443 lines
54 KiB
XML
443 lines
54 KiB
XML
<document id="3EF78F33E659975F0D210396B3538EAA" ID-DOI="10.5252/g2016n3a5" ID-ISSN="1638-9395" ID-Zenodo-Dep="5208548" ID-ZooBank="urn:lsid:zoobank.org:pub:EC4EA93F-AB85-4C7F-BA09-9CB0AB2A1B38" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629142939015" checkinUser="felipe" docAuthor="Perna, Rafael La & Vazzana, Angelo" docDate="2016" docId="D313D00CCE66FFD9338DB109FC4EFDCF" docLanguage="en" docName="Geodiversitas.38.3.451-461.pdf" docOrigin="Geodiversitas 38 (3)" docSource="http://dx.doi.org/10.5252/g2016n3a5" docStyle="DocumentStyle:4DA100580BA89F561A6D37CADC50A8FD.3:Geodiversitas.2015-2017.journal_article.type2" docStyleId="4DA100580BA89F561A6D37CADC50A8FD" docStyleName="Geodiversitas.2015-2017.journal_article.type2" docStyleVersion="3" docTitle="Marginella seguenzai Perna & Vazzana 2016, n. sp." docType="treatment" docUuid="936AFB6B-77AB-41E4-BB35-65EFC0A95E39" docUuidSource="ZooBank" docVersion="4" lastPageNumber="458" masterDocId="2F2AA874CE62FFDE3057B53DFF91FFFF" masterDocTitle="On the last occurrence of Marginella Lamarck, 1799 (Gastropoda, Marginellidae) in the Mediterranean: description of a new species from the Early Pleistocene and paleoceanographic implications" masterLastPageNumber="461" masterPageNumber="451" pageNumber="455" updateTime="1699061339206" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="7D4C1A13C6F51CFA304F45EB4AE2AA11">On the last occurrence of Marginella Lamarck, 1799 (Gastropoda, Marginellidae) in the Mediterranean: description of a new species from the Early Pleistocene and paleoceanographic implications</mods:title>
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<mods:roleTerm id="AFAB9DD64C64981ED01ADE195C7D23AA">Author</mods:roleTerm>
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<mods:namePart id="A82E6775157A918D4417D57438FAD48C">Perna, Rafael La</mods:namePart>
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<mods:name id="F48658BF14DE83ABEFBBE72B249D31C1" type="personal">
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<mods:roleTerm id="140652457B786CE4283417DD3EA57BD5">Author</mods:roleTerm>
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<mods:namePart id="467250680BC563CF04CDBDD7C97634D5">Vazzana, Angelo</mods:namePart>
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<mods:typeOfResource id="93CF3AFA12FE4D46D9DDF6C480AD3256">text</mods:typeOfResource>
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<mods:title id="9FBA533181E3E93F5FBEA4AE8198B9E6">Geodiversitas</mods:title>
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<mods:part id="5A446A8CF4E584F404717CDEA026ED45">
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<mods:date id="00A9432AF7ACFD14C8850DF54E54ADD5">2016</mods:date>
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<mods:number id="494006B6BC5F4573391EF8BAB380F001">2016-09-30</mods:number>
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<mods:number id="4952DBD593AD6DCD904699090BB0868D">38</mods:number>
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<mods:url id="7AF5C45EDEF2C4F8221D65EBE6AC750A">http://dx.doi.org/10.5252/g2016n3a5</mods:url>
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<mods:identifier id="6A89DA76F80190B0D99033DCFAD8232A" type="DOI">10.5252/g2016n3a5</mods:identifier>
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<treatment id="D313D00CCE66FFD9338DB109FC4EFDCF" LSID="urn:lsid:zoobank.org:act:936AFB6B-77AB-41E4-BB35-65EFC0A95E39" httpUri="http://treatment.plazi.org/id/D313D00CCE66FFD9338DB109FC4EFDCF" lastPageId="7" lastPageNumber="458" pageId="4" pageNumber="455">
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<subSubSection id="13A03291CE66FFDA338DB109FA93FBB0" box="[986,1282,1076,1103]" pageId="4" pageNumber="455" type="nomenclature">
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<paragraph id="5B05611ACE66FFDA338DB109FA93FBB0" blockId="4.[986,1282,1076,1135]" box="[986,1282,1076,1103]" pageId="4" pageNumber="455">
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<heading id="004DD676CE66FFDA338DB109FA93FBB0" box="[986,1282,1076,1103]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="455" reason="2">
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<taxonomicName id="9CBA1A99CE66FFDA338DB109FB51FBB1" ID-CoL="735MG" authority="Perna & Vazzana, 2016" authorityName="Perna & Vazzana" authorityYear="2016" box="[986,1216,1076,1103]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="455" phylum="Mollusca" rank="species" species="seguenzai" status="sp. nov.">
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<emphasis id="69CEBD08CE66FFDA338DB109FB51FBB1" bold="true" box="[986,1216,1076,1103]" italics="true" pageId="4" pageNumber="455">Marginella seguenzai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72FD0073CE66FFDA3490B108FA93FBB0" box="[1223,1282,1077,1103]" pageId="4" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="13A03291CE66FFDA33ACB169FB71FB90" box="[1019,1248,1108,1135]" pageId="4" pageNumber="455" type="description">
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<paragraph id="5B05611ACE66FFDA33ACB169FB71FB90" blockId="4.[986,1282,1076,1135]" box="[1019,1248,1108,1135]" pageId="4" pageNumber="455">
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(
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<figureCitation id="C3817D9FCE66FFDA3454B169FBD6FB91" box="[1027,1095,1108,1134]" captionStart="FIG" captionStartId="3.[306,317,1728,1746]" captionTargetBox="[132,1455,216,1689]" captionTargetId="figure-147@3.[132,1455,216,1689]" captionTargetPageId="3" captionText="FIG. 2. — Sample of 36 shells used for morphometry and shape analysis, with identification number. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5208552" httpUri="https://zenodo.org/record/5208552/files/figure.png" pageId="4" pageNumber="455">Figs 2</figureCitation>
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;
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<figureCitation id="C3817D9FCE66FFDA3404B169FBCEFB90" box="[1107,1119,1108,1135]" captionStart="FIG" captionStartId="6.[132,143,1701,1719]" captionTargetBox="[137,1450,215,1664]" captionTargetId="figure-144@6.[136,1450,214,1664]" captionTargetPageId="6" captionText="FIG. 5. — Marginella seguenzai n. sp.: A-C, holotype, H 9.3 mm (MZB60202); D, paratype 1, H 10.6 mm (MZB60203); E-G, paratype 2, H 9.7 mm (MZB60203); H, I, paratype 3, H 9.0 mm (MZB60203); J, paratype 4, H 8.4 mm (MBMPRC); L, paratype 5, H 11.8 mm (MBMPRC). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5208558" httpUri="https://zenodo.org/record/5208558/files/figure.png" pageId="4" pageNumber="455">5</figureCitation>
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A-L; 6A, B)
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</paragraph>
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</subSubSection>
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<subSubSection id="13A03291CE66FFDA337AB1A8FB8CFACE" pageId="4" pageNumber="455" type="materials_examined">
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<paragraph id="5B05611ACE66FFDA337AB1A8FB8CFACE" blockId="4.[811,1457,1172,1329]" pageId="4" pageNumber="455">
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<materialsCitation id="EBD26B47CE66FFDA337AB1A8FB71FB52" box="[813,1248,1172,1197]" collectionCode="TYPE, MATERIAL" pageId="4" pageNumber="455" specimenCount="1" typeStatus="holotype">
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||
<collectionCode id="3DABF9DFCE66FFDA337AB1A8FC26FB54" box="[813,951,1173,1196]" pageId="4" pageNumber="455">TYPE MATERIAL</collectionCode>
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. —
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<typeStatus id="8401DFB8CE66FFDA3389B1A9FBA7FB53" box="[990,1078,1172,1196]" pageId="4" pageNumber="455" type="holotype">Holotype</typeStatus>
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||
and
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<specimenCount id="4DBCAA93CE66FFDA3436B1A9FB71FB52" box="[1121,1248,1172,1197]" pageId="4" pageNumber="455" type="generic" typeStatus="paratypes">five paratypes</specimenCount>
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||
</materialsCitation>
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||
.
