treatments-xml/data/FD/6E/47/FD6E4758FFD6587B457A506FEE5BE1B3.xml

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<mods:title id="7AD57D4A27DFEF5E4F64A1B5A747EDD6">Rhopalomenia glandulosa spec. nov., and the restoration of Entonomenia Leloup (Mollusca: Solenogastres)</mods:title>
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<subSubSection id="3DDDA5C5FFD6587D457A506FE821E366" box="[264,596,1548,1574]" pageId="10" pageNumber="53" type="nomenclature">
<paragraph id="7578F64EFFD6587D457A506FE821E366" blockId="10.[264,596,1548,1574]" box="[264,596,1548,1574]" pageId="10" pageNumber="53">
<heading id="2E304122FFD6587D457A506FE821E366" bold="true" box="[264,596,1548,1574]" fontSize="11" level="1" pageId="10" pageNumber="53" reason="1">
<taxonomicName id="B2C78DCDFFD6587D457A506FE821E366" ID-CoL="7NYSW" authority="Leloup, 1948" authorityName="Leloup." authorityYear="1948" box="[264,596,1548,1574]" class="Solenogastres" family="Rhopalomeniidae" genus="Entonomenia" kingdom="Animalia" order="Cavibelonia" pageId="10" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="47B32A5CFFD6587D457A506FE821E366" bold="true" box="[264,596,1548,1574]" pageId="10" pageNumber="53">
<emphasis id="47B32A5CFFD6587D457A506FEBDEE366" bold="true" box="[264,427,1548,1574]" italics="true" pageId="10" pageNumber="53">Entonomenia</emphasis>
<bibRefCitation id="11568BBFFFD6587D45C1506FE821E366" author="Leloup, E." box="[435,596,1548,1574]" pageId="10" pageNumber="53" pagination="1 - 11" refId="ref3938" refString="Leloup, E., 1948. Un nouveau solenogastre proneomeniide, Entonomenia atlantica g. nov. sp. nov. Bulletin du Museum royal d'Histoirs Naturelle Belgique, 24 (37), 1 - 11." type="journal article" year="1948">Leloup, 1948</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3DDDA5C5FFD6587C457A503FEF73E786" lastPageId="11" lastPageNumber="54" pageId="10" pageNumber="53" type="discussion">
<paragraph id="7578F64EFFD6587C457A503FE8CEE41E" blockId="10.[264,1322,1628,1894]" lastBlockId="11.[264,1323,284,1030]" lastPageId="11" lastPageNumber="54" pageId="10" pageNumber="53">
<taxonomicName id="B2C78DCDFFD6587D457A503FEBA0E336" authorityName="Salvini-Plawen" authorityYear="1978" box="[264,469,1628,1654]" class="Solenogastres" family="Rhopalomeniidae" kingdom="Animalia" order="Cavibelonia" pageId="10" pageNumber="53" phylum="Mollusca" rank="family">Rhopalomeniidae</taxonomicName>
with only one pair of foregut glandular organs, characterised by a duct with subepithelial/extraepithelial glandular cells (so­called type A); with separate mouth opening, without radula, with dorsoterminal sense organ, with unpaired secondary genital opening, without copulatory stylets or respiratory organs. Type species 
<taxonomicName id="B2C78DCDFFD6587D403150B7EBA8E256" authority="Leloup, 1948" authorityName="Leloup" authorityYear="1948" class="Solenogastres" family="Rhopalomeniidae" genus="Entonomenia" kingdom="Animalia" order="Cavibelonia" pageId="10" pageNumber="53" phylum="Mollusca" rank="species" species="atlantica">
<emphasis id="47B32A5CFFD6587D403150B7EB41E255" italics="true" pageId="10" pageNumber="53">Entonomenia atlantica</emphasis>
<bibRefCitation id="11568BBFFFD6587D454C509FEBA8E256" author="Leloup, E." box="[318,477,1788,1814]" pageId="10" pageNumber="53" pagination="1 - 11" refId="ref3938" refString="Leloup, E., 1948. Un nouveau solenogastre proneomeniide, Entonomenia atlantica g. nov. sp. nov. Bulletin du Museum royal d'Histoirs Naturelle Belgique, 24 (37), 1 - 11." type="journal article" year="1948">Leloup, 1948</bibRefCitation>
</taxonomicName>
(off 
<collectingRegion id="B70338ACFFD6587D466F509FE8A4E256" box="[541,721,1788,1814]" country="Spain" name="Canarias" pageId="10" pageNumber="53">Canary Islands</collectingRegion>
, 
<quantity id="B23F5BABFFD6587D4692509FE944E256" box="[736,817,1788,1814]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="10" pageNumber="53" unit="m" value="540.0">540 m</quantity>
; 
<typeStatus id="AA7C48ECFFD6587D4732509FE9D3E256" box="[832,934,1788,1814]" pageId="10" pageNumber="53" type="holotype">holotype</typeStatus>
as section series in MOM). Six other species originally classified under 
<taxonomicName id="B2C78DCDFFD6587D468E5147E9D3E27D" authorityName="Simroth" authorityYear="1893" box="[764,934,1828,1853]" class="Solenogastres" family="Rhopalomeniidae" genus="Rhopalomenia" kingdom="Animalia" order="Cavibelonia" pageId="10" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="47B32A5CFFD6587D468E5147E9D3E27D" box="[764,934,1828,1853]" italics="true" pageId="10" pageNumber="53">Rhopalomenia</emphasis>
</taxonomicName>
are herewith transferred to the restored genus, five of which were originally described by 
<bibRefCitation id="11568BBFFFD6587D47B5512FEEC9E226" author="Salvini-Plawen, L." box="[967,1212,1868,1894]" pageId="10" pageNumber="53" pagination="1 - 315" refId="ref4141" refString="Salvini-Plawen, L. v. (1978) Antarktische und subantarktische Solenogastres (Eine Monographie 1898 - 1974). Zoologica (Stuttgart), 128, 1 - 315." type="journal article" year="1978">Salvini­Plawen 1978</bibRefCitation>
(
<emphasis id="47B32A5CFFD6587C40A2512FEB4BE475" italics="true" lastPageId="11" lastPageNumber="54" pageId="10" pageNumber="53">E. carinata</emphasis>
, 
<taxonomicName id="B2C78DCDFFD7587C453E577FEB92E475" baseAuthorityName="Salvini-Plawen" baseAuthorityYear="1978" box="[332,487,284,309]" class="Solenogastres" family="Rhopalomeniidae" genus="Entonomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="tricarinata">E tricarinata</taxonomicName>
, 
<taxonomicName id="B2C78DCDFFD7587C4586577FE8DAE475" box="[500,687,284,309]" class="Solenogastres" family="Rhopalomeniidae" genus="Entonomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="sertularicola">E. sertularicola</taxonomicName>
, 
<taxonomicName id="B2C78DCDFFD7587C46CE577FE94FE475" baseAuthorityName="Salvini-Plawen" baseAuthorityYear="1978" box="[700,826,284,309]" class="Solenogastres" family="Rhopalomeniidae" genus="Entonomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="cristata">E. cristata</taxonomicName>
, 
<taxonomicName id="B2C78DCDFFD7587C4734577FEE2FE475" baseAuthorityName="Salvini-Plawen" baseAuthorityYear="1978" box="[838,1114,284,309]" class="Solenogastres" family="Rhopalomeniidae" genus="Entonomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="rhynchopharyngeata">E. rhynchopharyngeata</taxonomicName>
) and 
<taxonomicName id="B2C78DCDFFD7587C40ED577FE8C2E41E" authority="(Handl &amp; Salvini-Plawen 2002)" authorityName="Handl &amp; Salvini-Plawen" authorityYear="2002" baseAuthorityName="Handl &amp; Salvini-Plawen" baseAuthorityYear="2002" class="Solenogastres" family="Rhopalomeniidae" genus="Entonomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="microporata">
<emphasis id="47B32A5CFFD7587C40ED577FEB4DE41D" italics="true" pageId="11" pageNumber="54">E. microporata</emphasis>
(
<bibRefCitation id="11568BBFFFD7587C453A5727E8DAE41E" author="Handl, Cl. &amp; Salvini-Plawen, L." box="[328,687,324,350]" pageId="11" pageNumber="54" pagination="423 - 450" refId="ref3794" refString="Handl, Cl. &amp; Salvini-Plawen, L. v., 2002. New records of Solenogastres-Cavibelonia (Mollusca) from Norwegian fjords and shelf waters including three new species. Sarsia, 87, 423 - 450." type="journal article" year="2002">Handl &amp; Salvini­Plawen 2002</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="7578F64EFFD7587C454A570FEE52E766" blockId="11.[264,1323,284,1030]" pageId="11" pageNumber="54">
<taxonomicName id="B2C78DCDFFD7587C454A570FE9F2E4C6" authority="Garcia-Alvarez &amp; Salvini-Plawen, 2001" authorityName="Garcia-Alvarez &amp; Salvini-Plawen" authorityYear="2001" box="[312,903,364,390]" class="Solenogastres" family="Rhopalomeniidae" genus="Urgorria" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="47B32A5CFFD7587C454A570FEBD4E4C5" box="[312,417,364,389]" italics="true" pageId="11" pageNumber="54">Urgorria</emphasis>
<bibRefCitation id="11568BBFFFD7587C45DE570FE9F2E4C6" author="Garcia-Alvarez, O. &amp; Salvini-Plawen, L." box="[428,903,364,390]" pageId="11" pageNumber="54" pagination="183 - 189" refId="ref3690" refString="Garcia-Alvarez, O. &amp; Salvini-Plawen, L. v., 2001. Urgorria compostelana gen. et sp. nov. (Mollusca, Solenogastres, Rhopalomeniidae), a new species from off Galicia, Northwest of Spain. Sarsia, 86, 183 - 189." type="journal article" year="2001">García­Álvarez &amp; Salvini­Plawen, 2001</bibRefCitation>
</taxonomicName>
= 
<taxonomicName id="B2C78DCDFFD7587C47DF570FEE0CE4C6" authorityName="Salvini-Plawen" authorityYear="1978" box="[941,1145,364,390]" class="Solenogastres" family="Rhopalomeniidae" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="family">Rhopalomeniidae</taxonomicName>
with only one pair of foregut glandular organs, characterised by epithelial/intraepithelial glandular cells (so­called 
<typeStatus id="AA7C48ECFFD7587C45F057DFEBC1E496" box="[386,436,444,470]" pageId="11" pageNumber="54">type</typeStatus>
C), and with a common atrio­buccal cavity; other characters as in 
<taxonomicName id="B2C78DCDFFD7587C40CD57DFEB3BE4BD" authorityName="Simroth" authorityYear="1893" class="Solenogastres" family="Rhopalomeniidae" genus="Rhopalomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="47B32A5CFFD7587C40CD57DFEB3BE4BD" italics="true" pageId="11" pageNumber="54">Rhopalomenia</emphasis>
</taxonomicName>
above. 
<typeStatus id="AA7C48ECFFD7587C45CB5787EB87E4BE" box="[441,498,484,510]" pageId="11" pageNumber="54">Type</typeStatus>
species 
<taxonomicName id="B2C78DCDFFD7587C46165787EB30E766" authority="Garcia-Alvarez &amp; Salvini-Plawen, 2001" authorityName="Garcia-Alvarez &amp; Salvini-Plawen" authorityYear="2001" class="Solenogastres" family="Rhopalomeniidae" genus="Urgorria" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="compostelana">
<emphasis id="47B32A5CFFD7587C46165787E908E4BD" box="[612,893,484,509]" italics="true" pageId="11" pageNumber="54">Urgorria compostelana</emphasis>
<bibRefCitation id="11568BBFFFD7587C47FE5787EB30E766" author="Garcia-Alvarez, O. &amp; Salvini-Plawen, L." pageId="11" pageNumber="54" pagination="183 - 189" refId="ref3690" refString="Garcia-Alvarez, O. &amp; Salvini-Plawen, L. v., 2001. Urgorria compostelana gen. et sp. nov. (Mollusca, Solenogastres, Rhopalomeniidae), a new species from off Galicia, Northwest of Spain. Sarsia, 86, 183 - 189." type="journal article" year="2001">García­Álvarez &amp; Salvini­Plawen, 2001</bibRefCitation>
</taxonomicName>
. The genus also includes 
<taxonomicName id="B2C78DCDFFD7587C461E546FEE56E766" authority="Salvini-Plawen, 2003" authorityName="Salvini-Plawen" authorityYear="2003" box="[620,1059,524,550]" class="Solenogastres" family="Rhopalomeniidae" genus="Urgorria" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="monoplicata">
<emphasis id="47B32A5CFFD7587C461E546FE955E765" box="[620,800,524,549]" italics="true" pageId="11" pageNumber="54">U. monoplicata</emphasis>
Salvini­Plawen, 2003
</taxonomicName>
.
</paragraph>
<paragraph id="7578F64EFFD7587C454A5457EF73E786" blockId="11.[264,1323,284,1030]" pageId="11" pageNumber="54">
<taxonomicName id="B2C78DCDFFD7587C454A5457E8F2E70E" authority="Nierstrasz, 1902" authorityName="Nierstrasz" authorityYear="1902" box="[312,647,564,590]" class="Solenogastres" family="Rhopalomeniidae" genus="Dinomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="47B32A5CFFD7587C454A5457EBCFE70D" box="[312,442,564,589]" italics="true" pageId="11" pageNumber="54">Dinomenia</emphasis>
Nierstrasz, 1902
</taxonomicName>
= 
<taxonomicName id="B2C78DCDFFD7587C46C45457E9F1E70E" authorityName="Salvini-Plawen" authorityYear="1978" box="[694,900,564,590]" class="Solenogastres" family="Rhopalomeniidae" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="family">Rhopalomeniidae</taxonomicName>
with one pair of foregut glandular organs with subepithelial/extraepithelial glandular cells draining through a distinct duct (
<typeStatus id="AA7C48ECFFD7587C456254E7EB31E7DE" box="[272,324,644,670]" pageId="11" pageNumber="54">type</typeStatus>
A), with a distichous radula and with copulatory stylets; terminal sense organ present. To date only the 
<typeStatus id="AA7C48ECFFD7587C465D54CFE814E786" box="[559,609,684,710]" pageId="11" pageNumber="54">type</typeStatus>
species 
<taxonomicName id="B2C78DCDFFD7587C46B754CFEEFDE786" authority="Nierstrasz, 1902" authorityName="Nierstrasz" authorityYear="1902" box="[709,1160,684,710]" class="Solenogastres" family="Rhopalomeniidae" genus="Dinomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="hubrechti">
<emphasis id="47B32A5CFFD7587C46B754CFE9CBE785" box="[709,958,684,709]" italics="true" pageId="11" pageNumber="54">Dinomenia hubrechti</emphasis>
Nierstrasz, 1902
</taxonomicName>
, is known.
</paragraph>
</subSubSection>
<subSubSection id="3DDDA5C5FFD7587C454A54B7EF73E626" pageId="11" pageNumber="54" type="description">
<paragraph id="7578F64EFFD7587C454A54B7EF73E626" blockId="11.[264,1323,284,1030]" pageId="11" pageNumber="54">
<taxonomicName id="B2C78DCDFFD7587C454A54B7E838E7AE" authority="Heath, 1911" authorityName="Heath. Only" authorityYear="1911" box="[312,589,724,750]" class="Solenogastres" family="Rhopalomeniidae" genus="Driomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="47B32A5CFFD7587C454A54B7EBC2E7AD" box="[312,439,724,749]" italics="true" pageId="11" pageNumber="54">Driomenia</emphasis>
Heath, 1911
</taxonomicName>
= 
<taxonomicName id="B2C78DCDFFD7587C461F54B7E94EE7AE" authorityName="Salvini-Plawen" authorityYear="1978" box="[621,827,724,750]" class="Solenogastres" family="Rhopalomeniidae" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="family">Rhopalomeniidae</taxonomicName>
with one pair of foregut glandular organs with duct and subepithelial/extraepithelial glandular cells (
<typeStatus id="AA7C48ECFFD7587C47C4549FE99FE656" box="[950,1002,764,790]" pageId="11" pageNumber="54">type</typeStatus>
A), without terminal sense organ, pericardium with a pair of ventro­anterior diverticles; other characters as in 
<taxonomicName id="B2C78DCDFFD7587C457A552FEBD1E625" authorityName="Leloup." authorityYear="1948" box="[264,420,844,869]" class="Solenogastres" family="Rhopalomeniidae" genus="Entonomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="47B32A5CFFD7587C457A552FEBD1E625" box="[264,420,844,869]" italics="true" pageId="11" pageNumber="54">Entonomenia</emphasis>
</taxonomicName>
above. Only the 
<typeStatus id="AA7C48ECFFD7587C4607552FE8D2E626" box="[629,679,844,870]" pageId="11" pageNumber="54">type</typeStatus>
species 
<taxonomicName id="B2C78DCDFFD7587C4778552FEEF2E626" authority="Heath, 1911" authorityName="Heath" authorityYear="1911" box="[778,1159,844,870]" class="Solenogastres" family="Rhopalomeniidae" genus="Driomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="pacifica">
<emphasis id="47B32A5CFFD7587C4778552FE99BE625" box="[778,1006,844,869]" italics="true" pageId="11" pageNumber="54">Driomenia pacifica</emphasis>
Heath, 1911
</taxonomicName>
, is known.
</paragraph>
</subSubSection>
<subSubSection id="3DDDA5C5FFD7587C454A5517EB8FE13E" pageId="11" pageNumber="54" type="reference_group">
<paragraph id="7578F64EFFD7587C454A5517E9E5E146" blockId="11.[264,1323,284,1030]" pageId="11" pageNumber="54">
<taxonomicName id="B2C78DCDFFD7587C454A5517E837E6CE" ID-CoL="7PDN8" authority="Thiele, 1894" authorityName="Thiele" authorityYear="1894" box="[312,578,884,910]" class="Solenogastres" family="Rhopalomeniidae" genus="Pruvotia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="47B32A5CFFD7587C454A5517EBEBE6CD" box="[312,414,884,909]" italics="true" pageId="11" pageNumber="54">Pruvotia</emphasis>
<bibRefCitation id="11568BBFFFD7587C45D85517E837E6CE" author="Thiele, L." box="[426,578,884,910]" pageId="11" pageNumber="54" pagination="222 - 302" refId="ref4415" refString="Thiele, L. (1894) Beitrage zur vergleichenden Anatomie der Amphineuren. I. Uber einige Neapeler Solenogastres. Zeitschrift fur wissenschaftliche Zoologie, 58, 222 - 302." type="journal article" year="1894">Thiele, 1894</bibRefCitation>
</taxonomicName>
= 
<taxonomicName id="B2C78DCDFFD7587C461A5517E943E6CE" ID-CoL="7NKVS" authorityName="Salvini-Plawen" authorityYear="1978" box="[616,822,884,910]" class="Solenogastres" family="Rhopalomeniidae" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="family">Rhopalomeniidae</taxonomicName>
of somewhat uncertain status due to the presence of two (respiratory?) folds in the pallial cavity and a lack of information on foregut glandular organs; other characters as in 
<taxonomicName id="B2C78DCDFFD7587C474A55A7EE3BE69E" ID-CoL="7NXL9" authority="Heath. Only" authorityName="Heath. Only" authorityYear="1911" box="[824,1102,964,990]" class="Solenogastres" family="Rhopalomeniidae" genus="Driomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="47B32A5CFFD7587C474A55A7E9C2E69D" box="[824,951,964,989]" italics="true" pageId="11" pageNumber="54">Driomenia</emphasis>
Heath. Only
</taxonomicName>
the 
<typeStatus id="AA7C48ECFFD7587C40F355A7EEC6E69E" box="[1153,1203,964,990]" pageId="11" pageNumber="54">type</typeStatus>
species is known, described as 
<taxonomicName id="B2C78DCDFFD7587C458C558FE9F9E146" authority="Pruvot, 1891" authorityName="Pruvot" authorityYear="1891" box="[510,908,1004,1030]" class="Solenogastres" family="Proneomeniidae" genus="Proneomenia" kingdom="Animalia" order="Cavibelonia" pageId="11" pageNumber="54" phylum="Mollusca" rank="species" species="sopita">
<emphasis id="47B32A5CFFD7587C458C558FE89CE145" box="[510,745,1004,1029]" italics="true" pageId="11" pageNumber="54">Proneomenia sopita</emphasis>
<bibRefCitation id="11568BBFFFD7587C4683558FE9F9E146" author="Pruvot, G." box="[753,908,1004,1030]" pageId="11" pageNumber="54" pagination="699 - 810" refId="ref4008" refString="Pruvot, G., 1891. Sur l'organisation de quelques neomeniens des cotes de France. Archives de Zoologie experimentale et generale, serie 2, 9, 699 - 810." type="journal article" year="1891">Pruvot, 1891</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="7578F64EFFD7587C457A5207EB8FE13E" blockId="11.[264,506,1124,1150]" box="[264,506,1124,1150]" pageId="11" pageNumber="54">
<emphasis id="47B32A5CFFD7587C457A5207EB8FE13E" bold="true" box="[264,506,1124,1150]" pageId="11" pageNumber="54">Acknowledgements</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="3DDDA5C5FFD7587B457A52CCEE5BE1B3" lastPageId="12" lastPageNumber="55" pageId="11" pageNumber="54" type="description">
<paragraph id="7578F64EFFD7587C457A52CCEBE9E1B1" blockId="11.[264,1323,1199,1265]" pageId="11" pageNumber="54">
The authors are very grateful to Dr. Michael Stachowitsch (
<collectingRegion id="B70338ACFFD7587C478052CCEE3CE189" box="[1010,1097,1199,1225]" country="Austria" name="Wien" pageId="11" pageNumber="54">Vienna</collectingRegion>
) for polishing the English text.
</paragraph>
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