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<document id="09CE84B33FB4AB35AC12C23561E01F56" ID="10.11646/zootaxa.4127.1.3" ID-CLB-Dataset="38760" ID-DOI="10.11646/zootaxa.4127.1.3" ID-GBIF-Dataset="0f8ad304-a17d-47b8-8885-0ec006a35f79" ID-ISSN="1175-5326" ID-Zenodo-Dep="266084" ID-ZooBank="B24B4A1B-208F-42A9-94FE-563FBED43742" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1466577261773" checkinUser="plazi" docAuthor="Obando, Ranulfo González, García Aldrete, Alfonso N. &amp; Carrejo, Nancy" docDate="2016" docId="03EF87F9FFE1FFA6FF6EFD24E5F265F9" docLanguage="en" docName="zootaxa.4127.1.3.pdf" docOrigin="Zootaxa 4127 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladiopsocus gregoriorum Obando, Aldrete &amp; Carrejo, 2016, n. sp." docType="treatment" docVersion="9" masterDocId="FFD6FF81FFE6FFA1FFF9FF90E626624E" masterDocTitle="New species of Cladiopsocus Roesler (Psocodea: Psocoptera: Psocomorpha: Cladiopsocidae) from the Colombian Amazonia" updateTime="1698663090508" updateUser="ExternalLinkService">
<mods:mods id="B7A6934736B9A1CCF0E05B63587D219B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1DE76206560252F85C142CB6A3E05B2C">New species of Cladiopsocus Roesler (Psocodea: Psocoptera: Psocomorpha: Cladiopsocidae) from the Colombian Amazonia</mods:title>
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<mods:name id="40B5104E5BF8B09B113A8364C9BC8F77" type="personal">
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<mods:namePart id="7655ADE5F26F53C88E3BB01BED5C60C9">Obando, Ranulfo González</mods:namePart>
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<mods:namePart id="DAE8D09A58CC7BA3B512C640A788C912">García Aldrete, Alfonso N.</mods:namePart>
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<mods:namePart id="ABE9C8BBEB5E7578FCAB7635A572A8CC">Carrejo, Nancy</mods:namePart>
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<mods:title id="56F632135D99FF9F31974F75EEF0BFE1">Zootaxa</mods:title>
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<mods:date id="3CD1643E57AD052C7F040E8F72B7E915">2016</mods:date>
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<mods:identifier id="BFFF951045A5D9253831E1598E7D098C" type="DOI">10.11646/zootaxa.4127.1.3</mods:identifier>
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<treatment id="03EF87F9FFE1FFA6FF6EFD24E5F265F9" ID-DOI="http://doi.org/10.5281/zenodo.6079720" ID-GBIF-Taxon="121061527" ID-Zenodo-Dep="6079720" LSID="urn:lsid:plazi:treatment:03EF87F9FFE1FFA6FF6EFD24E5F265F9" httpUri="http://treatment.plazi.org/id/03EF87F9FFE1FFA6FF6EFD24E5F265F9" pageId="7">
<subSubSection id="C35C6564FFE1FFA6FF6EFD24E70160A1" pageId="7" type="nomenclature">
<paragraph id="8BF936EFFFE1FFA6FF6EFD24E44E6080" blockId="7.[151,616,692,751]" box="[151,616,692,718]" pageId="7">
<heading id="D0B18183FFE1FFA6FF6EFD24E44E6080" bold="true" box="[151,616,692,718]" fontSize="11" level="1" pageId="7" reason="1">
<emphasis id="B932EAFDFFE1FFA6FF6EFD24E44E6080" bold="true" box="[151,616,692,718]" pageId="7">
<taxonomicName id="4C464D6CFFE1FFA6FF6EFD24E7FF6080" box="[151,473,692,718]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="7" phylum="Arthropoda" rank="species" species="gregoriorum" status="sp. nov.">
<emphasis id="B932EAFDFFE1FFA6FF6EFD24E7FF6080" bold="true" box="[151,473,692,718]" italics="true" pageId="7">Cladiopsocus gregoriorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2015786FFE1FFA6FE19FD24E4046080" box="[480,546,692,718]" pageId="7" rank="species">n. sp.</taxonomicNameLabel>
Male
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936EFFFE1FFA6FF6EFD46E70160A1" blockId="7.[151,616,692,751]" box="[151,295,726,751]" pageId="7">
(
<figureCitation id="137D2A6AFFE1FFA6FF66FD46E73960A1" box="[159,287,726,751]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Figs 2228</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C6564FFE1FFA6FF6EFC8EE273643D" pageId="7" type="diagnosis">
<paragraph id="8BF936EFFFE1FFA6FF6EFC8EE47F6189" blockId="7.[151,1436,798,1975]" pageId="7">
<emphasis id="B932EAFDFFE1FFA6FF6EFC8EE7376179" bold="true" box="[151,273,798,823]" pageId="7">Diagnosis.</emphasis>
Belonging in species group “2”, subgroup 4, in the classification of
<bibRefCitation id="EFD74B1EFFE1FFA6FBEEFC8EE2FF6179" author="Eertmoed" box="[1047,1241,798,823]" pageId="7" refString="Eertmoed, G. E. (1986) The redefinition of Cladiopsocus (Psocoptera: Cladiopsocidae). Studies on Neotropical Fauna and Environment, 21 (4), 207 - 229." type="journal article" year="1986">Eertmoed (1986)</bibRefCitation>
. In this subgroup it corresponds to species with phallosome with internal parameres vestigial, or not united posteriorly. Unlike
<taxonomicName id="4C464D6CFFE1FFA6FA7BFCD3E7006131" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="7" phylum="Arthropoda" rank="species" species="deliciasensis" status="sp. nov.">
<emphasis id="B932EAFDFFE1FFA6FA7BFCD3E7006131" italics="true" pageId="7">C. deliciasensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2015786FFE1FFA6FED7FCF6E7486131" box="[302,366,870,895]" pageId="7" rank="species">
<emphasis id="B932EAFDFFE1FFA6FED7FCF6E7486131" bold="true" box="[302,366,870,895]" pageId="7">n. sp.</emphasis>
</taxonomicNameLabel>
, it does not have the paraproctal prong with a tapered basal tooth, and unlike
<emphasis id="B932EAFDFFE1FFA6FB1FFCF7E73C61ED" italics="true" pageId="7">
<taxonomicName id="4C464D6CFFE1FFA6FB1FFCF7E3506130" box="[1254,1398,871,894]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="7" phylum="Arthropoda" rank="species" species="prionotus">C. prionotus</taxonomicName>
,
<taxonomicName id="4C464D6CFFE1FFA6FA7BFCF7E73C61ED" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="7" phylum="Arthropoda" rank="species" species="mocoaensis" status="sp. nov.">C. mocoaensis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A2015786FFE1FFA6FEDDFC1AE74061ED" box="[292,358,906,931]" pageId="7" rank="species">
<emphasis id="B932EAFDFFE1FFA6FEDDFC1AE74061ED" bold="true" box="[292,358,906,931]" pageId="7">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C464D6CFFE1FFA6FE81FC1BE7CA61ED" box="[376,492,907,931]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="7" phylum="Arthropoda" rank="species" species="garciai">
<emphasis id="B932EAFDFFE1FFA6FE81FC1BE7CA61ED" box="[376,492,907,931]" italics="true" pageId="7">C. garciai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C464D6CFFE1FFA6FDD3FC1BE4DC61ED" box="[554,762,906,931]" class="Insecta" family="Cladiopsocidae" genus="Cladiopsocus" kingdom="Animalia" order="Psocodea" pageId="7" phylum="Arthropoda" rank="species" species="multimaculatus">
<emphasis id="B932EAFDFFE1FFA6FDD3FC1BE4DC61ED" box="[554,762,906,931]" italics="true" pageId="7">C. multimaculatus</emphasis>
</taxonomicName>
, the epiproct has two tooth-like short projections distally, along the longitudinal midline (
<figureCitation id="137D2A6AFFE1FFA6FE01FC3EE46A6189" box="[504,588,942,967]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="8BF936EFFFE1FFA6FF3EFC42E6C96745" blockId="7.[151,1436,798,1975]" pageId="7">
<emphasis id="B932EAFDFFE1FFA6FF3EFC42E72C61A5" bold="true" box="[199,266,978,1003]" pageId="7">Color</emphasis>
. Head (
<figureCitation id="137D2A6AFFE1FFA6FE9EFC42E79D61A5" box="[359,443,978,1003]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 24</figureCitation>
) dark brown, anteclypeus and labrum pale brown. Compound eyes dark brown, ocelli hyaline. Antennae pale brown, flagellomeres with clear distal ends; maxillary palps pale brown. Meso- and metanotum brown, thoracic pleura and legs light brown, tibia and tarsi dark brown. Forewing membrane hyaline, with brown spots at wing margin; pterostigma with pale brown spot distally on costal area; wing margin with brown spots between the ends of veins R, M and Cu1 (
<figureCitation id="137D2A6AFFE1FFA6FD03FBF2E5686635" box="[762,846,1122,1147]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 22</figureCitation>
); veins brown, with brown spots at ends of R, M and Cu1a. Hindwing hyaline with a smoked spot on vein ends, and on wing margin between M and Cu1 (
<figureCitation id="137D2A6AFFE1FFA6FAC0FB16E3A966D1" box="[1337,1423,1158,1183]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 23</figureCitation>
). Abdomen cream, with light brown bands, clunium and hypandrium pale brown; phallosome pale brown; epiproct cream, with two little tubercles dark brown; paraprocts cream, paraproctal prong dark brown, more pigmented distally.
</paragraph>
<paragraph id="8BF936EFFFE1FFA6FF3EFA86E273643D" blockId="7.[151,1436,798,1975]" pageId="7">
<emphasis id="B932EAFDFFE1FFA6FF3EFA86E7476761" bold="true" box="[199,353,1302,1327]" pageId="7">Morphology.</emphasis>
As in diagnosis plus the following: head vertex slightly concave, almost on the same level with the top margin of the compound eyes (
<figureCitation id="137D2A6AFFE1FFA6FDBCFAAAE4BF671D" box="[581,665,1338,1363]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 24</figureCitation>
). D is 60% the length of the head in front view, almost as wide as the interocular distance (1.15); with interommatidial setae. Forewings (
<figureCitation id="137D2A6AFFE1FFA6FC5CFACEE5DA6739" box="[933,1020,1374,1399]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 22</figureCitation>
), membrane of pterostigma without setae; without spur veins. Hypandrium (
<figureCitation id="137D2A6AFFE1FFA6FD99FA12E49567D5" box="[608,691,1410,1435]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 27</figureCitation>
) broad, shield-shaped, rounded distally, with medium sized setae and lateral macrosetae. Epiproct trapeziform (
<figureCitation id="137D2A6AFFE1FFA6FD60FA36E4CA67F1" box="[665,748,1446,1471]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 25</figureCitation>
), posterior border broadly rounded, three medium sized setae in mesal field, and field of medium sized setae and macrosetae distally. Paraprocts (
<figureCitation id="137D2A6AFFE1FFA6FBC2FA5AE2AA67AD" box="[1083,1164,1482,1507]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 26</figureCitation>
) with abundant medium sized setae and macrosetae as illustrated; mesal prong stout, strongly sclerotized, acuminate distally; sensory fields with 18 trichobothria on basal rosettes. Phallosome sclerotized, slightly concave anteriorly, wider posteriorly, deeply cleft in the middle; external parameres broad, membranous, bearing pores, endophallic lobes triangular; arms of internal parameres separate, distally pointed, thick, well sclerotized (
<figureCitation id="137D2A6AFFE1FFA6FC0DF9CAE26F643D" box="[1012,1097,1626,1651]" captionStart="FIGURES 22 28" captionStartId="8.[151,264,1948,1971]" captionTargetBox="[161,1434,195,1918]" captionTargetId="figure@8.[152,1435,195,1926]" captionTargetPageId="8" captionText="FIGURES 22 28. Cladiopsocus gregoriorum n. sp. Male. 22. Forewing. 23. Hindwing. 24. Front view of head. 25. Epiproct. 26. Left paraproct. 27. Hypandrium. 28. Phallosome. Scales in mm." httpUri="https://zenodo.org/record/266088/files/figure.png" pageId="7">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C6564FFE1FFA6FF3EF9EEE25D6521" pageId="7" type="description">
<paragraph id="8BF936EFFFE1FFA6FF3EF9EEE7246491" blockId="7.[151,1436,798,1975]" pageId="7">
<emphasis id="B932EAFDFFE1FFA6FF3EF9EEE75B64D9" bold="true" box="[199,381,1662,1687]" pageId="7">Measurements.</emphasis>
FW: 2375, HW: 1825, F: 520, T: 940, t1: 380, t2: 50, t3: 63, ctt1: 22, f1: 390, f2: 285, f3: 240, f4: 217, f5: 140, f6: 130, f7: 113, f8: 110, f9: 98, f10: 63, f11: 82, Mx4: 155, IO: 195, D: 336, d: 225, IO/d: 0.87, PO: 0.67.
</paragraph>
<paragraph id="8BF936EFFFE1FFA6FF3EF97AE25D6521" blockId="7.[151,1436,798,1975]" pageId="7">
<emphasis id="B932EAFDFFE1FFA6FF3EF97AE788654D" bold="true" box="[199,430,1770,1795]" pageId="7">Specimens studied.</emphasis>
<typeStatus id="54FD884DFFE1FFA6FE41F97AE406654D" box="[440,544,1770,1795]" pageId="7" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F351767FFFE1FFA6FD8AF97BE52A654D" box="[627,780,1771,1795]" name="Colombia" pageId="7">COLOMBIA</collectingCountry>
. Amazonas, Leticia, San Martín de Amacayacu. Maloca trail,
<geoCoordinate id="EE725028FFE1FFA6FF21F89FE7436569" box="[216,357,1807,1831]" direction="south" orientation="latitude" pageId="7" precision="1" value="-3.7288334">3º4343.8”S</geoCoordinate>
:
<geoCoordinate id="EE725028FFE1FFA6FE8EF89FE4016568" box="[375,551,1807,1831]" direction="west" orientation="longitude" pageId="7" precision="1" value="-70.313866">70º1849.9” W</geoCoordinate>
,
<specimenCount id="9D40FD66FFE1FFA6FDC1F89FE4596568" box="[568,639,1807,1831]" pageId="7" type="generic">
<quantity id="4CBE9B0AFFE1FFA6FDC1F89FE45D6568" box="[568,635,1807,1831]" metricMagnitude="1" metricUnit="m" metricValue="9.3" pageId="7" unit="m" value="93.0">93 m</quantity>
.
</specimenCount>
<date id="FFF8102FFFE1FFA6FD75F89EE56A6569" box="[652,844,1806,1831]" pageId="7" value="2015-08-14" valueMax="2015-08-15" valueMin="2015-08-14">1415.VIII.2015</date>
. MUSENUV slide code 26138, N. Carrejo &amp; R. Gregorio.
<typeStatus id="54FD884DFFE1FFA6FEF3F8A3E75A6505" box="[266,380,1843,1867]" pageId="7" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D40FD66FFE1FFA6FE7FF8A3E7F26505" box="[390,468,1842,1867]" pageId="7" type="male">1 male</specimenCount>
, Agua Blanca trail,
<geoCoordinate id="EE725028FFE1FFA6FD4DF8A3E5666505" box="[692,832,1843,1867]" direction="south" orientation="latitude" pageId="7" precision="1" value="-3.705278">3º4219.0”S</geoCoordinate>
:
<geoCoordinate id="EE725028FFE1FFA6FCB5F8A2E5D36504" box="[844,1013,1842,1867]" direction="west" orientation="longitude" pageId="7" precision="1" value="-70.34058">70º2026.1” W</geoCoordinate>
,
<specimenCount id="9D40FD66FFE1FFA6FBF9F8A3E2676504" box="[1024,1089,1843,1867]" pageId="7" type="generic">
<quantity id="4CBE9B0AFFE1FFA6FBF9F8A3E21A6504" box="[1024,1084,1843,1867]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="7" unit="m" value="70.0">70 m</quantity>
.
</specimenCount>
<date id="FFF8102FFFE1FFA6FBBEF8A3E3216505" box="[1095,1287,1842,1867]" pageId="7" value="2015-08-12" valueMax="2015-08-13" valueMin="2015-08-12">1213.VIII.2015</date>
. MUSENUV slide code 26139, R. González, N. Calderón &amp; O. Saenz. Led light trap on forest canopy.
</paragraph>
</subSubSection>
<subSubSection id="C35C6564FFE1FFA6FF3EF8EAE5F265F9" pageId="7" type="etymology">
<paragraph id="8BF936EFFFE1FFA6FF3EF8EAE5F265F9" blockId="7.[151,1436,798,1975]" pageId="7">
<emphasis id="B932EAFDFFE1FFA6FF3EF8EAE76865DD" bold="true" box="[199,334,1914,1939]" pageId="7">Etymology.</emphasis>
This species is gratefully dedicated to José and Robinson Gregorio, guide and collaborator, respectively, to the explorations in San Martín de Amacayacu, Amazonas.
</paragraph>
</subSubSection>
</treatment>
</document>