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<materialsCitation id="EBD26B47CE66FFDA34B1B1A9FBB7FB39" accessionNumber="MZB60202" collectionCode="MZB" pageId="4" pageNumber="455" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="8401DFB8CE66FFDA34B1B1A9FAD0FB53" box="[1254,1345,1172,1196]" pageId="4" pageNumber="455" type="holotype">Holotype</typeStatus>
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:H
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||
<quantity id="9C42CCFFCE66FFDA3537B1A9FA3AFB53" box="[1376,1451,1172,1196]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="4" pageNumber="455" unit="mm" value="9.3">9.3 mm</quantity>
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, D
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||
<quantity id="9C42CCFFCE66FFDA3311B193FC04FB39" box="[838,917,1198,1222]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="455" unit="mm" value="5.3">5.3 mm</quantity>
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||
(
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||
<collectionCode id="3DABF9DFCE66FFDA33F4B193FC4DFB39" box="[931,988,1198,1222]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="455" type="Museum">MZB</collectionCode>
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||
60202)
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||
</materialsCitation>
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||
. —
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||
<materialsCitation id="EBD26B47CE66FFDA3406B192FBA4FB1E" accessionNumber="MZB60203" collectionCode="MZB" pageId="4" pageNumber="455" specimenCount="2" typeStatus="paratype">
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||
<typeStatus id="8401DFB8CE66FFDA3406B192FB21FB38" box="[1105,1200,1199,1223]" pageId="4" pageNumber="455" type="paratype">Paratypes</typeStatus>
|
||
:
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||
<typeStatus id="8401DFB8CE66FFDA34EDB192FA9CFB38" box="[1210,1293,1199,1223]" pageId="4" pageNumber="455" type="paratype">Paratype</typeStatus>
|
||
1, H
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||
<quantity id="9C42CCFFCE66FFDA351CB193FA38FB39" box="[1355,1449,1198,1222]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="4" pageNumber="455" unit="mm" value="10.6">10.6 mm</quantity>
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||
, D
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||
<quantity id="9C42CCFFCE66FFDA331CB1F4FC0FFB1E" box="[843,926,1225,1249]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="4" pageNumber="455" unit="mm" value="5.9">5.9 mm</quantity>
|
||
(
|
||
<collectionCode id="3DABF9DFCE66FFDA33F9B1F4FC7BFB1E" box="[942,1002,1225,1249]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="455" type="Museum">MZB</collectionCode>
|
||
60203)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="EBD26B47CE66FFDA3414B1F7FC26FB04" accessionNumber="MZB60203" collectionCode="MZB" pageId="4" pageNumber="455" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="8401DFB8CE66FFDA3414B1F7FB0AFB1D" box="[1091,1179,1226,1250]" pageId="4" pageNumber="455" type="paratype">Paratype</typeStatus>
|
||
2, H
|
||
<quantity id="9C42CCFFCE66FFDA3489B1F4FAA5FB1E" box="[1246,1332,1225,1249]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="4" pageNumber="455" unit="mm" value="9.7">9.7 mm</quantity>
|
||
, D
|
||
<quantity id="9C42CCFFCE66FFDA3509B1F4FA20FB1E" box="[1374,1457,1225,1249]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="4" pageNumber="455" unit="mm" value="5.4">5.4 mm</quantity>
|
||
(
|
||
<collectionCode id="3DABF9DFCE66FFDA3361B1DEFCFEFB04" box="[822,879,1251,1275]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="455" type="Museum">MZB</collectionCode>
|
||
60203)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="EBD26B47CE66FFDA3395B1D9FAE2FB04" accessionNumber="MZB60203" box="[962,1395,1251,1276]" collectionCode="MZB" pageId="4" pageNumber="455" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="8401DFB8CE66FFDA3395B1D9FB84FB03" box="[962,1045,1252,1276]" pageId="4" pageNumber="455" type="paratype">Paratype</typeStatus>
|
||
3, H 9.0, D
|
||
<quantity id="9C42CCFFCE66FFDA34C0B1DEFB75FB04" box="[1175,1252,1251,1275]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="455" unit="mm" value="5.3">5.3 mm</quantity>
|
||
(
|
||
<collectionCode id="3DABF9DFCE66FFDA34A6B1DEFABBFB04" box="[1265,1322,1251,1275]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="455" type="Museum">MZB</collectionCode>
|
||
60203)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="EBD26B47CE66FFDA352AB1D9FB54FAE9" collectionCode="MBMPRC" pageId="4" pageNumber="455" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="8401DFB8CE66FFDA352AB1D9FCC7FAE9" pageId="4" pageNumber="455" type="paratype">Paratype</typeStatus>
|
||
4, H
|
||
<quantity id="9C42CCFFCE66FFDA33DDB1C3FC46FAEA" box="[906,983,1278,1302]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="4" pageNumber="455" unit="mm" value="8.4">8.4 mm</quantity>
|
||
, D
|
||
<quantity id="9C42CCFFCE66FFDA33A0B1C3FBD3FAEA" box="[1015,1090,1278,1302]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="455" unit="mm" value="4.8">4.8 mm</quantity>
|
||
(
|
||
<collectionCode id="3DABF9DFCE66FFDA341AB1C3FB2CFAE9" box="[1101,1213,1278,1302]" pageId="4" pageNumber="455">MBMPRC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="EBD26B47CE66FFDA349BB1C3FB88FACE" collectionCode="MBMPRC" pageId="4" pageNumber="455" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="8401DFB8CE66FFDA349BB1C3FA8FFAE9" box="[1228,1310,1278,1302]" pageId="4" pageNumber="455" type="paratype">Paratype</typeStatus>
|
||
5, H
|
||
<quantity id="9C42CCFFCE66FFDA3506B1C3FA3DFAEA" box="[1361,1452,1278,1302]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="4" pageNumber="455" unit="mm" value="11.8">11.8 mm</quantity>
|
||
, D
|
||
<quantity id="9C42CCFFCE66FFDA3312B025FC02FACF" box="[837,915,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="4" pageNumber="455" unit="mm" value="6.7">6.7 mm</quantity>
|
||
(
|
||
<collectionCode id="3DABF9DFCE66FFDA33F7B024FB83FACE" box="[928,1042,1305,1329]" pageId="4" pageNumber="455">MBMPRC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A03291CE66FFDA337AB06DFB73FA7E" pageId="4" pageNumber="455" type="etymology">
