treatments-xml/data/03/C9/AC/03C9AC691E4EFF8B649FFA47B62FEE0D.xml
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<document id="A4B3B340F7A22A8058FFAA05D949E822" ID-CLB-Dataset="26462" ID-DOI="10.11646/zootaxa.4619.1.1" ID-GBIF-Dataset="9aaf6711-748d-4067-a759-26d37802b7fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3248363" ID-ZooBank="CA6AEB18-2F97-449C-AE34-E1509DFFC841" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560832679272" checkinUser="plazi" docAuthor="Lee, Bee Yan, Forges, Bertrand Richer De &amp; Ng, Peter K. L." docDate="2019" docId="03C9AC691E4EFF8B649FFA47B62FEE0D" docLanguage="en" docName="zootaxa.4619.1.1.pdf" docOrigin="Zootaxa 4619 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crocydocinus vanuatu Lee &amp; Forges &amp; Ng 2019, n. sp." docType="treatment" docVersion="16" lastPageNumber="34" masterDocId="FFF0D4111E6EFFAA6408FFAEB427EC58" masterDocTitle="Deep-sea spider crabs of the family Epialtidae MacLeay, 1838, from Papua New Guinea, with a redefinition of Tunepugettia Ng, Komai &amp; Sato, 2017, and descriptions of two new genera (Crustacea: Decapoda: Brachyura: Majoidea)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="33" updateTime="1698735225898" updateUser="ExternalLinkService">
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<mods:title id="4C41EB9113171592CBD6856B8DF26410">Deep-sea spider crabs of the family Epialtidae MacLeay, 1838, from Papua New Guinea, with a redefinition of Tunepugettia Ng, Komai &amp; Sato, 2017, and descriptions of two new genera (Crustacea: Decapoda: Brachyura: Majoidea)</mods:title>
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<mods:namePart id="3E247CC1F1BB11D1A033A928552A73D0">Lee, Bee Yan</mods:namePart>
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<mods:namePart id="F33601CF2273B513A9E764CA983FD1FF">Forges, Bertrand Richer De</mods:namePart>
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<mods:namePart id="CD2D8BD0FB1EE0BB1EC04D829901C83B">Ng, Peter K. L.</mods:namePart>
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<treatment id="03C9AC691E4EFF8B649FFA47B62FEE0D" ID-GBIF-Taxon="158290204" LSID="urn:lsid:plazi:treatment:03C9AC691E4EFF8B649FFA47B62FEE0D" httpUri="http://treatment.plazi.org/id/03C9AC691E4EFF8B649FFA47B62FEE0D" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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<paragraph id="8BDF1D7F1E4EFF8A649FFA47B5D5EA5B" blockId="32.[151,498,1513,1575]" box="[151,498,1513,1540]" pageId="32" pageNumber="33">
<heading id="D097AA131E4EFF8A649FFA47B5D5EA5B" bold="true" box="[151,498,1513,1540]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="B914C16D1E4EFF8A649FFA47B5D5EA5B" bold="true" box="[151,498,1513,1540]" pageId="32" pageNumber="33">
<taxonomicName id="4C6066FC1E4EFF8A649FFA47B58FEA5C" ID-CoL="84DVT" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[151,424,1513,1540]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="B914C16D1E4EFF8A649FFA47B58FEA5C" bold="true" box="[151,424,1513,1540]" italics="true" pageId="32" pageNumber="33">Crocydocinus vanuatu</emphasis>
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<taxonomicNameLabel id="A2277C161E4EFF8A65B8FA47B5D5EA5B" box="[432,498,1513,1539]" pageId="32" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C37A4EF41E4EFF8A649FF9A3B5C8EA7F" box="[151,495,1549,1575]" pageId="32" pageNumber="33" type="description">
<paragraph id="8BDF1D7F1E4EFF8A649FF9A3B5C8EA7F" blockId="32.[151,498,1513,1575]" box="[151,495,1549,1575]" pageId="32" pageNumber="33">
(
<figureCitation id="135B01FA1E4EFF8A6497F9A3B537EA7F" box="[159,272,1549,1575]" captionStart="FIGURE 14" captionStartId="30.[151,250,1652,1677]" captionTargetBox="[151,1436,474,1633]" captionTargetId="figure@30.[151,1436,474,1633]" captionTargetPageId="30" captionText="FIGURE 14. Overall dorsal view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248391/files/figure.png" pageId="32" pageNumber="33">Figs. 14C</figureCitation>
,
<figureCitation id="135B01FA1E4EFF8A6513F9A3B56BEA7F" box="[283,332,1549,1575]" captionStart="FIGURE 16" captionStartId="32.[151,250,1334,1359]" captionTargetBox="[151,1436,186,1315]" captionTargetId="figure@32.[151,1436,186,1315]" captionTargetPageId="32" captionText="FIGURE 16. Overall ventral view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248395/files/figure.png" pageId="32" pageNumber="33">16C</figureCitation>
,
<figureCitation id="135B01FA1E4EFF8A6550F9A3B5ACEA7F" box="[344,395,1549,1575]" captionStart="FIGURE 17" captionStartId="33.[151,250,1804,1829]" captionTargetBox="[184,1405,624,1785]" captionTargetId="figure@33.[184,1405,624,1786]" captionTargetPageId="33" captionText="FIGURE 17. Lateral view of carapace.A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Ile de la Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam; E, C. panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; F, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; G, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248397/files/figure.png" pageId="32" pageNumber="33">17G</figureCitation>
,
<figureCitation id="135B01FA1E4EFF8A659EF9A3B597EA7F" box="[406,432,1549,1575]" captionStart="FIGURE 19" captionStartId="36.[151,250,1890,1915]" captionTargetBox="[281,1307,176,1860]" captionTargetId="figure@36.[243,1344,166,1870]" captionTargetPageId="36" captionText="FIGURE 19. AD, Crocydocinus decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam, left G1. A, ventral view; B, ventral view of distal portion; C, dorsal view; D, dorsal view of distal portion. EH, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu, left G1. E, ventral view; F, ventral view of distal portion; G, dorsal view; H, dorsal view of distal portion." httpUri="https://zenodo.org/record/3248401/files/figure.png" pageId="32" pageNumber="33">19</figureCitation>
EH)
</paragraph>
</subSubSection>
<subSubSection id="C37A4EF41E4EFF8A649FF9E2B130EA3C" box="[151,1303,1611,1637]" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="8BDF1D7F1E4EFF8A649FF9E2B130EA3C" blockId="32.[151,1303,1611,1637]" box="[151,1303,1611,1637]" pageId="32" pageNumber="33">
<treatmentCitationGroup id="AB703A511E4EFF8A649FF9E2B130EA3C" box="[151,1303,1611,1637]" pageId="32" pageNumber="33">
<taxonomicName id="4C6066FC1E4EFF8A649FF9E2B72EEA3C" ID-CoL="7YTGL" authority="- Richer de Forges &amp; Ng 2013: 473" authorityName="Richer de Forges &amp; Ng" authorityPageNumber="473" authorityYear="2013" box="[151,777,1611,1637]" class="Malacostraca" family="Epialtidae" genus="Rochinia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="B914C16D1E4EFF8A649FF9E2B4D5EA3C" box="[151,242,1612,1636]" italics="true" pageId="32" pageNumber="33">Rochinia</emphasis>
aff.
