185 lines
13 KiB
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185 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.496.9169" ID-GBIF-Dataset="7bfcdf18-2d67-4da7-a171-bb9408e77bbf" ID-PMC="PMC4410158" ID-Pensoft-Pub="1313-2970-496-85" ID-PubMed="25931956" ID-ZBK="DE6EE3CABEFB4D20A41151743759BCB8" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-496-85" ModsDocOrigin="ZooKeys 496" ModsDocTitle="Description of a new species of Wormaldia from Sardinia and a new Drusus species from the Western Balkans (Trichoptera, Philopotamidae, Limnephilidae)" checkinTime="1451244436679" checkinUser="pensoft" docAuthor="Vitecek, Simon, Previsic, Ana, Kucinic, Mladen, Balint, Miklos, Keresztes, Lujza, Waringer, Johann, Pauls, Steffen U., Malicky, Hans & Graf, Wolfram" docDate="2015" docId="D415F900BB1E4CEEEEB212A612145C90" docLanguage="en" docName="ZooKeys 496: 85-103" docOrigin="ZooKeys 496" docSource="http://dx.doi.org/10.3897/zookeys.496.9169" docTitle="Wormaldia sarda Graf & Malicky, sp. n." docType="treatment" docUuid="F02C5CF5-9043-463F-809B-FCD5D2B8FBD2" docUuidSource="ZooBank" docVersion="4" lastPageNumber="87" masterDocId="0708A031FFEA6B79FE22B61FFF8C775A" masterDocTitle="Description of a new species of Wormaldia from Sardinia and a new Drusus species from the Western Balkans (Trichoptera, Philopotamidae, Limnephilidae)" masterLastPageNumber="103" masterPageNumber="85" pageNumber="86" updateTime="1668160203181" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of a new species of Wormaldia from Sardinia and a new Drusus species from the Western Balkans (Trichoptera, Philopotamidae, Limnephilidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vitecek, Simon</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Previsic, Ana</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kucinic, Mladen</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balint, Miklos</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Keresztes, Lujza</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Waringer, Johann</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pauls, Steffen U.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Malicky, Hans</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Graf, Wolfram</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>496</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>85</mods:start>
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<mods:end>103</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.496.9169</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.496.9169</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-496-85</mods:identifier>
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<mods:identifier type="ZBK">DE6EE3CABEFB4D20A41151743759BCB8</mods:identifier>
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<mods:identifier type="ZooBank">DE6EE3CABEFB4D20A41151743759BCB8</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152059634" LSID="urn:lsid:zoobank.org:act:F02C5CF5-9043-463F-809B-FCD5D2B8FBD2" httpUri="http://treatment.plazi.org/id/D415F900BB1E4CEEEEB212A612145C90" lastPageId="2" lastPageNumber="87" pageId="1" pageNumber="86">
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<subSubSection pageId="1" pageNumber="86" type="multiple">
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<paragraph pageId="1" pageNumber="86">Taxon classification Animalia Trichoptera Philopotamidae</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="86" type="nomenclature">
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<paragraph pageId="1" pageNumber="86">
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<taxonomicName LSID="http://zoobank.org/F02C5CF5-9043-463F-809B-FCD5D2B8FBD2" authority="Graf & Malicky" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sarda" order="Trichoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="sarda">Wormaldia sarda Graf & Malicky</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="86">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="86" type="material examined">
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<paragraph pageId="1" pageNumber="86">Material examined.</paragraph>
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<paragraph pageId="1" pageNumber="86">
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Holotype. 1 male pupa, holotype: Sardinia, Gola di Gorruppo;
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="40.185368">40°11.122'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="9.501734">9°30.104'E</geoCoordinate>
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; 350 m a.s.l.; 28.03.2001; leg. Monika Hess, Ulrich Heckes; currently in coll. W. Graf, will deposited in the Biologiezentrum des
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<normalizedToken originalValue="Oberösterreichischen">Oberoesterreichischen</normalizedToken>
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Landesmuseums, Linz, Austria.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="86" type="type locality">
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<paragraph pageId="1" pageNumber="86">Type locality.</paragraph>
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<paragraph pageId="1" pageNumber="86">Italy, Sardinia.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="87" pageId="1" pageNumber="86" type="diagnosis">
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<paragraph pageId="1" pageNumber="86">Diagnosis.</paragraph>
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<paragraph lastPageId="2" lastPageNumber="87" pageId="1" pageNumber="86">
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Morphology of the male terminalia suggests placement of the new species in
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="genus">
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<pageBreakToken pageId="2" pageNumber="87" start="start">Wormaldia</pageBreakToken>
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</taxonomicName>
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. The species is unique in the European
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<taxonomicName lsidName="" order="Trichoptera" pageId="2" pageNumber="87" rank="order">Trichoptera</taxonomicName>
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fauna, and easily differentiated from all other
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="genus">Wormaldia</taxonomicName>
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species by the combination of the following characters: (1) presence of median subtriangular protrusion in the distal half of the harpago, (2) membraneous dorsoproximal portion and trilobate lateral portions of segment X, and (3) distinct sclerotized structures visible on the invaginated phallus.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="87" type="description">
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<paragraph pageId="2" pageNumber="87">Description.</paragraph>
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<paragraph pageId="2" pageNumber="87">
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Adults (in pupa). Habitus dark, sclerites and tergites brown; cephalic and thoracic setal areas pale; cephalic, thoracic and abdominal setation dark brown; legs light brown, proximally darker; haustellum and intersegmental integument pale cream. Wings brown mottled with golden patches. Male maxillary palp 5-segmented. Spurformula 2
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<normalizedToken originalValue="–4–">-4-</normalizedToken>
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4 in males.