|
||
<paragraph id="5B05611ACE66FFDA337AB06DFB73FA7E" blockId="4.[813,1455,1359,1409]" pageId="4" pageNumber="455">ETYMOLOGY. — After Giuseppe Seguenza (Messina 1833-1889), eminent Italian geologist and paleontologist.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A03291CE66FFDA337AB09CFC33F98A" pageId="4" pageNumber="455" type="materials_examined">
|
||
<paragraph id="5B05611ACE66FFDA337AB09CFC21FA2C" blockId="4.[812,1455,1440,1491]" pageId="4" pageNumber="455">
|
||
<collectionCode id="3DABF9DFCE66FFDA337AB09CFC1EFA48" box="[813,911,1441,1464]" pageId="4" pageNumber="455">MATERIAL</collectionCode>
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||
EXAMINED. — Over 80 shells, all from the
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||
<typeStatus id="8401DFB8CE66FFDA3563B09CFACFFA46" box="[1332,1374,1441,1465]" pageId="4" pageNumber="455">type</typeStatus>
|
||
horizon (
|
||
<collectionCode id="3DABF9DFCE66FFDA3364B086FC34FA2C" box="[819,933,1467,1491]" pageId="4" pageNumber="455">MBMPRC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5B05611ACE66FFDA337AB0CFFC1CF9DB" blockId="4.[811,1455,1521,1572]" pageId="4" pageNumber="455">
|
||
<collectionCode id="3DABF9DFCE66FFDA337AB0CFFC2DF9F7" box="[813,956,1522,1545]" pageId="4" pageNumber="455">
|
||
<typeStatus id="8401DFB8CE66FFDA337AB0CFFCCCF9F8" box="[813,861,1522,1545]" pageId="4" pageNumber="455">TYPE</typeStatus>
|
||
HORIZON
|
||
</collectionCode>
|
||
. — Bed b in the outcrop of
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<taxonomicName id="9CBA1A99CE66FFDA3493B0CCFAC8F9F6" authority=", Reggio" authorityName="Reggio" box="[1220,1369,1521,1545]" class="Gastropoda" family="Camaenidae" genus="Gallina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="455" phylum="Mollusca" rank="genus">Gallina, Reggio</taxonomicName>
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||
<collectingRegion id="997EAFF8CE66FFDA3509B0CCFA3EF9F6" box="[1374,1455,1521,1545]" country="Italy" name="Calabria" pageId="4" pageNumber="455">Calabria</collectingRegion>
|
||
(
|
||
<figureCitation id="C3817D9FCE66FFDA3362B331FC10F9DB" box="[821,897,1548,1572]" captionStart="FIG" captionStartId="2.[132,143,891,909]" captionTargetBox="[132,1455,215,849]" captionTargetId="figure-588@2.[132,1455,215,848]" captionTargetPageId="2" captionText="FIG. 1. — A, Map of the Messina Strait;B, location of the outcrop;C, schematic stratigraphy of the outcrop (asterisk indicates the sampled bed).Abbreviations:see text." figureDoi="http://doi.org/10.5281/zenodo.5208550" httpUri="https://zenodo.org/record/5208550/files/figure.png" pageId="4" pageNumber="455">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5B05611ACE66FFDA337AB37EFC33F98A" blockId="4.[813,1455,1602,1653]" pageId="4" pageNumber="455">
|
||
<collectionCode id="3DABF9DFCE66FFDA337AB37EFC56F9A6" box="[813,967,1603,1626]" pageId="4" pageNumber="455">
|
||
<typeStatus id="8401DFB8CE66FFDA337AB37EFCCEF9A7" box="[813,863,1603,1626]" pageId="4" pageNumber="455">TYPE</typeStatus>
|
||
LOCALITY
|
||
</collectionCode>
|
||
. —
|
||
<taxonomicName id="9CBA1A99CE66FFDA33AEB37FFB08F9A5" authority=", Reggio" authorityName="Reggio" box="[1017,1177,1602,1626]" class="Gastropoda" family="Camaenidae" genus="Gallina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="455" phylum="Mollusca" rank="genus">Gallina, Reggio</taxonomicName>
|
||
<collectingRegion id="997EAFF8CE66FFDA34F5B37FFB6AF9A5" box="[1186,1275,1602,1626]" country="Italy" name="Calabria" pageId="4" pageNumber="455">Calabria</collectingRegion>
|
||
, Calabrian, Early Pleistocene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A03291CE66FFDA337AB3A9FC65F8CF" pageId="4" pageNumber="455" type="diagnosis">
|
||
<paragraph id="5B05611ACE66FFDA337AB3A9FC65F8CF" blockId="4.[811,1457,1683,1840]" pageId="4" pageNumber="455">
|
||
DIAGNOSIS. — A small
|
||
<taxonomicName id="9CBA1A99CE66FFDA344BB3AEFB41F954" box="[1052,1232,1683,1707]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="455" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="69CEBD08CE66FFDA344BB3AEFB15F954" box="[1052,1156,1683,1707]" italics="true" pageId="4" pageNumber="455">Marginella</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, biconical, moderately inflated, with last whorl making about 85% of shell height, maximum diameter at shell mid-height. Apex blunt, spire low, with two flat whorls. Aperture narrow, elongate, somewhat parallel-sided, with four columellar plications. Outer lip internally smooth, thickened by a wide outer rim.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A03291CE66FFDB337AB272FDC3F91C" lastPageId="5" lastPageNumber="456" pageId="4" pageNumber="455" type="distribution">
|
||
<paragraph id="5B05611ACE66FFDA337AB272FCEBF87E" blockId="4.[811,1457,1870,2027]" pageId="4" pageNumber="455">
|
||
DISTRIBUTION. — The new species is only known from the
|
||
<typeStatus id="8401DFB8CE66FFDA35D2B272FA3EF898" box="[1413,1455,1871,1895]" pageId="4" pageNumber="455">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="5B05611ACE66FFDA337AB2B9FA3EF814" blockId="4.[811,1457,1870,2027]" pageId="4" pageNumber="455">
|
||
Because of the allochthonous character of the fauna from
|
||
<taxonomicName id="9CBA1A99CE66FFDA3537B2BEFA3AF864" box="[1376,1451,1923,1947]" class="Gastropoda" family="Camaenidae" genus="Gallina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="455" phylum="Mollusca" rank="genus">Gallina</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CBA1A99CE66FFDA337BB2A2FC38F84A" authority="Perna & Vazzana, 2016" authorityName="Perna & Vazzana" authorityYear="2016" box="[812,937,1950,1974]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="455" phylum="Mollusca" rank="species" species="seguenzai" status="sp. nov.">
|
||
<emphasis id="69CEBD08CE66FFDA337BB2A2FC38F84A" box="[812,937,1950,1974]" italics="true" pageId="4" pageNumber="455">M. seguenzai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72FD0073CE66FFDA33E2B2A3FC7CF84A" box="[949,1005,1950,1973]" pageId="4" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
|
||
could be a bathyal species, or a shelf species whose shells were transported to bathyal depths.
