8D4E24FFF11E52FF5DFE6DE8104B8C" bold="true" box="[199,522,474,500]" pageId="4" pageNumber="81">Geographical distribution:</emphasis>
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The
<specimenCount id="EEFF59BFFFF11E52FDD0FE6CE8EE4B8D" box="[586,756,475,501]" pageId="4" pageNumber="81" type="generic">two specimens</specimenCount>
came from
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, near
<collectingMunicipality id="1822084CFFF11E52FBE5FE6CEED24B8D" box="[1151,1224,475,501]" pageId="4" pageNumber="81">Puerto</collectingMunicipality>
<collectingRegion id="3A3D5CD4FFF11E52FB48FE6CEF4E4B8D" box="[1234,1364,475,501]" isEnumeration="true" pageId="4" pageNumber="81">Loja, Napo</collectingRegion>
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system,
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Amazon
<location id="FD26C4EDFFF11E52FECDFE48EB8F4861" LSID="urn:lsid:plazi:treatment:70502320FFF71E52FF0DFDF1EAD048FD:FD26C4EDFFF11E52FECDFE48EB8F4861" box="[343,405,511,537]" county="Aguarico River" latitude="-0.8825" longLatPrecision="21" longitude="-75.225" municipality="Puerto" name="River Basin" pageId="4" pageNumber="81" stateProvince="Loja, Napo">River</location>
</location>
Basin
</location>
, approx.
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<geoCoordinate id="9DCDF4F1FFF11E52FD48FE49E97B4861" box="[722,865,510,537]" degrees="75" direction="west" minutes="13" orientation="longitude" pageId="4" pageNumber="81" precision="15" seconds="30" value="-75.225">75°13’30”W</geoCoordinate>
(
<figureCitation id="60C28EB3FFF11E52FCEAFE48E9AE4862" box="[880,948,511,538]" captionStart="FIGURE 6" captionStartId="10.[151,250,1868,1892]" captionTargetBox="[151,1435,455,1842]" captionTargetId="figure-139@10.[151,1435,453,1844]" captionTargetPageId="10" captionText="FIGURE 6. Map of Ecuador showing the geographic distribution of Pseudohemiodon species. P. almendarizi, red star, type locality, P. apithanos, yellow triangles, P. lamina, black circles. Some symbols may represent more than one lot." figureDoi="http://doi.org/10.5281/zenodo.6488201" httpUri="https://zenodo.org/record/6488201/files/figure.png" pageId="4" pageNumber="81">Fig. 6</figureCitation>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="B0E3C1BDFFF11E52FF5DFD94EAD048FD" pageId="4" pageNumber="81" type="etymology">
<paragraph id="F8469236FFF11E52FF5DFD94EAD048FD" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="81">
<emphasis id="CA8D4E24FFF11E52FF5DFD94EB484845" bold="true" box="[199,338,547,573]" pageId="4" pageNumber="81">Etymology:</emphasis>
The specific name honors Ana de Lourdes Almendáriz, in recognition of her significant contributions to the
<collectingCountry id="80EED2A6FFF11E52FE86FDF0EB634819" box="[284,377,583,609]" name="Ecuador" pageId="4" pageNumber="81">Ecuador</collectingCountry>
herpetofauna knowledge, and her enthusiasm and friendship for many years. Noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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