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<document id="2339A9A10970D5871247463465A09E6C" ID-CLB-Dataset="7470" ID-DOI="10.11646/zootaxa.4948.3.4" ID-GBIF-Dataset="6bfd846c-0304-4330-8941-95c406630230" ID-ISSN="1175-5326" ID-Zenodo-Dep="4629100" ID-ZooBank="8E0BF97A-D9E1-4F2E-BB63-82A6ED47E5DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616485698777" checkinUser="plazi" docAuthor="Xu, Deliang, Webb, Michael D., Shang, Suqin &amp; Zhang, Yalin" docDate="2021" docId="597A878EFFBEFF93FF66F69F864FF2F9" docLanguage="en" docName="zootaxa.4948.3.4.pdf" docOrigin="Zootaxa 4948 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hecaloidella Osborn 1934" docType="treatment" docVersion="6" lastPageNumber="384" masterDocId="A543FFF6FFBCFF90FFF1F728870FF30F" masterDocTitle="Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species" masterLastPageNumber="403" masterPageNumber="381" pageNumber="383" updateTime="1698934487255" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="235ABACF5FADB8240F811E123FF7FA10">Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species</mods:title>
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<mods:namePart id="8C30C4944A88619258BD33F4CAF8257A">Xu, Deliang</mods:namePart>
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<mods:affiliation id="AA872077AA64C72F8FA5D079242387B2">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; 18710305867 @ 163. com; https: // orcid. org / 0000 - 0003 - 0106 - 5570</mods:affiliation>
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<mods:namePart id="96BB9D2A9E235C838C220F312F997D04">Webb, Michael D.</mods:namePart>
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<mods:affiliation id="26104EB88F334329CC252D5A764666BF">Department of Life Sciences, Natural History Museum, Cromwell Road, London, ME 15 6 QX, UK &amp; mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
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<mods:namePart id="34FDB374CB2363F6E404E917FE3E0D18">Shang, Suqin</mods:namePart>
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<mods:affiliation id="47860DC517955267FBE7C4FED0BD7AE1">Biological Engineering Laboratory of Crop Diseases and Pests of Gansu Province, College of Plant Protection, Gansu Agricultural University, Lanzhou, Gansu 730070, China &amp; shangsq @ gsau. edu. cn; https: // orcid. org / 0000 - 0001 - 9376 - 0735</mods:affiliation>
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<mods:namePart id="79B5A2BBE941483C4F306005E6FCCBF8">Zhang, Yalin</mods:namePart>
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<mods:affiliation id="44435713B18EC1BDEBB749EC24E818CF">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; yalinzh @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0002 - 1204 - 9181</mods:affiliation>
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<subSubSection id="99C96513FFBEFF92FF66F69F8538F2DE" box="[151,567,439,466]" pageId="2" pageNumber="383" type="nomenclature">
<paragraph id="D16C3698FFBEFF92FF66F69F8538F2DE" blockId="2.[151,567,439,466]" box="[151,567,439,466]" pageId="2" pageNumber="383">
<heading id="8A2481F4FFBEFF92FF66F69F8538F2DE" bold="true" box="[151,567,439,466]" fontSize="11" level="1" pageId="2" pageNumber="383" reason="1">
<emphasis id="E3A7EA8AFFBEFF92FF66F69F8538F2DE" bold="true" box="[151,567,439,466]" pageId="2" pageNumber="383">
Genus
<taxonomicName id="16D34D1BFFBEFF92FF1FF69F8538F2DE" ID-CoL="8HKBY" authority="Osborn, 1934" authorityName="Osborn" authorityYear="1934" box="[238,567,439,466]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="E3A7EA8AFFBEFF92FF1FF69F8688F2DD" bold="true" box="[238,391,439,466]" italics="true" pageId="2" pageNumber="383">Hecaloidella</emphasis>
