202 lines
20 KiB
XML
202 lines
20 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5584834" ID-GBIF-Dataset="b9f36fb1-1c9d-4af8-96ca-d57973b94862" ID-GBIF-Taxon="192710751" ID-ISSN="1997-3500" ID-Zenodo-Dep="5584834" approvalRequired="1" checkinTime="1619623649741" checkinUser="donat" docAuthor="Seifert, Bernhard" docDate="2021" docId="F52B87F65E216156FF74DD2EFC301E78" docLanguage="en" docName="myrmecNews.31_133-179_printable.pdf" docOrigin="Myrmecological News 31" docSource="http://zoobank.org/0e55c0d7-531a-48d7-a078-148b96bd461d" docTitle="Formica major NYLANDER 1849" docType="treatment" docVersion="2" lastPageNumber="156" masterDocId="0912FF8E5E366141FFC9DA41FF91193E" masterDocTitle="A taxonomic revision of the Palaearctic members of the Formica rufa group (Hymenoptera: Formicidae) - the famous mound-building red wood ants" masterLastPageNumber="179" masterPageNumber="133" pageId="23" pageNumber="156" updateTime="1643400318318" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of the Palaearctic members of the Formica rufa group (Hymenoptera: Formicidae) - the famous mound-building red wood ants</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Seifert, Bernhard</mods:namePart>
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<mods:title>Myrmecological News</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-28</mods:number>
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<mods:detail type="volume">
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<mods:number>31</mods:number>
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<mods:extent unit="page">
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<mods:start>133</mods:start>
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<mods:end>179</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.25849/myrmecol.news_031:133</mods:identifier>
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<mods:identifier type="GBIF-Dataset">b9f36fb1-1c9d-4af8-96ca-d57973b94862</mods:identifier>
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<mods:identifier type="ISSN">1997-3500</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5582216</mods:identifier>
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<mods:identifier type="ZooBank">0E55C0D7-531A-48D7-A078-148B96BD461D</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5584834" ID-GBIF-Taxon="192710751" ID-Zenodo-Dep="5584834" LSID="urn:lsid:plazi:treatment:F52B87F65E216156FF74DD2EFC301E78" httpUri="http://treatment.plazi.org/id/F52B87F65E216156FF74DD2EFC301E78" lastPageNumber="156" pageId="23" pageNumber="156">
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<subSubSection box="[189,602,1903,1926]" pageId="23" pageNumber="156" type="nomenclature">
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<paragraph blockId="23.[151,773,1649,2023]" box="[189,602,1903,1926]" pageId="23" pageNumber="156">
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<treatmentCitation box="[189,602,1903,1926]" pageId="23" pageNumber="156">
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<taxonomicName authority="NYLANDER, 1849" authorityName="NYLANDER" authorityYear="1849" box="[189,602,1903,1926]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="major">
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<emphasis bold="true" box="[189,602,1903,1926]" pageId="23" pageNumber="156">
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<emphasis bold="true" box="[189,384,1903,1926]" italics="true" pageId="23" pageNumber="156">Formica major</emphasis>
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<bibRefCitation author="NYLANDER" box="[390,602,1904,1925]" firstAuthor="NYLANDER" journalOrPublisher="Acta Societatis Scientiarum Fennicae" pageId="23" pageNumber="156" pagination="25 - 48" part="3" refId="ref46234" refString="NYLANDER, W. 1849: Additamentum alterum adnotationum in monographiam formicarum borealium. - Acta Societatis Scientiarum Fennicae 3: 25 - 48." title="Additamentum alterum adnotationum in monographiam formicarum borealium" type="journal article" year="1849">NYLANDER, 1849</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</treatmentCitation>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,728,1936,1958]" pageId="23" pageNumber="156" type="reference_group">
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<paragraph blockId="23.[151,773,1649,2023]" box="[151,728,1936,1958]" pageId="23" pageNumber="156">
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<treatmentCitation box="[151,510,1937,1958]" pageId="23" pageNumber="156">
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<taxonomicName authority="NYLANDER, 1849" authorityName="NYLANDER" authorityYear="1849" box="[151,510,1937,1958]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="major">
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<emphasis box="[151,318,1937,1958]" italics="true" pageId="23" pageNumber="156">Formica major</emphasis>
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<bibRefCitation author="NYLANDER" box="[324,510,1937,1958]" firstAuthor="NYLANDER" journalOrPublisher="Acta Societatis Scientiarum Fennicae" pageId="23" pageNumber="156" pagination="25 - 48" part="3" refId="ref46234" refString="NYLANDER, W. 1849: Additamentum alterum adnotationum in monographiam formicarum borealium. - Acta Societatis Scientiarum Fennicae 3: 25 - 48." title="Additamentum alterum adnotationum in monographiam formicarum borealium" type="journal article" year="1849">NYLANDER, 1849</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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[
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<typeStatus box="[522,572,1937,1958]" pageId="23" pageNumber="156">type</typeStatus>
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investigation]
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="156" type="discussion">
