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<document id="F23E471E080D16681CFD7272D0DD1427" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D50766630F852C6ECC8FDA273050433" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onthophagus transisthmius Howden &amp; Young 1981" docType="treatment" docVersion="8" lastPageNumber="163" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="163" updateTime="1731448046962" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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(original description). Type locality:
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:
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: Gamboa. Name-bearing type:
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(
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)
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(list). —
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Medina
<emphasis id="A78D1B6230F852C6EF65FD5475A30233" box="[626,687,754,780]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
2001: 140
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(checklist for
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). —
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(checklist for
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). —
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<bibRefCitation id="F168BA8130F852C6ECEEFCB274930210" author="Pulido-Herrera L. A. &amp; Zunino M." box="[505,927,789,815]" pageId="162" pageNumber="163" refId="ref136342" refString="Pulido-Herrera L. A. &amp; Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. &amp; Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">Pulido-Herrera &amp; Zunino 2007: 117</bibRefCitation>
</treatmentCitation>
(catalogue). —
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(key, cited for
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). —
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Chamorro
<emphasis id="A78D1B6230F852C6EFA5FC9E75E0026D" box="[690,748,824,850]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
2018: 97
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(cited for
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). —
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Rossini
<emphasis id="A78D1B6230F852C6E9F4FC9E7212026D" box="[1251,1310,824,850]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
2018a: 547
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, 552553, 563564, figs 2ad, hi, 5 (key, diagnosis, redescription, remark);
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<bibRefCitation id="F168BA8130F852C6E990FCFC73F3024A" author="Rossini M. &amp; Vaz-de-Mello F. Z. &amp; Zunino M." box="[1159,1279,859,885]" firstAuthor="b" pageId="162" pageNumber="163" pagination="1 - 21" refId="ref137851" refString="Rossini M., Vaz-de-Mello F. Z. &amp; Zunino M. 2018 b. Toward a comprehensive taxonomic revision of the &quot; hirculus &quot; group of American Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae). European Journal of Taxonomy 432: 1 - 21. https: // doi. org / 10.5852 / ejt. 2018.432" type="journal article" year="2018">2018b: 10</bibRefCitation>
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(
<taxonomicName id="52F9BCF330F852C6E806FCFC727A024A" ID-CoL="7XNRQ" authorityName="Guerin-Meneville" authorityYear="1855" box="[1297,1398,859,885]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="osculatii">
<emphasis id="A78D1B6230F852C6E806FCFC727A024A" box="[1297,1398,859,885]" italics="true" pageId="162" pageNumber="163">osculatii</emphasis>
</taxonomicName>
complex). —
<treatmentCitation id="1458E16130F852C6EC48FCD9759902A6" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[351,661,894,921]" httpUri="http://treatment.plazi.org/id/412EF25DA0D2A64A84E2AEEE39C88C60" page="198" pageId="162" pageNumber="163" year="2019">
<bibRefCitation id="F168BA8130F852C6EC48FCD9759902A6" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[351,661,894,921]" pageId="162" pageNumber="163" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
Chamorro
<emphasis id="A78D1B6230F852C6ECCAFCD8751402A7" box="[477,536,894,920]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
2019: 198
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(catalogue).
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</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30F852C6EDAAFC6775FB0547" pageId="162" pageNumber="163" type="distribution">
<paragraph id="9546C77030F852C6EDAAFC67765A02E5" blockId="162.[189,342,960,986]" box="[189,342,960,986]" pageId="162" pageNumber="163">
<heading id="CE0E701C30F852C6EDAAFC67765A02E5" bold="true" box="[189,342,960,986]" fontSize="11" level="3" pageId="162" pageNumber="163" reason="3">
<emphasis id="A78D1B6230F852C6EDAAFC67765A02E5" bold="true" box="[189,342,960,986]" pageId="162" pageNumber="163">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030F852C6EDAAFC4874790537" blockId="162.[189,885,1006,1033]" box="[189,885,1006,1033]" pageId="162" pageNumber="163">
<collectingCountry id="EDEE87E030F852C6EDAAFC4876110536" box="[189,285,1007,1033]" name="Panama" pageId="162" pageNumber="163">Panama</collectingCountry>
,
<collectingCountry id="EDEE87E030F852C6EC3EFC4976AC0537" box="[297,416,1006,1032]" name="Colombia" pageId="162" pageNumber="163">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E030F852C6ECBDFC49752A0537" box="[426,550,1006,1032]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E030F852C6EF25FC49759A0537" box="[562,662,1006,1032]" name="Ecuador" pageId="162" pageNumber="163">Ecuador</collectingCountry>
,
<collectingCountry id="EDEE87E030F852C6EFB6FC4875D60536" box="[673,730,1007,1033]" name="Peru" pageId="162" pageNumber="163">Peru</collectingCountry>
, and
<collectingCountry id="EDEE87E030F852C6EE00FC49747D0537" box="[791,881,1006,1032]" name="Bolivia" pageId="162" pageNumber="163">Bolivia</collectingCountry>
.
