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<document id="80302693A6D9E6D0D80E4CA980247C1A" ID-DOI="10.1649/0010-065X(2000)054[0511:ROTGHT]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4891355" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622574133316" checkinUser="carolina" docAuthor="Mermudes, José Ricardo M. &amp; Napp, Dilma Solange" docDate="2000" docId="03DFFE4DFFB3FFAAFD8CFACAFC4EFDD8" docLanguage="en" docName="TheColeopBull.54.4.511-519.pdf.imf" docOrigin="The Coleopterists Bulletin 54 (4)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Haenkea TIPPMANN 1953" docType="treatment" docVersion="3" lastPageNumber="513" masterDocId="FFE68635FFB3FFA8FF86FFF7FFE0FFD8" masterDocTitle="Review Of The Genus Haenkea Tippmann (Coleoptera, Cerambycidae, Cerambycinae, Cleomenini)" masterLastPageNumber="519" masterPageNumber="511" pageNumber="511" updateTime="1698952553901" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="ED4F0CE2339AC4DB7B2A01FFAEAE0487">Review Of The Genus Haenkea Tippmann (Coleoptera, Cerambycidae, Cerambycinae, Cleomenini)</mods:title>
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<mods:namePart id="C48B7116A772E653881DE91BEC90FA70">Mermudes, José Ricardo M.</mods:namePart>
<mods:affiliation id="7AD1ADEB4D99E3583E52BB5F9499350C">Departamento de Zoologia Universidade Federal do Parana´, Caixa Postal 19020 81531 ­ 990 Curitiba, Paraná, BRAZIL</mods:affiliation>
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<mods:namePart id="0C7B24BFB4CFE0E9DC8B0669BAEBD22B">Napp, Dilma Solange</mods:namePart>
<mods:affiliation id="79DBD8B9783E9BCDE6A8B926E688E737">Departamento de Zoologia Universidade Federal do Parana´, Caixa Postal 19020 81531 ­ 990 Curitiba, Paraná, BRAZIL</mods:affiliation>
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<mods:title id="4D15E3B95FEFEF9B8499CD95A9B6B17C">The Coleopterists Bulletin</mods:title>
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<mods:date id="950228510176DC769D6BC7D8846752CC">2000</mods:date>
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<mods:number id="C9BE349820B788B26D4D8667B0F5273A">54</mods:number>
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<paragraph id="8BC94F5BFFB3FFA8FD8CFACAFD3BFA89" blockId="0.[522,731,1340,1362]" box="[522,731,1340,1362]" pageId="0" pageNumber="511">
<heading id="D081F837FFB3FFA8FD8CFACAFD3BFA89" box="[522,731,1340,1362]" centered="true" fontSize="9" level="2" pageId="0" pageNumber="511" reason="2">
<taxonomicName id="4C7634D8FFB3FFA8FD8CFACAFD3BFA89" ID-CoL="4SJ9" authority="Tippmann" authorityName="TIPPMANN" authorityYear="1953" box="[522,731,1340,1362]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="511" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB3FFA8FD8CFACAFD87FA8A" box="[522,615,1341,1362]" italics="true" pageId="0" pageNumber="511">Haenkea</emphasis>
Tippmann
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</heading>
</paragraph>
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<subSubSection id="C36C1CD0FFB3FFA8FF55FA93FEA1FA4C" pageId="0" pageNumber="511" type="reference_group">
<paragraph id="8BC94F5BFFB3FFA8FF55FA93FEA1FA4C" blockId="0.[211,1043,1380,1454]" pageId="0" pageNumber="511">
<taxonomicName id="4C7634D8FFB3FFA8FF55FA93FDF5FAA1" ID-CoL="4SJ9" authority="Tippman 1953: 220" authorityName="TIPPMANN" authorityPageNumber="220" authorityYear="1953" box="[211,533,1380,1401]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="511" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB3FFA8FF55FA93FED0FAA1" box="[211,304,1380,1401]" italics="true" pageId="0" pageNumber="511">Haenkea</emphasis>
Tippman 1953: 220
</taxonomicName>
; Monné 1993: 78; Monné and Giesbert 1995: 126.
</paragraph>
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<subSubSection id="C36C1CD0FFB3FFA8FF55FA6EFC55FA76" box="[211,949,1433,1454]" pageId="0" pageNumber="511" type="materials_examined">
<paragraph id="8BC94F5BFFB3FFA8FF55FA6EFC55FA76" blockId="0.[211,1043,1380,1454]" box="[211,949,1433,1454]" pageId="0" pageNumber="511">
<typeStatus id="54CDF1F9FFB3FFA8FF55FA6EFEE8FA76" box="[211,264,1433,1454]" pageId="0" pageNumber="511">Type</typeStatus>
species:
<taxonomicName id="4C7634D8FFB3FFA8FEEAFA6EFD2EFA76" authority="Tippmann 1953" authorityName="Tippmann" authorityYear="1953" box="[364,718,1433,1454]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="511" phylum="Arthropoda" rank="species" species="zishkai">
<emphasis id="B9029349FFB3FFA8FEEAFA6EFDFBFA76" box="[364,539,1433,1454]" italics="true" pageId="0" pageNumber="511">Haenkea zishkai</emphasis>
<bibRefCitation id="EFE732AAFFB3FFA8FDA5FA6EFD2EFA76" author="Tippmann" box="[547,718,1433,1454]" pageId="0" pageNumber="511" pagination="181 - 228" refId="ref4516" refString="Tippmann, F. F. 1953. Studien uber neotropishe longicornier I (Coleoptera, Cerambycidae). Dusenia 4: 181 - 228." type="journal article" year="1953">Tippmann 1953</bibRefCitation>
</taxonomicName>
, original designation.
</paragraph>
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<subSubSection id="C36C1CD0FFB3FFA9FF6CFA35FD52FBCE" lastPageId="1" lastPageNumber="512" pageId="0" pageNumber="511" type="description">
<paragraph id="8BC94F5BFFB3FFA8FF6CFA35FBF3F9ED" blockId="0.[211,1044,1474,1589]" pageId="0" pageNumber="511">
<emphasis id="B9029349FFB3FFA8FF6CFA35FE82FA0D" bold="true" box="[234,354,1474,1493]" pageId="0" pageNumber="511">Description.</emphasis>
Head elongate. Frons as wide as long, oblique, impressed on each side near clypeus, carinate laterally. Genae at least
<emphasis id="B9029349FFB3FFA8FD16FA2EFD40FA34" bold="true" box="[656,672,1497,1516]" pageId="0" pageNumber="511"></emphasis>
as long as lower eye lobes. Eyes very finely granulate, deeply emarginate; lower lobes large, prominent, somewhat extending onto frons; upper lobes well separated, about
<emphasis id="B9029349FFB3FFA8FD0CF9FEFD7AF9C4" bold="true" box="[650,666,1545,1564]" pageId="0" pageNumber="511"></emphasis>
as wide as lower lobes. Clypeus large, about
<emphasis id="B9029349FFB3FFA8FE88F9D6FEFEF9EC" bold="true" box="[270,286,1569,1588]" pageId="0" pageNumber="511"></emphasis>
as long as frons, clypeal suture angulate. Antennal tubercles moderately prom­
</paragraph>
<paragraph id="8BC94F5BFFB3FFA8FF55F99FFD64F972" blockId="0.[211,326,1640,1657]" box="[211,644,1640,1706]" lastBlockId="0.[609,644,1687,1706]" pageId="0" pageNumber="511">
<superScript id="7C03E213FFB3FFA8FF55F99FFF39F9AA" attach="left" box="[211,217,1640,1650]" fontSize="4" pageId="0" pageNumber="511">1</superScript>
Contribution 511
</paragraph>
<paragraph id="8BC94F5BFFB2FFA9FF4DFEE2FD52FBCE" blockId="1.[203,1037,277,1046]" pageId="1" pageNumber="512">
inent, rounded. Mandibles nearly straight, bent only at apex, acute, fringed along inner border. Apical segment of palps cylindrical, attenuate at base and apex. Antennae opaque, 11­segmented, conspicuously serrate, extending to about middle of elytra. Scape short, subcylindrical, feebly enlarged apicad, longer than segment 3, dorsally with a well impressed furrow from base to apex. Segments 311 short; 36 impressed on four sides, 35 subequal, 6 the longest, pyramidal, strongly expanded at outer apex; 710 flattened, subequal or slightly longer than 35. Prothorax longer than broad, subcylindrical. Prosternum almost impunctate, subglabrous, inconspicuously transversely rugose; sexual punctation formed by a single coarse, deep, rounded puncture on each side at apical third (
<figureCitation id="134D53DEFFB2FFA9FE81FE1AFE86FDD8" box="[263,358,493,512]" captionStart="Figs" captionStartId="4.[234,277,1658,1677]" captionTargetBox="[227,1027,272,1637]" captionTargetId="figure-10@4.[227,1027,272,1637]" captionTargetPageId="4" captionText="Figs. 45. Haenkea zischkai, male. 4) prosternum, 453; 5) detail of the sexual puncture of the prosternum, 8103; sp 5 sexual puncture." figureDoi="http://doi.org/10.5281/zenodo.4891359" httpUri="https://zenodo.org/record/4891359/files/figure.png" pageId="1" pageNumber="512">Figs. 45</figureCitation>
); intercoxal process declivous, narrowed between procoxae, expanded apically. Anterior coxal cavities rounded at sides, closed behind. Mesosternum with intercoxal process prominent, vertical in front, 1.5 times as wide as mesocoxa, parallelsided, truncate at apex. Intermediate coxal cavities closed to epimera. Mesepisterna somewhat swollen. Metepisterna narrow, parallel­sided. Scutellum small, subtriangular. Elytra relatively short and broad, strongly flattened, somewhat widened behind middle. Each elytron with two costae; one strong, extending from humerus to apex above vertical epipleuron, one weakly prominent, sinuous, extending from inside humerus to apical third; punctures coarse, arranged in four well separated rows on apical
<emphasis id="B9029349FFB2FFA9FCF3FD59FC65FD19" bold="true" box="[885,901,686,705]" pageId="1" pageNumber="512"></emphasis>
, two between humeral and discal costae and two between suture and costae; surface opaque, densely clothed with pale, silky, appressed pubescence, neither obscuring punctures nor costae; suture and costae sparsely asperate punctate, each puncture with a long, erect, pale hair. Epipleuron longitudinally impressed with a prominent, longitudinal carina, strongly asperate punctate, each puncture bearing a moderately short, stiff seta. Elytral apices barely declivous, conjointly rounded truncate. Hind legs almost twice as long as prolegs. Femora pedunculate, suddenly clavate at apical third. Peduncles bicarinate and sulcate, sparsely, slightly asperate punctate. Meso­ and metafemora acutely bidentate at apices. Metafemora exceeding elytral apices by about club length. Tibiae bicarinate and sulcate on both sides, somewhat quadrangular, the four edges sparsely asperate punctate, with long, sparse hairs. Tibial spurs short, the inner longer. First segment of metatarsi about
<emphasis id="B9029349FFB2FFA9FC7DFC4EFBEBFC14" bold="true" box="[1019,1035,953,972]" pageId="1" pageNumber="512"></emphasis>
longer than the following two combined. Abdomen somewhat oval. First urosternite at least a little longer than the following two combined, intercoxal process broad, rounded apically. Fifth urosternite rounded truncate at apex.
</paragraph>
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<subSubSection id="C36C1CD0FFB2FFAAFF63FBC7FC4EFDD8" lastPageId="2" lastPageNumber="513" pageId="1" pageNumber="512" type="discussion">
<paragraph id="8BC94F5BFFB2FFA9FF63FBC7FEBBF9F2" blockId="1.[203,1035,1072,1659]" pageId="1" pageNumber="512">
<emphasis id="B9029349FFB2FFA9FF63FBC7FEB1FB9D" bold="true" box="[229,337,1072,1093]" pageId="1" pageNumber="512">Remarks.</emphasis>
Among the neotropical genera of Cleomenini with short, serrate antennae and cylindrical, elongate prothorax,
<taxonomicName id="4C7634D8FFB2FFA9FD40FBBCFCC3FBB8" authorityName="TIPPMANN" authorityYear="1953" box="[710,803,1099,1120]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB2FFA9FD40FBBCFCC3FBB8" box="[710,803,1099,1120]" italics="true" pageId="1" pageNumber="512">Haenkea</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C7634D8FFB2FFA9FF4DFB91FCEAFBA3" authority="Chevrolat (1859)" authorityName="Chevrolat" authorityYear="1859" box="[203,778,1126,1147]" class="Insecta" family="Cerambycidae" genus="Dihammaphora" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB2FFA9FF4DFB91FE91FBA3" box="[203,369,1126,1147]" italics="true" pageId="1" pageNumber="512">Dihammaphora</emphasis>
<bibRefCitation id="EFE732AAFFB2FFA9FEFAFB91FDD4FBA3" author="Chevrolat" box="[380,564,1126,1147]" pageId="1" pageNumber="512" pagination="50 - 54" refId="ref4196" refString="Chevrolat, L. A. 1859. Description d'un genre noveau etabli aux depens de plusieurs especes de Rhopalophora de Dejean. Arcana Naturae 1: 50 - 54." type="journal article" year="1859">Chevrolat (1859)</bibRefCitation>
(Mermudes, 1999)
</taxonomicName>
. At present,
<taxonomicName id="4C7634D8FFB2FFA9FC1DFB91FEE9FB4E" authorityName="Chevrolat" authorityYear="1859" class="Insecta" family="Cerambycidae" genus="Dihammaphora" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB2FFA9FC1DFB91FEE9FB4E" italics="true" pageId="1" pageNumber="512">Dihammaphora</emphasis>
</taxonomicName>
includes 37 species (Monné 1993; Chemsak and Noguera 1993;
<bibRefCitation id="EFE732AAFFB2FFA9FC5EFB76FEAFFB69" author="Mermudes" pageId="1" pageNumber="512" pagination="97 - 100" refId="ref4359" refString="Mermudes, J. R. M. 1998. Nova especie de Dihammaphora Chevrolat, 1859 da Colombia (Coleoptera, Cerambycidae, Cerambycinae, Cleomenini). Iheringia, Serie Zoologia 85: 97 - 100." type="journal article" year="1998">Mermudes 1998</bibRefCitation>
) and it has never been revised. The main characters that distinguish
<taxonomicName id="4C7634D8FFB2FFA9FE88FB40FE8BFB14" authorityName="TIPPMANN" authorityYear="1953" box="[270,363,1207,1228]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB2FFA9FE88FB40FE8BFB14" box="[270,363,1207,1228]" italics="true" pageId="1" pageNumber="512">Haenkea</emphasis>
</taxonomicName>
from
<taxonomicName id="4C7634D8FFB2FFA9FE35FB40FDABFB14" authorityName="Chevrolat" authorityYear="1859" box="[435,587,1207,1228]" class="Insecta" family="Cerambycidae" genus="Diammaphora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB2FFA9FE35FB40FDABFB14" box="[435,587,1207,1228]" italics="true" pageId="1" pageNumber="512">Diammaphora</emphasis>
</taxonomicName>
are the sixth antennal segment with pyramidal shape, longer than the remainder segments, and the sexual punctation formed by a single, coarse puncture on each side of the prosternum, features probably unique in
<emphasis id="B9029349FFB2FFA9FE22FAFFFDE8FAC5" box="[420,520,1288,1309]" italics="true" pageId="1" pageNumber="512">
<taxonomicName id="4C7634D8FFB2FFA9FE22FAFFFDE4FAC5" authorityName="TIPPMANN" authorityYear="1953" box="[420,516,1288,1309]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">Haenkea</taxonomicName>
.
</emphasis>
In addition, the head is elongate, the antennal segments 35 and 811 are short and similarly sized, the metafemora exceed the elytral apices by the club length and the lateral margins of the elytra are strongly asperate.
<taxonomicName id="4C7634D8FFB2FFA9FE13FAAFFDDBFAB5" authorityName="Chevrolat" authorityYear="1859" box="[405,571,1368,1389]" class="Insecta" family="Cerambycidae" genus="Dihammaphora" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB2FFA9FE13FAAFFDDBFAB5" box="[405,571,1368,1389]" italics="true" pageId="1" pageNumber="512">Dihammaphora</emphasis>
</taxonomicName>
includes small­sized species presenting a short head, with the sixth antennal segment subserrate and the sexual punctation, if present, formed by a large, micropunctate patch on each side of the prosternum. In addition, in
<taxonomicName id="4C7634D8FFB2FFA9FE6AFA5EFD72FA66" authorityName="Chevrolat" authorityYear="1859" box="[492,658,1449,1470]" class="Insecta" family="Cerambycidae" genus="Dihammaphora" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB2FFA9FE6AFA5EFD72FA66" box="[492,658,1449,1470]" italics="true" pageId="1" pageNumber="512">Dihammaphora</emphasis>
</taxonomicName>
the antennal segments 35 are subcylindrical and longer than the following ones, which are short and somewhat swollen; the femora are not dentate apically, the hind clubs at most barely exceeding the elytral apices; the lateral margins of the elytra are weakly asperate or not.
</paragraph>
<paragraph id="8BC94F5BFFB2FFAAFF63F9C7FC40FE3D" blockId="1.[203,1035,1072,1659]" lastBlockId="2.[211,1043,277,512]" lastPageId="2" lastPageNumber="513" pageId="1" pageNumber="512">
Presently, the species of
<taxonomicName id="4C7634D8FFB2FFA9FE6FF9C7FDA6F99D" authorityName="TIPPMANN" authorityYear="1953" box="[489,582,1584,1605]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B9029349FFB2FFA9FE6FF9C7FDA6F99D" box="[489,582,1584,1605]" italics="true" pageId="1" pageNumber="512">Haenkea</emphasis>
</taxonomicName>
show a disjunct distribution (
<figureCitation id="134D53DEFFB2FFA9FCFBF9C7FC5DF99D" box="[893,957,1584,1605]" captionStart="Fig" captionStartId="5.[226,260,1116,1135]" captionTargetBox="[207,1031,274,1093]" captionTargetId="figure-12@5.[204,1034,272,1096]" captionTargetPageId="5" captionText="Fig. 6. Distribution records of Haenkea zischkai (triangles), H. thoracica (squares) and H. atra (circles)." figureDoi="http://doi.org/10.5281/zenodo.4891363" httpUri="https://zenodo.org/record/4891363/files/figure.png" pageId="1" pageNumber="512">Fig. 6</figureCitation>
), being associated with tropical rain forests in South and Central
<taxonomicName id="4C7634D8FFB2FFA9FCDEF9BDFEEBF9A2" authority="(Hueck 1972)" baseAuthorityName="Hueck" baseAuthorityYear="1972" class="Insecta" family="Cerambycidae" genus="America" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="genus">America (Hueck 1972)</taxonomicName>
.
<taxonomicName id="4C7634D8FFB2FFA9FE93F991FE2DF9A3" authorityName="Tippmann" authorityYear="1953" box="[277,461,1638,1659]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="512" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis id="B9029349FFB2FFA9FE93F991FE2DF9A3" box="[277,461,1638,1659]" italics="true" pageId="1" pageNumber="512">Haenkea zischkai</emphasis>
</taxonomicName>
occurs along the Amazon basin, from near the eastern slopes of the Andes in
<collectingCountry id="F3610FCBFFB1FFAAFE57FEE2FDFEFEF2" box="[465,542,277,298]" name="Bolivia" pageId="2" pageNumber="513">Bolivia</collectingCountry>
and
<collectingCountry id="F3610FCBFFB1FFAAFDD1FEE2FD6AFEF2" box="[599,650,277,298]" name="Peru" pageId="2" pageNumber="513">Peru</collectingCountry>
.
<emphasis id="B9029349FFB1FFAAFD11FEE1FC89FEF3" box="[663,873,278,299]" italics="true" pageId="2" pageNumber="513">
<taxonomicName id="4C7634D8FFB1FFAAFD11FEE1FC85FEF3" authorityName="Pronotum" baseAuthorityName="Chevrolat" baseAuthorityYear="1855" box="[663,869,278,299]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="513" phylum="Arthropoda" rank="species" species="thoracica">Haenkea thoracica</taxonomicName>
,
</emphasis>
described from
<collectingCountry id="F3610FCBFFB1FFAAFF55FEC7FEA2FE9D" box="[211,322,304,325]" name="Venezuela" pageId="2" pageNumber="513">Venezuela</collectingCountry>
, is distributed northwards through
<collectingCountry id="F3610FCBFFB1FFAAFD36FEC7FCE3FE9D" box="[688,771,304,325]" name="Panama" pageId="2" pageNumber="513">Panama</collectingCountry>
and
<collectingCountry id="F3610FCBFFB1FFAAFCB1FEC7FC4CFE9D" box="[823,940,304,325]" name="Costa Rica" pageId="2" pageNumber="513">Costa Rica</collectingCountry>
, probably associated with Caribbean rain forest.
<emphasis id="B9029349FFB1FFAAFDE8FEBCFCE5FEB8" box="[622,773,331,352]" italics="true" pageId="2" pageNumber="513">
<taxonomicName id="4C7634D8FFB1FFAAFDE8FEBCFCE1FEB8" baseAuthorityName="Chevrolat" baseAuthorityYear="1855" box="[622,769,331,352]" class="Insecta" family="Cerambycidae" genus="Haenkea" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="513" phylum="Arthropoda" rank="species" species="atra">Haenkea atra</taxonomicName>
,
</emphasis>
on the other hand, is the only representative occurring in the Atlantic forest, in eastern
<collectingCountry id="F3610FCBFFB1FFAAFC14FE92FC34FEA2" box="[914,980,357,378]" name="Brazil" pageId="2" pageNumber="513">Brazil</collectingCountry>
, with some examples having been found in subtropical forest of
<collectingRegion id="49B281B9FFB1FFAAFCC2FE77FC6BFE4D" box="[836,907,384,405]" country="Brazil" name="Parana" pageId="2" pageNumber="513">Paraná</collectingRegion>
River basin, which is an extension of the Atlantic forest. This southeast/northwest disjunction in the Neotropical region is congruent to some extent with those found by Amorim and Pires (1996),
<bibRefCitation id="EFE732AAFFB1FFAAFD9EFE27FD29FE3D" author="Camargo" box="[536,713,464,485]" pageId="2" pageNumber="513" pagination="107 - 121" refId="ref4063" refString="Camargo, J. M. 1966. Meliponini Neotropicais (Apinae, Apidae, Hymenoptera): Biogeografia historica [pp. 107 - 121]. In: Anais do II Encontro sobre Abelhas. Universidade de Sao Paulo, Ribeirao Preto. 351 pp." type="book chapter" year="1966">Camargo (1966)</bibRefCitation>
, and
<bibRefCitation id="EFE732AAFFB1FFAAFC83FE27FC7CFE3D" author="Grazia" box="[773,924,464,485]" pageId="2" pageNumber="513" pagination="43 - 48" refId="ref4250" refString="Grazia, J. 1997. Cladistic analysis of the Evoplitus genus group of Pentatomini (Heteroptera: Pentatomidae). Journal of Comparative Biology 2: 43 - 48." type="journal article" year="1997">Grazia (1997)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC94F5BFFB1FFAAFF6BFE1CFC4EFDD8" blockId="2.[211,1043,277,512]" box="[237,942,491,512]" pageId="2" pageNumber="513">Immatures are unknown and larval hosts have not been recorded.</paragraph>
</subSubSection>
</treatment>
</document>