<emphasis id="B914C16D1E4EFF8A6529F9E2B5B2EA3C" box="[289,405,1612,1636]" italics="true" pageId="32" pageNumber="33">brevirostris</emphasis>
<treatmentCitation id="0AC13B6E1E4EFF8A65A7F9E5B72EEA3C" author="Richer de Forges, B. &amp; Ng, P. K. L." box="[431,777,1611,1637]" page="473" pageId="32" pageNumber="33" year="2013">
<bibRefCitation id="EFF1608E1E4EFF8A65A7F9E5B72EEA3C" author="Richer de Forges, B. &amp; Ng, P. K. L." box="[431,777,1611,1637]" pageId="32" pageNumber="33" pagination="467 - 483" refId="ref30088" refString="Richer de Forges, B. &amp; Ng, P. K. L. (2013) On a collection of spider crabs of the genera Rochinia A. Milne-Edwards, 1875 and Naxioides A. Milne-Edwards, 1865 (Crustacea, Brachyura, Majoidea, Epialtidae) from Mozambique Channel, Solomon, Vanuatu and Philippine Islands, with description of a new species of Rochinia. In: Ahyong, S. T., Chan, T. Y., Corbari, L. &amp; Ng, P. K. L. (Eds.), Tropical Deep-sea Benthos 27. Museum national d'Histoire naturelle, Paris, pp. 467 - 483." type="book chapter" year="2013">Richer de Forges &amp; Ng 2013: 473</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, fig. 4A [not
<taxonomicName id="4C6066FC1E4EFF8A6798F9E3B12BEA3C" ID-CoL="84FWM" authority="Doflein, 1904" authorityName="Doflein" authorityYear="1904" box="[912,1292,1611,1636]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="B914C16D1E4EFF8A6798F9E3B053EA3C" box="[912,1140,1612,1636]" italics="true" pageId="32" pageNumber="33">Hyastenus brevirostris</emphasis>
<bibRefCitation id="EFF1608E1E4EFF8A6072F9E5B12BEA3C" author="Doflein, F." box="[1146,1292,1611,1636]" pageId="32" pageNumber="33" pagination="1 - 58" refId="ref27028" refString="Doflein, F. (1904) Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Vol. 6. Gustav Fischer, Jena, pp. 1 - 6 + i-xiv + 1 - 314 pp., figs. I-LVIII, atlas (pls. 1 - 58)." type="book chapter" year="1904">Doflein, 1904</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37A4EF41E4EFF8A649FF93CB792EA88" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BDF1D7F1E4EFF8A649FF93CB792EA88" blockId="32.[151,1437,1682,2033]" pageId="32" pageNumber="33">
<emphasis id="B914C16D1E4EFF8A649FF93CB558EAF5" bold="true" box="[151,383,1682,1709]" pageId="32" pageNumber="33">Material examined.</emphasis>
<materialsCitation id="3B0817221E4EFF8A658DF93DB792EA88" ID-GBIF-Occurrence="2268190909" collectingDate="2006-10-13" collectionCode="ZRC, SANTO" country="Vanuatu" county="Expedition" elevation="583" location="Big Bay" municipality="Espiritu Santo Island" pageId="32" pageNumber="33" specimenCode="ZRC 2011.1047, SANTO 2006" specimenCount="1" typeStatus="holotype">
<emphasis id="B914C16D1E4EFF8A658DF93DB5DCEAF5" bold="true" box="[389,507,1683,1709]" pageId="32" pageNumber="33">
<typeStatus id="54DBA3DD1E4EFF8A658DF93DB5D1EAF5" box="[389,502,1683,1709]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (16.0 × 11.0 mm) (
<specimenCode id="DBC6B5041E4EFF8A6704F93DB7E2EAF5" box="[780,965,1683,1709]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="32" pageNumber="33">ZRC 2011.1047</specimenCode>
), stn AT89,
<location id="8EBF4BA41E4EFF8A604EF93CB085EAF5" LSID="urn:lsid:plazi:treatment:03C9AC691E4EFF8B649FFA47B62FEE0D:8EBF4BA41E4EFF8A604EF93CB085EAF5" box="[1094,1186,1682,1709]" country="Vanuatu" county="Expedition" municipality="Espiritu Santo Island" name="Big Bay" pageId="32" pageNumber="33">Big Bay</location>
,
<collectingMunicipality id="6BBB87051E4EFF8A60A5F93CB1BFEAF4" box="[1197,1432,1682,1709]" pageId="32" pageNumber="33">Espiritu Santo Island</collectingMunicipality>
,
<collectingCountry id="F3775DEF1E4EFF8A649FF919B4D1EA89" box="[151,246,1719,1745]" name="Vanuatu" pageId="32" pageNumber="33">Vanuatu</collectingCountry>
, 630
<elevation id="004DFA4C1E4EFF8A6532F918B5A5EA89" box="[314,386,1718,1745]" metricMagnitude="2" metricUnit="m" metricValue="5.83" pageId="32" pageNumber="33" unit="m" value="583.0">
<quantity id="4C98B09A1E4EFF8A6532F918B5A5EA89" box="[314,386,1718,1745]" metricMagnitude="2" metricUnit="m" metricValue="5.83" pageId="32" pageNumber="33" unit="m" value="583.0">583 m</quantity>
</elevation>
, coll.
<specimenCode id="DBC6B5041E4EFF8A65CEF919B645EA88" box="[454,610,1718,1745]" collectionCode="SANTO" pageId="32" pageNumber="33">SANTO 2006</specimenCode>
<collectingCounty id="62BE65F31E4EFF8A6661F918B6C0EA88" box="[617,743,1718,1744]" pageId="32" pageNumber="33">Expedition</collectingCounty>
,
<collectingDate id="EF9AC2571E4EFF8A66F9F919B795EA88" box="[753,946,1718,1745]" pageId="32" pageNumber="33" value="2006-10-13">
<date id="FFDE3BBF1E4EFF8A66F9F919B795EA88" box="[753,946,1718,1745]" pageId="32" pageNumber="33" value="2006-10-13">13 October 2006</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37A4EF41E4EFF8B64CFF974B0CDEC8E" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BDF1D7F1E4EFF8B64CFF974B0CDEC8E" blockId="32.[151,1437,1682,2033]" lastBlockId="33.[151,1436,153,597]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="B914C16D1E4EFF8A64CFF974B565EAAD" bold="true" box="[199,322,1754,1781]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
Carapace pyriform, covered with thick layer of setae, surface smooth when denuded (
<figureCitation id="135B01FA1E4EFF8A612EF974B1B7EAAD" box="[1318,1424,1754,1781]" captionStart="FIGURE 14" captionStartId="30.[151,250,1652,1677]" captionTargetBox="[151,1436,474,1633]" captionTargetId="figure@30.[151,1436,474,1633]" captionTargetPageId="30" captionText="FIGURE 14. Overall dorsal view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248391/files/figure.png" pageId="32" pageNumber="33">Fig. 14C</figureCitation>
). Pseudorostral spines straight, thick, short, diverging, V-shaped. Supraorbital eave fused to carapace; pre-orbital angle blunt; postorbital lobe cup-like, fused with hepatic spine, base swollen. Carapace with regions defined; gastric region swollen; epibranchial region slight swell; posterior region with small granule medially (
<figureCitation id="135B01FA1E4EFF8A60B8F8E8B131EB39" box="[1200,1302,1862,1889]" captionStart="FIGURE 14" captionStartId="30.[151,250,1652,1677]" captionTargetBox="[151,1436,474,1633]" captionTargetId="figure@30.[151,1436,474,1633]" captionTargetPageId="30" captionText="FIGURE 14. Overall dorsal view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248391/files/figure.png" pageId="32" pageNumber="33">Fig. 14C</figureCitation>
); 3 granules along lateral margin of carapace on margin of branchial region (
<figureCitation id="135B01FA1E4EFF8A676EF8C4B7E8EBDD" box="[870,975,1898,1925]" captionStart="FIGURE 17" captionStartId="33.[151,250,1804,1829]" captionTargetBox="[184,1405,624,1785]" captionTargetId="figure@33.[184,1405,624,1786]" captionTargetPageId="33" captionText="FIGURE 17. Lateral view of carapace.A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Ile de la Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam; E, C. panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; F, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; G, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248397/files/figure.png" pageId="32" pageNumber="33">Fig. 17G</figureCitation>
). Antennal flagellum slightly longer than pseudorostral spines. Basal antennal article longer than broad, distal angle rounded. Buccal frame covered by third maxilliped. Pterygostomial region with 3 granules on outer margin (
<figureCitation id="135B01FA1E4EFF8A678DF81CB7CDEB95" box="[901,1002,1970,1997]" captionStart="FIGURE 16" captionStartId="32.[151,250,1334,1359]" captionTargetBox="[151,1436,186,1315]" captionTargetId="figure@32.[151,1436,186,1315]" captionTargetPageId="32" captionText="FIGURE 16. Overall ventral view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248395/files/figure.png" pageId="32" pageNumber="33">Fig. 16C</figureCitation>
). Chelipeds slender, covered with setae except on fingers. Ambulatory legs slender, covered with layer of setae except on dactylus; P2 longest. Male tho- racic sternum depressed anteriorly; sternites 3 and 4 with concave surface, margin constricted with curved margin (
<figureCitation id="135B01FA1E4FFF8B64A8FF15B521EC8E" box="[160,262,187,214]" captionStart="FIGURE 16" captionStartId="32.[151,250,1334,1359]" captionTargetBox="[151,1436,186,1315]" captionTargetId="figure@32.[151,1436,186,1315]" captionTargetPageId="32" captionText="FIGURE 16. Overall ventral view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248395/files/figure.png" pageId="33" pageNumber="34">Fig. 16C</figureCitation>
). Male pleon broad triangular (
<figureCitation id="135B01FA1E4FFF8B666BFF15B6EEEC8E" box="[611,713,187,214]" captionStart="FIGURE 16" captionStartId="32.[151,250,1334,1359]" captionTargetBox="[151,1436,186,1315]" captionTargetId="figure@32.[151,1436,186,1315]" captionTargetPageId="32" captionText="FIGURE 16. Overall ventral view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248395/files/figure.png" pageId="33" pageNumber="34">Fig. 16C</figureCitation>
). G1 straight, with distal tip sharp (
<figureCitation id="135B01FA1E4FFF8B605DFF15B083EC8E" box="[1109,1188,187,214]" captionStart="FIGURE 19" captionStartId="36.[151,250,1890,1915]" captionTargetBox="[281,1307,176,1860]" captionTargetId="figure@36.[243,1344,166,1870]" captionTargetPageId="36" captionText="FIGURE 19. AD, Crocydocinus decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam, left G1. A, ventral view; B, ventral view of distal portion; C, dorsal view; D, dorsal view of distal portion. EH, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu, left G1. E, ventral view; F, ventral view of distal portion; G, dorsal view; H, dorsal view of distal portion." httpUri="https://zenodo.org/record/3248401/files/figure.png" pageId="33" pageNumber="34">Fig. 19</figureCitation>
EH).
</paragraph>
</subSubSection>
<subSubSection id="C37A4EF41E4FFF8B64CFFF70B5FCED43" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BDF1D7F1E4FFF8B64CFFF70B5FCED43" blockId="33.[151,1436,153,597]" pageId="33" pageNumber="34">
<emphasis id="B914C16D1E4FFF8B64CFFF70B569ECA0" bold="true" box="[199,334,222,248]" pageId="33" pageNumber="34">Etymology.</emphasis>
The species is named after the country from which the specimen was collected from. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C37A4EF41E4FFF8B64CFFE8AB62FEE0D" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BDF1D7F1E4FFF8B64CFFE8AB62FEE0D" blockId="33.[151,1436,153,597]" pageId="33" pageNumber="34">
<emphasis id="B914C16D1E4FFF8B64CFFE8AB51CED66" bold="true" box="[199,315,292,318]" pageId="33" pageNumber="34">Remarks.</emphasis>
<taxonomicName id="4C6066FC1E4FFF8B654AFE8AB619ED65" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[322,574,292,318]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="B914C16D1E4FFF8B654AFE8AB619ED65" box="[322,574,292,318]" italics="true" pageId="33" pageNumber="34">Crocydocinus vanuatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E4FFF8B664DFE8AB6A3ED66" box="[581,644,292,318]" pageId="33" pageNumber="34" rank="species">
<emphasis id="B914C16D1E4FFF8B664DFE8AB6A3ED66" bold="true" box="[581,644,292,318]" pageId="33" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
was identified as
<taxonomicName id="4C6066FC1E4FFF8B6747FE8AB03BED66" authorityName="Doflein" authorityYear="1904" box="[847,1052,291,318]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="B914C16D1E4FFF8B6747FE8AB747ED65" box="[847,864,292,317]" italics="true" pageId="33" pageNumber="34">R</emphasis>
. aff.
<emphasis id="B914C16D1E4FFF8B6793FE8AB03BED66" box="[923,1052,292,318]" italics="true" pageId="33" pageNumber="34">brevirostris</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF1608E1E4FFF8B604DFE8DB1BBED66" author="Richer de Forges, B. &amp; Ng, P. K. L." box="[1093,1436,291,318]" pageId="33" pageNumber="34" pagination="467 - 483" refId="ref30088" refString="Richer de Forges, B. &amp; Ng, P. K. L. (2013) On a collection of spider crabs of the genera Rochinia A. Milne-Edwards, 1875 and Naxioides A. Milne-Edwards, 1865 (Crustacea, Brachyura, Majoidea, Epialtidae) from Mozambique Channel, Solomon, Vanuatu and Philippine Islands, with description of a new species of Rochinia. In: Ahyong, S. T., Chan, T. Y., Corbari, L. &amp; Ng, P. K. L. (Eds.), Tropical Deep-sea Benthos 27. Museum national d'Histoire naturelle, Paris, pp. 467 - 483." type="book chapter" year="2013">Richer de Forges &amp; Ng (2013)</bibRefCitation>
but they noted it differed from the Bay of Bengal specimen as illustrated by Griffin &amp; Tranter (1986: fig. 57c) in several aspects. Comparison of the present material with the
<typeStatus id="54DBA3DD1E4FFF8B673FFEC7B740EDDB" box="[823,871,361,387]" pageId="33" pageNumber="34">type</typeStatus>
specimen of
<taxonomicName id="4C6066FC1E4FFF8B67F2FEC7B169EDDB" authority="(Doflein, 1904)" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Doflein" baseAuthorityYear="1904" box="[1018,1358,361,387]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="brevirostris" status="comb. nov.">
<emphasis id="B914C16D1E4FFF8B67F2FEC7B02AEDDB" box="[1018,1037,361,387]" italics="true" pageId="33" pageNumber="34">C</emphasis>
.
<emphasis id="B914C16D1E4FFF8B6012FEC7B0BCEDDB" box="[1050,1179,361,387]" italics="true" pageId="33" pageNumber="34">brevirostris</emphasis>
(
<bibRefCitation id="EFF1608E1E4FFF8B60A0FEC7B161EDDB" author="Doflein, F." box="[1192,1350,361,387]" pageId="33" pageNumber="34" pagination="1 - 58" refId="ref27028" refString="Doflein, F. (1904) Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Vol. 6. Gustav Fischer, Jena, pp. 1 - 6 + i-xiv + 1 - 314 pp., figs. I-LVIII, atlas (pls. 1 - 58)." type="book chapter" year="1904">Doflein, 1904</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2277C161E4FFF8B615CFEC7B4EFEDFE" pageId="33" pageNumber="34" rank="species">
<emphasis id="B914C16D1E4FFF8B615CFEC7B4EFEDFE" bold="true" pageId="33" pageNumber="34">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="135B01FA1E4FFF8B64DFFE25B560EDFE" box="[215,327,395,422]" captionStart="FIGURE 13" captionStartId="25.[151,250,1863,1888]" captionTargetBox="[151,1436,690,1844]" captionTargetId="figure@25.[151,1436,690,1844]" captionTargetPageId="25" captionText="FIGURE 13. Overall dorsal view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248389/files/figure.png" pageId="33" pageNumber="34">Figs. 13B</figureCitation>
,
<figureCitation id="135B01FA1E4FFF8B655AFE25B5A4EDFE" box="[338,387,395,422]" captionStart="FIGURE 15" captionStartId="31.[151,250,1906,1931]" captionTargetBox="[151,1436,751,1887]" captionTargetId="figure@31.[151,1436,751,1887]" captionTargetPageId="31" captionText="FIGURE 15. Overall ventral view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248393/files/figure.png" pageId="33" pageNumber="34">15B</figureCitation>
,
<figureCitation id="135B01FA1E4FFF8B6587FE22B5E7EDFE" box="[399,448,396,422]" captionStart="FIGURE 17" captionStartId="33.[151,250,1804,1829]" captionTargetBox="[184,1405,624,1785]" captionTargetId="figure@33.[184,1405,624,1786]" captionTargetPageId="33" captionText="FIGURE 17. Lateral view of carapace.A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Ile de la Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam; E, C. panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; F, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; G, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248397/files/figure.png" pageId="33" pageNumber="34">17B</figureCitation>
), revealed more differences, justifying treating them as two separate species.
<taxonomicName id="4C6066FC1E4FFF8B6117FE22B538ED90" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="B914C16D1E4FFF8B6117FE22B538ED90" italics="true" pageId="33" pageNumber="34">Crocydocinus vanuatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E4FFF8B652EFE00B543ED90" box="[294,356,430,456]" pageId="33" pageNumber="34" rank="species">
<emphasis id="B914C16D1E4FFF8B652EFE00B543ED90" bold="true" box="[294,356,430,456]" pageId="33" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
lacks all the distinct large granules (
<figureCitation id="135B01FA1E4FFF8B66F5FE00B746ED90" box="[765,865,430,456]" captionStart="FIGURE 14" captionStartId="30.[151,250,1652,1677]" captionTargetBox="[151,1436,474,1633]" captionTargetId="figure@30.[151,1436,474,1633]" captionTargetPageId="30" captionText="FIGURE 14. Overall dorsal view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248391/files/figure.png" pageId="33" pageNumber="34">Fig. 14C</figureCitation>
) seen in
<taxonomicName id="4C6066FC1E4FFF8B67CDFE01B040ED90" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Doflein" baseAuthorityYear="1904" box="[965,1127,430,457]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="brevirostris" status="comb. nov.">
<emphasis id="B914C16D1E4FFF8B67CDFE01B7FFED91" box="[965,984,431,457]" italics="true" pageId="33" pageNumber="34">C</emphasis>
.
<emphasis id="B914C16D1E4FFF8B67EEFE00B040ED90" box="[998,1127,430,456]" italics="true" pageId="33" pageNumber="34">brevirostris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E4FFF8B6065FE00B0CAED90" box="[1133,1261,430,456]" pageId="33" pageNumber="34" rank="species">
<emphasis id="B914C16D1E4FFF8B6065FE00B0CAED90" bold="true" box="[1133,1261,430,456]" pageId="33" pageNumber="34">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="135B01FA1E4FFF8B60F4FE00B145ED90" box="[1276,1378,430,456]" captionStart="FIGURE 13" captionStartId="25.[151,250,1863,1888]" captionTargetBox="[151,1436,690,1844]" captionTargetId="figure@25.[151,1436,690,1844]" captionTargetPageId="25" captionText="FIGURE 13. Overall dorsal view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248389/files/figure.png" pageId="33" pageNumber="34">Fig. 13B</figureCitation>
); has only weak lateral branchial spines (
<figureCitation id="135B01FA1E4FFF8B662BFE7FB6A0EDB3" box="[547,647,465,491]" captionStart="FIGURE 14" captionStartId="30.[151,250,1652,1677]" captionTargetBox="[151,1436,474,1633]" captionTargetId="figure@30.[151,1436,474,1633]" captionTargetPageId="30" captionText="FIGURE 14. Overall dorsal view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248391/files/figure.png" pageId="33" pageNumber="34">Fig. 14C</figureCitation>
) (versus distinct lateral branchial spines in
<taxonomicName id="4C6066FC1E4FFF8B6064FE7FB129EDB3" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Doflein" baseAuthorityYear="1904" box="[1132,1294,465,491]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="brevirostris" status="comb. nov.">
<emphasis id="B914C16D1E4FFF8B6064FE7FB058EDB3" box="[1132,1151,465,491]" italics="true" pageId="33" pageNumber="34">C</emphasis>
.
<emphasis id="B914C16D1E4FFF8B6085FE7FB129EDB3" box="[1165,1294,465,491]" italics="true" pageId="33" pageNumber="34">brevirostris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E4FFF8B611CFE7FB1B3EDB3" box="[1300,1428,465,491]" pageId="33" pageNumber="34" rank="species">
<emphasis id="B914C16D1E4FFF8B611CFE7FB1B3EDB3" bold="true" box="[1300,1428,465,491]" pageId="33" pageNumber="34">comb. nov.</emphasis>
</taxonomicNameLabel>
;
<figureCitation id="135B01FA1E4FFF8B649FFE5DB4DAEE56" box="[151,253,499,526]" captionStart="FIGURE 13" captionStartId="25.[151,250,1863,1888]" captionTargetBox="[151,1436,690,1844]" captionTargetId="figure@25.[151,1436,690,1844]" captionTargetPageId="25" captionText="FIGURE 13. Overall dorsal view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248389/files/figure.png" pageId="33" pageNumber="34">Fig. 13B</figureCitation>
); and the basal antennal article has a straight outer margin (
<figureCitation id="135B01FA1E4FFF8B67ADFE5DB02DEE56" box="[933,1034,499,526]" captionStart="FIGURE 16" captionStartId="32.[151,250,1334,1359]" captionTargetBox="[151,1436,186,1315]" captionTargetId="figure@32.[151,1436,186,1315]" captionTargetPageId="32" captionText="FIGURE 16. Overall ventral view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248395/files/figure.png" pageId="33" pageNumber="34">Fig. 16C</figureCitation>
) (versus basal antennal article with slightly convex outer margin in
<taxonomicName id="4C6066FC1E4FFF8B65F7FDB9B686EE68" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Doflein" baseAuthorityYear="1904" box="[511,673,534,561]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="brevirostris" status="comb. nov.">
<emphasis id="B914C16D1E4FFF8B65F7FDB9B635EE69" box="[511,530,535,561]" italics="true" pageId="33" pageNumber="34">C</emphasis>
.
<emphasis id="B914C16D1E4FFF8B6628FDB8B686EE68" box="[544,673,534,560]" italics="true" pageId="33" pageNumber="34">brevirostris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E4FFF8B66A0FDB8B70EEE68" box="[680,809,534,560]" pageId="33" pageNumber="34" rank="species">
<emphasis id="B914C16D1E4FFF8B66A0FDB8B70EEE68" bold="true" box="[680,809,534,560]" pageId="33" pageNumber="34">comb. nov.</emphasis>
</taxonomicNameLabel>
;
<figureCitation id="135B01FA1E4FFF8B6730FDB8B7B9EE68" box="[824,926,533,560]" captionStart="FIGURE 15" captionStartId="31.[151,250,1906,1931]" captionTargetBox="[151,1436,751,1887]" captionTargetId="figure@31.[151,1436,751,1887]" captionTargetPageId="31" captionText="FIGURE 15. Overall ventral view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248393/files/figure.png" pageId="33" pageNumber="34">Fig. 15B</figureCitation>
).
<taxonomicName id="4C6066FC1E4FFF8B67BAFDB8B089EE68" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[946,1198,534,560]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vanuatu" status="sp. nov.">
<emphasis id="B914C16D1E4FFF8B67BAFDB8B089EE68" box="[946,1198,534,560]" italics="true" pageId="33" pageNumber="34">Crocydocinus vanuatu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E4FFF8B60BEFDB8B0D2EE68" box="[1206,1269,534,560]" pageId="33" pageNumber="34" rank="species">
<emphasis id="B914C16D1E4FFF8B60BEFDB8B0D2EE68" bold="true" box="[1206,1269,534,560]" pageId="33" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
is only known from its
<typeStatus id="54DBA3DD1E4FFF8B64FEFD95B501EE0D" box="[246,294,571,597]" pageId="33" pageNumber="34">type</typeStatus>
locality in
<collectingCountry id="F3775DEF1E4FFF8B65ADFD95B622EE0D" box="[421,517,571,597]" name="Vanuatu" pageId="33" pageNumber="34">Vanuatu</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>