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<document id="2BEE3A38379606407CFBD0E6C335E40D" ID-DOI="10.11646/zootaxa.5570.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734655" ID-ZooBank="1995C18B-1993-4DC4-B8AB-E4891F148A1E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737736390657" checkinUser="plazi" docAuthor="Dayananda, Naruwan &amp; Benjamin, Suresh P." docDate="2025" docId="039D5914FF94FFA9FF4DF8E7FCB7FE2F" docLanguage="en" docName="zootaxa.5570.1.4.pdf" docOrigin="Zootaxa 5570 (1)" docSource="https://doi.org/10.11646/zootaxa.5570.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mallinella milkyway Dayananda &amp; Benjamin, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="106" masterDocId="FFA4216CFF97FFAFFFDAFFE9FFD8FFFA" masterDocTitle="Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka" masterLastPageNumber="118" masterPageNumber="100" pageNumber="103" updateTime="1737736825942" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="884E17C329252DD627397D3A32592139">Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka</mods:title>
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<mods:namePart id="7E8605E16511F558C4C1F27C2636E10B">Dayananda, Naruwan</mods:namePart>
<mods:affiliation id="C19D651C5A46C8B309869D3C03216C29">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka.</mods:affiliation>
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<mods:namePart id="3590E8EADA29DCCB6B22458ABF00443A">Benjamin, Suresh P.</mods:namePart>
<mods:affiliation id="C326A98C1F580340A2831F3BB770E080">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka.</mods:affiliation>
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<mods:title id="8A018FE66C0D5FCB3F4ED995A0E29CAE">Zootaxa</mods:title>
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<mods:date id="B745158086454CF8DCE54490E431B2E1">2025</mods:date>
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<treatment id="039D5914FF94FFA9FF4DF8E7FCB7FE2F" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F" httpUri="http://treatment.plazi.org/id/039D5914FF94FFA9FF4DF8E7FCB7FE2F" lastPageId="6" lastPageNumber="106" pageId="3" pageNumber="103">
<subSubSection id="C32EBB89FF94FFACFF4DF8E7FE2BF8D3" box="[151,499,1806,1833]" pageId="3" pageNumber="103" type="nomenclature">
<paragraph id="8B8BE802FF94FFACFF4DF8E7FE2BF8D3" blockId="3.[151,537,1806,1869]" box="[151,499,1806,1833]" pageId="3" pageNumber="103">
<heading id="D0C35F6EFF94FFACFF4DF8E7FE2BF8D3" bold="true" box="[151,499,1806,1833]" fontSize="11" level="1" pageId="3" pageNumber="103" reason="1">
<emphasis id="B9403410FF94FFACFF4DF8E7FE2BF8D3" bold="true" box="[151,499,1806,1833]" pageId="3" pageNumber="103">
<taxonomicName id="4C349381FF94FFACFF4DF8E7FE56F8D3" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[151,398,1806,1833]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="milkyway" status="sp. nov.">
<emphasis id="B9403410FF94FFACFF4DF8E7FE56F8D3" bold="true" box="[151,398,1806,1833]" italics="true" pageId="3" pageNumber="103">Mallinella milkyway</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273896BFF94FFACFE4FF8E6FE2BF8D3" box="[405,499,1807,1833]" pageId="3" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF94FFACFF4DF8DAFDC1F8B7" box="[151,537,1843,1869]" pageId="3" pageNumber="103" type="description">
<paragraph id="8B8BE802FF94FFACFF4DF8DAFDC1F8B7" blockId="3.[151,537,1806,1869]" box="[151,537,1843,1869]" pageId="3" pageNumber="103">
<figureCitation id="130FF487FF94FFACFF4DF8DAFF06F8B7" box="[151,222,1843,1869]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="3" pageNumber="103">Figs 2</figureCitation>
,
<figureCitation id="130FF487FF94FFACFF30F8DAFEE5F8B7" box="[234,317,1843,1869]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="3" pageNumber="103">6B, 6G</figureCitation>
,
<figureCitation id="130FF487FF94FFACFE93F8DAFE44F8B7" box="[329,412,1843,1869]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="3" pageNumber="103">7B, 7G</figureCitation>
,
<figureCitation id="130FF487FF94FFACFE72F8DAFE13F8B7" box="[424,459,1843,1869]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="3" pageNumber="103">8B</figureCitation>
,
<figureCitation id="130FF487FF94FFACFE0DF8DAFDC1F8B7" box="[471,537,1843,1869]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="3" pageNumber="103">9CD</figureCitation>
</paragraph>
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<subSubSection id="C32EBB89FF94FFAAFF4DF892FC03FD7F" lastPageId="5" lastPageNumber="105" pageId="3" pageNumber="103" type="materials_examined">
<paragraph id="8B8BE802FF94FFACFF4DF892FD25F827" blockId="3.[151,1436,1914,2013]" pageId="3" pageNumber="103">
<emphasis id="B9403410FF94FFACFF4DF892FE9FF86E" bold="true" box="[151,327,1914,1941]" pageId="3" pageNumber="103">Type material.</emphasis>
<materialsCitation id="3B5CE25FFF94FFACFE8AF892FB68F843" collectingDate="2010-02-02" collectorName="S. Batuwita" country="Sri Lanka" county="Kandy District" elevation="1360" latitude="7.361111" location="Knuckles Range" longLatPrecision="21" longitude="80.833336" municipality="Corbett's Gap" pageId="3" pageNumber="103" specimenCount="1" specimenCount-male="1" stateProvince="Central" typeStatus="holotype">
<emphasis id="B9403410FF94FFACFE8AF892FCB4F86F" bold="true" box="[336,876,1914,1944]" pageId="3" pageNumber="103">
<typeStatus id="548F56A0FF94FFACFE8AF892FE65F86F" box="[336,445,1915,1941]" pageId="3" pageNumber="103" type="holotype">Holotype</typeStatus>
<specimenCount id="9D32238BFF94FFACFE1CF895FE05F862" box="[454,477,1916,1944]" count="1" pageId="3" pageNumber="103" type="male"></specimenCount>
:
<collectingCountry id="F323A892FF94FFACFE37F893FD56F86E" box="[493,654,1914,1940]" name="Sri Lanka" pageId="3" pageNumber="103">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF94FFACFD46F893FCB4F86F" bold="true" box="[668,876,1914,1941]" italics="true" pageId="3" pageNumber="103">
<collectingRegion id="49F026E0FF94FFACFD46F893FCBDF86F" box="[668,869,1914,1941]" country="Sri Lanka" name="Central" pageId="3" pageNumber="103">Central Province</collectingRegion>
:
</emphasis>
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<collectingCounty id="62EA908EFF94FFACFCAFF892FBF8F86F" box="[885,1056,1915,1941]" pageId="3" pageNumber="103">Kandy District</collectingCounty>
,
<collectingMunicipality id="6BEF7278FF94FFACFBF7F892FB15F86F" box="[1069,1229,1915,1941]" pageId="3" pageNumber="103">Corbetts Gap</collectingMunicipality>
,
<location id="8EEBBED9FF94FFACFB01F892FA40F86F" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF94FFACFB01F892FA40F86F" box="[1243,1432,1915,1941]" country="Sri Lanka" county="Kandy District" latitude="7.361111" longLatPrecision="21" longitude="80.833336" municipality="Corbett's Gap" name="Knuckles Range" pageId="3" pageNumber="103" stateProvince="Central">Knuckles Range</location>
,
<geoCoordinate id="EE008EC5FF94FFACFF4DF876FEFCF843" box="[151,292,1951,1977]" degrees="07" direction="north" minutes="21" orientation="latitude" pageId="3" pageNumber="103" precision="15" seconds="40" value="7.361111">07°2140N</geoCoordinate>
<geoCoordinate id="EE008EC5FF94FFACFEF4F877FE62F843" box="[302,442,1950,1977]" degrees="80" direction="east" minutes="50" orientation="longitude" pageId="3" pageNumber="103" precision="15" seconds="00" value="80.833336">80°5000E</geoCoordinate>
,
<quantity id="4CCC45E7FF94FFACFE12F876FDF9F843" box="[456,545,1951,1977]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="3" pageNumber="103" unit="m" value="1360.0">
<elevation id="00190F31FF94FFACFE12F876FDF9F843" box="[456,545,1951,1977]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="3" pageNumber="103" unit="m" value="1360.0">1360 m</elevation>
</quantity>
,
<date id="FF8ACEC2FF94FFACFDF5F876FD37F843" box="[559,751,1951,1977]" pageId="3" pageNumber="103" value="2010-02-02">
<collectingDate id="EFCE372AFF94FFACFDF5F876FD37F843" box="[559,751,1951,1977]" pageId="3" pageNumber="103" value="2010-02-02">2 February 2010</collectingDate>
</date>
, leg.
<collectorName id="26C18DD4FF94FFACFCF4F876FC6BF843" box="[814,947,1951,1977]" pageId="3" pageNumber="103">S. Batuwita</collectorName>
<emphasis id="B9403410FF94FFACFC66F876FC37F843" box="[956,1007,1951,1977]" italics="true" pageId="3" pageNumber="103">et al</emphasis>
. (IFS_ZOD_36)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF94FFACFB66F876FD21F827" collectingDate="2010-02-02" collectorName="S. Batuwita" country="Sri Lanka" county="Kandy District" elevation="1360" latitude="7.361111" location="Knuckles Range" longLatPrecision="21" longitude="80.833336" municipality="Corbett's Gap" pageId="3" pageNumber="103" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Paratypes" typeStatus="paratype">
<emphasis id="B9403410FF94FFACFB66F876FAE5F843" bold="true" box="[1212,1341,1951,1977]" pageId="3" pageNumber="103">
<typeStatus id="548F56A0FF94FFACFB66F876FAE0F843" box="[1212,1336,1951,1977]" pageId="3" pageNumber="103" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D32238BFF94FFACFA9CF876FAB1F843" box="[1350,1385,1951,1977]" count="2" pageId="3" pageNumber="103" type="male">2♂</specimenCount>
<specimenCount id="9D32238BFF94FFACFAA8F876FA4DF843" box="[1394,1429,1951,1977]" count="1" pageId="3" pageNumber="103" type="female">1♀</specimenCount>
, with same data as for holotype (IFS_ZOD_34, 35, 37)
</materialsCitation>
.
</paragraph>
<caption id="DF4BB88AFF93FFABFF4DF938FC55F81F" ID-DOI="http://doi.org/10.5281/zenodo.14734664" ID-Zenodo-Dep="14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="4" pageNumber="104" startId="4.[151,250,1745,1769]" targetBox="[256,1330,187,1717]" targetPageId="4" targetType="figure">
<paragraph id="8B8BE802FF93FFABFF4DF938FC55F81F" blockId="4.[151,1437,1745,2021]" pageId="4" pageNumber="104">
<emphasis id="B9403410FF93FFABFF4DF938FECDF913" bold="true" box="[151,277,1745,1769]" pageId="4" pageNumber="104">FIGURE 2.</emphasis>
<taxonomicName id="4C349381FF93FFABFEC4F938FE28F913" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[286,496,1745,1769]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="104" phylum="Arthropoda" rank="species" species="milkyway" status="sp. nov.">
<emphasis id="B9403410FF93FFABFEC4F938FE28F913" box="[286,496,1745,1769]" italics="true" pageId="4" pageNumber="104">Mallinella milkyway</emphasis>
</taxonomicName>
<emphasis id="B9403410FF93FFABFE22F938FD93F913" bold="true" box="[504,587,1745,1769]" pageId="4" pageNumber="104">
<taxonomicNameLabel id="A273896BFF93FFABFE22F938FD93F913" box="[504,587,1745,1769]" pageId="4" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8BE802FF92FFAAFF1DFF7FFC03FD7F" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DFF7FFE27FF4A" bold="true" box="[199,511,150,176]" pageId="5" pageNumber="105">Other material examined.</emphasis>
<materialsCitation id="3B5CE25FFF92FFAAFDD2FF7FFE40FEBB" collectingDate="2010-08-24" collectingDateMax="2019-02-16" collectingDateMin="2010-08-24" collectorName="Benjamin &amp; Athukorala" country="Sri Lanka" elevation="180" latitude="7.859444" location="Matale District" longLatPrecision="21" longitude="80.67444" pageId="5" pageNumber="105" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Central">
<emphasis id="B9403410FF92FFAAFDD2FF7FFC5BFF4B" bold="true" box="[520,899,150,177]" pageId="5" pageNumber="105">
<collectingCountry id="F323A892FF92FFAAFDD2FF7FFD70FF4A" box="[520,680,150,176]" name="Sri Lanka" pageId="5" pageNumber="105">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF92FFAAFD6DFF7FFC5BFF4B" bold="true" box="[695,899,150,177]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFD6DFF7FFCA5FF4B" box="[695,893,150,177]" country="Sri Lanka" name="Central" pageId="5" pageNumber="105">Central province</collectingRegion>
:
</emphasis>
</emphasis>
<location id="8EEBBED9FF92FFAAFC56FF7EFBE2FF4B" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFC56FF7EFBE2FF4B" box="[908,1082,151,177]" country="Sri Lanka" latitude="7.859444" longLatPrecision="21" longitude="80.67444" name="Matale District" pageId="5" pageNumber="105" stateProvince="Central">Matale District</location>
