The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution Author Rosa, Cecilia Lozano De La 36CFEA67-71C5-4F3C-97F8-B57E43C811CC Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela. lozanoceci@gmail.com Author Cupello, Mario BDB03C18-7095-4EAA-8BDD-03CB4F79676D Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A & M University, College Station, TX, 77843, USA. mcupello@hotmail.com Author Vaz-De-Mello, Fernando Z. 2BC093C2-3E2B-466B-B31C-9F3D9FA844BC Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil. vazdemello@gmail.com text European Journal of Taxonomy 2024 2024-10-11 959 1 1 272 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401 journal article 305167 10.5852/ejt.2024.959.2677 bf132c8e-dbc7-4f3d-96e9-1a0b153e765a 2118-9773 13945464 30872C13-516F-42FA-AFA7-30ADC6BF1BAF Ontherus sulcator ( Fabricius, 1775 ) Fig. 58B Scarabaeus sulcator Fabricius, 1775: 27 (original description). Type locality: French Guiana . Name-bearing type: according to Zimsen (1964) , three syntypes in ZMUK, but deemed lost without further explanation by Génier (1996) . We ourselves have made no attempt to locate them, and so they may well be in ZMUK as Zimsen said. Copris nisus Castelnau, 1840: 79 (original description). Type locality: French Guiana . Name-bearing type: lectotype (MVMA), designated by Génier (1996) , not examined. Copris ovalipennis Blanchard, 1846: 180 (original description). Type locality: Argentina : Corrientes . Name-bearing type: lectotype (MNHN), designated by Génier (1996) , not examined. Copris senegalensis Gillet, 1910: 15 (original description). Type locality: Unknown. Originally said to be Senegal (“ Sénégal ”), but this is incorrect ( Gillet 1911a ). Name-bearing type: syntypes (RBINS) ( Gillet 1910 ), not examined. Scarabaeus sulcator Olivier 1789: 142 (redescription). — Fabricius 1792: 61 (redescription). Copris sulcator Fabricius 1801: 53 (redescription). — Olivier, 1790: 168 (redescription). — Martínez 1959: 71 (cited). Copris nisus Blanchard, 1846: 180 (comments). — Burmeister 1874: 126 (list for Argentina ). — Martínez 1959: 71 (cited as synonym of Ontherus sulcator ). Ontherus sulcator Erichson 1847b: 761 (comments). — Harold 1869c: 59 (comments); 1869d: 1008 (catalogue). — Burmeister 1874: 126 (list for Argentina ). — Gillet 1911a: 319 (list); 1911b: 58 (catalogue). — Bruch 1915: 541 (list). — Lucas 1920: 459 (catalogue). — Luederwaldt 1930: 106 (comments); 1931a: 371 , 385, 389, 392, 408, 411, 416 (key, synonymy). — Pessôa & Lane 1941: 456 , 458 (key). — Blackwelder 1944: 206 (checklist). — Martínez 1959: 71 (catalogue). — Vulcano & Pereira, 1967: 583 (key). — Forsyth & Gill 1993: 70 (as O. suclator [sic], cited for Guyana ). — Escobar 2000: 209 (list). — Vaz-de-Mello 2000: 194 (checklist for Brazil ). — Medina et al. 2001: 139 (checklist for Colombia ). — Hamel-Leigue et al. 2006: 16 (inventory, cited for Bolivia ). — Feer 2008: 62 (ecology, list). — Larsen et al. 2008: 1294 (list). — Krajcik 2012: 174 (checklist, cited for Cayenne). — Boilly & Vaz-de-Mello 2013: 112 ( Fig. 43 , key). — Ferrer-Paris et al. 2013: 109 (list). — Ratcliffe et al. 2015: 197 (checklist, cited for Peru ). Fig. 58. Species distribution. A . Ontherus pubens Génier, 1996 . B . Ontherus sulcator ( Fabricius, 1775 ) . Black circle = literature data. Copris ovalipennis Harold 1869a: 503 (comments); 1869c: 59 (comments). — Burmeister 1874: 126 (list for Argentina ). — Luederwaldt 1931a: 386 (cited as synonym of Ontherus nisus ). — Martínez 1959: 71 (cited as synonym of Ontherus sulcator ). Ontherus nisus Harold 1869d: 1008 (catalogue, cited as synonym of O. sulcator ). — Gillet 1911b: 58 (catalog, cited as synonym of O. sulcator ). — Luederwaldt 1931a: 387 , 392, 399, 400, 409, 411, 417 (redescription, distribution, key). — Pessôa & Lane 1941: 457 , 458 (key, redescription). — Blackwelder 1944: 206 (cited as synonym of O. sulcator ). — Lange 1947: 310 (list). — Martínez 1959: 71 (cited as synonym of O. sulcator ). Pinotus ovalipennis Harold 1869d: 1008 (catalogue, cited for Argentina : Corrientes ). Ontherus ovalipennis Gillet 1911b: 58 (catalog, cited as synonym of O. sulcator ). — Luederwaldt 1931a: 387 (cited as synonym of O. nisus ). — Pessôa & Lane 1941: 458 (cited as synonym of O. nisus ). — Blackwelder 1944: 206 (cited as synonym of O. sulcator ). — Martínez 1959: 71 (cited as synonym of O. sulcator ). Copris senegalensis Gillet 1911a: 319 (cited as synonym of O. sulcator ); 1911b: 58 (catalog, cited as synonym of O. sulcator ). — Bruch 1915: 541 (list, cited as synonym of O. sulcator ). — Martínez 1959: 72 (cited as synonym of O. sulcator ). Ontherus sulcator Luederwaldt 1931a: 364 ( type species). Ontherus senegalensis Luederwaldt 1931a: 386 , 387 (cited as synonym of Ontherus nisus ). — Pessôa & Lane 1941: 458 (cited as synonym of O. nisus ). — Blackwelder 1944: 206 (cited as synonym of O. sulcator ). Ontherus ( Ontherus ) sulcator Génier 1996: 74‒76 (revision). — Hielkema & Hielkema 2019: 57 (catalogue for the Guianas). Distribution Colombia , Venezuela , Trinidad and Tobago , Guyana , Suriname , French Guiana, Brazil , Ecuador , Peru , Bolivia , Paraguay , Argentina , and Uruguay ( Génier 1996 ). Subregions of Venezuela System of hills and low piedmont mountains of the Guiana Shield, System of low mountains and hills Imataca-Cuyuní of Northeast Guyana Shield, and Guiana Shield. Literature records Luederwaldt 1931a: 387 , 409, 417 ( Venezuela ). — Pessôa & Lane 1941: 457–458 ( Venezuela ). — Vulcano & Pereira 1967: 583 ( Venezuela ). — Génier 1996: 74 ( Venezuela : Bolívar ). — Larsen et al. 2008: 1294 ( Venezuela : Bolívar ). — Ferrer-Paris et al. 2013: 109 ( Venezuela : Bolívar : Isla de Anacoco).