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<paragraph blockId="23.[151,773,1649,2023]" lastBlockId="23.[815,1437,977,2023]" pageId="23" pageNumber="156">
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This taxon was described from the environs of Helsingfors (Helsinki). Three
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<typeStatus box="[387,473,2001,2022]" pageId="23" pageNumber="156">syntype</typeStatus>
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workers on one pin labelled “Zool. Mus. H:fors Spec. typ. No 5423
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<taxonomicName authority="Nyl." authorityName="NYLANDER" authorityYear="1849" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="major">Formica major Nyl.</taxonomicName>
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“, “major Nyl piniphila Schenck rufa auctt nec Bondr.”, “Mus. Hels. N:o 2676” do not carry an information on the sampling locality but the specimens fully match the morphological description of
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<bibRefCitation author="NYLANDER" box="[1094,1290,1105,1127]" firstAuthor="NYLANDER" journalOrPublisher="Acta Societatis Scientiarum Fennicae" pageId="23" pageNumber="156" pagination="25 - 48" part="3" refId="ref46234" refString="NYLANDER, W. 1849: Additamentum alterum adnotationum in monographiam formicarum borealium. - Acta Societatis Scientiarum Fennicae 3: 25 - 48." title="Additamentum alterum adnotationum in monographiam formicarum borealium" type="journal article" year="1849">NYLANDER (1849)</bibRefCitation>
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. Considering their numeric data and by subjective impression, the type series of
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<taxonomicName authorityName="NYLANDER" authorityYear="1849" box="[916,1011,1169,1190]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="major">
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<emphasis box="[916,1011,1169,1190]" italics="true" pageId="23" pageNumber="156">F. major</emphasis>
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</taxonomicName>
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cannot be hybrids
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<taxonomicName authorityName="FOERSTER" authorityYear="1850" box="[1226,1437,1169,1190]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="polyctena">
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<emphasis box="[1226,1437,1169,1190]" italics="true" pageId="23" pageNumber="156">Formica polyctena</emphasis>
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</taxonomicName>
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×
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<taxonomicName authorityName="LINNAEUS" authorityYear="1761" box="[839,887,1201,1222]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="rufa">
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<emphasis box="[839,887,1201,1222]" italics="true" pageId="23" pageNumber="156">rufa</emphasis>
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</taxonomicName>
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but represents either
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<taxonomicName authorityName="FOERSTER" authorityYear="1850" box="[1129,1264,1201,1222]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="polyctena">
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<emphasis box="[1129,1264,1201,1222]" italics="true" pageId="23" pageNumber="156">F. polyctena</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="YARROW" authorityYear="1955" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="aquilonia">
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<emphasis italics="true" pageId="23" pageNumber="156">Formica aquilonia</emphasis>
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</taxonomicName>
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×
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<taxonomicName authorityName="FOERSTER" authorityYear="1850" box="[901,1010,1233,1254]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="polyctena">
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<emphasis box="[901,1010,1233,1254]" italics="true" pageId="23" pageNumber="156">polyctena</emphasis>
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</taxonomicName>
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hybrids or backcrosses. This morphology-based idea is fully in line with the existence of large hybridization and introgression zone in southern
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<collectingCountry box="[1350,1436,1297,1319]" name="Finland" pageId="23" pageNumber="156">Finland</collectingCountry>
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shown by nuDNA data (
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<bibRefCitation author="BERESFORD" box="[1086,1339,1329,1351]" etAl="& al." firstAuthor="BERESFORD" journalOrPublisher="Molecular Ecology" pageId="23" pageNumber="156" pagination="4013 - 4026" part="26" refId="ref44707" refString="BERESFORD, J., ELIAS, M., PLUCKROSE, L., SUNDSTROM, L., BUTLIN, R. K., PAMILO, P. & KULMUNI, J. 2017: Widespread hybridization within mound-building wood ants in Southern Finland results in cytonuclear mismatches and potential for sex-specific hybrid breakdown. - Molecular Ecology 26: 4013 - 4026." title="Widespread hybridization within mound-building wood ants in Southern Finland results in cytonuclear mismatches and potential for sex-specific hybrid breakdown" type="journal article" year="2017">BERESFORD & al. 2017</bibRefCitation>
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). Run as wild-cards in a two-class LDA considering the characters
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<collectionCode box="[816,848,1393,1414]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="23" pageNumber="156">CS</collectionCode>
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, CL / CW
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<subScript attach="right" box="[949,987,1402,1416]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, SL /
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<collectionCode box="[1046,1075,1393,1414]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="23" pageNumber="156">CS</collectionCode>
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<subScript attach="left" box="[1075,1113,1402,1416]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, nCH
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<subScript attach="left" box="[1173,1211,1402,1416]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, OccHL
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<subScript attach="left" box="[1297,1335,1402,1416]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, nGu
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<subScript attach="left" box="[1392,1430,1402,1416]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, GuHL
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<subScript attach="left" box="[887,927,1434,1448]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, nPn
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<subScript attach="left" box="[994,1034,1434,1448]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, mPnHL
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<subScript attach="left" box="[1145,1185,1434,1448]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, nMes
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<subScript attach="left" box="[1268,1308,1434,1448]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, nMet
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<subScript attach="left" box="[1389,1429,1434,1448]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, MetHL
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<subScript attach="right" box="[894,932,1466,1480]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, nPr
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<subScript attach="left" box="[980,1018,1466,1480]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, SL / Smax
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<subScript attach="left" box="[1137,1175,1466,1480]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, EyeHL
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<subScript attach="left" box="[1259,1297,1466,1480]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, and nSc
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<subScript attach="left" box="[1392,1430,1466,1480]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, the type sample is allocated with p = 0.4867 to
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<taxonomicName authorityName="FOERSTER" authorityYear="1850" box="[1306,1437,1489,1510]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="polyctena">
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<emphasis box="[1306,1437,1489,1510]" italics="true" pageId="23" pageNumber="156">F. polyctena</emphasis>
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</taxonomicName>
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and with p = 0.5133 to the hybrid cluster. The marginal position in the hybrid cluster suggests
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<taxonomicName authorityName="NYLANDER" authorityYear="1849" box="[1240,1334,1553,1574]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="major">
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<emphasis box="[1240,1334,1553,1574]" italics="true" pageId="23" pageNumber="156">F. major</emphasis>
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</taxonomicName>
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to represent a backcross of a
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<taxonomicName authorityName="YARROW" authorityYear="1955" box="[1040,1171,1585,1606]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="aquilonia">
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<emphasis box="[1040,1171,1585,1606]" italics="true" pageId="23" pageNumber="156">F. aquilonia</emphasis>
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</taxonomicName>
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×
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<taxonomicName authorityName="FOERSTER" authorityYear="1850" box="[1196,1305,1585,1606]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="polyctena">
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<emphasis box="[1196,1305,1585,1606]" italics="true" pageId="23" pageNumber="156">polyctena</emphasis>
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</taxonomicName>
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hybrid with
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<taxonomicName authorityName="FOERSTER" authorityYear="1850" box="[816,948,1617,1638]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="polyctena">
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<emphasis box="[816,948,1617,1638]" italics="true" pageId="23" pageNumber="156">F. polyctena</emphasis>
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</taxonomicName>
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. A PCA plot considering the characters
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<collectionCode box="[1367,1400,1617,1638]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="23" pageNumber="156">CS</collectionCode>
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, SL /
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<collectionCode box="[833,862,1649,1670]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="23" pageNumber="156">CS</collectionCode>
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<subScript attach="left" box="[862,900,1658,1672]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, OccHL
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<subScript attach="left" box="[988,1026,1658,1672]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, GuHL
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<subScript attach="left" box="[1105,1143,1658,1672]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, SL / Smax
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<subScript attach="left" box="[1267,1305,1658,1672]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, EyeHL
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<subScript attach="left" box="[1391,1429,1658,1672]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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, and nSc
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<subScript attach="left" box="[902,940,1690,1704]" fontSize="6" pageId="23" pageNumber="156">1750</subScript>
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provides the same impression (
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<figureCitation box="[1280,1355,1681,1703]" captionStart="Fig" captionStartId="23.[816,852,625,647]" captionTargetBox="[847,1414,173,604]" captionTargetId="graphics-932@23.[887,1409,177,564]" captionTargetPageId="23" captionText="Fig.24: Position of the type samples of Formica major NY- LANDER, 1849 (MA) and Formica constricta KARAVAJEV, 1929 (CO) in a principal component analysis considering 58 nest samples of Formica polyctena (black squares) and 27 nest samples of Formica aquilonia × polyctena or backcrosses (white dots). The F. polyctena sample deeply placed within the hybrid cluster is aberrant and cannot be a hybrid for zoogeographical reasons. Seven phenotypic characters were considered." figureDoi="http://doi.org/10.5281/zenodo.5582255" httpUri="https://zenodo.org/record/5582255/files/figure.png" pageId="23" pageNumber="156">Fig.24</figureCitation>
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). These investigations considered 27 nest samples and
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<specimenCount box="[1307,1437,1713,1735]" pageId="23" pageNumber="156" type="worker">148 workers</specimenCount>
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of
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<taxonomicName authorityName="YARROW" authorityYear="1955" box="[842,973,1745,1766]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="aquilonia">
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<emphasis box="[842,973,1745,1766]" italics="true" pageId="23" pageNumber="156">F. aquilonia</emphasis>
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</taxonomicName>
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×
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<taxonomicName authorityName="FOERSTER" authorityYear="1850" box="[998,1107,1745,1766]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="polyctena">
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<emphasis box="[998,1107,1745,1766]" italics="true" pageId="23" pageNumber="156">polyctena</emphasis>
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</taxonomicName>
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hybrids or backcrosses and 57 nest samples with
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<specimenCount box="[1024,1158,1777,1799]" pageId="23" pageNumber="156" type="worker">217 workers</specimenCount>
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of
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<taxonomicName authorityName="FOERSTER" authorityYear="1850" box="[1195,1332,1777,1798]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="polyctena">
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<emphasis box="[1195,1332,1777,1798]" italics="true" pageId="23" pageNumber="156">F. polyctena</emphasis>
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</taxonomicName>
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. See also section “
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<treatmentCitation box="[912,1286,1808,1830]" pageId="23" pageNumber="156">
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<taxonomicName authority="YARROW, 1955" authorityName="YARROW" authorityYear="1955" box="[912,1286,1808,1830]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="156" phylum="Arthropoda" rank="species" species="aquilonia">
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<emphasis box="[912,1120,1809,1830]" italics="true" pageId="23" pageNumber="156">Formica aquilonia</emphasis>
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<bibRefCitation author="YARROW" box="[1128,1286,1808,1830]" firstAuthor="YARROW" journalOrPublisher="Transactions of the Society for British Entomology" pageId="23" pageNumber="156" pagination="1 - 48" part="12" refId="ref47744" refString="YARROW, I. H. H. 1955: The British ants allied to Formica rufa L. - Transactions of the Society for British Entomology 12: 1 - 48." title="The British ants allied to Formica rufa L" type="journal article" year="1955">YARROW, 1955</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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” (p. 158) and
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<figureCitation box="[816,924,1840,1863]" captionStart="Fig" captionStartId="24.[151,187,625,647]" captionTargetBox="[157,781,174,617]" captionTargetId="graphics-742@24.[201,763,176,582]" captionText="Fig.25: Position of the type samples of Formica major NY- LANDER, 1849 (MA) and Formica constricta KARAVAJEV,1929 (CO) in a linear discriminant analysis considering 58 nest samples of Formica polyctena (black squares), 27 nest samples of Formica aquilonia × polyctena or backcrosses (white dots), and 75 nest samples of F. aquilonia (black rhombs). The type samples were run aswild-cards.Sixteen phenotypiccharacters were considered." figureDoi="http://doi.org/10.5281/zenodo.5582257" httpUri="https://zenodo.org/record/5582257/files/figure.png" pageId="23" pageNumber="156">Figure 25</figureCitation>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |