Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species Author Xu, Deliang 0000-0003-0106-5570 Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A & F University, Yangling, Shaanxi 712100, China & 18710305867 @ 163. com; https: // orcid. org / 0000 - 0003 - 0106 - 5570 Author Webb, Michael D. 0000-0002-1312-6142 Department of Life Sciences, Natural History Museum, Cromwell Road, London, ME 15 6 QX, UK & mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142 mdwebb04@gmail.com Author Shang, Suqin 0000-0001-9376-0735 Biological Engineering Laboratory of Crop Diseases and Pests of Gansu Province, College of Plant Protection, Gansu Agricultural University, Lanzhou, Gansu 730070, China & shangsq @ gsau. edu. cn; https: // orcid. org / 0000 - 0001 - 9376 - 0735 shangsq@gsau.edu.cn Author Zhang, Yalin 0000-0002-1204-9181 Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A & F University, Yangling, Shaanxi 712100, China & yalinzh @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0002 - 1204 - 9181 yalinzh@nwsuaf.edu.cn text Zootaxa 2021 2021-03-22 4948 3 381 403 journal article 7470 10.11646/zootaxa.4948.3.4 6bfd846c-0304-4330-8941-95c406630230 1175-5326 4629100 8E0BF97A-D9E1-4F2E-BB63-82A6ED47E5DC H . longipenis Zhang & Webb sp. nov. Figs. 1K , 6G–J Description. Body length (including tegmina), male: 6.2–6.3mm . Colour as in H . borneoensis . Male genitalia similar to H . borneoensis but aedeagal shaft lacking a sinuate apex and the gonopore arising slightly basad of mid-length on ventral surface. Material examined. Holotype : ♂ , Malaysia , Sarawak, Foot of Mt Dulit , junction of rivers Tinjar and Lejok , at light, 25.ix.1932 , B.M. Hobby & A.W. Moore , Oxford University Expedition , NHMUK 010591763 ( BMNH ) . Paratype : 1♂ , same data as holotype except River Kapah , tributary of River Tinjar ( BMNH ) . FIGURE 3. H. nitida , A–D, holotype, E–I, paratype (Samoa); J–M, Maluku; A–B, head and thorax, dorsal and lateral view; C, face; D, forewing; E, male pygofer; F, style; G, subgenital plate, valve and style, dorsal view; H, connective, dorsal view; I, aedeagus, caudal view; J, aedeagus and connective, lateral view; K, aedeagus, lateral view; L, apex of connective, lateral view; M, connective, dorsal view. FIGURE 4. Male genitalia of H. nitida species group; A–E, H. caudospina ; A–B, aedeagus, lateral and caudal view; C–D, connective, lateral and dorsal view; E, style; F–J, H. nudapenis ; F–G, aedeagus, lateral and caudal view; H–I, connective, lateral and dorsal view; J, style; K–O, H. pacifica ; K–L, aedeagus, lateral and caudal view; M–N, connective, lateral and dorsal view; O, style. FIGURE 5. H. borneoensis ; A–B, habitus, dorsal and lateral view; C, face; D, head and thorax, anterior view; E, male genital capsule, lateral view; F, valve and subgenital plate, dorsal view; G, style and connective, dorsal view; H, style; I, aedeagus and connective, lateral view; J–K, aedeagus, anterior and caudal view. FIGURE 6. Male genitalia and female pregenital sternite of H. borneoensis species group; A–B, H. borneoensis , A, style; B, connective and aedeagus, lateral view; C–D, H. brevipenis ; C, style; D, aedeagus and connective, lateral view; E–F, H. brevispina ; E, style; F, aedeagus and connective, lateral view; G–J, H. longipenis ; G, style; H, subgenital pate and valve, ventral view; I, aedeagus and connective, lateral view; J, connective, dorsal view; K–O, H. quadrispina ; K, style; L, connective, dorsal view; M, aedeagus, dorsal view; N, subgenital plate, dorsal view; O, aedeagus and connective, lateral view; P–U, H. spinosa ; P, style; Q, connective, dorsal view; R, aedeagus, dorsal view; S, aedeagus and connective, lateral view; T, subgenital pate and valve, ventral view; U, female pregenital sternite. Etymology. This species is named for its relatively long aedeagal shaft. Remarks. This species is similar to H. borneoensis and from the same locality. It can be distinguished from this and other species by its relatively long aedeagal shaft with gonopore arising at approximately mid-length of shaft on ventral surface. See also Remarks under H. borneoensis .