<bibRefCitation id="B5424B69FFBEFF92FE7EF69F8538F2DE" author="Osborn, H." box="[399,567,439,465]" pageId="2" pageNumber="383" pagination="163 - 192" refId="ref12261" refString="Osborn, H. (1934) Hemiptera - Cicadellidae (Jassidae). Insects of Samoa and other Samoan terrestrial Arthropoda, II (4), 163 - 192." type="journal article" year="1934">Osborn, 1934</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="99C96513FFBEFF92FF66F6D486D1F11B" box="[151,478,508,532]" pageId="2" pageNumber="383" type="reference_group">
<paragraph id="D16C3698FFBEFF92FF66F6D486D1F11B" blockId="2.[151,782,508,563]" box="[151,478,508,532]" pageId="2" pageNumber="383">
<treatmentCitationGroup id="F1C311B6FFBEFF92FF66F6D486D1F11B" box="[151,478,508,532]" pageId="2" pageNumber="383">
<treatmentCitation id="50721089FFBEFF92FF66F6D486D1F11B" author="Osborn, H." box="[151,478,508,532]" page="173" pageId="2" pageNumber="383" year="1934">
<taxonomicName id="16D34D1BFFBEFF92FF66F6D486D1F11B" ID-CoL="8HKBY" authority="Osborn, 1934: 173" authorityName="Osborn" authorityPageNumber="173" authorityYear="1934" box="[151,478,508,532]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="genus">
<emphasis id="E3A7EA8AFFBEFF92FF66F6D48615F11B" box="[151,282,508,532]" italics="true" pageId="2" pageNumber="383">Hecaloidella</emphasis>
<bibRefCitation id="B5424B69FFBEFF92FED1F6D486D1F11B" author="Osborn, H." box="[288,478,508,532]" pageId="2" pageNumber="383" pagination="163 - 192" refId="ref12261" refString="Osborn, H. (1934) Hemiptera - Cicadellidae (Jassidae). Insects of Samoa and other Samoan terrestrial Arthropoda, II (4), 163 - 192." type="journal article" year="1934">Osborn, 1934: 173</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="99C96513FFBEFF92FF66F5338401F13D" box="[151,782,538,563]" pageId="2" pageNumber="383" type="materials_examined">
<paragraph id="D16C3698FFBEFF92FF66F5338401F13D" blockId="2.[151,782,508,563]" box="[151,782,538,563]" pageId="2" pageNumber="383">
<typeStatus id="0E68883AFFBEFF92FF66F53387C6F13C" box="[151,201,539,563]" pageId="2" pageNumber="383">Type</typeStatus>
species:
<taxonomicName id="16D34D1BFFBEFF92FED7F533851CF13D" authority="Osborn, 1934" authorityName="Osborn" authorityYear="1934" box="[294,531,538,562]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="E3A7EA8AFFBEFF92FED7F5338637F13D" box="[294,312,539,562]" italics="true" pageId="2" pageNumber="383">H</emphasis>
.
<emphasis id="E3A7EA8AFFBEFF92FEB4F532868FF13D" box="[325,384,538,562]" italics="true" pageId="2" pageNumber="383">nitida</emphasis>
<bibRefCitation id="B5424B69FFBEFF92FE76F532851CF13D" author="Osborn, H." box="[391,531,538,562]" pageId="2" pageNumber="383" pagination="163 - 192" refId="ref12261" refString="Osborn, H. (1934) Hemiptera - Cicadellidae (Jassidae). Insects of Samoa and other Samoan terrestrial Arthropoda, II (4), 163 - 192." type="journal article" year="1934">Osborn, 1934</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
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<subSubSection id="99C96513FFBEFF92FF66F5488248F77D" pageId="2" pageNumber="383" type="diagnosis">
<paragraph id="D16C3698FFBEFF92FF66F5488562F078" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="383">
<emphasis id="E3A7EA8AFFBEFF92FF66F548861DF175" bold="true" box="[151,274,608,634]" pageId="2" pageNumber="383">Diagnosis.</emphasis>
Head anteriorly foliaceous, anterior margin rim-like with a pair of marginal carina; vertex longer medially than next to eyes, slightly transversely sulcate between anterior corners of eyes, smooth or finely obliquely striate to sulcus or to anterior margin, transversely striate each side of margin; ocelli on marginal rim close to corresponding eye; face convex in profile, dorsally concave with fine transverse striations extending laterally onto ocellocular area and then ventrally to antennal ledge, remaining ventral part of face shagreen; antennae short, arising approximately lower than mid-height of eyes in facial view; anterior tentorial branches Y-shaped with asymmetric arms. Pronotum with sides short, disc finely transversely striate or smooth. Forewing with medial and outer subapical cells closed, inner subapical cell open.
</paragraph>
<paragraph id="D16C3698FFBEFF92FF36F4A8866FF741" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="383">
Male subgenital plates without macrosetae, with fine dorsolateral setae, sometimes with digitate apex crenulate. Connective with apices of arms close or touching, sometimes fused but rarely widely spaced (
<taxonomicName id="16D34D1BFFBEFF92FB41F48D825FF0B1" box="[1200,1360,932,958]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="E3A7EA8AFFBEFF92FB41F48D825FF0B1" box="[1200,1360,932,958]" italics="true" pageId="2" pageNumber="383">H. bimaculata</emphasis>
</taxonomicName>
); stem short to long and usually extended caudally into a long distal process, the latter usually bifurcate basally or apically, rarely stem without long extension (
<taxonomicName id="16D34D1BFFBEFF92FDC3F4C585A3F709" box="[562,684,1005,1030]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="E3A7EA8AFFBEFF92FDC3F4C585A3F709" box="[562,684,1005,1030]" italics="true" pageId="2" pageNumber="383">H. spinosa</emphasis>
</taxonomicName>
and
<taxonomicName id="16D34D1BFFBEFF92FD15F4C58444F709" box="[740,843,1004,1030]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="E3A7EA8AFFBEFF92FD15F4C58444F709" box="[740,843,1004,1030]" italics="true" pageId="2" pageNumber="383">H. brevis</emphasis>
</taxonomicName>
). Aedeagus situated either dorsad of connective and connected to it by membrane or articulated with connective; with basal apodeme short to long, sometimes with a lobe on each side.
</paragraph>
<paragraph id="D16C3698FFBEFF92FF36F3708248F77D" blockId="2.[151,1437,608,2039]" box="[199,1351,1112,1139]" pageId="2" pageNumber="383">Female genitalia with first valvulae with strigate sculpture dorsally; second valvulae with serrate teeth.</paragraph>
</subSubSection>
<subSubSection id="99C96513FFBEFF93FF36F35483A8F3BE" lastPageId="3" lastPageNumber="384" pageId="2" pageNumber="383" type="description">
<paragraph id="D16C3698FFBEFF92FF36F354852FF628" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="383">
<emphasis id="E3A7EA8AFFBEFF92FF36F3548656F799" bold="true" box="[199,345,1148,1174]" pageId="2" pageNumber="383">Description.</emphasis>
Small to medium sized leafhoppers. Sordid or brownish yellow to stramineous. Vertex with or without brown marking. Pronotum and mesonotum laterally sometimes with one or two lateral brown spots, respectively. Forewings brownish hyaline usually with brown spot at apex of claval veins and clavus and usually more heavily marked with brown on veins of fifth apical cells and wing apex, rarely forewing darker brown with outer half of clavus yellow (
<taxonomicName id="16D34D1BFFBEFF92FE62F2258500F628" box="[403,527,1293,1319]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="E3A7EA8AFFBEFF92FE62F2258500F628" box="[403,527,1293,1319]" italics="true" pageId="2" pageNumber="383">H. pacifica</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="D16C3698FFBEFF92FF36F218846CF5BD" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="383">
Head very slightly wider than pronotum, anteriorly foliaceous with anterior margin rim-like with a pair of marginal carina; vertex approximately one and a half times as long medially than next to eyes (distinctly longer in
<taxonomicName id="16D34D1BFFBEFF92FA70F27D861AF69D" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="E3A7EA8AFFBEFF92FA70F27D829AF661" box="[1409,1429,1365,1390]" italics="true" pageId="2" pageNumber="383">H</emphasis>
.
<emphasis id="E3A7EA8AFFBEFF92FF66F250861AF69D" box="[151,277,1400,1426]" italics="true" pageId="2" pageNumber="383">bimaculata</emphasis>
</taxonomicName>
) and slightly transversely sulcate between anterior corners of eyes, smooth or finely obliquely striate to sulcus or to anterior margin, transversely striate each side of margin; ocelli on marginal rim close to corresponding eye; face convex in profile (except
<taxonomicName id="16D34D1BFFBEFF92FDD1F2E985D8F6D5" box="[544,727,1472,1498]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="brunifasciata">
<emphasis id="E3A7EA8AFFBEFF92FDD1F2E985D8F6D5" box="[544,727,1472,1498]" italics="true" pageId="2" pageNumber="383">H. brunifasciata</emphasis>
</taxonomicName>
), dorsally concave with fine transverse striations extending laterally onto ocellocular area and then ventrally to antennal ledge, remaining ventral part of face shagreen; antennae short, less than half body length, arising approximately at mid-height of eyes in facial view (higher in
<taxonomicName id="16D34D1BFFBEFF92FAE2F12187C0F549" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="brunifasciata">
<emphasis id="E3A7EA8AFFBEFF92FAE2F12187C0F549" italics="true" pageId="2" pageNumber="383">H. brunifasciata</emphasis>
</taxonomicName>
); anterior tentorial branches Y-shaped with asymmetric arms. Pronotum sides relatively short, carinate; disc finely transversely striate or smooth. Forewing with three subapical cells, inner open. Fore femur with AM1 present, IC row with relatively thick setae, AV setae absent. Fore tibia with dorsal setal formula 1+4. Hind femur with apical setal formula 2+2+1. Abdominal basal apodemes without lobes.
</paragraph>
<paragraph id="D16C3698FFBEFF92FF36F1948239F4F9" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="383">
Male pygofer tapered to conically rounded apex with several macrosetae distally, without processes. Segment X simple, rarely extended ventrally (
<taxonomicName id="16D34D1BFFBEFF92FDB5F1C985F0F5F5" box="[580,767,1760,1786]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="brunifasciata">
<emphasis id="E3A7EA8AFFBEFF92FDB5F1C985F0F5F5" box="[580,767,1760,1786]" italics="true" pageId="2" pageNumber="383">H. brunifasciata</emphasis>
</taxonomicName>
). Subgenital plates wide basally, thereafter lateral margin week to strongly incurved and then distally evenly tapered to apex or apex digitate, sometimes digitate apex crenulate (
<figureCitation id="49E82A1DFFBEFF92FF3EF0008625F44C" box="[207,298,1832,1859]" captionStart="FIGURE 6" captionStartId="14.[151,250,1787,1811]" captionTargetBox="[234,1347,188,1755]" captionTargetId="figure-19@14.[232,1356,181,1763]" captionTargetPageId="14" captionText="FIGURE 6. Male genitalia and female pregenital sternite of H. borneoensis species group; AB, H. borneoensis, A, style; B, connective and aedeagus, lateral view; CD, H. brevipenis; C, style; D, aedeagus and connective, lateral view; EF, H. brevispina; E, style; F, aedeagus and connective, lateral view; GJ, H. longipenis; G, style; H, subgenital pate and valve, ventral view; I, aedeagus and connective, lateral view; J, connective, dorsal view; KO, H. quadrispina; K, style; L, connective, dorsal view; M, aedeagus, dorsal view; N, subgenital plate, dorsal view; O, aedeagus and connective, lateral view; PU, H. spinosa; P, style; Q, connective, dorsal view; R, aedeagus, dorsal view; S, aedeagus and connective, lateral view; T, subgenital pate and valve, ventral view; U, female pregenital sternite." figureDoi="http://doi.org/10.5281/zenodo.4629112" httpUri="https://zenodo.org/record/4629112/files/figure.png" pageId="2" pageNumber="383">Fig. 6H</figureCitation>
); without macrosetae, with fine dorsolateral setae. Style with apical process hook-like, preapical lobe variable in length. Connective with apices of arms close or touching, sometimes fused but rarely widely spaced (
<taxonomicName id="16D34D1BFFBEFF92FA70F065861AF485" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="bimaculata">
<emphasis id="E3A7EA8AFFBEFF92FA70F065861AF485" italics="true" pageId="2" pageNumber="383">H. bimaculata</emphasis>
</taxonomicName>
); stem short to long and usually extended caudally into a long distal process, the latter usually bifurcate basally or apically, rarely stem without long extension (
<taxonomicName id="16D34D1BFFBEFF92FCFBF0BD848DF4A1" box="[778,898,1941,1966]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="E3A7EA8AFFBEFF92FCFBF0BD848DF4A1" box="[778,898,1941,1966]" italics="true" pageId="2" pageNumber="383">H. spinosa</emphasis>
</taxonomicName>
and
<taxonomicName id="16D34D1BFFBEFF92FC48F0BD8310F4A1" box="[953,1055,1940,1966]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="E3A7EA8AFFBEFF92FC48F0BD8310F4A1" box="[953,1055,1940,1966]" italics="true" pageId="2" pageNumber="383">H. brevis</emphasis>
</taxonomicName>
). Aedeagus situated either dorsad of connective and connected to it by membrane or articulated with connective; with or without processes, gonopore apical or apical on ventral surface; with basal apodeme short to long, sometimes with a lobe on each side.
</paragraph>
<paragraph id="D16C3698FFBFFF93FF36F7BF83A8F3BE" blockId="3.[151,1437,151,502]" box="[199,1191,151,177]" pageId="3" pageNumber="384">Female first valvulae with strigate sculpture dorsally; second valvulae with serrate teeth.</paragraph>
</subSubSection>
<subSubSection id="99C96513FFBFFF93FF36F79285C3F3DA" box="[199,716,186,213]" pageId="3" pageNumber="384" type="distribution">
<paragraph id="D16C3698FFBFFF93FF36F79285C3F3DA" blockId="3.[151,1437,151,502]" box="[199,716,186,213]" pageId="3" pageNumber="384">
<emphasis id="E3A7EA8AFFBFFF93FF36F7928656F3DB" bold="true" box="[199,345,186,212]" pageId="3" pageNumber="384">Distribution</emphasis>
. Oriental and Australian Region.
</paragraph>
</subSubSection>
<subSubSection id="99C96513FFBFFF93FF36F7F7864FF2F9" pageId="3" pageNumber="384" type="discussion">
<paragraph id="D16C3698FFBFFF93FF36F7F7864FF2F9" blockId="3.[151,1437,151,502]" pageId="3" pageNumber="384">
<emphasis id="E3A7EA8AFFBFFF93FF36F7F78634F3F6" bold="true" box="[199,315,223,249]" pageId="3" pageNumber="384">Remarks.</emphasis>
<taxonomicName id="16D34D1BFFBFFF93FEB2F7F786DBF3F6" authorityName="Osborn" authorityYear="1934" box="[323,468,223,249]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="E3A7EA8AFFBFFF93FEB2F7F786DBF3F6" box="[323,468,223,249]" italics="true" pageId="3" pageNumber="384">Hecaloidella</emphasis>
</taxonomicName>
can be distinguished by its convex face in profile (
<figureCitation id="49E82A1DFFBFFF93FBE4F7F78363F3F6" box="[1045,1132,223,249]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[191,1403,188,1905]" captionTargetId="figure-17@7.[179,1408,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. H. bimaculata; AB, habitus, dorsal and lateral view; C, face, D, head and thorax, anterior view; E, male genital capsule, lateral view; F, style; G, valve, subgenital plates, styles and connective, dorsal view; H, aedeagus, lateral view; I aedeagus, caudal view; J, anal tube, lateral view; K, connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4629104" httpUri="https://zenodo.org/record/4629104/files/figure.png" pageId="3" pageNumber="384">Fig. 2B</figureCitation>
) and sculpture on the head and pronotum (see Diagnosis) and in the male genitalia by lacking macrosetae on the subgenital plate, the connective with a posterior extension (except
<taxonomicName id="16D34D1BFFBFFF93FDBFF60F85C8F24F" box="[590,711,295,320]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="384" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="E3A7EA8AFFBFFF93FDBFF60F85C8F24F" box="[590,711,295,320]" italics="true" pageId="3" pageNumber="384">H. spinosa</emphasis>
</taxonomicName>
and
<taxonomicName id="16D34D1BFFBFFF93FD0FF60F846BF24E" box="[766,868,295,321]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="384" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="E3A7EA8AFFBFFF93FD0FF60F846BF24E" box="[766,868,295,321]" italics="true" pageId="3" pageNumber="384">H. brevis</emphasis>
</taxonomicName>
) and the connective frequently separated from the aedeagus; all but the first of these characters are shared with some or all
<taxonomicName id="16D34D1BFFBFFF93FC48F66383F5F26A" authority="(Paraboloponina)" authorityName="Ishihara" authorityYear="1953" baseAuthorityName="Paraboloponina" box="[953,1274,331,357]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="384" phylum="Arthropoda" rank="tribe" tribe="Drabescini">Drabescini (Paraboloponina)</taxonomicName>
(see Introduction). Within the genus most species fall into three groups (see below for details). One species (
<taxonomicName id="16D34D1BFFBFFF93FB2BF647829CF286" box="[1242,1427,367,393]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="384" phylum="Arthropoda" rank="species" species="brunifasciata">
<emphasis id="E3A7EA8AFFBFFF93FB2BF64783E1F287" box="[1242,1262,367,392]" italics="true" pageId="3" pageNumber="384">H</emphasis>
.
<emphasis id="E3A7EA8AFFBFFF93FB0FF647829CF286" box="[1278,1427,367,393]" italics="true" pageId="3" pageNumber="384">brunifasciata</emphasis>
</taxonomicName>
) is only tentatively included in the genus (
<emphasis id="E3A7EA8AFFBFFF93FD84F6BB841DF2A2" box="[629,786,403,429]" italics="true" pageId="3" pageNumber="384">incertae sedis</emphasis>
) (see species Remarks for details). Only one species,
<taxonomicName id="16D34D1BFFBFFF93FA70F6BB87FEF2DE" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="384" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="E3A7EA8AFFBFFF93FA70F6BB829AF2A3" box="[1409,1429,403,428]" italics="true" pageId="3" pageNumber="384">H</emphasis>
.
<emphasis id="E3A7EA8AFFBFFF93FF66F69F87FEF2DE" box="[151,241,439,465]" italics="true" pageId="3" pageNumber="384">pacifica</emphasis>
</taxonomicName>
is widespread in the Pacific (on cloves) and nearly all other species are restricted to Borneo or
<collectingRegion id="1317F87AFFBFFF93FAC6F69F8293F2DE" box="[1335,1436,439,465]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="384">Sulawesi</collectingRegion>
(see checklist).
</paragraph>
</subSubSection>
</treatment>
</document>