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<mods:title id="66E69625EC9B6E2672D377B7F02EB990">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="BF87246B69DB1D3F58EDB9FF2F84E3A9">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="BEE9D9EEA13ABBE8EE6E1395E7E03639">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:namePart id="91B361777DE2F8416939AEF065A35766">Cupello, Mario</mods:namePart>
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<mods:affiliation id="B134CD71E6D2092476E3D07EDF4AAF7C">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="ABD9FA28ED2DAB3FE5858512151089A3" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="A3B8A9ACA23B0F8A73317D9E7D6743A4">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="096CDA217176BCFB534296206DEE9767">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<treatment id="1D50766630CF52F3EF54FC70727B0716" ID-DOI="http://doi.org/10.5281/zenodo.14154703" ID-GBIF-Taxon="246297851" ID-Zenodo-Dep="14154703" LSID="urn:lsid:plazi:treatment:1D50766630CF52F3EF54FC70727B0716" httpUri="http://treatment.plazi.org/id/1D50766630CF52F3EF54FC70727B0716" lastPageId="151" lastPageNumber="152" pageId="149" pageNumber="150">
<subSubSection id="DDE394FB30CF52F1EF54FC7074FC02CD" box="[579,1008,983,1010]" pageId="149" pageNumber="150" type="nomenclature">
<paragraph id="9546C77030CF52F1EF54FC7074FC02CD" blockId="149.[579,1008,983,1045]" box="[579,1008,983,1010]" pageId="149" pageNumber="150">
<heading id="CE0E701C30CF52F1EF54FC7074FC02CD" box="[579,1008,983,1010]" centered="true" fontSize="11" level="2" pageId="149" pageNumber="150" reason="2">
<taxonomicName id="52F9BCF330CF52F1EF54FC7074FC02CD" ID-CoL="7ZQ4B" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[579,1008,983,1010]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CF52F1EF54FC70741302CD" bold="true" box="[579,799,983,1010]" italics="true" pageId="149" pageNumber="150">Ontherus sulcator</emphasis>
(
<bibRefCitation id="F168BA8130CF52F1EE39FC7074E402CD" author="Fabricius J. C." box="[814,1000,983,1010]" pageId="149" pageNumber="150" refId="ref118353" refString="Fabricius J. C. 1775. Systema entomologiae, sistens insectorvm classes, ordines, genera, species, adiectis synonymis, locis, descriptionibvs, observationibvs. Kortius, Flensburg and Leipzig [Flensbvrgi et Lipsiae]. https: // doi. org / 10.5962 / bhl. title. 36510" type="book" year="1775">Fabricius, 1775</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30CF52F1EFF1FC5D7441052A" box="[742,845,1018,1045]" pageId="149" pageNumber="150" type="description">
<paragraph id="9546C77030CF52F1EFF1FC5D7441052A" blockId="149.[579,1008,983,1045]" box="[742,845,1018,1045]" pageId="149" pageNumber="150">
<figureCitation id="0DC2DBF530CF52F1EFF1FC5D7441052A" box="[742,845,1018,1045]" captionStart="Fig" captionStartId="150.[189,232,1963,1989]" captionTargetBox="[274,1302,278,1887]" captionTargetId="figure-18@150.[262,1325,265,1913]" captionTargetPageId="150" captionText="Fig. 58. Species distribution. A. Ontherus pubens Génier, 1996. B. Ontherus sulcator (Fabricius, 1775). Black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945625" httpUri="https://zenodo.org/record/13945625/files/figure.png" pageId="149" pageNumber="150">Fig. 58B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30CF52F3EDAAFB9B753E0573" lastPageId="151" lastPageNumber="152" pageId="149" pageNumber="150" type="reference_group">
<paragraph id="9546C77030CF52F1EDAAFB9B757405FF" blockId="149.[189,1399,1084,1462]" pageId="149" pageNumber="150">
<treatmentCitationGroup id="B5E9E05E30CF52F1EDAAFB9B757405FF" pageId="149" pageNumber="150">
<treatmentCitation id="1458E16130CF52F1EDAAFB9B75AE0569" author="Fabricius J. C." box="[189,674,1084,1111]" page="27" pageId="149" pageNumber="150" year="1775">
<taxonomicName id="52F9BCF330CF52F1EDAAFB9B75AE0569" ID-CoL="7YY98" authority="Fabricius, 1775: 27" authorityName="Fabricius" authorityPageNumber="27" authorityYear="1775" box="[189,674,1084,1111]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CF52F1EDAAFB9B76BC0569" box="[189,432,1084,1110]" italics="true" pageId="149" pageNumber="150">Scarabaeus sulcator</emphasis>
<bibRefCitation id="F168BA8130CF52F1ECADFB9B75AE0569" author="Fabricius J. C." box="[442,674,1084,1111]" pageId="149" pageNumber="150" refId="ref118353" refString="Fabricius J. C. 1775. Systema entomologiae, sistens insectorvm classes, ordines, genera, species, adiectis synonymis, locis, descriptionibvs, observationibvs. Kortius, Flensburg and Leipzig [Flensbvrgi et Lipsiae]. https: // doi. org / 10.5962 / bhl. title. 36510" type="book" year="1775">Fabricius, 1775: 27</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E030CF52F1E972FB9B72160569" box="[1125,1306,1084,1110]" name="French Guiana" pageId="149" pageNumber="150">French Guiana</collectingCountry>
. Name-bearing type: according to
<treatmentCitation id="1458E16130CF52F1EF37FBF875DC0546" author="Zimsen E." box="[544,720,1119,1145]" pageId="149" pageNumber="150" year="1964">
<bibRefCitation id="F168BA8130CF52F1EF37FBF875DC0546" author="Zimsen E." box="[544,720,1119,1145]" pageId="149" pageNumber="150" refId="ref142090" refString="Zimsen E. 1964. The Type Series of I. C. Fabricius. Munksgaard, Copenhagen." type="book" year="1964">Zimsen (1964)</bibRefCitation>
</treatmentCitation>
,
<specimenCount id="83FF0CF930CF52F1EFCDFBF8748E0545" box="[730,898,1119,1146]" count="3" pageId="149" pageNumber="150" type="generic" typeStatus="syntype">three syntypes</specimenCount>
in ZMUK, but deemed lost without further explanation by
<treatmentCitation id="1458E16130CF52F1EC88FB25754505A3" author="Genier F." box="[415,585,1154,1180]" pageId="149" pageNumber="150" year="1996">
<bibRefCitation id="F168BA8130CF52F1EC88FB25754505A3" author="Genier F." box="[415,585,1154,1180]" pageId="149" pageNumber="150" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier (1996)</bibRefCitation>
</treatmentCitation>
. We ourselves have made no attempt to locate them, and so they may well be in ZMUK as Zimsen said.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CF52F1EDAAFB6E732B0439" blockId="149.[189,1399,1084,1462]" pageId="149" pageNumber="150">
<treatmentCitationGroup id="B5E9E05E30CF52F1EDAAFB6E732B0439" pageId="149" pageNumber="150">
<treatmentCitation id="1458E16130CF52F1EDAAFB6E754705DC" author="Castelnau" box="[189,587,1224,1251]" page="79" pageId="149" pageNumber="150" year="1840">
<taxonomicName id="52F9BCF330CF52F1EDAAFB6E754705DC" ID-CoL="7VDLM" authority="Castelnau, 1840: 79" authorityName="Castelnau" authorityPageNumber="79" authorityYear="1840" box="[189,587,1224,1251]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="nisus">
<emphasis id="A78D1B6230CF52F1EDAAFB6E765F05DD" box="[189,339,1225,1251]" italics="true" pageId="149" pageNumber="150">Copris nisus</emphasis>
<bibRefCitation id="F168BA8130CF52F1EC4AFB6F754705DC" author="Castelnau" box="[349,587,1224,1251]" pageId="149" pageNumber="150" refId="ref113936" refString="Castelnau, Comte de 1840. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes. Histoire naturelle des insectes coleopteres. Tome deuxieme. Dumenil, Paris. https: // doi. org / 10.5962 / bhl. title. 47104" type="book" year="1840">Castelnau, 1840: 79</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E030CF52F1E919FB6F73CE05DD" box="[1038,1218,1224,1250]" name="French Guiana" pageId="149" pageNumber="150">French Guiana</collectingCountry>
. Name-bearing type:
<typeStatus id="4A4279D230CF52F1EC38FB4B76970439" box="[303,411,1260,1286]" pageId="149" pageNumber="150" type="lectotype">lectotype</typeStatus>
(MVMA), designated by
<treatmentCitation id="1458E16130CF52F1EFDBFB4B74780439" author="Genier F." box="[716,884,1260,1286]" pageId="149" pageNumber="150" year="1996">
<bibRefCitation id="F168BA8130CF52F1EFDBFB4B74780439" author="Genier F." box="[716,884,1260,1286]" pageId="149" pageNumber="150" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier (1996)</bibRefCitation>
</treatmentCitation>
, not examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CF52F1EDAAFAA873C20473" blockId="149.[189,1399,1084,1462]" pageId="149" pageNumber="150">
<treatmentCitationGroup id="B5E9E05E30CF52F1EDAAFAA873C20473" pageId="149" pageNumber="150">
<treatmentCitation id="1458E16130CF52F1EDAAFAA875A70416" author="Blanchard E." box="[189,683,1295,1321]" page="180" pageId="149" pageNumber="150" year="1846">
<taxonomicName id="52F9BCF330CF52F1EDAAFAA875A70416" ID-CoL="7VDLN" authority="Blanchard, 1846: 180" authorityName="Blanchard" authorityPageNumber="180" authorityYear="1846" box="[189,683,1295,1321]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="ovalipennis">
<emphasis id="A78D1B6230CF52F1EDAAFAA876920416" box="[189,414,1295,1321]" italics="true" pageId="149" pageNumber="150">Copris ovalipennis</emphasis>
<bibRefCitation id="F168BA8130CF52F1ECBFFAA875A70416" author="Blanchard E." box="[424,683,1295,1321]" pageId="149" pageNumber="150" pagination="153 - 168" refId="ref111318" refString="Blanchard E. 1846. [livraison 79: 153 - 168, pl. 28; livraison 80: 169 - 184, pl. 31]. In: Blanchard E. &amp; Brulle G. A. 1835 - 1846. Voyage dans l'Amerique Meridionale (le Bresil, la Republique orientale de L'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833, par Alcide d'Orbigny. Tome Sixieme. 2. e partie: Insectes. P. Bertrand, Paris, and Ve. Lerrault, Strasbourg. [Both livraisons date from 1846. For details on the publication date of livraison 79, see Cupello et al. (2023 a: 89 - 90); the earliest evidence of publication of livraison 80 dates from 18 Dec. 1846 (Neal Evenhuis, pers. com. to MC, 3 Sep. 2019).]" type="journal article" year="1846">Blanchard, 1846: 180</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E030CF52F1E979FAA873E40416" box="[1134,1256,1295,1321]" name="Argentina" pageId="149" pageNumber="150">Argentina</collectingCountry>
:
<collectingRegion id="573D099230CF52F1E9E2FAA8727F0416" box="[1269,1395,1295,1321]" country="Argentina" name="Corrientes" pageId="149" pageNumber="150">Corrientes</collectingRegion>
. Name-bearing type:
<typeStatus id="4A4279D230CF52F1ECCCFA95754B0473" box="[475,583,1330,1356]" pageId="149" pageNumber="150" type="lectotype">lectotype</typeStatus>
(MNHN), designated by
<treatmentCitation id="1458E16130CF52F1EE63FA9573100473" author="Genier F." box="[884,1052,1330,1356]" pageId="149" pageNumber="150" year="1996">
<bibRefCitation id="F168BA8130CF52F1EE63FA9573100473" author="Genier F." box="[884,1052,1330,1356]" pageId="149" pageNumber="150" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier (1996)</bibRefCitation>
</treatmentCitation>
, not examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CF52F1EDAAFAF175350489" blockId="149.[189,1399,1084,1462]" pageId="149" pageNumber="150">
<treatmentCitationGroup id="B5E9E05E30CF52F1EDAAFAF175350489" pageId="149" pageNumber="150">
<treatmentCitation id="1458E16130CF52F1EDAAFAF17567044F" author="Gillet J. J. E." box="[189,619,1365,1392]" page="15" pageId="149" pageNumber="150" year="1910">
<taxonomicName id="52F9BCF330CF52F1EDAAFAF17567044F" ID-CoL="7VDMF" authority="Gillet, 1910: 15" authorityName="Gillet" authorityPageNumber="15" authorityYear="1910" box="[189,619,1365,1392]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="senegalensis">
<emphasis id="A78D1B6230CF52F1EDAAFAF176A50450" box="[189,425,1365,1392]" italics="true" pageId="149" pageNumber="150">Copris senegalensis</emphasis>
<bibRefCitation id="F168BA8130CF52F1ECA6FAF27567044F" author="Gillet J. J. E." box="[433,619,1365,1392]" pageId="149" pageNumber="150" pagination="1 - 31" refId="ref121131" refString="Gillet J. J. E. 1910. Especes nouvelles du genre Copris et releve synonymique des especes decrites a ce jour. Notes from the Leyden Museum 32: 1 - 31. Available from https: // repository. naturalis. nl / pub / 508947 [accessed 16 Feb. 2024]." type="journal article" year="1910">Gillet, 1910: 15</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality: Unknown. Originally said to be
<collectingCountry id="EDEE87E030CF52F1EC18FADF766004AD" box="[271,364,1400,1426]" name="Senegal" pageId="149" pageNumber="150">Senegal</collectingCountry>
(“
<collectingCountry id="EDEE87E030CF52F1EC90FADF76E104AD" box="[391,493,1400,1426]" name="Senegal" pageId="149" pageNumber="150">Sénégal</collectingCountry>
”), but this is incorrect (
<treatmentCitation id="1458E16130CF52F1EE17FADF749404AC" author="Gillet J. J. E." box="[768,920,1400,1427]" pageId="149" pageNumber="150" year="1911">
<bibRefCitation id="F168BA8130CF52F1EE17FADF749404AC" author="Gillet J. J. E." box="[768,920,1400,1427]" pageId="149" pageNumber="150" pagination="315 - 319" refId="ref121188" refString="Gillet J. J. E. 1911 a. Lamellicornes coprophages nouveaux ou peu connus d'Amerique du Sud. Annales de la Societe entomologique de Belgique 55: 315 - 319. https: // doi. org / 10.5962 / bhl. part. 2978" type="journal article" year="1911">Gillet 1911a</bibRefCitation>
</treatmentCitation>
). Name-bearing type:
<typeStatus id="4A4279D230CF52F1E988FADE720A04AC" box="[1183,1286,1401,1427]" pageId="149" pageNumber="150" type="syntype">syntypes</typeStatus>
(RBINS) (
<treatmentCitation id="1458E16130CF52F1EDE4FA3B76720489" author="Gillet J. J. E." box="[243,382,1436,1462]" pageId="149" pageNumber="150" year="1910">
<bibRefCitation id="F168BA8130CF52F1EDE4FA3B76720489" author="Gillet J. J. E." box="[243,382,1436,1462]" pageId="149" pageNumber="150" pagination="1 - 31" refId="ref121131" refString="Gillet J. J. E. 1910. Especes nouvelles du genre Copris et releve synonymique des especes decrites a ce jour. Notes from the Leyden Museum 32: 1 - 31. Available from https: // repository. naturalis. nl / pub / 508947 [accessed 16 Feb. 2024]." type="journal article" year="1910">Gillet 1910</bibRefCitation>
</treatmentCitation>
), not examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CF52F1EDAAFA7A721704C8" blockId="149.[189,1399,1501,1985]" box="[189,1307,1501,1528]" pageId="149" pageNumber="150">
<treatmentCitationGroup id="B5E9E05E30CF52F1EDAAFA7A721704C8" box="[189,1307,1501,1528]" pageId="149" pageNumber="150">
<taxonomicName id="52F9BCF330CF52F1EDAAFA7A76A104C8" ID-CoL="7YY98" authorityName="Fabricius" authorityYear="1775" box="[189,429,1501,1527]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CF52F1EDAAFA7A76A104C8" box="[189,429,1501,1527]" italics="true" pageId="149" pageNumber="150">Scarabaeus sulcator</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CF52F1ECDCFA7A759704C8" author="Olivier G. A." box="[459,667,1501,1527]" page="142" pageId="149" pageNumber="150" year="1789">
<bibRefCitation id="F168BA8130CF52F1ECDCFA7A759704C8" author="Olivier G. A." box="[459,667,1501,1527]" pageId="149" pageNumber="150" refId="ref133896" refString="Olivier G. A. 1789. N ° 3. Scarabe. Scarabaeus. In: Olivier G. A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome Premier. Baudouin, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 61905" type="book" year="1789">Olivier 1789: 142</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130CF52F1EE94FA7A735204C8" author="Fabricius J. C." box="[899,1118,1501,1527]" page="61" pageId="149" pageNumber="150" year="1792">
<bibRefCitation id="F168BA8130CF52F1EE94FA7A735204C8" author="Fabricius J. C." box="[899,1118,1501,1527]" pageId="149" pageNumber="150" refId="ref118573" refString="Fabricius J. C. 1792. Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tom. I. Pars II. Christ. Gottl. Proft., Copenhagen [Hafniae]. [Publication date following Bousquet 2016.] https: // doi. org / 10.5962 / bhl. title. 122153" type="journal volume" year="1792">Fabricius 1792: 61</bibRefCitation>
</treatmentCitation>
(redescription).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CF52F1EDAAF9A676A10701" blockId="149.[189,1399,1501,1985]" pageId="149" pageNumber="150">
<treatmentCitationGroup id="B5E9E05E30CF52F1EDAAF9A676A10701" pageId="149" pageNumber="150">
<taxonomicName id="52F9BCF330CF52F1EDAAF9A676780725" box="[189,372,1536,1563]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CF52F1EDAAF9A676780725" box="[189,372,1536,1563]" italics="true" pageId="149" pageNumber="150">Copris sulcator</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CF52F1EC87F9A775670724" author="Fabricius J. C." box="[400,619,1536,1563]" page="53" pageId="149" pageNumber="150" year="1801">
<bibRefCitation id="F168BA8130CF52F1EC87F9A775670724" author="Fabricius J. C." box="[400,619,1536,1563]" pageId="149" pageNumber="150" refId="ref118644" refString="Fabricius J. C. 1801. Systema eleutheratorum, secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Tomus I. Bibliopolii Academici Novi, Kiel [Kiliae]. [Publication date following Bousquet 2016.]" type="journal volume" year="1801">Fabricius 1801: 53</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130CF52F1EE46F9A7732B0725" author="Olivier G. A." box="[849,1063,1536,1563]" page="168" pageId="149" pageNumber="150" year="1790">
<bibRefCitation id="F168BA8130CF52F1EE46F9A7732B0725" author="Olivier G. A." box="[849,1063,1536,1563]" pageId="149" pageNumber="150" pagination="1 - 368" refId="ref133979" refString="Olivier G. A. 1790. [livraison 41: 1 - 368]. In: Olivier G. A. 1790 - 1791. Encyclopedie methodique, ou par ordre de matieres. Histoire naturelle. Insectes. Tome cinquieme. Paris, Panckoucke. [Publication date following Bousquet 2016.] https: // doi. org / 10.5962 / bhl. title. 82248" type="journal article" year="1790">Olivier, 1790: 168</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130CF52F1E81AF9A6765D0701" author="Martinez A." page="71" pageId="149" pageNumber="150" year="1959">
<bibRefCitation id="F168BA8130CF52F1E81AF9A6765D0701" author="Martinez A." pageId="149" pageNumber="150" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 71</bibRefCitation>
</treatmentCitation>
(cited).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CF52F1EDAAF9E074A007BB" blockId="149.[189,1399,1501,1985]" pageId="149" pageNumber="150">
<treatmentCitationGroup id="B5E9E05E30CF52F1EDAAF9E074A007BB" pageId="149" pageNumber="150">
<taxonomicName id="52F9BCF330CF52F1EDAAF9E07658075E" ID-CoL="7VDLM" authorityName="Castelnau" authorityYear="1840" box="[189,340,1607,1633]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="nisus">
<emphasis id="A78D1B6230CF52F1EDAAF9E07658075E" box="[189,340,1607,1633]" italics="true" pageId="149" pageNumber="150">Copris nisus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CF52F1EC6EF9E07572075E" author="Blanchard E." box="[377,638,1607,1633]" page="180" pageId="149" pageNumber="150" year="1846">
<bibRefCitation id="F168BA8130CF52F1EC6EF9E07572075E" author="Blanchard E." box="[377,638,1607,1633]" pageId="149" pageNumber="150" pagination="153 - 168" refId="ref111318" refString="Blanchard E. 1846. [livraison 79: 153 - 168, pl. 28; livraison 80: 169 - 184, pl. 31]. In: Blanchard E. &amp; Brulle G. A. 1835 - 1846. Voyage dans l'Amerique Meridionale (le Bresil, la Republique orientale de L'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833, par Alcide d'Orbigny. Tome Sixieme. 2. e partie: Insectes. P. Bertrand, Paris, and Ve. Lerrault, Strasbourg. [Both livraisons date from 1846. For details on the publication date of livraison 79, see Cupello et al. (2023 a: 89 - 90); the earliest evidence of publication of livraison 80 dates from 18 Dec. 1846 (Neal Evenhuis, pers. com. to MC, 3 Sep. 2019).]" type="journal article" year="1846">Blanchard, 1846: 180</bibRefCitation>
</treatmentCitation>
(comments). —
<treatmentCitation id="1458E16130CF52F1EE46F9E07354075E" author="Burmeister H." box="[849,1112,1607,1633]" page="126" pageId="149" pageNumber="150" year="1874">
<bibRefCitation id="F168BA8130CF52F1EE46F9E07354075E" author="Burmeister H." box="[849,1112,1607,1633]" pageId="149" pageNumber="150" pagination="120 - 133" refId="ref113171" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133. Available from https: // www. biodiversitylibrary. org / page / 9236010 [accessed 14 Feb. 2024]." type="journal article" year="1874">Burmeister 1874: 126</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E030CF52F1E9DFF9E0724E075E" box="[1224,1346,1607,1633]" name="Argentina" pageId="149" pageNumber="150">Argentina</collectingCountry>
). —
<treatmentCitation id="1458E16130CF52F1EDFDF9CD76CF07BB" author="Martinez A." box="[234,451,1642,1669]" page="71" pageId="149" pageNumber="150" year="1959">
<bibRefCitation id="F168BA8130CF52F1EDFDF9CD76CF07BB" author="Martinez A." box="[234,451,1642,1669]" pageId="149" pageNumber="150" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 71</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CF52F1EFD0F9CD749707BB" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[711,923,1642,1668]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CF52F1EFD0F9CD749707BB" box="[711,923,1642,1668]" italics="true" pageId="149" pageNumber="150">Ontherus sulcator</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CF52F3EDAAF92A73780079" blockId="149.[189,1399,1501,1985]" lastBlockId="151.[189,1399,265,1100]" lastPageId="151" lastPageNumber="152" pageId="149" pageNumber="150">
<treatmentCitationGroup id="B5E9E05E30CF52F3EDAAF92A73780079" lastPageId="151" lastPageNumber="152" pageId="149" pageNumber="150">
<taxonomicName id="52F9BCF330CF52F1EDAAF92A769E0798" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[189,402,1677,1703]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CF52F1EDAAF92A769E0798" box="[189,402,1677,1703]" italics="true" pageId="149" pageNumber="150">Ontherus sulcator</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CF52F1ECA5F92A75AB0798" author="Erichson W. F." box="[434,679,1677,1703]" page="761" pageId="149" pageNumber="150" year="1847">
<bibRefCitation id="F168BA8130CF52F1ECA5F92A75AB0798" author="Erichson W. F." box="[434,679,1677,1703]" pageId="149" pageNumber="150" pagination="1 - 968" refId="ref117862" refString="Erichson W. F. 1847 b. Naturgeschichte des Insecten Deutschland. I. Coleoptera, Scaphidilia, Scarabaeides. Erste Antheilung, Dritter Band, Erste Lieferung. Nicolaischen Buchhandlung, Berlin. 1 - 968. https: // doi. org / 10.5962 / bhl. title. 50882" type="book chapter" year="1847">Erichson 1847b: 761</bibRefCitation>
</treatmentCitation>
(comments). —
<treatmentCitation id="1458E16130CF52F1EE67F92A73320797" author="Harold E." box="[880,1086,1677,1704]" page="59" pageId="149" pageNumber="150" year="1869">
<bibRefCitation id="F168BA8130CF52F1EE67F92A73320797" author="Harold E." box="[880,1086,1677,1704]" pageId="149" pageNumber="150" pagination="46 - 70" refId="ref123583" refString="Harold E. 1869 c. Ueber Coprophage Lamellicornien mit besonderer Berucksichtigung der Pariser Sammlungen. Coleopterologische Hefte 5: 46 - 70. Available from https: // www. biodiversitylibrary. org / page / 16069721 [accessed 16 Feb. 2024]." type="journal article" year="1869">Harold 1869c: 59</bibRefCitation>
</treatmentCitation>
(comments);
<treatmentCitation id="1458E16130CF52F1E9F4F92A727B0798" author="Harold E." box="[1251,1399,1677,1704]" page="1008" pageId="149" pageNumber="150" year="1869">
<bibRefCitation id="F168BA8130CF52F1E9F4F92A727B0798" author="Harold E." box="[1251,1399,1677,1704]" firstAuthor="d" pageId="149" pageNumber="150" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">1869d: 1008</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130CF52F1ECBFF91775A307F5" author="Burmeister H." box="[424,687,1712,1739]" page="126" pageId="149" pageNumber="150" year="1874">
<bibRefCitation id="F168BA8130CF52F1ECBFF91775A307F5" author="Burmeister H." box="[424,687,1712,1739]" pageId="149" pageNumber="150" pagination="120 - 133" refId="ref113171" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133. Available from https: // www. biodiversitylibrary. org / page / 9236010 [accessed 14 Feb. 2024]." type="journal article" year="1874">Burmeister 1874: 126</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E030CF52F1EE0AF917749B07F5" box="[797,919,1712,1738]" name="Argentina" pageId="149" pageNumber="150">Argentina</collectingCountry>
