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<document id="FAC05DCF2336E6DE4AD02D9762ECE832" ID-CLB-Dataset="7470" ID-DOI="10.11646/zootaxa.4948.3.4" ID-GBIF-Dataset="6bfd846c-0304-4330-8941-95c406630230" ID-ISSN="1175-5326" ID-Zenodo-Dep="4629100" ID-ZooBank="8E0BF97A-D9E1-4F2E-BB63-82A6ED47E5DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616485698777" checkinUser="plazi" docAuthor="Xu, Deliang, Webb, Michael D., Shang, Suqin &amp; Zhang, Yalin" docDate="2021" docId="597A878EFFB6FF9FFF66F1B0841FF212" docLanguage="en" docName="zootaxa.4948.3.4.pdf" docOrigin="Zootaxa 4948 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hecaloidella longipenis Zhang &amp; Webb 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="396" masterDocId="A543FFF6FFBCFF90FFF1F728870FF30F" masterDocTitle="Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species" masterLastPageNumber="403" masterPageNumber="381" pageNumber="391" updateTime="1698934487255" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="97DE7876583D52149F3443C1C6AF7481" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4CF80280FF6FEBDBE1ADB55D7871FD1E">
<mods:title id="398AFBDA0564F49E084A1625A516776A">Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species</mods:title>
</mods:titleInfo>
<mods:name id="74D01BB73BDFAB796962CDCDE6CD932D" type="personal">
<mods:role id="75A5AC13D544FE6D4B082FDA9C36525B">
<mods:roleTerm id="A16614F0A90BF12DDC5749491727D1A8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B855CBF383CA7211EDC362CF3556A0AE">Xu, Deliang</mods:namePart>
<mods:nameIdentifier id="5D4A737CBEB5F50278D270E942516C84" type="ORCID">0000-0003-0106-5570</mods:nameIdentifier>
<mods:affiliation id="25C392623BA7946DEAEBED13B7966C07">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; 18710305867 @ 163. com; https: // orcid. org / 0000 - 0003 - 0106 - 5570</mods:affiliation>
</mods:name>
<mods:name id="769173E8AC1BA89D3226DDC88CA8F140" type="personal">
<mods:role id="F0B16DACBA9CBDFABE4825374E7ABF95">
<mods:roleTerm id="5924FF7D4798930D034F88BC228D4AC7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5BCFF7C6BA34F578E3C45972F8FCED6">Webb, Michael D.</mods:namePart>
<mods:nameIdentifier id="D03E4AE85A8E74516809716DA270455B" type="ORCID">0000-0002-1312-6142</mods:nameIdentifier>
<mods:affiliation id="7052427393E5221A55EAD784AC1F98E7">Department of Life Sciences, Natural History Museum, Cromwell Road, London, ME 15 6 QX, UK &amp; mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
<mods:nameIdentifier id="6419AF0A06AC29B581537DD361731B7F" type="email">mdwebb04@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="DE618A67EF860EE135597F515BFA3148" type="personal">
<mods:role id="C8DA8CE241838BDEAA6E7318579A54E5">
<mods:roleTerm id="67B850455B4C61AF4C8C2A4CCE561736">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13A94DA8A3723FAE834C28DCC0948066">Shang, Suqin</mods:namePart>
<mods:nameIdentifier id="DE16FA97680B5586C687DF8C6DBE447E" type="ORCID">0000-0001-9376-0735</mods:nameIdentifier>
<mods:affiliation id="AB00D00006E3B2DCB56F57DCBE1891F2">Biological Engineering Laboratory of Crop Diseases and Pests of Gansu Province, College of Plant Protection, Gansu Agricultural University, Lanzhou, Gansu 730070, China &amp; shangsq @ gsau. edu. cn; https: // orcid. org / 0000 - 0001 - 9376 - 0735</mods:affiliation>
