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<mods:title id="3388453DE9A4BBB5BBA93859A348D0EE">Scydmaenus Latreille of Australia: Revision of species in subgenera Choleropsis Franz, Kingius Franz, Mascarensia Franz, Parallomicrus Franz, Scottiscydmaenus Franz, and description of Ascydmaenus subgen. n. (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<emphasis id="84E56153FFB04A0627ACB3DBFCDDE782" bold="true" box="[151,770,438,465]" pageId="69" pageNumber="70">
<taxonomicName id="7191C6C2FFB04A0627ACB3DBFD56E782" authority="(King)" authorityName="Jałoszyński" authorityYear="2023" baseAuthorityName="Jałoszyński" baseAuthorityYear="2023" box="[151,649,438,465]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="armitagei" status="comb. nov." subGenus="Tetramelus">
<emphasis id="84E56153FFB04A0627ACB3DBFECCE783" bold="true" box="[151,275,439,465]" italics="true" pageId="69" pageNumber="70">Euconnus</emphasis>
(
<emphasis id="84E56153FFB04A06261FB3DAFE75E783" bold="true" box="[292,426,438,465]" italics="true" pageId="69" pageNumber="70">Tetramelus</emphasis>
)
<emphasis id="84E56153FFB04A062680B3DAFDF0E782" bold="true" box="[443,559,438,464]" italics="true" pageId="69" pageNumber="70">armitagei</emphasis>
(King)
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,
<taxonomicNameLabel id="9FD6DC28FFB04A0625ACB3DAFCDDE782" box="[663,770,438,464]" pageId="69" pageNumber="70" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
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<treatmentCitationGroup id="96819A6FFFB04A0627ACB397FDE3E441" box="[151,572,507,531]" pageId="69" pageNumber="70">
<treatmentCitation id="37309B50FFB04A0627ACB397FDE8E441" author="King, R. L." box="[151,567,507,531]" page="98" pageId="69" pageNumber="70" year="1864">
<taxonomicName id="7191C6C2FFB04A0627ACB397FDE8E441" authority="King, 1864: 98" authorityName="King" authorityPageNumber="98" authorityYear="1864" box="[151,567,507,531]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="armitagei">
<emphasis id="84E56153FFB04A0627ACB397FE49E440" box="[151,406,507,531]" italics="true" pageId="69" pageNumber="70">Heterognathus armitagei</emphasis>
<bibRefCitation id="D200C0B0FFB04A0626A7B397FDE8E441" author="King, R. L." box="[412,567,507,531]" pageId="69" pageNumber="70" pagination="91 - 99" refId="ref51270" refString="King, R. L. (1864) On the Scydmaenides of New South Wales. Transactions of the Entomological Society of New South Wales, 1, 91 - 99, pls. 6 - 7." type="journal article" year="1864">King, 1864: 98</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="B62EBD41FFB04A0627ACB075FD34E463" blockId="69.[151,747,507,598]" box="[151,747,537,561]" pageId="69" pageNumber="70">
<treatmentCitationGroup id="96819A6FFFB04A0627ACB075FD34E463" box="[151,747,537,561]" pageId="69" pageNumber="70">
<taxonomicName id="7191C6C2FFB04A0627ACB075FDE1E463" ID-CoL="4W3PR" authority="(King)" baseAuthorityName="King" baseAuthorityYear="1864" box="[151,574,537,561]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="armitagei" subGenus="Cholerus">
<emphasis id="84E56153FFB04A0627ACB075FECBE463" box="[151,276,537,561]" italics="true" pageId="69" pageNumber="70">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFB04A062618B075FE5FE463" box="[291,384,537,561]" italics="true" pageId="69" pageNumber="70">Cholerus</emphasis>
)
<emphasis id="84E56153FFB04A0626B4B076FE2DE463" box="[399,498,538,561]" italics="true" pageId="69" pageNumber="70">armitagei</emphasis>
(King)
</taxonomicName>
;
<treatmentCitation id="37309B50FFB04A062572B075FD38E463" author="Csiki, E." box="[585,743,537,561]" page="72" pageId="69" pageNumber="70" year="1919">
<bibRefCitation id="D200C0B0FFB04A062572B075FD38E463" author="Csiki, E." box="[585,743,537,561]" pageId="69" pageNumber="70" pagination="70" refId="ref49615" refString="Csiki, E. (1919) Pars 70: Scydmaenidae. In: Schenkling, S. &amp; Junk, W. (Eds.), Coleopterorum Catalogus. Vol. 7. W. Junk, Ber- lin, pp. 1 - 106." type="book chapter" year="1919">Csiki, 1919: 72</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="B62EBD41FFB04A0627ACB057FE9CE404" blockId="69.[151,747,507,598]" box="[151,323,571,598]" pageId="69" pageNumber="70">
(
<figureCitation id="2EAAA1C4FFB04A0627A4B057FEE4E404" box="[159,315,571,598]" captionStart="FIGURES 187188" captionStartId="69.[151,264,1620,1644]" captionTargetBox="[151,1435,802,1597]" captionTargetId="figure-143@69.[151,1435,802,1597]" captionTargetPageId="69" captionText="FIGURES 187188. Syntypes of Euconnus (Tetramelus) armitagei (King), comb. n. (=Heterognathus armitagei). © Natalie Tees (AMS)." figureDoi="http://doi.org/10.5281/zenodo.10144721" httpUri="https://zenodo.org/record/10144721/files/figure.png" pageId="69" pageNumber="70">Figs 187188</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFB04A0627ACB0E8FBFAE558" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="B62EBD41FFB04A0627ACB0E8FBFAE558" blockId="69.[151,1436,643,778]" pageId="69" pageNumber="70">
<emphasis id="84E56153FFB04A0627ACB0E8FBC3E4CC" bold="true" box="[151,1052,643,670]" pageId="69" pageNumber="70">
Type specimens studied as high-quality photographs. Presumable
<typeStatus id="692A03E3FFB04A062497B0E8FBC8E4CC" box="[940,1047,644,670]" pageId="69" pageNumber="70" type="syntype">syntypes</typeStatus>
:
</emphasis>
2 exx. (
<figureCitation id="2EAAA1C4FFB04A062343B0EFFB08E4CC" box="[1144,1239,643,670]" captionStart="FIGURES 187188" captionStartId="69.[151,264,1620,1644]" captionTargetBox="[151,1435,802,1597]" captionTargetId="figure-143@69.[151,1435,802,1597]" captionTargetPageId="69" captionText="FIGURES 187188. Syntypes of Euconnus (Tetramelus) armitagei (King), comb. n. (=Heterognathus armitagei). © Natalie Tees (AMS)." figureDoi="http://doi.org/10.5281/zenodo.10144721" httpUri="https://zenodo.org/record/10144721/files/figure.png" pageId="69" pageNumber="70">Fig. 187</figureCitation>
) mounted on one card, “K23213” [brownish, handwritten], “K 197769” {white, printed],
<materialsCitation id="06F9B71CFFB04A0624E8B0C4FEECE4B4" collectionCode="AMS" pageId="69" pageNumber="70" specimenCount="1" typeStatus="syntype">
<typeStatus id="692A03E3FFB04A0624E7B0C4FBB2E490" box="[988,1133,680,706]" pageId="69" pageNumber="70" type="paratype">PARATYPE</typeStatus>
” [blue, printed] (
<collectionCode id="D0802584FFB04A06220EB0C4FAA8E490" box="[1333,1399,680,706]" pageId="69" pageNumber="70">AMS</collectionCode>
);
<specimenCount id="A09776C8FFB04A0622B5B0C4FF66E4B4" count="1" pageId="69" pageNumber="70" type="generic">1 ex.</specimenCount>
(
