437 lines
58 KiB
XML
437 lines
58 KiB
XML
<document id="6F4D1192F8D6444E0D40785CAEFB2024" ID-DOI="10.5281/zenodo.12808306" ID-ISSN="1942-1354" ID-Zenodo-Dep="12808306" ID-ZooBank="50F22DF8-FE88-4A8F-806D-1046E47C617A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721837329056" checkinUser="felipe" docAuthor="Huchet, Jean-Bernard" docDate="2021" docId="03DF99647918293973E1426BFAFCD5BB" docLanguage="en" docName="InsectaMundi.2021.0893.1-12.pdf" docOrigin="Insecta Mundi 2021 (893)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Nothochodaeus huxleyi Huchet 2021, new species" docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="FFE6E11C791F293273214314FFCCD423" masterDocTitle="Two new species of Nothochodaeus Nikolajev, 2005 from Palawan Island, Philippines (Coleoptera: Scarabaeoidea: Ochodaeidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="6" updateTime="1721840243770" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="4BD92A49B4F101C33F7F8E5C3B12CAC0">Two new species of Nothochodaeus Nikolajev, 2005 from Palawan Island, Philippines (Coleoptera: Scarabaeoidea: Ochodaeidae)</mods:title>
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<mods:name id="3BBF55E83BCD52D4141A104EE0CD204F" type="personal">
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<mods:roleTerm id="1DADBDFF6CDDB1BB6FED74C5802959F7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="95ECE963E27652FC3588C1EB59C8A1B8">Huchet, Jean-Bernard</mods:namePart>
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<mods:affiliation id="1F9A3A17684EEEEF90F239B047158673">Institut Systématique, Evolution, Biodiversité (ISYEB), CNRS, MNHN, SU, EPHE, UA. Muséum national d’Histoire naturelle, CP 50, 45 rue Buffon, F- 75005 Paris, France.</mods:affiliation>
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<mods:typeOfResource id="754B0912362DD4FB4BBBE2326A99674C">text</mods:typeOfResource>
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<mods:title id="A623639A49C5E8918ACD4B5B538B49F6">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="EC327577EFC502D3E2C74F549512AC98">
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<mods:date id="9A8D0FCDF5B2EA61084FA8CADBA46955">2021</mods:date>
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<mods:number id="AA10DFD2269633938614ECFB60F9498A">2021-11-19</mods:number>
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<mods:number id="A3D62BB0147B6816BE04ACB89409BFCE">2021</mods:number>
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<mods:detail id="B31CF9EFC148F67C97161B5D84B9D62A" type="issue">
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<mods:number id="DD61D7D8FC81DCDA7012F6040C3A3074">893</mods:number>
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<mods:start id="AA31192CA3917E963D5DBE627ADA8110">1</mods:start>
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<mods:classification id="463B95533A103876CF6AAD9DDA4C24C8">journal article</mods:classification>
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<mods:identifier id="E0B806E94AAD21F5A1315C5B2759D552" type="DOI">10.5281/zenodo.12808306</mods:identifier>
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<mods:identifier id="9037784AC57A9BA678686E170E7713C2" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="31894392650FFCBA109F9F816951BE3D" type="Zenodo-Dep">12808306</mods:identifier>
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<mods:identifier id="30C2584374FFC8C8B993EB99AB56BEB5" type="ZooBank">50F22DF8-FE88-4A8F-806D-1046E47C617A</mods:identifier>
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<treatment id="03DF99647918293973E1426BFAFCD5BB" ID-DOI="http://doi.org/10.5281/zenodo.12808326" ID-Zenodo-Dep="12808326" LSID="urn:lsid:plazi:treatment:03DF99647918293973E1426BFAFCD5BB" httpUri="http://treatment.plazi.org/id/03DF99647918293973E1426BFAFCD5BB" lastPageId="11" lastPageNumber="10" pageId="7" pageNumber="6">
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<subSubSection id="C36C7BF97918293573E1426BFBFFD5BF" box="[192,1075,383,412]" pageId="7" pageNumber="6" type="nomenclature">
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<paragraph id="8BC928727918293573E1426BFBFFD5BF" blockId="7.[192,1075,383,412]" box="[192,1075,383,412]" pageId="7" pageNumber="6">
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<heading id="D0819F1E7918293573E1426BFBFFD5BF" box="[192,1075,383,412]" fontSize="12" level="2" pageId="7" pageNumber="6" reason="2">
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<emphasis id="B902F4607918293573E1426BFBFFD5BF" bold="true" box="[192,1075,383,412]" pageId="7" pageNumber="6">
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<taxonomicName id="4C7653F17918293573E1426BFD86D5BF" ID-CoL="8SSNG" authority="Huchet" authorityName="Huchet" authorityYear="2021" box="[192,586,383,412]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="huxleyi" status="sp. nov.">
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<emphasis id="B902F4607918293573E1426BFE29D5BF" bold="true" box="[192,485,383,412]" italics="true" pageId="7" pageNumber="6">Nothochodaeus huxleyi</emphasis>
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Huchet
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</taxonomicName>
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,
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<taxonomicNameLabel id="A231491B7918293571784294FD22D5BF" box="[601,750,383,412]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="134D34F77918293571DC426BFC6ED5BF" box="[765,930,383,412]" captionStart-0="Figure 6" captionStart-1="Figures 7–8" captionStart-2="Figures 9–10" captionStart-3="Figures 11–13" captionStartId-0="7.[193,269,1865,1891]" captionStartId-1="8.[192,279,1866,1892]" captionStartId-2="10.[192,279,1866,1892]" captionStartId-3="11.[192,278,1153,1179]" captionTargetBox-0="[549,1084,1335,1846]" captionTargetBox-1="[289,1345,219,1833]" captionTargetBox-2="[357,1362,226,1824]" captionTargetBox-3="[323,1307,462,1118]" captionTargetId-0="figure-496@7.[549,1084,1335,1848]" captionTargetId-1="figure-30@8.[218,1414,213,1836]" captionTargetId-2="figure-10@10.[330,1379,213,1827]" captionTargetId-3="figure-86@11.[305,1327,457,1124]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="Figure 6.Nothochodaeus. huxleyi Huchet, new species, male holotype, head in frontal view showing the U-shaped frontoclypeal carina (Photo J.-B. Huchet, MNHN)." captionText-1="Figures 7–8. Nothochodaeus huxleyi Huchet, new species, male holotype. 7) Habitus, dorsal view. 8) Habitus, latero-dorsal view (Photos J.-B. Huchet, MNHN)." captionText-2="Figures 9–10. Nothochodaeus huxleyi Huchet, new species, female allotype. 9) Habitus, dorsal view. 10) Habitus, latero-dorsal view (Photos J.-B. Huchet, MNHN)." captionText-3="Figures 11–13. Nothochodaeus sp., male genitalia. 11) N. huxleyi Huchet, new species, aedeagus in latero-dorsal view. 12) N. borneensis Ochi, Kon and Masumoto, 2013, aedeagus in latero-dorsal view (insets illustrating a closeup of left paramere). 13) N. huxleyi Huchet, new species, genital segment (urite IX) (Photos J.-B. Huchet, MNHN)." figureDoi-0="http://doi.org/10.5281/zenodo.12808318" figureDoi-1="http://doi.org/10.5281/zenodo.12808322" figureDoi-2="http://doi.org/10.5281/zenodo.12808324" figureDoi-3="http://doi.org/10.5281/zenodo.12808328" httpUri-0="https://zenodo.org/record/12808318/files/figure.png" httpUri-1="https://zenodo.org/record/12808322/files/figure.png" httpUri-2="https://zenodo.org/record/12808324/files/figure.png" httpUri-3="https://zenodo.org/record/12808328/files/figure.png" pageId="7" pageNumber="6">Fig. 6–11, 13</figureCitation>
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,
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<figureCitation id="134D34F779182935708E426BFC30D5BF" box="[943,1020,383,412]" captionStart="Figures 16–19" captionStartId="12.[192,279,975,1001]" captionTargetBox="[336,1296,216,944]" captionTargetId="figure-90@12.[336,1296,216,944]" captionTargetPageId="12" captionText="Figures 16–19. Nothochodaeus sp., clypeofrontal carina and protibia, comparizon. 16) N. huxleyi Huchet, n. sp., holotype male, clypeofrontal carina, frontal view.17) N. borneensis, male, clypeofrontal carina, frontal view.18) N. huxleyi Huchet, n. sp., holotype male, right protibia, dorsal view. 19) N. borneensis, male, right protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12808332" httpUri="https://zenodo.org/record/12808332/files/figure.png" pageId="7" pageNumber="6">16, 18</figureCitation>
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,
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<figureCitation id="134D34F779182935772B426BFBE7D5BF" box="[1034,1067,383,412]" captionStart="Figure 20" captionStartId="12.[192,268,1866,1892]" captionTargetBox="[336,1295,1121,1837]" captionTargetId="figure-92@12.[336,1295,1121,1837]" captionTargetPageId="12" captionText="Figure 20.Distribution map of Nothochodaeus minotaurus Huchet, new species (red dots), and N. huxleyi Huchet, new species (yellow dots) on Palawan Island (the type locality indicated with a star)." figureDoi="http://doi.org/10.5281/zenodo.12808334" httpUri="https://zenodo.org/record/12808334/files/figure.png" pageId="7" pageNumber="6">20</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C7BF97918293573E142B8FB68D7A2" pageId="7" pageNumber="6" type="materials_examined">
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<paragraph id="8BC928727918293573E142B8FEA2D653" blockId="7.[191,1441,427,897]" pageId="7" pageNumber="6">
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<emphasis id="B902F4607918293573E142B8FEA7D5E6" bold="true" box="[192,363,427,454]" pageId="7" pageNumber="6">Type material.</emphasis>
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<materialsCitation id="3B1E222F79182935725542BFFEA2D653" collectingDate="2016-05" collectionCode="CJBH, TYPE" collectorName="S. J. Soriano & Coll. J. - B. & Huchet" country="Philippines" determinerName="J. - B. Huchet" location="Roxas" pageId="7" pageNumber="6" specimenCount="2" stateProvince="Palawan" typeStatus="holotype">
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<typeStatus id="54CD96D079182935725542BFFE10D5E6" box="[372,476,427,453]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="ED67B0B779182935710D42B8FDB8D5E5" box="[556,628,428,454]" pageId="7" pageNumber="6">CJBH</collectionCode>
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), labeled: a) rectangular, white paper, printed: “
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<collectingCountry id="F36168E27918293577B442B8FAEDD5E5" box="[1173,1313,428,454]" name="Philippines" pageId="7" pageNumber="6">PHILIPPINES</collectingCountry>
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,
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<collectingRegion id="49B2E69079182935761042BFFA42D5E6" box="[1329,1422,427,453]" country="Philippines" name="Palawan" pageId="7" pageNumber="6">Palawan</collectingRegion>
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/
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<location id="8EA97EA97918293573E142DAFEC9D5CB" LSID="urn:lsid:plazi:treatment:03DF99647918293973E1426BFAFCD5BB:8EA97EA97918293573E142DAFEC9D5CB" box="[192,261,462,488]" country="Philippines" name="Roxas" pageId="7" pageNumber="6" stateProvince="Palawan">Roxas</location>
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,
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<date id="FFC80EB279182935723D42DAFEBED5C4" box="[284,370,461,488]" pageId="7" pageNumber="6" value="2016-05">
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||
<collectingDate id="EF8CF75A79182935723D42DAFEBED5C4" box="[284,370,461,488]" pageId="7" pageNumber="6" value="2016-05">V. 2016</collectingDate>
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||
</date>
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||
/
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<collectingMethod id="523750657918293572AF42D9FD54D5CB" box="[398,664,461,488]" pageId="7" pageNumber="6">Flight interception trap</collectingMethod>
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/
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<collectorName id="26834DA479182935719542DAFC8CD5C4" box="[692,832,461,488]" pageId="7" pageNumber="6">S. J. Soriano</collectorName>
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lgt”; b) rectangular, white paper, printed: “
|
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<collectorName id="26834DA479182935761342D9FED9D629" pageId="7" pageNumber="6">Coll. J.-B. Huchet</collectorName>
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”; c) rectangular, red paper: “
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||
<collectionCode id="ED67B0B779182935717442E4FD52D629" box="[597,670,496,522]" pageId="7" pageNumber="6">TYPE</collectionCode>
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”; d) rectangular, red paper, printed: “
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<taxonomicName id="4C7653F179182935771F42E4FB2AD629" authorityName="Nikolajev" authorityYear="2005" box="[1086,1254,496,522]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B902F46079182935771F42E4FB2AD629" box="[1086,1254,496,522]" italics="true" pageId="7" pageNumber="6">Nothochodaeus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C7653F17918293577DF42E4FA83D629" authorityName="Huchet" authorityYear="2021" box="[1278,1359,496,522]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="huxleyi" status="sp. nov.">
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<emphasis id="B902F4607918293577DF42E4FA83D629" box="[1278,1359,496,522]" italics="true" pageId="7" pageNumber="6">huxleyi</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A231491B79182935767442E4FA5CD629" box="[1365,1424,496,522]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
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/
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<typeStatus id="54CD96D07918293573E04106FE9ED60F" box="[193,338,530,556]" pageId="7" pageNumber="6" type="holotype">HOLOTYPE</typeStatus>
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/
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<determinerName id="7E15C4B67918293572474106FE21D60F" box="[358,493,530,556]" pageId="7" pageNumber="6">J.-B. Huchet</determinerName>
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det. 2021”. Genitalia stored in a small glycerol vial, pinned under the specimen. Note: currently in the author collection, this specimen is intended to be deposited at the Muséum National d’Histoire naturelle, Paris.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BC92872791829357221416CFB68D7A2" blockId="7.[191,1441,427,897]" pageId="7" pageNumber="6">
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<materialsCitation id="3B1E222F791829357221416CFBC3D697" collectingDate="2016-06" collectionCode="CJBH, VI" collectorName="S. J. Soriano & Coll. J. - B. & Huchet" country="Philippines" location="Roxas" pageId="7" pageNumber="6" specimenCount="1" specimenCount-female="1" stateProvince="Palawan" typeStatus="allotype">
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||
<specimenCount id="9D70E3FB791829357221416CFE4CD6B1" box="[256,384,632,658]" count="1" pageId="7" pageNumber="6" type="female">One female</specimenCount>
|
||
<typeStatus id="54CD96D07918293572A7416CFE13D6B1" box="[390,479,632,658]" pageId="7" pageNumber="6" type="allotype">allotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED67B0B77918293572CD416CFDFED6B1" box="[492,562,632,658]" pageId="7" pageNumber="6">CJBH</collectionCode>
|
||
) labeled: a) “
|
||
<collectingCountry id="F36168E27918293571E0416CFC81D6B1" box="[705,845,632,658]" name="Philippines" pageId="7" pageNumber="6">PHILIPPINES</collectingCountry>
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||
,
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||
<collectingRegion id="49B2E690791829357078416CFC7AD6B1" box="[857,950,632,658]" country="Philippines" name="Palawan" pageId="7" pageNumber="6">Palawan</collectingRegion>
|
||
/
|
||
<location id="8EA97EA97918293570EA416DFBDCD6B0" LSID="urn:lsid:plazi:treatment:03DF99647918293973E1426BFAFCD5BB:8EA97EA97918293570EA416DFBDCD6B0" box="[971,1040,633,659]" country="Philippines" name="Roxas" pageId="7" pageNumber="6" stateProvince="Palawan">Roxas</location>
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||
,
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||
<date id="FFC80EB279182935773B416DFBB5D6B1" box="[1050,1145,632,659]" pageId="7" pageNumber="6" value="2016-06">
|
||
<collectingDate id="EF8CF75A79182935773B416DFBB5D6B1" box="[1050,1145,632,659]" pageId="7" pageNumber="6" value="2016-06">
|
||
<collectionCode id="ED67B0B779182935773B416DFBF6D6B0" box="[1050,1082,633,659]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="6">VI</collectionCode>
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||
. 2016
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||
</collectingDate>
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</date>
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||
/
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||
<collectingMethod id="523750657918293577AF416CFA5DD6B0" box="[1166,1425,632,659]" pageId="7" pageNumber="6">Flight interception trap</collectingMethod>
|
||
/
|
||
<collectorName id="26834DA479182935739E418EFE80D697" box="[191,332,666,693]" pageId="7" pageNumber="6">S. J. Soriano</collectorName>
|
||
lgt” b) rectangular, white paper, printed: “
|
||
<collectorName id="26834DA4791829357018418EFBC9D697" box="[825,1029,666,693]" pageId="7" pageNumber="6">Coll. J.-B. Huchet</collectorName>
|
||
”
|
||
</materialsCitation>
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||
;
|
||
<materialsCitation id="3B1E222F79182935773C418EFE94D739" collectingDate="2021-09" collectionCode="CJBH" collectorName="Coll. J. - B. & Huchet" country="Philippines" determinerName="J. - B. Huchet" location="Tinitian" municipality="Roxas" pageId="7" pageNumber="6" specimenCount="2" specimenCount-female="1" stateProvince="Palawan" typeStatus="paratype">
|
||
c) rectangular, red paper, printed: “
|
||
<taxonomicName id="4C7653F17918293573EA41A8FEBFD6F5" authorityName="Nikolajev" authorityYear="2005" box="[203,371,700,726]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902F4607918293573EA41A8FEBFD6F5" box="[203,371,700,726]" italics="true" pageId="7" pageNumber="6">Nothochodaeus</emphasis>
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||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7653F17918293572AA41A8FE10D6F5" authorityName="Huchet" authorityYear="2021" box="[395,476,700,726]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="huxleyi" status="sp. nov.">
|
||
<emphasis id="B902F4607918293572AA41A8FE10D6F5" box="[395,476,700,726]" italics="true" pageId="7" pageNumber="6">huxleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231491B7918293572C241A9FDD3D6F4" box="[483,543,701,727]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
/
|
||
<typeStatus id="54CD96D079182935711741A9FD71D6F4" box="[566,701,701,727]" pageId="7" pageNumber="6" type="paratype">PARATYPE</typeStatus>
|
||
/
|
||
<determinerName id="7E15C4B67918293571F441A9FC91D6F5" box="[725,861,700,727]" pageId="7" pageNumber="6">J.-B. Huchet</determinerName>
|
||
det. 2021”.
|
||
<specimenCount id="9D70E3FB7918293570FE41A8FBADD6F5" box="[991,1121,700,726]" count="1" pageId="7" pageNumber="6" type="female">One female</specimenCount>
|
||
<typeStatus id="54CD96D079182935774941A9FB05D6F4" box="[1128,1225,701,727]" pageId="7" pageNumber="6" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="ED67B0B77918293577F941A8FAD2D6F5" box="[1240,1310,700,726]" pageId="7" pageNumber="6">CJBH</collectionCode>
|
||
) labeled: a) “
|
||
<collectingCountry id="F36168E27918293573EA41CBFE9BD6DA" box="[203,343,735,761]" name="Philippines" pageId="7" pageNumber="6">PHILIPPINES</collectingCountry>
|
||
,
|
||
<collectingRegion id="49B2E69079182935724541CAFE0DD6DB" box="[356,449,734,760]" country="Philippines" name="Palawan" pageId="7" pageNumber="6">Palawan</collectingRegion>
|
||
/
|
||
<collectingMunicipality id="6BADB2087918293572F841CBFDD2D6DA" box="[473,542,735,761]" pageId="7" pageNumber="6">Roxas</collectingMunicipality>
|
||
,
|
||
<location id="8EA97EA979182935710B41CAFD48D6DB" LSID="urn:lsid:plazi:treatment:03DF99647918293973E1426BFAFCD5BB:8EA97EA979182935710B41CAFD48D6DB" box="[554,644,734,760]" country="Philippines" municipality="Roxas" name="Tinitian" pageId="7" pageNumber="6" stateProvince="Palawan">Tinitian</location>
|
||
/
|
||
<date id="FFC80EB27918293571BC41CBFD30D6DA" box="[669,764,735,761]" pageId="7" pageNumber="6" value="2021-09">
|
||
<collectingDate id="EF8CF75A7918293571BC41CBFD30D6DA" box="[669,764,735,761]" pageId="7" pageNumber="6" value="2021-09">IX. 2021</collectingDate>
|
||
</date>
|
||
/ local collector” b) rectangular, white paper, printed: “
|
||
<collectorName id="26834DA479182935764B41CAFE82D739" pageId="7" pageNumber="6">Coll. J.-B. Huchet</collectorName>
|
||
”
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1E222F7918293572474015FB3AD77C" collectionCode="CJBH, VIII" collectorName="Coll. J. - B. & Huchet" country="Philippines" county="Northern" determinerName="J. - B. Huchet" location="Mt. Magara" municipality="Roxas" pageId="7" pageNumber="6" specimenCount="2" specimenCount-female="1" stateProvince="Palawan" typeStatus="paratype">
|
||
c) rectangular, red paper, printed: “
|
||
<taxonomicName id="4C7653F17918293571DA4014FC6FD739" authorityName="Nikolajev" authorityYear="2005" box="[763,931,768,794]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902F4607918293571DA4014FC6FD739" box="[763,931,768,794]" italics="true" pageId="7" pageNumber="6">Nothochodaeus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7653F179182935709E4014FBDCD739" authorityName="Huchet" authorityYear="2021" box="[959,1040,768,794]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="huxleyi" status="sp. nov.">
|
||
<emphasis id="B902F46079182935709E4014FBDCD739" box="[959,1040,768,794]" italics="true" pageId="7" pageNumber="6">huxleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231491B7918293577384015FB9BD738" box="[1049,1111,769,795]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
/
|
||
<typeStatus id="54CD96D07918293577534015FB35D738" box="[1138,1273,769,795]" pageId="7" pageNumber="6" type="paratype">PARATYPE</typeStatus>
|
||
/
|
||
<determinerName id="7E15C4B67918293576374015FA6CD739" box="[1302,1440,768,795]" pageId="7" pageNumber="6">J.-B. Huchet</determinerName>
|
||
det. 2021”.
|
||
<specimenCount id="9D70E3FB7918293572604037FE0AD71E" box="[321,454,803,829]" count="1" pageId="7" pageNumber="6" type="female">One female</specimenCount>
|
||
<typeStatus id="54CD96D07918293572F14037FDFDD71E" box="[464,561,803,829]" pageId="7" pageNumber="6" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="ED67B0B77918293571654037FD46D71E" box="[580,650,803,829]" pageId="7" pageNumber="6">CJBH</collectionCode>
|
||
) labeled: a) “
|
||
<collectingCountry id="F36168E27918293570064037FC7FD71E" box="[807,947,803,829]" name="Philippines" pageId="7" pageNumber="6">PHILIPPINES</collectingCountry>
|
||
,
|
||
<collectingCounty id="62A850FE7918293570E24037FBE1D71E" box="[963,1069,803,829]" pageId="7" pageNumber="6">Northern</collectingCounty>
|
||
<collectingRegion id="49B2E6907918293577194037FB59D71E" box="[1080,1173,803,829]" country="Philippines" name="Palawan" pageId="7" pageNumber="6">Palawan</collectingRegion>
|
||
/
|
||
<collectingMunicipality id="6BADB2087918293577924037FB34D71E" box="[1203,1272,803,829]" pageId="7" pageNumber="6">Roxas</collectingMunicipality>
|
||
,
|
||
<location id="8EA97EA97918293576264037FA40D71E" LSID="urn:lsid:plazi:treatment:03DF99647918293973E1426BFAFCD5BB:8EA97EA97918293576264037FA40D71E" box="[1287,1420,803,829]" country="Philippines" county="Northern" municipality="Roxas" name="Mt. Magara" pageId="7" pageNumber="6" stateProvince="Palawan">Mt. Magara</location>
|
||
/
|
||
<date id="FFC80EB27918293573E14051FE9FD77C" box="[192,339,837,863]" pageId="7" pageNumber="6" value="2021-08-21">
|
||
21-
|
||
<collectionCode id="ED67B0B77918293573C24051FEDBD77C" box="[227,279,837,863]" pageId="7" pageNumber="6">VIII</collectionCode>
|
||
.2021
|
||
</date>
|
||
,
|
||
<collectingMethod id="5237506579182935727F4051FE08D77C" box="[350,452,837,863]" pageId="7" pageNumber="6">light trap</collectingMethod>
|
||
/ local collector”. b) rectangular, white paper, printed: “
|
||
<collectorName id="26834DA47918293577064051FB20D77C" box="[1063,1260,837,863]" pageId="7" pageNumber="6">Coll. J.-B. Huchet</collectorName>
|
||
”
|
||
</materialsCitation>
|
||
; c) rectangular, red paper, printed: “
|
||
<taxonomicName id="4C7653F17918293572814073FD84D7A2" authorityName="Nikolajev" authorityYear="2005" box="[416,584,871,897]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902F4607918293572814073FD84D7A2" box="[416,584,871,897]" italics="true" pageId="7" pageNumber="6">Nothochodaeus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7653F179182935717F4073FD63D7A2" authorityName="Huchet" authorityYear="2021" box="[606,687,871,897]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="huxleyi" status="sp. nov.">
|
||
<emphasis id="B902F46079182935717F4073FD63D7A2" box="[606,687,871,897]" italics="true" pageId="7" pageNumber="6">huxleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231491B7918293571944073FD3CD7A2" box="[693,752,871,897]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
/
|
||
<typeStatus id="54CD96D07918293570274073FC41D7A2" box="[774,909,871,897]" pageId="7" pageNumber="6" type="paratype">PARATYPE</typeStatus>
|
||
/
|
||
<determinerName id="7E15C4B67918293570824073FBE6D7A2" box="[931,1066,871,897]" pageId="7" pageNumber="6">J.-B. Huchet</determinerName>
|
||
det. 2021”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C7BF97918293573E14085FD2BD031" pageId="7" pageNumber="6" type="diagnosis">
|
||
<paragraph id="8BC928727918293573E14085FD2BD031" blockId="7.[192,1440,913,1042]" pageId="7" pageNumber="6">
|
||
<emphasis id="B902F4607918293573E14085FEF0D788" bold="true" box="[192,316,913,939]" pageId="7" pageNumber="6">Diagnosis.</emphasis>
|
||
This species has the body robust, strongly convex, densely pubescent, the head with a strongly procurved “U” shaped raised carina distinctly tuberculate at both ends. Overall color orange with localized dark patterns on pronotum and elytra. The underside and legs are pale yellow; the antennae orange-yellow except the club, dark-brown. The stridulatory peg is present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C7BF97918293B73E14735FC65D1D8" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6" type="description">
|
||
<paragraph id="8BC928727918293B73E14735FA51D125" blockId="7.[191,1441,1057,1289]" lastBlockId="9.[191,1443,214,1287]" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6">
|
||
<emphasis id="B902F4607918293573E14735FE9ED018" bold="true" box="[192,338,1057,1083]" pageId="7" pageNumber="6">Description.</emphasis>
|
||
<typeStatus id="54CD96D07918293572794735FE0CD018" box="[344,448,1057,1083]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
|
||
male (
|
||
<figureCitation id="134D34F779182935712D4735FDA5D018" box="[524,617,1057,1083]" captionStart-0="Figure 6" captionStart-1="Figures 7–8" captionStartId-0="7.[193,269,1865,1891]" captionStartId-1="8.[192,279,1866,1892]" captionTargetBox-0="[549,1084,1335,1846]" captionTargetBox-1="[289,1345,219,1833]" captionTargetId-0="figure-496@7.[549,1084,1335,1848]" captionTargetId-1="figure-30@8.[218,1414,213,1836]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figure 6.Nothochodaeus. huxleyi Huchet, new species, male holotype, head in frontal view showing the U-shaped frontoclypeal carina (Photo J.-B. Huchet, MNHN)." captionText-1="Figures 7–8. Nothochodaeus huxleyi Huchet, new species, male holotype. 7) Habitus, dorsal view. 8) Habitus, latero-dorsal view (Photos J.-B. Huchet, MNHN)." figureDoi-0="http://doi.org/10.5281/zenodo.12808318" figureDoi-1="http://doi.org/10.5281/zenodo.12808322" httpUri-0="https://zenodo.org/record/12808318/files/figure.png" httpUri-1="https://zenodo.org/record/12808322/files/figure.png" pageId="7" pageNumber="6">Fig. 6–8</figureCitation>
|
||
,
|
||
<figureCitation id="134D34F77918293571554736FD74D01F" box="[628,696,1058,1084]" captionStart="Figures 11–13" captionStartId="11.[192,278,1153,1179]" captionTargetBox="[323,1307,462,1118]" captionTargetId="figure-86@11.[305,1327,457,1124]" captionTargetPageId="11" captionText="Figures 11–13. Nothochodaeus sp., male genitalia. 11) N. huxleyi Huchet, new species, aedeagus in latero-dorsal view. 12) N. borneensis Ochi, Kon and Masumoto, 2013, aedeagus in latero-dorsal view (insets illustrating a closeup of left paramere). 13) N. huxleyi Huchet, new species, genital segment (urite IX) (Photos J.-B. Huchet, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.12808328" httpUri="https://zenodo.org/record/12808328/files/figure.png" pageId="7" pageNumber="6">11, 13</figureCitation>
|
||
,
|
||
<figureCitation id="134D34F77918293571E54735FD2ED018" box="[708,738,1057,1083]" captionStart="Figures 16–19" captionStartId="12.[192,279,975,1001]" captionTargetBox="[336,1296,216,944]" captionTargetId="figure-90@12.[336,1296,216,944]" captionTargetPageId="12" captionText="Figures 16–19. Nothochodaeus sp., clypeofrontal carina and protibia, comparizon. 16) N. huxleyi Huchet, n. sp., holotype male, clypeofrontal carina, frontal view.17) N. borneensis, male, clypeofrontal carina, frontal view.18) N. huxleyi Huchet, n. sp., holotype male, right protibia, dorsal view. 19) N. borneensis, male, right protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12808332" httpUri="https://zenodo.org/record/12808332/files/figure.png" pageId="7" pageNumber="6">16</figureCitation>
|
||
).
|
||
<emphasis id="B902F4607918293571D44736FC83D01F" bold="true" box="[757,847,1058,1084]" pageId="7" pageNumber="6">Length:</emphasis>
|
||
<quantity id="4C8E85977918293570744736FC69D01F" box="[853,933,1058,1084]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="6" unit="mm" value="11.0">11 mm</quantity>
|
||
(from the apex of the mandibles to the apical part of the tergite VIII).
|
||
<emphasis id="B902F4607918293572EA4757FDD1D07E" bold="true" box="[459,541,1091,1117]" pageId="7" pageNumber="6">Width:</emphasis>
|
||
5.9 mm (greater width of the pronotum).
|
||
<emphasis id="B902F4607918293570CC4750FBE7D07D" bold="true" box="[1005,1067,1092,1118]" pageId="7" pageNumber="6">Head</emphasis>
|
||
: Head transverse, sub-hexagonal, entirely orange except for the frontoclypeal carina which is darkened. Frontoclypeal carina strongly raised, in inverted “U” shape, ended on both sides by a small, well individualized tubercle (
|
||
<figureCitation id="134D34F7791829357764479CFB49D081" box="[1093,1157,1160,1186]" captionStart="Figure 6" captionStartId="7.[193,269,1865,1891]" captionTargetBox="[549,1084,1335,1846]" captionTargetId="figure-496@7.[549,1084,1335,1848]" captionTargetPageId="7" captionText="Figure 6.Nothochodaeus. huxleyi Huchet, new species, male holotype, head in frontal view showing the U-shaped frontoclypeal carina (Photo J.-B. Huchet, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.12808318" httpUri="https://zenodo.org/record/12808318/files/figure.png" pageId="7" pageNumber="6">Fig. 6</figureCitation>
|
||
,
|
||
<figureCitation id="134D34F77918293577B0479CFB63D081" box="[1169,1199,1160,1186]" captionStart="Figures 16–19" captionStartId="12.[192,279,975,1001]" captionTargetBox="[336,1296,216,944]" captionTargetId="figure-90@12.[336,1296,216,944]" captionTargetPageId="12" captionText="Figures 16–19. Nothochodaeus sp., clypeofrontal carina and protibia, comparizon. 16) N. huxleyi Huchet, n. sp., holotype male, clypeofrontal carina, frontal view.17) N. borneensis, male, clypeofrontal carina, frontal view.18) N. huxleyi Huchet, n. sp., holotype male, right protibia, dorsal view. 19) N. borneensis, male, right protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12808332" httpUri="https://zenodo.org/record/12808332/files/figure.png" pageId="7" pageNumber="6">16</figureCitation>
|
||
). Clypeus trapezoidal in outline, strongly obliquely sloping forward, the anterior margin finely denticulate, the interval between these small denticles with a small yellow-orange bristle directed forward. Posterior declivity of clypeus relatively abrupt behind the carina, flanked on both sides by the clypeofrontal suture, slightly impressed and extending laterally up to the level of the anteromedial angle of the eye. Head surface shiny, covered with tight but not contiguous medium setiferous granules; a median longitudinal area, smooth, from the base to the middle of the head in front. Labrum transverse, dorsally convex, hyaline, long pubescent, strongly emarginate in the middle front, with a distinct raised transverse ridge in the middle. Eyes moderate in size, globose, strongly produced laterad with spaced long setae along their inner edge. Mandibles subequal, falciform, slightly concave dorsally, the surface matt, distinctly wrinkled on a microreticulate background. Mentum oblong, subquadrangular, weakly depressed in front. Antenna 10-segmented, the scape and funicle yellow-orange, pubescent, the club shiny, dark-brown except for narrow whitish apical part.
|
||
<emphasis id="B902F4607916293B710442D6FD69D5FF" bold="true" box="[549,677,450,476]" pageId="9" pageNumber="8">Pronotum:</emphasis>
|
||
Transverse, strongly convex, entirely margined, distinctly emarginate behind the head with a thin hyaline membrane in front, the lateral edges and base fringed with setae. Front angles prominent, projecting forward, the posterior ones obtusely rounded. Pronotal surface with a very dense punctation, consisting of small, tightly packed, medium-sized setiferous rounded granules with yellow-orange setae erect upwards. The setae located laterally at the front angles distinctly longer, black, curved upwards. A short median longitudinal furrow at basal third, extending forward to the anterior margin as a thin, darkened line. Pronotum uniformly orange with two black paramedian spots along the front margin, two other spots more broadly spaced along the base, and a distinct darkened fovea in the middle of each side.
|
||
<emphasis id="B902F4607916293B77B541BBFB2FD6EA" bold="true" box="[1172,1251,687,713]" pageId="9" pageNumber="8">Elytra:</emphasis>
|
||
Transverse, dull, densely pubescent, their color coarsely similar to that of the pronotum. Striae well-marked, relatively shallow, consisting of aligned sunken medium points separated by about 1–1.5× their diameter. Intervals almost flat, punctation strong and tight consisting of small setose granules on a microreticulate background, the minute setae erected, distinctly darkened. Humeral callus well developed. Elytral color patterns as follow: a thin transverse dark band on the fourth interstria along the front margin, a round black spot located on the humeral callus and a transverse median dark band interrupted in the middle, merging laterally with the next band located in the apical third of the elytra. Scutellum elongate, triangular in outline, the lateral edges darkened, the disc slightly depressed longitudinally in the middle.
|
||
<emphasis id="B902F4607916293B713040A8FD40D7F5" bold="true" box="[529,652,956,982]" pageId="9" pageNumber="8">Abdomen:</emphasis>
|
||
Strongly convex with six visible ventrites (III–VIII). Surface smooth, shiny, microreticulate; ventrites with minute setigerous tubercles along their front margin, the disc with medium setae roughly aligned. Tergite VIII (pygidium) brownish-black, pubescent, the punctation consisting of small close setose granules embedded on a microreticulate background; a lightened transverse band along the upper edge. Tergite VII (propygidium), entirely orange with trapezoidal interlocking mechanism. Stridulatory apparatus (sternite VI) present.
|
||
<emphasis id="B902F4607916293B72F34771FDC3D05C" bold="true" box="[466,527,1125,1151]" pageId="9" pageNumber="8">Legs:</emphasis>
|
||
Protibia quadridentate externally, the median tooth (third) barely visible, the basal tooth very reduced, spinose, pointed; inner apical angle of protibia with a short, acute pollex directed anteriorly (
|
||
<figureCitation id="134D34F77916293B73E947BDFED7D0E0" box="[200,283,1193,1219]" captionStart="Figures 16–19" captionStartId="12.[192,279,975,1001]" captionTargetBox="[336,1296,216,944]" captionTargetId="figure-90@12.[336,1296,216,944]" captionTargetPageId="12" captionText="Figures 16–19. Nothochodaeus sp., clypeofrontal carina and protibia, comparizon. 16) N. huxleyi Huchet, n. sp., holotype male, clypeofrontal carina, frontal view.17) N. borneensis, male, clypeofrontal carina, frontal view.18) N. huxleyi Huchet, n. sp., holotype male, right protibia, dorsal view. 19) N. borneensis, male, right protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12808332" httpUri="https://zenodo.org/record/12808332/files/figure.png" pageId="9" pageNumber="8">Fig. 18</figureCitation>
|
||
). Femur without accessory teeth, their surface with two parallel rows of setose punctures.
|
||
<emphasis id="B902F4607916293B760D47BDFA6CD0E0" bold="true" box="[1324,1440,1193,1219]" pageId="9" pageNumber="8">Genitalia:</emphasis>
|
||
Aedeagus with phallobase dorsoventrally curved, acuminate apically; parameres short, symmetrical, distinctly divergent at apex, the apical lobe ovoid. Endophallus well developed, unarmed (lacking endophallites) (
|
||
<figureCitation id="134D34F77916293B766147F8FA43D125" box="[1344,1423,1260,1286]" captionStart="Figures 11–13" captionStartId="11.[192,278,1153,1179]" captionTargetBox="[323,1307,462,1118]" captionTargetId="figure-86@11.[305,1327,457,1124]" captionTargetPageId="11" captionText="Figures 11–13. Nothochodaeus sp., male genitalia. 11) N. huxleyi Huchet, new species, aedeagus in latero-dorsal view. 12) N. borneensis Ochi, Kon and Masumoto, 2013, aedeagus in latero-dorsal view (insets illustrating a closeup of left paramere). 13) N. huxleyi Huchet, new species, genital segment (urite IX) (Photos J.-B. Huchet, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.12808328" httpUri="https://zenodo.org/record/12808328/files/figure.png" pageId="9" pageNumber="8">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0978FA7918293573E0445DFD39D3A6" ID-DOI="http://doi.org/10.5281/zenodo.12808318" ID-Zenodo-Dep="12808318" httpUri="https://zenodo.org/record/12808318/files/figure.png" pageId="7" pageNumber="6" startId="7.[193,269,1865,1891]" targetBox="[549,1084,1335,1846]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8BC928727918293573E0445DFD39D3A6" blockId="7.[191,1440,1865,1926]" pageId="7" pageNumber="6">
|
||
<emphasis id="B902F4607918293573E0445DFEE4D347" bold="true" box="[193,296,1865,1892]" pageId="7" pageNumber="6">Figure 6.</emphasis>
|
||
<emphasis id="B902F46079182935720B445EFE15D347" box="[298,473,1866,1892]" italics="true" pageId="7" pageNumber="6">
|
||
<taxonomicName id="4C7653F179182935720B445EFE19D347" authorityName="Nikolajev" authorityYear="2005" box="[298,469,1866,1892]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">Nothochodaeus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C7653F17918293572FC445EFD44D347" authority="Huchet" authorityName="Huchet" authorityYear="2021" box="[477,648,1866,1892]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="huxleyi" status="sp. nov.">
|
||
<emphasis id="B902F4607918293572FC445EFDE2D347" box="[477,558,1866,1892]" italics="true" pageId="7" pageNumber="6">huxleyi</emphasis>
|
||
Huchet
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A231491B7918293571B3445EFCDAD347" box="[658,790,1866,1892]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
|
||
, male holotype, head in frontal view showing the U-shaped frontoclypeal carina (Photo J.-B. Huchet, MNHN).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0978FA7917293A73E1445EFD25D3A5" ID-DOI="http://doi.org/10.5281/zenodo.12808322" ID-Zenodo-Dep="12808322" httpUri="https://zenodo.org/record/12808322/files/figure.png" pageId="8" pageNumber="7" startId="8.[192,279,1866,1892]" targetBox="[289,1345,219,1833]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BC928727917293A73E1445EFD25D3A5" blockId="8.[192,1440,1866,1927]" pageId="8" pageNumber="7">
|
||
<emphasis id="B902F4607917293A73E1445EFE9DD347" bold="true" box="[192,337,1866,1892]" pageId="8" pageNumber="7">Figures 7–8.</emphasis>
|
||
<taxonomicName id="4C7653F17917293A727B445EFD74D347" authority="Huchet" authorityName="Huchet" authorityYear="2021" box="[346,696,1866,1892]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="huxleyi" status="sp. nov.">
|
||
<emphasis id="B902F4607917293A727B445EFD90D347" box="[346,604,1866,1892]" italics="true" pageId="8" pageNumber="7">Nothochodaeus huxleyi</emphasis>
|
||
Huchet
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A231491B7917293A71E7445FFC82D347" box="[710,846,1866,1893]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
|
||
, male holotype.
|
||
<emphasis id="B902F4607917293A772D445EFBE8D347" bold="true" box="[1036,1060,1866,1892]" pageId="8" pageNumber="7">7)</emphasis>
|
||
Habitus, dorsal view.
|
||
<emphasis id="B902F4607917293A7603445EFAF6D347" bold="true" box="[1314,1338,1866,1892]" pageId="8" pageNumber="7">8)</emphasis>
|
||
Habitus, latero-dorsal view (Photos J.-B. Huchet, MNHN).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC928727916293B73E14602FC65D1D8" blockId="9.[190,1441,1302,1531]" pageId="9" pageNumber="8">
|
||
<emphasis id="B902F4607916293B73E14602FE67D113" bold="true" box="[192,427,1302,1328]" pageId="9" pageNumber="8">Sexual dimorphism.</emphasis>
|
||
The female
|
||
<typeStatus id="54CD96D07916293B710C4603FD55D112" box="[557,665,1303,1329]" pageId="9" pageNumber="8" type="paratype">paratypes</typeStatus>
|
||
show little variation from the male, and negligible sexual dimorphism. The area located behind by the clypeofrontal carina is flat-topped within the female, somewhat reminiscent of the clypeal horn of males of the genus
|
||
<taxonomicName id="4C7653F17916293B7151464EFC76D157" authority="Huchet, 2017" authorityName="Huchet" authorityYear="2017" box="[624,954,1370,1396]" class="Insecta" family="Ochodaeidae" genus="Ceratochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902F4607916293B7151464EFCD2D157" box="[624,798,1370,1396]" italics="true" pageId="9" pageNumber="8">Ceratochodaeus</emphasis>
|
||
<bibRefCitation id="EFE755837916293B7004464EFC76D157" author="Huchet JB" box="[805,954,1370,1396]" pageId="9" pageNumber="8" pagination="93 - 101" refId="ref5893" refString="Huchet JB. 2017. Un nouveau sous-genre et une nouvelle espece d'Ochodaeidae des Philippines (Coleoptera, Scarabaeoidea). Coleopteres 23 (9): 93 - 101." type="journal article" year="2017">Huchet, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
(this area is obliquely truncated backwards within the male). Finally, overall body coloring of the female may appear distinctly darker than in males (notably the underside, entirely dark orange-brown within the female
|
||
<typeStatus id="54CD96D07916293B70554689FC01D194" box="[884,973,1437,1463]" pageId="9" pageNumber="8" type="allotype">allotype</typeStatus>
|
||
whereas it is entirely pale yellow in male
|
||
<typeStatus id="54CD96D07916293B73E346ABFEEAD1FA" box="[194,294,1471,1497]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
, except for the abdomen, partly darkened). Similar color variation patterns between both sexes have been reported for the close species
|
||
<taxonomicName id="4C7653F17916293B716746F5FC69D1D8" authority="(Ochi et al. 2013)" baseAuthorityName="Ochi" baseAuthorityYear="2013" box="[582,933,1505,1531]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="B902F4607916293B716746F5FD16D1D8" box="[582,730,1505,1531]" italics="true" pageId="9" pageNumber="8">N. borneensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE755837916293B71C946F5FC52D1D8" author="Ochi T & Kon M & Masumoto K." box="[744,926,1505,1531]" pageId="9" pageNumber="8" pagination="309 - 326" refId="ref6191" refString="Ochi T, Kon M, Masumoto K. 2013. Six new taxa of the genus Nothochodaeus Nikolajev from the Malay Peninsula, Sumatra and Borneo (Coleoptera, Scarabaeoidea, Ochodaeidae). Japanese Journal of systematic entomology 19 (2): 309 - 326." type="journal article" year="2013">Ochi et al. 2013</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C7BF97916293B73E1451FFDC6D24B" pageId="9" pageNumber="8" type="etymology">
|
||
<paragraph id="8BC928727916293B73E1451FFDC6D24B" blockId="9.[191,1440,1546,1640]" pageId="9" pageNumber="8">
|
||
<emphasis id="B902F4607916293B73E1451FFE88D206" bold="true" box="[192,324,1547,1573]" pageId="9" pageNumber="8">Etymology.</emphasis>
|
||
Dedicated to the late English biologist Thomas Henry Huxley (1825–1895) who modified the famous Wallace’s Line, including
|
||
<collectingRegion id="49B2E6907916293B72FC4538FDF6D265" box="[477,570,1580,1606]" country="Philippines" name="Palawan" pageId="9" pageNumber="8">Palawan</collectingRegion>
|
||
as a component of the Asian biogeographic realm and separating this island from the oceanic
|
||
<collectingCountry id="F36168E27916293B72A4455AFDCAD24B" box="[389,518,1614,1640]" name="Philippines" pageId="9" pageNumber="8">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C7BF97916293B73E14563FCFFD2B1" box="[192,819,1655,1682]" pageId="9" pageNumber="8" type="distribution">
|
||
<paragraph id="8BC928727916293B73E14563FCFFD2B1" blockId="9.[192,819,1655,1682]" box="[192,819,1655,1682]" pageId="9" pageNumber="8">
|
||
<emphasis id="B902F4607916293B73E14563FE97D2B2" bold="true" box="[192,347,1655,1681]" pageId="9" pageNumber="8">Distribution.</emphasis>
|
||
North
|
||
<collectingRegion id="49B2E6907916293B728A456CFDC4D2B1" box="[427,520,1656,1682]" country="Philippines" name="Palawan" pageId="9" pageNumber="8">Palawan</collectingRegion>
|
||
(El Nido, Roxas) (
|
||
<figureCitation id="134D34F77916293B71F7456CFCEBD2B1" box="[726,807,1656,1682]" captionStart="Figure 20" captionStartId="12.[192,268,1866,1892]" captionTargetBox="[336,1295,1121,1837]" captionTargetId="figure-92@12.[336,1295,1121,1837]" captionTargetPageId="12" captionText="Figure 20.Distribution map of Nothochodaeus minotaurus Huchet, new species (red dots), and N. huxleyi Huchet, new species (yellow dots) on Palawan Island (the type locality indicated with a star)." figureDoi="http://doi.org/10.5281/zenodo.12808334" httpUri="https://zenodo.org/record/12808334/files/figure.png" pageId="9" pageNumber="8">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C7BF97916293973E145B6FAFCD5BB" lastPageId="11" lastPageNumber="10" pageId="9" pageNumber="8" type="discussion">
|
||
<paragraph id="8BC928727916293B73E145B6FBA8D3A5" blockId="9.[192,1441,1697,1927]" pageId="9" pageNumber="8">
|
||
<emphasis id="B902F4607916293B73E145B6FEE3D29F" bold="true" box="[192,303,1698,1724]" pageId="9" pageNumber="8">Remarks.</emphasis>
|
||
<taxonomicName id="4C7653F17916293B721445B6FDCDD298" authority="Huchet" authorityName="Huchet" authorityYear="2021" box="[309,513,1697,1723]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="huxleyi" status="new species">
|
||
<emphasis id="B902F4607916293B721445B6FE6BD298" box="[309,423,1697,1723]" italics="true" pageId="9" pageNumber="8">N. huxleyi</emphasis>
|
||
Huchet
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A231491B7916293B712D45B6FD5ED298" box="[524,658,1697,1724]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
|
||
, presents close affinities with
|
||
<taxonomicName id="4C7653F17916293B70FD45B6FF3AD2FE" authority="Ochi, Kon and Masumoto, 2013" authorityName="Ochi, Kon and Masumoto" authorityYear="2013" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="B902F4607916293B70FD45B6FBBCD298" box="[988,1136,1697,1723]" italics="true" pageId="9" pageNumber="8">N. borneensis</emphasis>
|
||
<bibRefCitation id="EFE755837916293B775645B5FF3AD2FE" author="Ochi T & Kon M & Masumoto K." pageId="9" pageNumber="8" pagination="309 - 326" refId="ref6191" refString="Ochi T, Kon M, Masumoto K. 2013. Six new taxa of the genus Nothochodaeus Nikolajev from the Malay Peninsula, Sumatra and Borneo (Coleoptera, Scarabaeoidea, Ochodaeidae). Japanese Journal of systematic entomology 19 (2): 309 - 326." type="journal article" year="2013">Ochi, Kon and Masumoto, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
from Borneo, but also with
|
||
<taxonomicName id="4C7653F17916293B711445D0FC00D2FE" authority="(Wiedemann, 1823)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[565,972,1731,1757]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="xanthomelas">
|
||
<emphasis id="B902F4607916293B711445D0FD2ED2FE" box="[565,738,1731,1757]" italics="true" pageId="9" pageNumber="8">N. xanthomelas</emphasis>
|
||
(Wiedemann, 1823)
|
||
</taxonomicName>
|
||
, inhabiting Java and Sumatra, pointing to a common biogeographical history between these two regions. Other geographically more distant species also appear to be morphologically closely related to these taxa. These include
|
||
<taxonomicName id="4C7653F17916293B7721441CFAAED302" authority="(Fairmaire, 1897)" baseAuthorityName="Fairmaire" baseAuthorityYear="1897" box="[1024,1378,1799,1825]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="grandiceps">
|
||
<emphasis id="B902F4607916293B7721441CFB59D302" box="[1024,1173,1799,1825]" italics="true" pageId="9" pageNumber="8">N. grandiceps</emphasis>
|
||
(Fairmaire, 1897)
|
||
</taxonomicName>
|
||
from continental
|
||
<collectingCountry id="F36168E27916293B7268443CFE42D361" box="[329,398,1832,1858]" name="China" pageId="9" pageNumber="8">China</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C7653F17916293B72BF443DFC0AD361" authority="(Ochi, Masumoto and Li, 2006)" baseAuthorityName="Ochi, Masumoto and Li" baseAuthorityYear="2006" box="[414,966,1832,1859]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="sakaii" subSpecies="sakaii">
|
||
<emphasis id="B902F4607916293B72BF443DFD81D361" box="[414,589,1832,1858]" italics="true" pageId="9" pageNumber="8">N. sakaii sakaii</emphasis>
|
||
(Ochi, Masumoto and Li, 2006)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F36168E27916293B7730443CFBA8D361" box="[1041,1124,1832,1858]" name="Taiwan" pageId="9" pageNumber="8">Taiwan</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C7653F17916293B7787443DFE25D346" authority="Masumoto and Ochi, 2015" authorityName="Masumoto and Ochi" authorityYear="2015" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="sakaii" subSpecies="khaoyaiensis">
|
||
<emphasis id="B902F4607916293B7787443DFA6CD361" box="[1190,1440,1832,1858]" italics="true" pageId="9" pageNumber="8">N. sakaii khaoyaiensis</emphasis>
|
||
Masumoto and Ochi, 2015
|
||
</taxonomicName>
|
||
inhabiting
|
||
<collectingCountry id="F36168E27916293B7148445EFD03D347" box="[617,719,1866,1892]" name="Thailand" pageId="9" pageNumber="8">Thailand</collectingCountry>
|
||
. We propose here to group all these morphologically related species into a distinct species complex designated as “
|
||
<taxonomicName id="4C7653F17916293B71D04479FC51D3A5" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[753,925,1900,1926]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="xanthomelas">
|
||
<emphasis id="B902F4607916293B71D04479FC51D3A5" box="[753,925,1900,1926]" italics="true" pageId="9" pageNumber="8">N. xanthomelas</emphasis>
|
||
</taxonomicName>
|
||
species complex”.
|
||
</paragraph>
|
||
<caption id="DF0978FA7915293873E1445EFD25D3A5" ID-DOI="http://doi.org/10.5281/zenodo.12808324" ID-Zenodo-Dep="12808324" httpUri="https://zenodo.org/record/12808324/files/figure.png" pageId="10" pageNumber="9" startId="10.[192,279,1866,1892]" targetBox="[357,1362,226,1824]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8BC928727915293873E1445EFD25D3A5" blockId="10.[192,1440,1866,1927]" pageId="10" pageNumber="9">
|
||
<emphasis id="B902F4607915293873E1445EFE90D347" bold="true" box="[192,348,1866,1892]" pageId="10" pageNumber="9">Figures 9–10.</emphasis>
|
||
<taxonomicName id="4C7653F1791529387243445EFD77D347" authority="Huchet" authorityName="Huchet" authorityYear="2021" box="[354,699,1866,1892]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="huxleyi" status="sp. nov.">
|
||
<emphasis id="B902F460791529387243445EFDADD347" box="[354,609,1866,1892]" italics="true" pageId="10" pageNumber="9">Nothochodaeus huxleyi</emphasis>
|
||
Huchet
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A231491B7915293871E7445FFC87D347" box="[710,843,1866,1893]" pageId="10" pageNumber="9" rank="species">new species</taxonomicNameLabel>
|
||
, female allotype.
|
||
<emphasis id="B902F46079152938772A445EFBE8D347" bold="true" box="[1035,1060,1866,1892]" pageId="10" pageNumber="9">9)</emphasis>
|
||
Habitus, dorsal view.
|
||
<emphasis id="B902F460791529387637445EFAF0D347" bold="true" box="[1302,1340,1866,1892]" pageId="10" pageNumber="9">10)</emphasis>
|
||
Habitus, latero-dorsal view (Photos J.-B. Huchet, MNHN).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC9287279142939722143C3FAFCD5BB" blockId="11.[191,1442,214,408]" pageId="11" pageNumber="10">
|
||
<taxonomicName id="4C7653F179142939722143C3FE03D4D3" authority="Huchet" authorityName="Huchet" authorityYear="2021" box="[256,463,214,240]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="huxleyi" status="new species">
|
||
<emphasis id="B902F46079142939722143C3FEB8D4D3" box="[256,372,214,240]" italics="true" pageId="11" pageNumber="10">N. huxleyi</emphasis>
|
||
Huchet
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A231491B7914293972FD43C2FDAED4D3" box="[476,610,214,240]" pageId="11" pageNumber="10" rank="species">new species</taxonomicNameLabel>
|
||
can be separated from
|
||
<taxonomicName id="4C7653F179142939705143C3FBCAD4D3" authorityName="Ochi, Kon and Masumoto" authorityYear="2013" box="[880,1030,214,240]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="B902F46079142939705143C3FBCAD4D3" box="[880,1030,214,240]" italics="true" pageId="11" pageNumber="10">N. borneensis</emphasis>
|
||
</taxonomicName>
|
||
by the conformation of the clypeus, distinctly more abruptly sloping forward, the clypeofrontal carina more strongly arched in male (
|
||
<figureCitation id="134D34F77914293977D243E3FAA7D532" box="[1267,1387,247,273]" captionStart="Figures 16–19" captionStartId="12.[192,279,975,1001]" captionTargetBox="[336,1296,216,944]" captionTargetId="figure-90@12.[336,1296,216,944]" captionTargetPageId="12" captionText="Figures 16–19. Nothochodaeus sp., clypeofrontal carina and protibia, comparizon. 16) N. huxleyi Huchet, n. sp., holotype male, clypeofrontal carina, frontal view.17) N. borneensis, male, clypeofrontal carina, frontal view.18) N. huxleyi Huchet, n. sp., holotype male, right protibia, dorsal view. 19) N. borneensis, male, right protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12808332" httpUri="https://zenodo.org/record/12808332/files/figure.png" pageId="11" pageNumber="10">Fig. 16–17</figureCitation>
|
||
), the pronotal and elytral patterns of coloration, and the scutellum uniformly orange (always black or darkened within
|
||
<taxonomicName id="4C7653F179142939739E422FFE9ED577" authorityName="Ochi, Kon and Masumoto" authorityYear="2013" box="[191,338,314,340]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="B902F46079142939739E422FFE9ED577" box="[191,338,314,340]" italics="true" pageId="11" pageNumber="10">N. borneensis</emphasis>
|
||
</taxonomicName>
|
||
as indicated by
|
||
<bibRefCitation id="EFE75583791429397120422EFD05D576" author="Ochi T & Kon M & Masumoto K." box="[513,713,314,341]" pageId="11" pageNumber="10" pagination="309 - 326" refId="ref6191" refString="Ochi T, Kon M, Masumoto K. 2013. Six new taxa of the genus Nothochodaeus Nikolajev from the Malay Peninsula, Sumatra and Borneo (Coleoptera, Scarabaeoidea, Ochodaeidae). Japanese Journal of systematic entomology 19 (2): 309 - 326." type="journal article" year="2013">Ochi et al. (2013))</bibRefCitation>
|
||
(
|
||
<figureCitation id="134D34F77914293971F7422EFC83D576" box="[726,847,314,341]" captionStart="Figures 14–15" captionStartId="11.[194,281,1866,1892]" captionTargetBox="[263,1365,1304,1846]" captionTargetId="figure-9@11.[258,1375,1299,1849]" captionTargetPageId="11" captionText="Figures 14–15. Nothochodaeus borneensis Ochi, Kon and Masumoto, 2013. 14) Male, dorsal view (coll. MNHN). 15) Female, dorsal view (coll. MNHN) (Photos J.-B. Huchet, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.12808330" httpUri="https://zenodo.org/record/12808330/files/figure.png" pageId="11" pageNumber="10">Fig. 14–15</figureCitation>
|
||
). Protibiae broader in both sexes, 1st tooth longer and more strongly curved outward, 2nd very large (distinctly shorter and less developed within
|
||
<taxonomicName id="4C7653F17914293977E94249FA97D555" authorityName="Ochi, Kon and Masumoto" authorityYear="2013" box="[1224,1371,348,374]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="B902F4607914293977E94249FA97D555" box="[1224,1371,348,374]" italics="true" pageId="11" pageNumber="10">N. borneensis</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="134D34F77914293976554248FEC5D5BB" captionStart="Figures 16–19" captionStartId="12.[192,279,975,1001]" captionTargetBox="[336,1296,216,944]" captionTargetId="figure-90@12.[336,1296,216,944]" captionTargetPageId="12" captionText="Figures 16–19. Nothochodaeus sp., clypeofrontal carina and protibia, comparizon. 16) N. huxleyi Huchet, n. sp., holotype male, clypeofrontal carina, frontal view.17) N. borneensis, male, clypeofrontal carina, frontal view.18) N. huxleyi Huchet, n. sp., holotype male, right protibia, dorsal view. 19) N. borneensis, male, right protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12808332" httpUri="https://zenodo.org/record/12808332/files/figure.png" pageId="11" pageNumber="10">Fig. 18–19</figureCitation>
|
||
). Finally, these two species can be separated by the conformation of male genitalia (
|
||
<figureCitation id="134D34F779142939778B426AFAE8D5BB" box="[1194,1316,382,408]" captionStart="Figures 11–13" captionStartId="11.[192,278,1153,1179]" captionTargetBox="[323,1307,462,1118]" captionTargetId="figure-86@11.[305,1327,457,1124]" captionTargetPageId="11" captionText="Figures 11–13. Nothochodaeus sp., male genitalia. 11) N. huxleyi Huchet, new species, aedeagus in latero-dorsal view. 12) N. borneensis Ochi, Kon and Masumoto, 2013, aedeagus in latero-dorsal view (insets illustrating a closeup of left paramere). 13) N. huxleyi Huchet, new species, genital segment (urite IX) (Photos J.-B. Huchet, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.12808328" httpUri="https://zenodo.org/record/12808328/files/figure.png" pageId="11" pageNumber="10">Fig. 11–12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |