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245 lines
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<document id="F23E471E080D16681CFD7272D0DD1427" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario & Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D50766630F852C6ECC8FDA273050433" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onthophagus transisthmius Howden & Young 1981" docType="treatment" docVersion="8" lastPageNumber="163" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="163" updateTime="1731448046962" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B696A4803D9D0CD928B625E641978BF8">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="3C150F2993491C7405B723F68147EE0B">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="E902747FC54F6BCB2355BE417394CF71">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:nameIdentifier id="7FB77D4C36522EF9264F9E0D5903DF09" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A2DB930945809E9C35E1A57966505A17">Cupello, Mario</mods:namePart>
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<mods:affiliation id="77969460B1B8A60BD5854169BDF7BE9F">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A & M University, College Station, TX, 77843, USA.</mods:affiliation>
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<mods:nameIdentifier id="D714AE60CA05AC04244A38D333130A0B" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="02157FB4EA4C893AEB3ED0E250DF3147">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="9F75899E32A64FADF4C09F1078C30D89">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="FE94939985B6EDC122E36B162AA39074" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:title id="12EAAAF792F2D2DFB3CB197915DCA5F0">European Journal of Taxonomy</mods:title>
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<mods:date id="4D8D85493EA13A100258AF7D32A6D82A">2024</mods:date>
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<mods:number id="B98639CB24641A11B32D86CBB4F394B8">2024-10-11</mods:number>
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<treatment id="1D50766630F852C6ECC8FDA273050433" ID-DOI="http://doi.org/10.5281/zenodo.13945853" ID-Zenodo-Dep="13945853" LSID="urn:lsid:plazi:treatment:1D50766630F852C6ECC8FDA273050433" httpUri="http://treatment.plazi.org/id/1D50766630F852C6ECC8FDA273050433" lastPageNumber="163" pageId="162" pageNumber="163">
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<subSubSection id="DDE394FB30F852C6ECC8FDA273590320" box="[479,1109,517,544]" pageId="162" pageNumber="163" type="nomenclature">
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<paragraph id="9546C77030F852C6ECC8FDA273590320" blockId="162.[479,1109,517,578]" box="[479,1109,517,544]" pageId="162" pageNumber="163">
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<taxonomicName id="52F9BCF330F852C6ECC8FDA273590320" ID-CoL="7XPCX" authority="Howden & Young, 1981" authorityName="Howden & Young" authorityYear="1981" box="[479,1109,517,544]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="transisthmius">
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<emphasis id="A78D1B6230F852C6ECC8FDA2743C031F" bold="true" box="[479,816,517,544]" italics="true" pageId="162" pageNumber="163">Onthophagus transisthmius</emphasis>
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<bibRefCitation id="F168BA8130F852C6EE20FDA273590320" author="Howden H. F. & Young O. P." box="[823,1109,517,544]" pageId="162" pageNumber="163" pagination="1 - 204" refId="ref124824" refString="Howden H. F. & Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden & Young, 1981</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DDE394FB30F852C6EFF1FD8F7441037D" box="[742,845,552,578]" pageId="162" pageNumber="163" type="description">
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<paragraph id="9546C77030F852C6EFF1FD8F7441037D" blockId="162.[479,1109,517,578]" box="[742,845,552,578]" pageId="162" pageNumber="163">