, NIFS arboretum,
<geoCoordinate id="EE008EC5FF92FFAAFAD5FF7FFA45FF4B" box="[1295,1437,150,177]" degrees="07" direction="north" minutes="51" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="34" value="7.859444">07°5134N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFF4DFF52FEFCFF2F" box="[151,292,187,213]" degrees="80" direction="east" minutes="40" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="28" value="80.67444">80°4028E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFEF5FF52FEAFFF2F" box="[303,375,187,213]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="105" unit="m" value="180.0">
<elevation id="00190F31FF92FFAAFEF5FF52FEAFFF2F" box="[303,375,187,213]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="105" unit="m" value="180.0">180 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFE58FF52FE7EFF2F" box="[386,422,187,213]" count="1" pageId="5" pageNumber="105" type="male">1♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFE68FF52FDEEFF2F" box="[434,566,187,213]" pageId="5" pageNumber="105" value="2013-07-07">
<collectingDate id="EFCE372AFF92FFAAFE68FF52FDEEFF2F" box="[434,566,187,213]" pageId="5" pageNumber="105" value="2013-07-07">7 July 2013</collectingDate>
</date>
, leg. SP.
<collectorName id="26C18DD4FF92FFAAFD43FF52FCDEFF2F" box="[665,774,187,213]" pageId="5" pageNumber="105">Benjamin</collectorName>
<emphasis id="B9403410FF92FFAAFCD6FF52FCE5FF2F" box="[780,829,187,213]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_25);
<specimenCount id="9D32238BFF92FFAAFBDFFF52FBF2FF2F" box="[1029,1066,187,213]" count="3" pageId="5" pageNumber="105" type="male">3♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFBEFFF52FB30FF2F" box="[1077,1256,187,213]" pageId="5" pageNumber="105" value="2010-08-24">
<collectingDate id="EFCE372AFF92FFAAFBEFFF52FB30FF2F" box="[1077,1256,187,213]" pageId="5" pageNumber="105" value="2010-08-24">24 August 2010</collectingDate>
</date>
, NP.
<collectorName id="26C18DD4FF92FFAAFAC4FF52FA44FF2F" box="[1310,1436,187,213]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFF4DFF36FF13FF03" box="[151,203,223,249]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_41, 210, 211);
<specimenCount id="9D32238BFF92FFAAFDD6FF36FDE8FF03" box="[524,560,223,249]" count="3" pageId="5" pageNumber="105" type="female">3♀</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFDE5FF36FD27FF03" box="[575,767,223,249]" pageId="5" pageNumber="105" value="2016-01-20">
<collectingDate id="EFCE372AFF92FFAAFDE5FF36FD27FF03" box="[575,767,223,249]" pageId="5" pageNumber="105" value="2016-01-20">20 January 2016</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFCB4FF36FC34FF03" box="[878,1004,223,249]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFC2CFF36FBF1FF03" box="[1014,1065,223,249]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_132, 133, 134);
<specimenCount id="9D32238BFF92FFAAFAA3FF36FA44FF03" box="[1401,1436,223,249]" count="1" pageId="5" pageNumber="105" type="male">1♂</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFF1CFEEAFF31FEE7" box="[198,233,259,285]" count="1" pageId="5" pageNumber="105" type="female">1♀</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFF2DFEEAFE19FEE7" box="[247,449,259,285]" pageId="5" pageNumber="105" value="2019-02-16">
<collectingDate id="EFCE372AFF92FFAAFF2DFEEAFE19FEE7" box="[247,449,259,285]" pageId="5" pageNumber="105" value="2019-02-16">16 February 2019</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFDF3FEEAFD7FFEE7" box="[553,679,259,285]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFD74FEEAFD07FEE7" box="[686,735,259,285]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_182, 183);
<specimenCount id="9D32238BFF92FFAAFC34FEEAFBCAFEE7" box="[1006,1042,259,285]" count="1" pageId="5" pageNumber="105" type="male">1♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFBC4FEEAFB6EFEE7" box="[1054,1206,258,285]" pageId="5" pageNumber="105" value="2015-05-28">
<collectingDate id="EFCE372AFF92FFAAFBC4FEEAFB6EFEE7" box="[1054,1206,258,285]" pageId="5" pageNumber="105" value="2015-05-28">28 May 2015</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFAC4FEEAFA44FEE7" box="[1310,1436,259,285]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFF4DFECEFF12FEBB" box="[151,202,295,321]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_209)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF92FFAAFE7EFECFFBAEFE9F" collectingDate="2010-05-24" collectorName="Athukorala" country="Sri Lanka" elevation="30" latitude="8.170834" location="Wanatha Villu" longLatPrecision="21" longitude="79.875" municipality="Puttalam District" pageId="5" pageNumber="105" specimenCount="6" specimenCount-female="6" stateProvince="North Western">
<emphasis id="B9403410FF92FFAAFE7EFECFFD1FFEBB" bold="true" box="[420,711,294,321]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFE7EFECFFD18FEBB" box="[420,704,294,321]" country="Sri Lanka" name="North Western" pageId="5" pageNumber="105">North Western Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BEF7278FF92FFAAFD0AFECEFC4BFEBB" box="[720,915,295,321]" pageId="5" pageNumber="105">Puttalam District</collectingMunicipality>
,
<location id="8EEBBED9FF92FFAAFC45FECEFB9CFEBB" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFC45FECEFB9CFEBB" box="[927,1092,295,321]" country="Sri Lanka" latitude="8.170834" longLatPrecision="21" longitude="79.875" municipality="Puttalam District" name="Wanatha Villu" pageId="5" pageNumber="105" stateProvince="North Western">Wanatha Villu</location>
,
<geoCoordinate id="EE008EC5FF92FFAAFB8AFECFFB05FEBB" box="[1104,1245,294,321]" degrees="08" direction="north" minutes="10" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="15" value="8.170834">08°1015N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFB3CFECFFAABFEBB" box="[1254,1395,294,321]" degrees="79" direction="east" minutes="52" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="30" value="79.875">79°5230E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFA5AFECEFF77FE9F" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="105" unit="m" value="30.0">
<elevation id="00190F31FF92FFAAFA5AFECEFF77FE9F" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="105" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFF66FEA2FF38FE9F" box="[188,224,331,357]" count="6" pageId="5" pageNumber="105" type="female">6♀</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFF37FEA2FE5FFE9F" box="[237,391,331,357]" pageId="5" pageNumber="105" value="2010-05-24">
<collectingDate id="EFCE372AFF92FFAAFF37FEA2FE5FFE9F" box="[237,391,331,357]" pageId="5" pageNumber="105" value="2010-05-24">24 May 2010</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFE28FEA2FDA8FE9F" box="[498,624,331,357]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFDA2FEA2FD72FE9F" box="[632,682,331,357]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_23, 53, 142, 143, 144, 145)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF92FFAAFB5BFEA3FDDCFE57" collectingDate="2012-01-10" collectorName="Athukorala" country="Sri Lanka" elevation="56" latitude="8.8" location="Padaviya" longLatPrecision="21" longitude="80.75" municipality="Anuradhapura District" pageId="5" pageNumber="105" specimenCount="3" specimenCount-female="3" stateProvince="North Central">
<emphasis id="B9403410FF92FFAAFB5BFEA3FA44FE9F" bold="true" box="[1153,1436,330,357]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFB5BFEA3FA4DFE9F" box="[1153,1429,330,357]" country="Sri Lanka" name="North Central" pageId="5" pageNumber="105">North Central Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BEF7278FF92FFAAFF4DFE86FE42FE73" box="[151,410,367,393]" pageId="5" pageNumber="105">Anuradhapura District</collectingMunicipality>
,
<location id="8EEBBED9FF92FFAAFE70FE86FDCCFE73" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFE70FE86FDCCFE73" box="[426,532,367,393]" country="Sri Lanka" latitude="8.8" longLatPrecision="21" longitude="80.75" municipality="Anuradhapura District" name="Padaviya" pageId="5" pageNumber="105" stateProvince="North Central">Padaviya</location>
,
<geoCoordinate id="EE008EC5FF92FFAAFDFEFE86FD69FE73" box="[548,689,367,393]" degrees="08" direction="north" minutes="48" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="00" value="8.8">08°4800N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFD67FE87FC92FE73" box="[701,842,366,393]" degrees="80" direction="east" minutes="45" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="00" value="80.75">80°4500E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFC80FE87FC42FE73" box="[858,922,366,393]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="105" unit="m" value="56.0">
<elevation id="00190F31FF92FFAAFC80FE87FC42FE73" box="[858,922,366,393]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="105" unit="m" value="56.0">56 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFC71FE86FC17FE73" box="[939,975,367,393]" count="3" pageId="5" pageNumber="105" type="female">3♀</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFC3BFE86FB7FFE73" box="[993,1191,367,393]" pageId="5" pageNumber="105" value="2012-01-10">
<collectingDate id="EFCE372AFF92FFAAFC3BFE86FB7FFE73" box="[993,1191,367,393]" pageId="5" pageNumber="105" value="2012-01-10">10 January 2012</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFAC4FE86FA44FE73" box="[1310,1436,367,393]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFF4DFE7AFF17FE57" box="[151,207,403,429]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_158,159,160)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF92FFAAFDCEFE7AFA40FE2B" collectingDate="2023-09-26" collectorName="Dayananda" country="Sri Lanka" elevation="9" latitude="8.9366665" location="Beach Side" longLatPrecision="21" longitude="81.006386" municipality="Trincomalee District" pageId="5" pageNumber="105" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Eastern">
<emphasis id="B9403410FF92FFAAFDCEFE7AFD34FE57" bold="true" box="[532,748,403,429]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFDCEFE7AFD3DFE57" box="[532,741,403,429]" country="Sri Lanka" name="Eastern" pageId="5" pageNumber="105">Eastern Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BEF7278FF92FFAAFD23FE7AFC32FE57" box="[761,1002,403,429]" pageId="5" pageNumber="105">Trincomalee District</collectingMunicipality>
, Arisimale
<location id="8EEBBED9FF92FFAAFBAFFE7AFB25FE57" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFBAFFE7AFB25FE57" box="[1141,1277,403,429]" country="Sri Lanka" latitude="8.9366665" longLatPrecision="21" longitude="81.006386" municipality="Trincomalee District" name="Beach Side" pageId="5" pageNumber="105" stateProvince="Eastern">Beach Side</location>
,
<geoCoordinate id="EE008EC5FF92FFAAFAD5FE7BFA44FE57" box="[1295,1436,402,429]" degrees="08" direction="north" minutes="56" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="12" value="8.9366665">08°5612N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFF4DFE5EFEFBFE2B" box="[151,291,439,465]" degrees="81" direction="east" minutes="00" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="23" value="81.006386">81°0023E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFEE8FE5EFEBAFE2B" box="[306,354,439,465]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="105" unit="m" value="9.0">
<elevation id="00190F31FF92FFAAFEE8FE5EFEBAFE2B" box="[306,354,439,465]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="105" unit="m" value="9.0">9 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFEAAFE5EFE4BFE2B" box="[368,403,439,465]" count="2" pageId="5" pageNumber="105" type="female">2♀</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFE15FE5EFE2DFE2B" box="[463,501,439,465]" count="2" pageId="5" pageNumber="105" type="male">2♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFDD9FE5EFD3DFE2B" box="[515,741,439,465]" pageId="5" pageNumber="105" value="2023-09-26">
<collectingDate id="EFCE372AFF92FFAAFDD9FE5EFD3DFE2B" box="[515,741,439,465]" pageId="5" pageNumber="105" value="2023-09-26">26 September 2023</collectingDate>
</date>
, leg. NG.
<collectorName id="26C18DD4FF92FFAAFC85FE5EFC05FE2B" box="[863,989,439,465]" pageId="5" pageNumber="105">Dayananda</collectorName>
<emphasis id="B9403410FF92FFAAFC3CFE5EFBC2FE2B" box="[998,1050,439,465]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_205, 206, 207, 208)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF92FFAAFF4DFE33FC0EFD7F" collectingDate="2016-09-20" collectingDateMax="2023-10-04" collectingDateMin="2016-09-20" collectorName="Dayananda" country="Sri Lanka" elevation="4" latitude="9.62" location="Mandaitivu" longLatPrecision="21" longitude="79.97806" municipality="Jaffna District" pageId="5" pageNumber="105" specimenCount="16" specimenCount-female="5" specimenCount-male="11" stateProvince="Nothern Province">
<emphasis id="B9403410FF92FFAAFF4DFE33FEABFE0F" bold="true" box="[151,371,474,501]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFF4DFE33FEB4FE0F" box="[151,364,474,501]" country="Sri Lanka" name="Nothern Province" pageId="5" pageNumber="105">Nothern Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BEF7278FF92FFAAFEA4FE32FDFCFE0F" box="[382,548,475,501]" pageId="5" pageNumber="105">Jaffna District</collectingMunicipality>
,
<location id="8EEBBED9FF92FFAAFDE9FE32FD6FFE0F" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFDE9FE32FD6FFE0F" box="[563,695,475,501]" country="Sri Lanka" latitude="9.62" longLatPrecision="21" longitude="79.97806" municipality="Jaffna District" name="Mandaitivu" pageId="5" pageNumber="105" stateProvince="Nothern Province">Mandaitivu</location>
,
<geoCoordinate id="EE008EC5FF92FFAAFD1CFE32FC8CFE0F" box="[710,852,475,501]" degrees="09" direction="north" minutes="37" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="30" value="9.625">09°3730N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFC85FE33FC34FE0F" box="[863,1004,474,501]" degrees="79" direction="east" minutes="59" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="50" value="79.997215">79°5950E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFC21FE32FBF4FE0F" box="[1019,1068,475,501]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="105" unit="m" value="4.0">
<elevation id="00190F31FF92FFAAFC21FE32FBF4FE0F" box="[1019,1068,475,501]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="105" unit="m" value="4.0">4 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFBE6FE32FB87FE0F" box="[1084,1119,475,501]" count="1" pageId="5" pageNumber="105" type="female">1♀</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFB44FE32FB1BFE0F" box="[1182,1219,475,501]" count="1" pageId="5" pageNumber="105" type="male">1♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFB09FE32FF0AFDE3" pageId="5" pageNumber="105" value="2016-09-20" valueMax="2016-09-22" valueMin="2016-09-20">
<collectingDate id="EFCE372AFF92FFAAFB09FE32FF0AFDE3" pageId="5" pageNumber="105" value="2016-09-20" valueMax="2016-09-22" valueMin="2016-09-20">2022 September 2016</collectingDate>
</date>
, leg. SP. Benjamin
<emphasis id="B9403410FF92FFAAFE76FE16FE05FDE3" box="[428,477,511,537]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_180, 181);
<specimenCount id="9D32238BFF92FFAAFD37FE16FCCAFDE3" box="[749,786,511,537]" count="4" pageId="5" pageNumber="105" type="male">4♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFCC4FE16FC06FDE3" box="[798,990,511,537]" pageId="5" pageNumber="105" value="2023-10-03">
<collectingDate id="EFCE372AFF92FFAAFCC4FE16FC06FDE3" box="[798,990,511,537]" pageId="5" pageNumber="105" value="2023-10-03">03 October 2023</collectingDate>
</date>
, NG. Dayananda
<emphasis id="B9403410FF92FFAAFB7FFE16FB0EFDE3" box="[1189,1238,511,537]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_224, 225, 226, 227); Chettypulam Road side,
<geoCoordinate id="EE008EC5FF92FFAAFDB1FDCAFD20FDC7" box="[619,760,547,573]" degrees="09" direction="north" minutes="37" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="48" value="9.63">09°3748N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFCD8FDCBFC56FDC7" box="[770,910,546,573]" degrees="79" direction="east" minutes="54" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="45" value="79.9125">79°5445E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFC46FDCAFC13FDC7" box="[924,971,547,573]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="105" unit="m" value="6.0">
<elevation id="00190F31FF92FFAAFC46FDCAFC13FDC7" box="[924,971,547,573]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="105" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFC03FDCAFC24FDC7" box="[985,1020,547,573]" count="1" pageId="5" pageNumber="105" type="female">1♀</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFBEDFDCAFB84FDC7" box="[1079,1116,547,573]" count="3" pageId="5" pageNumber="105" type="male">3♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFBB0FDCAFAF5FDC7" box="[1130,1325,547,573]" pageId="5" pageNumber="105" value="2023-10-04">
<collectingDate id="EFCE372AFF92FFAAFBB0FDCAFAF5FDC7" box="[1130,1325,547,573]" pageId="5" pageNumber="105" value="2023-10-04">04 October 2023</collectingDate>
</date>
, leg. NG. Dayananda
<emphasis id="B9403410FF92FFAAFEC1FDAEFE94FD9B" box="[283,332,583,609]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_220,221,222,223); Allaipiddy,
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<geoCoordinate id="EE008EC5FF92FFAAFC13FDAFFB8EFD9B" box="[969,1110,582,609]" degrees="79" direction="east" minutes="58" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="41" value="79.97806">79°5841E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFBBBFDAEFB55FD9B" box="[1121,1165,583,609]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="105" unit="m" value="6.0">
<elevation id="00190F31FF92FFAAFBBBFDAEFB55FD9B" box="[1121,1165,583,609]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="105" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFB42FDAEFB63FD9B" box="[1176,1211,583,609]" count="3" pageId="5" pageNumber="105" type="female">3♀</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFB2AFDAEFACDFD9B" box="[1264,1301,583,609]" count="3" pageId="5" pageNumber="105" type="male">3♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFAFAFDAEFF0AFD7F" pageId="5" pageNumber="105" value="2023-10-04">
<collectingDate id="EFCE372AFF92FFAAFAFAFDAEFF0AFD7F" pageId="5" pageNumber="105" value="2023-10-04">04 October 2023</collectingDate>
</date>
, leg. NG.