). —
<treatmentCitation id="1458E16130CF52F1EEC1F91773A507F4" author="Gillet J. J. E." box="[982,1193,1712,1739]" page="319" pageId="149" pageNumber="150" year="1911">
<bibRefCitation id="F168BA8130CF52F1EEC1F91773A507F4" author="Gillet J. J. E." box="[982,1193,1712,1739]" pageId="149" pageNumber="150" pagination="315 - 319" refId="ref121188" refString="Gillet J. J. E. 1911 a. Lamellicornes coprophages nouveaux ou peu connus d'Amerique du Sud. Annales de la Societe entomologique de Belgique 55: 315 - 319. https: // doi. org / 10.5962 / bhl. part. 2978" type="journal article" year="1911">Gillet 1911a: 319</bibRefCitation>
</treatmentCitation>
(list);
<treatmentCitation id="1458E16130CF52F1E9E9F917727A07F4" author="Gillet J. J. E." box="[1278,1398,1712,1739]" page="58" pageId="149" pageNumber="150" year="1911">
<bibRefCitation id="F168BA8130CF52F1E9E9F917727A07F4" author="Gillet J. J. E." box="[1278,1398,1712,1739]" firstAuthor="b" pageId="149" pageNumber="150" pagination="315 - 319" refId="ref121188" refString="Gillet J. J. E. 1911 a. Lamellicornes coprophages nouveaux ou peu connus d'Amerique du Sud. Annales de la Societe entomologique de Belgique 55: 315 - 319. https: // doi. org / 10.5962 / bhl. part. 2978" type="journal article" year="1911">1911b: 58</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130CF52F1ECBCF973757607D1" author="Bruch C." box="[427,634,1747,1774]" page="541" pageId="149" pageNumber="150" year="1915">
<bibRefCitation id="F168BA8130CF52F1ECBCF973757607D1" author="Bruch C." box="[427,634,1747,1774]" pageId="149" pageNumber="150" pagination="538 - 573" refId="ref112550" refString="Bruch C. 1915. Suplemento al catalogo sistematico de los coleopteros de la Republica Argentina. I (addenda, corrigenda y resumen). Revista del Museo de La Plata 19 (2): 538 - 573. Available from http: // sedici. unlp. edu. ar / handle / 10915 / 155145 [accessed 14 Feb. 2024]." type="journal article" year="1915">Bruch 1915: 541</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130CF52F1EFEDF97374CB07D1" author="Lucas R." box="[762,967,1747,1774]" page="459" pageId="149" pageNumber="150" year="1920">
<bibRefCitation id="F168BA8130CF52F1EFEDF97374CB07D1" author="Lucas R." box="[762,967,1747,1774]" pageId="149" pageNumber="150" refId="ref128561" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Nicolaische, Berlin." type="book" year="1920">Lucas 1920: 459</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130CF52F1E984F973761A062E" author="Luederwaldt H." page="106" pageId="149" pageNumber="150" year="1930">
<bibRefCitation id="F168BA8130CF52F1E984F973761A062E" author="Luederwaldt H." pageId="149" pageNumber="150" pagination="105 - 108" refId="ref128717" refString="Luederwaldt H. 1930. Especies novas de Ontherus. Archivos do Instituto Biologico 3: 105 - 108." type="journal article" year="1930">Luederwaldt 1930: 106</bibRefCitation>
</treatmentCitation>
(comments);
<treatmentCitation id="1458E16130CF52F1ECA8F9507544062E" author="Luederwaldt H." box="[447,584,1783,1809]" page="371" pageId="149" pageNumber="150" year="1931">
<bibRefCitation id="F168BA8130CF52F1ECA8F9507544062E" author="Luederwaldt H." box="[447,584,1783,1809]" firstAuthor="a" pageId="149" pageNumber="150" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">1931a: 371</bibRefCitation>
</treatmentCitation>
, 385, 389, 392, 408, 411, 416 (key, synonymy). —
<treatmentCitation id="1458E16130CF52F1E9D6F9507668060A" author="Pessoa S. B. &amp; Lane F." page="456" pageId="149" pageNumber="150" year="1941">
<bibRefCitation id="F168BA8130CF52F1E9D6F9507668060A" author="Pessoa S. B. &amp; Lane F." pageId="149" pageNumber="150" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 456</bibRefCitation>
</treatmentCitation>
, 458 (key). —
<treatmentCitation id="1458E16130CF52F1EF0FF8BD7427060B" author="Blackwelder R. E." box="[536,811,1818,1844]" page="206" pageId="149" pageNumber="150" year="1944">
<bibRefCitation id="F168BA8130CF52F1EF0FF8BD7427060B" author="Blackwelder R. E." box="[536,811,1818,1844]" pageId="149" pageNumber="150" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="1458E16130CF52F1EEF3F8BD73B1060B" author="Martinez A." box="[996,1213,1818,1845]" page="71" pageId="149" pageNumber="150" year="1959">
<bibRefCitation id="F168BA8130CF52F1EEF3F8BD73B1060B" author="Martinez A." box="[996,1213,1818,1845]" pageId="149" pageNumber="150" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 71</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130CF52F1EDFCF89A755B0667" author="Vulcano M. A. &amp; Pereira F. S." box="[235,599,1853,1880]" page="583" pageId="149" pageNumber="150" year="1967">
<bibRefCitation id="F168BA8130CF52F1EDFCF89A755B0667" author="Vulcano M. A. &amp; Pereira F. S." box="[235,599,1853,1880]" pageId="149" pageNumber="150" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira, 1967: 583</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130CF52F1EFCEF89A730E0668" author="Forsyth A. B. &amp; Gill B. D." box="[729,1026,1853,1879]" page="70" pageId="149" pageNumber="150" year="1993">
<bibRefCitation id="F168BA8130CF52F1EFCEF89A730E0668" author="Forsyth A. B. &amp; Gill B. D." box="[729,1026,1853,1879]" pageId="149" pageNumber="150" refId="ref119487" refString="Forsyth A. B. &amp; Gill B. D. 1993. Invertebrate indicators. In: Foster R. B., Emmons L. H., Freed P., Forsyth A. B., Hoffman B. &amp; Gill B. D. (eds) A Biological Assessment of the Kanuku Mountain Region of Southwestern Guyana: 32 - 33 + appendix 6. Rapid Assessment Program, RAP Working Papers. Conservation International, Washington, D. C." type="journal volume" year="1993">Forsyth &amp; Gill 1993: 70</bibRefCitation>
</treatmentCitation>
(as
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<emphasis id="A78D1B6230CF52F1E92EF89973CC0668" box="[1081,1216,1853,1880]" italics="true" pageId="149" pageNumber="150">O. suclator</emphasis>
</taxonomicName>
[sic], cited for
<collectingCountry id="EDEE87E030CF52F1EDFDF8C676470644" box="[234,331,1889,1915]" name="Guyana" pageId="149" pageNumber="150">Guyana</collectingCountry>
). —
<treatmentCitation id="1458E16130CF52F1EC95F8C775520644" author="Escobar F." box="[386,606,1888,1915]" page="209" pageId="149" pageNumber="150" year="2000">
<bibRefCitation id="F168BA8130CF52F1EC95F8C775520644" author="Escobar F." box="[386,606,1888,1915]" pageId="149" pageNumber="150" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 209</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130CF52F1EFC7F8C774E20644" author="Vaz-de-Mello F. Z." box="[720,1006,1888,1915]" page="194" pageId="149" pageNumber="150" year="2000">
<bibRefCitation id="F168BA8130CF52F1EFC7F8C774E20644" author="Vaz-de-Mello F. Z." box="[720,1006,1888,1915]" pageId="149" pageNumber="150" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 194</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030CF52F1E98FF8C773E90645" box="[1176,1253,1888,1914]" name="Brazil" pageId="149" pageNumber="150">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E16130CF52F1E80AF8C776A006A1" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." page="139" pageId="149" pageNumber="150" year="2001">
<bibRefCitation id="F168BA8130CF52F1E80AF8C776A006A1" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." pageId="149" pageNumber="150" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B6230CF52F1EDFDF823762406A1" box="[234,296,1924,1950]" italics="true" pageId="149" pageNumber="150">et al.</emphasis>
2001: 139
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030CF52F1EF48F82375D406A1" box="[607,728,1924,1950]" name="Colombia" pageId="149" pageNumber="150">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130CF52F1EE01F823737006A2" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[790,1148,1923,1950]" page="16" pageId="149" pageNumber="150" year="2006">
<bibRefCitation id="F168BA8130CF52F1EE01F823737006A2" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[790,1148,1923,1950]" pageId="149" pageNumber="150" pagination="1 - 18" refId="ref122830" refString="Hamel-Leigue A., Mann D. J., Vaz-de-Mello F. Z. &amp; Herzog S. K. 2006. Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeinae) de Bolivia: primera compilacion de los generos y especies registrados para el pais. Revista Boliviana de Ecologia y Conservacion Ambiental 20: 1 - 18." type="journal article" year="2006">
Hamel-Leigue
<emphasis id="A78D1B6230CF52F1EEDDF823730406A1" box="[970,1032,1924,1950]" italics="true" pageId="149" pageNumber="150">et al.</emphasis>
2006: 16
</bibRefCitation>
</treatmentCitation>
(inventory, cited for
<collectingCountry id="EDEE87E030CF52F1EDFDF800764A06FE" box="[234,326,1959,1985]" name="Bolivia" pageId="149" pageNumber="150">Bolivia</collectingCountry>
). —
<treatmentCitation id="1458E16130CF52F1EC97F800752906FE" author="Feer F." box="[384,549,1959,1985]" page="62" pageId="149" pageNumber="150" year="2008">
<bibRefCitation id="F168BA8130CF52F1EC97F800752906FE" author="Feer F." box="[384,549,1959,1985]" pageId="149" pageNumber="150" pagination="49 - 62" refId="ref118745" refString="Feer F. 2008. Responses of dung beetle assemblages to characteristics of rain forest edges. Ecotropica 14: 49 - 62." type="journal article" year="2008">Feer 2008: 62</bibRefCitation>
</treatmentCitation>
(ecology, list). —
<treatmentCitation id="1458E16130CF52F1EE11F800732506FE" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." box="[774,1065,1959,1985]" page="1294" pageId="149" pageNumber="150" year="2008">
<bibRefCitation id="F168BA8130CF52F1EE11F800732506FE" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." box="[774,1065,1959,1985]" pageId="149" pageNumber="150" pagination="1288 - 1298" refId="ref127038" refString="Larsen T. H., Lopera A. &amp; Forsyth A. 2008. Understanding trait-dependent community disassembly: dung beetles, density functions, and forest fragmentation. Conservation Biology 22 (5): 1288 - 1298. https: // doi. org / 10.1111 / j. 1523 - 1739.2008.00969. x" type="journal article" year="2008">
Larsen
<emphasis id="A78D1B6230CF52F1EE77F80F749006FE" box="[864,924,1959,1985]" italics="true" pageId="149" pageNumber="150">et al.</emphasis>
2008: 1294
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130CF52F1E988F800727A06FE" author="Krajcik M." box="[1183,1398,1959,1985]" page="174" pageId="149" pageNumber="150" year="2012">
<bibRefCitation id="F168BA8130CF52F1E988F800727A06FE" author="Krajcik M." box="[1183,1398,1959,1985]" pageId="149" pageNumber="150" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 174</bibRefCitation>
</treatmentCitation>
(checklist, cited for Cayenne). —
<treatmentCitation id="1458E16130CD52F3EF6AFEAE7301001C" author="Boilly O. &amp; Vaz-de-Mello F. Z." box="[637,1037,265,291]" page="112" pageId="151" pageNumber="152" year="2013">
<bibRefCitation id="F168BA8130CD52F3EF6AFEAE7301001C" author="Boilly O. &amp; Vaz-de-Mello F. Z." box="[637,1037,265,291]" pageId="151" pageNumber="152" pagination="103 - 112" refId="ref111991" refString="Boilly O. &amp; Vaz-de-Mello F. Z. 2013. Les Scarabaeinae de Guyane: cle illustree des genres. ACOREP- France 7: 103 - 112." type="journal article" year="2013">Boilly &amp; Vaz-de-Mello 2013: 112</bibRefCitation>
</treatmentCitation>
(
<figureCitation id="0DC2DBF530CD52F3E90AFEAE7379001C" box="[1053,1141,265,291]" captionStart="Fig" captionStartId="109.[189,232,1963,1989]" captionTargetBox="[276,1305,278,1887]" captionTargetId="figure-12@109.[265,1323,265,1923]" captionTargetPageId="109" captionText="Fig. 43. Species distribution. A. Dichotomius satanas (Harold, 1867). B. Dichotomius tristis (Luederwaldt, 1923). Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945583" httpUri="https://zenodo.org/record/13945583/files/figure.png" pageId="151" pageNumber="152">Fig. 43</figureCitation>
, key). —
<treatmentCitation id="1458E16130CD52F3E9FCFEAE76AF0079" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." page="109" pageId="151" pageNumber="152" year="2013">
<bibRefCitation id="F168BA8130CD52F3E9FCFEAE76AF0079" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." pageId="151" pageNumber="152" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B6230CD52F3EDFCFE8A762A0079" box="[235,294,300,326]" italics="true" pageId="151" pageNumber="152">et al.</emphasis>
2013: 109
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130CD52F3EF02FE8B74360079" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." box="[533,826,300,327]" page="197" pageId="151" pageNumber="152" year="2015">
<bibRefCitation id="F168BA8130CD52F3EF02FE8B74360079" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." box="[533,826,300,327]" pageId="151" pageNumber="152" pagination="186 - 207" refId="ref137018" refString="Ratcliffe B. C., Jameson M. L., Figueroa L., Cave R. D., Paulsen M. J., Cano E. B., Beza-Beza C., Jimenez- Ferbans L., Reyes-Castillo P. 2015. Beetles (Coleoptera) of Peru: a survey of the families. Journal of the Kansas Entomological Society 88 (2): 186 - 207." type="journal article" year="2015">
Ratcliffe
<emphasis id="A78D1B6230CD52F3EF95FE8A75B10079" box="[642,701,300,326]" italics="true" pageId="151" pageNumber="152">et al.</emphasis>
2015: 197
</bibRefCitation>
</treatmentCitation>
(checklist, cited for
<collectingCountry id="EDEE87E030CD52F3E93BFE8B736B0079" box="[1068,1127,300,326]" name="Peru" pageId="151" pageNumber="152">Peru</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<caption id="C18697F830CC52F2EDAAF80C751E06D7" ID-DOI="http://doi.org/10.5281/zenodo.13945625" ID-Zenodo-Dep="13945625" httpUri="https://zenodo.org/record/13945625/files/figure.png" pageId="150" pageNumber="151" startId="150.[189,232,1963,1989]" targetBox="[274,1302,278,1887]" targetPageId="150">
<paragraph id="9546C77030CC52F2EDAAF80C751E06D7" blockId="150.[188,1398,1963,2025]" pageId="150" pageNumber="151">
<emphasis id="A78D1B6230CC52F2EDAAF80C761606FA" bold="true" box="[189,282,1963,1989]" pageId="150" pageNumber="151">Fig. 58.</emphasis>
Species distribution.
<emphasis id="A78D1B6230CC52F2EF03F80C752506FA" bold="true" box="[532,553,1963,1989]" pageId="150" pageNumber="151">A</emphasis>
.
<taxonomicName id="52F9BCF330CC52F2EF2FF80C74AC06FA" authority="Genier, 1996" authorityName="Genier" authorityYear="1996" box="[568,928,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="species" species="pubens">
<emphasis id="A78D1B6230CC52F2EF2FF80C75F306FA" box="[568,767,1963,1989]" italics="true" pageId="150" pageNumber="151">Ontherus pubens</emphasis>
<bibRefCitation id="F168BA8130CC52F2EE12F80C74AC06FA" author="Genier F." box="[773,928,1963,1989]" pageId="150" pageNumber="151" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier, 1996</bibRefCitation>
</taxonomicName>
.
<emphasis id="A78D1B6230CC52F2EEBEF80C74B106FA" bold="true" box="[937,957,1963,1989]" pageId="150" pageNumber="151">B</emphasis>
.
<taxonomicName id="52F9BCF330CC52F2EEDBF80C727E06F9" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[972,1394,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CC52F2EEDBF80C739306FA" box="[972,1183,1963,1989]" italics="true" pageId="150" pageNumber="151">Ontherus sulcator</emphasis>
(
<bibRefCitation id="F168BA8130CC52F2E9BAF80C726506F9" author="Fabricius J. C." box="[1197,1385,1963,1990]" pageId="150" pageNumber="151" refId="ref118353" refString="Fabricius J. C. 1775. Systema entomologiae, sistens insectorvm classes, ordines, genera, species, adiectis synonymis, locis, descriptionibvs, observationibvs. Kortius, Flensburg and Leipzig [Flensbvrgi et Lipsiae]. https: // doi. org / 10.5962 / bhl. title. 36510" type="book" year="1775">Fabricius, 1775</bibRefCitation>
)
</taxonomicName>
. Black circle = literature data.
</paragraph>
</caption>
<paragraph id="9546C77030CD52F3EDAAFEF77437008F" blockId="151.[189,1399,265,1100]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAFEF77437008F" pageId="151" pageNumber="152">
<taxonomicName id="52F9BCF330CD52F3EDAAFEF776900056" ID-CoL="7VDLN" authorityName="Blanchard" authorityYear="1846" box="[189,412,335,362]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="ovalipennis">
<emphasis id="A78D1B6230CD52F3EDAAFEF776900056" box="[189,412,335,362]" italics="true" pageId="151" pageNumber="152">Copris ovalipennis</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CD52F3ECADFEE8759A0055" author="Harold E." box="[442,662,335,362]" page="503" pageId="151" pageNumber="152" year="1869">
<bibRefCitation id="F168BA8130CD52F3ECADFEE8759A0055" author="Harold E." box="[442,662,335,362]" pageId="151" pageNumber="152" pagination="493 - 512" refId="ref123508" refString="Harold E. 1869 a. Note sur quelques Coprides du Mexique. Annales de la Societe Entomologique de France, Quatrieme Serie 9: 493 - 512. Available from https: // www. biodiversitylibrary. org / page / 32547053 [accessed 14 Feb. 2024]." type="journal article" year="1869">Harold 1869a: 503</bibRefCitation>
</treatmentCitation>
(comments); 1869c: 59 (comments). —
<treatmentCitation id="1458E16130CD52F3E961FEE8727B0056" author="Burmeister H." box="[1142,1399,335,361]" page="126" pageId="151" pageNumber="152" year="1874">
<bibRefCitation id="F168BA8130CD52F3E961FEE8727B0056" author="Burmeister H." box="[1142,1399,335,361]" pageId="151" pageNumber="152" pagination="120 - 133" refId="ref113171" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133. Available from https: // www. biodiversitylibrary. org / page / 9236010 [accessed 14 Feb. 2024]." type="journal article" year="1874">Burmeister 1874: 126</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E030CD52F3EC5FFED576CE00B3" box="[328,450,370,396]" name="Argentina" pageId="151" pageNumber="152">Argentina</collectingCountry>
). —
<treatmentCitation id="1458E16130CD52F3ECEDFED5741500B3" author="Luederwaldt H." box="[506,793,370,397]" page="386" pageId="151" pageNumber="152" year="1931">
<bibRefCitation id="F168BA8130CD52F3ECEDFED5741500B3" author="Luederwaldt H." box="[506,793,370,397]" pageId="151" pageNumber="152" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 386</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3E909FED573C300B3" ID-CoL="7VDLM" baseAuthorityName="Kirsch" baseAuthorityYear="1871" box="[1054,1231,370,396]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="nisus">
<emphasis id="A78D1B6230CD52F3E909FED573C300B3" box="[1054,1231,370,396]" italics="true" pageId="151" pageNumber="152">Ontherus nisus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3E81AFED4765E008F" author="Martinez A." page="71" pageId="151" pageNumber="152" year="1959">
<bibRefCitation id="F168BA8130CD52F3E81AFED4765E008F" author="Martinez A." pageId="151" pageNumber="152" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 71</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EF41FE317426008F" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[598,810,406,432]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EF41FE317426008F" box="[598,810,406,432]" italics="true" pageId="151" pageNumber="152">Ontherus sulcator</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFE1E75E7035F" blockId="151.[189,1399,265,1100]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAFE1E75E7035F" pageId="151" pageNumber="152">
<taxonomicName id="52F9BCF330CD52F3EDAAFE1E767C00EC" ID-CoL="7VDLM" baseAuthorityName="Kirsch" baseAuthorityYear="1871" box="[189,368,441,467]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="nisus">
<emphasis id="A78D1B6230CD52F3EDAAFE1E767C00EC" box="[189,368,441,467]" italics="true" pageId="151" pageNumber="152">Ontherus nisus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CD52F3EC85FE1E758F00EC" author="Harold E." box="[402,643,441,467]" page="1008" pageId="151" pageNumber="152" year="1869">
<bibRefCitation id="F168BA8130CD52F3EC85FE1E758F00EC" author="Harold E." box="[402,643,441,467]" pageId="151" pageNumber="152" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1008</bibRefCitation>
</treatmentCitation>
(catalogue, cited as synonym of
<taxonomicName id="52F9BCF330CD52F3E903FE1E739500EC" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[1044,1177,441,467]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3E903FE1E739500EC" box="[1044,1177,441,467]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3E9CDFE1E760600C8" author="Gillet J. J. E." page="58" pageId="151" pageNumber="152" year="1911">
<bibRefCitation id="F168BA8130CD52F3E9CDFE1E760600C8" author="Gillet J. J. E." pageId="151" pageNumber="152" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 58</bibRefCitation>
</treatmentCitation>
(catalog, cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EF7AFE7A75FD00C9" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[621,753,476,503]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EF7AFE7A75FD00C9" box="[621,753,476,503]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3EE39FE7B734300C9" author="Luederwaldt H." box="[814,1103,476,502]" page="387" pageId="151" pageNumber="152" year="1931">
<bibRefCitation id="F168BA8130CD52F3EE39FE7B734300C9" author="Luederwaldt H." box="[814,1103,476,502]" pageId="151" pageNumber="152" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 387</bibRefCitation>
</treatmentCitation>
, 392, 399, 400, 409, 411, 417 (redescription, distribution, key). —
<treatmentCitation id="1458E16130CD52F3EFCAFDA773170325" author="Pessoa S. B. &amp; Lane F." box="[733,1051,511,538]" page="457" pageId="151" pageNumber="152" year="1941">
<bibRefCitation id="F168BA8130CD52F3EFCAFDA773170325" author="Pessoa S. B. &amp; Lane F." box="[733,1051,511,538]" pageId="151" pageNumber="152" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 457</bibRefCitation>
</treatmentCitation>
, 458 (key, redescription). —
<treatmentCitation id="1458E16130CD52F3EDFCFD8576F30303" author="Blackwelder R. E." box="[235,511,546,573]" page="206" pageId="151" pageNumber="152" year="1944">
<bibRefCitation id="F168BA8130CD52F3EDFCFD8576F30303" author="Blackwelder R. E." box="[235,511,546,573]" pageId="151" pageNumber="152" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EE1DFD8474820303" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[778,910,546,573]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EE1DFD8474820303" box="[778,910,546,573]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3EED9FD84739B0302" author="Lange R. B." box="[974,1175,547,573]" page="310" pageId="151" pageNumber="152" year="1947">
<bibRefCitation id="F168BA8130CD52F3EED9FD84739B0302" author="Lange R. B." box="[974,1175,547,573]" pageId="151" pageNumber="152" pagination="305 - 315" refId="ref126831" refString="Lange R. B. 1947. Ensaio da zoogeografia dos Scarabaeidae do Parana com algumas notas etnoecologicas. Arquivos Museu Paranaense 6: 305 - 315." type="journal article" year="1947">Lange 1947: 310</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130CD52F3E81AFD84765F035F" author="Martinez A." page="71" pageId="151" pageNumber="152" year="1959">
<bibRefCitation id="F168BA8130CD52F3E81AFD84765F035F" author="Martinez A." pageId="151" pageNumber="152" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 71</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EF41FDE175D6035F" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[598,730,582,608]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EF41FDE175D6035F" box="[598,730,582,608]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFDCD73BB03BC" blockId="151.[189,1399,265,1100]" box="[189,1207,617,643]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAFDCD73BB03BC" box="[189,1207,617,643]" pageId="151" pageNumber="152">
<taxonomicName id="52F9BCF330CD52F3EDAAFDCD76A903BC" authorityName="Luederwaldt" authorityYear="1931" box="[189,421,617,643]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="ovalipennis">
<emphasis id="A78D1B6230CD52F3EDAAFDCD76A903BC" box="[189,421,617,643]" italics="true" pageId="151" pageNumber="152">Pinotus ovalipennis</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CD52F3ECD4FDCE75A303BC" author="Harold E." box="[451,687,617,643]" page="1008" pageId="151" pageNumber="152" year="1869">
<bibRefCitation id="F168BA8130CD52F3ECD4FDCE75A303BC" author="Harold E." box="[451,687,617,643]" pageId="151" pageNumber="152" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1008</bibRefCitation>
</treatmentCitation>
(catalogue, cited for
<collectingCountry id="EDEE87E030CD52F3EEB0FDCE732D03BC" box="[935,1057,617,643]" name="Argentina" pageId="151" pageNumber="152">Argentina</collectingCountry>
:
<collectingRegion id="573D099230CD52F3E93BFDCE73A003BC" box="[1068,1196,617,643]" country="Argentina" name="Corrientes" pageId="151" pageNumber="152">Corrientes</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFD2B753E022F" blockId="151.[189,1399,265,1100]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAFD2B753E022F" pageId="151" pageNumber="152">
<taxonomicName id="52F9BCF330CD52F3EDAAFD2B76B70399" ID-CoL="7VDLN" box="[189,443,652,678]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="ovalipennis">
<emphasis id="A78D1B6230CD52F3EDAAFD2B76B70399" box="[189,443,652,678]" italics="true" pageId="151" pageNumber="152">Ontherus ovalipennis</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CD52F3ECC9FD2B75AE0398" author="Gillet J. J. E." box="[478,674,652,679]" page="58" pageId="151" pageNumber="152" year="1911">
<bibRefCitation id="F168BA8130CD52F3ECC9FD2B75AE0398" author="Gillet J. J. E." box="[478,674,652,679]" pageId="151" pageNumber="152" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 58</bibRefCitation>
</treatmentCitation>
(catalog, cited as synonym of
<taxonomicName id="52F9BCF330CD52F3E90EFD2A73AC0399" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[1049,1184,652,679]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3E90EFD2A73AC0399" box="[1049,1184,652,679]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3E9F5FD2B767D03F6" author="Luederwaldt H." page="387" pageId="151" pageNumber="152" year="1931">
<bibRefCitation id="F168BA8130CD52F3E9F5FD2B767D03F6" author="Luederwaldt H." pageId="151" pageNumber="152" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 387</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EF90FD1775E703F6" ID-CoL="7VDLM" baseAuthorityName="Kirsch" baseAuthorityYear="1871" box="[647,747,688,714]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="nisus">
<emphasis id="A78D1B6230CD52F3EF90FD1775E703F6" box="[647,747,688,714]" italics="true" pageId="151" pageNumber="152">O. nisus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3EE27FD17736003F5" author="Pessoa S. B. &amp; Lane F." box="[816,1132,687,714]" page="458" pageId="151" pageNumber="152" year="1941">
<bibRefCitation id="F168BA8130CD52F3EE27FD17736003F5" author="Pessoa S. B. &amp; Lane F." box="[816,1132,687,714]" pageId="151" pageNumber="152" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 458</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EDFCFD74764603D3" ID-CoL="7VDLM" baseAuthorityName="Kirsch" baseAuthorityYear="1871" box="[235,330,723,749]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="nisus">
<emphasis id="A78D1B6230CD52F3EDFCFD74764603D3" box="[235,330,723,749]" italics="true" pageId="151" pageNumber="152">O. nisus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3EC91FD75759A03D3" author="Blackwelder R. E." box="[390,662,722,749]" page="206" pageId="151" pageNumber="152" year="1944">
<bibRefCitation id="F168BA8130CD52F3EC91FD75759A03D3" author="Blackwelder R. E." box="[390,662,722,749]" pageId="151" pageNumber="152" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EE81FD74731503D3" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[918,1049,722,749]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EE81FD74731503D3" box="[918,1049,722,749]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3E941FD74722103D2" author="Martinez A." box="[1110,1325,722,749]" page="71" pageId="151" pageNumber="152" year="1959">
<bibRefCitation id="F168BA8130CD52F3E941FD74722103D2" author="Martinez A." box="[1110,1325,722,749]" pageId="151" pageNumber="152" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 71</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EC8AFD51752D022F" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[413,545,758,784]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EC8AFD51752D022F" box="[413,545,758,784]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFCBE75E70246" blockId="151.[189,1399,265,1100]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAFCBE75E70246" pageId="151" pageNumber="152">
<taxonomicName id="52F9BCF330CD52F3EDAAFCBE76A4020C" ID-CoL="7VDMF" authorityName="Gillet" authorityYear="1910" box="[189,424,793,819]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="senegalensis">
<emphasis id="A78D1B6230CD52F3EDAAFCBE76A4020C" box="[189,424,793,819]" italics="true" pageId="151" pageNumber="152">Copris senegalensis</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CD52F3ECD1FCBE759F020C" author="Gillet J. J. E." box="[454,659,793,819]" page="319" pageId="151" pageNumber="152" year="1911">
<bibRefCitation id="F168BA8130CD52F3ECD1FCBE759F020C" author="Gillet J. J. E." box="[454,659,793,819]" pageId="151" pageNumber="152" pagination="315 - 319" refId="ref121188" refString="Gillet J. J. E. 1911 a. Lamellicornes coprophages nouveaux ou peu connus d'Amerique du Sud. Annales de la Societe entomologique de Belgique 55: 315 - 319. https: // doi. org / 10.5962 / bhl. part. 2978" type="journal article" year="1911">Gillet 1911a: 319</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EE80FCBE7317020C" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[919,1051,793,819]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EE80FCBE7317020C" box="[919,1051,793,819]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
); 1911b: 58 (catalog, cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EC96FC9A75080269" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[385,516,828,855]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EC96FC9A75080269" box="[385,516,828,855]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3EF53FC9B74070268" author="Bruch C." box="[580,779,828,855]" page="541" pageId="151" pageNumber="152" year="1915">
<bibRefCitation id="F168BA8130CD52F3EF53FC9B74070268" author="Bruch C." box="[580,779,828,855]" pageId="151" pageNumber="152" pagination="538 - 573" refId="ref112550" refString="Bruch C. 1915. Suplemento al catalogo sistematico de los coleopteros de la Republica Argentina. I (addenda, corrigenda y resumen). Revista del Museo de La Plata 19 (2): 538 - 573. Available from http: // sedici. unlp. edu. ar / handle / 10915 / 155145 [accessed 14 Feb. 2024]." type="journal article" year="1915">Bruch 1915: 541</bibRefCitation>
</treatmentCitation>
(list, cited as synonym of
<taxonomicName id="52F9BCF330CD52F3E95EFC9A73C10269" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[1097,1229,828,855]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3E95EFC9A73C10269" box="[1097,1229,828,855]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3E81AFC9B765F0246" author="Martinez A." page="72" pageId="151" pageNumber="152" year="1959">
<bibRefCitation id="F168BA8130CD52F3E81AFC9B765F0246" author="Martinez A." pageId="151" pageNumber="152" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 72</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EF41FCC775D60246" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[598,730,863,890]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EF41FCC775D60246" box="[598,730,863,890]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFC25747302A3" blockId="151.[189,1399,265,1100]" box="[189,895,898,925]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAFC25747302A3" box="[189,895,898,925]" pageId="151" pageNumber="152">
<taxonomicName id="52F9BCF330CD52F3EDAAFC25769E02A3" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[189,402,898,924]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EDAAFC25769E02A3" box="[189,402,898,924]" italics="true" pageId="151" pageNumber="152">Ontherus sulcator</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CD52F3ECB8FC2575C202A3" author="Luederwaldt H." box="[431,718,898,925]" page="364" pageId="151" pageNumber="152" year="1931">
<bibRefCitation id="F168BA8130CD52F3ECB8FC2575C202A3" author="Luederwaldt H." box="[431,718,898,925]" pageId="151" pageNumber="152" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 364</bibRefCitation>
</treatmentCitation>
(
<typeStatus id="4A4279D230CD52F3EFCAFC24741D02A2" box="[733,785,899,925]" pageId="151" pageNumber="152">type</typeStatus>
species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFC01751E0539" blockId="151.[189,1399,265,1100]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAFC01751E0539" pageId="151" pageNumber="152">
<taxonomicName id="52F9BCF330CD52F3EDAAFC0176C002FF" ID-CoL="7VDMF" box="[189,460,934,960]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="senegalensis">
<emphasis id="A78D1B6230CD52F3EDAAFC0176C002FF" box="[189,460,934,960]" italics="true" pageId="151" pageNumber="152">Ontherus senegalensis</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CD52F3ECE1FC01742802FF" author="Luederwaldt H." box="[502,804,934,960]" page="386" pageId="151" pageNumber="152" year="1931">
<bibRefCitation id="F168BA8130CD52F3ECE1FC01742802FF" author="Luederwaldt H." box="[502,804,934,960]" pageId="151" pageNumber="152" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 386</bibRefCitation>
</treatmentCitation>
, 387 (cited as synonym of
<taxonomicName id="52F9BCF330CD52F3E993FC01723602FF" ID-CoL="7VDLM" baseAuthorityName="Kirsch" baseAuthorityYear="1871" box="[1156,1338,934,960]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="nisus">
<emphasis id="A78D1B6230CD52F3E993FC01723602FF" box="[1156,1338,934,960]" italics="true" pageId="151" pageNumber="152">Ontherus nisus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3EDFCFC6E752402DB" author="Pessoa S. B. &amp; Lane F." box="[235,552,969,996]" page="458" pageId="151" pageNumber="152" year="1941">
<bibRefCitation id="F168BA8130CD52F3EDFCFC6E752402DB" author="Pessoa S. B. &amp; Lane F." box="[235,552,969,996]" pageId="151" pageNumber="152" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 458</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EE56FC6E74A902DC" ID-CoL="7VDLM" baseAuthorityName="Kirsch" baseAuthorityYear="1871" box="[833,933,969,995]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="nisus">
<emphasis id="A78D1B6230CD52F3EE56FC6E74A902DC" box="[833,933,969,995]" italics="true" pageId="151" pageNumber="152">O. nisus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130CD52F3EEFCFC6E720802DC" author="Blackwelder R. E." box="[1003,1284,969,995]" page="206" pageId="151" pageNumber="152" year="1944">
<bibRefCitation id="F168BA8130CD52F3EEFCFC6E720802DC" author="Blackwelder R. E." box="[1003,1284,969,995]" pageId="151" pageNumber="152" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330CD52F3EC6AFC4A750D0539" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[381,513,1004,1031]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator">
<emphasis id="A78D1B6230CD52F3EC6AFC4A750D0539" box="[381,513,1004,1031]" italics="true" pageId="151" pageNumber="152">O. sulcator</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFBA8753E0573" blockId="151.[189,1399,265,1100]" pageId="151" pageNumber="152">
<treatmentCitationGroup id="B5E9E05E30CD52F3EDAAFBA8753E0573" pageId="151" pageNumber="152">
<taxonomicName id="52F9BCF330CD52F3EDAAFBA8752D0516" ID-CoL="7ZQ4B" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[189,545,1039,1065]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="sulcator" subGenus="Ontherus">
<emphasis id="A78D1B6230CD52F3EDAAFBA876260516" box="[189,298,1039,1065]" italics="true" pageId="151" pageNumber="152">Ontherus</emphasis>
(
<emphasis id="A78D1B6230CD52F3EC28FBA876A00516" box="[319,428,1039,1065]" italics="true" pageId="151" pageNumber="152">Ontherus</emphasis>
)
<emphasis id="A78D1B6230CD52F3ECD6FBA8752D0516" box="[449,545,1039,1065]" italics="true" pageId="151" pageNumber="152">sulcator</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130CD52F3EF51FBA874340516" author="Genier F." box="[582,824,1039,1065]" page="74" pageId="151" pageNumber="152" year="1996">
<bibRefCitation id="F168BA8130CD52F3EF51FBA874340516" author="Genier F." box="[582,824,1039,1065]" pageId="151" pageNumber="152" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 7476</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="1458E16130CD52F3EEE7FBA8727A0515" ID-DOI="http://doi.org/10.5281/zenodo.3684673" ID-Zenodo-Dep="3684673" author="Hielkema A. J. &amp; Hielkema M. A." box="[1008,1398,1039,1066]" httpUri="http://treatment.plazi.org/id/03D7FF1CFFF1FFD7FF48FD6F338BF893" page="57" pageId="151" pageNumber="152" year="2019">
<bibRefCitation id="F168BA8130CD52F3EEE7FBA8727A0515" author="Hielkema A. J. &amp; Hielkema M. A." box="[1008,1398,1039,1066]" pageId="151" pageNumber="152" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 57</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30CD52F3EDAAFBD374B4044C" pageId="151" pageNumber="152" type="distribution">
<paragraph id="9546C77030CD52F3EDAAFBD3765A05B1" blockId="151.[189,342,1140,1166]" box="[189,342,1140,1166]" pageId="151" pageNumber="152">
<heading id="CE0E701C30CD52F3EDAAFBD3765A05B1" bold="true" box="[189,342,1140,1166]" fontSize="11" level="3" pageId="151" pageNumber="152" reason="3">
<emphasis id="A78D1B6230CD52F3EDAAFBD3765A05B1" bold="true" box="[189,342,1140,1166]" pageId="151" pageNumber="152">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFB05746705DF" blockId="151.[189,1398,1186,1248]" pageId="151" pageNumber="152">
<collectingCountry id="EDEE87E030CD52F3EDAAFB0576380583" box="[189,308,1186,1212]" name="Colombia" pageId="151" pageNumber="152">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E030CD52F3EC57FB0576B00583" box="[320,444,1186,1212]" name="Venezuela" pageId="151" pageNumber="152">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E030CD52F3ECD0FB0575B30583" box="[455,703,1186,1212]" name="Trinidad and Tobago" pageId="151" pageNumber="152">Trinidad and Tobago</collectingCountry>
,
<collectingCountry id="EDEE87E030CD52F3EFDCFB0474250582" box="[715,809,1187,1213]" name="Guyana" pageId="151" pageNumber="152">Guyana</collectingCountry>
,
<collectingCountry id="EDEE87E030CD52F3EE21FB0574A50583" box="[822,937,1186,1212]" name="Suriname" pageId="151" pageNumber="152">Suriname</collectingCountry>
, French Guiana,
<collectingCountry id="EDEE87E030CD52F3E964FB0573B20583" box="[1139,1214,1186,1212]" name="Brazil" pageId="151" pageNumber="152">Brazil</collectingCountry>
,
<collectingCountry id="EDEE87E030CD52F3E9DDFB0572220583" box="[1226,1326,1186,1212]" name="Ecuador" pageId="151" pageNumber="152">Ecuador</collectingCountry>
,
<collectingCountry id="EDEE87E030CD52F3E82DFB04727F0582" box="[1338,1395,1187,1213]" name="Peru" pageId="151" pageNumber="152">Peru</collectingCountry>
,
<collectingCountry id="EDEE87E030CD52F3EDA9FB61761B05DF" box="[190,279,1222,1248]" name="Bolivia" pageId="151" pageNumber="152">Bolivia</collectingCountry>
,
<collectingCountry id="EDEE87E030CD52F3EC34FB61769D05DF" box="[291,401,1222,1248]" name="Paraguay" pageId="151" pageNumber="152">Paraguay</collectingCountry>
,
<collectingCountry id="EDEE87E030CD52F3EC8CFB61751F05DF" box="[411,531,1222,1248]" name="Argentina" pageId="151" pageNumber="152">Argentina</collectingCountry>
, and
<collectingCountry id="EDEE87E030CD52F3EF47FB6175BB05DF" box="[592,695,1222,1248]" name="Uruguay" pageId="151" pageNumber="152">Uruguay</collectingCountry>
(
<bibRefCitation id="F168BA8130CD52F3EFD1FB61745205DF" author="Genier F." box="[710,862,1222,1248]" pageId="151" pageNumber="152" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996</bibRefCitation>
).
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFAA076E1041E" blockId="151.[189,493,1287,1313]" box="[189,493,1287,1313]" pageId="151" pageNumber="152">
<heading id="CE0E701C30CD52F3EDAAFAA076E1041E" bold="true" box="[189,493,1287,1313]" fontSize="11" level="3" pageId="151" pageNumber="152" reason="3">
<emphasis id="A78D1B6230CD52F3EDAAFAA076E1041E" bold="true" box="[189,493,1287,1313]" pageId="151" pageNumber="152">
Subregions of
<collectingCountry id="EDEE87E030CD52F3EC78FAA076E1041E" box="[367,493,1287,1313]" name="Venezuela" pageId="151" pageNumber="152">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFA9174B4044C" blockId="151.[189,1399,1334,1395]" pageId="151" pageNumber="152">
System of hills and low piedmont mountains of the Guiana Shield, System of low mountains and hills Imataca-Cuyuní of Northeast
<collectingCountry id="EDEE87E030CD52F3EF0BFAFE757B044C" box="[540,631,1369,1395]" name="Guyana" pageId="151" pageNumber="152">Guyana</collectingCountry>
Shield, and Guiana Shield.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30CD52F3EDAAFA3D727B0716" pageId="151" pageNumber="152" type="reference_group">
<paragraph id="9546C77030CD52F3EDAAFA3D76A8048B" blockId="151.[189,420,1434,1460]" box="[189,420,1434,1460]" pageId="151" pageNumber="152">
<heading id="CE0E701C30CD52F3EDAAFA3D76A8048B" bold="true" box="[189,420,1434,1460]" fontSize="11" level="3" pageId="151" pageNumber="152" reason="3">
<emphasis id="A78D1B6230CD52F3EDAAFA3D76A8048B" bold="true" box="[189,420,1434,1460]" pageId="151" pageNumber="152">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030CD52F3EDAAFA6E727B0716" blockId="151.[189,1399,1481,1578]" pageId="151" pageNumber="152">
<bibRefCitation id="F168BA8130CD52F3EDAAFA6E76EB04DC" author="Luederwaldt H." box="[189,487,1481,1507]" pageId="151" pageNumber="152" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 387</bibRefCitation>
, 409, 417 (
<collectingCountry id="EDEE87E030CD52F3EF61FA6E75FA04DC" box="[630,758,1481,1507]" name="Venezuela" pageId="151" pageNumber="152">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130CD52F3EE21FA6E73A304DB" author="Pessoa S. B. &amp; Lane F." box="[822,1199,1481,1508]" pageId="151" pageNumber="152" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 457458</bibRefCitation>
(
<collectingCountry id="EDEE87E030CD52F3E9D5FA6E724E04DC" box="[1218,1346,1481,1507]" name="Venezuela" pageId="151" pageNumber="152">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130CD52F3EDAAFA4B75100738" author="Vulcano M. A. &amp; Pereira F. S." box="[189,540,1516,1543]" pageId="151" pageNumber="152" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 583</bibRefCitation>
(
<collectingCountry id="EDEE87E030CD52F3EF39FA4B75A20739" box="[558,686,1516,1542]" name="Venezuela" pageId="151" pageNumber="152">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130CD52F3EFFEFA4B74A70739" author="Genier F." box="[745,939,1516,1542]" pageId="151" pageNumber="152" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 74</bibRefCitation>
(
<collectingCountry id="EDEE87E030CD52F3EEABFA4B73370739" box="[956,1083,1516,1542]" name="Venezuela" pageId="151" pageNumber="152">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230CD52F3E95FFA4B73A90739" box="[1096,1189,1516,1542]" country="Venezuela" name="Bolivar" pageId="151" pageNumber="152">Bolívar</collectingRegion>
). —
<bibRefCitation id="F168BA8130CD52F3E9F6FA4A764D0716" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." pageId="151" pageNumber="152" pagination="1288 - 1298" refId="ref127038" refString="Larsen T. H., Lopera A. &amp; Forsyth A. 2008. Understanding trait-dependent community disassembly: dung beetles, density functions, and forest fragmentation. Conservation Biology 22 (5): 1288 - 1298. https: // doi. org / 10.1111 / j. 1523 - 1739.2008.00969. x" type="journal article" year="2008">
Larsen
<emphasis id="A78D1B6230CD52F3E82BFA4A727B0739" box="[1340,1399,1516,1542]" italics="true" pageId="151" pageNumber="152">et al.</emphasis>
2008: 1294
</bibRefCitation>
(
<collectingCountry id="EDEE87E030CD52F3EC58F9A876C20716" box="[335,462,1551,1577]" name="Venezuela" pageId="151" pageNumber="152">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230CD52F3ECC0F9A875380716" box="[471,564,1551,1577]" country="Venezuela" name="Bolivar" pageId="151" pageNumber="152">Bolívar</collectingRegion>
). —
<bibRefCitation id="F168BA8130CD52F3EF7DF9A874BC0716" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[618,944,1551,1577]" pageId="151" pageNumber="152" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B6230CD52F3EFEDF9B774380716" box="[762,820,1551,1577]" italics="true" pageId="151" pageNumber="152">et al.</emphasis>
2013: 109
</bibRefCitation>
(
<collectingCountry id="EDEE87E030CD52F3EEA9F9A873310716" box="[958,1085,1551,1577]" name="Venezuela" pageId="151" pageNumber="152">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230CD52F3E951F9A873AF0716" box="[1094,1187,1551,1577]" country="Venezuela" name="Bolivar" pageId="151" pageNumber="152">Bolívar</collectingRegion>
:
<collectingCountry id="EDEE87E030CD52F3E9BAF9A873DB0716" box="[1197,1239,1551,1577]" name="Iceland" pageId="151" pageNumber="152">Isla</collectingCountry>
de Anacoco).
</paragraph>
</subSubSection>
</treatment>
</document>