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<figureCitation id="0DC2DBF530F852C6EFF1FD8F7441037D" box="[742,845,552,578]" captionStart="Fig" captionStartId="163.[188,231,1963,1989]" captionTargetBox="[274,1307,278,1896]" captionTargetId="figure-11@163.[262,1323,265,1914]" captionTargetPageId="163" captionText="Fig. 62. Species distribution. A. Onthophagus rhinophyllus Harold, 1868.B. Onthophagus transisthmius Howden & Young, 1981. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945635" httpUri="https://zenodo.org/record/13945635/files/figure.png" pageId="162" pageNumber="163">Fig. 62B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DDE394FB30F852C6EDAAFDCD742502A7" pageId="162" pageNumber="163" type="reference_group">
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<paragraph id="9546C77030F852C6EDAAFDCD73560398" blockId="162.[188,1399,618,680]" pageId="162" pageNumber="163">
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<treatmentCitationGroup id="B5E9E05E30F852C6EDAAFDCD73560398" pageId="162" pageNumber="163">
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<treatmentCitation id="1458E16130F852C6EDAAFDCD745203BB" author="Howden H. F. & Young O. P." box="[189,862,618,644]" page="106" pageId="162" pageNumber="163" year="1981">
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<taxonomicName id="52F9BCF330F852C6EDAAFDCD745203BB" ID-CoL="7XPCX" authority="Howden & Young, 1981: 106" authorityName="Howden & Young" authorityPageNumber="106" authorityYear="1981" box="[189,862,618,644]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="transisthmius">
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<emphasis id="A78D1B6230F852C6EDAAFDCD750C03BB" box="[189,512,618,644]" italics="true" pageId="162" pageNumber="163">Onthophagus transisthmius</emphasis>
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<bibRefCitation id="F168BA8130F852C6EF10FDCD745203BB" author="Howden H. F. & Young O. P." box="[519,862,618,644]" pageId="162" pageNumber="163" pagination="1 - 204" refId="ref124824" refString="Howden H. F. & Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden & Young, 1981: 106</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(original description). Type locality:
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<collectingCountry id="EDEE87E030F852C6E806FDCD727F03BB" box="[1297,1395,618,644]" name="Panama" pageId="162" pageNumber="163">Panama</collectingCountry>
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:
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<collectingRegion id="573D099230F852C6EDABFD2A76040398" box="[188,264,653,679]" country="Panama" name="Colon" pageId="162" pageNumber="163">Colón</collectingRegion>
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:
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<collectingRegion id="573D099230F852C6EC03FD2A766C0398" box="[276,352,653,679]" country="Panama" name="Colon" pageId="162" pageNumber="163">Colón</collectingRegion>
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: Gamboa. Name-bearing type:
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<typeStatus id="4A4279D230F852C6EFD9FD2A74390398" box="[718,821,653,679]" pageId="162" pageNumber="163" type="holotype">holotype</typeStatus>
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(USNM), not examined.
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</paragraph>
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<paragraph id="9546C77030F852C6EDAAFD69742502A7" blockId="162.[189,1399,718,921]" pageId="162" pageNumber="163">
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<treatmentCitationGroup id="B5E9E05E30F852C6EDAAFD69742502A7" pageId="162" pageNumber="163">
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<taxonomicName id="52F9BCF330F852C6EDAAFD6975B303D7" ID-CoL="7XPCX" authorityName="Howden & Young" authorityYear="1981" box="[189,703,718,745]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="transisthmius" subGenus="Onthophagus">
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<emphasis id="A78D1B6230F852C6EDAAFD69765703D7" box="[189,347,718,744]" italics="true" pageId="162" pageNumber="163">Onthophagus</emphasis>
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(
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<emphasis id="A78D1B6230F852C6EC78FD69750103D7" box="[367,525,718,744]" italics="true" pageId="162" pageNumber="163">Onthophagus</emphasis>
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)
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<emphasis id="A78D1B6230F852C6EF36FD6975B303D7" box="[545,703,718,744]" italics="true" pageId="162" pageNumber="163">transisthmius</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="1458E16130F852C6EFE6FD69735C03D7" author="Zunino M. & Halffter G." box="[753,1104,718,745]" page="161" pageId="162" pageNumber="163" year="1997">
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<bibRefCitation id="F168BA8130F852C6EFE6FD69735C03D7" author="Zunino M. & Halffter G." box="[753,1104,718,745]" pageId="162" pageNumber="163" pagination="157 - 178" refId="ref142302" refString="Zunino M. & Halffter G. 1997. Sobre Onthophagus Latreille, 1802 americanos (Coleoptera: Scarabaeidae: Scarabaeinae). Elytron 11: 157 - 178." type="journal article" year="1997">Zunino & Halffter 1997: 161</bibRefCitation>
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</treatmentCitation>
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(list). —
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<treatmentCitation id="1458E16130F852C6E9DCFD6977E50233" author="Escobar F." page="209" pageId="162" pageNumber="163" year="2000">
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<bibRefCitation id="F168BA8130F852C6E9DCFD6977E50233" author="Escobar F." pageId="162" pageNumber="163" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. & Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 209</bibRefCitation>
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</treatmentCitation>
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(list for
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<collectingCountry id="EDEE87E030F852C6EC40FD5576DC0233" box="[343,464,754,780]" name="Colombia" pageId="162" pageNumber="163">Colombia</collectingCountry>
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). —
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<treatmentCitation id="1458E16130F852C6EF19FD55743E0233" author="Medina C. A. & Lopera-Toro A. & Vitolo A. & Gill B." box="[526,818,754,780]" page="140" pageId="162" pageNumber="163" year="2001">
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<bibRefCitation id="F168BA8130F852C6EF19FD55743E0233" author="Medina C. A. & Lopera-Toro A. & Vitolo A. & Gill B." box="[526,818,754,780]" pageId="162" pageNumber="163" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. & Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
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Medina
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<emphasis id="A78D1B6230F852C6EF65FD5475A30233" box="[626,687,754,780]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
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2001: 140
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</bibRefCitation>
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</treatmentCitation>
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(checklist for
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<collectingCountry id="EDEE87E030F852C6EEF2FD5573520233" box="[997,1118,754,780]" name="Colombia" pageId="162" pageNumber="163">Colombia</collectingCountry>
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). —
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<treatmentCitation id="1458E16130F852C6E98BFD55727A0233" author="Ratcliffe B. C." box="[1180,1398,754,780]" page="17" pageId="162" pageNumber="163" year="2002">