<mods:nameIdentifier id="AAFC13AEF4F4F047F94FB4B5841DDCD2" type="email">shangsq@gsau.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="272BD356B4E1BD59F848A50388D2ED94" type="personal">
<mods:role id="401E9E6F2DB0D7D9BFDBA7928BD0A6AE">
<mods:roleTerm id="980332621BFBA07819FAC9DAFCEC686C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF9D7AC88950F99172F95D993ADAB6B5">Zhang, Yalin</mods:namePart>
<mods:nameIdentifier id="E4D4FDC84E825CD411A0FB5A8F1FC8B4" type="ORCID">0000-0002-1204-9181</mods:nameIdentifier>
<mods:affiliation id="10E40AC3A7151CDA26280A52F526F696">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; yalinzh @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0002 - 1204 - 9181</mods:affiliation>
<mods:nameIdentifier id="3B7BC7E3C51698045FFAF02AF7658DC8" type="email">yalinzh@nwsuaf.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="574689412602BA2D0D63FDA592D77078">text</mods:typeOfResource>
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<mods:titleInfo id="FF45C35AD452AF4130E59B02A7B887E9">
<mods:title id="0BCC10D9F616B28C85834FBB195711BB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1799255BD3B0A983D3DFBA7EB8B7894A">
<mods:date id="E2F9AAF7B8EE39CCE326F399454FD976">2021</mods:date>
<mods:detail id="EADC4C718F535FD6331CB4BF0C7638C8" type="pubDate">
<mods:number id="97AFF9DA841259675AFBC531C855AFE5">2021-03-22</mods:number>
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<mods:number id="CD8EE70E3B8A22DBE06C31850BE7B908">4948</mods:number>
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<mods:classification id="C309390FEDE5B94DA6E7A5496ED83639">journal article</mods:classification>
<mods:identifier id="5FFCDDDDE570F5A517DA4E7DD876FF1E" type="CLB-Dataset">7470</mods:identifier>
<mods:identifier id="0016CA5FB1BC1C15E848198DE7F8A23B" type="DOI">10.11646/zootaxa.4948.3.4</mods:identifier>
<mods:identifier id="CC04E7A16CF5B606A97A3A336D2BF071" type="GBIF-Dataset">6bfd846c-0304-4330-8941-95c406630230</mods:identifier>
<mods:identifier id="698201723E4CD39C2D3DA6C2134A5E93" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="734561EB5EE950FD69EBE7DF3F268847" type="Zenodo-Dep">4629100</mods:identifier>
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<treatment id="597A878EFFB6FF9FFF66F1B0841FF212" ID-DOI="http://doi.org/10.5281/zenodo.4629128" ID-GBIF-Taxon="180877874" ID-Zenodo-Dep="4629128" LSID="urn:lsid:plazi:treatment:597A878EFFB6FF9FFF66F1B0841FF212" httpUri="http://treatment.plazi.org/id/597A878EFFB6FF9FFF66F1B0841FF212" lastPageId="15" lastPageNumber="396" pageId="10" pageNumber="391">
<subSubSection id="99C96513FFB6FF9AFF66F1B08567F5BD" box="[151,616,1687,1714]" pageId="10" pageNumber="391" type="nomenclature">
<paragraph id="D16C3698FFB6FF9AFF66F1B08567F5BD" blockId="10.[151,616,1687,1750]" box="[151,616,1687,1714]" pageId="10" pageNumber="391">
<heading id="8A2481F4FFB6FF9AFF66F1B08567F5BD" bold="true" box="[151,616,1687,1714]" fontSize="11" level="1" pageId="10" pageNumber="391" reason="1">
<emphasis id="E3A7EA8AFFB6FF9AFF66F1B08567F5BD" bold="true" box="[151,616,1687,1714]" pageId="10" pageNumber="391">
<taxonomicName id="16D34D1BFFB6FF9AFF66F1B0850CF5BD" ID-CoL="8KK4K" authority="Zhang &amp; Webb" authorityName="Zhang &amp; Webb" authorityYear="2021" box="[151,515,1687,1714]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="longipenis" status="sp. nov.">
<emphasis id="E3A7EA8AFFB6FF9AFF66F1B087A1F5BD" bold="true" box="[151,174,1688,1714]" italics="true" pageId="10" pageNumber="391">H</emphasis>
.
<emphasis id="E3A7EA8AFFB6FF9AFF4CF1BF8634F5BD" bold="true" box="[189,315,1687,1714]" italics="true" pageId="10" pageNumber="391">longipenis</emphasis>
Zhang &amp; Webb
</taxonomicName>
<taxonomicNameLabel id="F89457F1FFB6FF9AFDFBF1B08567F5BD" box="[522,616,1688,1714]" pageId="10" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="99C96513FFB6FF9AFF66F19485A6F485" pageId="10" pageNumber="391" type="description">
<paragraph id="D16C3698FFB6FF9AFF66F194864EF5D9" blockId="10.[151,616,1687,1750]" box="[151,321,1724,1750]" pageId="10" pageNumber="391">
<figureCitation id="49E82A1DFFB6FF9AFF66F19487F5F5D9" box="[151,250,1724,1750]" captionStart="FIGURE 1" captionStartId="6.[151,250,1870,1894]" captionTargetBox="[223,1380,183,1845]" captionTargetId="figure-19@6.[205,1382,181,1846]" captionTargetPageId="6" captionText="FIGURE 1. Habitus images of Hecaloidella species; A, H. bimaculata (holotype); B, H. caudospina (holotype); C, H. nitida (holotype); D, H. nudapenis (holotype); E–G, H. pacifica (paratypes and immature); H, H. borneoensis (holotype); I, H. brevipenis (holotype); J, H. brevispina (holotype); K, H. longipenis (holotype); L, H. quadrispina (holotype); M, H. spinosa (holotype); N, H. brevis (holotype); O, H. brunnifacialis (topotype); P, H. knighti (holotype); Q, H. quadripicta (paratype); R, H. sandakanensis (holotype); S, H. singaporensis (paratype); T, H. brunifasciata (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4629102" httpUri="https://zenodo.org/record/4629102/files/figure.png" pageId="10" pageNumber="391">Figs. 1K</figureCitation>
,
<figureCitation id="49E82A1DFFB6FF9AFEF7F194864EF5D9" box="[262,321,1724,1750]" captionStart="FIGURE 6" captionStartId="14.[151,250,1787,1811]" captionTargetBox="[234,1347,188,1755]" captionTargetId="figure-19@14.[232,1356,181,1763]" captionTargetPageId="14" captionText="FIGURE 6. Male genitalia and female pregenital sternite of H. borneoensis species group; A–B, H. borneoensis, A, style; B, connective and aedeagus, lateral view; C–D, H. brevipenis; C, style; D, aedeagus and connective, lateral view; E–F, H. brevispina; E, style; F, aedeagus and connective, lateral view; G–J, H. longipenis; G, style; H, subgenital pate and valve, ventral view; I, aedeagus and connective, lateral view; J, connective, dorsal view; K–O, H. quadrispina; K, style; L, connective, dorsal view; M, aedeagus, dorsal view; N, subgenital plate, dorsal view; O, aedeagus and connective, lateral view; P–U, H. spinosa; P, style; Q, connective, dorsal view; R, aedeagus, dorsal view; S, aedeagus and connective, lateral view; T, subgenital pate and valve, ventral view; U, female pregenital sternite." figureDoi="http://doi.org/10.5281/zenodo.4629112" httpUri="https://zenodo.org/record/4629112/files/figure.png" pageId="10" pageNumber="391">6G–J</figureCitation>
</paragraph>
<paragraph id="D16C3698FFB6FF9AFF66F02C8477F411" blockId="10.[151,1437,1796,2039]" box="[151,888,1796,1823]" pageId="10" pageNumber="391">
<emphasis id="E3A7EA8AFFB6FF9AFF66F02C8626F411" bold="true" box="[151,297,1796,1822]" pageId="10" pageNumber="391">Description.</emphasis>
Body length (including tegmina), male:
<quantity id="162B9B7DFFB6FF9AFD00F02C847BF411" box="[753,884,1796,1822]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.3" metricValueMin="6.2" pageId="10" pageNumber="391" unit="mm" value="6.25" valueMax="6.3" valueMin="6.2">6.2–6.3mm</quantity>
.
</paragraph>
<paragraph id="D16C3698FFB6FF9AFF36F0008507F44C" blockId="10.[151,1437,1796,2039]" box="[199,520,1832,1859]" pageId="10" pageNumber="391">
Colour as in
<taxonomicName id="16D34D1BFFB6FF9AFEA6F001850FF44D" box="[343,512,1832,1859]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="E3A7EA8AFFB6FF9AFEA6F0018664F44D" box="[343,363,1833,1858]" italics="true" pageId="10" pageNumber="391">H</emphasis>
.
<emphasis id="E3A7EA8AFFB6FF9AFE88F000850FF44D" box="[377,512,1832,1858]" italics="true" pageId="10" pageNumber="391">borneoensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D16C3698FFB6FF9AFF36F06485A6F485" blockId="10.[151,1437,1796,2039]" pageId="10" pageNumber="391">
Male genitalia similar to
<taxonomicName id="16D34D1BFFB6FF9AFE1BF0658599F469" box="[490,662,1868,1895]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="E3A7EA8AFFB6FF9AFE1BF06586F1F469" box="[490,510,1869,1894]" italics="true" pageId="10" pageNumber="391">H</emphasis>
.
<emphasis id="E3A7EA8AFFB6FF9AFDFEF0648599F469" box="[527,662,1868,1894]" italics="true" pageId="10" pageNumber="391">borneoensis</emphasis>
</taxonomicName>
but aedeagal shaft lacking a sinuate apex and the gonopore arising slightly basad of mid-length on ventral surface.
</paragraph>
</subSubSection>
<subSubSection id="99C96513FFB6FF9AFF36F0BC83A9F4F9" pageId="10" pageNumber="391" type="materials_examined">
<paragraph id="D16C3698FFB6FF9AFF36F0BC83A9F4F9" blockId="10.[151,1437,1796,2039]" pageId="10" pageNumber="391">
<emphasis id="E3A7EA8AFFB6FF9AFF36F0BC86BDF4A1" bold="true" box="[199,434,1940,1966]" pageId="10" pageNumber="391">Material examined.</emphasis>
<materialsCitation id="61BB3CC5FFB6FF9AFE4BF0BC8296F4DD" ID-GBIF-Occurrence="3068500326" collectingDate="1932-09-25" collectionCode="BMNH" country="Malaysia" county="Lejok" location="Foot of Mt Dulit" municipality="Tinjar" pageId="10" pageNumber="391" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="0E68883AFFB6FF9AFE4BF0BC8528F4A1" box="[442,551,1940,1966]" pageId="10" pageNumber="391" type="holotype">Holotype</typeStatus>
:
<specimenCount id="C7D5FD11FFB6FF9AFDC2F0BD8545F4A0" box="[563,586,1941,1967]" pageId="10" pageNumber="391" type="male">♂</specimenCount>
,
<collectingCountry id="A9C47608FFB6FF9AFDA6F0BC85CFF4A1" box="[599,704,1940,1966]" name="Malaysia" pageId="10" pageNumber="391">Malaysia</collectingCountry>
, Sarawak,
<location id="D40C6043FFB6FF9AFCCCF0BD84F1F4A1" LSID="urn:lsid:plazi:treatment:597A878EFFB6FF9FFF66F1B0841FF212:D40C6043FFB6FF9AFCCCF0BD84F1F4A1" box="[829,1022,1940,1967]" country="Malaysia" county="Lejok" municipality="Tinjar" name="Foot of Mt Dulit" pageId="10" pageNumber="391">Foot of Mt Dulit</location>
, junction of rivers
<collectingMunicipality id="3108ACE2FFB6FF9AFB26F0BC8215F4A1" box="[1239,1306,1940,1966]" pageId="10" pageNumber="391">Tinjar</collectingMunicipality>
and
<collectingCounty id="380D4E14FFB6FF9AFAA5F0BC8297F4A1" box="[1364,1432,1940,1966]" pageId="10" pageNumber="391">Lejok</collectingCounty>
, at light,
<date id="A56D1058FFB6FF9AFF09F090867DF4DC" box="[248,370,1976,2003]" pageId="10" pageNumber="391" value="1932-09-25">
<collectingDate id="B529E9B0FFB6FF9AFF09F090867DF4DC" box="[248,370,1976,2003]" pageId="10" pageNumber="391" value="1932-09-25">25.ix.1932</collectingDate>
</date>
,
<collectorName id="7C26534EFFB6FF9AFE8EF0918501F4DD" box="[383,526,1976,2003]" pageId="10" pageNumber="391">B.M. Hobby</collectorName>
&amp;
<collectorName id="7C26534EFFB6FF9AFDC5F09085CAF4DC" box="[564,709,1976,2003]" pageId="10" pageNumber="391">A.W. Moore</collectorName>
,
<collectorName id="7C26534EFFB6FF9AFD20F0908327F4DD" box="[721,1064,1976,2002]" pageId="10" pageNumber="391">Oxford University Expedition</collectorName>
, NHMUK 010591763 (
<collectionCode id="B7C2AE5DFFB6FF9AFAC7F090829FF4DD" box="[1334,1424,1976,2002]" pageId="10" pageNumber="391">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="61BB3CC5FFB6FF9AFF66F0F583ADF4F9" ID-GBIF-Occurrence="3068500307" box="[151,1186,2012,2039]" collectingDate="1932-09-25" collectionCode="BMNH" country="Malaysia" county="Lejok" location="River Kapah" municipality="River Tinjar" pageId="10" pageNumber="391" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="0E68883AFFB6FF9AFF66F0F587F2F4F8" box="[151,253,2013,2039]" pageId="10" pageNumber="391" type="paratype">Paratype</typeStatus>
:
<specimenCount id="C7D5FD11FFB6FF9AFEF9F0F48623F4F9" box="[264,300,2012,2038]" pageId="10" pageNumber="391" type="male">1♂</specimenCount>
, same data as holotype except
<location id="D40C6043FFB6FF9AFD76F0F48417F4F9" LSID="urn:lsid:plazi:treatment:597A878EFFB6FF9FFF66F1B0841FF212:D40C6043FFB6FF9AFD76F0F48417F4F9" box="[647,792,2012,2038]" municipality="River Tinjar" name="River Kapah" pageId="10" pageNumber="391">River Kapah</location>
, tributary of
<collectingMunicipality id="3108ACE2FFB6FF9AFC58F0F4833FF4F9" box="[937,1072,2012,2038]" pageId="10" pageNumber="391">River Tinjar</collectingMunicipality>
(
<collectionCode id="B7C2AE5DFFB6FF9AFBB1F0F48395F4F9" box="[1088,1178,2012,2038]" pageId="10" pageNumber="391">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="85AC6610FFB7FF9BFF66F05C86BFF4F7" ID-DOI="http://doi.org/10.5281/zenodo.4629106" ID-Zenodo-Dep="4629106" httpUri="https://zenodo.org/record/4629106/files/figure.png" pageId="11" pageNumber="392" startId="11.[151,250,1908,1932]" targetBox="[202,1391,194,1881]" targetPageId="11">
<paragraph id="D16C3698FFB7FF9BFF66F05C86BFF4F7" blockId="11.[151,1436,1908,2040]" pageId="11" pageNumber="392">
<emphasis id="E3A7EA8AFFB7FF9BFF66F05C861BF483" bold="true" box="[151,276,1908,1932]" pageId="11" pageNumber="392">FIGURE 3.</emphasis>
<taxonomicName id="16D34D1BFFB7FF9BFEEDF05D8677F483" authorityName="Osborn" authorityYear="1934" box="[284,376,1908,1932]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="392" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="E3A7EA8AFFB7FF9BFEEDF05D8677F483" box="[284,376,1908,1932]" italics="true" pageId="11" pageNumber="392">H. nitida</emphasis>
</taxonomicName>
, A–D, holotype, E–I, paratype (Samoa); J–M, Maluku; A–B, head and thorax, dorsal and lateral view; C, face; D, forewing; E, male pygofer; F, style; G, subgenital plate, valve and style, dorsal view; H, connective, dorsal view; I, aedeagus, caudal view; J, aedeagus and connective, lateral view; K, aedeagus, lateral view; L, apex of connective, lateral view; M, connective, dorsal view.
</paragraph>
</caption>
<caption id="85AC6610FFB0FF9CFF66F04F87E2F4E4" ID-DOI="http://doi.org/10.5281/zenodo.4629108" ID-Zenodo-Dep="4629108" httpUri="https://zenodo.org/record/4629108/files/figure.png" pageId="12" pageNumber="393" startId="12.[151,250,1895,1919]" targetBox="[198,1374,189,1861]" targetPageId="12">
<paragraph id="D16C3698FFB0FF9CFF66F04F87E2F4E4" blockId="12.[151,1436,1895,2028]" pageId="12" pageNumber="393">
<emphasis id="E3A7EA8AFFB0FF9CFF66F04F861DF470" bold="true" box="[151,274,1895,1919]" pageId="12" pageNumber="393">FIGURE 4.</emphasis>
Male genitalia of
<taxonomicName id="16D34D1BFFB0FF9CFE3BF040852BF470" authorityName="Osborn" authorityYear="1934" box="[458,548,1895,1919]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="E3A7EA8AFFB0FF9CFE3BF040852BF470" box="[458,548,1895,1919]" italics="true" pageId="12" pageNumber="393">H. nitida</emphasis>
</taxonomicName>
species group; A–E,
<taxonomicName id="16D34D1BFFB0FF9CFD0BF0408482F470" box="[762,909,1895,1919]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="caudospina">
<emphasis id="E3A7EA8AFFB0FF9CFD0BF0408482F470" box="[762,909,1895,1919]" italics="true" pageId="12" pageNumber="393">H. caudospina</emphasis>
</taxonomicName>
; A–B, aedeagus, lateral and caudal view; C–D, connective, lateral and dorsal view; E, style; F–J,
<taxonomicName id="16D34D1BFFB0FF9CFD85F0A485F3F4AC" box="[628,764,1931,1955]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="nudapenis">
<emphasis id="E3A7EA8AFFB0FF9CFD85F0A485F3F4AC" box="[628,764,1931,1955]" italics="true" pageId="12" pageNumber="393">H. nudapenis</emphasis>
</taxonomicName>
; F–G, aedeagus, lateral and caudal view; H–I, connective, lateral and dorsal view; J, style; K–O,
<taxonomicName id="16D34D1BFFB0FF9CFE26F0988547F4C8" box="[471,584,1967,1991]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="E3A7EA8AFFB0FF9CFE26F0988547F4C8" box="[471,584,1967,1991]" italics="true" pageId="12" pageNumber="393">H. pacifica</emphasis>
</taxonomicName>
; K–L, aedeagus, lateral and caudal view; M–N, connective, lateral and dorsal view; O, style.
</paragraph>
</caption>
<caption id="85AC6610FFB1FF9DFF66F0B0843BF4F6" ID-DOI="http://doi.org/10.5281/zenodo.4629110" ID-Zenodo-Dep="4629110" httpUri="https://zenodo.org/record/4629110/files/figure.png" pageId="13" pageNumber="394" startId="13.[151,250,1944,1968]" targetBox="[184,1406,185,1907]" targetPageId="13">
<paragraph id="D16C3698FFB1FF9DFF66F0B0843BF4F6" blockId="13.[151,1437,1944,2041]" pageId="13" pageNumber="394">
<emphasis id="E3A7EA8AFFB1FF9DFF66F0B0861CF4BE" bold="true" box="[151,275,1944,1969]" pageId="13" pageNumber="394">FIGURE 5.</emphasis>
<taxonomicName id="16D34D1BFFB1FF9DFEEBF0B186BBF4BE" box="[282,436,1945,1969]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="394" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="E3A7EA8AFFB1FF9DFEEBF0B186BBF4BE" box="[282,436,1945,1969]" italics="true" pageId="13" pageNumber="394">H. borneoensis</emphasis>
</taxonomicName>
; A–B, habitus, dorsal and lateral view; C, face; D, head and thorax, anterior view; E, male genital capsule, lateral view; F, valve and subgenital plate, dorsal view; G, style and connective, dorsal view; H, style; I, aedeagus and connective, lateral view; J–K, aedeagus, anterior and caudal view.
</paragraph>
</caption>
<caption id="85AC6610FFB2FF9EFF66F1D38545F4E4" ID-DOI="http://doi.org/10.5281/zenodo.4629112" ID-Zenodo-Dep="4629112" httpUri="https://zenodo.org/record/4629112/files/figure.png" pageId="14" pageNumber="395" startId="14.[151,250,1787,1811]" targetBox="[234,1347,188,1755]" targetPageId="14">
<paragraph id="D16C3698FFB2FF9EFF66F1D38545F4E4" blockId="14.[151,1437,1787,2027]" pageId="14" pageNumber="395">
<emphasis id="E3A7EA8AFFB2FF9EFF66F1D3861BF41C" bold="true" box="[151,276,1787,1811]" pageId="14" pageNumber="395">FIGURE 6.</emphasis>
Male genitalia and female pregenital sternite of
<taxonomicName id="16D34D1BFFB2FF9EFCF9F1D484ADF41C" box="[776,930,1787,1811]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="395" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="E3A7EA8AFFB2FF9EFCF9F1D484ADF41C" box="[776,930,1787,1811]" italics="true" pageId="14" pageNumber="395">H. borneoensis</emphasis>
</taxonomicName>
species group; A–B,
<taxonomicName id="16D34D1BFFB2FF9EFB71F1D48215F41C" box="[1152,1306,1787,1811]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="395" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="E3A7EA8AFFB2FF9EFB71F1D48215F41C" box="[1152,1306,1787,1811]" italics="true" pageId="14" pageNumber="395">H. borneoensis</emphasis>
</taxonomicName>
, A, style; B, connective and aedeagus, lateral view; C–D,
<taxonomicName id="16D34D1BFFB2FF9EFD97F00885E0F438" box="[614,751,1823,1847]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="395" phylum="Arthropoda" rank="species" species="brevipenis">
<emphasis id="E3A7EA8AFFB2FF9EFD97F00885E0F438" box="[614,751,1823,1847]" italics="true" pageId="14" pageNumber="395">H. brevipenis</emphasis>
</taxonomicName>
; C, style; D, aedeagus and connective, lateral view; E–F,
<taxonomicName id="16D34D1BFFB2FF9EFAB0F00887C1F454" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="395" phylum="Arthropoda" rank="species" species="brevispina">
<emphasis id="E3A7EA8AFFB2FF9EFAB0F00887C1F454" italics="true" pageId="14" pageNumber="395">H. brevispina</emphasis>
</taxonomicName>
; E, style; F, aedeagus and connective, lateral view; G–J,
<taxonomicName id="16D34D1BFFB2FF9EFCF5F06C8483F454" box="[772,908,1859,1883]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="395" phylum="Arthropoda" rank="species" species="longipenis">
<emphasis id="E3A7EA8AFFB2FF9EFCF5F06C8483F454" box="[772,908,1859,1883]" italics="true" pageId="14" pageNumber="395">H. longipenis</emphasis>
</taxonomicName>
; G, style; H, subgenital pate and valve, ventral view; I, aedeagus and connective, lateral view; J, connective, dorsal view; K–O,
<taxonomicName id="16D34D1BFFB2FF9EFC79F040832EF470" box="[904,1057,1895,1919]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="395" phylum="Arthropoda" rank="species" species="quadrispina">
<emphasis id="E3A7EA8AFFB2FF9EFC79F040832EF470" box="[904,1057,1895,1919]" italics="true" pageId="14" pageNumber="395">H. quadrispina</emphasis>
</taxonomicName>
; K, style; L, connective, dorsal view; M, aedeagus, dorsal view; N, subgenital plate, dorsal view; O, aedeagus and connective, lateral view; P–U,
<taxonomicName id="16D34D1BFFB2FF9EFB24F0A4824FF4AC" box="[1237,1344,1932,1955]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="395" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="E3A7EA8AFFB2FF9EFB24F0A4824FF4AC" box="[1237,1344,1932,1955]" italics="true" pageId="14" pageNumber="395">H. spinosa</emphasis>
</taxonomicName>
; P, style; Q, connective, dorsal view; R, aedeagus, dorsal view; S, aedeagus and connective, lateral view; T, subgenital pate and valve, ventral view; U, female pregenital sternite.
</paragraph>
</caption>
<subSubSection id="99C96513FFB3FF9FFF36F7BF84E5F3BE" box="[199,1002,151,177]" pageId="15" pageNumber="396" type="etymology">
<paragraph id="D16C3698FFB3FF9FFF36F7BF84E5F3BE" blockId="15.[151,1436,151,285]" box="[199,1002,151,177]" pageId="15" pageNumber="396">
<emphasis id="E3A7EA8AFFB3FF9FFF36F7BF8641F3BE" bold="true" box="[199,334,151,177]" pageId="15" pageNumber="396">Etymology.</emphasis>
This species is named for its relatively long aedeagal shaft.
</paragraph>
</subSubSection>
<subSubSection id="99C96513FFB3FF9FFF36F793841FF212" pageId="15" pageNumber="396" type="discussion">
<paragraph id="D16C3698FFB3FF9FFF36F793841FF212" blockId="15.[151,1436,151,285]" pageId="15" pageNumber="396">
<emphasis id="E3A7EA8AFFB3FF9FFF36F7938634F3DA" bold="true" box="[199,315,187,213]" pageId="15" pageNumber="396">Remarks.</emphasis>
This species is similar to
<taxonomicName id="16D34D1BFFB3FF9FFDA6F793840FF3DA" box="[599,768,187,213]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="396" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="E3A7EA8AFFB3FF9FFDA6F793840FF3DA" box="[599,768,187,213]" italics="true" pageId="15" pageNumber="396">H. borneoensis</emphasis>
</taxonomicName>
and from the same locality. It can be distinguished from this and other species by its relatively long aedeagal shaft with gonopore arising at approximately mid-length of shaft on ventral surface. See also Remarks under
<taxonomicName id="16D34D1BFFB3FF9FFD91F62B8406F212" box="[608,777,259,285]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="396" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="E3A7EA8AFFB3FF9FFD91F62B8406F212" box="[608,777,259,285]" italics="true" pageId="15" pageNumber="396">H. borneoensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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