</paragraph>
<paragraph id="9546C77030F852C6EDAAFB9776E10575" blockId="162.[189,493,1072,1098]" box="[189,493,1072,1098]" pageId="162" pageNumber="163">
<heading id="CE0E701C30F852C6EDAAFB9776E10575" bold="true" box="[189,493,1072,1098]" fontSize="11" level="3" pageId="162" pageNumber="163" reason="3">
<emphasis id="A78D1B6230F852C6EDAAFB9776E10575" bold="true" box="[189,493,1072,1098]" pageId="162" pageNumber="163">
Subregions of
<collectingCountry id="EDEE87E030F852C6EC78FB9776E10575" box="[367,493,1072,1098]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030F852C6EDAAFBF975FB0547" blockId="162.[189,759,1118,1144]" box="[189,759,1118,1144]" pageId="162" pageNumber="163">Plains, Serrania de Perija, and Andes mountains.</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30F852C6EDAAFB0773050433" pageId="162" pageNumber="163" type="reference_group">
<paragraph id="9546C77030F852C6EDAAFB0776A80585" blockId="162.[189,420,1184,1210]" box="[189,420,1184,1210]" pageId="162" pageNumber="163">
<heading id="CE0E701C30F852C6EDAAFB0776A80585" bold="true" box="[189,420,1184,1210]" fontSize="11" level="3" pageId="162" pageNumber="163" reason="3">
<emphasis id="A78D1B6230F852C6EDAAFB0776A80585" bold="true" box="[189,420,1184,1210]" pageId="162" pageNumber="163">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030F852C6EDAAFB6973050433" blockId="162.[189,1399,1230,1292]" pageId="162" pageNumber="163">
<bibRefCitation id="F168BA8130F852C6EDAAFB6976F905D6" author="Rossini M. &amp; Vaz-de-Mello F. Z. &amp; Zunino M." box="[189,501,1230,1257]" pageId="162" pageNumber="163" pagination="541 - 586" refId="ref137777" refString="Rossini M., Vaz-de-Mello F. Z. &amp; Zunino M. 2018 a. A taxonomic revision of the New World Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae) of the osculatii species-complex, with description of two new species from South America. Journal of Natural History 52 (9 - 10): 541 - 586. https: // doi. org / 10.1080 / 00222933.2018.1437230" type="journal article" year="2018">
Rossini
<emphasis id="A78D1B6230F852C6EC37FB68765305D7" box="[288,351,1230,1256]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
2018a: 552
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, 563 (
<collectingCountry id="EDEE87E030F852C6EF53FB6975CF05D7" box="[580,707,1230,1256]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230F852C6EFC6FB69742205D7" box="[721,814,1230,1256]" country="Venezuela" name="Tachira" pageId="162" pageNumber="163">Táchira</collectingRegion>
,
<collectingRegion id="573D099230F852C6EE2AFB6874B305D6" box="[829,959,1231,1257]" country="Venezuela" name="Portuguesa" pageId="162" pageNumber="163">Portuguesa</collectingRegion>
and
<collectingRegion id="573D099230F852C6E916FB69734805D7" box="[1025,1092,1230,1256]" country="Venezuela" name="Zulia" pageId="162" pageNumber="163">Zulia</collectingRegion>
); 2018b: 11 (two species reaching eastern Amazonia). —
<bibRefCitation id="F168BA8130F852C6EF2FFB5574620433" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[568,878,1266,1292]" pageId="162" pageNumber="163" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
Chamorro
<emphasis id="A78D1B6230F852C6EFA1FB5575FD0433" box="[694,753,1266,1292]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
2019: 198
</bibRefCitation>
(
<collectingCountry id="EDEE87E030F852C6EE6AFB5574F10433" box="[893,1021,1266,1292]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
).
</paragraph>
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