<figureCitation id="2EAAA1C4FFB04A0627F3B0A7FEF5E4B4" box="[200,298,715,742]" captionStart="FIGURES 187188" captionStartId="69.[151,264,1620,1644]" captionTargetBox="[151,1435,802,1597]" captionTargetId="figure-143@69.[151,1435,802,1597]" captionTargetPageId="69" captionText="FIGURES 187188. Syntypes of Euconnus (Tetramelus) armitagei (King), comb. n. (=Heterognathus armitagei). © Natalie Tees (AMS)." figureDoi="http://doi.org/10.5281/zenodo.10144721" httpUri="https://zenodo.org/record/10144721/files/figure.png" pageId="69" pageNumber="70">Fig. 188</figureCitation>
)
</materialsCitation>
, with labels illustrated in
<figureCitation id="2EAAA1C4FFB04A06256DB0A7FD68E4B4" box="[598,695,715,742]" captionStart="FIGURES 189205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="69" pageNumber="70">Fig. 205</figureCitation>
: “K23212” [brownish, handwritten], “Het / armitagei” [brownish, handwritten], “K 197770” [white, printed],
<materialsCitation id="06F9B71CFFB04A0625BBB09CFBFFE558" box="[640,1056,751,778]" collectionCode="AMS" pageId="69" pageNumber="70" specimenCount="1" typeStatus="paratype">
<typeStatus id="692A03E3FFB04A0625B2B09CFCFDE558" box="[649,802,752,778]" pageId="69" pageNumber="70" type="holotype">HOLOTYPE</typeStatus>
” [red, printed] (
<collectionCode id="D0802584FFB04A0624EDB09CFBC7E558" box="[982,1048,752,778]" pageId="69" pageNumber="70">AMS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="E2EEEDC9FFB04A0627ACB438FEC5E0C3" ID-DOI="http://doi.org/10.5281/zenodo.10144721" ID-Zenodo-Dep="10144721" httpUri="https://zenodo.org/record/10144721/files/figure.png" pageId="69" pageNumber="70" startId="69.[151,264,1620,1644]" targetBox="[151,1435,802,1597]" targetPageId="69" targetType="figure">
<paragraph id="B62EBD41FFB04A0627ACB438FEC5E0C3" blockId="69.[151,1436,1620,1681]" pageId="69" pageNumber="70">
<emphasis id="84E56153FFB04A0627ACB438FEB0E03E" bold="true" box="[151,367,1620,1644]" pageId="69" pageNumber="70">FIGURES 187188.</emphasis>
Syntypes of
<taxonomicName id="7191C6C2FFB04A0626C3B439FC42E03E" authority="(King)" authorityName="Jałoszyński" authorityYear="2023" baseAuthorityName="Jałoszyński" baseAuthorityYear="2023" box="[504,925,1620,1645]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="armitagei" status="comb. nov." subGenus="Tetramelus">
<emphasis id="84E56153FFB04A0626C3B439FD83E03E" box="[504,604,1621,1644]" italics="true" pageId="69" pageNumber="70">Euconnus</emphasis>
(
<emphasis id="84E56153FFB04A062556B438FD02E03E" box="[621,733,1620,1644]" italics="true" pageId="69" pageNumber="70">Tetramelus</emphasis>
)
<emphasis id="84E56153FFB04A0625D6B439FC8FE03E" box="[749,848,1621,1644]" italics="true" pageId="69" pageNumber="70">armitagei</emphasis>
(King)
</taxonomicName>
,
<taxonomicNameLabel id="9FD6DC28FFB04A062492B438FBDCE03E" box="[937,1027,1620,1644]" pageId="69" pageNumber="70" rank="species">comb. n.</taxonomicNameLabel>
(=
<taxonomicName id="7191C6C2FFB04A06231AB438FAFEE03E" authorityName="King" authorityYear="1864" box="[1057,1313,1620,1644]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="armitagei">
<emphasis id="84E56153FFB04A06231AB438FAFEE03E" box="[1057,1313,1620,1644]" italics="true" pageId="69" pageNumber="70">Heterognathus armitagei</emphasis>
</taxonomicName>
). © Natalie Tees (AMS).
</paragraph>
</caption>
<subSubSection id="FE8BEECAFFB04A0627FCB4D3FAB2E1E3" pageId="69" pageNumber="70" type="discussion">
<paragraph id="B62EBD41FFB04A0627FCB4D3FAB2E1E3" blockId="69.[151,1437,1727,1969]" pageId="69" pageNumber="70">
<emphasis id="84E56153FFB04A0627FCB4D3FEE4E08B" bold="true" box="[199,315,1727,1753]" pageId="69" pageNumber="70">Remarks.</emphasis>
Already
<bibRefCitation id="D200C0B0FFB04A06269AB4D3FDF2E08B" author="King, R. L." box="[417,557,1727,1753]" pageId="69" pageNumber="70" pagination="91 - 99" refId="ref51270" refString="King, R. L. (1864) On the Scydmaenides of New South Wales. Transactions of the Entomological Society of New South Wales, 1, 91 - 99, pls. 6 - 7." type="journal article" year="1864">King (1864)</bibRefCitation>
, who described this species, noted that in contrast to all remaining species that he placed in
<taxonomicName id="7191C6C2FFB04A062612B48FFE11E0AF" authorityName="King" authorityYear="1864" box="[297,462,1763,1789]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="84E56153FFB04A062612B48FFE11E0AF" box="[297,462,1763,1789]" italics="true" pageId="69" pageNumber="70">Heterognathus</emphasis>
</taxonomicName>
, this one has a tetramerous antennal club. He noted: “I place it however in this genus with some hesitation, as I have not yet obtained a specimen for dissection”. In addition, he described “
<emphasis id="84E56153FFB04A06222EB56BFE53E117" italics="true" pageId="69" pageNumber="70">thorace fossula transversa basali</emphasis>
”, i.e., the pronotum with a transverse antebasal groove, which is a character not known in any
<taxonomicName id="7191C6C2FFB04A0627ACB523FEFEE13B" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[151,289,1871,1897]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="84E56153FFB04A0627ACB523FEFEE13B" box="[151,289,1871,1897]" italics="true" pageId="69" pageNumber="70">Scydmaenus</emphasis>
</taxonomicName>
, but commonly found among Stenichnini. If the specimens preserved at AMS are indeed those studied by King, they can be unambiguously placed in
<taxonomicName id="7191C6C2FFB04A062593B518FC72E1DF" authorityName="Motschulsky" authorityYear="1869" box="[680,941,1907,1933]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="subGenus" subGenus="Tetramelus">
<emphasis id="84E56153FFB04A062593B518FCC9E1DF" box="[680,790,1908,1933]" italics="true" pageId="69" pageNumber="70">Euconnus</emphasis>
(
<emphasis id="84E56153FFB04A06241CB51FFC7DE1DF" box="[807,930,1907,1933]" italics="true" pageId="69" pageNumber="70">Tetramelus</emphasis>
)
</taxonomicName>
. If not, a placement in the subgenus is putative. However, the original description allows for excluding this species from
<taxonomicName id="7191C6C2FFB04A0624C4B5FBFB56E1E3" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[1023,1161,1943,1969]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="84E56153FFB04A0624C4B5FBFB56E1E3" box="[1023,1161,1943,1969]" italics="true" pageId="69" pageNumber="70">Scydmaenus</emphasis>
</taxonomicName>
(and
<taxonomicName id="7191C6C2FFB04A0623F2B5FBFABEE1E3" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[1225,1377,1943,1969]" pageId="69" pageNumber="4" rank="subGenus" status="subgen. nov." subGenus="Scydmaenini">Scydmaenini</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>