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<bibRefCitation id="F168BA8130F852C6E98BFD55727A0233" author="Ratcliffe B. C." box="[1180,1398,754,780]" firstAuthor="Ratcliffe" pageId="162" pageNumber="163" pagination="1 - 48" refId="ref136799" refString="Ratcliffe B. C. 2002. A checklist of the Scarabaeoidea of Panama. Zootaxa 32: 1 - 48. https: // doi. org / 10.11646 / zootaxa. 32.1.1" type="journal article" year="2002">Ratcliffe 2002: 17</bibRefCitation>
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</treatmentCitation>
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(checklist for
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<collectingCountry id="EDEE87E030F852C6EC48FCB276CD0210" box="[351,449,789,815]" name="Panama" pageId="162" pageNumber="163">Panama</collectingCountry>
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). —
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<treatmentCitation id="1458E16130F852C6ECEEFCB274930210" author="Pulido-Herrera L. A. & Zunino M." box="[505,927,789,815]" page="117" pageId="162" pageNumber="163" year="2007">
|
|||
|
|
<bibRefCitation id="F168BA8130F852C6ECEEFCB274930210" author="Pulido-Herrera L. A. & Zunino M." box="[505,927,789,815]" pageId="162" pageNumber="163" refId="ref136342" refString="Pulido-Herrera L. A. & Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. & Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">Pulido-Herrera & Zunino 2007: 117</bibRefCitation>
|
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|
|
</treatmentCitation>
|
|||
|
|
(catalogue). —
|
|||
|
|
<treatmentCitation id="1458E16130F852C6E94BFCB277D6026C" author="Delgado L. & Curoe D. J." page="65" pageId="162" pageNumber="163" year="2014">
|
|||
|
|
<bibRefCitation id="F168BA8130F852C6E94BFCB277D6026C" author="Delgado L. & Curoe D. J." pageId="162" pageNumber="163" pagination="61 - 67" refId="ref116462" refString="Delgado L. & Curoe D. J. 2014. Panamanian Onthophagus (Coleoptera: Scarabaeidae): description of a new species, and a revised key to the species. Florida Entomologist 97 (1): 61 - 67. https: // doi. org / 10.1653 / 024.097.0107" type="journal article" year="2014">Delgado & Curoe 2014: 65</bibRefCitation>
|
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|
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</treatmentCitation>
|
|||
|
|
(key, cited for
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EC84FC9F76F9026D" box="[403,501,824,850]" name="Panama" pageId="162" pageNumber="163">Panama</collectingCountry>
|
|||
|
|
). —
|
|||
|
|
<treatmentCitation id="1458E16130F852C6EF27FC9F7452026D" author="Chamorro W. & Marin-Armijos D. & Granda V. & Vaz-de-Mello F. Z." box="[560,862,824,850]" page="97" pageId="162" pageNumber="163" year="2018">
|
|||
|
|
<bibRefCitation id="F168BA8130F852C6EF27FC9F7452026D" author="Chamorro W. & Marin-Armijos D. & Granda V. & Vaz-de-Mello F. Z." box="[560,862,824,850]" pageId="162" pageNumber="163" pagination="72 - 100" refId="ref114035" refString="Chamorro W., Marin-Armijos D., Granda V. & Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
|
|||
|
|
Chamorro
|
|||
|
|
<emphasis id="A78D1B6230F852C6EFA5FC9E75E0026D" box="[690,748,824,850]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
|
|||
|
|
2018: 97
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(cited for
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EEC9FC9F7349026D" box="[990,1093,824,850]" name="Ecuador" pageId="162" pageNumber="163">Ecuador</collectingCountry>
|
|||
|
|
). —
|
|||
|
|
<treatmentCitation id="1458E16130F852C6E997FC9F77E20249" author="Rossini M. & Vaz-de-Mello F. Z. & Zunino M." httpUri="http://treatment.plazi.org/id/266887BDFF86A132010AFCF1FDECB29F" page="547" pageId="162" pageNumber="163" year="2018">
|
|||
|
|
<bibRefCitation id="F168BA8130F852C6E997FC9F77E20249" author="Rossini M. & Vaz-de-Mello F. Z. & Zunino M." pageId="162" pageNumber="163" pagination="541 - 586" refId="ref137777" refString="Rossini M., Vaz-de-Mello F. Z. & Zunino M. 2018 a. A taxonomic revision of the New World Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae) of the osculatii species-complex, with description of two new species from South America. Journal of Natural History 52 (9 - 10): 541 - 586. https: // doi. org / 10.1080 / 00222933.2018.1437230" type="journal article" year="2018">
|
|||
|
|
Rossini
|
|||
|
|
<emphasis id="A78D1B6230F852C6E9F4FC9E7212026D" box="[1251,1310,824,850]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
|
|||
|
|
2018a: 547
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
, 552–553, 563‒564, figs 2a‒d, h‒i, 5 (key, diagnosis, redescription, remark);
|
|||
|
|
<treatmentCitation id="1458E16130F852C6E990FCFC73F3024A" author="Rossini M. & Vaz-de-Mello F. Z. & Zunino M." box="[1159,1279,859,885]" httpUri="http://treatment.plazi.org/id/266887BDFF86A132010AFCF1FDECB29F" page="10" pageId="162" pageNumber="163" year="2018">
|
|||
|
|
<bibRefCitation id="F168BA8130F852C6E990FCFC73F3024A" author="Rossini M. & Vaz-de-Mello F. Z. & Zunino M." box="[1159,1279,859,885]" firstAuthor="b" pageId="162" pageNumber="163" pagination="1 - 21" refId="ref137851" refString="Rossini M., Vaz-de-Mello F. Z. & Zunino M. 2018 b. Toward a comprehensive taxonomic revision of the " hirculus " group of American Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae). European Journal of Taxonomy 432: 1 - 21. https: // doi. org / 10.5852 / ejt. 2018.432" type="journal article" year="2018">2018b: 10</bibRefCitation>
|
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|
|
</treatmentCitation>
|
|||
|
|
(
|
|||
|
|
<taxonomicName id="52F9BCF330F852C6E806FCFC727A024A" ID-CoL="7XNRQ" authorityName="Guerin-Meneville" authorityYear="1855" box="[1297,1398,859,885]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="osculatii">
|
|||
|
|
<emphasis id="A78D1B6230F852C6E806FCFC727A024A" box="[1297,1398,859,885]" italics="true" pageId="162" pageNumber="163">osculatii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
complex). —
|
|||
|
|
<treatmentCitation id="1458E16130F852C6EC48FCD9759902A6" author="Chamorro W. & Marin-Armijos D. & Asenjo A. & Vaz-De-Mello F. Z." box="[351,661,894,921]" httpUri="http://treatment.plazi.org/id/412EF25DA0D2A64A84E2AEEE39C88C60" page="198" pageId="162" pageNumber="163" year="2019">
|
|||
|
|
<bibRefCitation id="F168BA8130F852C6EC48FCD9759902A6" author="Chamorro W. & Marin-Armijos D. & Asenjo A. & Vaz-De-Mello F. Z." box="[351,661,894,921]" pageId="162" pageNumber="163" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. & Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
|
|||
|
|
Chamorro
|
|||
|
|
<emphasis id="A78D1B6230F852C6ECCAFCD8751402A7" box="[477,536,894,920]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
|
|||
|
|
2019: 198
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(catalogue).
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DDE394FB30F852C6EDAAFC6775FB0547" pageId="162" pageNumber="163" type="distribution">
|
|||
|
|
<paragraph id="9546C77030F852C6EDAAFC67765A02E5" blockId="162.[189,342,960,986]" box="[189,342,960,986]" pageId="162" pageNumber="163">
|
|||
|
|
<heading id="CE0E701C30F852C6EDAAFC67765A02E5" bold="true" box="[189,342,960,986]" fontSize="11" level="3" pageId="162" pageNumber="163" reason="3">
|
|||
|
|
<emphasis id="A78D1B6230F852C6EDAAFC67765A02E5" bold="true" box="[189,342,960,986]" pageId="162" pageNumber="163">Distribution</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9546C77030F852C6EDAAFC4874790537" blockId="162.[189,885,1006,1033]" box="[189,885,1006,1033]" pageId="162" pageNumber="163">
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EDAAFC4876110536" box="[189,285,1007,1033]" name="Panama" pageId="162" pageNumber="163">Panama</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EC3EFC4976AC0537" box="[297,416,1006,1032]" name="Colombia" pageId="162" pageNumber="163">Colombia</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6ECBDFC49752A0537" box="[426,550,1006,1032]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EF25FC49759A0537" box="[562,662,1006,1032]" name="Ecuador" pageId="162" pageNumber="163">Ecuador</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EFB6FC4875D60536" box="[673,730,1007,1033]" name="Peru" pageId="162" pageNumber="163">Peru</collectingCountry>
|
|||
|
|
, and
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EE00FC49747D0537" box="[791,881,1006,1032]" name="Bolivia" pageId="162" pageNumber="163">Bolivia</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9546C77030F852C6EDAAFB9776E10575" blockId="162.[189,493,1072,1098]" box="[189,493,1072,1098]" pageId="162" pageNumber="163">
|
|||
|
|
<heading id="CE0E701C30F852C6EDAAFB9776E10575" bold="true" box="[189,493,1072,1098]" fontSize="11" level="3" pageId="162" pageNumber="163" reason="3">
|
|||
|
|
<emphasis id="A78D1B6230F852C6EDAAFB9776E10575" bold="true" box="[189,493,1072,1098]" pageId="162" pageNumber="163">
|
|||
|
|
Subregions of
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EC78FB9776E10575" box="[367,493,1072,1098]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9546C77030F852C6EDAAFBF975FB0547" blockId="162.[189,759,1118,1144]" box="[189,759,1118,1144]" pageId="162" pageNumber="163">Plains, Serrania de Perija, and Andes mountains.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DDE394FB30F852C6EDAAFB0773050433" pageId="162" pageNumber="163" type="reference_group">
|
|||
|
|
<paragraph id="9546C77030F852C6EDAAFB0776A80585" blockId="162.[189,420,1184,1210]" box="[189,420,1184,1210]" pageId="162" pageNumber="163">
|
|||
|
|
<heading id="CE0E701C30F852C6EDAAFB0776A80585" bold="true" box="[189,420,1184,1210]" fontSize="11" level="3" pageId="162" pageNumber="163" reason="3">
|
|||
|
|
<emphasis id="A78D1B6230F852C6EDAAFB0776A80585" bold="true" box="[189,420,1184,1210]" pageId="162" pageNumber="163">Literature records</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9546C77030F852C6EDAAFB6973050433" blockId="162.[189,1399,1230,1292]" pageId="162" pageNumber="163">
|
|||
|
|
<bibRefCitation id="F168BA8130F852C6EDAAFB6976F905D6" author="Rossini M. & Vaz-de-Mello F. Z. & Zunino M." box="[189,501,1230,1257]" pageId="162" pageNumber="163" pagination="541 - 586" refId="ref137777" refString="Rossini M., Vaz-de-Mello F. Z. & Zunino M. 2018 a. A taxonomic revision of the New World Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae) of the osculatii species-complex, with description of two new species from South America. Journal of Natural History 52 (9 - 10): 541 - 586. https: // doi. org / 10.1080 / 00222933.2018.1437230" type="journal article" year="2018">
|
|||
|
|
Rossini
|
|||
|
|
<emphasis id="A78D1B6230F852C6EC37FB68765305D7" box="[288,351,1230,1256]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
|
|||
|
|
2018a: 552
|
|||
|
|
</bibRefCitation>
|
|||
|
|
, 563 (
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EF53FB6975CF05D7" box="[580,707,1230,1256]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="573D099230F852C6EFC6FB69742205D7" box="[721,814,1230,1256]" country="Venezuela" name="Tachira" pageId="162" pageNumber="163">Táchira</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="573D099230F852C6EE2AFB6874B305D6" box="[829,959,1231,1257]" country="Venezuela" name="Portuguesa" pageId="162" pageNumber="163">Portuguesa</collectingRegion>
|
|||
|
|
and
|
|||
|
|
<collectingRegion id="573D099230F852C6E916FB69734805D7" box="[1025,1092,1230,1256]" country="Venezuela" name="Zulia" pageId="162" pageNumber="163">Zulia</collectingRegion>
|
|||
|
|
); 2018b: 11 (two species reaching eastern Amazonia). —
|
|||
|
|
<bibRefCitation id="F168BA8130F852C6EF2FFB5574620433" author="Chamorro W. & Marin-Armijos D. & Asenjo A. & Vaz-De-Mello F. Z." box="[568,878,1266,1292]" pageId="162" pageNumber="163" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. & Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
|
|||
|
|
Chamorro
|
|||
|
|
<emphasis id="A78D1B6230F852C6EFA1FB5575FD0433" box="[694,753,1266,1292]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
|
|||
|
|
2019: 198
|
|||
|
|
</bibRefCitation>
|
|||
|
|
(
|
|||
|
|
<collectingCountry id="EDEE87E030F852C6EE6AFB5574F10433" box="[893,1021,1266,1292]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|