treatments-xml/data/70/72/77/7072770BCC3C570C8A2A95B8C6E4EDFC.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1136.72744" ID-Pensoft-Pub="1313-2970-1136-1" ID-Pensoft-UUID="5D0E6613B1D05E4293ABEE4BFD1CA18A" ID-ZooBank="11C6BBFB339A4672AE662CE2B1E6321E" ModsDocID="1313-2970-1136-1" checkinTime="1671287389350" checkinUser="pensoft" docAuthor="Miller, Kelly B." docDate="2022" docId="7072770BCC3C570C8A2A95B8C6E4EDFC" docLanguage="en" docName="ZooKeys 1136: 1-56" docOrigin="ZooKeys 1136" docPubDate="2022-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1136.72744" docTitle="Desmopachria wolfei Miller 2022, sp. nov." docType="treatment" docUuid="854B6B76-1805-4601-8192-8772F6371F9E" docUuidSource="ZooBank" docVersion="2" id="5D0E6613B1D05E4293ABEE4BFD1CA18A" lastPageNumber="1" masterDocId="5D0E6613B1D05E4293ABEE4BFD1CA18A" masterDocTitle="Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus" masterLastPageNumber="56" masterPageNumber="1" pageNumber="1" updateTime="1671287691396" updateUser="ExternalLinkService">
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<mods:title>Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Kelly B.</mods:namePart>
<mods:affiliation>Department of Biology and Museum of Southwestern Biology University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
<mods:nameIdentifier type="email">kbmiller@unm.edu</mods:nameIdentifier>
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<mods:date>2022</mods:date>
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<mods:number>2022-12-16</mods:number>
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<mods:detail type="volume">
<mods:number>1136</mods:number>
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<mods:start>1</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1136.72744</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1136.72744</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:854B6B76-1805-4601-8192-8772F6371F9E" httpUri="http://treatment.plazi.org/id/7072770BCC3C570C8A2A95B8C6E4EDFC" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/854B6B76-1805-4601-8192-8772F6371F9E" authority="Miller, 2022" authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria wolfei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wolfei" status="sp. nov.">Desmopachria wolfei</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figures 46–50" captionStartId="F4" captionText="Figures 46 - 50. Desmopachria wolfei. 46 habitus 47 male median lobe, right lateral aspect 48 male median lobe, ventral aspect 49 male median lobe and right lateral lobe, ventral aspect 50 right lateral lobe, right lateral aspect. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1">Figures 46-50</figureCitation>
<figureCitation captionStart="Figures 79, 80" captionStartId="F9" captionText="Figures 79, 80. Desmopachria species distributions. 79 Desmopachria nitida species group and Desmopachria portmanni-aldessa species group 80 Desmopachria portmanni-aldessa species group." figureDoi="10.3897/zookeys.1136.72744.figures79-80" httpUri="https://binary.pensoft.net/fig/782461" pageId="0" pageNumber="1">, 79</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Venezuela, Bolivar State, Rio Aponwao at Highway 10, 
<geoCoordinate degrees="5" direction="north" minutes="50" orientation="latitude" precision="1" seconds="49.2" value="5.847">5°50'49.2&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="62" direction="west" minutes="28" orientation="longitude" precision="1" seconds="2.4" value="-62.467335">62°28'2.4&quot;W</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species belongs to the 
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria nitida" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitida">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria nitida</emphasis>
</taxonomicName>
group sensu 
<bibRefCitation DOI="https://doi.org/10.1649/0010-065X(2001)055[0219:DONSOD]2.0.CO;2" author="Miller, KB" journalOrPublisher="Entomologica Scandinavica" pageId="0" pageNumber="1" refId="B11" refString="Miller, KB, 2001. Descriptions of new species of Desmopachria Babington, 1841 (Coleoptera: Dytiscidae: Hydroporinae: Hyphydrini) with a reassessment of the subgenera and species groups and a synopsis of the species. The Coleopterists Bulletin 55: 219-240. https://doi.org/10.1649/0010-065X(2001)055[0219:DONSOD]2.0.CO;2" title="Descriptions of new species of Desmopachria Babington, 1841 (Coleoptera: Dytiscidae: Hydroporinae: Hyphydrini) with a reassessment of the subgenera and species groups and a synopsis of the species. The Coleopterists Bulletin 55: 219 - 240." url="https://doi.org/10.1649/0010-065X(2001)055[0219:DONSOD]2.0.CO;2" year="2001">Miller (2001)</bibRefCitation>
because of presence of deeply bifid male lateral lobes (Fig. 
<figureCitation captionStart="Figures 46–50" captionStartId="F4" captionText="Figures 46 - 50. Desmopachria wolfei. 46 habitus 47 male median lobe, right lateral aspect 48 male median lobe, ventral aspect 49 male median lobe and right lateral lobe, ventral aspect 50 right lateral lobe, right lateral aspect. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1">50</figureCitation>
). Within the group, the species has a bilaterally symmetrical, but extremely complicated male median lobe that is very unique in shape, medially distinctly constricted, apically broadly truncate, and with an apicomedial emargination, among other shape characteristics (Figs 
<figureCitation captionStart="Figures 46–50" captionStartId="F4" captionText="Figures 46 - 50. Desmopachria wolfei. 46 habitus 47 male median lobe, right lateral aspect 48 male median lobe, ventral aspect 49 male median lobe and right lateral lobe, ventral aspect 50 right lateral lobe, right lateral aspect. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1">47-50</figureCitation>
). Dorsal coloration is simple with the head and pronotum orange and elytron brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Measurements.</emphasis>
TL = 1.8-1.9 mm, GW = 1.2-1.3 mm, PW = 0.9-1.0 mm, HW = 0.6-0.7 mm, EW = 0.3 mm, TL/GW = 1.4-1.5, HW/EW = 1.9-2.0. Body broad, elongate oval, laterally broadly curved, lateral margins continuous between pronotum and elytron, body broadest across elytra at midlength of body (Fig. 
<figureCitation captionStart="Figures 46–50" captionStartId="F4" captionText="Figures 46 - 50. Desmopachria wolfei. 46 habitus 47 male median lobe, right lateral aspect 48 male median lobe, ventral aspect 49 male median lobe and right lateral lobe, ventral aspect 50 right lateral lobe, right lateral aspect. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1">46</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
(Fig. 
<figureCitation captionStart="Figures 46–50" captionStartId="F4" captionText="Figures 46 - 50. Desmopachria wolfei. 46 habitus 47 male median lobe, right lateral aspect 48 male median lobe, ventral aspect 49 male median lobe and right lateral lobe, ventral aspect 50 right lateral lobe, right lateral aspect. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1">46</figureCitation>
). Dorsal surface of head and pronotum evenly pale orange. Elytron dark orange, darker than surface of pronotum. Head appendages, legs and ventral surfaces orange to dark orange or orange-red.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sculpture and structure.</emphasis>
Head broad; anterior margin of clypeus evenly curved, flattened, margined with conspicuous, continuous narrow bead; surface of head shiny, finely and sparsely punctate; eyes large (HW/EW = 1.9-2.0); antennae short, scape and pedicel relatively large and rounded, flagellomere III long and slender, apically expanded, antennomeres IV-X short and broad, lobed at anterodorsal angle, antennomere XI elongate, apically pointed. Pronotum short, lateral margins short, sublinear with continuous narrow bead, slightly wider medially; surface shiny, impunctate medially, punctation denser along anterior and posterior margins, punctation fine. Elytron moderately broad, laterally broadly curved; surface shiny, more coarsely and evenly punctate than pronotum, punctation distinctive and prominent. Prosternum extremely short, longitudinally compressed, medially slightly carinate; prosternal process slender anteriorly, with distinctive, small medial tubercle, apical blade short and broad, basally transversely carinate, medially concave, apically broadly pointed. Metaventrite broad and evenly smoothly convex medially, surface shiny, very finely and sparsely; metaventrite wings extremely slender. Metacoxa with medial portion short, &lt;1/3 length of metaventrite medially, metacoxal lines divergent anteriorly; lateral portion of metacoxa extremely large, anteriorly strongly expanded; surface shiny, very finely and sparsely punctate. Metatrochanter large, subequal to length of ventral margin of metafemur; legs otherwise not noticeably modified. Abdomen with surfaces shiny and smooth, very finely and sparsely punctate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia.</emphasis>
Male genitalia complex; median lobe elongate in lateral aspect, broad basally, medially constricted, broadly sinuate, apex strongly curved dorsal, apically pointed (Fig. 
<figureCitation captionStart="Figures 46–50" captionStartId="F4" captionText="Figures 46 - 50. Desmopachria wolfei. 46 habitus 47 male median lobe, right lateral aspect 48 male median lobe, ventral aspect 49 male median lobe and right lateral lobe, ventral aspect 50 right lateral lobe, right lateral aspect. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1">47</figureCitation>
); median lobe in ventral aspect very broad, lateral margins subparallel in basal half, with distinct constriction medially due to distinct lateral emarginations, apical half with lateral margins curved to apicolateral angle, apex very broadly truncate with distinct medial emargination (Figs 
<figureCitation captionStart="Figures 46–50" captionStartId="F4" captionText="Figures 46 - 50. Desmopachria wolfei. 46 habitus 47 male median lobe, right lateral aspect 48 male median lobe, ventral aspect 49 male median lobe and right lateral lobe, ventral aspect 50 right lateral lobe, right lateral aspect. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1">48</figureCitation>
, 
<figureCitation captionStart="Figures 46–50" captionStartId="F4" captionText="Figures 46 - 50. Desmopachria wolfei. 46 habitus 47 male median lobe, right lateral aspect 48 male median lobe, ventral aspect 49 male median lobe and right lateral lobe, ventral aspect 50 right lateral lobe, right lateral aspect. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1">49</figureCitation>
). Lateral lobe in broad basally, apically conspicuously bifurcate, dorsal branch apically broadly rounded, ventral branch apically truncate with dorsal corner acutely pointed dorsad (Fig. 
<figureCitation captionStart="Figures 46–50" captionStartId="F4" captionText="Figures 46 - 50. Desmopachria wolfei. 46 habitus 47 male median lobe, right lateral aspect 48 male median lobe, ventral aspect 49 male median lobe and right lateral lobe, ventral aspect 50 right lateral lobe, right lateral aspect. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1">50</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sexual dimorphism.</emphasis>
No obvious sexual dimorphic features were discovered.
</paragraph>
<caption doi="10.3897/zookeys.1136.72744.figures46-50" httpUri="https://binary.pensoft.net/fig/782456" pageId="0" pageNumber="1" start="Figures 46–50" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 46-50.</emphasis>
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria wolfei." order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wolfei.">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria wolfei.</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">46</emphasis>
habitus 
<emphasis bold="true" pageId="0" pageNumber="1">47</emphasis>
male median lobe, right lateral aspect 
<emphasis bold="true" pageId="0" pageNumber="1">48</emphasis>
male median lobe, ventral aspect 
<emphasis bold="true" pageId="0" pageNumber="1">49</emphasis>
male median lobe and right lateral lobe, ventral aspect 
<emphasis bold="true" pageId="0" pageNumber="1">50</emphasis>
right lateral lobe, right lateral aspect. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Variation.</emphasis>
Some specimens are variously paler or darker in coloration than others.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This distinctive species is named 
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria wolfei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wolfei">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria wolfei</emphasis>
</taxonomicName>
for G.W. Wolfe, gifted coleopterist, exemplary water beetle biologist, exceptionally fine husband and father, and the 
<normalizedToken originalValue="author’s">author's</normalizedToken>
dear friend for many years.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known only from Bolivar State, Venezuela (Fig. 
<figureCitation captionStart="Figures 79, 80" captionStartId="F9" captionText="Figures 79, 80. Desmopachria species distributions. 79 Desmopachria nitida species group and Desmopachria portmanni-aldessa species group 80 Desmopachria portmanni-aldessa species group." figureDoi="10.3897/zookeys.1136.72744.figures79-80" httpUri="https://binary.pensoft.net/fig/782461" pageId="0" pageNumber="1">79</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype in MIZA, male labeled, &quot;VENEZUELA: Bolivar State 
<geoCoordinate degrees="5" direction="north" minutes="50" orientation="latitude" precision="1" seconds="49.2" value="5.847">5°50'49.2&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="62" direction="west" minutes="28" orientation="longitude" precision="1" seconds="2.4" value="-62.467335">62°28'2.4&quot;W</geoCoordinate>
, 1340 m Rio Aponwao @ Hwy 10 31.vii.2008; leg. A.Short, M. 
<normalizedToken originalValue="García">Garcia</normalizedToken>
AS-08-060a; small vegetated pool/ SM0827530 KUNHM-ENT [barcode label] / HOLOTYPE 
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria wolfei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wolfei">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria wolfei</emphasis>
</taxonomicName>
Miller, 2021 [red label with black line border].&quot; Paratypes, 23; 9 in MIZA and SEMC labeled same as holotype except with 
<normalizedToken originalValue="“…">&quot;...</normalizedToken>
PARATYPE 
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria wolfei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wolfei">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria wolfei</emphasis>
</taxonomicName>
Miller, 2021 [blue label with black line border]&quot; and different barcode labels (see Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Barcode label data for Desmopachria species." httpUri="http://table.plazi.org/id/B5FF9C5283064B11CB6B8D1966FB9A48" pageId="0" pageNumber="1" tableUuid="B5FF9C5283064B11CB6B8D1966FB9A48">1</tableCitation>
); 14 in SEMC labeled, &quot;VENEZUELA: Bolivar State 
<geoCoordinate degrees="4" direction="north" minutes="49.944" orientation="latitude" precision="1" value="4.8324">4°49.944'N</geoCoordinate>
, 
<geoCoordinate degrees="61" direction="west" minutes="3.813" orientation="longitude" precision="1" value="-61.06355">61°3.813'W</geoCoordinate>
, 890m ca. 25 km S. San Francisco 3.viii.2008; A.Short &amp; M. 
<normalizedToken originalValue="García">Garcia</normalizedToken>
AS-08-069; large marsh/...&quot;
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="checklist of species in the desmopachria nitida species group">
<paragraph pageId="0" pageNumber="1">
Checklist of species in the 
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria nitida" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitida">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria nitida</emphasis>
</taxonomicName>
species group
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K. B. Miller" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria annae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="annae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria annae</emphasis>
</taxonomicName>
Miller, 2005 - Bolivia
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria anauine" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anauine">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria anauine</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2018 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Makhan" authorityYear="2012" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria aschnae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aschnae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria aschnae</emphasis>
</taxonomicName>
Makhan, 2012 - Suriname. This species is hereby placed into the 
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria nitida" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitida">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria nitida</emphasis>
</taxonomicName>
species group based on published figures of bifid lateral lobes (
<bibRefCitation author="Makhan, D" journalOrPublisher="Calodema" pageId="0" pageNumber="1" pagination="1 - 6" refId="B7" refString="Makhan, D, 2012. Four new species of Desmopachria (Coleoptera: Dytiscidae) from Suriname. Calodema 235: 1 - 6" title="Four new species of Desmopachria (Coleoptera: Dytiscidae) from Suriname." volume="235" year="2012">Makhan 2012</bibRefCitation>
: fig. 4).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K. B. Miller" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria balionota" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="balionota">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria balionota</emphasis>
</taxonomicName>
Miller, 2005 - Peru, Brazil (Braga and Ferreira-Jr. 2010, 2014).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K.B.Miller &amp; Wolfe" authorityYear="2018" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria curseenae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="curseenae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria curseenae</emphasis>
</taxonomicName>
Miller &amp; Wolfe, 2018 - Suriname
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1989" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria darlingtoni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria darlingtoni</emphasis>
</taxonomicName>
Young, 1989 - Jamaica, Cuba, Haiti, Colombia
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K.B.Miller &amp; Wolfe" authorityYear="2018" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria delongi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="delongi">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria delongi</emphasis>
</taxonomicName>
Miller &amp; Wolfe, 2018 - Suriname
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria draco" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="draco">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria draco</emphasis>
</taxonomicName>
Miller, 1999 - Bolivia, Brazil (Braga and Ferreira-Jr. 2010)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K.B.Miller &amp; Wolfe" authorityYear="2018" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria gingerae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="gingerae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria gingerae</emphasis>
</taxonomicName>
Miller &amp; Wolfe, 2018 - Venezuela
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1986" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria granoides" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="granoides">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria granoides</emphasis>
</taxonomicName>
Young, 1986 - Brazil (Braga and Ferreira-Jr. 2014), Bolivia, Suriname, Venezuela, Trinidad
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K.B.Miller &amp; Wolfe" authorityYear="2018" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria gyrationi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="gyrationi">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria gyrationi</emphasis>
</taxonomicName>
Miller &amp; Wolfe, 2018 - Guyana
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K.B.Miller &amp; Wolfe" authorityYear="2018" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria hardyae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hardyae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria hardyae</emphasis>
</taxonomicName>
Miller &amp; Wolfe, 2018 - Guyana
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K.B.Miller &amp; Wolfe" authorityYear="2018" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria kemptonae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kemptonae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria kemptonae</emphasis>
</taxonomicName>
Miller &amp; Wolfe, 2018 - Venezuela
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria leptophallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="leptophallica">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria leptophallica</emphasis>
</taxonomicName>
Braga &amp; Ferreira-Jr., 2014 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1986" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria liosomata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="liosomata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria liosomata</emphasis>
</taxonomicName>
Young, 1986 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K.B.Miller &amp; Wolfe" authorityYear="2018" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lloydi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lloydi">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lloydi</emphasis>
</taxonomicName>
Miller &amp; Wolfe, 2018 - Bolivia
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1990" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria margarita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="margarita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria margarita</emphasis>
</taxonomicName>
Young, 1990 - Panama, Brazil? (Braga and Ferreira-Jr. 2014).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria nitida" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitida">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria nitida</emphasis>
</taxonomicName>
Babington, 1841 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1990" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria nitidoides" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidoides">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria nitidoides</emphasis>
</taxonomicName>
Young, 1990 - Paraguay
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Guignot" authorityYear="1950" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria phacoides" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="phacoides">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria phacoides</emphasis>
</taxonomicName>
Guignot, 1950 - Paraguay, Bolivia
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria psarammo" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="psarammo">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria psarammo</emphasis>
</taxonomicName>
Miller, 1999 - Bolivia
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria rhea" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rhea">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria rhea</emphasis>
</taxonomicName>
Miller, 1999 - Bolivia
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K.B.Miller &amp; Wolfe" authorityYear="2018" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria singhae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="singhae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria singhae</emphasis>
</taxonomicName>
Miller &amp; Wolfe, 2018 - Venezuela
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Zimmermann" authorityYear="1921" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria subnotata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="subnotata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria subnotata</emphasis>
</taxonomicName>
Zimmermann, 1921 - Brazil (Braga and Ferreira-Jr. 2010).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria subtilis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="subtilis">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria subtilis</emphasis>
</taxonomicName>
Sharp, 1882 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K.B.Miller &amp; Wolfe" authorityYear="2018" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria vohrae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vohrae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria vohrae</emphasis>
</taxonomicName>
Miller &amp; Wolfe, 2018 - Venezuela
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria wolfei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wolfei">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria wolfei</emphasis>
</taxonomicName>
sp. nov. - Venezuela
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1990" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria zelota" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zelota">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria zelota</emphasis>
</taxonomicName>
Young, 1990 - Brazil
</paragraph>
</subSubSection>
</treatment>
</document>