|
||
<taxonomicName id="9CBA1A99CE66FFDA34AAB285FAF4F82F" authorityName="Lamarck" authorityYear="1799" box="[1277,1381,1976,2000]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="455" phylum="Mollusca" rank="genus">
|
||
<emphasis id="69CEBD08CE66FFDA34AAB285FAF4F82F" box="[1277,1381,1976,2000]" italics="true" pageId="4" pageNumber="455">Marginella</emphasis>
|
||
</taxonomicName>
|
||
mainly has a shallow water distribution, but some species are widely dis-
|
||
</paragraph>
|
||
<paragraph id="5B05611ACE67FFDB317EB043FA34FA6F" blockId="5.[297,1445,1406,1424]" box="[297,1445,1406,1424]" pageId="5" pageNumber="456">–0.06 –0.03 0 0.03 0.06</paragraph>
|
||
<caption id="0FC53192CE67FFDB314BB08AFA85FA36" ID-DOI="http://doi.org/10.5281/zenodo.5208556" ID-Zenodo-Dep="5208556" box="[284,1300,1462,1481]" httpUri="https://zenodo.org/record/5208556/files/figure.png" pageId="5" pageNumber="456" startId="5.[284,295,1463,1481]" targetBox="[136,1452,215,1389]" targetPageId="5">
|
||
<paragraph id="5B05611ACE67FFDB314BB08AFA85FA36" blockId="5.[284,1300,1462,1481]" box="[284,1300,1462,1481]" pageId="5" pageNumber="456">FIG. 4. — PCA based on harmonic decomposition of shell outline: plane PCA1-PCA2. Identification numbers as in Figure 2</paragraph>
|
||
</caption>
|
||
<paragraph id="5B05611ACE67FFDB30D3B32FFDC3F91C" blockId="5.[131,776,1554,1763]" pageId="5" pageNumber="456">
|
||
tributed bathymetrically (e.g.
|
||
<bibRefCitation id="3F2B1CEBCE67FFDB31FDB32FFDE6F9D5" author="GOUD J. & NEEFS J." box="[426,631,1554,1578]" pageId="5" pageNumber="456" pagination="25 - 50" refId="ref6984" refString="GOUD J. & NEEFS J. 1996. - The larger marginelliform gastropods (Cystiscidae and Marginellidae) collected during the CANCAP and MAURITANIA expeditions in the southern part of the North Atlantic Ocean. The genera Persicula, Prunum, Dentimargo, Marginella and Glabella. Vita Marina 43 (3 - 4): 25 - 50." type="journal article" year="1996">Goud & Neefs 1996</bibRefCitation>
|
||
), while few are only known from bathyal depths (e.g.
|
||
<bibRefCitation id="3F2B1CEBCE67FFDB31A1B310FD01F9BB" author="COOMANS H. E." box="[502,656,1580,1605]" pageId="5" pageNumber="456" pagination="99 - 101" refId="ref6243" refString="COOMANS H. E. 1975. - Marginella orstomi, a new species from deeper water off the coast of West Africa (Gastropoda, Marginellidae). Bulletin Zoologisch Museum Universiteit Van Amsterdam 4 (12): 99 - 101." type="journal article" year="1975">Coomans 1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3F2B1CEBCE67FFDB32CEB310FE91F9A0" author="BOUCHET P. & WAREN A." pageId="5" pageNumber="456" pagination="123 - 296" refId="ref6057" refString="BOUCHET P. & WAREN A. 1985. - Revision of the Northeast Atlantic bathyal and abyssal Neogastropoda excluding Turridae (Mollusca, Gastropoda). Bollettino Malacologico suppl. 1: 123 - 296." type="journal article" year="1985">Bouchet & Warén 1985</bibRefCitation>
|
||
). Considering the general good preservation of shells and the prevalent composition of the molluscan assemblage, it can be assumed that
|
||
<taxonomicName id="9CBA1A99CE67FFDB3102B340FE40F96B" authorityName="Perna & Vazzana" authorityYear="2016" box="[341,465,1661,1684]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="seguenzai">
|
||
<emphasis id="69CEBD08CE67FFDB3102B340FE40F96B" box="[341,465,1661,1684]" italics="true" pageId="5" pageNumber="456">M. seguenzai</emphasis>
|
||
</taxonomicName>
|
||
n. sp was a deep water species. This hypothesis is also supported by close similarities occurring between the new species and a Recent deep water species from the NE Atlantic, as discussed in Distribution part.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A03291CE67FFDB30D3B22FFBCBF814" pageId="5" pageNumber="456" type="description">
|
||
<paragraph id="5B05611ACE67FFDB30D3B22FFE8AF8D5" blockId="5.[131,776,1810,2028]" box="[132,283,1810,1836]" pageId="5" pageNumber="456">DESCRIPTION</paragraph>
|
||
<paragraph id="5B05611ACE67FFDB30D3B20CFBCBF814" blockId="5.[131,776,1810,2028]" lastBlockId="5.[811,1457,1554,2027]" pageId="5" pageNumber="456">
|
||
Shell small, up to
|
||
<emphasis id="69CEBD08CE67FFDB310FB20FFEF8F8B4" box="[344,361,1842,1867]" italics="true" pageId="5" pageNumber="456">c.</emphasis>
|
||
<quantity id="9C42CCFFCE67FFDB3126B20FFE52F8B3" box="[369,451,1842,1868]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="456" unit="mm" value="12.0">12 mm</quantity>
|
||
in height, moderately solid, biconical, with last whorl making about 85% of total height; maximum diameter at about mid-height to slightly posterior; apex blunt. Protoconch not well distinct from teleoconch, smooth, formed by about one whorl. Spire low, consisting of about two flat-sided whorls. Sutures shallow, poorly impressed, distinct. Sometimes, the spire is slightly narrower, forming a stepped suture with last whorl. Last whorl regularly rounded, gently tapering anteriorly, rarely forming a poorly distinct basal constriction. Aperture elongate, narrow, somewhat parallel-sided, taking about 70% of shell height. Outer lip internally smooth; externally bordered by a well distinct, wide rim, making the lip uniformly thick; straight for most of its length, posteriorly arched; anterior notch shallow, moderately wide. Four columellar plications, all well distinct, not markedly prominent, the posterior one smaller. Sometimes a fifth false plication is present, as an elongate, weak tubercle or as a thin plait similar to a true plication. Columellar and parietal callus rather wide, forming a thin glossy lining near aperture, slightly thicker posteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="0FC53192CE64FFD830D3B398FB4EF932" ID-DOI="http://doi.org/10.5281/zenodo.5208558" ID-Zenodo-Dep="5208558" httpUri="https://zenodo.org/record/5208558/files/figure.png" pageId="6" pageNumber="457" startId="6.[132,143,1701,1719]" targetBox="[137,1450,215,1664]" targetPageId="6">
|
||
<paragraph id="5B05611ACE64FFD830D3B398FB4EF932" blockId="6.[132,1457,1700,1741]" pageId="6" pageNumber="457">
|
||
FIG. 5. —
|
||
<taxonomicName id="9CBA1A99CE64FFD83082B398FE12F948" authority="Perna & Vazzana, 2016" authorityName="Perna & Vazzana" authorityYear="2016" box="[213,387,1701,1719]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="457" phylum="Mollusca" rank="species" species="seguenzai" status="sp. nov.">
|
||
<emphasis id="69CEBD08CE64FFD83082B398FE12F948" box="[213,387,1701,1719]" italics="true" pageId="6" pageNumber="457">Marginella seguenzai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72FD0073CE64FFD831DFB398FE2AF948" box="[392,443,1701,1719]" pageId="6" pageNumber="457" rank="species">n. sp.</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="69CEBD08CE64FFD83195B398FE75F949" bold="true" box="[450,484,1700,1719]" pageId="6" pageNumber="457">A -C</emphasis>
|
||
, holotype, H 9.3 mm (MZB60202);
|
||
<emphasis id="69CEBD08CE64FFD8335DB398FC89F948" bold="true" box="[778,792,1701,1719]" pageId="6" pageNumber="457">D</emphasis>
|
||
, paratype 1, H 10.6 mm (MZB60203);
|
||
<emphasis id="69CEBD08CE64FFD8340CB398FBEDF949" bold="true" box="[1115,1148,1700,1719]" pageId="6" pageNumber="457">E -G</emphasis>
|
||
, paratype 2, H 9.7 mm (MZB60203);
|
||
<emphasis id="69CEBD08CE64FFD830D3B386FF03F932" bold="true" box="[132,146,1723,1741]" pageId="6" pageNumber="457">H</emphasis>
|
||
,
|
||
<emphasis id="69CEBD08CE64FFD830CBB386FF33F932" bold="true" box="[156,162,1723,1741]" pageId="6" pageNumber="457">I</emphasis>
|
||
, paratype 3, H 9.0 mm (MZB60203);
|
||
<emphasis id="69CEBD08CE64FFD83180B386FE70F932" bold="true" box="[471,481,1723,1741]" pageId="6" pageNumber="457">J</emphasis>
|
||
, paratype 4, H 8.4 mm (MBMPRC);
|
||
<emphasis id="69CEBD08CE64FFD83359B386FC88F932" bold="true" box="[782,793,1723,1741]" pageId="6" pageNumber="457">L</emphasis>
|
||
, paratype 5, H 11.8 mm (MBMPRC). Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="13A03291CE64FFD930D3B22FFE07F814" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457" type="discussion">
|
||
<paragraph id="5B05611ACE64FFD830D3B22FFF79F8D5" blockId="6.[131,776,1810,2028]" box="[132,232,1810,1836]" pageId="6" pageNumber="457">REMARKS</paragraph>
|
||
<paragraph id="5B05611ACE64FFD830D3B20CFC66F874" blockId="6.[131,776,1810,2028]" lastBlockId="6.[811,1457,1810,2028]" pageId="6" pageNumber="457">
|
||
The genus
|
||
<taxonomicName id="9CBA1A99CE64FFD830A2B20CFEF9F8B4" authorityName="Lamarck" authorityYear="1799" box="[245,360,1841,1867]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="457" phylum="Mollusca" rank="genus">
|
||
<emphasis id="69CEBD08CE64FFD830A2B20CFEF9F8B4" box="[245,360,1841,1867]" italics="true" pageId="6" pageNumber="457">Marginella</emphasis>
|
||
</taxonomicName>
|
||
typically includes species with brightly coloured shells, with a range of patterns (variously combined bands, flames, lineoles, blotches, dots, etc.) (
|
||
<bibRefCitation id="3F2B1CEBCE64FFD83234B24CFF51F854" author="GOUD J. & NEEFS J." pageId="6" pageNumber="457" pagination="25 - 50" refId="ref6984" refString="GOUD J. & NEEFS J. 1996. - The larger marginelliform gastropods (Cystiscidae and Marginellidae) collected during the CANCAP and MAURITANIA expeditions in the southern part of the North Atlantic Ocean. The genera Persicula, Prunum, Dentimargo, Marginella and Glabella. Vita Marina 43 (3 - 4): 25 - 50." type="journal article" year="1996">Goud & Neefs 1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3F2B1CEBCE64FFD83099B2ACFE1DF853" author="COSSIGNANI T." box="[206,396,1937,1964]" pageId="6" pageNumber="457" refId="ref6349" refString="COSSIGNANI T. 2006. - Marginellidae and Cystiscidae of the World. Atlas with over 2,600 photos. L'Informatore Piceno, 408 p." type="book" year="2006">Cossignani 2006</bibRefCitation>
|
||
), but several colourless species are also known, mainly from deep waters (
|
||
<bibRefCitation id="3F2B1CEBCE64FFD8326CB28CFF51F813" author="BOUCHET P. & WAREN A." pageId="6" pageNumber="457" pagination="123 - 296" refId="ref6057" refString="BOUCHET P. & WAREN A. 1985. - Revision of the Northeast Atlantic bathyal and abyssal Neogastropoda excluding Turridae (Mollusca, Gastropoda). Bollettino Malacologico suppl. 1: 123 - 296." type="journal article" year="1985">Bouchet & Warén 1985</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3F2B1CEBCE64FFD8309DB2ECFE32F813" author="GOUD J. & NEEFS J." box="[202,419,2001,2028]" pageId="6" pageNumber="457" pagination="25 - 50" refId="ref6984" refString="GOUD J. & NEEFS J. 1996. - The larger marginelliform gastropods (Cystiscidae and Marginellidae) collected during the CANCAP and MAURITANIA expeditions in the southern part of the North Atlantic Ocean. The genera Persicula, Prunum, Dentimargo, Marginella and Glabella. Vita Marina 43 (3 - 4): 25 - 50." type="journal article" year="1996">Goud& Neefs 1996</bibRefCitation>
|
||
). No preserved colour pattern was observed in the material of
|
||
<taxonomicName id="9CBA1A99CE64FFD83403B22FFB4AF8D4" authority="Perna & Vazzana, 2016" authorityName="Perna & Vazzana" authorityYear="2016" box="[1108,1243,1810,1835]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="457" phylum="Mollusca" rank="species" species="seguenzai" status="sp. nov.">
|
||
<emphasis id="69CEBD08CE64FFD83403B22FFB4AF8D4" box="[1108,1243,1810,1835]" italics="true" pageId="6" pageNumber="457">M. seguenzai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72FD0073CE64FFD834B5B22FFA8EF8D3" box="[1250,1311,1810,1836]" pageId="6" pageNumber="457" rank="species">n. sp.</taxonomicNameLabel>
|
||
, either under daily light, or under UV light, but the uniform milky aspect of the shell surface suggests a colourless, or lightly coloured, original condition.
|
||
</paragraph>
|
||
<paragraph id="5B05611ACE64FFD93314B2AFFE5AFC50" blockId="6.[811,1457,1810,2028]" lastBlockId="7.[130,777,821,2028]" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457">
|
||
In addition to the variability in size and shape, another variable character was observed: some shells (about 20%) show a “false fifth plication” (
|
||
<bibRefCitation id="3F2B1CEBCE64FFD8347CB2ECFAD8F813" author="COOVERT G. A. & COOVERT H. K." box="[1067,1353,2001,2028]" pageId="6" pageNumber="457" pagination="43 - 110" refId="ref6313" refString="COOVERT G. A. & COOVERT H. K. 1995. - Revision of the supraspecific classification of marginelliform gastropods. The Nautilus 109 (2 - 3): 43 - 110." type="journal article" year="1995">Coovert & Coovert, 1995</bibRefCitation>
|
||
). In most cases it consists of a weak, elongate tubercle near the fourth (posterior) plication (
|
||
<figureCitation id="C3817D9FCE65FFD93109B668FE3DFC90" box="[350,428,853,879]" captionStart="FIG" captionStartId="7.[132,143,707,725]" captionTargetBox="[132,775,215,669]" captionTargetId="figure-886@7.[132,775,215,669]" captionTargetPageId="7" captionText="FIG. 6. — Marginella seguenzai n. sp.: shells with a fifth false plication (arrows): A, poorly developed fifth plication; B, well developed fifth plication. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5208560" httpUri="https://zenodo.org/record/5208560/files/figure.png" pageId="7" pageNumber="458">Fig.6A</figureCitation>
|
||
). Rarely, the false plication is more elongate and stronger, resembling a true, internally continuous, columellar plication ((
|
||
<figureCitation id="C3817D9FCE65FFD93139B6A8FE2FFC50" box="[366,446,917,943]" captionStart="FIG" captionStartId="7.[132,143,707,725]" captionTargetBox="[132,775,215,669]" captionTargetId="figure-886@7.[132,775,215,669]" captionTargetPageId="7" captionText="FIG. 6. — Marginella seguenzai n. sp.: shells with a fifth false plication (arrows): A, poorly developed fifth plication; B, well developed fifth plication. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5208560" httpUri="https://zenodo.org/record/5208560/files/figure.png" pageId="7" pageNumber="458">Fig. 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="0FC53192CE65FFD930D3B7FEFF2AFCFD" ID-DOI="http://doi.org/10.5281/zenodo.5208560" ID-Zenodo-Dep="5208560" httpUri="https://zenodo.org/record/5208560/files/figure.png" pageId="7" pageNumber="458" startId="7.[132,143,707,725]" targetBox="[132,775,215,669]" targetPageId="7">
|
||
<paragraph id="5B05611ACE65FFD930D3B7FEFF2AFCFD" blockId="7.[132,777,707,770]" pageId="7" pageNumber="458">
|
||
FIG. 6. —
|
||
<taxonomicName id="9CBA1A99CE65FFD93084B7FEFEEEFD2A" authority="Perna & Vazzana, 2016" authorityName="Perna & Vazzana" authorityYear="2016" box="[211,383,707,725]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="seguenzai" status="sp. nov.">
|
||
<emphasis id="69CEBD08CE65FFD93084B7FEFEEEFD2A" box="[211,383,707,725]" italics="true" pageId="7" pageNumber="458">Marginella seguenzai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72FD0073CE65FFD931D3B7FEFE24FD2A" box="[388,437,707,725]" pageId="7" pageNumber="458" rank="species">n. sp.</taxonomicNameLabel>
|
||
: shells with a fifth false plication (arrows):
|
||
<emphasis id="69CEBD08CE65FFD930D3B7E4FF00FD14" bold="true" box="[132,145,729,747]" pageId="7" pageNumber="458">A</emphasis>
|
||
, poorly developed fifth plication;
|
||
<emphasis id="69CEBD08CE65FFD931F2B7E4FE23FD14" bold="true" box="[421,434,729,747]" pageId="7" pageNumber="458">B</emphasis>
|
||
, well developed fifth plication. Scale bar: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5B05611ACE65FFD930CCB688FD3BFBF0" blockId="7.[130,777,821,2028]" pageId="7" pageNumber="458">
|
||
In some cases, the transition between spire and last whorl appears somewhat stepped, the spire being slightly narrower than the adjacent last whorl, as seen in
|
||
<figureCitation id="C3817D9FCE65FFD93248B6C9FD37FBF0" box="[543,678,1012,1039]" captionStart="FIG" captionStartId="6.[132,143,1701,1719]" captionTargetBox="[137,1450,215,1664]" captionTargetId="figure-144@6.[136,1450,214,1664]" captionTargetPageId="6" captionText="FIG. 5. — Marginella seguenzai n. sp.: A-C, holotype, H 9.3 mm (MZB60202); D, paratype 1, H 10.6 mm (MZB60203); E-G, paratype 2, H 9.7 mm (MZB60203); H, I, paratype 3, H 9.0 mm (MZB60203); J, paratype 4, H 8.4 mm (MBMPRC); L, paratype 5, H 11.8 mm (MBMPRC). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5208558" httpUri="https://zenodo.org/record/5208558/files/figure.png" pageId="7" pageNumber="458">Figure 5H, I</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5B05611ACE65FFD930CCB128FDC2FB70" blockId="7.[130,777,821,2028]" pageId="7" pageNumber="458">
|
||
None of the
|
||
<taxonomicName id="9CBA1A99CE65FFD93149B129FE4DFBD1" box="[286,476,1044,1070]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="69CEBD08CE65FFD93149B129FE1FFBD1" box="[286,398,1044,1070]" italics="true" pageId="7" pageNumber="458">Marginella</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
known from the Pliocene of Estepona (
|
||
<bibRefCitation id="3F2B1CEBCE65FFD930A5B109FE23FBB1" author="MUNIZ SOLIS R." box="[242,434,1076,1102]" pageId="7" pageNumber="458" pagination="263 - 306" refId="ref7806" refString="MUNIZ SOLIS R. 2002. - Estudio de las Familias Cystiscidae Stimpson, 1865 y Marginellidae Fleming, 1828 (Gastropoda, Neogastropoda) del Plioceno de Estepona (Malaga, Espana). Pliocenica 2: 263 - 306." type="journal article" year="2002">Muñiz Solis 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3F2B1CEBCE65FFD931EBB109FD1EFBB0" author="LANDAU B. & LA PERNA R. & MARQUET R." box="[444,655,1076,1103]" pageId="7" pageNumber="458" pagination="22 - 60" refId="ref7334" refString="LANDAU B., LA PERNA R. & MARQUET R. 2006 a. - The Early Pliocene Gastropoda (Mollusca) of Estepona, Southern Spain. Part 10. Marginellidae, Cystiscidae. Palaeontos 9: 22 - 60." type="journal article" year="2006">
|
||
Landau
|
||
<emphasis id="69CEBD08CE65FFD93244B108FDD7FBB1" box="[531,582,1076,1102]" italics="true" pageId="7" pageNumber="458">et al.</emphasis>
|
||
2006a
|
||
</bibRefCitation>
|
||
) is particularly similar to the new species, all markedly differing in some respects (mainly size, shape and robustness).
|
||
</paragraph>
|
||
<paragraph id="5B05611ACE65FFD930CCB1A9FE07F814" blockId="7.[130,777,821,2028]" pageId="7" pageNumber="458">
|
||
Few of the many species known from Eastern Atlantic, deserve a comparison with the new species, such as
|
||
<taxonomicName id="9CBA1A99CE65FFD932CFB189FE1BFB11" authority="Jousseaume, 1884" authorityName="Jousseaume" authorityYear="1884" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="gloriosa">
|
||
<emphasis id="69CEBD08CE65FFD932CFB189FF5CFB11" italics="true" pageId="7" pageNumber="458">Marginella gloriosa</emphasis>
|
||
Jousseaume, 1884
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CBA1A99CE65FFD931C7B1E9FD32FB11" authority="Locard, 1897" authorityName="Locard" authorityYear="1897" box="[400,675,1236,1262]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="marocana">
|
||
<emphasis id="69CEBD08CE65FFD931C7B1E9FD84FB12" box="[400,533,1236,1261]" italics="true" pageId="7" pageNumber="458">M. marocana</emphasis>
|
||
Locard, 1897
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CBA1A99CE65FFD932FEB1E9FECBFAF1" authority="Dautzenberg, 1910" authorityName="Dautzenberg" authorityYear="1910" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="bavay">
|
||
<emphasis id="69CEBD08CE65FFD932FEB1E9FC96FB11" box="[681,775,1236,1262]" italics="true" pageId="7" pageNumber="458">M. bavay</emphasis>
|
||
Dautzenberg, 1910
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CBA1A99CE65FFD9313EB1C9FC92FAF1" authority="Bouchet & Waren, 1985" authorityName="Bouchet & Waren" authorityYear="1985" box="[361,771,1267,1294]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="aronnax">
|
||
<emphasis id="69CEBD08CE65FFD9313EB1C9FE79FAF2" box="[361,488,1268,1293]" italics="true" pageId="7" pageNumber="458">M. aronnax</emphasis>
|
||
<bibRefCitation id="3F2B1CEBCE65FFD931B9B1CEFC92FAF1" author="BOUCHET P. & WAREN A." box="[494,771,1267,1294]" pageId="7" pageNumber="458" pagination="123 - 296" refId="ref6057" refString="BOUCHET P. & WAREN A. 1985. - Revision of the Northeast Atlantic bathyal and abyssal Neogastropoda excluding Turridae (Mollusca, Gastropoda). Bollettino Malacologico suppl. 1: 123 - 296." type="journal article" year="1985">Bouchet & Warén, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CBA1A99CE65FFD930D1B029FE47FAD1" authority="(Bozzetti, 1995)" baseAuthorityName="Bozzetti" baseAuthorityYear="1995" box="[134,470,1299,1326]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="colomborum">
|
||
<emphasis id="69CEBD08CE65FFD930D1B029FEB9FAD2" box="[134,296,1299,1325]" italics="true" pageId="7" pageNumber="458">M. colomborum</emphasis>
|
||
(
|
||
<bibRefCitation id="3F2B1CEBCE65FFD93164B02EFE5FFAD1" author="BOZZETTI L." box="[307,462,1299,1326]" pageId="7" pageNumber="458" pagination="48 - 52" refId="ref6094" refString="BOZZETTI L. 1995. - Tre nuove specie della famiglia Marginellidae Fleming, 1828 (Gastropoda, Prosobranchia) from the Indian and Atlantic Ocean. World Shells 14: 48 - 52 and errata corrige 15: 62." type="journal article" year="1995">Bozzetti, 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9CBA1A99CE65FFD9325FB029FF2CFAB2" authority="Goud & Neefs, 1996" authorityName="Goud & Neefs" authorityYear="1996" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="gilva">
|
||
<emphasis id="69CEBD08CE65FFD9325FB029FDCCFAD2" box="[520,605,1299,1325]" italics="true" pageId="7" pageNumber="458">M. gilva</emphasis>
|
||
<bibRefCitation id="3F2B1CEBCE65FFD93233B02EFF2CFAB2" author="GOUD J. & NEEFS J." pageId="7" pageNumber="458" pagination="25 - 50" refId="ref6984" refString="GOUD J. & NEEFS J. 1996. - The larger marginelliform gastropods (Cystiscidae and Marginellidae) collected during the CANCAP and MAURITANIA expeditions in the southern part of the North Atlantic Ocean. The genera Persicula, Prunum, Dentimargo, Marginella and Glabella. Vita Marina 43 (3 - 4): 25 - 50." type="journal article" year="1996">Goud & Neefs, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
, but in most cases there are evident differences involving size and shape (convexity of spire whorls, spire elevation, shell slenderness, shape of outer lip, etc.). Of these species, only
|
||
<taxonomicName id="9CBA1A99CE65FFD930D1B0A9FEB4FA52" baseAuthorityName="Bozzetti" baseAuthorityYear="1995" box="[134,293,1427,1453]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="colomborum">
|
||
<emphasis id="69CEBD08CE65FFD930D1B0A9FEB4FA52" box="[134,293,1427,1453]" italics="true" pageId="7" pageNumber="458">M.colomborum</emphasis>
|
||
</taxonomicName>
|
||
shows an interesting overlap of characters with the new species. It was described from the Josephine Bank (
|
||
<bibRefCitation id="3F2B1CEBCE65FFD93283B08EFF63FA11" author="BOZZETTI L." pageId="7" pageNumber="458" pagination="48 - 52" refId="ref6094" refString="BOZZETTI L. 1995. - Tre nuove specie della famiglia Marginellidae Fleming, 1828 (Gastropoda, Prosobranchia) from the Indian and Atlantic Ocean. World Shells 14: 48 - 52 and errata corrige 15: 62." type="journal article" year="1995">Bozzetti 1995</bibRefCitation>
|
||
), a seamount in the North-East Atlantic, off Cape
|
||
<collectingCountry id="23AD218ACE65FFD930D2B0CEFE91F9F2" box="[133,256,1523,1549]" name="Saint Vincent and the Grenadines" pageId="7" pageNumber="458">St. Vincent</collectingCountry>
|
||
(
|
||
<bibRefCitation id="3F2B1CEBCE65FFD93147B0CEFE3FF9F2" author="GUBBAY S." box="[272,430,1523,1549]" pageId="7" pageNumber="458" refId="ref7045" refString="GUBBAY S. 2003. - Seamounts of the North East Atlantic. WWF / OASIS, Bremen, 38 p." type="book" year="2003">Gubbay 2003</bibRefCitation>
|
||
). Though originally assigned to the genus
|
||
<taxonomicName id="9CBA1A99CE65FFD930B9B32EFED2F9D3" authorityName="Herrmannsen" authorityYear="1852" box="[238,323,1555,1580]" class="Gastropoda" family="Marginellidae" genus="Prunum" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="genus">
|
||
<emphasis id="69CEBD08CE65FFD930B9B32EFED2F9D3" box="[238,323,1555,1580]" italics="true" pageId="7" pageNumber="458">Prunum</emphasis>
|
||
</taxonomicName>
|
||
, this species must be allocated in the genus
|
||
<taxonomicName id="9CBA1A99CE65FFD930D3B30FFF65F9B3" authorityName="Lamarck" authorityYear="1799" box="[132,244,1586,1612]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="genus">
|
||
<emphasis id="69CEBD08CE65FFD930D3B30FFF65F9B3" box="[132,244,1586,1612]" italics="true" pageId="7" pageNumber="458">Marginella</emphasis>
|
||
</taxonomicName>
|
||
. The species was examined on three shells from the
|
||
<typeStatus id="8401DFB8CE65FFD930D4B36EFF21F992" box="[131,176,1619,1645]" pageId="7" pageNumber="458">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="C3817D9FCE65FFD93143B36FFEC0F992" box="[276,337,1618,1645]" captionStart="FIG" captionStartId="8.[132,143,824,842]" captionTargetBox="[132,1455,215,781]" captionTargetId="figure-764@8.[132,1455,214,781]" captionTargetPageId="8" captionText="FIG. 7. — Marginella colomborum (Bozzetti,1995),Josephine Bank,Seamount 1 Exp.,St.DW37,255-270 m,36°42N,14°18W,4 Oct.1988 (MNHN):A-C, H 10.0 mm; D, H 9.8 mm. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5208562" httpUri="https://zenodo.org/record/5208562/files/figure.png" pageId="7" pageNumber="458">Fig. 7</figureCitation>
|
||
A-D): the shell is colourless (actually very light pinkish, with the outer lip white), height close to that of
|
||
<taxonomicName id="9CBA1A99CE65FFD930F7B3AEFEB4F953" authority="Perna & Vazzana, 2016" authorityName="Perna & Vazzana" authorityYear="2016" box="[160,293,1683,1708]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="seguenzai" status="sp. nov.">
|
||
<emphasis id="69CEBD08CE65FFD930F7B3AEFEB4F953" box="[160,293,1683,1708]" italics="true" pageId="7" pageNumber="458">M. seguenzai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72FD0073CE65FFD9317CB3AEFEF4F952" box="[299,357,1683,1709]" pageId="7" pageNumber="458" rank="species">n. sp.</taxonomicNameLabel>
|
||
also in the range (12.9-
|
||
<quantity id="9C42CCFFCE65FFD93237B3AFFD52F952" box="[608,707,1682,1709]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="7" pageNumber="458" unit="mm" value="11.3">11.3 mm</quantity>
|
||
in the original description, 10.0-
|
||
<quantity id="9C42CCFFCE65FFD931C1B38FFE7AF932" box="[406,491,1714,1741]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="7" pageNumber="458" unit="mm" value="8.8">8.8 mm</quantity>
|
||
in the examined material), outer lip moderately thick, making a broad curve posteriorly, as in the new species. A remarkable difference from
|
||
<taxonomicName id="9CBA1A99CE65FFD932D5B3CEFC97F8F3" authority="Perna & Vazzana, 2016" authorityName="Perna & Vazzana" authorityYear="2016" box="[642,774,1779,1804]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="seguenzai" status="sp. nov.">
|
||
<emphasis id="69CEBD08CE65FFD932D5B3CEFC97F8F3" box="[642,774,1779,1804]" italics="true" pageId="7" pageNumber="458">M. seguenzai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72FD0073CE65FFD930D3B22FFF2EF8D3" box="[132,191,1810,1836]" pageId="7" pageNumber="458" rank="species">n. sp.</taxonomicNameLabel>
|
||
is the last whorl convexity, being
|
||
<taxonomicName id="9CBA1A99CE65FFD93272B22FFD58F8D3" baseAuthorityName="Bozzetti" baseAuthorityYear="1995" box="[549,713,1810,1836]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="colomborum">
|
||
<emphasis id="69CEBD08CE65FFD93272B22FFD58F8D3" box="[549,713,1810,1836]" italics="true" pageId="7" pageNumber="458">M. colomborum</emphasis>
|
||
</taxonomicName>
|
||
more inflated(H/D
|
||
<quantity id="9C42CCFFCE65FFD93140B20FFE04F8B4" box="[279,405,1841,1868]" metricMagnitude="-2" metricUnit="m" metricValue="4.5212" metricValueMax="4.7244" metricValueMin="4.318" pageId="7" pageNumber="458" unit="in" value="1.78" valueMax="1.86" valueMin="1.7">1.70-1.86 in</quantity>
|
||
the examined material) than the new species, apparently without overlap.The shape of the aperture, rather parallel-sided in the new species and widening anteriorly in
|
||
<taxonomicName id="9CBA1A99CE65FFD930C9B2AFFEADF854" baseAuthorityName="Bozzetti" baseAuthorityYear="1995" box="[158,316,1937,1963]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="colomborum">
|
||
<emphasis id="69CEBD08CE65FFD930C9B2AFFEADF854" box="[158,316,1937,1963]" italics="true" pageId="7" pageNumber="458">M. colomborum</emphasis>
|
||
</taxonomicName>
|
||
, is another outstanding difference. Moreover,
|
||
<taxonomicName id="9CBA1A99CE65FFD930D5B28FFEB5F834" baseAuthorityName="Bozzetti" baseAuthorityYear="1995" box="[130,292,1969,1995]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="colomborum">
|
||
<emphasis id="69CEBD08CE65FFD930D5B28FFEB5F834" box="[130,292,1969,1995]" italics="true" pageId="7" pageNumber="458">M. colomborum</emphasis>
|
||
</taxonomicName>
|
||
has slightly convex spire whorls, whereas they are flat in the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A03291CE65FFD93314B5EAFC4EFDCF" pageId="7" pageNumber="458" type="reference_group">
|
||
<paragraph id="5B05611ACE65FFD93314B5EAFC4EFDCF" blockId="7.[811,1457,215,560]" pageId="7" pageNumber="458">
|
||
<taxonomicName id="9CBA1A99CE65FFD93314B5EAFBC5FF0E" ID-CoL="3Y5H8" baseAuthorityName="Bozzetti" baseAuthorityYear="1995" box="[835,1108,215,241]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="colomborum">
|
||
<emphasis id="69CEBD08CE65FFD93314B5EAFBC5FF0E" box="[835,1108,215,241]" italics="true" pageId="7" pageNumber="458">Marginella colomborum</emphasis>
|
||
</taxonomicName>
|
||
has been also recorded from Seine and Ampère, other NE Atlantic seamounts (
|
||
<bibRefCitation id="3F2B1CEBCE65FFD93521B5CAFC37FECD" author="BECK T. & METZGER T. & FREIWALD A." pageId="7" pageNumber="458" refId="ref5918" refString="BECK T., METZGER T. & FREIWALD A. 2005. - BIAS. Biodiversity Inventorial Atlas of Macrobenthic Seamount Animals. OASIS, Hamburg, 118 p." type="book" year="2005">
|
||
Beck
|
||
<emphasis id="69CEBD08CE65FFD93378B425FCF2FECE" box="[815,867,279,305]" italics="true" pageId="7" pageNumber="458">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
). On WoRMS,
|
||
<bibRefCitation id="3F2B1CEBCE65FFD93404B42AFB76FECD" author="GOFAS S." box="[1107,1255,279,306]" pageId="7" pageNumber="458" refId="ref6793" refString="GOFAS S. 2015. - Marginella colomborum (Bozzetti, 1995) in MolluscaBase (2015). Accessed through: World Register of Marine Species at http: // www. marinespecies. org / aphia. php? p = taxdetails & id = 140321 on 2015 - 07 - 17." type="url" year="2015">Gofas (2015)</bibRefCitation>
|
||
added: “Endemic of Josephine, Seine and Ampère seamounts; common on Josephine,
|
||
<quantity id="9C42CCFFCE65FFD933FDB46AFBBDFE8E" box="[938,1068,343,370]" metricMagnitude="2" metricUnit="m" metricValue="3.025" metricValueMax="3.35" metricValueMin="2.7" pageId="7" pageNumber="458" unit="m" value="302.5" valueMax="335.0" valueMin="270.0">270-335 m</quantity>
|
||
, rare on Seine and Ampère; a very similar if not identical species is found in the Italian Pleistocene”. The fossil species to which Gofas refers is
|
||
<taxonomicName id="9CBA1A99CE65FFD93536B4AAFCE5FE2F" ID-CoL="735MG" authority="Perna & Vazzana, 2016" authorityName="Perna & Vazzana" authorityYear="2016" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="seguenzai" status="sp. nov.">
|
||
<emphasis id="69CEBD08CE65FFD93536B4AAFCE5FE2F" italics="true" pageId="7" pageNumber="458">M. seguenzai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72FD0073CE65FFD9332EB48AFC27FE2E" box="[889,950,439,465]" pageId="7" pageNumber="458" rank="species">n. sp.</taxonomicNameLabel>
|
||
, of which he had examined some shells. Gofas’ view about the close similarity between
|
||
<taxonomicName id="9CBA1A99CE65FFD93556B4EAFA20FE0F" ID-CoL="3Y5H8" baseAuthorityName="Bozzetti" baseAuthorityYear="1995" box="[1281,1457,470,496]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="458" phylum="Mollusca" rank="species" species="colomborum">
|
||
<emphasis id="69CEBD08CE65FFD93556B4EAFA20FE0F" box="[1281,1457,470,496]" italics="true" pageId="7" pageNumber="458">M. colomborum</emphasis>
|
||
</taxonomicName>
|
||
and the fossil species is confirmed, but they are evidently distinct species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |