treatments-xml/data/F8/68/4D/F8684D7942F05F29B5D99020233C20B2.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.82461" ID-Pensoft-Pub="1313-2970-1110-1" ID-Pensoft-UUID="88D66028C7A752528DD9138A2105F895" ID-ZooBank="01302157EE404B9999DD3EC3377D929C" ModsDocID="1313-2970-1110-1" checkinTime="1657006639549" checkinUser="pensoft" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak, Lin, Aung &amp; Panha, Somsak" docDate="2022" docId="F8684D7942F05F29B5D99020233C20B2" docLanguage="en" docName="ZooKeys 1110: 1-37" docOrigin="ZooKeys 1110" docPubDate="2022-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1110.82461" docTitle="Burmochlamys albida Pholyotha &amp; Panha 2022, sp. nov." docType="treatment" docUuid="CF80F28E-FFF3-4C4E-A453-D383CA8FA1E2" docUuidSource="ZooBank" docVersion="1" id="88D66028C7A752528DD9138A2105F895" lastPageNumber="1" masterDocId="88D66028C7A752528DD9138A2105F895" masterDocTitle="Uncovering local endemism from southeastern Myanmar: description of the new karst-associated terrestrial snail genus Burmochlamys (Eupulmonata, Helicarionidae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="1" updateTime="1657006941096" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Uncovering local endemism from southeastern Myanmar: description of the new karst-associated terrestrial snail genus Burmochlamys (Eupulmonata, Helicarionidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pholyotha, Arthit</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Aung</mods:namePart>
<mods:affiliation>Fauna and Flora International, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1110</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>37</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1110.82461</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1110.82461</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1110-1</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:CF80F28E-FFF3-4C4E-A453-D383CA8FA1E2" httpUri="http://treatment.plazi.org/id/F8684D7942F05F29B5D99020233C20B2" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/CF80F28E-FFF3-4C4E-A453-D383CA8FA1E2" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Burmochlamys albida" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="albida" status="sp. nov.">Burmochlamys albida Pholyotha &amp; Panha</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the Salween River basin (known as Thanlwin River) in the southeastern Myanmar showing the sampling sites. Coloured triangles indicate the localities recorded for each Burmochlamys species. White circles indicate the localities recorded for Sophina species. White squares indicate the sampling localities with no records of both Sophina and Burmochlamys species." figureDoi="10.3897/zookeys.1110.82461.figure1" httpUri="https://binary.pensoft.net/fig/709567" pageId="0" pageNumber="1">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitat of some species of Burmochlamys gen. nov. in the karst basin of Hpa-An, Kayin State, Myanmar. A type locality of B. fasciola sp. nov. at Bardai Mountain and microhabitat structure of the karst wall B newly discovered locality of B. poongee at Kaw Ka Thaung Cave C type locality of B. albida sp. nov. at Waiponla Monastery D type locality of B. moulmeinica sp. nov. at Lun Nya Pagoda E type locality of B. versicolor sp. nov. at Bayin Nyi Cave. Photos by Ruttapon Srisonchai." figureDoi="10.3897/zookeys.1110.82461.figure2" httpUri="https://binary.pensoft.net/fig/709568" pageId="0" pageNumber="1">, 2C</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Living snails of Burmochlamys species A B. fasciola sp. nov. paratype CUMZ 14214 showing left and right lateral views B B. cassidula specimen CUMZ 14209 C B. poongee specimen CUMZ 14210 D B. albida sp. nov. paratype CUMZ 14212 E B. moulmeinica sp. nov. paratype CUMZ 14217 F B. versicolor sp. nov. paratype CUMZ 14219 G B. whitteni sp. nov. paratype CUMZ 14222." figureDoi="10.3897/zookeys.1110.82461.figure5" httpUri="https://binary.pensoft.net/fig/709571" pageId="0" pageNumber="1">, 5D</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells of Burmochlamys species A, B B. albida sp. nov. A holotype CUMZ 14211, and D paratype CUMZ 14212 C, D B. fasciola sp. nov. C holotype CUMZ 14213 D paratype CUMZ 14214 E, F B. moulmeinica sp. nov. E holotype CUMZ 14216 F paratype CUMZ 14217." figureDoi="10.3897/zookeys.1110.82461.figure7" httpUri="https://binary.pensoft.net/fig/709573" pageId="0" pageNumber="1">, 7A, B</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Representative SEM images of the shell of Burmochlamys species A-C B. cassidula specimen CUMZ 14209 A protoconch B body whorl C zoom-in view of body whorl D-F B. poongee specimen CUMZ 14210 D protoconch E body whorl F zoom-in view of body whorl G-I B. albida sp. nov. paratype CUMZ 14212 G protoconch H body whorl I zoom-in view of body whorl J-L B. fasciola sp. nov. paratype CUMZ 14214 J protoconch K body whorl L zoom-in view of body whorl." figureDoi="10.3897/zookeys.1110.82461.figure8" httpUri="https://binary.pensoft.net/fig/709574" pageId="0" pageNumber="1">, 8G-I</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. General view of the genital system of Burmochlamys species A B. cassidula specimen CUMZ 14209 B B. poongee specimen CUMZ 14210 C B. albida sp. nov. paratype CUMZ 14212. White arrow indicates the end of the penis. Red-dotted line indicates vas deferens missing." figureDoi="10.3897/zookeys.1110.82461.figure9" httpUri="https://binary.pensoft.net/fig/709575" pageId="0" pageNumber="1">, 9C</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Representative SEM images of the radula of Burmochlamys species A B. cassidula specimen CUMZ 14209 B B. poongee specimen CUMZ 14210 C B. albida sp. nov. paratype CUMZ 14212 D B. fasciola sp. nov. paratype CUMZ 14214. Central tooth indicated by ' C'; yellow colour indicates central tooth row." figureDoi="10.3897/zookeys.1110.82461.figure11" httpUri="https://binary.pensoft.net/fig/709577" pageId="0" pageNumber="1">, 11C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
Type material. 
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
</emphasis>
: CUMZ 14211 (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells of Burmochlamys species A, B B. albida sp. nov. A holotype CUMZ 14211, and D paratype CUMZ 14212 C, D B. fasciola sp. nov. C holotype CUMZ 14213 D paratype CUMZ 14214 E, F B. moulmeinica sp. nov. E holotype CUMZ 14216 F paratype CUMZ 14217." figureDoi="10.3897/zookeys.1110.82461.figure7" httpUri="https://binary.pensoft.net/fig/709573" pageId="0" pageNumber="1">7A</figureCitation>
; width 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.5" unit="mm" value="4.5">4.5 mm</quantity>
, height 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.7" unit="mm" value="3.7">3.7 mm</quantity>
). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
: Same locality as holotype: CUMZ 14212 (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells of Burmochlamys species A, B B. albida sp. nov. A holotype CUMZ 14211, and D paratype CUMZ 14212 C, D B. fasciola sp. nov. C holotype CUMZ 14213 D paratype CUMZ 14214 E, F B. moulmeinica sp. nov. E holotype CUMZ 14216 F paratype CUMZ 14217." figureDoi="10.3897/zookeys.1110.82461.figure7" httpUri="https://binary.pensoft.net/fig/709573" pageId="0" pageNumber="1">7B</figureCitation>
; width 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.4" unit="mm" value="4.4">4.4 mm</quantity>
, height 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.2" unit="mm" value="3.2">3.2 mm</quantity>
), NHMUK (two shells).
</paragraph>
<caption doi="10.3897/zookeys.1110.82461.figure7" httpUri="https://binary.pensoft.net/fig/709573" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
Shells of 
<taxonomicName authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
species 
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<taxonomicName genus="B." lsidName="B. albida" pageId="0" pageNumber="1" rank="species" species="albida">
<emphasis italics="true" pageId="0" pageNumber="1">B. albida</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<typeStatus>holotype</typeStatus>
CUMZ 14211, and 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<typeStatus>paratype</typeStatus>
CUMZ 14212 
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
<taxonomicName genus="B." lsidName="B. fasciola" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis italics="true" pageId="0" pageNumber="1">B. fasciola</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<typeStatus>holotype</typeStatus>
CUMZ 14213 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<typeStatus>paratype</typeStatus>
CUMZ 14214 
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
<taxonomicName genus="B." lsidName="B. moulmeinica" pageId="0" pageNumber="1" rank="species" species="moulmeinica">
<emphasis italics="true" pageId="0" pageNumber="1">B. moulmeinica</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<typeStatus>holotype</typeStatus>
CUMZ 14216 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<typeStatus>paratype</typeStatus>
CUMZ 14217.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Limestone outcrop at Waiponla Monastery, Hpa-An, Kayin State, Myanmar (
<geoCoordinate degrees="16" direction="north" minutes="56" orientation="latitude" precision="1" seconds="07.4" value="16.935389">16°56'07.4&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="97" direction="east" minutes="42" orientation="longitude" precision="1" seconds="56.8" value="97.715775">97°42'56.8&quot;E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell globose and milky-white with a yellow tinge, rarely with a faintly yellowish brown peripheral band. Animal pale fleshy grey with brown or yellow tinge and five mantle extensions. Genitalia with rather short epiphallic caecum attached by a very large and thick penial retractor muscle and rather short vagina.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
(Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells of Burmochlamys species A, B B. albida sp. nov. A holotype CUMZ 14211, and D paratype CUMZ 14212 C, D B. fasciola sp. nov. C holotype CUMZ 14213 D paratype CUMZ 14214 E, F B. moulmeinica sp. nov. E holotype CUMZ 14216 F paratype CUMZ 14217." figureDoi="10.3897/zookeys.1110.82461.figure7" httpUri="https://binary.pensoft.net/fig/709573" pageId="0" pageNumber="1">7A, B</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Representative SEM images of the shell of Burmochlamys species A-C B. cassidula specimen CUMZ 14209 A protoconch B body whorl C zoom-in view of body whorl D-F B. poongee specimen CUMZ 14210 D protoconch E body whorl F zoom-in view of body whorl G-I B. albida sp. nov. paratype CUMZ 14212 G protoconch H body whorl I zoom-in view of body whorl J-L B. fasciola sp. nov. paratype CUMZ 14214 J protoconch K body whorl L zoom-in view of body whorl." figureDoi="10.3897/zookeys.1110.82461.figure8" httpUri="https://binary.pensoft.net/fig/709574" pageId="0" pageNumber="1">8G-I</figureCitation>
). Shell globose, small (width up to 4.5 mm, height up to 3.7 mm), thin, semi-translucent. Colour milky-white with a yellow tinge and without or rarely with faintly yellowish brown band above periphery. Protoconch and teleoconch surface with similar sculpture. Surface of body whorl with little distinct spiral furrows at regular intervals, crossed by very faintly undulating radial lines (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Representative SEM images of the shell of Burmochlamys species A-C B. cassidula specimen CUMZ 14209 A protoconch B body whorl C zoom-in view of body whorl D-F B. poongee specimen CUMZ 14210 D protoconch E body whorl F zoom-in view of body whorl G-I B. albida sp. nov. paratype CUMZ 14212 G protoconch H body whorl I zoom-in view of body whorl J-L B. fasciola sp. nov. paratype CUMZ 14214 J protoconch K body whorl L zoom-in view of body whorl." figureDoi="10.3897/zookeys.1110.82461.figure8" httpUri="https://binary.pensoft.net/fig/709574" pageId="0" pageNumber="1">8G-I</figureCitation>
). Whorls 6- 
<normalizedToken originalValue="6½">61/2</normalizedToken>
, increasing regularly; suture shallowly impressed; spire much elevated; last whorl well-rounded. Aperture obliquely crescent-shaped; peristome simple. Columellar margin simple, slightly reflected near umbilicus. Umbilicus open, narrow, and deep.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Genital organs</emphasis>
(Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. General view of the genital system of Burmochlamys species A B. cassidula specimen CUMZ 14209 B B. poongee specimen CUMZ 14210 C B. albida sp. nov. paratype CUMZ 14212. White arrow indicates the end of the penis. Red-dotted line indicates vas deferens missing." figureDoi="10.3897/zookeys.1110.82461.figure9" httpUri="https://binary.pensoft.net/fig/709575" pageId="0" pageNumber="1">9C</figureCitation>
). Atrium (at) rather short. Penis (p) moderate, cylindrical, and with penial sheath covering more than half of its length. Epiphallus (e1+e2) ca. three-sevenths of penis length; e1 cylindrical and small diameter smaller than penis and e2; e2 bulbous and ca. half of e1 length. Epiphallic caecum (ec) rather short, cylindrical, and with very large and thick penial retractor muscle (prm) attached at tip. Vas deferens (vd) thin tube. Dart apparatus large, long cylindrical, and located on atrium close to genital orifice. Vagina (v) rather short, cylindrical-shaped. Gametolytic sac (gs) bulbous; gametolytic duct (gd) cylindrical, moderate, ca. half of penis length. Free oviduct (fo) ca. one-quarter of penis length, cylindrical, and encircled with thick tissue near vagina.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" lsidName="Radula" order="Porellales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Radula</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Representative SEM images of the radula of Burmochlamys species A B. cassidula specimen CUMZ 14209 B B. poongee specimen CUMZ 14210 C B. albida sp. nov. paratype CUMZ 14212 D B. fasciola sp. nov. paratype CUMZ 14214. Central tooth indicated by ' C'; yellow colour indicates central tooth row." figureDoi="10.3897/zookeys.1110.82461.figure11" httpUri="https://binary.pensoft.net/fig/709577" pageId="0" pageNumber="1">11C</figureCitation>
). Generally resembles 
<taxonomicName genus="B." lsidName="B. cassidula" pageId="0" pageNumber="1" rank="species" species="cassidula">
<emphasis italics="true" pageId="0" pageNumber="1">B. cassidula</emphasis>
</taxonomicName>
. Teeth arranged in wide U-shaped rows, each row consisting of ~ 55 teeth. Central tooth monocuspid, large, and ovate spatulate shape; with two smaller triangular-shaped teeth. Laterals and marginals not differentiated, monocuspid, large spatulate shape and then gradually become narrower, elongate, rather more pointed cusp, and with one smaller triangular-shaped tooth. Some outermost teeth with a small, pointed cusp at the inner side.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">External appearance</emphasis>
(Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Living snails of Burmochlamys species A B. fasciola sp. nov. paratype CUMZ 14214 showing left and right lateral views B B. cassidula specimen CUMZ 14209 C B. poongee specimen CUMZ 14210 D B. albida sp. nov. paratype CUMZ 14212 E B. moulmeinica sp. nov. paratype CUMZ 14217 F B. versicolor sp. nov. paratype CUMZ 14219 G B. whitteni sp. nov. paratype CUMZ 14222." figureDoi="10.3897/zookeys.1110.82461.figure5" httpUri="https://binary.pensoft.net/fig/709571" pageId="0" pageNumber="1">5D</figureCitation>
). Living animal with reticulated skin, pale freshy-grey with a brown or yellow tinge. Five well-developed mantle extensions, with similar colour to body. Sole divided into three parts longitudinally; caudal foss and caudal horn well developed, similar colour to body.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet 
<emphasis italics="true" pageId="0" pageNumber="1">albida</emphasis>
is from the Latin word 
<emphasis italics="true" pageId="0" pageNumber="1">albidus</emphasis>
meaning white. It refers to the whitish shell, which characterises this species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys albida" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="albida">
<emphasis italics="true" pageId="0" pageNumber="1">Burmochlamys albida</emphasis>
</taxonomicName>
sp. nov. is endemic to a limestone outcrop at Waiponla Monastery. The surrounding paddy fields usually become flooded during the monsoon season (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the Salween River basin (known as Thanlwin River) in the southeastern Myanmar showing the sampling sites. Coloured triangles indicate the localities recorded for each Burmochlamys species. White circles indicate the localities recorded for Sophina species. White squares indicate the sampling localities with no records of both Sophina and Burmochlamys species." figureDoi="10.3897/zookeys.1110.82461.figure1" httpUri="https://binary.pensoft.net/fig/709567" pageId="0" pageNumber="1">1</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitat of some species of Burmochlamys gen. nov. in the karst basin of Hpa-An, Kayin State, Myanmar. A type locality of B. fasciola sp. nov. at Bardai Mountain and microhabitat structure of the karst wall B newly discovered locality of B. poongee at Kaw Ka Thaung Cave C type locality of B. albida sp. nov. at Waiponla Monastery D type locality of B. moulmeinica sp. nov. at Lun Nya Pagoda E type locality of B. versicolor sp. nov. at Bayin Nyi Cave. Photos by Ruttapon Srisonchai." figureDoi="10.3897/zookeys.1110.82461.figure2" httpUri="https://binary.pensoft.net/fig/709568" pageId="0" pageNumber="1">2C</figureCitation>
). In addition, this new species is also sympatric with the limestone karst-restricted species, 
<taxonomicName authorityName="Sutcharit &amp; Panha" authorityYear="2020" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" lsidName="Sophina salweenica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="salweenica">
<emphasis italics="true" pageId="0" pageNumber="1">Sophina salweenica</emphasis>
</taxonomicName>
Sutcharit &amp; Panha, 2020.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Among the whitish-shelled species (see Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of shell, radula, genital system, and distribution of all members of Burmochlamys gen. nov. in Myanmar." httpUri="http://table.plazi.org/id/A3EF01A52C7ECD353E45707D9F7BAEB8" pageId="0" pageNumber="1" tableUuid="A3EF01A52C7ECD353E45707D9F7BAEB8">2</tableCitation>
), 
<taxonomicName genus="B." lsidName="B. albida" pageId="0" pageNumber="1" rank="species" species="albida">
<emphasis italics="true" pageId="0" pageNumber="1">B. albida</emphasis>
</taxonomicName>
sp. nov. has a very indistinctly narrow yellowish brown peripheral band, whereas 
<taxonomicName genus="B." lsidName="B. cassidula" pageId="0" pageNumber="1" rank="species" species="cassidula">
<emphasis italics="true" pageId="0" pageNumber="1">B. cassidula</emphasis>
</taxonomicName>
has a wide yellowish brown peripheral band, 
<taxonomicName genus="B." lsidName="B. fasciola" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis italics="true" pageId="0" pageNumber="1">B. fasciola</emphasis>
</taxonomicName>
sp. nov. has a narrow yellowish brown peripheral band, and 
<taxonomicName genus="B." lsidName="B. versicolor" pageId="0" pageNumber="1" rank="species" species="versicolor">
<emphasis italics="true" pageId="0" pageNumber="1">B. versicolor</emphasis>
</taxonomicName>
sp. nov. has yellowish brown body whorl near the aperture and wider umbilicus than the other preceding species. Anatomically, 
<taxonomicName genus="B." lsidName="B. albida" pageId="0" pageNumber="1" rank="species" species="albida">
<emphasis italics="true" pageId="0" pageNumber="1">B. albida</emphasis>
</taxonomicName>
sp. nov. has a large and thick penial retractor muscle, rather slender vagina, and simple at the tip of dart apparatus. In comparison, 
<taxonomicName genus="B." lsidName="B. cassidula" pageId="0" pageNumber="1" rank="species" species="cassidula">
<emphasis italics="true" pageId="0" pageNumber="1">B. cassidula</emphasis>
</taxonomicName>
has a thin penial retractor muscle, large and short vagina, and soft at the tip of dart apparatus, while 
<taxonomicName genus="B." lsidName="B. fasciola" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis italics="true" pageId="0" pageNumber="1">B. fasciola</emphasis>
</taxonomicName>
sp. nov. has a thin penial retractor muscle, very short vagina, and solid at the tip of dart apparatus. In addition, 
<taxonomicName genus="B." lsidName="B. versicolor" pageId="0" pageNumber="1" rank="species" species="versicolor">
<emphasis italics="true" pageId="0" pageNumber="1">B. versicolor</emphasis>
</taxonomicName>
sp. nov. has a thin penial retractor muscle, rather slender vagina, and relatively smaller dart apparatus with soft at the tip.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys albida" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="albida">
<emphasis italics="true" pageId="0" pageNumber="1">Burmochlamys albida</emphasis>
</taxonomicName>
sp. nov. differs from the remaining 
<taxonomicName authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
species by having a milky-white shell. While most other species have a pale to dark brownish shell without a peripheral band (see Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of shell, radula, genital system, and distribution of all members of Burmochlamys gen. nov. in Myanmar." httpUri="http://table.plazi.org/id/A3EF01A52C7ECD353E45707D9F7BAEB8" pageId="0" pageNumber="1" tableUuid="A3EF01A52C7ECD353E45707D9F7BAEB8">2</tableCitation>
). Regardless of the shell colour and genitalia, this new species differs from 
<taxonomicName genus="B." lsidName="B. cauisa" pageId="0" pageNumber="1" rank="species" species="cauisa">
<emphasis italics="true" pageId="0" pageNumber="1">B. cauisa</emphasis>
</taxonomicName>
and 
<taxonomicName genus="B." lsidName="B. perpaula" pageId="0" pageNumber="1" rank="species" species="perpaula">
<emphasis italics="true" pageId="0" pageNumber="1">B. perpaula</emphasis>
</taxonomicName>
by spire, body whorl, and aperture. In comparison, 
<taxonomicName genus="B." lsidName="B. cauisa" pageId="0" pageNumber="1" rank="species" species="cauisa">
<emphasis italics="true" pageId="0" pageNumber="1">B. cauisa</emphasis>
</taxonomicName>
has slightly elevated spire, well-rounded last whorl, and rather broad crescent-shaped aperture, while 
<taxonomicName genus="B." lsidName="B. perpaula" pageId="0" pageNumber="1" rank="species" species="perpaula">
<emphasis italics="true" pageId="0" pageNumber="1">B. perpaula</emphasis>
</taxonomicName>
has slightly elevated spire, rather more convex body whorl, and narrower crescent-shaped aperture.
</paragraph>
</subSubSection>
</treatment>
</document>