519 lines
38 KiB
XML
519 lines
38 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.884.30988" ID-GBIF-Dataset="a4b13748-99a9-41ed-8964-29d76df19769" ID-PMC="PMC6834763" ID-Pensoft-Pub="1313-2970-884-23" ID-Pensoft-UUID="6251848C807E5E0896A9C39CC22ECD19" ID-PubMed="31723324" ID-ZooBank="F1B4B6A8288C4CE9AFEF7DE08DBE9CC0" ModsDocID="1313-2970-884-23" checkinTime="1572509043610" checkinUser="pensoft" docAuthor="Dong, Yan, Law, Michelle Man Suet, Jiang, Jibao B. & Qiu, Jiangping P." docDate="2019" docId="6A0577B9A1BE5C3EADE92FB3D3446525" docLanguage="en" docName="ZooKeys 884: 23-42" docOrigin="ZooKeys 884" docSource="http://dx.doi.org/10.3897/zookeys.884.30988" docTitle="Amynthas maximus Qiu & Dong, sp. nov." docType="treatment" docUuid="E84CFBE5-4FF8-4F53-B49A-233EC5D04298" docUuidSource="ZooBank" docVersion="7" id="6251848C807E5E0896A9C39CC22ECD19" lastPageNumber="23" masterDocId="6251848C807E5E0896A9C39CC22ECD19" masterDocTitle="Three new species and one subspecies of the Amynthas corticis - group from Guangxi Zhuang Autonomous Region, China (Oligochaeta, Megascolecidae)" masterLastPageNumber="42" masterPageNumber="23" pageNumber="23" updateTime="1668167990170" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Three new species and one subspecies of the Amynthas corticis - group from Guangxi Zhuang Autonomous Region, China (Oligochaeta, Megascolecidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Dong, Yan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Law, Michelle Man Suet</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Jiang, Jibao B.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Qiu, Jiangping P.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2019</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>884</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>23</mods:start>
|
||
<mods:end>42</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.884.30988</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.884.30988</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-884-23</mods:identifier>
|
||
<mods:identifier type="ZooBank">F1B4B6A8288C4CE9AFEF7DE08DBE9CC0</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">6251848C807E5E0896A9C39CC22ECD19</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="159932978" LSID="urn:lsid:zoobank.org:act:E84CFBE5-4FF8-4F53-B49A-233EC5D04298" httpUri="http://treatment.plazi.org/id/6A0577B9A1BE5C3EADE92FB3D3446525" lastPageNumber="23" pageId="0" pageNumber="23">
|
||
<subSubSection pageId="0" pageNumber="23" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="23">
|
||
<taxonomicName LSID="6A0577B9-A1BE-5C3E-ADE9-2FB3D3446525" authority="Qiu & Dong" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas maximus" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="maximus">Amynthas maximus Qiu & Dong</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="23">sp. nov.</taxonomicNameLabel>
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A Ventral view showing spermathecal pores, female pores and male pores of Amynthas maximus sp. nov. B spermathecae of Amynthas maximus sp. nov. C illustration of the details of the male pore region." figureDoi="10.3897/zookeys.884.30988.figure2" httpUri="https://binary.pensoft.net/fig/351025" pageId="0" pageNumber="23" tableDoi="10.3897/zookeys.884.30988.figure2">Figure 2</figureCitation>
|
||
<tableCitation captionStartId="T2" httpUri="http://table.plazi.org/id/A93DF62E0DCD243A376D501E1E680C61" pageId="0" pageNumber="23" tableUuid="A93DF62E0DCD243A376D501E1E680C61">, Table 2</tableCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="23" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="23">Material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="23">Holotype</emphasis>
|
||
:1 clitellate (C-GX201304-01A): China, Guangxi Zhuang Autonomous Region, Shiwan Mountain Nature Reserve (
|
||
<geoCoordinate degrees="21.50299" direction="north" orientation="latitude" precision="1" value="21.50299">21.50299°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="107.3035" direction="east" orientation="longitude" precision="5" value="107.3035">107.3035°E</geoCoordinate>
|
||
), 449 m asl, black sandy soil under bryophytes in a subtropical evergreen forest, 12 May 2013, JP Qiu, Y Hong, JB Jiang, LL Zhang, Y Dong legit.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="23">Paratypes</emphasis>
|
||
: 8 clitellates (C-GX201304-01B): same date as for holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="23" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="23">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
Dimensions 145-170 mm by 5.8-6.2 mm at clitellum, clitellum taupe in 2/5
|
||
<normalizedToken originalValue="XIV–XVI">XIV-XVI</normalizedToken>
|
||
, 78-101 segments. First dorsal pore in 13/14. Setae numbering 33-38 at III, 32-36 at V, 29-33 at VIII, 18-22 at XX, 50-65 at XXV; 9-13 between male pores; setae between spermathecal pores numbering 14-16 at VI, 10-14 at VII, and 18-22 at VIII. Four pairs of spermathecal pores ventrally in 5/6-8/9. Seven or eight (three specimens have seven papillae, and four specimens have eight papillae) postsetal genital papillae arranged in two rows in
|
||
<normalizedToken originalValue="VI–IX">VI-IX</normalizedToken>
|
||
, 0.33 circumference ventrally apart from each other. One pair of male pores in XVIII, each on the top of a central, round porophore surrounded by three or four circular ridges, with one presetal indented-topped genital papilla medial of each porophore. Ampulla elongate oval, stout duct as long as 3/5 ampulla. Diverticulum slightly shorter than main pouch, a little twist in the middle, terminal 2/5 dilated into a swollen, club-shaped seminal chamber. Prostate glands well developed.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.884.30988.figure2" httpUri="https://binary.pensoft.net/fig/351025" pageId="0" pageNumber="23" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="23">
|
||
<emphasis bold="true" pageId="0" pageNumber="23">Figure 2.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="23">A</emphasis>
|
||
Ventral view showing spermathecal pores, female pores and male pores of
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas maximus" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Amynthas maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="23">B</emphasis>
|
||
spermathecae of
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas maximus" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Amynthas maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="23">C</emphasis>
|
||
illustration of the details of the male pore region.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-Table-UUID="A93DF62E0DCD243A376D501E1E680C61" httpUri="http://table.plazi.org/id/A93DF62E0DCD243A376D501E1E680C61" pageId="0" pageNumber="23" start="Table 2" startId="T2">
|
||
<paragraph pageId="0" pageNumber="23">
|
||
<emphasis bold="true" pageId="0" pageNumber="23">Table 2.</emphasis>
|
||
A comparison of characters of
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName lsidName="A. dorsualis" pageId="0" pageNumber="23" rank="species" species="dorsualis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. dorsualis</emphasis>
|
||
</taxonomicName>
|
||
, 2013,
|
||
<taxonomicName lsidName="A. carnosus" pageId="0" pageNumber="23" rank="species" species="carnosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. carnosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="A. corticis" pageId="0" pageNumber="23" rank="species" species="corticis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. corticis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. wulinensis" pageId="0" pageNumber="23" rank="species" species="wulinensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. wulinensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
<table pageId="0" pageNumber="23">
|
||
<tr pageId="0" pageNumber="23">
|
||
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">Characteristics</th>
|
||
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<taxonomicName lsidName="A. dorsualis" pageId="0" pageNumber="23" rank="species" species="dorsualis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. dorsualis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<taxonomicName lsidName="A. carnosus" pageId="0" pageNumber="23" rank="species" species="carnosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. carnosus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<taxonomicName lsidName="A. corticis" pageId="0" pageNumber="23" rank="species" species="corticis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. corticis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<taxonomicName lsidName="A. wulinensis" pageId="0" pageNumber="23" rank="species" species="wulinensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. wulinensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Body length (mm)</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">145-170</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">121-?</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">110-340</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">45-170</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">128-174</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Body width (mm)</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">5.8-6.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">2.7-?</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">4.0-9.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">3.0-6.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">5.6-6.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Pigment</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">dorsum</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Light purple brown before clitellum, from light purple brown to brown after</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Dark grey before clitellum, dark brown after</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Dark brown or purple</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Greenish brown</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Whitish purple</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">ventrum</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">No pigment before clitellum, yellowish after</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Light grey before VII, no pigment after</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Dark brown or purple</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">No</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Whitish gray</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">First dorsal pore</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">13/14</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">13/14</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">12/13</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">10/11 or 12/13, usually at 11/12</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">11/12</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Clitellum locality</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
2/5
|
||
<normalizedToken originalValue="XIV–XVI">XIV-XVI</normalizedToken>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1/10XIV-7/10XVI</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<normalizedToken originalValue="XIV–XVI">XIV-XVI</normalizedToken>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<normalizedToken originalValue="XIV–XVI">XIV-XVI</normalizedToken>
|
||
, occasionally shorter
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<normalizedToken originalValue="XIV–XVI">XIV-XVI</normalizedToken>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Spermathecal pores</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">4 pairs, in 5/6-8/9, 0.33C</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">4 pairs, in 5/6-8/9, dorsally, 0.6C</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">4 pairs in 5/6-8/9 or 3 pairs in 6/7-8/9, 0.33C</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">4 pairs, 5/6-8/9, 0.33C</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">4 pairs, 5/6-8/9, ventral, 0.29C</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Male pores</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Middle, round, surrounded by 3-4 circular ridges, 0.4C</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Slightly raised, glandular, surrounded by 5-6 elliptic circular folds, 0.33C</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Round or elliptic</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Small, circular to transverse elliptical disc, 0.24-0.30C</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Round or oval-shaped on setal line with depressed center, 2-3 circular folds, 0.24-0.28C</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Papillae preclitellum</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
7-8 post-setal indented-topped genital papillae arranged in two rows in
|
||
<normalizedToken originalValue="VI–IX">VI-IX</normalizedToken>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Invisible</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">8 papillae just overhead 8 spermathecal pores, 2 pairs preclitellar arranged on VIII and IX</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Paired presetal and/or postsetal in some or all, near spermathecal pores</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Absent</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Papillae postclitellum</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">2 presetal indented-topped genital papillae medial of porophore</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Invisible</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">2 paired presetal genital on XVIII and IX, 1 pair postsetal genital on XVIII</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Present or absent, occasionally one or more near male pore</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Oval-shaped, medial to male pore in each of XVII and XIX, occasionally XX</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Prostate glands</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<normalizedToken originalValue="XVII–XIX">XVII-XIX</normalizedToken>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<normalizedToken originalValue="XVI–XX">XVI-XX</normalizedToken>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Well developed</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<normalizedToken originalValue="XVII–XX">XVII-XX</normalizedToken>
|
||
, rudimentary or absent
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">
|
||
<normalizedToken originalValue="XV–XX">XV-XX</normalizedToken>
|
||
, racemose, follicular
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Spermathecae</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">About 1.6 mm long, ampulla long-oval; duct as long as 3/5 ampulla</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">About 2.2 mm long, ampulla heart-shaped; duct as long as 2/5 ampulla</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Ampulla oval or pear-shaped, duct equal to or slightly shorter than ampulla</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Ampulla ovoid</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Very short and stout stalk</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Diverticulum</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Shorter, lightly twist in middle, terminal 2/5, swollen, club-shaped seminal chamber</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Shorter, terminal 1/5, ovoid plump seminal chamber</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">One-third to half of ampulla, slender stalk and a wider seminal chamber</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Blunt ovioid, straight stalk</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Oval, shining white seminal chamber, a slender and straight stalk</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="23">
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Accessory glands</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">1 or 2 stalked accessory glands observed near ventral median line in VI, VII, VIII, IX</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Invisible</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Stalked, coelomic, bound down to parietes or retained within body wall</td>
|
||
<td colspan="1" pageId="0" pageNumber="23" rowspan="1">Paired in XVII and XIX, sessile, flowery</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="23" type="description">
|
||
<paragraph pageId="0" pageNumber="23">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="23">External characters</emphasis>
|
||
: Light purple-brown pigment on pre-clitellum dorsum, no pigment on ventrum. Pigment from light purple-brown to brown on post-clitellum dorsum, light yellowish on ventrum. Clitellum taupe in 2/5
|
||
<normalizedToken originalValue="XIV–XVI">XIV-XVI</normalizedToken>
|
||
. Dimensions 160 mm by 6.0 mm at clitellum, 92 segments. Prostomium
|
||
<normalizedToken originalValue="½">1/2</normalizedToken>
|
||
epilobous. First dorsal pore in 13/14. Setae numbering 36 at III, 34 at V, 31 at VIII, 20 at XX, 60 at XXV; 12 between male pores; Setae numbering 15 at VI, 12 at VII, 20 at VIII between spermathecal pores. Setae formula: AA = 1.1-1.4AB, ZZ = 1.2-2.0ZY. Clitellum annular, yellowish, in 2/5
|
||
<normalizedToken originalValue="XIV–XVI">XIV-XVI</normalizedToken>
|
||
, setae not visible externally. Four pairs of spermathecal pores in 5/6-8/9, ventral, eye-like, 0.4 circumference ventrally apart from each other. Seven or eight (three specimens have seven papillae, and four specimens have eight papillae) postsetal genital papillae arranged in two rows in
|
||
<normalizedToken originalValue="VI–X">VI-X</normalizedToken>
|
||
, 0.33 circumference ventrally apart from each other. One pair of male pores in XVIII, 0.4 circumference apart ventrally, each on the top of a central, round porophore surrounded by three or four circular ridges, with one presetal indented-topped genital papilla medial of each porophore (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of known species in Guangxi Zhuang Autonomous Region and sampling points of this paper." figureDoi="10.3897/zookeys.884.30988.figure1" httpUri="https://binary.pensoft.net/fig/351024" pageId="0" pageNumber="23" tableDoi="10.3897/zookeys.884.30988.figure1">Figure 1A</figureCitation>
|
||
). Single female pore in XIV, ovoid.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
<emphasis bold="true" pageId="0" pageNumber="23">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Internal characters</emphasis>
|
||
.
|
||
</emphasis>
|
||
Septa 5/6-7/8, 10/11-13/14 thick and muscular, 8/9-9/10 absent. Gizzard bucket-shaped, in
|
||
<normalizedToken originalValue="VIII–X">VIII-X</normalizedToken>
|
||
. Intestine enlarged distinctly from XV onwards. Intestinal caeca paired in XXVII, extending anteriorly to XXII, transition state, ventral margin smooth, four pointed saccules in dorsal margin. Four pairs of esophageal hearts in
|
||
<normalizedToken originalValue="X–XIII">X-XIII</normalizedToken>
|
||
, developed. Ovaries in XIII. Four pairs of spermathecae in
|
||
<normalizedToken originalValue="VI–IX">VI-IX</normalizedToken>
|
||
, short, approx. 1.6 mm long, ampulla elongate-oval; duct as long as 3/5 ampulla. Diverticulum slightly shorter than main pouch (ampulla and duct), a little twist in middle, terminal 2/5 dilated into a swollen, club-shaped seminal chamber (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of known species in Guangxi Zhuang Autonomous Region and sampling points of this paper." figureDoi="10.3897/zookeys.884.30988.figure1" httpUri="https://binary.pensoft.net/fig/351024" pageId="0" pageNumber="23" tableDoi="10.3897/zookeys.884.30988.figure1">Figure 1B</figureCitation>
|
||
). One or two stalked accessory glands observed near ventral median line in
|
||
<normalizedToken originalValue="VI–IX">VI-IX</normalizedToken>
|
||
. Holandric: two pairs of testis sacs in X and XI, separated from each other, well developed. Two pairs of seminal vesicles in XI and XII, developed. Prostate glands undeveloped, inserting in XVIII and extending from
|
||
<normalizedToken originalValue="XVII–XIX">XVII-XIX</normalizedToken>
|
||
, coarsely lobate, prostatic duct I-shaped, of uniform thickness. No accessory glands observed in male pore region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="23" type="etymology">
|
||
<paragraph pageId="0" pageNumber="23">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">The species is named after its large accessory glands observed in the spermathecal area.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="23" type="remarks">
|
||
<paragraph pageId="0" pageNumber="23">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas maximus" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Amynthas maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. keys to the
|
||
<taxonomicName lsidName="corticis" pageId="0" pageNumber="23" rank="species" species="corticis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">corticis</emphasis>
|
||
</taxonomicName>
|
||
-group in
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1095-8312.1972.tb00694.x" author="Sims, R" journalOrPublisher="Biological Journal of the Linnean Society" pageId="0" pageNumber="23" pagination="169 - 268" refId="B21" refString="Sims, R, Easton, E, 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society 4: 169 - 268" title="A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition." url="https://doi.org/10.1111/j.1095-8312.1972.tb00694.x" volume="4" year="1972">Sims and Easton (1972)</bibRefCitation>
|
||
with four pairs of spermathecal pores intersegmentally in 5/6-8/9.
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas maximus" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Amynthas maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas carnosus" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="carnosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Amynthas carnosus</emphasis>
|
||
</taxonomicName>
|
||
(Goto & Hatai, 1899) as re-described by
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4179.3.7" author="Chang, CH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="23" pagination="495 - 529" refId="B4" refString="Chang, CH, Snyder, BA, Szlavecz, K, 2016. Asian pheretimoid earthworms in North America north of Mexico: an illustrated key to the genera Amynthas, Metaphire, Pithemera, and Polypheretima (Clitellata: Megascolecidae). Zootaxa 4179: 495 - 529" title="Asian pheretimoid earthworms in North America north of Mexico: an illustrated key to the genera Amynthas, Metaphire, Pithemera, and Polypheretima (Clitellata: Megascolecidae)." url="https://doi.org/10.11646/zootaxa.4179.3.7" volume="4179" year="2016">Chang et al. (2016)</bibRefCitation>
|
||
with respect to body size, the distance between spermathecal pores and male pores, shorter diverticulum than main spermathecal axis, and no accessary glands near prostates. In contrast, the pigment on its ventrum is lighter than
|
||
<taxonomicName lsidName="A. carnosus" pageId="0" pageNumber="23" rank="species" species="carnosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. carnosus</emphasis>
|
||
</taxonomicName>
|
||
and other differences include the first dorsal pore, clitellum location, spermathecal pores, and male pores characters. The first dorsal pore in
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is located in 13/14, versus 12/13 in
|
||
<taxonomicName lsidName="A. carnosus" pageId="0" pageNumber="23" rank="species" species="carnosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. carnosus</emphasis>
|
||
</taxonomicName>
|
||
; the clitellum occupies less than three segments; four pairs of spermathecal pores while sometimes three pairs in
|
||
<taxonomicName lsidName="A. carnosus" pageId="0" pageNumber="23" rank="species" species="carnosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. carnosus</emphasis>
|
||
</taxonomicName>
|
||
; the porophore is surrounded by three or four circular ridges, but no ridges are present in
|
||
<taxonomicName lsidName="A. carnosus" pageId="0" pageNumber="23" rank="species" species="carnosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. carnosus</emphasis>
|
||
</taxonomicName>
|
||
; several accessory glands observed in the spermathecal region in the new species but none in
|
||
<taxonomicName lsidName="A. carnosus" pageId="0" pageNumber="23" rank="species" species="carnosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. carnosus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
We also compare the new species with
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas corticis" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="corticis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Amynthas corticis</emphasis>
|
||
</taxonomicName>
|
||
(Kinberg, 1867) which has been recognized as the typical species in the
|
||
<taxonomicName lsidName="corticis" pageId="0" pageNumber="23" rank="species" species="corticis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">corticis</emphasis>
|
||
</taxonomicName>
|
||
-group. They share several common characters such as body size, pigment, clitellum extent, setal number, and both have stalked accessory glands. Other than that, the first dorsal pore in the new species is in 13/14, but in 10/11 or 12/13, usually in 11/12 in
|
||
<taxonomicName lsidName="A. corticis" pageId="0" pageNumber="23" rank="species" species="corticis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. corticis</emphasis>
|
||
</taxonomicName>
|
||
. The diverticulum of
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. has a small twist in the middle compared with
|
||
<taxonomicName lsidName="A. corticis" pageId="0" pageNumber="23" rank="species" species="corticis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. corticis</emphasis>
|
||
</taxonomicName>
|
||
, which has a long stalk. Moreover,
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. always exhibits genital markings in the male pore region, whereas in
|
||
<taxonomicName lsidName="A. corticis" pageId="0" pageNumber="23" rank="species" species="corticis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. corticis</emphasis>
|
||
</taxonomicName>
|
||
, these markings are occasionally absent.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
We further compare the new species with another species
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas dorsualis" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="dorsualis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Amynthas dorsualis</emphasis>
|
||
</taxonomicName>
|
||
Sun & Qiu, 2013 described from Hainan, China, and its clitellum also occupies fewer than three segments. In our results, we find the two species share several common characters, including the first dorsal pore location, setal formula, male pore characters, and in the diverticulum being shorter than the main pouch. However, the morphological dissimilarity of the two species is substantial. For instance, the locations of spermathecal pores are different between
|
||
<taxonomicName lsidName="A. dorsualis" pageId="0" pageNumber="23" rank="species" species="dorsualis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. dorsualis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. In
|
||
<taxonomicName lsidName="A. dorsualis" pageId="0" pageNumber="23" rank="species" species="dorsualis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. dorsualis</emphasis>
|
||
</taxonomicName>
|
||
, the spermathecal pores are located on the dorsum, while the pores are located on the ventrum in
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. The distance between male pore is shorter on the ventral side in
|
||
<taxonomicName lsidName="A. dorsualis" pageId="0" pageNumber="23" rank="species" species="dorsualis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. dorsualis</emphasis>
|
||
</taxonomicName>
|
||
than
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas maximus" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Amynthas maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. In addition,
|
||
<taxonomicName lsidName="A. dorsualis" pageId="0" pageNumber="23" rank="species" species="dorsualis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. dorsualis</emphasis>
|
||
</taxonomicName>
|
||
has no genital markings and no accessory glands, whereas
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. exhibits genital markings near the spermathecal pores and the male pores region, and stalked accessory glands are present in spermathecal pores region.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="23">
|
||
The body size of
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName lsidName="A. wulinensis" pageId="0" pageNumber="23" rank="species" species="wulinensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. wulinensis</emphasis>
|
||
</taxonomicName>
|
||
described from Taiwan Island. But the other characters of
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differ from
|
||
<taxonomicName lsidName="A. wulinensis" pageId="0" pageNumber="23" rank="species" species="wulinensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. wulinensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas maximus" order="Opisthopora" pageId="0" pageNumber="23" phylum="Annelida" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">Amynthas maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. has no pigment before clitellum, yellowish after clitellum on dorsum, first dorsal pore in 13/14, 0.33C between spermathecal pores, seven or eight indented-topped genital papillae in
|
||
<normalizedToken originalValue="VI–IX">VI-IX</normalizedToken>
|
||
and accessory glands in spermathecal pores region. In contrast,
|
||
<taxonomicName lsidName="A. wulinensis" pageId="0" pageNumber="23" rank="species" species="wulinensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. wulinensis</emphasis>
|
||
</taxonomicName>
|
||
has whitish gray on dorsum, first dorsal pore in 11/12, 0.29C between spermathecal pores, no genital papillae observed in spermathecal pores region, and accessory glands observed in male pores region. Table
|
||
<tableCitation captionStartId="T2" httpUri="http://table.plazi.org/id/A93DF62E0DCD243A376D501E1E680C61" pageId="0" pageNumber="23" tableUuid="A93DF62E0DCD243A376D501E1E680C61">2</tableCitation>
|
||
shows the comparison of characters of
|
||
<taxonomicName lsidName="A. maximus" pageId="0" pageNumber="23" rank="species" species="maximus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. maximus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. with
|
||
<taxonomicName lsidName="A. dorsualis" pageId="0" pageNumber="23" rank="species" species="dorsualis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. dorsualis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="A. carnosus" pageId="0" pageNumber="23" rank="species" species="carnosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. carnosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="A. corticis" pageId="0" pageNumber="23" rank="species" species="corticis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. corticis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. wulinensis" pageId="0" pageNumber="23" rank="species" species="wulinensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="23">A. wulinensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |