treatments-xml/data/C8/74/B2/C874B2ED48A4AD9AE678AAE68BD4C2A0.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.806.27320" ID-GBIF-Dataset="eef5c293-43ff-4179-8eaa-e5285c9a6250" ID-PMC="PMC6302077" ID-Pensoft-Pub="1313-2970-806-17" ID-PubMed="30588155" ID-ZBK="A312BA3989FF4D43B6E206F880B956E6" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-806-17" ModsDocOrigin="ZooKeys 806" ModsDocTitle="A survey of basal insects (Microcoryphia and Zygentoma) from subterranean environments of Iran, with description of three new species" checkinTime="1545315501544" checkinUser="pensoft" docAuthor="Molero, Rafael, Tahami, Mohadeseh Sadat, Gaju, Miquel &amp; Sadeghi, Saber" docDate="2018" docId="C874B2ED48A4AD9AE678AAE68BD4C2A0" docLanguage="en" docName="ZooKeys 806: 17-46" docOrigin="ZooKeys 806" docSource="http://dx.doi.org/10.3897/zookeys.806.27320" docTitle="Haslundiella iranica Gaju, Molero, Tahami &amp; Sadeghi, sp. n." docType="treatment" docUuid="DF55D262-64C2-47CE-8ED9-0A8834DA25DD" docUuidSource="ZooBank" docVersion="5" lastPageNumber="17" masterDocId="7643FFD3DC7D6740A920CB1350370210" masterDocTitle="A survey of basal insects (Microcoryphia and Zygentoma) from subterranean environments of Iran, with description of three new species" masterLastPageNumber="46" masterPageNumber="17" pageNumber="17" updateTime="1668166586086" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A survey of basal insects (Microcoryphia and Zygentoma) from subterranean environments of Iran, with description of three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Molero, Rafael</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tahami, Mohadeseh Sadat</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gaju, Miquel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sadeghi, Saber</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>806</mods:number>
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<mods:start>17</mods:start>
<mods:end>46</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.806.27320</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.806.27320</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-806-17</mods:identifier>
<mods:identifier type="ZBK">A312BA3989FF4D43B6E206F880B956E6</mods:identifier>
<mods:identifier type="ZooBank">A312BA3989FF4D43B6E206F880B956E6</mods:identifier>
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<treatment ID-GBIF-Taxon="154125734" LSID="urn:lsid:zoobank.org:act:DF55D262-64C2-47CE-8ED9-0A8834DA25DD" httpUri="http://treatment.plazi.org/id/C874B2ED48A4AD9AE678AAE68BD4C2A0" lastPageNumber="17" pageId="0" pageNumber="17">
<subSubSection pageId="0" pageNumber="17" type="nomenclature">
<paragraph pageId="0" pageNumber="17">
<taxonomicName LSID="http://zoobank.org/DF55D262-64C2-47CE-8ED9-0A8834DA25DD" authority="Gaju, Molero, Tahami &amp; Sadeghi" class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Microcoryphia" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica Gaju, Molero, Tahami &amp; Sadeghi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="17">sp. n.</taxonomicNameLabel>
Figs 2, 3, 4, 5, 6; Tables 1, 2
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="type material">
<paragraph pageId="0" pageNumber="17">Type material.</paragraph>
<paragraph pageId="0" pageNumber="17">
Holotype (MNCN): male, body length = 8.5 mm, preserved in alcohol and partially mounted on slide, both vial and slide labelled &quot;Mirza Cave, Rafsanjan, Kerman Province, Iran. 20.VI.2015. Cat. Types N. 2834&quot;; &quot;HOLOTYPE ♂ 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n., des. Gaju, Molero, Tahami &amp; Sadeghi, 2018&quot;. Paratypes (3 ex.): 1 male (UCO): same data as holotype, preserved in alcohol and partially mounted on slide, Ref. M1655a. 1 female juvenile, preserved in alcohol (Fig. 2): same locality and date, Ref. M1655b. 1 female (ZM-CBSU): from Khane Khoda cave, Heart, Yazd Province, Iran. 30.IX.2015, preserved in alcohol and partially mounted on slide (#C2636).
</paragraph>
<caption pageId="0" pageNumber="17">
<paragraph pageId="0" pageNumber="17">
Figure 2. 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n., habitus (female specimen from Mirza cave).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="diagnosis">
<paragraph pageId="0" pageNumber="17">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName family="Machilidae" lsidName="" pageId="0" pageNumber="17" rank="family">Machilidae</taxonomicName>
of medium size, about 8-9 mm. Body unpigmented. Compound eyes wider than long (l/w = 0.65), lateral ocelli in sublateral position. Antennae shorter than body length, distal chains with a lower number of annuli compared with the remaining species of the genus. Coxal styli on middle and hind legs. Urosternites with 1+1 eversible vesicles; sternites with their posterior angle slightly obtuse, about 95°. Male without special chaetotaxy on maxillary palps and legs. Proximal part of the penis less than 1.3 times longer than the distal part. Female ovipositor with 1+58 divisions.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="description">
<paragraph pageId="0" pageNumber="17">Description.</paragraph>
<paragraph pageId="0" pageNumber="17">Habitus of the new species. All specimens, although coming from two different locations (about 300 km apart from each other), are very similar because their whitish, unpigmented bodies (Fig. 2). Body length of male 8.5 mm, female 9 mm. Body and appendages covered with scales and completely devoid of pigment. Paracercus and cerci broken. Dorsal scales pattern unknown. Meso and metathorax as usual in the order, not especially humped. Antennae shorter than the body, the specimen with longest antennae, a subadult female 7.5 mm long, with antennae of 3.9 mm (Fig. 2). Compound eyes black (Fig. 3, from adult female, those of males not clearly visible), wider than long (l/w: 0.65 and cl/l: 0.53). Frons not especially protruded between paired ocelli, which are brownish, sublateral, transversally ovoid: w/l: 2.29; not especially small (w.ocellus/w.eye: 0.55).</paragraph>
<caption pageId="0" pageNumber="17">
<paragraph pageId="0" pageNumber="17">
Figure 3. 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n., female from Khane Khoda cave. A head frontal view B head, lateral view C frontal view of compound eyes D frontal view of right eye and lateral ocellus. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="17">Description of males. Head: Antennae broken in the holotype (preserved length: 3.6 mm), scapus (Fig. 4A) not especially long (l/w: 1.52); distal chains with 10, 11 annuli (Fig. 4B); each annulus with one circle of bristles, slightly longer than the diameter of the annuli (Fig. 4C), some of them straight and strong others thinner and curled; the penultimate annuli of each chain with two specialized basiconic sensilla. Male maxillary palp not modified and without special chaetotaxy (Fig. 4D, E), all articles of similar length, only the third article slightly shorter (l 3rd/l 7th = 0.88). Hyaline spines in articles 5, 6 and 7 typical, low in number (2, 6 and 7 respectively); article ratios shown in Table 1. Labial palp typical, third article not specially enlarged (Fig. 4F); sensorial cones typical, not very numerous (Fig. 4G), surrounded by strong bristles slightly longer than these cones (in the female this character is clearer).</paragraph>
<paragraph pageId="0" pageNumber="17">Thorax: Second and third pair of legs with a stylus on the coxa, their length about 2/3 of the coxa (0.63). Fore femur not especially enlarged (Fig. 4H); hind tibiae longer than fore and mid tibiae (Figs 4H-J); all legs without special chaetotaxy, only with spines inserted ventrally on femora, tibiae and tarsi (Fig. 4K and Table 2); these spines have their distal part dark.</paragraph>
<caption pageId="0" pageNumber="17">
<paragraph pageId="0" pageNumber="17">
Figure 4. 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n., male holotype (Mirza cave). A antennal scapus, pedicellus and basal annuli of flagellum B antennal distal chains C chaetotaxy of apical annuli of antennal distal chain D maxillary palp E seventh article of maxillary palp F labial palp G field of sensorial cones of third article of labial palp H fore leg I mid leg J hind leg K tarsus of hind leg. Scale bars: 0.1 mm (A, B, 
<normalizedToken originalValue="D–K">D-K</normalizedToken>
); 50 
<normalizedToken originalValue="μm">μm</normalizedToken>
(C).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="17">Abdomen: urosternites with medial sternite (Fig. 5A), slightly obtusangle (95°); with one pair of eversible vesicles on urocoxites I-VII; styli present on segments II-IX, the terminal spine of each stylus not so long; styli with strong subterminal spiniform setae (Fig. 5B); stylus/coxite and spine/stylus ratios shown in Table 1. Coxites VII-IX with hyaline spines (Fig. 5C, F and Table 1). Male genitalia without parameres VIII; penis and parameres IX completely covered by coxites IX, surpassing slightly half the coxite (Fig. 5E); parameres with 1+6-7 divisions (Fig. 5G), not surpassing the tip of penis; proximal part of penis longer than distal part, ratio pp/pd: 1.26. Penis opening subterminal (Fig. 5H).</paragraph>
<caption pageId="0" pageNumber="17">
<paragraph pageId="0" pageNumber="17">
Figure 5. 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n.: male holotype (A, B, E, G, H), male paratype from Mirza cave (C, D, F) and female paratype (
<normalizedToken originalValue="I–L">I-L</normalizedToken>
). A fifth urosternite B fifth abdominal stylus C eighth urosternite D detail of lateral spines of eighth coxite E eighth and ninth urosternites with male genitalia F detail of spines in coxite IX G penis and nine parameres H penis opening I antennal scapus and pedicellus J labial palp K third labial palp article L preserved part of maxillary palp. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption pageId="0" pageNumber="17">
<paragraph pageId="0" pageNumber="17">
Table 1. Comparison of several morphometric parameters of 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n. with the remaining species of the genus. Length: body length from frons until the end of tenth tergite; eyes cl/l: eye contact line/eye length; eye l/w: length/width of eye; paired ocelli w/l: width/length; w ocelli/w eye: width ocelli/width eye; l ant: length antennae; scapus l/w: scapus length/width; ant d.c.: number of annuli of antennal distal chains; l cerci: length cercus; l p.cercus: length paracercus; mx-p7-p6-p5-p4-p3-p2/mx-p7: length of maxillary palp articles 7 
<normalizedToken originalValue="–6–5–4–3–">-6-5-4-3-</normalizedToken>
2/length of maxillary palp 7; sp mx-p 7 and sp mx-p 6: spines on the dorsal side of maxillary palp articles 7th and 6th; l Ti1: length of fore tibia; Fe1 (l/w): length/width of fore femur; l st-cx-L2-L3: length of coxal stylus from second and third legs respectively; l st/l cx-L2-L3: length of coxal stylus/ length of coxa for second and third legs respectively; angle of st abd V: angle of urosternite V; st/cx abd V, VIII, IX: length stylus without spine/length coxite of urosternites V, VIII and IX; sp/st abd V, VIII, IX: length terminal spine/length of stylus without spine V, VIII and IX; pp/pd penis: length proximal /length distal part of penis; parameres: number of divisions of parameres; g-VIII, IX: number of divisions of gonapophysis VIII and IX. Data from 
<taxonomicName lsidName="H. steinitzi" pageId="0" pageNumber="17" rank="species" species="steinitzi">H. steinitzi</taxonomicName>
and 
<taxonomicName lsidName="H. nisensis" pageId="0" pageNumber="17" rank="species" species="nisensis">H. nisensis</taxonomicName>
obtained from text or drawings of original descriptions (those calculated from drawings are marked with bold).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="17">
<table pageId="0" pageNumber="17">
<tr pageId="0" pageNumber="17">
<th colspan="3" pageId="0" pageNumber="17" rowspan="1">Males</th>
<th colspan="3" pageId="0" pageNumber="17" rowspan="1">Females</th>
</tr>
<tr pageId="0" pageNumber="17">
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. steinitzi" pageId="0" pageNumber="17" rank="species" species="steinitzi">H. steinitzi</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. nisensis" pageId="0" pageNumber="17" rank="species" species="nisensis">H. nisensis</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. iranica" pageId="0" pageNumber="17" rank="species" species="iranica">H. iranica</taxonomicName>
n. sp.
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. steinitzi" pageId="0" pageNumber="17" rank="species" species="steinitzi">H. steinitzi</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. nisensis" pageId="0" pageNumber="17" rank="species" species="nisensis">H. nisensis</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. iranica" pageId="0" pageNumber="17" rank="species" species="iranica">H. iranica</taxonomicName>
n. sp.
</th>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.5</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.73</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">2.33</td>
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">2.25</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.5</td>
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.54</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">1.54</td>
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">1</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">1.42</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">2.23</td>
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">1.46</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.77</td>
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.88</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">1.31</td>
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.88</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">2.17</td>
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">2.26</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.21</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.11</td>
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">0.09</td>
</tr>
<tr pageId="0" pageNumber="17">
<td colspan="1" pageId="0" pageNumber="17" rowspan="1">ca 1.87</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="17">Description of females. The only adult female has most of its appendages broken and some of them are lost, specifically right antennae, left maxillary palp, fore legs and left middle leg.</paragraph>
<paragraph pageId="0" pageNumber="17">Head: As described in habitus section. Antennae broken, only the basal part of the left one is preserved; scapus similar to that of the male (Fig. 5I). Maxillary palp broken, the three basal articles preserved shown in Fig. 5L. Labial palp (Fig. 5J) as in male, but the strong setae surrounding the sensorial cones of the third article seem stronger (Fig. 5K).</paragraph>
<paragraph pageId="0" pageNumber="17">Thorax: Middle and hind legs (Fig. 6A, B) similar to that of male, but slightly bigger; without special chaetotaxy, only ventral spines on femora, tibiae and tarsi (Table 2).</paragraph>
<paragraph pageId="0" pageNumber="17">Abdomen: sternites typical of the genus, slightly obtusangle (95°), slightly bigger than those of male (Fig. 6C); styli of coxite similar to those of males (Fig. 6D). Coxite VII modified, with a terminal inner projection (Fig. 6E); one spine in its outer side. Coxites VIII and IX typical, with one spine in the outer side of the former (Fig. 6F) and two spines in the inner part of the later (Fig. 6G). Ovipositor of tertiary type, with 1 + 58 divisions, not attaining the apex of styli IX (Fig. 6G); gonapophysis VIII (Fig. 6H, I) with conspicuous chaetotaxy in the 22 distal divisions (36-58) and gonapophysis IX (Fig. 6J, K) in the 19 distal divisions (39-58), in the remaining divisions the chaetotaxy consists of very small bristles (if they are present).</paragraph>
<caption pageId="0" pageNumber="17">
<paragraph pageId="0" pageNumber="17">
Figure 6. 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n.: female paratype. A mid leg B hind leg C fifth urosternite D fifth abdominal stylus E seventh urosternite F eighth urocoxite and gonapophysis G ninth urocoxite and gonapophysis H medial part of eighth gonapophysis (divisions 36-45) I distal part of eighth gonapophysis (divisions 53-58) J medial part of ninth gonapophysis (divisions 37-46) K distal part of ninth gonapophysis (divisions 53-58). Scale bars: 0.25 mm (
<normalizedToken originalValue="A–C">A-C</normalizedToken>
, E); 0.2 mm (F, G); 0.1 mm (D, 
<normalizedToken originalValue="H–K">H-K</normalizedToken>
).
</paragraph>
</caption>
<caption ID-Table-UUID="6DAF75E8B7F398ABDC2FA8508A8C304A" httpUri="http://table.plazi.org/id/6DAF75E8B7F398ABDC2FA8508A8C304A" pageId="0" pageNumber="17">
<paragraph pageId="0" pageNumber="17">
Table 2. Comparison of the numbers of spines in different articles of legs of 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n. with the remaining species of the genus. sp Fe1, 2, 3: number of spines on fore, mid and hind femur; sp Ti1, 2, 3: number of spines on fore, mid and hind tibia; sp Ta1-1, 2, 3: number of spines on the three articles of fore tarsus (the same for mind tarsus -sp-Ta2- and hind tarsus -sp-Ta3).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="17">
<table pageId="0" pageNumber="17">
<tr pageId="0" pageNumber="17">
<th colspan="3" pageId="0" pageNumber="17" rowspan="1">Males</th>
<th colspan="3" pageId="0" pageNumber="17" rowspan="1">Females</th>
</tr>
<tr pageId="0" pageNumber="17">
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. steinitzi" pageId="0" pageNumber="17" rank="species" species="steinitzi">H. steinitzi</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. nisensis" pageId="0" pageNumber="17" rank="species" species="nisensis">H. nisensis</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. iranica" pageId="0" pageNumber="17" rank="species" species="iranica">H. iranica</taxonomicName>
n. sp.
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. steinitzi" pageId="0" pageNumber="17" rank="species" species="steinitzi">H. steinitzi</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. nisensis" pageId="0" pageNumber="17" rank="species" species="nisensis">H. nisensis</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="17" rowspan="1">
<taxonomicName lsidName="H. iranica" pageId="0" pageNumber="17" rank="species" species="iranica">H. iranica</taxonomicName>
n. sp.
</th>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="discussion">
<paragraph pageId="0" pageNumber="17">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="17">
The first species described of the genus 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="genus">Haslundiella</taxonomicName>
Janetschek, 1954 was found in Palestine and was named as 
<taxonomicName class="Insecta" family="Machilidae" genus="Praemachilis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praemachilis steinitzi" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="steinitzi">Praemachilis steinitzi</taxonomicName>
(Wygodzinsky, 1942). In their description 
<bibRefCitation author="Wygodzinsky, P" journalOrPublisher="Revista Brasileira de Biologia" pageId="0" pageNumber="17" pagination="29 - 46" title="Second contribution towards the knowledge of Diplura and Thysanura from Palestine." volume="2" year="1942">Wygodzinsky (1942)</bibRefCitation>
suggested that &quot;The remarkable formation of maxillary palp and the genitalia of the male might, in the future, lead to establish a new genus within the 
<taxonomicName lsidName="" pageId="0" pageNumber="17" rank="subfamily" subfamily="Praemachilinae">Praemachilinae</taxonomicName>
&quot;. 
<bibRefCitation author="Janetschek, H" journalOrPublisher="Eos" pageId="0" pageNumber="17" pagination="163 - 314" title="Ueber Felsenspringer der mittelmeerlaender (Thysanura, Machilidae)." volume="30" year="1954">Janetschek (1954)</bibRefCitation>
erected the genus 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="genus">Haslundiella</taxonomicName>
, including in it only 
<taxonomicName lsidName="H. steinitzi" pageId="0" pageNumber="17" rank="species" species="steinitzi">H. steinitzi</taxonomicName>
. 
<bibRefCitation author="Kaplin, VG" journalOrPublisher="Nasekomye Mongolii" pageId="0" pageNumber="17" pagination="16 - 61" title="New data on the fauna of Thysanura of Mongolia, Kasakhstan and Middle Asia. [In Russian]." volume="8" year="1982">Kaplin (1982)</bibRefCitation>
described the second species of this genus (
<taxonomicName lsidName="H. nisensis" pageId="0" pageNumber="17" rank="species" species="nisensis">H. nisensis</taxonomicName>
Kaplin, 1982) from Turkmenistan. Now, 
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n. is described from Iran, geographically placed between the two former species.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName class="Insecta" family="Machilidae" genus="Haslundiella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haslundiella iranica" order="Archaeognatha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="iranica">Haslundiella iranica</taxonomicName>
sp. n., can be distinguished from the other species by several characters, with the most remarkable being the significant absence of special chaetotaxy on the maxillary palps and the legs of males. Moreover, some other characters can be mentioned: Antennae are shorter than body length; the shape of the maxillary palp is different because all articles are similar in length, showing clear differences with 
<taxonomicName lsidName="H. steinitzi" pageId="0" pageNumber="17" rank="species" species="steinitzi">H. steinitzi</taxonomicName>
and 
<taxonomicName lsidName="H. nisensis" pageId="0" pageNumber="17" rank="species" species="nisensis">H. nisensis</taxonomicName>
(Table 1); and legs are similar in shape but with very different chaetotaxy. The male genitalia of 
<taxonomicName lsidName="H. iranica" pageId="0" pageNumber="17" rank="species" species="iranica">H. iranica</taxonomicName>
is clearly different from that of 
<taxonomicName lsidName="H. steinitzi" pageId="0" pageNumber="17" rank="species" species="steinitzi">H. steinitzi</taxonomicName>
and similar to 
<taxonomicName lsidName="H. nisensis" pageId="0" pageNumber="17" rank="species" species="nisensis">H. nisensis</taxonomicName>
, although penis ratios are different to those of both previously described species. The female ovipositor does not surpass the styli IX, as in 
<taxonomicName lsidName="H. nisensis" pageId="0" pageNumber="17" rank="species" species="nisensis">H. nisensis</taxonomicName>
, although with lower number of divisions, meanwhile that of 
<taxonomicName lsidName="H. steinitzi" pageId="0" pageNumber="17" rank="species" species="steinitzi">H. steinitzi</taxonomicName>
has a low number of divisions but surpasses the styli IX.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="distribution">
<paragraph pageId="0" pageNumber="17">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="17">Only known from the two localities in Kerman and Yazd provinces in Iran (see Fig. 1).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="etymology">
<paragraph pageId="0" pageNumber="17">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="17">
The name 
<taxonomicName genus="Hexapoda" lsidName="Hexapoda iranica" pageId="0" pageNumber="17" rank="species" species="iranica">iranica</taxonomicName>
is a genitive case of the name Iran, the country where the new species is found.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="habitat">
<paragraph pageId="0" pageNumber="17">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="17">
The specimens were found and collected close to the entrance of the cave where it was still darker and more humid than the outside (somewhere between the endogean and parahypogean zones) and 
<normalizedToken originalValue="couldn’t">couldn't</normalizedToken>
be found deeper in as the cave is a complex of horizontal (about 50 m from the entrance) and then vertical (a 50 m of a vertical pit) passages reaching a big hall at the end.
</paragraph>
</subSubSection>
</treatment>
</document>