treatments-xml/data/F5/A4/FB/F5A4FBB14EA35564709B05F227352B22.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.689.12021" ID-GBIF-Dataset="907713bd-c7f2-4f8e-a75e-b7f9c958865f" ID-PMC="PMC5672588" ID-Pensoft-Pub="1313-2970-689-1" ID-PubMed="29200920" ID-ZBK="2F40BF4AD35F4CC697D5976EC201E652" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-689-1" ModsDocOrigin="ZooKeys 689" ModsDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" checkinTime="1502744367222" checkinUser="pensoft" docAuthor="Lackner, Tomas &amp; Leschen, Richard A. B." docDate="2017" docId="F5A4FBB14EA35564709B05F227352B22" docLanguage="en" docName="ZooKeys 689: 1-263" docOrigin="ZooKeys 689" docSource="http://dx.doi.org/10.3897/zookeys.689.12021" docTitle="Saprinus (Saprinus) rarus Lackner &amp; Leschen, 2017, sp. n." docType="treatment" docUuid="7DF7647D-3AE5-4071-A734-C9C4EAB9920F" docUuidSource="ZooBank" docVersion="6" lastPageNumber="154" masterDocId="D302FFB6FFA66345FFBAFF89311EFFDE" masterDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" masterLastPageNumber="263" masterPageNumber="1" pageNumber="150" updateTime="1668164684569" updateUser="ExternalLinkService">
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<mods:title>A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lackner, Tomas</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Leschen, Richard A. B.</mods:namePart>
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<mods:date>2017</mods:date>
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<mods:number>689</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.689.12021</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.689.12021</mods:identifier>
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<treatment ID-GBIF-Taxon="132648672" LSID="urn:lsid:zoobank.org:act:7DF7647D-3AE5-4071-A734-C9C4EAB9920F" httpUri="http://treatment.plazi.org/id/F5A4FBB14EA35564709B05F227352B22" lastPageId="153" lastPageNumber="154" pageId="149" pageNumber="150">
<subSubSection pageId="149" pageNumber="150" type="nomenclature">
<paragraph pageId="149" pageNumber="150">
<taxonomicName LSID="http://zoobank.org/7DF7647D-3AE5-4071-A734-C9C4EAB9920F" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) rarus" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="rarus" subGenus="Saprinus">
<pageBreakToken pageId="149" pageNumber="150" start="start">Saprinus</pageBreakToken>
(Saprinus) rarus
</taxonomicName>
<taxonomicNameLabel pageId="149" pageNumber="150">sp. n.</taxonomicNameLabel>
Figs 528, 529-537, 538-546, 764
</paragraph>
</subSubSection>
<subSubSection pageId="149" pageNumber="150" type="type locality">
<paragraph pageId="149" pageNumber="150">Type locality.</paragraph>
<paragraph pageId="149" pageNumber="150">Australia: New South Wales: N of Sydney: Pearl Beach at Broken Bay.</paragraph>
<caption pageId="149" pageNumber="150">
<paragraph pageId="149" pageNumber="150">
Figure 528. 
<taxonomicName class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) rarus" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="rarus" subGenus="Saprinus">Saprinus (Saprinus) rarus</taxonomicName>
sp. n. habitus, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="149" pageNumber="150" type="type material">
<paragraph pageId="149" pageNumber="150">Type material examined.</paragraph>
<paragraph pageId="149" pageNumber="150">
Holotype, ♂, side-mounted on a triangular card, right metatibia broken off, glued to the same triangular card as the specimen, with male genitalia dismembered, glued to the same mounting card as the specimen, with the following labels: &quot;Australia, / N of Sydney, / Pearl Beach at Broken Bay / 7.iii.1997 / leg. D. Scherbakov&quot; (printed); with a consecutive label: &quot;Coll. of / A. Sokolov&quot; (hand-written); with consecutive label: &quot; 
<taxonomicName class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
(s.str.) 
<taxonomicName class="Insecta" family="Corydiidae" genus="Histeroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Histeroidea rarus" order="Blattodea" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="rarus">rarus</taxonomicName>
/ n. spec. HOLOTYPE / Det. T. Lackner &amp; R. / Leschen 2013&quot; (red label, hand-written) (MAMU).
</paragraph>
<paragraph pageId="149" pageNumber="150">
Paratype, ♀, side-mounted on a triangular card, right mesotarsus broken off, glued to the same triangular card as the specimen with the following labels: &quot;Blackdown / 
<normalizedToken originalValue="T’">T'</normalizedToken>
l Q. / 14.5. [19]81 / N.W. Rodd&quot; (printed-written); followed by: 
<normalizedToken originalValue="“♀”">&quot;♀&quot;</normalizedToken>
(beige label, printed); followed by: &quot;Australian Museum / K 270235&quot; (printed); followed by: &quot; 
<taxonomicName class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
(s.l.) / sp. / G. Arriagada det. 1990&quot; (black-framed label, printed-written); &quot;followed by: 
<normalizedToken originalValue="“10-115”">&quot;10-115&quot;</normalizedToken>
(light-yellow pencil written label); followed by: &quot; 
<taxonomicName class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
(s.str.) 
<taxonomicName class="Insecta" family="Corydiidae" genus="Histeroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Histeroidea rarus" order="Blattodea" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="rarus">rarus</taxonomicName>
/ n. spec. PARATYPE / Det. T. Lackner &amp; R. / Leschen 2013&quot; (red label, hand-written); followed by: &quot;Photographed by / B. Rhode&quot; (yellow label, printed) (MAMU); paratype, ♂, mounted on a mounting card, with the following labels: &quot; 
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-33.601944">33°36'07&quot;S</geoCoordinate>
; 
<geoCoordinate direction="east" orientation="longitude" precision="15" value="151.32138">151°19'17&quot;E</geoCoordinate>
/ AUSTRALIA: New South Wales, Palm Beach, 15. / VIII.1962, G.R.Wearne / Host: 
<taxonomicName class="Insecta" family="Termitidae" genus="Nasutitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nasutitermes walkeri" order="Blattodea" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="walkeri">Nasutitermes walkeri</taxonomicName>
(Hill)&quot; (printed); followed by: &quot;Australia: N.S.W / Palm Beach, / 15.viii.1962&quot; (printed-written); followed by: &quot;Coll., Host det. / G.R. Wearne&quot; (written); followed by: &quot;Host: / 
<taxonomicName class="Insecta" family="Termitidae" genus="Nasutitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nasutitermes" order="Blattodea" pageId="149" pageNumber="150" phylum="Arthropoda" rank="genus">Nasutitermes</taxonomicName>
/ 
<taxonomicName class="Insecta" family="Corydiidae" genus="Histeroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Histeroidea walkeri" order="Blattodea" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="walkeri">walkeri</taxonomicName>
&quot; (written); followed by: &quot; 
<taxonomicName class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus rarus" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="rarus">Saprinus rarus</taxonomicName>
/ sp.n. PARATYPE / det. T. Lackner 2016&quot; (red label, written) (CPK).
</paragraph>
</subSubSection>
<subSubSection pageId="149" pageNumber="150" type="biology_ecology">
<paragraph pageId="149" pageNumber="150">Biology.</paragraph>
<paragraph pageId="149" pageNumber="150">
Collected from the nest of the arboreal Tree termite (
<taxonomicName class="Insecta" family="Termitidae" genus="Nasutitermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nasutitermes walkeri" order="Blattodea" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="walkeri">Nasutitermes walkeri</taxonomicName>
(Hill, 1942)). Based on the morphology (thickened and dilated antennal scape, tibiae) and collection circumstances, 
<taxonomicName class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus rarus" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="rarus">Saprinus rarus</taxonomicName>
is presumed to be a specialized termitophile. This is the first record of a termitophilic 
<taxonomicName lsidName="" pageId="149" pageNumber="150" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
from the Australopacific Region and only the third case of termitophily in the subfamily in general (the two other taxa are: African 
<taxonomicName class="Insecta" family="Histeridae" genus="Pilisaprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilisaprinus verschureni" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="verschureni">Pilisaprinus verschureni</taxonomicName>
(
<normalizedToken originalValue="Thérond">Therond</normalizedToken>
, 1959) ihabiting dead termitaria of the genus 
<taxonomicName class="Insecta" family="Termitidae" genus="Macrotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrotermes" order="Blattodea" pageId="149" pageNumber="150" phylum="Arthropoda" rank="genus">Macrotermes</taxonomicName>
(
<taxonomicName family="Termitidae" lsidName="" pageId="149" pageNumber="150" rank="family">Termitidae</taxonomicName>
) recorded from Congo, Ivory Coast and Benin and 
<taxonomicName class="Insecta" family="Histeridae" genus="Nannolepidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannolepidius braunsi" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="braunsi">Nannolepidius braunsi</taxonomicName>
(Bickhardt, 1921) found in nests of 
<taxonomicName class="Insecta" family="Hodotermitidae" genus="Hodotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hodotermes" order="Blattodea" pageId="149" pageNumber="150" phylum="Arthropoda" rank="genus">Hodotermes</taxonomicName>
termites (
<taxonomicName family="Hodotermitidae" lsidName="" pageId="149" pageNumber="150" rank="family">Hodotermitidae</taxonomicName>
) in the Cape Region of South Africa, respectively).
</paragraph>
</subSubSection>
<subSubSection pageId="149" pageNumber="150" type="distribution">
<paragraph pageId="149" pageNumber="150">Distribution.</paragraph>
<paragraph pageId="149" pageNumber="150">This species is known only from three Australian specimens: two males collected near Sydney (New South Wales) and a female collected in Blackdown Tableland National Park, near Rockhampton (Queensland) (Fig. 764).</paragraph>
</subSubSection>
<subSubSection pageId="149" pageNumber="150" type="etymology">
<paragraph pageId="149" pageNumber="150">Etymology.</paragraph>
<paragraph pageId="149" pageNumber="150">
The specific epithet ' 
<taxonomicName class="Insecta" family="Corydiidae" genus="Histeroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Histeroidea rarus" order="Blattodea" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="rarus">rarus</taxonomicName>
' refers to the scarcity of this beetle in collections.
</paragraph>
</subSubSection>
<subSubSection pageId="149" pageNumber="150" type="diagnosis">
<paragraph pageId="149" pageNumber="150">Diagnosis.</paragraph>
<paragraph pageId="149" pageNumber="150">
<taxonomicName class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus rarus" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="rarus">Saprinus rarus</taxonomicName>
has fused parameres (Fig. 545); its elongate body (Fig. 528), dilated tibiae (Figs 536-537), thickened and dilated antennal scape (Fig. 529) and large, almost circular antennal clubs (Fig. 530) will differentiate this species from other species in the genus.
</paragraph>
<caption pageId="149" pageNumber="150">
<paragraph pageId="149" pageNumber="150">
Figures 529-537. 529 
<taxonomicName class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) rarus" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="rarus" subGenus="Saprinus">Saprinus (Saprinus) rarus</taxonomicName>
sp. n. head, dorsal view 530 antennal club, ventral view 531 mentum, ventral view 532 propygidium + pygidium 533 prosternum 534 mesoventrite 535 lateral disc of metaventrite + metepisternum 536 protibia, ventral view 537 metatibia, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="153" lastPageNumber="154" pageId="149" pageNumber="150" type="description">
<paragraph pageId="149" pageNumber="150">Description.</paragraph>
<paragraph pageId="149" pageNumber="150">Body length: PEL: 3.50 mm; APW: 1.50 mm; PPW: 2.50 mm; EL: 2.25 mm; EW: 2.90 mm (only one specimen was measured). Body (Fig. 528) elongate, convex, pronotum distinctly narrower than elytra, pronotum piceous, shining, elytra dark blue with metallic tinge; legs, mouthparts and antennal scape+funicle chestnut brown, antennal club lighter, amber colored, becoming progressively lighter apically.</paragraph>
<paragraph pageId="150" pageNumber="151">
<pageBreakToken pageId="150" pageNumber="151" start="start">Antennal</pageBreakToken>
scape (Fig. 529) strongly triangularly dilated, thickened, punctate, almost asetose (setae worn off?); antennal club (Fig. 530) large, almost circular, slightly depressed dorso-ventrally, entirely covered in dense short sensilla in the female paratype, while in the male holotype the lower third of the antennal club asetose, intermingled with sparse longer erect sensilla; sensory structures of antennal club not examined.
</paragraph>
<paragraph pageId="150" pageNumber="151">Mandibles with rounded outer margin, acutely pointed, sub-apical tooth on inner margin of left mandible not examined; labrum slightly depressed medially, labral setae not examined (worn off?); terminal labial palpomere elongated, its width about one-third its length; mentum (Fig. 531) almost square-shaped, anterior margin medially with prominent notch, surface around it with four long setae, anterior angles each with two shorter setae; lateral margins with a single row of even shorter sparse ramose setae, disc on apical half (roughly) with few scattered setae of various sizes (none of these setae are as long as those in the anterior corners or near median notch), basal half (roughly) asetose; terminal maxillary palpomere elongated, its width about one-third half its length, other mouthparts not examined.</paragraph>
<paragraph pageId="151" pageNumber="152">
<pageBreakToken pageId="151" pageNumber="152" start="start">Clypeus</pageBreakToken>
(Fig. 529) rectangular, flattened, with slight median depression, rounded laterally, anterior angles slightly convex, with dense shallow fine punctures; frontal stria (Fig. 529) complete; frontal disc (Fig. 529) anteriorly flattened, entirely punctate, punctures shallow, separated by about their own to twice their diameter, postero-median part of frontal disc with distinct fovea; eyes flattened, visible from above.
</paragraph>
<paragraph pageId="151" pageNumber="152">Pronotal sides (Fig. 528) widest on basal third, gradually convergent apically, apical angles prominent, marginal pronotal stria complete, carinate laterally, weakened behind head; disc almost entirely punctate, punctures round, deep, medially separated by their own to several times their diameter, laterally creating a band of denser and coarser punctation, between it and lateral pronotal margin a narrow impunctate band present; pronotal hypomeron glabrous; scutellum very small.</paragraph>
<paragraph pageId="151" pageNumber="152">Elytral epipleura finely punctate; marginal epipleural stria thin; marginal elytral stria well impressed, thin, continuous along elytral apex as apical elytral stria; humeral elytral stria joined with inner subhumeral stria creating thus a complimentary dorsal elytral stria; four dorsal elytral striae 1-4 present, first the longest, slightly surpassing elytral half, second and third each slightly shorter, fourth stria the shortest, present as a short fragment on basal elytral sixth; sutural elytral stria abbreviated basally, in the female paratype present as a short fragment on (roughly) basal elytral third; in the male holotype entirely missing. Entire elytral disc very coarsely and densely punctate, punctures separated approximately by their diameter, between them another kind of much finer sparser punctures present, interspaces between punctures imbricate; before apical elytral stria punctation weakens, becomes much finer and sparser.</paragraph>
<paragraph pageId="151" pageNumber="152">Propygidium (Fig. 532) almost completely exposed, covered with dense but shallow punctures separated by less than half their own diameter; punctation of pygidium (Fig. 532) sparser, but still very dense, punctures separated by less than their diameter, becoming sparser and finer near pygidial apex, interspaces imbricate.</paragraph>
<paragraph pageId="151" pageNumber="152">Anterior margin of median portion of prosternum (Fig. 533) straight; marginal prosternal stria inconspicuous; prosternal process between carinal prosternal striae flattened, in sparse fine punctures, laterally with sparse small oval punctures, interspaces imbricate; carinal prosternal striae (Fig. 533) carinate, slightly divergent anteriorly and united in front under a rounded loop; lateral prosternal striae carinate, convergent anteriorly, attaining prosternal process near united apices of carinal prosternal striae.</paragraph>
<paragraph pageId="151" pageNumber="152">Discal marginal mesoventral stria laterally well impressed, medially interrupted (Fig. 534); disc with sparse punctures of various sizes separated by several times their diameter; meso-metaventral suture fine; meso-metaventral sutural stria impressed as a row of large shallow punctures; intercoxal disc of metaventrite medially in female with shallow depression, while in the male this depression is deeper; larger punctures appear mostly along median longitudinal line and behind metacoxae, rest of metaventrite with sparse microscopic punctation. Lateral metaventral stria (Fig. 535) well impressed, almost straight, ending short of metacoxa; lateral disc of metaventrite (Fig. 535) depressed, with round shallow large punctures; metepisternum evenly punctate with even denser punctation; lateral metepisternal stria (Fig. 535) present, deeply impressed, absent on fused metepimeron.</paragraph>
<paragraph pageId="152" pageNumber="153">
<pageBreakToken pageId="152" pageNumber="153" start="start">Intercoxal</pageBreakToken>
disc of first abdominal ventrite completely striate laterally; surface of disc with scattered punctation, punctures becoming sparser and finer medially.
</paragraph>
<paragraph lastPageId="153" lastPageNumber="154" pageId="152" pageNumber="153">
Protibia (Fig. 536) dilated, outer margin with three short triangular teeth topped by short denticle, followed by another short denticle; setae of outer row very short, sparsely paced; setae of median row inconspicuous; protarsal groove very shallow, al 
<pageBreakToken pageId="153" pageNumber="154" start="start">most</pageBreakToken>
non-existent; protibial spur tiny, growing out from anterior protibial margin; anterior protibial margin ventrally with three tiny denticles; anterior protibial stria complete, very fine; outer part of posterior surface of protibia (Fig. 536) knobby; median part of posterior surface glabrous, separated from outer part by a definite fine stria with minuscule setae; posterior protibial stria complete, with regular minuscule setae turning into two minuscule denticles apically; inner margin with single row of sparse microscopic setae.
</paragraph>
<paragraph pageId="153" pageNumber="154">Mesotibia dilated, outer margin with 5 widely spaced short denticles growing in size apically; setae of outer row sparse, minuscule; setae of median row inconspicuous; posterior mesotibial stria not examined; anterior surface of mesotibia convex medially; anterior mesotibial stria complete; mesotibial spur short; claws of apical tarsomere short, less than half its length; metatibia (Fig. 537) basically similar to mesotibia, but denticles of outer margin even sparser than those of mesotibia and metatibial spur longer.</paragraph>
<paragraph pageId="153" pageNumber="154">Male genitalia. Eighth sternite (Figs 538-539) completely fused medially, apically with asetose velum covered with pseudo-pores, apex of 8th sternite with several short setae. Eighth tergite (Fig. 539) basally with deep emargination, apex only slightly emarginate; 8th sternite and tergite fused (Fig. 542). Ninth tergite (Figs 543-544) medially with strong sclerotization, with pseudo-pores. Apex of spiculum gastrale (Fig. 540) strongly sclerotized, basal end outwardly arcuate. Aedeagus with parameres separated on apical half (Fig. 545), curved, apex of parameres with setae (Fig. 546).</paragraph>
<caption pageId="153" pageNumber="154">
<paragraph pageId="153" pageNumber="154">
Figures 538-546. 538 
<taxonomicName class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) rarus" order="Coleoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="rarus" subGenus="Saprinus">Saprinus (Saprinus) rarus</taxonomicName>
sp. n. male terminalia: 8th sternite + 8th tergite, ventral view 539 ditto, dorsal view 540 male terminalia: spiculum gastrale, ventral view 541 ditto, lateral view 542 male terminalia: 8th sternite + 8th tergite, lateral view 543 male terminalia: 9th + 10th tergite, dorsal view 544 ditto, lateral view 545 male terminalia: aedeagus, dorsal view 546 ditto, lateral view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>