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<document id="A0A9492484FC0C3F9F4D039C68E8AFFF" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FFEA84138CFBFF7FD809FE12" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes alboniger" docType="treatment" docVersion="7" lastPageNumber="120" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="120" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="7068C7A275C0D55F5720FF9F8D58DC12">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<mods:namePart id="FBE965E4FDC971F094CFE0997A688D66">Kluge, Nikita J.</mods:namePart>
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<treatment id="039787A6FFEA84138CFBFF7FD809FE12" ID-DOI="http://doi.org/10.5281/zenodo.3811799" ID-GBIF-Taxon="163420775" ID-Zenodo-Dep="3811799" LSID="urn:lsid:plazi:treatment:039787A6FFEA84138CFBFF7FD809FE12" httpUri="http://treatment.plazi.org/id/039787A6FFEA84138CFBFF7FD809FE12" lastPageNumber="120" pageId="119" pageNumber="120">
<subSubSection id="C324653BFFEA84138CFBFF7FDA5BFF16" box="[151,357,153,180]" pageId="119" pageNumber="120" type="nomenclature">
<paragraph id="8B8136B0FFEA84138CFBFF7FDA5BFF16" blockId="119.[151,357,153,180]" box="[151,357,153,180]" pageId="119" pageNumber="120">
<heading id="D0C981DCFFEA84138CFBFF7FDA5BFF16" bold="true" box="[151,357,153,180]" fontSize="10" level="2" pageId="119" pageNumber="120" reason="2">
<emphasis id="B94AEAA2FFEA84138CFBFF7FDA5BFF16" bold="true" box="[151,357,153,180]" pageId="119" pageNumber="120">
Group
<taxonomicName id="4C3E4D33FFEA84138C9DFF7FDA5BFF16" box="[241,357,153,180]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="alboniger">
<emphasis id="B94AEAA2FFEA84138C9DFF7FDA5BFF16" bold="true" box="[241,357,153,180]" italics="true" pageId="119" pageNumber="120">alboniger</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFEA84138CFBFF07D89EFEE6" pageId="119" pageNumber="120" type="diagnosis">
<paragraph id="8B8136B0FFEA84138CFBFF07D89EFEE6" blockId="119.[151,1437,225,432]" pageId="119" pageNumber="120">
<emphasis id="B94AEAA2FFEA84138CFBFF07DA28FF5E" bold="true" box="[151,278,225,252]" pageId="119" pageNumber="120">Definition.</emphasis>
Larva of the single species placed here, differs from all other
<taxonomicName id="4C3E4D33FFEA84138FB5FF07DF67FF59" authorityName="Ulmer" authorityYear="1920" box="[985,1113,225,251]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAA2FFEA84138FB5FF07DF67FF59" box="[985,1113,225,251]" italics="true" pageId="119" pageNumber="120">Thraulodes</emphasis>
</taxonomicName>
by wider anterior transverse setal row on labrum (
<figureCitation id="13052A35FFEA84138DEEFEE3DADEFE82" box="[386,480,261,288]" captionStart="FIGURES 563568" captionStartId="122.[151,264,1809,1834]" captionTargetBox="[151,1436,198,1782]" captionTargetId="figure@122.[151,1436,198,1782]" captionTargetPageId="122" captionText="FIGURES 563568. Thraulodes alboniger sp. n. 563, labrum; 564565 and 566567, male imaginal genitalia of two different specimens; 568, fragment of imaginal caudalius (563, 566568, holotype).Abbreviations: anter.r., anterior transverse setal row; poster.r., posterior transverse setal row; other arrow show six sensilla separating five denticles." figureDoi="http://doi.org/10.5281/zenodo.3737046" httpUri="https://zenodo.org/record/3737046/files/figure.png" pageId="119" pageNumber="120">Fig. 563</figureCitation>
) and opposite bent of tergalii (
<figureCitation id="13052A35FFEA84138F5EFEE3D8EFFE82" box="[818,977,261,288]" captionStart="FIGURES 578588" captionStartId="124.[151,264,1896,1921]" captionTargetBox="[151,1429,258,1747]" captionTargetId="figure@124.[151,1436,249,1757]" captionTargetPageId="124" captionText="FIGURES 578588. Thraulodes alboniger sp. n. 578, hind wing; 579, genitalia of male imago in ventral view; 580, dorsal margin of styliger and penis of another specimen; 581, subimaginal exuviae of genitalia; 582, penis in lateral view; 583, posterior margin of IX sternum of female larva; 584, the same of male larva with protogonostyli and protopenis; 585588, tergalii I, III, VI and VII (578, 580581, 584588, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737050" httpUri="https://zenodo.org/record/3737050/files/figure.png" pageId="119" pageNumber="120">Figs 585588</figureCitation>
). Imago differs from all other
<taxonomicName id="4C3E4D33FFEA84138971FEE3DEA3FEBD" authorityName="Ulmer" authorityYear="1920" box="[1309,1437,261,287]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAA2FFEA84138971FEE3DEA3FEBD" box="[1309,1437,261,287]" italics="true" pageId="119" pageNumber="120">Thraulodes</emphasis>
</taxonomicName>
by very compact black preapical band on hind femur (
<figureCitation id="13052A35FFEA84138E99FECFD8AAFEE6" box="[757,916,297,324]" captionStart="FIGURES 569577" captionStartId="123.[151,264,1886,1911]" captionTargetBox="[151,1436,214,1850]" captionTargetId="figure@123.[151,1436,214,1850]" captionTargetPageId="123" captionText="FIGURES 569577. Thraulodes alboniger sp. n. 569, female subimago extracted from larva; 570, apex of fore wing; 571573, abdominal terga of three different male imagines; 574575, two male imagines in lateral view; 576, head and thorax of male imago in dorsal view; 577, right half of subimaginal exuviae of mesonotum (570, 572, 577, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737048" httpUri="https://zenodo.org/record/3737048/files/figure.png" pageId="119" pageNumber="120">Figs 574, 575</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFEA84138CABFEABD809FE12" pageId="119" pageNumber="120" type="discussion">
<paragraph id="8B8136B0FFEA84138CABFEABD809FE12" blockId="119.[151,1437,225,432]" pageId="119" pageNumber="120">
<emphasis id="B94AEAA2FFEA84138CABFEABDA59FECA" bold="true" box="[199,359,333,360]" pageId="119" pageNumber="120">Composition.</emphasis>
One species,
<taxonomicName id="4C3E4D33FFEA84138E6DFEABD9A4FEC5" box="[513,666,333,359]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="alboniger" status="sp. nov.">
<emphasis id="B94AEAA2FFEA84138E6DFEABD9A4FEC5" box="[513,666,333,359]" italics="true" pageId="119" pageNumber="120">Th. alboniger</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFEA84138ECCFEA8D9DEFECA" bold="true" box="[672,736,334,360]" pageId="119" pageNumber="120">
<taxonomicNameLabel id="A27957D9FFEA84138ECCFEA8D9DEFECA" box="[672,736,334,360]" pageId="119" pageNumber="120" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is described here. Variability of imaginal coloration (
<figureCitation id="13052A35FFEA8413895FFEABDBFCFE2E" captionStart="FIGURES 569577" captionStartId="123.[151,264,1886,1911]" captionTargetBox="[151,1436,214,1850]" captionTargetId="figure@123.[151,1436,214,1850]" captionTargetPageId="123" captionText="FIGURES 569577. Thraulodes alboniger sp. n. 569, female subimago extracted from larva; 570, apex of fore wing; 571573, abdominal terga of three different male imagines; 574575, two male imagines in lateral view; 576, head and thorax of male imago in dorsal view; 577, right half of subimaginal exuviae of mesonotum (570, 572, 577, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737048" httpUri="https://zenodo.org/record/3737048/files/figure.png" pageId="119" pageNumber="120">Figs 574, 575</figureCitation>
) allows to assume that the
<typeStatus id="54858812FFEA84138D95FE94D917FE2E" box="[505,553,370,396]" pageId="119" pageNumber="120">type</typeStatus>
series consists of more than one species. To verify this, it is necessary to rear from larvae imagines of the light form (
<figureCitation id="13052A35FFEA84138E3FFE73D814FE12" box="[595,810,405,432]" captionStart="FIGURES 569577" captionStartId="123.[151,264,1886,1911]" captionTargetBox="[151,1436,214,1850]" captionTargetId="figure@123.[151,1436,214,1850]" captionTargetPageId="123" captionText="FIGURES 569577. Thraulodes alboniger sp. n. 569, female subimago extracted from larva; 570, apex of fore wing; 571573, abdominal terga of three different male imagines; 574575, two male imagines in lateral view; 576, head and thorax of male imago in dorsal view; 577, right half of subimaginal exuviae of mesonotum (570, 572, 577, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737048" httpUri="https://zenodo.org/record/3737048/files/figure.png" pageId="119" pageNumber="120">Figs 571, 573, 574</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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