<collectorName id="26C18DD4FF92FFAAFE9EFD82FE1AFD7F" box="[324,450,619,645]" pageId="5" pageNumber="105">Dayananda</collectorName>
<emphasis id="B9403410FF92FFAAFE13FD82FE22FD7F" box="[457,506,619,645]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS ZOD_214, 215, 219, 216, 217, 218)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF92FFAAFF1DFD66FAE0FD53" box="[199,1336,655,681]" pageId="5" pageNumber="105" type="etymology">
<paragraph id="8B8BE802FF92FFAAFF1DFD66FAE0FD53" blockId="5.[151,1437,150,2013]" box="[199,1336,655,681]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DFD66FE96FD53" bold="true" box="[199,334,655,681]" pageId="5" pageNumber="105">Etymology.</emphasis>
The new species is named after the chocolate MilkyWay. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF92FFAAFF1DFD5BFCDFFC17" pageId="5" pageNumber="105" type="diagnosis">
<paragraph id="8B8BE802FF92FFAAFF1DFD5BFCDFFC17" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DFD5BFE9AFD36" bold="true" box="[199,322,690,716]" pageId="5" pageNumber="105">Diagnosis.</emphasis>
<taxonomicName id="4C349381FF92FFAAFE92FD5AFDF6FD37" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[328,558,691,717]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="milkyway" status="sp. nov.">
<emphasis id="B9403410FF92FFAAFE92FD5AFDF6FD37" box="[328,558,691,717]" italics="true" pageId="5" pageNumber="105">Mallinella milkyway</emphasis>
</taxonomicName>
<emphasis id="B9403410FF92FFAAFDEEFD5AFD47FD37" bold="true" box="[564,671,691,717]" pageId="5" pageNumber="105">
<taxonomicNameLabel id="A273896BFF92FFAAFDEEFD5AFD55FD37" box="[564,653,691,717]" pageId="5" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
s
</emphasis>
hare features such as U-shaped epigynal plate, tear-shaped PVS, and tremendously lengthened insemination ducts with the members of the redimita-group. Within the redimita-group, males of
<taxonomicName id="4C349381FF92FFAAFF21FD12FE52FCEF" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[251,394,763,789]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="milkyway" status="sp. nov.">
<emphasis id="B9403410FF92FFAAFF21FD12FE52FCEF" box="[251,394,763,789]" italics="true" pageId="5" pageNumber="105">M. milkyway</emphasis>
</taxonomicName>
<emphasis id="B9403410FF92FFAAFE55FD12FE30FCEF" bold="true" box="[399,488,763,789]" pageId="5" pageNumber="105">
<taxonomicNameLabel id="A273896BFF92FFAAFE55FD12FE30FCEF" box="[399,488,763,789]" pageId="5" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are most similar to the males of
<taxonomicName id="4C349381FF92FFAAFC97FD12FC17FCEF" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[845,975,763,789]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="redimita">
<emphasis id="B9403410FF92FFAAFC97FD12FC17FCEF" box="[845,975,763,789]" italics="true" pageId="5" pageNumber="105">M. redimita</emphasis>
</taxonomicName>
as both share flange-like RTA, bifurcated embolus, sharp apical ridge directed posteromesad and a triangular apico-retrolateral fold with a blunt apex but can be separated from it by the rami of the embolus with blunt apices (sharply pointed in
<taxonomicName id="4C349381FF92FFAAFBBEFCAAFB31FCA7" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[1124,1257,835,861]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="redimita">
<emphasis id="B9403410FF92FFAAFBBEFCAAFB31FCA7" box="[1124,1257,835,861]" italics="true" pageId="5" pageNumber="105">M. redimita</emphasis>
</taxonomicName>
) and lanceolate dorsal scutum (cf.
<bibRefCitation id="EFA595F3FF92FFAAFEBDFC8EFD5AFC7B" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[359,642,871,897]" pageId="5" pageNumber="105" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF92FFAAFDD2FC8EFD98FC7B" box="[520,576,871,897]" italics="true" pageId="5" pageNumber="105">et al.</emphasis>
2012
</bibRefCitation>
: figs. 706707 and
<figureCitation id="130FF487FF92FFAAFC87FC8EFBD5FC7B" box="[861,1037,871,897]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs. 2A, 2FH</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFBC2FC8EFBE4FC7B" box="[1048,1084,871,897]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">8B</figureCitation>
). Females are also most similar to the females of
<taxonomicName id="4C349381FF92FFAAFE86FC62FE3EFC5F" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[348,486,907,933]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="redimata">
<emphasis id="B9403410FF92FFAAFE86FC62FE3EFC5F" box="[348,486,907,933]" italics="true" pageId="5" pageNumber="105">M. redimata</emphasis>
</taxonomicName>
as both share U-shaped epigynal plate, and elongated insemination ducts but can be separated from it by globular spermathecae (truncated spermathecae in
<taxonomicName id="4C349381FF92FFAAFC60FC46FBE6FC33" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[954,1086,943,969]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="redimita">
<emphasis id="B9403410FF92FFAAFC60FC46FBE6FC33" box="[954,1086,943,969]" italics="true" pageId="5" pageNumber="105">M. redimita</emphasis>
</taxonomicName>
) (cf.
<bibRefCitation id="EFA595F3FF92FFAAFBA6FC46FA40FC33" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[1148,1432,943,969]" pageId="5" pageNumber="105" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF92FFAAFAC7FC46FA8DFC33" box="[1309,1365,943,969]" italics="true" pageId="5" pageNumber="105">et al.</emphasis>
2012
</bibRefCitation>
: figs. 694699, 701-703 and
<figureCitation id="130FF487FF92FFAAFE09FC3AFD8FFC17" box="[467,599,979,1005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs. 2CD</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFDB8FC3AFD70FC17" box="[610,680,979,1005]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="5" pageNumber="105">7BG</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFD69FC3AFD23FC17" box="[691,763,979,1005]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="5" pageNumber="105">9CD</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF92FFA9FF1DFC1FFB7CFE77" lastPageId="6" lastPageNumber="106" pageId="5" pageNumber="105" type="description">
<paragraph id="8B8BE802FF92FFAAFF1DFC1FFE70F9F3" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DFC1FFE81FBEA" bold="true" box="[199,345,1014,1040]" pageId="5" pageNumber="105">Description.</emphasis>
<emphasis id="B9403410FF92FFAAFE84FC1EFE40FBEB" box="[350,408,1015,1041]" italics="true" pageId="5" pageNumber="105">Male</emphasis>
in alcohol (
<typeStatus id="548F56A0FF92FFAAFDCCFC1EFDA5FBEB" box="[534,637,1015,1041]" pageId="5" pageNumber="105" type="holotype">holotype</typeStatus>
;
<figureCitation id="130FF487FF92FFAAFD5DFC1EFD06FBEB" box="[647,734,1015,1041]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2A</figureCitation>
). Body length 3.96; carapace 2.25 long, 1.76 wide; opisthosoma 1.71 long, 1.36 wide. Habitus as in
<figureCitation id="130FF487FF92FFAAFDF0FBF2FD5EFBCF" box="[554,646,1051,1077]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2A</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFD49FBF2FD3EFBCF" box="[659,742,1051,1077]" captionStart="FIGURE 11" captionStartId="16.[151,250,771,795]" captionTargetBox="[151,1436,181,747]" captionTargetId="figure-17@16.[151,1436,181,747]" captionTargetPageId="16" captionText="FIGURE 11. Photographs of live Mallinella milkyway sp. nov. from Sri Lanka, paratype male from Arisimale Beach Side (IFS_ZOD_207). Photographs by SP. Benjamin" figureDoi="http://doi.org/10.5281/zenodo.14734687" httpUri="https://zenodo.org/record/14734687/files/figure.png" pageId="5" pageNumber="105">11AB</figureCitation>
. Carapace ovoid, granulate, brown, slightly darker anteriorly; fovea red-brown (
<figureCitation id="130FF487FF92FFAAFEBEFBD6FE18FBA3" box="[356,448,1087,1113]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2A</figureCitation>
). Clypeus light brown, 0.42 high. Eye sizes and inter-distances: AME 0.13, ALE 0.12, PME 0.12, PLE 0.12; AMEAME 0.08, AMEPME 0.15, AMEALE 0.12, ALEALE 0.55, PMEPME 0.12, PMEPLE 0.20, PLEPLE 0.68, ALEPLE 0.06. MOA 0.41 long, front width 0.34, back width 0.38. Chilum unipartite, trapezoid. Chelicerae 0.78 long, brown, without teeth. Endites yellow-brown. Labium triangular, yellow-brown, 0.23 long, 0.38 wide. Sternum brown, with sparse black setae, 1.01 long, 0.93 wide. Lateral margins of sternum with small semi-circular pits in front of each coxa of third and fourth pair of legs (
<figureCitation id="130FF487FF92FFAAFB50FB1AFB3CFAF7" box="[1162,1252,1267,1293]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="5" pageNumber="105">Fig. 6G</figureCitation>
). Legs generally yellowish, except for brown femora, proximal tubercle on anterior femora (
<figureCitation id="130FF487FF92FFAAFC21FAFEFB8FFACB" box="[1019,1111,1303,1329]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="5" pageNumber="105">Fig. 6B</figureCitation>
). Measurements of palp and legs: pedipalp (right) 2.78 (0.97 + 0.45 + 0.47 + 0.89) I 6.70 (1.60 + 0.50 + 1.75 + 1.50+ 1.35), II 5.90 (1.45 + 0.45 + 1.30 + 1.40 + 1.30), III 6.25 (1.40 + 0.35 + 1.25 + 1.35 + 1.90), IV 8.17 (1.95 + 0.50+ 1.85 + 2.15 + 1.72). Leg formula: 4132. Opisthosoma oval, covered with black short setae, with lanceolate dorsal scutum. Dorsum of opisthosoma dark sepia. Pattern on dorsum of opisthosoma represented by a paired of large anterior patches followed by irregular arrangement of pale spots. Posterior ventral spines short, tear-shaped (
<figureCitation id="130FF487FF92FFAAFB6DFA22FAC9FA1F" box="[1207,1297,1483,1509]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2E</figureCitation>
). Spinnerets yellow-brown (
<figureCitation id="130FF487FF92FFAAFE9EFA06FE43F9F3" box="[324,411,1519,1545]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF92FFAAFF1DF9FAFAB2F91B" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DF9FAFF24F9D7" box="[199,252,1555,1581]" italics="true" pageId="5" pageNumber="105">Palp</emphasis>
(
<figureCitation id="130FF487FF92FFAAFED7F9FAFE52F9D7" box="[269,394,1555,1581]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2FH</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFE4DF9FAFE62F9D7" box="[407,442,1555,1581]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">8B</figureCitation>
). RTA flange-like in retrolateral view, broad at base, gradually tapered towards sharply pointed apex (
<figureCitation id="130FF487FF92FFAAFEEDF9DEFE4DF9AB" box="[311,405,1591,1617]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2G</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFE78F9DEFE1DF9AB" box="[418,453,1591,1617]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">8B</figureCitation>
). VTA subtriangular blunt apex. Tegular spine minute (
<figureCitation id="130FF487FF92FFAAFBEFF9DEFB55F9AB" box="[1077,1165,1591,1617]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">Fig. 8B</figureCitation>
). Cymbial fold shallow, less than half the length of cymbium (
<figureCitation id="130FF487FF92FFAAFD91F9B2FD7FF98F" box="[587,679,1627,1653]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2G</figureCitation>
). TA with sharply pointed apical ridge directed posteromesad with triangular apicoprotrolateral process having blunt apex, directed mesad (
<figureCitation id="130FF487FF92FFAAFC75F996FBDCF963" box="[943,1028,1663,1689]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">Fig. 8B</figureCitation>
). Embolic base aligned in retrolateral direction, anterior membranous part broad, triangular (
<figureCitation id="130FF487FF92FFAAFCDEF94AFCBBF947" box="[772,867,1699,1725]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2G</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFCAAF94AFC4BF947" box="[880,915,1699,1725]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">8B</figureCitation>
). Embolus blade-like, broadest medially, subterminally divided, both rami slender, with blunt tips (
<figureCitation id="130FF487FF92FFAAFD2FF92EFC95F91B" box="[757,845,1735,1761]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">Fig. 8B</figureCitation>
). Conductor beak-shaped, sharply pointed apex.
</paragraph>
<paragraph id="8B8BE802FF92FFA9FF1DF902FC75FF2F" blockId="5.[151,1437,150,2013]" lastBlockId="6.[151,1437,150,469]" lastPageId="6" lastPageNumber="106" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DF902FEC3F8FF" box="[199,283,1771,1797]" italics="true" pageId="5" pageNumber="105">Female</emphasis>
(
<typeStatus id="548F56A0FF92FFAAFEF2F902FE56F8FF" box="[296,398,1771,1797]" pageId="5" pageNumber="105" type="paratype">paratype</typeStatus>
;
<figureCitation id="130FF487FF92FFAAFE4DF902FE35F8FF" box="[407,493,1771,1797]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2B</figureCitation>
). Body length 4.98; carapace 2.24 long, 1.54 wide; opisthosoma 2.74 long, 1.91 wide. Habitus and details as in male except for the following (
<figureCitation id="130FF487FF92FFAAFCDAF8E6FC83F8D3" box="[768,859,1807,1833]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2B</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFCBCF8E6FC63F8D3" box="[870,955,1807,1833]" captionStart="FIGURE 10" captionStartId="15.[151,250,1923,1947]" captionTargetBox="[151,1435,181,1900]" captionTargetId="figure-25@15.[151,1435,181,1900]" captionTargetPageId="15" captionText="FIGURE 10. Photographs of live Mallinella spp. described from Sri Lanka. AB Mallinella milkyway sp. nov., paratype female from Arisimale Beach Side (IFS_ZOD_205). CE Mallinella dhanahami sp. nov., paratype female from Gilimale Forest Reserve (IFS_ZOD_235). Photographs by SP. Benjamin." figureDoi="http://doi.org/10.5281/zenodo.14734683" httpUri="https://zenodo.org/record/14734683/files/figure.png" pageId="5" pageNumber="105">10AB</figureCitation>
). Carapace with smooth tegument. Clypeus 0.50 high. Eye sizes and inter-distances: AME 0.10, ALE 0.09, PME 0.09, PLE 0.09; AMEAME 0.08, AMEPME 0.14, AMEALE 0.12, ALEALE 0.41, PMEPME 0.12, PMEPLE 0.16, PLEPLE 0.58, ALEPLE 0.05. MOA 0.36 long, front width 0.27, back width 0.31. Chelicerae 0.75 long. Labium 0.25 long, 0.39 wide. Sternum with shallow semi-circular pits, 0.94 long, 0.89 wide Legs with proximal tubercle on anterior femora. Measurements of palp and legs: palp (right) 1.94 (0.64 + 0.36 + 0.39 + 0.55), I 4.85 (1.25 + 0.50 + 1.10 + 1.15 + 0.85), II 4.75 (1.15 + 0.45 + 1.05 + 1.15 + 0.95), III 5.00 (1.25 + 0.45+ 1.00 + 1.35 + 0.95), IV 6.25 (1.50 + 0.55 + 1.80 + 1.25 + 1.15). Leg formula: 4132. Dorsum of opisthosoma without scutum (
<figureCitation id="130FF487FF91FFA9FC92FF52FC78FF2F" box="[840,928,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="6" pageNumber="106">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF91FFA9FF1DFF09FB5FFE9D" blockId="6.[151,1437,150,469]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFF09FEE9FF00" box="[199,305,224,250]" italics="true" pageId="6" pageNumber="106">Genitalia</emphasis>
(
<figureCitation id="130FF487FF91FFA9FEE4FF09FE62FF00" box="[318,442,224,250]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="6" pageNumber="106">Figs 2CD</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FE19FF09FDCBFF00" box="[451,531,224,250]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">7B, 7G</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FDC7FF09FDBDFF00" box="[541,613,224,250]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="6" pageNumber="106">9CD</figureCitation>
). Epigynal plate; narrow, U-shaped, prominent anterior median depression, weakly sclerotized. Lateral borders retracted (
<figureCitation id="130FF487FF91FFA9FD70FEEDFCD2FEE4" box="[682,778,260,286]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="6" pageNumber="106">Figs 2C</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FCC3FEECFCE4FEE5" box="[793,828,261,287]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">7B</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FC91FEEDFCB7FEE4" box="[843,879,260,286]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="6" pageNumber="106">9C</figureCitation>
). Insemination ducts elongated, slightly curving inwardly (
<figureCitation id="130FF487FF91FFA9FED1FEC0FEB2FEB9" box="[267,362,297,323]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">Figs 7G</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FEA2FEC0FE46FEB9" box="[376,414,297,323]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="6" pageNumber="106">9D</figureCitation>
). Spermathecae globular in shape, slightly widened distally. Insemination ducts short and straight extend posteriorly from the copulatory openings. Fertilization ducts as in
<figureCitation id="130FF487FF91FFA9FBF0FEA4FB5BFE9D" box="[1066,1155,333,359]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">Fig. 7G</figureCitation>
.
</paragraph>
<paragraph id="8B8BE802FF91FFA9FF1DFE9BFB7CFE77" blockId="6.[151,1437,150,469]" box="[199,1188,370,397]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFE9BFEE7FE76" bold="true" box="[199,319,370,396]" pageId="6" pageNumber="106">Variation.</emphasis>
Male (n=22): body length 3.904.02. Female (n=21): body length 5.005.19.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF91FFA9FF1DFE7FFCB7FE2F" pageId="6" pageNumber="106" type="distribution">
<paragraph id="8B8BE802FF91FFA9FF1DFE7FFCB7FE2F" blockId="6.[151,1437,150,469]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFE7FFEB9FE4A" bold="true" box="[199,353,406,432]" pageId="6" pageNumber="106">Distribution.</emphasis>
<collectingCountry id="F323A892FF91FFA9FEBCFE7FFE0BFE4A" box="[358,467,406,432]" name="Sri Lanka" pageId="6" pageNumber="106">Sri Lanka</collectingCountry>
(Matale District, Jaffna District, Anuradhapura District, Trincomalee District, Puttalam District, Kandy District) (
<figureCitation id="130FF487FF91FFA9FE6DFE52FDD3FE2F" box="[439,523,443,469]" captionStart="FIGURE 12" captionStartId="17.[151,250,1133,1157]" captionText="FIGURE 12. Distribution map of known Mallinella species of Sri Lanka." pageId="6" pageNumber="106">Fig. 12</figureCitation>
). Possibly also present in
<collectingCountry id="F323A892FF91FFA9FCF4FE52FCB3FE2F" box="[814,875,443,469]" name="India" pageId="6" pageNumber="106">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>