344 lines
40 KiB
XML
344 lines
40 KiB
XML
<document id="649C87D4E3F81897EE2F50895E29575D" ID-CLB-Dataset="46398" ID-DOI="10.5281/zenodo.201335" ID-GBIF-Dataset="13326814-0c76-4aff-b6f0-b5c2c057b15c" ID-ISSN="1175-5326" ID-Zenodo-Dep="201335" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460386445923" checkinUser="plazi" docAuthor="Yin, Zi-Wei, Li, Li-Zhen & Zhao, Mei-Jun" docDate="2011" docId="D34D87F5B42BFFBDFF3E16C4FD40F9FC" docLanguage="en" docName="zt03016p050.pdf" docOrigin="Zootaxa 3016" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Batriscenellus Jeannel 1958" docType="treatment" docVersion="10" lastPageNumber="39" masterDocId="2F74FF8DB42AFFBFFFA9143DFF44FF88" masterDocTitle="Batriscenellus Jeannel (Coleoptera, Staphylinidae, Pselaphinae) redefined, with notes on the Chinese species" masterLastPageNumber="50" masterPageNumber="37" pageNumber="38" updateTime="1698297671878" updateUser="ExternalLinkService">
|
||
<mods:mods id="A3A4DD120A2514F98A0F5F8A97E991AE" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="075AFB8D4DFA906EDC796E0C470C00C5">
|
||
<mods:title id="B329D00C7BEB53E1C5A2FFF839F65026">Batriscenellus Jeannel (Coleoptera, Staphylinidae, Pselaphinae) redefined, with notes on the Chinese species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="DCF14511E10CDA9CF0C96F341847E014" type="personal">
|
||
<mods:role id="B9E932F2B9A34AD3FC783AF119AFABB8">
|
||
<mods:roleTerm id="BB2B4C07D481F8AEC39A9DC82BACECF6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="897BE58B353F43CA5692826F890829FB">Yin, Zi-Wei</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="C9313E01DAD2F94720D872F2950F16C3" type="personal">
|
||
<mods:role id="30516A285320C6C80CCAEA9543D2FB19">
|
||
<mods:roleTerm id="E23761D26F73B50034B586EB78031BD8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="556EBE38410812C3D54CF0B6862AFC60">Li, Li-Zhen</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="41819B968015B39FC306F9A17C472B97" type="personal">
|
||
<mods:role id="33F7B81BA2F5E776991F57E6E422B5E9">
|
||
<mods:roleTerm id="74B85A9236671AB836359D6D75E14407">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D38FC33C7AFEA110DFBF840897986475">Zhao, Mei-Jun</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D194E9C9E743C6B892CB6CBF95F87C1B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="5A92136AF115E6CD9DD3FC65394B6062" type="host">
|
||
<mods:titleInfo id="EBDE5C5D6C4FEF479C8B13DDB95CAB9D">
|
||
<mods:title id="9716A87DDFC76D7353F2FEC91A1372C3">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="C8C18EB06D1AF81ABA219798A9BE21B9">
|
||
<mods:date id="1BD0BB394AF18EADAA7D4C19B163E9A8">2011</mods:date>
|
||
<mods:detail id="CC70F54D5CA24F216468621166A25B33" type="volume">
|
||
<mods:number id="8DBC6C75255EB652F789977DD5E97C7A">3016</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="9F2332903054791E2E403E7ACB08E93B" unit="page">
|
||
<mods:start id="FFACCF890A68E68A5BE840FEB5745189">37</mods:start>
|
||
<mods:end id="8664AC141582AD9EB1FA0CDC24BDF9E6">50</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="7897A8FFD07FD3D9643744B6C101B5B5">journal article</mods:classification>
|
||
<mods:identifier id="EFB098EAAA49A355F15A36E6926B334F" type="CLB-Dataset">46398</mods:identifier>
|
||
<mods:identifier id="01CD7A8519AF5A0E5B0E87790204573E" type="DOI">10.5281/zenodo.201335</mods:identifier>
|
||
<mods:identifier id="2974418B7EEC6DDEF5252EBB3D5CD257" type="GBIF-Dataset">13326814-0c76-4aff-b6f0-b5c2c057b15c</mods:identifier>
|
||
<mods:identifier id="D9F4B52A47114B18B93BEA088DE214BC" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="87526A94E4A38DE3B95280C5C3E83E0A" type="Zenodo-Dep">201335</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="D34D87F5B42BFFBDFF3E16C4FD40F9FC" ID-DOI="http://doi.org/10.5281/zenodo.6184817" ID-GBIF-Taxon="119415589" ID-Zenodo-Dep="6184817" LSID="urn:lsid:plazi:treatment:D34D87F5B42BFFBDFF3E16C4FD40F9FC" httpUri="http://treatment.plazi.org/id/D34D87F5B42BFFBDFF3E16C4FD40F9FC" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
|
||
<subSubSection id="13FE6568B42BFFBEFF3E16C4FEA0FC9A" box="[151,484,761,786]" pageId="1" pageNumber="38" type="nomenclature">
|
||
<paragraph id="5B5B36E3B42BFFBEFF3E16C4FEA0FC9A" blockId="1.[151,484,761,786]" box="[151,484,761,786]" pageId="1" pageNumber="38">
|
||
<heading id="0013818FB42BFFBEFF3E16C4FEA0FC9A" bold="true" box="[151,484,761,786]" fontSize="10" level="2" pageId="1" pageNumber="38" reason="2">
|
||
<taxonomicName id="9CE44D60B42BFFBEFF3E16C4FEA0FC9A" ID-CoL="8X758" authority="Jeannel, 1958" authorityName="Jeannel" authorityYear="1958" box="[151,484,761,786]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B42BFFBEFF3E16C4FEA0FC9A" bold="true" box="[151,484,761,786]" pageId="1" pageNumber="38">
|
||
<emphasis id="6990EAF1B42BFFBEFF3E16C4FE78FC9A" bold="true" box="[151,316,761,786]" italics="true" pageId="1" pageNumber="38">Batriscenellus</emphasis>
|
||
<bibRefCitation id="3F754B12B42BFFBEFEEB16C7FEA0FC9A" author="Jeannel" box="[322,484,761,786]" pageId="1" pageNumber="38" refString="Jeannel, R. (1958) Revision des Pselaphides du Japon. Memoires du Museum National d'Histoire Naturelle, (N. S., Serie A, Zoologie), 18, 1 - 138." type="journal article" year="1958">Jeannel, 1958</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13FE6568B42BFFBEFF3E177FFCDCFCD1" box="[151,920,833,858]" pageId="1" pageNumber="38" type="materials_examined">
|
||
<paragraph id="5B5B36E3B42BFFBEFF3E177FFCDCFCD1" blockId="1.[151,1437,833,2009]" box="[151,920,833,858]" pageId="1" pageNumber="38">
|
||
<emphasis id="6990EAF1B42BFFBEFF3E177FFE76FCD2" bold="true" box="[151,306,833,858]" pageId="1" pageNumber="38">
|
||
<typeStatus id="845F8841B42BFFBEFF3E177FFF96FCD2" box="[151,210,834,858]" pageId="1" pageNumber="38">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="9CE44D60B42BFFBEFE90177FFD39FCD1" authority="Sharp, 1883" authorityName="Sharp" authorityYear="1883" box="[313,637,833,857]" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="fragilis">
|
||
<emphasis id="6990EAF1B42BFFBEFE90177FFEAAFCD1" box="[313,494,833,857]" italics="true" pageId="1" pageNumber="38">Batrisus fragilis</emphasis>
|
||
<bibRefCitation id="3F754B12B42BFFBEFE5C177CFD39FCD1" author="Sharp" box="[501,637,833,857]" pageId="1" pageNumber="38" refString="Sharp, D. (1883) Revision of the Pselaphidae of Japan. Transactions of the Entomological Society of London, 1883 (3), 291 - 331." type="journal article" year="1883">Sharp, 1883</bibRefCitation>
|
||
</taxonomicName>
|
||
(by original designation)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13FE6568B42BFFBEFF6C1758FB5EFBB9" pageId="1" pageNumber="38" type="diagnosis">
|
||
<paragraph id="5B5B36E3B42BFFBEFF6C1758FB5EFBB9" blockId="1.[151,1437,833,2009]" pageId="1" pageNumber="38">
|
||
<emphasis id="6990EAF1B42BFFBEFF6C1758FE7BFCF6" bold="true" box="[197,319,869,894]" pageId="1" pageNumber="38">Diagnosis.</emphasis>
|
||
Head lacking frontal rostrum, lacking semicircular sulcus connecting vertexal foveae, median longitudinal carina distinct to absent, when present separated from frons by transverse sulcus that may vary in depth; antennomere I with subconical trichome at anterolateral margin. Pronotum with distinct median and lateral longitudinal sulci, transverse antebasal sulcus continuous or interrupted near base of median longitudinal sulcus, lacking antebasal spines. Elytra each with two basal foveae, subhumeral foveae present, discal stria extending beyond midpoint of elytral length. Visible tergite I longer than remaining tergites combined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13FE6568B42BFFBDFF6C1000FE36FB60" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="description">
|
||
<paragraph id="5B5B36E3B42BFFBEFF6C1000FB20FB6D" blockId="1.[151,1437,833,2009]" pageId="1" pageNumber="38">
|
||
<emphasis id="6990EAF1B42BFFBEFF6C1000FE37FBDE" bold="true" box="[197,371,1085,1110]" pageId="1" pageNumber="38">Redescription.</emphasis>
|
||
Body yellowish brown to reddish brown, length 1.70–2.34. Head rectangular, slightly wider than long to slightly elongate, antennal tubercles prominent; vertexal foveae nude, located posterior to point level with anterior margin of eyes; with 11 antennomeres, clubs formed by apical three antennomeres; ocular-mandibular carinae present; eyes round and prominent; maxillary palpi with palpomere I minute, II elongate and pedunculate, III short, nearly triangular, IV large, fusiform; with gular carina and single gular fovea.
|
||
</paragraph>
|
||
<paragraph id="5B5B36E3B42BFFBEFF6C10CFFA67FAA5" blockId="1.[151,1437,833,2009]" pageId="1" pageNumber="38">Pronotum about as long as wide, with four sulci, one pair of lateral antebasal foveae and two pairs of basolateral foveae; lacking spines; paranotal carinae present; with lateral procoxal foveae and subcoxal foveae.</paragraph>
|
||
<paragraph id="5B5B36E3B42BFFBEFF6C1104FA29FAD9" blockId="1.[151,1437,833,2009]" box="[197,1389,1337,1361]" pageId="1" pageNumber="38">Elytra usually wider than long, narrowed at base; sutural and marginal striae and lateroapical cleft present.</paragraph>
|
||
<paragraph id="5B5B36E3B42BFFBEFF6C1160FBD3FA35" blockId="1.[151,1437,833,2009]" pageId="1" pageNumber="38">Thorax with median mesoventral foveae widely forked, lateral mesoventral foveae straightly inserted; lateral mesocoxal foveae present; lateral metaventral foveae small, metaventral apex slightly notched. Legs slender, fore and mid tarsomeres II and III subequal in length, hind tarsomeres II slightly longer than III.</paragraph>
|
||
<paragraph id="5B5B36E3B42BFFBEFF6C11F4FBADF9C5" blockId="1.[151,1437,833,2009]" pageId="1" pageNumber="38">Abdomen with lateral margins subparallel to broadly rounded, apex narrowed and rounded; tergite IV largest, with mediobasal impression between mediobasal foveae, basolateral foveae with wide openings, discal carinae present; tergites V–VII successively narrower, V–VI about subequal in length, VII about as long as V and VI combined; sternite IV about as long as V–VII combined, with mediobasal and large basolateral foveae.</paragraph>
|
||
<paragraph id="5B5B36E3B42BFFBEFF6C1264FB8AF955" blockId="1.[151,1437,833,2009]" pageId="1" pageNumber="38">Males with sexual characters present on mesotibiae, metafemora, tergites IV–VI, and on sternite IV. Aedeagus strongly asymmetric, median lobe with large basal capsule; parameres fused and strongly reduced, attached on ventral side of capsule, membranous; ventral lobe broad at base, narrowed towards apex; dorsal apophysis inserted at dorsal side of basal capsule, on right side, usually strongly curved leftward, basal foramen large.</paragraph>
|
||
<paragraph id="5B5B36E3B42BFFBEFF6C12D4FC98F8C1" blockId="1.[151,1437,833,2009]" pageId="1" pageNumber="38">Females with eyes smaller, lacking modifications; genitalia basically provided with sternite IX and genital plate, sternite IX divided into two lobes, one usually paired and attached to another larger transverse one, genital plate narrower than sternite IX, with paired lateral arms narrowed apically.</paragraph>
|
||
<paragraph id="5B5B36E3B42BFFBDFF6C1368FE48FD2F" blockId="1.[151,1437,833,2009]" lastBlockId="2.[151,1437,151,1653]" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
|
||
<emphasis id="6990EAF1B42BFFBEFF6C1368FEF6F8E5" bold="true" box="[197,434,1877,1902]" pageId="1" pageNumber="38">Comparative notes.</emphasis>
|
||
<taxonomicName id="9CE44D60B42BFFBEFE151368FD18F8E5" box="[444,604,1877,1901]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B42BFFBEFE151368FD18F8E5" box="[444,604,1877,1901]" italics="true" pageId="1" pageNumber="38">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
is allied to a number of genera (three Asian, two African) centered on
|
||
<taxonomicName id="9CE44D60B42BFFBEFF3E1344FE8EF819" authority="Raffray. Current" authorityName="Raffray. Current" box="[151,458,1913,1937]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B42BFFBEFF3E1344FE49F819" box="[151,269,1913,1937]" italics="true" pageId="1" pageNumber="38">Batrisiella</emphasis>
|
||
Raffray. Current
|
||
</taxonomicName>
|
||
definitions of several group members are insufficient and problematical. Some of them are heavily relied upon the position of male sexual characters.
|
||
<taxonomicName id="9CE44D60B42BFFBEFCFD13A0FCB0F83D" box="[852,1012,1949,1973]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B42BFFBEFCFD13A0FCB0F83D" box="[852,1012,1949,1973]" italics="true" pageId="1" pageNumber="38">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
was defined by the male sexual modifications present on tergites VI–VII (
|
||
<bibRefCitation id="3F754B12B42BFFBEFDE213FCFC5EF851" author="Jeannel" box="[587,794,1985,2009]" pageId="1" pageNumber="38" refString="Jeannel, R. (1958) Revision des Pselaphides du Japon. Memoires du Museum National d'Histoire Naturelle, (N. S., Serie A, Zoologie), 18, 1 - 138." type="journal article" year="1958">Jeannel, 1958: 60</bibRefCitation>
|
||
), and was redefined by
|
||
<bibRefCitation id="3F754B12B42BFFBEFB9D13FFFA64F851" author="Nomura" box="[1076,1312,1985,2009]" pageId="1" pageNumber="38" refString="Nomura, S. (1991) Systematic study on the genus Batrisoplisus and its allied genera from Japan (Coleoptera, Pselaphidae). Esakia, 30, 1 - 462." type="journal article" year="1991">Nomura (1991: 299)</bibRefCitation>
|
||
to include presence of a subconical trichome on the anterolateral margin of the first antennomere.
|
||
<taxonomicName id="9CE44D60B428FFBDFBDA14A5FBADFF38" box="[1139,1257,152,176]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFBDA14A5FBADFF38" box="[1139,1257,152,176]" italics="true" pageId="2" pageNumber="39">Batrisiella</emphasis>
|
||
</taxonomicName>
|
||
was defined by the male sexual modifications being usually present on tergite IV, and antennomere I lacking modified setae (
|
||
<bibRefCitation id="3F754B12B428FFBDFAF61481FE48FF7F" author="Lobl" pageId="2" pageNumber="39" refString="Lobl, I. & Kurbatov, S. A. (2001) The Batrisini of Sri Lanka (Coleoptera: Staphylinidae: Pselaphinae). Revue Suisse de Zoologie, 108 (3), 559 - 697." type="journal article" year="2001">Löbl, 2001: 671</bibRefCitation>
|
||
). However, the definitions of the two genera were confused when specimens of
|
||
<taxonomicName id="9CE44D60B428FFBDFB3F14DDFADCFF7F" authority="Lobl" authorityName="Lobl" box="[1174,1432,223,248]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="aurita">
|
||
<emphasis id="6990EAF1B428FFBDFB3F14DDFA1CFF70" box="[1174,1368,224,248]" italics="true" pageId="2" pageNumber="39">Batrisiella aurita</emphasis>
|
||
Löbl
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CE44D60B428FFBDFF3E1539FEF4FE94" authority="Nomura" authorityName="Nomura" box="[151,432,260,284]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="6990EAF1B428FFBDFF3E1539FE0FFE94" box="[151,331,260,284]" italics="true" pageId="2" pageNumber="39">Batrisiella satoi</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CE44D60B428FFBDFE121539FC65FE94" authority="Nomura" authorityName="Nomura" box="[443,801,260,284]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="subalpicola">
|
||
<emphasis id="6990EAF1B428FFBDFE121539FDFFFE94" box="[443,699,260,284]" italics="true" pageId="2" pageNumber="39">Batrisiella subalpicola</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
, and some undescribed species became available for this study. Males of these
|
||
<taxonomicName id="9CE44D60B428FFBDFE221515FD45FEC8" box="[395,513,296,320]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFE221515FD45FEC8" box="[395,513,296,320]" italics="true" pageId="2" pageNumber="39">Batrisiella</emphasis>
|
||
</taxonomicName>
|
||
species and two undescribed species have a sexual patch on tergite IV, while they also have antennomere I bearing subconical setae, and one undescribed species has strongly thickened metafemora in the male. We found it hard to apply the current generic definitions to place these species either in
|
||
<taxonomicName id="9CE44D60B428FFBDFB55154DFAD8FE00" box="[1276,1436,368,392]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFB55154DFAD8FE00" box="[1276,1436,368,392]" italics="true" pageId="2" pageNumber="39">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="9CE44D60B428FFBDFF1F15A9FE68FE24" box="[182,300,404,428]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFF1F15A9FE68FE24" box="[182,300,404,428]" italics="true" pageId="2" pageNumber="39">Batrisiella</emphasis>
|
||
</taxonomicName>
|
||
. As the male secondary sexual characters of
|
||
<taxonomicName id="9CE44D60B428FFBDFC8615A9FCCBFE24" box="[815,911,404,428]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
|
||
may be highly variable within a single genus, and their use as group characters are usually unreliable (
|
||
<bibRefCitation id="3F754B12B428FFBDFC9C158AFCB3FE47" author="Lobl" box="[821,1015,439,463]" pageId="2" pageNumber="39" refString="Lobl, I. & Kurbatov, S. A. (2001) The Batrisini of Sri Lanka (Coleoptera: Staphylinidae: Pselaphinae). Revue Suisse de Zoologie, 108 (3), 559 - 697." type="journal article" year="2001">Löbl, 2001: 566</bibRefCitation>
|
||
), we therefore choose the presence/ absence of a trichome on antennomere I to serve as the only criterion of value in separating the two genera, and the form and position of the male sexual characters are considered useless at the generic level. We are relieved that pictures of the first antennomere of the
|
||
<typeStatus id="845F8841B428FFBDFD851618FD18FDB5" box="[556,604,549,573]" pageId="2" pageNumber="39">type</typeStatus>
|
||
species of both genera,
|
||
<taxonomicName id="9CE44D60B428FFBDFCCE1619FBFEFDB4" authority="Sharp" authorityName="Sharp" box="[871,1210,548,572]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="fragilis">
|
||
<emphasis id="6990EAF1B428FFBDFCCE1619FB1BFDB4" box="[871,1119,548,572]" italics="true" pageId="2" pageNumber="39">Batriscenellus fragilis</emphasis>
|
||
(Sharp)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9CE44D60B428FFBDFB461619FE18FDD7" authority="Raffray" authorityName="Raffray" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="caviventris">
|
||
<emphasis id="6990EAF1B428FFBDFB461619FFAFFDE8" italics="true" pageId="2" pageNumber="39">Batrisiella caviventris</emphasis>
|
||
(Raffray)
|
||
</taxonomicName>
|
||
, were available for our use, thanks to the collaboration of Shûhei Nomura. As a result,
|
||
<taxonomicName id="9CE44D60B428FFBDFA8C1675FE5EFD0C" authority="Lobl" authorityName="Lobl" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="aurita">
|
||
<emphasis id="6990EAF1B428FFBDFA8C1675FF9FFD0C" italics="true" pageId="2" pageNumber="39">Batrisiella aurita</emphasis>
|
||
Löbl
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CE44D60B428FFBDFE8C1651FD78FD0C" authority="Nomura" authorityName="Nomura" box="[293,572,620,644]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="6990EAF1B428FFBDFE8C1651FE9DFD0C" box="[293,473,620,644]" italics="true" pageId="2" pageNumber="39">Batrisiella satoi</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9CE44D60B428FFBDFDDA1651FC92FD0C" authority="Nomura" authorityName="Nomura" box="[627,982,620,644]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="subalpicola">
|
||
<emphasis id="6990EAF1B428FFBDFDDA1651FC36FD0C" box="[627,882,620,644]" italics="true" pageId="2" pageNumber="39">Batrisiella subalpicola</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
are moved to
|
||
<taxonomicName id="9CE44D60B428FFBDFBDF1651FA52FD0C" box="[1142,1302,620,644]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFBDF1651FA52FD0C" box="[1142,1302,620,644]" italics="true" pageId="2" pageNumber="39">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="6990EAF1B428FFBDFA8C1651FE41FD20" bold="true" pageId="2" pageNumber="39">New combinations</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5B5B36E3B428FFBDFF6E1688FD9FFC40" blockId="2.[151,1437,151,1653]" pageId="2" pageNumber="39">
|
||
<bibRefCitation id="3F754B12B428FFBDFF6E1688FE31FD44" author="Nomura" box="[199,373,692,716]" pageId="2" pageNumber="39" refString="Nomura, S. (1995) Descriptions of a new genus and a new species of the tribe Batrisini from Japan (Coleoptera, Pselaphidae). Esakia, 35: 123 - 128." type="journal article" year="1995">Nomura (1995)</bibRefCitation>
|
||
described the monotypic genus
|
||
<taxonomicName id="9CE44D60B428FFBDFD4C1689FCAFFD44" authority="Nomura" authorityName="Nomura" box="[741,1003,692,716]" class="Insecta" family="Staphylinidae" genus="Babascenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFD4C1689FCC3FD44" box="[741,903,692,716]" italics="true" pageId="2" pageNumber="39">Babascenellus</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
from Honshu and the Ryukyu Islands of
|
||
<collectingCountry id="23F37673B428FFBDFF1F16E5FFB2FD67" box="[182,246,728,751]" name="Japan" pageId="2" pageNumber="39">Japan</collectingCountry>
|
||
(three identified specimens examined in the present study). As stated, it is separated from
|
||
<taxonomicName id="9CE44D60B428FFBDFB5D16E5FAD0FD78" box="[1268,1428,728,752]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFB5D16E5FAD0FD78" box="[1268,1428,728,752]" italics="true" pageId="2" pageNumber="39">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CE44D60B428FFBDFF3E16C1FE49FC9C" box="[151,269,764,788]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFF3E16C1FE49FC9C" box="[151,269,764,788]" italics="true" pageId="2" pageNumber="39">Batrisiella</emphasis>
|
||
</taxonomicName>
|
||
, and two African genera,
|
||
<taxonomicName id="9CE44D60B428FFBDFD8616C1FC50FC9C" authority="Jeannel" authorityName="Jeannel" box="[559,788,764,788]" class="Insecta" family="Staphylinidae" genus="Arthromelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFD8616C1FDFEFC9C" box="[559,698,764,788]" italics="true" pageId="2" pageNumber="39">Arthromelus</emphasis>
|
||
Jeannel
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9CE44D60B428FFBDFCE316C1FB53FC9C" authority="Jeannel" authorityName="Jeannel" box="[842,1047,764,788]" class="Insecta" family="Staphylinidae" genus="Seydelites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFCE316C1FCFEFC9C" box="[842,954,764,788]" italics="true" pageId="2" pageNumber="39">Seydelites</emphasis>
|
||
Jeannel
|
||
</taxonomicName>
|
||
, by the strongly protruding anterolateral margin of antennomeres I in both sexes, and by the dorsal apophysis of the aedeagus not curving at the base. Since the dorsal apophyses of at least several
|
||
<taxonomicName id="9CE44D60B428FFBDFD331779FC54FCD4" box="[666,784,836,860]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFD331779FC54FCD4" box="[666,784,836,860]" italics="true" pageId="2" pageNumber="39">Batrisiella</emphasis>
|
||
</taxonomicName>
|
||
species seem have a straight base (
|
||
<bibRefCitation id="3F754B12B428FFBDFB371779FACBFCD4" author="Lobl" box="[1182,1423,836,860]" pageId="2" pageNumber="39" refString="Lobl, I. & Kurbatov, S. A. (2001) The Batrisini of Sri Lanka (Coleoptera: Staphylinidae: Pselaphinae). Revue Suisse de Zoologie, 108 (3), 559 - 697." type="journal article" year="2001">Löbl, 2001: 683, 688</bibRefCitation>
|
||
), the character of the curved base of the dorsal apophysis could hardly be considered as a synapomorphy of
|
||
<taxonomicName id="9CE44D60B428FFBDFF3E17B1FE49FC2C" box="[151,269,908,932]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFF3E17B1FE49FC2C" box="[151,269,908,932]" italics="true" pageId="2" pageNumber="39">Batrisiella</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="9CE44D60B428FFBDFE8717B1FE8AFC2C" box="[302,462,908,932]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFE8717B1FE8AFC2C" box="[302,462,908,932]" italics="true" pageId="2" pageNumber="39">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="9CE44D60B428FFBDFE5917B1FD38FC2C" box="[496,636,908,932]" class="Insecta" family="Staphylinidae" genus="Arthromelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFE5917B1FD38FC2C" box="[496,636,908,932]" italics="true" pageId="2" pageNumber="39">Arthromelus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="9CE44D60B428FFBDFD3517B1FC48FC2C" box="[668,780,908,932]" class="Insecta" family="Staphylinidae" genus="Seydelites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFD3517B1FC48FC2C" box="[668,780,908,932]" italics="true" pageId="2" pageNumber="39">Seydelites</emphasis>
|
||
</taxonomicName>
|
||
. For now we accept the placement of
|
||
<taxonomicName id="9CE44D60B428FFBDFB6C17B1FA23FC2C" box="[1221,1383,908,932]" class="Insecta" family="Staphylinidae" genus="Babascenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFB6C17B1FA23FC2C" box="[1221,1383,908,932]" italics="true" pageId="2" pageNumber="39">Babascenellus</emphasis>
|
||
</taxonomicName>
|
||
as a genus on the basis of its peculiar antennal structure.
