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<mods:title id="98E953092AAEE4C360381F103AEB585E">Decapod Crustacea of the Californian and Oregonian Zoogeographic Provinces 3371</mods:title>
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<taxonomicName id="8ACCB1C7FFA3B32C44D1FA66CA3F0EE5" ID-CoL="84WXR" authority="Rathbun, 1904" authorityName="Rathbun" authorityYear="1904" box="[151,569,1455,1481]" class="Malacostraca" family="Alpheidae" genus="Betaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="harrimani">
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<emphasis id="7FB81656FFA3B32C44D1FA66C97C0EE5" bold="true" box="[151,378,1455,1481]" italics="true" pageId="63" pageNumber="64">Betaeus harrimani</emphasis>
<bibRefCitation id="295DB7B5FFA3B32C45C7FA66CA3F0EE5" author="Rathbun, M. J." box="[385,569,1455,1481]" pageId="63" pageNumber="64" pagination="1 - 219" refId="ref234490" refString="Rathbun, M. J. (1904) Decapod crustaceans of the northwest coast of North America. Harriman Alaska Expedition, 10, 1 - 219." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
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<paragraph id="4D73CA44FFA3B32C44D1FA1BC9B80EC6" blockId="63.[151,569,1455,1514]" box="[151,446,1490,1514]" pageId="63" pageNumber="64">
(
<figureCitation id="D5F7D6C1FFA3B32C44D9FA1BC9B00EC6" box="[159,438,1490,1514]" captionStart="FIGURE 15" captionStartId="64.[151,250,1853,1876]" captionTargetBox="[195,1387,183,1832]" captionTargetId="figure-139@64.[177,1401,161,1832]" captionTargetPageId="64" captionText="FIGURE 15. Family Alpheidae. A, B, G, K, O, P, Q. Betaeus harrimani Rathbun, 1904; A, frontal region in dorsal view; B, scaphocerite; G, tail fan; K, right cheliped of male; O, pereopod 3; P, left chela of female; Q, right chela of female. C, D, H, J, L, R Betaeus longidactylus Lockington, 1877; C, frontal region in dorsal view; D, scaphocerite; H, tail fan; J, left chela of male; L, left cheliped of female; R, right chela of female. E, F, I, M, N. Betaeus ensenadaensis Glassell, 1938; E, frontal region in dorsal view; F, scaphocerite; I, tail fan; M, right cheliped of mature male; N, right chela of female or immature male. Scale =1 mm. From Hart 1964." pageId="63" pageNumber="64">Figs. 15A, B, G, K, OQ</figureCitation>
)
</paragraph>
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<treatmentCitation id="CC6DEC55FFA3B32C44D1F9D2CA220D1C" author="Rathbun, M. J." box="[151,548,1562,1584]" page="108" pageId="63" pageNumber="64" year="1904">
<taxonomicName id="8ACCB1C7FFA3B32C44D1F9D2CA220D1C" ID-CoL="84WXR" authority="Rathbun, 1904: 108" authorityName="Rathbun" authorityPageNumber="108" authorityYear="1904" box="[151,548,1562,1584]" class="Malacostraca" family="Alpheidae" genus="Betaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="harrimani">
<emphasis id="7FB81656FFA3B32C44D1F9D2C9550D1C" box="[151,339,1562,1584]" italics="true" pageId="63" pageNumber="64">Betaeus harrimani</emphasis>
<bibRefCitation id="295DB7B5FFA3B32C451CF9D3CA220D1C" author="Rathbun, M. J." box="[346,548,1562,1584]" pageId="63" pageNumber="64" pagination="1 - 219" refId="ref234490" refString="Rathbun, M. J. (1904) Decapod crustaceans of the northwest coast of North America. Harriman Alaska Expedition, 10, 1 - 219." type="journal article" year="1904">Rathbun, 1904: 108</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 49. —
<treatmentCitation id="CC6DEC55FFA3B32C46DAF9D2CB300D1C" author="Hart, J. F. L." box="[668,822,1562,1584]" page="435" pageId="63" pageNumber="64" year="1964">
<bibRefCitation id="295DB7B5FFA3B32C46DAF9D2CB300D1C" author="Hart, J. F. L." box="[668,822,1562,1584]" pageId="63" pageNumber="64" pagination="431 - 466" refId="ref224439" refString="Hart, J. F. L. (1964) Shrimps of the genus Betaeus on the Pacific coast of North America with descriptions of three new species. Proceedings of the United States National Museum, 115, 431 - 466." type="journal article" year="1964">Hart 1964: 435</bibRefCitation>
</treatmentCitation>
, pl. 1, figs. 116, 2931, 3739, pl. 1. —
<treatmentCitation id="CC6DEC55FFA3B32C409AF9D3CD9F0D1C" author="Kozloff, E. N." box="[1244,1433,1562,1584]" page="165" pageId="63" pageNumber="64" year="1974">
<bibRefCitation id="295DB7B5FFA3B32C409AF9D3CD9F0D1C" author="Kozloff, E. N." box="[1244,1433,1562,1584]" pageId="63" pageNumber="64" refId="ref228405" refString="Kozloff, E. N. (1974) Keys to the Marine Invertebrates of Puget Sound, the San Juan Archipelago, and Adjacent Regions. University of Washington Press, Seattle, 226 pp." type="book" year="1974">Kozloff 1974: 165</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="CC6DEC55FFA3B32C44AEF9F0C9FB0D63" author="Word, J. &amp; Charwat, D." box="[232,509,1593,1615]" page="53" pageId="63" pageNumber="64" year="1976">
<bibRefCitation id="295DB7B5FFA3B32C44AEF9F0C9FB0D63" author="Word, J. &amp; Charwat, D." box="[232,509,1593,1615]" pageId="63" pageNumber="64" refId="ref241447" refString="Word, J. &amp; Charwat, D. (1976) Invertebrates of Southern California Coastal Waters. II. Natantia. Southern California Coastal Water Research Project, El Segundo, California. 238 pp." type="book" year="1976">Word &amp; Charwat 1976: 53</bibRefCitation>
</treatmentCitation>
. — Butler 980: 151. —
<treatmentCitation id="CC6DEC55FFA3B32C46BFF9F0CC140D63" author="Chace, F. A. Jr. &amp; Abbott, D. P." box="[761,1042,1593,1615]" page="571" pageId="63" pageNumber="64" year="1980">
<bibRefCitation id="295DB7B5FFA3B32C46BFF9F0CC140D63" author="Chace, F. A. Jr. &amp; Abbott, D. P." box="[761,1042,1593,1615]" pageId="63" pageNumber="64" pagination="567 - 576" refId="ref219414" refString="Chace, F. A. Jr. &amp; Abbott, D. P. (1980) Caridea: the shrimps. In: Morris, R. H., Abbott, D. P. &amp; Haderlie, E. C. (Eds.) Intertidal Invertebrates of California. Stanford University Press, Stanford, California, pp. 567 - 576." type="book chapter" year="1980">Chace &amp; Abbott 1980: 571</bibRefCitation>
</treatmentCitation>
, fig. 23.4 —
<treatmentCitation id="CC6DEC55FFA3B32C40DAF9F0CD7E0D63" author="Wicksten, M. K." box="[1180,1400,1593,1615]" page="188" pageId="63" pageNumber="64" year="1984">
<bibRefCitation id="295DB7B5FFA3B32C40DAF9F0CD7E0D63" author="Wicksten, M. K." box="[1180,1400,1593,1615]" pageId="63" pageNumber="64" pagination="186 - 190" refId="ref239496" refString="Wicksten, M. K. (1984 a) New records of snapping shrimps (family Alpheidae) from California. Proceedings of the Biological Society of Washington, 97, 186 - 190." type="journal article" year="1984">Wicksten 1984a: 188</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="CC6DEC55FFA3B32C4481F991C96B0D42" author="Jensen, G. C." box="[199,365,1624,1646]" page="43" pageId="63" pageNumber="64" year="1995">
<bibRefCitation id="295DB7B5FFA3B32C4481F991C96B0D42" author="Jensen, G. C." box="[199,365,1624,1646]" pageId="63" pageNumber="64" refId="ref227177" refString="Jensen, G. C. (1995) Pacific Coast Crabs and Shrimps. Sea Challengers, Monterey, California, 87 pp." type="book" year="1995">Jensen 1995: 43</bibRefCitation>
</treatmentCitation>
, fig. 67. —
<treatmentCitation id="CC6DEC55FFA3B32C45A3F991CAC20D42" author="Kuris, A. M. &amp; Sadeghian, P. &amp; Carlton, J. T." box="[485,708,1624,1646]" page="637" pageId="63" pageNumber="64" year="2007">
<bibRefCitation id="295DB7B5FFA3B32C45A3F991CAC20D42" author="Kuris, A. M. &amp; Sadeghian, P. &amp; Carlton, J. T." box="[485,708,1624,1646]" pageId="63" pageNumber="64" pagination="636 - 656" refId="ref228858" refString="Kuris, A. M., Sadeghian, P. &amp; Carlton, J. T. (2007) Keys to Decapod Crustacea. In: Carlton, J. T. (Ed.) The Light and Smith Manual: Intertidal Invertebrates Central California to Oregon. University of California Press, Berkeley, 4 th ed., pp. 636 - 656." type="book chapter" year="2007">
Kuris
<emphasis id="7FB81656FFA3B32C4662F990CA510D42" box="[548,599,1624,1646]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
2007: 637
</bibRefCitation>
</treatmentCitation>
, pl. 317 B, E, l, M.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="05D699CFFFA3B35244D1F953C9E40A30" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64" type="diagnosis">
<paragraph id="4D73CA44FFA3B32C44D1F953CA200C85" blockId="63.[151,1436,1690,2032]" pageId="63" pageNumber="64">
<emphasis id="7FB81656FFA3B32C44D1F953C9170D9F" bold="true" box="[151,273,1690,1715]" inLineHeading="true" pageId="63" pageNumber="64" reason="1">Diagnosis.</emphasis>
Front of carapace slightly curved, depressed anteriorly. Stylocerite reaching almost to end of second segment of antennular peduncle. Scaphocerite broad, lateral tooth exceeding blade, reaching past middle of last segment of antennular peduncle. Anterior margin of carapace with 2 shallow sinuses. Pereopods 1 with fingers 0.5 times as long as palm, with or without gape. Pereopods 2 slender, with 5 carpal articles; article 1 equal in length to next 3 together. Pereopods 3, 4 slender, somewhat flattened, with thin, curved dactyls. Pereopod 5 similar to 3, 4 but with bands of setae forming brush on distal half of propodus. Pleura of abdominal somites 13 rounded, pleura of 4, 5 angled. Telson with 2 pairs spines on dorsal surface, 2 spines at each posteolateral angle, posterior margin deeply curved. Total length
<quantity id="8A3467A1FFA3B32C4596F858CA240C85" box="[464,546,1937,1961]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="63" pageNumber="64" unit="mm" value="35.0">35 mm</quantity>
.
</paragraph>
<paragraph id="4D73CA44FFA3B32C4481F87DCAFE0CC3" blockId="63.[151,1436,1690,2032]" pageId="63" pageNumber="64">
<emphasis id="7FB81656FFA3B32C4481F87DC96F0CE1" bold="true" box="[199,361,1972,1997]" inLineHeading="true" pageId="63" pageNumber="64" reason="1">Color in life.</emphasis>
Color determined by distribution of blue and red chromatophores: translucent, reddish to purplish, pale green; turning blue at night (
<bibRefCitation id="295DB7B5FFA3B32C4633F81ECAEA0CC3" author="Hart, J. F. L." box="[629,748,2007,2031]" pageId="63" pageNumber="64" pagination="431 - 466" refId="ref224439" refString="Hart, J. F. L. (1964) Shrimps of the genus Betaeus on the Pacific coast of North America with descriptions of three new species. Proceedings of the United States National Museum, 115, 431 - 466." type="journal article" year="1964">Hart 1964</bibRefCitation>
).
</paragraph>
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<paragraph id="4D73CA44FFDCB35344D1F8F4C9730CC0" blockId="64.[151,1436,1853,2028]" pageId="64" pageNumber="65">
<emphasis id="7FB81656FFDCB35344D1F8F4C9270C78" bold="true" box="[151,289,1853,1876]" pageId="64" pageNumber="65">FIGURE 15.</emphasis>
Family
<taxonomicName id="8ACCB1C7FFDCB353453EF8F4C9E20C7F" authorityName="Rafinesque" authorityYear="1815" box="[376,484,1853,1875]" class="Malacostraca" family="Alpheidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
. A, B, G, K, O, P, Q.
<taxonomicName id="8ACCB1C7FFDCB3534682F8F7CC220C7F" authority="Rathbun, 1904" authorityName="Rathbun" authorityYear="1904" box="[708,1060,1853,1875]" class="Malacostraca" family="Alpheidae" genus="Betaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="harrimani">
<emphasis id="7FB81656FFDCB3534682F8F7CB840C7F" box="[708,898,1853,1875]" italics="true" pageId="64" pageNumber="65">Betaeus harrimani</emphasis>
<bibRefCitation id="295DB7B5FFDCB35347CDF8F4CC220C7F" author="Rathbun, M. J." box="[907,1060,1853,1875]" pageId="64" pageNumber="65" pagination="1 - 219" refId="ref234490" refString="Rathbun, M. J. (1904) Decapod crustaceans of the northwest coast of North America. Harriman Alaska Expedition, 10, 1 - 219." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
</taxonomicName>
; A, frontal region in dorsal view; B, scaphocerite; G, tail fan; K, right cheliped of male; O, pereopod 3; P, left chela of female; Q, right chela of female. C, D, H, J, L, R
<taxonomicName id="8ACCB1C7FFDCB353448FF8B5CA620CBC" authority="Lockington, 1877" authorityName="Lockington" authorityYear="1877" box="[201,612,1914,1937]" class="Malacostraca" family="Alpheidae" genus="Betaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="longidactylus">
<emphasis id="7FB81656FFDCB353448FF8B5C9AE0CBD" box="[201,424,1915,1937]" italics="true" pageId="64" pageNumber="65">Betaeus longidactylus</emphasis>
Lockington, 1877
</taxonomicName>
; C, frontal region in dorsal view; D, scaphocerite; H, tail fan; J, left chela of male; L, left cheliped of female; R, right chela of female. E, F, I, M, N.
<taxonomicName id="8ACCB1C7FFDCB3534702F853CCCD0C83" authority="Glassell, 1938" authorityName="Glassell" authorityYear="1938" box="[836,1227,1945,1967]" class="Malacostraca" family="Alpheidae" genus="Betaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="ensenadaensis">
<emphasis id="7FB81656FFDCB3534702F853CC280C83" box="[836,1070,1945,1967]" italics="true" pageId="64" pageNumber="65">Betaeus ensenadaensis</emphasis>
<bibRefCitation id="295DB7B5FFDCB3534073F850CCCD0C83" author="Glassell, S. A." box="[1077,1227,1945,1967]" pageId="64" pageNumber="65" pagination="411 - 454" refId="ref222874" refString="Glassell, S. A. (1938) New and obscure decapod Crustacea from the West American coast. Transactions of the San Diego Society of Natural History, 8, 411 - 454." type="journal article" year="1938">Glassell, 1938</bibRefCitation>
</taxonomicName>
; E, frontal region in dorsal view; F, scaphocerite; I, tail fan; M, right cheliped of mature male; N, right chela of female or immature male. Scale =1 mm. From
<bibRefCitation id="295DB7B5FFDCB353454EF81EC9770CC0" author="Hart, J. F. L." box="[264,369,2006,2028]" pageId="64" pageNumber="65" pagination="431 - 466" refId="ref224439" refString="Hart, J. F. L. (1964) Shrimps of the genus Betaeus on the Pacific coast of North America with descriptions of three new species. Proceedings of the United States National Museum, 115, 431 - 466." type="journal article" year="1964">Hart 1964</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="4D73CA44FFDDB3524481FF5EC9F60BFF" blockId="65.[151,1436,151,284]" pageId="65" pageNumber="66">
<emphasis id="7FB81656FFDDB3524481FF5EC9B80B9C" bold="true" box="[199,446,151,176]" inLineHeading="true" pageId="65" pageNumber="66" reason="1">Habitat and depth.</emphasis>
In pools, among oysters, in burrows of
<taxonomicName id="8ACCB1C7FFDDB3524796FF51CCC80B83" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[976,1230,152,176]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="pugettensis">
<emphasis id="7FB81656FFDDB3524796FF51CCC80B83" box="[976,1230,152,176]" italics="true" pageId="65" pageNumber="66">Upogebia pugettensis</emphasis>
</taxonomicName>
and
<taxonomicName id="8ACCB1C7FFDDB352415FFF50C9290BF8" baseAuthorityName="Dana" baseAuthorityYear="1854" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="californiensis">
<emphasis id="7FB81656FFDDB352415FFF50C9290BF8" italics="true" pageId="65" pageNumber="66">Neotrypaea californiensis</emphasis>
</taxonomicName>
, intertidal zones.
</paragraph>
<paragraph id="4D73CA44FFDDB3524481FF29C9E40A30" blockId="65.[151,1436,151,284]" pageId="65" pageNumber="66">
<emphasis id="7FB81656FFDDB3524481FF29C91C0BD4" bold="true" box="[199,282,224,248]" inLineHeading="true" pageId="65" pageNumber="66" reason="1">Range.</emphasis>
<materialsCitation id="FDA4C019FFDDB3524562FF16C9DB0A30" county="Sitka" location="eastern Pacific" municipality="Harbor" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Alaska">
<collectingCounty id="A412B2C8FFDDB3524562FF16C9660BDB" box="[292,352,223,247]" pageId="65" pageNumber="66">Sitka</collectingCounty>
,
<collectingRegion id="8F0804A6FFDDB3524529FF16C9BB0BDB" box="[367,445,223,247]" country="United States of America" name="Alaska" pageId="65" pageNumber="66">Alaska</collectingRegion>
to
<collectingRegion id="8F0804A6FFDDB35245AEFF29CA4C0BDB" box="[488,586,224,247]" country="United Kingdom" name="Newport" pageId="65" pageNumber="66">Newport</collectingRegion>
<collectingMunicipality id="AD17503EFFDDB3524612FF16CAA30BDB" box="[596,677,223,247]" pageId="65" pageNumber="66">Harbor</collectingMunicipality>
,
<collectingRegion id="8F0804A6FFDDB35246F5FF16CB2E0BDB" box="[691,808,223,247]" country="United States of America" name="California" pageId="65" pageNumber="66">California</collectingRegion>
.
<typeStatus id="927774E6FFDDB3524770FF29CB680BD4" box="[822,878,224,248]" pageId="65" pageNumber="66">Type</typeStatus>
locality Sitka,
<collectingRegion id="8F0804A6FFDDB3524064FF16CC750BDB" box="[1058,1139,223,247]" country="United States of America" name="Alaska" pageId="65" pageNumber="66">Alaska</collectingRegion>
. This is the northernmost alpheid in the
<location id="48139C9FFFDDB3524570FECDC9DB0A30" LSID="urn:lsid:plazi:treatment:C5657B52FFA3B35244D1FA66C9E40A30:48139C9FFFDDB3524570FECDC9DB0A30" box="[310,477,260,284]" county="Sitka" municipality="Harbor" name="eastern Pacific" pageId="65" pageNumber="66" stateProvince="Alaska">eastern Pacific</location>
</materialsCitation>
.
</paragraph>
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