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</paragraph>
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<paragraph pageId="2" pageNumber="87">
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Male genitalia (Fig. 1
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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). Segment IX in lateral view subrectangular, bulging anteriad; dorsal quarter reduced to a narrow transverse bridge, ventral 3/4ers broad (Fig. 1A). Segment X in lateral view trilobate: unpaired dorsal lobe strongly convex with a bicuspid apex, dorsoproximally membraneous; 1 lateromedian lobe, subovate, pointed on either side; 1 ventral lobe, posteriad, pointed on either side (Fig. 1A, B). Superior appendages suboval, curved dorsad in lateral view, flat with a rounded apex in dorsal and ventral view (Fig. 1A, C, D). Invaginated phallus terminally with a dorsal pair of sclerotized, laterad divergent tines and a ventral sclerotized plate; internally with 4 distinct tines (Fig. 1A, D). Coxopodite subovate in lateral view, ventrally with a sharp mediolaterad ridge (Fig. 1A, C). Harpago subovate in lateral view, in ventral view distally with a median subtriangular serrated protrusion flattened dorsoventrally (Fig. 1A, D).
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</paragraph>
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<caption pageId="2" pageNumber="87">
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<paragraph pageId="2" pageNumber="87">
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Figure 1. Male genitalia of
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sarda" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="sarda">Wormaldia sarda</taxonomicName>
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sp. n. A right lateral view, intact B right lateral view, superior appendage removed C ventral view D dorsal view.
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</paragraph>
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</caption>
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<paragraph pageId="2" pageNumber="87">
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Mature pupa (Fig. 2
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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). Mandibles tubular, dilated at the apex (Fig. 2E,F). Abdominal dorsal sclerites as in Fig. 2D.
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</paragraph>
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<caption pageId="2" pageNumber="87">
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<paragraph pageId="2" pageNumber="87">
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Figure 2. Pupal characteristics of
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sarda" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="sarda">Wormaldia sarda</taxonomicName>
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sp. n.,
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Philopotamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopotamus montanus" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="montanus">Philopotamus montanus</taxonomicName>
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, and
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="genus">Wormaldia</taxonomicName>
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spp. A generalized pupal abdomen in dorsal view, depicting the position of the dorsal sclerites B dorsal sclerites of
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Philopotamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopotamus montanus" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="montanus">Philopotamus montanus</taxonomicName>
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C dorsal sclerites of
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia occipitalis" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="occipitalis">Wormaldia occipitalis</taxonomicName>
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D dorsal sclerites of
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sarda" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="sarda">Wormaldia sarda</taxonomicName>
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sp. n. E head of
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sarda" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="sarda">Wormaldia sarda</taxonomicName>
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pupa in left lateral view F head of
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sarda" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="sarda">Wormaldia sarda</taxonomicName>
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pupa in ventral view G pupal mandibles of
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Philopotamus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopotamus montanus" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="montanus">Philopotamus montanus</taxonomicName>
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in ventral view H pupal mandibles of
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<taxonomicName class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia copiosa" order="Trichoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="copiosa">Wormaldia copiosa</taxonomicName>
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in ventral view. Scale bars: 100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(B); 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(C, D).
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</paragraph>
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</caption>
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<paragraph pageId="2" pageNumber="87">Female and larva unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="87" type="etymology">
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<paragraph pageId="2" pageNumber="87">Etymology.</paragraph>
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<paragraph pageId="2" pageNumber="87">The species epithet refers to the island of Sardinia, the type locality.</paragraph>
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</subSubSection>
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</treatment>
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</document> |