|
||
</paragraph>
|
||
<paragraph id="5B5B36E3B428FFBDFF6E17E8FE36FB60" blockId="2.[151,1437,151,1653]" pageId="2" pageNumber="39">
|
||
We are not familiar with the two African genera
|
||
<taxonomicName id="9CE44D60B428FFBDFD5D17E9FC3BFC64" box="[756,895,980,1004]" class="Insecta" family="Staphylinidae" genus="Arthromelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFD5D17E9FC3BFC64" box="[756,895,980,1004]" italics="true" pageId="2" pageNumber="39">Arthromelus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9CE44D60B428FFBDFC1117E9FB6CFC64" box="[952,1064,980,1004]" class="Insecta" family="Staphylinidae" genus="Seydelites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFC1117E9FB6CFC64" box="[952,1064,980,1004]" italics="true" pageId="2" pageNumber="39">Seydelites</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="9CE44D60B428FFBDFB9E17E9FBE9FC64" box="[1079,1197,980,1004]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFB9E17E9FBE9FC64" box="[1079,1197,980,1004]" italics="true" pageId="2" pageNumber="39">Batrisiella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9CE44D60B428FFBDFB4F17E9FA35FC64" box="[1254,1393,980,1004]" class="Insecta" family="Staphylinidae" genus="Arthromelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFB4F17E9FA35FC64" box="[1254,1393,980,1004]" italics="true" pageId="2" pageNumber="39">Arthromelus</emphasis>
|
||
</taxonomicName>
|
||
are separated only by geographical ranges, and were suggested as probable synonyms by
|
||
<bibRefCitation id="3F754B12B428FFBDFBCE17C4FA17FB87" author="Nomura" box="[1127,1363,1015,1040]" pageId="2" pageNumber="39" refString="Nomura, S. (2003) Two new species of Batrisiella (Coleoptera, Staphylinidae, Pselaphinae) from the alpine area of Sichuan, China. Special Bulletin of the Japanese Society of Coleopterology, 6, 165 - 172." type="journal article" year="2003">Nomura (2003: 166)</bibRefCitation>
|
||
. As to
|
||
<taxonomicName id="9CE44D60B428FFBDFF3E1021FE43FBBC" box="[151,263,1052,1076]" class="Insecta" family="Staphylinidae" genus="Seydelites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFF3E1021FE43FBBC" box="[151,263,1052,1076]" italics="true" pageId="2" pageNumber="39">Seydelites</emphasis>
|
||
</taxonomicName>
|
||
, we have only seen a picture of
|
||
<taxonomicName id="9CE44D60B428FFBDFDC21020FC66FBBC" authority="Jeannel" authorityName="Jeannel" box="[619,802,1052,1076]" class="Insecta" family="Staphylinidae" genus="Seydelites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="franzi">
|
||
<emphasis id="6990EAF1B428FFBDFDC21020FD3DFBBC" box="[619,633,1053,1076]" italics="true" pageId="2" pageNumber="39">S</emphasis>
|
||
.
|
||
<emphasis id="6990EAF1B428FFBDFD2F1021FD83FBBC" box="[646,711,1052,1076]" italics="true" pageId="2" pageNumber="39">franzi</emphasis>
|
||
Jeannel
|
||
</taxonomicName>
|
||
(Nomura, pers. com.). Males of this species have the lateral margins of tergite IV strongly expanded laterally, but based on the picture we failed to find any consistent character that could serve as a criterion for separating it from
|
||
<taxonomicName id="9CE44D60B428FFBDFCBD1059FCE4FBF4" box="[788,928,1124,1148]" class="Insecta" family="Staphylinidae" genus="Arthromelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFCBD1059FCE4FBF4" box="[788,928,1124,1148]" italics="true" pageId="2" pageNumber="39">Arthromelus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="9CE44D60B428FFBDFC6A1059FB7DFBF4" box="[963,1081,1124,1148]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFC6A1059FB7DFBF4" box="[963,1081,1124,1148]" italics="true" pageId="2" pageNumber="39">Batrisiella</emphasis>
|
||
</taxonomicName>
|
||
. Accurate placement of the two genera must require further examination of the
|
||
<typeStatus id="845F8841B428FFBDFD0110B4FD9CFB29" box="[680,728,1161,1185]" pageId="2" pageNumber="39">type</typeStatus>
|
||
species. To determine either the justification or the inappropriateness of the generic status of these related taxa, a phylogenetic analysis based on a wide range of
|
||
<taxonomicName id="9CE44D60B428FFBDFB431091FA0EFB4C" box="[1258,1354,1196,1220]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
|
||
genera is probably needed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13FE6568B428FFBDFF6C10C9FD40F9FC" pageId="2" pageNumber="39" type="discussion">
|
||
<paragraph id="5B5B36E3B428FFBDFF6C10C9FD40F9FC" blockId="2.[151,1437,151,1653]" pageId="2" pageNumber="39">
|
||
<emphasis id="6990EAF1B428FFBDFF6C10C9FE7DFA84" bold="true" box="[197,313,1268,1292]" pageId="2" pageNumber="39">Remarks.</emphasis>
|
||
Within
|
||
<taxonomicName id="9CE44D60B428FFBDFE3A10C9FD77FA84" box="[403,563,1268,1292]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFE3A10C9FD77FA84" box="[403,563,1268,1292]" italics="true" pageId="2" pageNumber="39">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
, four subgenera are currently recognized. Characters used to separate subgenera rely mostly upon the presence/absence and position of male sexual characters on the abdominal tergites. However, the subgeneric concepts were formed largely based on geographically restricted material. Most described species of
|
||
<taxonomicName id="9CE44D60B428FFBDFEA6115DFEEBFAF0" box="[271,431,1376,1400]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFEA6115DFEEBFAF0" box="[271,431,1376,1400]" italics="true" pageId="2" pageNumber="39">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
are from
|
||
<collectingCountry id="23F37673B428FFBDFDB6115CFD26FAF0" box="[543,610,1377,1400]" name="Japan" pageId="2" pageNumber="39">Japan</collectingCountry>
|
||
, and only a few (e.g.
|
||
<taxonomicName id="9CE44D60B428FFBDFCF1115CFCA7FAF0" box="[856,995,1376,1400]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="6990EAF1B428FFBDFCF1115CFC2DFAF0" box="[856,873,1377,1400]" italics="true" pageId="2" pageNumber="39">B</emphasis>
|
||
.
|
||
<emphasis id="6990EAF1B428FFBDFCD1115DFCA7FAF0" box="[888,995,1376,1400]" italics="true" pageId="2" pageNumber="39">orientalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CE44D60B428FFBDFC5B115CFB28FAF0" box="[1010,1132,1377,1400]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="vicarius">
|
||
<emphasis id="6990EAF1B428FFBDFC5B115CFB47FAF0" box="[1010,1027,1377,1400]" italics="true" pageId="2" pageNumber="39">B</emphasis>
|
||
.
|
||
<emphasis id="6990EAF1B428FFBDFBBB115CFB28FAF0" box="[1042,1132,1377,1400]" italics="true" pageId="2" pageNumber="39">vicarius</emphasis>
|
||
</taxonomicName>
|
||
) are known from the adjacent Russian Far East, the Korean peninsula, and Northeast
|
||
<collectingCountry id="23F37673B428FFBDFCE111B9FCCFFA14" box="[840,907,1412,1436]" name="China" pageId="2" pageNumber="39">China</collectingCountry>
|
||
(except for one record of
|
||
<taxonomicName id="9CE44D60B428FFBDFB1211B8FA03FA14" box="[1211,1351,1412,1436]" class="Insecta" family="Staphylinidae" genus="Batrisiella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="6990EAF1B428FFBDFB1211B8FB88FA14" box="[1211,1228,1413,1436]" italics="true" pageId="2" pageNumber="39">B</emphasis>
|
||
.
|
||
<emphasis id="6990EAF1B428FFBDFB7511B9FA03FA14" box="[1244,1351,1412,1436]" italics="true" pageId="2" pageNumber="39">orientalis</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="23F37673B428FFBDFAC611B9FF8EFA37" name="Taiwan" pageId="2" pageNumber="39">Taiwan</collectingCountry>
|
||
). The subgeneric system does not apply well to material from outside this region. Species that have male sexual characters located on different parts of the body cannot be placed in any current subgenus of
|
||
<taxonomicName id="9CE44D60B428FFBDFB5511F1FAD8FA6C" box="[1276,1436,1484,1508]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFB5511F1FAD8FA6C" box="[1276,1436,1484,1508]" italics="true" pageId="2" pageNumber="39">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
as they are defined. We do not believe there is any justification for describing a number of additional subgenera as substitutes for species groups, and here we synonymize
|
||
<taxonomicName id="9CE44D60B428FFBDFC8B1229FB6CF9A4" authority="Jeannel" authorityName="Jeannel" box="[802,1064,1556,1580]" class="Insecta" family="Staphylinidae" genus="Scaioscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFC8B1229FC83F9A4" box="[802,967,1556,1580]" italics="true" pageId="2" pageNumber="39">Scaioscenellus</emphasis>
|
||
Jeannel
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9CE44D60B428FFBDFB9F1229FA22F9A4" authority="Nomura" authorityName="Nomura" box="[1078,1382,1556,1580]" class="Insecta" family="Staphylinidae" genus="Nipponoscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFB9F1229FBB9F9A4" box="[1078,1277,1556,1580]" italics="true" pageId="2" pageNumber="39">Nipponoscenellus</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="9CE44D60B428FFBDFF3E1205FEF4F9D8" authority="Nomura" authorityName="Nomura" box="[151,432,1592,1616]" class="Insecta" family="Staphylinidae" genus="Batriscenellinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFF3E1205FE09F9D8" box="[151,333,1592,1616]" italics="true" pageId="2" pageNumber="39">Batriscenellinus</emphasis>
|
||
Nomura
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="9CE44D60B428FFBDFE461205FDCBF9D8" box="[495,655,1592,1616]" class="Insecta" family="Staphylinidae" genus="Batriscenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6990EAF1B428FFBDFE461205FDCBF9D8" box="[495,655,1592,1616]" italics="true" pageId="2" pageNumber="39">Batriscenellus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="6990EAF1B428FFBDFD341205FC0BF9D8" bold="true" box="[669,847,1592,1616]" pageId="2" pageNumber="39">New synonyms</emphasis>
|
||
. The genus is divided to six newly proposed species groups which are defined below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |