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<mods:title id="CD88E5879E8D2DF8C14F798A32705137">First record of a hydrozoan (Cnidaria, Hydrozoa) growing on a sigalionid scaleworm (Annelida, Sigalionidae)</mods:title>
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<mods:namePart id="8188C7931566FCBFF3CE06BB4F169C4F">Cruz-Gómez, Christopher</mods:namePart>
<mods:affiliation id="38BE26DEF27DE154360143B63108C1BF">Departamento de Sistemática y Ecología Acuática, El Colegio de la Frontera Sur, Chetumal, Quintana Roo, 77014, Mexico</mods:affiliation>
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<mods:namePart id="F1B4433FA1A71B23710F243F17F4DC43">Mendoza-Becerril, María A.</mods:namePart>
<mods:affiliation id="43243F44B52839805EC530324D1DCF46">Departamento de Sistemática y Ecología Acuática, El Colegio de la Frontera Sur, Chetumal, Quintana Roo, 77014, Mexico</mods:affiliation>
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<taxonomicName id="4C204D67FF9AFF972FCAB8DDFE12FA0B" ID-CoL="68KZJ" authority="Wright, 1859" authorityName="Wright" authorityYear="1859" box="[151,505,1446,1473]" class="Hydrozoa" family="Bougainvilliidae" genus="Bimeria" kingdom="Animalia" order="Anthoathecata" pageId="1" pageNumber="388" phylum="Cnidaria" rank="species" species="vestita">
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<emphasis id="B954EAF6FF9AFF972FCAB8DDFEA6FA0A" bold="true" box="[151,333,1447,1473]" italics="true" pageId="1" pageNumber="388">Bimeria vestita</emphasis>
Wright, 1859
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<figureCitation id="131B2A61FF9AFF972FCAB8B1FEF1FA2E" box="[151,282,1483,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Figs. 1AH</figureCitation>
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<emphasis id="B954EAF6FF9AFF972FCABB68FE68F9E7" bold="true" box="[151,387,1554,1580]" pageId="1" pageNumber="388">Material examined.</emphasis>
<materialsCitation id="3B483CB9FF9AFF972ED0BB68FF44F99A" collectionCode="LACM-AHF" country="Panama" elevation="45" latitude="7.9652777" location="Secas Islands" longLatPrecision="21" longitude="-82.00083" municipality="Gulf" pageId="1" pageNumber="388" specimenCode="LACM-AHF 1576" specimenCount="1" stateProvince="Chiriqui">
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<specimenCode id="DB869E9FFF9AFF972ED0BB68FD8CF9E7" box="[397,615,1554,1580]" collectionCode="LACM-AHF" pageId="1" pageNumber="388">LACM-AHF 1576</specimenCode>
a
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,
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,
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of
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,
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,
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,
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<elevation id="000DD1D7FF9AFF972ADDBB68FF44F99A" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="388" unit="m" value="45.0">45 m</elevation>
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,
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<collectionCode id="ED31AE21FF9AFF972FE1BB4DFF20F99A" box="[188,203,1591,1617]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="388">R</collectionCode>
/
<collectionCode id="ED31AE21FF9AFF972F8BBB4DFF0EF99A" box="[214,229,1591,1617]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="388" type="Museum">V</collectionCode>
Velero III
</emphasis>
, Sta. 250-34, February 22, 1934, on the 5
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right anterior scale of
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<emphasis id="B954EAF6FF9AFF972B62BB4DFACBF99A" box="[1087,1312,1591,1617]" italics="true" pageId="1" pageNumber="388">Pelogenia fimbriata</emphasis>
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, the fertile colony was
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above the dorsal line of the worm and extended to the lateral sides of the scale about
<quantity id="4CD89B01FF9AFF972A60BB20FA73F9BE" box="[1341,1432,1626,1653]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="388" unit="mm" value="0.5">0.5 mm</quantity>
; proximally, over posterior scales and the dorsal midline, distally, right above the chaetae.
</paragraph>
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<paragraph id="8B9F36E4FF9AFF972F9ABBD8FE29F85E" blockId="1.[151,1436,1554,2013]" pageId="1" pageNumber="388">
<emphasis id="B954EAF6FF9AFF972F9ABBD8FEB2F977" bold="true" box="[199,345,1698,1724]" pageId="1" pageNumber="388">Description.</emphasis>
Mature colony small (
<quantity id="4CD89B01FF9AFF972D0CBBD8FD4CF976" box="[593,679,1698,1725]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="388" unit="mm" value="1.5">1.5 mm</quantity>
high), stolonal, monosiphonic, arising from creeping tubular stolons (
<figureCitation id="131B2A61FF9AFF972FC2BBBDFEC9F92A" box="[159,290,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Figs. 1AE</figureCitation>
). Bilayered exoskeleton formed by perisarc and exosarc. Perisarc continuous from hydrorhiza to tentacle base unwrinkled at the base of hydrocaulus. Exosarc thin, thinner over hydranth, tentacle base, and hypostome, forming a pseudohydrotheca (
<figureCitation id="131B2A61FF9AFF972EBABA75FD60F8E2" box="[487,651,1807,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Figs. 1D, F, H</figureCitation>
). Hydranth vasiform, merging with short pedicel. Hypostome conical, 1012 tentacles in two very close whorls (
<figureCitation id="131B2A61FF9AFF972D20BA49FD38F886" box="[637,723,1843,1869]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Fig. 1F</figureCitation>
). Gonophore fixed sporosac on hydrorhiza, pedicellate, ovoid, without radial canals, enveloped in the thin exoskeleton (
<figureCitation id="131B2A61FF9AFF972C79BA2DFC90F8BA" box="[804,891,1879,1905]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Fig. 1E</figureCitation>
). Nematocysts of tentacles: microbasic eurytele and desmoneme (
<figureCitation id="131B2A61FF9AFF972E01BA01FE5EF85E" box="[348,437,1915,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[286,1301,179,1848]" captionTargetId="figure-23@2.[286,1301,179,1848]" captionTargetPageId="2" captionText="FIGURE 1. Epibiosis of Bimeria vestita Wright, 1859 on Pelogenia fimbriata (Hartman, 1939) (LACM-AHF Poly 1576a). A.Anterior end of P. fimbriata, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates B. vestita attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca." pageId="1" pageNumber="388">Fig. 1G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A656FFF9AFF972F9ABAE4FB35F816" pageId="1" pageNumber="388" type="distribution">
<paragraph id="8B9F36E4FF9AFF972F9ABAE4FB35F816" blockId="1.[151,1436,1554,2013]" pageId="1" pageNumber="388">
<emphasis id="B954EAF6FF9AFF972F9ABAE4FE8AF873" bold="true" box="[199,353,1950,1976]" pageId="1" pageNumber="388">Distribution.</emphasis>
Circumglobal in temperate and tropical waters (
<bibRefCitation id="EFB14B15FF9AFF972CC8BAE5FBC3F872" author="Calder, D. R." box="[917,1064,1951,1977]" pageId="1" pageNumber="388" pagination="1 - 72" refId="ref3033" refString="Calder, D. R. (2013) Some shallow-water hydroids (Cnidaria: Hydrozoa) from the central east coast of Florida, USA. Zootaxa, 3648 (1), 1 - 72. https: // doi. org / 10.11646 / zootaxa. 3648.1.1" type="journal article" year="2013">Calder 2013</bibRefCitation>
). Records in the Eastern Pacific Ocean are restricted to
<collectingCountry id="F3377674FF9AFF972EC6BAB9FE1AF816" box="[411,497,1987,2013]" name="Mexico" pageId="1" pageNumber="388">Mexico</collectingCountry>
and
<collectingCountry id="F3377674FF9AFF972D7ABAB9FD6BF816" box="[551,640,1987,2013]" name="Panama" pageId="1" pageNumber="388">Panama</collectingCountry>
(
<bibRefCitation id="EFB14B15FF9AFF972DD3BAB9FCCFF816" author="Fraser, C. M." box="[654,804,1987,2013]" pageId="1" pageNumber="388" pagination="1 - 105" refId="ref3705" refString="Fraser, C. M. (1938 a) Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pacific Expeditions, 4, 1 - 105." type="journal article" year="1938">Fraser 1938a</bibRefCitation>
,
<bibRefCitation id="EFB14B15FF9AFF972C72BAB9FC92F816" author="Fraser, C. M." box="[815,889,1987,2013]" pageId="1" pageNumber="388" pagination="129 - 153" refId="ref3735" refString="Fraser, C. M. (1938 b) Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pacific Expeditions, 4, 129 - 153." type="journal article" year="1938">1938b</bibRefCitation>
;
<bibRefCitation id="EFB14B15FF9AFF972CD9BAB9FB38F816" author="Mendoza-Becerril, M. A. &amp; Estrada-Gonzalez, M. C. &amp; Mazariegos-Villarreal, A. &amp; Restrepo-Avedano, L. &amp; Villar-Beltran, R. D. &amp; Aguero, J. &amp; Cunha, A. F." box="[900,1235,1987,2013]" pageId="1" pageNumber="388" pagination="1 - 37" refId="ref4477" refString="Mendoza-Becerril, M. A., Estrada-Gonzalez, M. C., Mazariegos-Villarreal, A., Restrepo-Avedano, L., Villar-Beltran, R. D., Aguero, J. &amp; Cunha, A. F. (2020) Taxonomy and diversity of Hydrozoa (Cnidaria, Medusozoa) of La Paz Bay, Gulf of California. Zootaxa, 4808 (1), 1 - 37. https: // doi. org / 10.11646 / zootaxa. 4808.1.1" type="journal article" year="2020">
Mendoza-Becerril
<emphasis id="B954EAF6FF9AFF972B0ABAB9FB64F816" box="[1111,1167,1987,2013]" italics="true" pageId="1" pageNumber="388">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5F666CFF99FF942FCABA28FC36F831" pageId="2" pageNumber="389" startId="2.[151,250,1874,1898]" targetBox="[286,1301,179,1848]" targetPageId="2" targetType="figure">
<paragraph id="8B9F36E4FF99FF942FCABA28FC36F831" blockId="2.[151,1437,1874,2043]" pageId="2" pageNumber="389">
<emphasis id="B954EAF6FF99FF942FCABA28FA76F8A1" bold="true" box="[151,1437,1874,1899]" pageId="2" pageNumber="389">
FIGURE 1. Epibiosis of
<taxonomicName id="4C204D67FF99FF942EC1BA28FD20F8A1" authority="Wright, 1859" authorityName="Wright" authorityYear="1859" box="[412,715,1874,1898]" class="Hydrozoa" family="Bougainvilliidae" genus="Bimeria" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="389" phylum="Cnidaria" rank="species" species="vestita">
<emphasis id="B954EAF6FF99FF942EC1BA28FDD3F8A1" bold="true" box="[412,568,1874,1898]" italics="true" pageId="2" pageNumber="389">Bimeria vestita</emphasis>
Wright, 1859
</taxonomicName>
on
<taxonomicName id="4C204D67FF99FF942DACBA28FA7CF8A1" authority="(Hartman, 1939) (LACM-AHF Poly 1576 a)" authorityName="LACM-AHF Poly" authorityYear="1576" baseAuthorityName="Hartman" baseAuthorityYear="1939" box="[753,1431,1874,1899]" class="Polychaeta" family="Sigalionidae" genus="Pelogenia" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="389" phylum="Annelida" rank="species" species="fimbriata">
<emphasis id="B954EAF6FF99FF942DACBA28FC2BF8A1" bold="true" box="[753,960,1874,1898]" italics="true" pageId="2" pageNumber="389">Pelogenia fimbriata</emphasis>
(
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) (LACM-AHF Poly 1576a)
</taxonomicName>
.
</emphasis>
A. Anterior end of
<taxonomicName id="4C204D67FF99FF942E0CBA0DFE2DF845" authorityName="LACM-AHF Poly" authorityYear="1576" baseAuthorityName="Hartman" baseAuthorityYear="1939" box="[337,454,1910,1934]" class="Polychaeta" family="Sigalionidae" genus="Pelogenia" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="389" phylum="Annelida" rank="species" species="fimbriata">
<emphasis id="B954EAF6FF99FF942E0CBA0DFE2DF845" box="[337,454,1910,1934]" italics="true" pageId="2" pageNumber="389">P. fimbriata</emphasis>
</taxonomicName>
, dorsal view, the asterisk indicates the prostomium, and the arrowhead indicates
<taxonomicName id="4C204D67FF99FF942BBABA0DFAA8F845" authorityName="Wright" authorityYear="1859" box="[1255,1347,1911,1934]" class="Hydrozoa" family="Bougainvilliidae" genus="Bimeria" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="389" phylum="Cnidaria" rank="species" species="vestita">
<emphasis id="B954EAF6FF99FF942BBABA0DFAA8F845" box="[1255,1347,1911,1934]" italics="true" pageId="2" pageNumber="389">B. vestita</emphasis>
</taxonomicName>
attached to the scale. B. Hydroid colony on the surface of the scale still attached to the dorsum, side view. C. Anterior scale removed. D. Close-up of hydrants. E. Close-up of the colony, arrowhead indicates gonophore. F. Detail of tentacles. G. Nematocyst microbasic eurytele. H. SEM of a hydrant, arrowhead indicates pseudohydrotheca.
</paragraph>
</caption>
<subSubSection id="C33A656FFF98FF952F9ABDEDFE75FED6" pageId="3" pageNumber="390" type="discussion">
<paragraph id="8B9F36E4FF98FF952F9ABDEDFE75FED6" blockId="3.[151,1436,151,285]" pageId="3" pageNumber="390">
<emphasis id="B954EAF6FF98FF952F9ABDEDFED0FF7A" bold="true" box="[199,315,151,177]" pageId="3" pageNumber="390">Remarks.</emphasis>
See
<bibRefCitation id="EFB14B15FF98FF952E2FBDEDFDE4FF7A" author="Calder, D. R." box="[370,527,151,177]" pageId="3" pageNumber="390" pagination="1 - 107" refId="ref2956" refString="Calder, D. R. (1988) Shallow-water hydroids of Bermuda: the athecatae. Royal Ontario Museum Life Sciences Contributions, 148, 1 - 107." type="journal article" year="1988">Calder (1988)</bibRefCitation>
and
<bibRefCitation id="EFB14B15FF98FF952D1BBDEDFCE3FF7A" author="Schuchert, P." box="[582,776,151,177]" pageId="3" pageNumber="390" pagination="195 - 396" refId="ref5294" refString="Schuchert, P. (2007) The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Filifera Part 2. Le Revue Suisse de Zoologie, 114, 195 - 396. https: // doi. org / 10.5962 / bhl. part. 80395" type="journal article" year="2007">Schuchert (2007)</bibRefCitation>
for taxonomic details and synonyms of this species. There are currently five species of the genus
<taxonomicName id="4C204D67FF98FF952D0BBDC1FC99FF1E" authority="(WoRMS 2024)" baseAuthorityName="WoRMS" baseAuthorityYear="2024" box="[598,882,187,213]" class="Hydrozoa" family="Bougainvilliidae" genus="Bimeria" kingdom="Animalia" order="Anthoathecata" pageId="3" pageNumber="390" phylum="Cnidaria" rank="genus">
<emphasis id="B954EAF6FF98FF952D0BBDC1FD5BFF1F" box="[598,688,187,212]" italics="true" pageId="3" pageNumber="390">Bimeria</emphasis>
(WoRMS 2024)
</taxonomicName>
. The species
<taxonomicName id="4C204D67FF98FF952B50BDC1FB9CFF1F" authorityName="Wright" authorityYear="1859" box="[1037,1143,187,212]" class="Hydrozoa" family="Bougainvilliidae" genus="Bimeria" kingdom="Animalia" order="Anthoathecata" pageId="3" pageNumber="390" phylum="Cnidaria" rank="species" species="vestita">
<emphasis id="B954EAF6FF98FF952B50BDC1FB9CFF1F" box="[1037,1143,187,212]" italics="true" pageId="3" pageNumber="390">B. vestita</emphasis>
</taxonomicName>
is distinguished from the remainder of the genus species by the proximal part of its tentacles being ensheathed with an exoskeleton (
<bibRefCitation id="EFB14B15FF98FF952A0DBDA5FF39FED6" author="Calder, D. R." pageId="3" pageNumber="390" pagination="1 - 107" refId="ref2956" refString="Calder, D. R. (1988) Shallow-water hydroids of Bermuda: the athecatae. Royal Ontario Museum Life Sciences Contributions, 148, 1 - 107." type="journal article" year="1988">Calder 1988</bibRefCitation>
;
<bibRefCitation id="EFB14B15FF98FF952F83BC79FE79FED6" author="Schuchert, P." box="[222,402,259,285]" pageId="3" pageNumber="390" pagination="195 - 396" refId="ref5294" refString="Schuchert, P. (2007) The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Filifera Part 2. Le Revue Suisse de Zoologie, 114, 195 - 396. https: // doi. org / 10.5962 / bhl. part. 80395" type="journal article" year="2007">Schuchert 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,351 @@
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<heading id="D0C881D5FFE8857AFF26F485FD69F941" bold="true" box="[151,668,1759,1786]" fontSize="11" level="1" pageId="6" pageNumber="537" reason="1">
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<emphasis id="B94BEAABFFE8857AFF26F485FD69F941" bold="true" box="[151,668,1759,1786]" pageId="6" pageNumber="537">
<emphasis id="B94BEAABFFE8857AFF26F485FEDAF942" bold="true" box="[151,303,1759,1786]" italics="true" pageId="6" pageNumber="537">Scydmaenus</emphasis>
(
<emphasis id="B94BEAABFFE8857AFEF1F4BAFEBEF942" bold="true" box="[320,331,1760,1786]" italics="true" pageId="6" pageNumber="537">s</emphasis>
.
<emphasis id="B94BEAABFFE8857AFEEBF4BAFE8CF942" bold="true" box="[346,377,1760,1786]" italics="true" pageId="6" pageNumber="537">str</emphasis>
.)
<emphasis id="B94BEAABFFE8857AFE20F485FE1EF941" bold="true" box="[401,491,1759,1785]" italics="true" pageId="6" pageNumber="537">vestitus</emphasis>
(
<bibRefCitation id="EFAE4B48FFE8857AFE4AF485FD66F941" author="Sharp, D." box="[507,659,1759,1785]" pageId="6" pageNumber="537" pagination="105 - 130" refId="ref5511" refString="Sharp, D. (1874) The Pselaphidae and Scydmaenidae of Japan. Transactions of the Entomological Society of London, 1, 105 - 130. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00160. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
)
</emphasis>
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</paragraph>
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Chinese common name:
<emphasis id="B94BEAABFFE8857AFE1CF55EFDE8F8A3" bold="true" box="[429,541,1796,1819]" pageId="6" pageNumber="537">AE毛ẅm</emphasis>
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(
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)
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<emphasis id="B94BEAABFFE8857AFF26F537FEBEF83C" box="[151,331,1901,1924]" italics="true" pageId="6" pageNumber="537">Eumicrus vestitus</emphasis>
<bibRefCitation id="EFAE4B48FFE8857AFEE0F536FDF7F83C" author="Sharp, D." box="[337,514,1900,1924]" pageId="6" pageNumber="537" pagination="105 - 130" refId="ref5511" refString="Sharp, D. (1874) The Pselaphidae and Scydmaenidae of Japan. Transactions of the Entomological Society of London, 1, 105 - 130. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00160. x" type="journal article" year="1874">Sharp, 1874: 126</bibRefCitation>
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;
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locality:
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<emphasis id="B94BEAABFFE8857AFF26F5D1FEE1F81B" box="[151,276,1931,1955]" italics="true" pageId="6" pageNumber="537">Scydmaenus</emphasis>
(
<emphasis id="B94BEAABFFE8857AFE95F5D1FEDBF81A" box="[292,302,1931,1954]" italics="true" pageId="6" pageNumber="537">s</emphasis>
.
<emphasis id="B94BEAABFFE8857AFE8DF5D1FEA2F81A" box="[316,343,1931,1954]" italics="true" pageId="6" pageNumber="537">str</emphasis>
.)
<emphasis id="B94BEAABFFE8857AFEDCF5D1FE4CF81A" box="[365,441,1931,1954]" italics="true" pageId="6" pageNumber="537">vestitus</emphasis>
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<bibRefCitation id="EFAE4B48FFE8857AFE79F5D1FD8AF81B" author="Reitter, E." box="[456,639,1931,1955]" pageId="6" pageNumber="537" pagination="265 - 268" refId="ref5471" refString="Reitter, E. (1913) H. Sauter's Formosa-Ausbeute. Scydmaenidae. Entomologische Mitteilungen, 2 (9), 265 - 268. https: // doi. org / 10.5962 / bhl. part. 14989" type="journal article" year="1913">Reitter 1913: 267</bibRefCitation>
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;
<treatmentCitation id="0A9E10A8FFE8857AFD3BF5D1FCC3F81B" author="Franz, H." box="[650,822,1930,1955]" page="109" pageId="6" pageNumber="537" year="1985">
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;
<treatmentCitation id="0A9E10A8FFE8857AFCF3F5D1FB8AF81B" author="Schulke, M. &amp; Smetana, A." box="[834,1151,1930,1955]" page="898" pageId="6" pageNumber="537" year="2015">
<bibRefCitation id="EFAE4B48FFE8857AFCF3F5D1FB8AF81B" author="Schulke, M. &amp; Smetana, A." box="[834,1151,1930,1955]" pageId="6" pageNumber="537" pagination="304 - 1134" refId="ref5580" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae Latreille, 1802, In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2 / 1 - 2. Revised and Updated Edition. Hydrophiloidea-Staphylinoidea. Brill, Leiden, pp. 304 - 1134." type="book chapter" year="2015">Schülke &amp; Smetana 2015: 898</bibRefCitation>
</treatmentCitation>
; OKeefe &amp; Li, J.-K. 1999: 160.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8036B9FFE8857AFF26F592FAD0F858" blockId="6.[151,1437,1900,2016]" box="[151,1317,1992,2016]" pageId="6" pageNumber="537">
<treatmentCitationGroup id="AB2F1197FFE8857AFF26F592FAD0F858" box="[151,1317,1992,2016]" pageId="6" pageNumber="537">
<treatmentCitation id="0A9E10A8FFE8857AFF26F592FDB8F858" author="Franz, H." box="[151,589,1992,2016]" page="24" pageId="6" pageNumber="537" year="1988">
<taxonomicName id="4C3F4D3AFFE8857AFF26F592FDB8F858" ID-CoL="4W43T" authority="Franz, 1988: 24" authorityName="Franz" authorityPageNumber="24" authorityYear="1988" box="[151,589,1992,2016]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="kunmingensis">
<emphasis id="B94BEAABFFE8857AFF26F592FE50F858" box="[151,421,1992,2016]" italics="true" pageId="6" pageNumber="537">Scydmaenus kunmingensis</emphasis>
<bibRefCitation id="EFAE4B48FFE8857AFE1AF592FDB8F858" author="Franz, H." box="[427,589,1992,2016]" pageId="6" pageNumber="537" pagination="23 - 25" refId="ref4934" refString="Franz, H. (1988) Zwei neue Scydmaenus - Arten aus sud-China (Coleoptera, Scydmaenidae). Elytron, 2, 23 - 25." type="journal article" year="1988">Franz, 1988: 24</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0A9E10A8FFE8857AFDE9F592FCA3F858" author="O'Keefe, S. T. &amp; Li, J. - K." box="[600,854,1992,2016]" page="160" pageId="6" pageNumber="537" year="1999">
<bibRefCitation id="EFAE4B48FFE8857AFDE9F592FCA3F858" author="O'Keefe, S. T. &amp; Li, J. - K." box="[600,854,1992,2016]" pageId="6" pageNumber="537" pagination="150 - 162" refId="ref5354" refString="O'Keefe, S. T. &amp; Li, J. - K. (1999) Review of the Scydmaenidae (Coleoptera: Scydmaenidae) of eastern Asia, with particular reference to Scydmaenus, and description of the first scydmaenid from Hainan Island, China. Journal of New York Entomological Society, 106 (4), 150 - 162. [1998]" type="journal article" year="1999">OKeefe &amp; Li 1999: 160</bibRefCitation>
</treatmentCitation>
.
<typeStatus id="5484881BFFE8857AFCEEF592FC64F858" box="[863,913,1992,2016]" pageId="6" pageNumber="537">Type</typeStatus>
locality:
<collectingRegion id="49FBF85BFFE8857AFC41F592FBB6F867" box="[1008,1091,1992,2015]" country="China" name="Yunnan" pageId="6" pageNumber="537">Yünnan</collectingRegion>
, Kunming.
<emphasis id="B94BEAABFFE8857AFB75F592FAD0F858" bold="true" box="[1220,1317,1992,2016]" pageId="6" pageNumber="537">
<taxonomicNameLabel id="A27857D0FFE8857AFB75F592FAD0F858" box="[1220,1317,1992,2016]" pageId="6" pageNumber="537">Syn. nov.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8036B9FFE9857BFF26F2CDFEAEFF75" blockId="7.[151,1437,150,205]" pageId="7" pageNumber="538">
<treatmentCitationGroup id="AB2F1197FFE9857BFF26F2CDFEAEFF75" pageId="7" pageNumber="538">
<taxonomicName id="4C3F4D3AFFE9857BFF26F2CDFE07FF17" ID-CoL="4W43T" authorityName="Franz" authorityYear="1988" box="[151,498,151,175]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="kunmingensis">
<emphasis id="B94BEAABFFE9857BFF26F2CDFEE1FF17" box="[151,276,151,175]" italics="true" pageId="7" pageNumber="538">Scydmaenus</emphasis>
(
<emphasis id="B94BEAABFFE9857BFE92F2CDFED8FF16" box="[291,301,151,174]" italics="true" pageId="7" pageNumber="538">s</emphasis>
.
<emphasis id="B94BEAABFFE9857BFE89F2CDFEA6FF16" box="[312,339,151,174]" italics="true" pageId="7" pageNumber="538">str</emphasis>
.)
<emphasis id="B94BEAABFFE9857BFED9F2CDFE07FF17" box="[360,498,151,175]" italics="true" pageId="7" pageNumber="538">kunmingensis</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A9E10A8FFE9857BFE4FF2CDFD79FF17" author="Vit, S." box="[510,652,151,175]" page="226" pageId="7" pageNumber="538" year="2004">
<bibRefCitation id="EFAE4B48FFE9857BFE4FF2CDFD79FF17" author="Vit, S." box="[510,652,151,175]" pageId="7" pageNumber="538" pagination="223 - 228" refId="ref5709" refString="Vit, S. (2004) Scydmaenidae [except Cyrtoscydmini, Cephenniini]. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 206, 223 - 228." type="journal article" year="2004">Vít 2004: 226</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9E10A8FFE9857BFD27F2CDFC39FF17" author="Schulke, M. &amp; Smetana, A." box="[662,972,150,175]" page="897" pageId="7" pageNumber="538" year="2015">
<bibRefCitation id="EFAE4B48FFE9857BFD27F2CDFC39FF17" author="Schulke, M. &amp; Smetana, A." box="[662,972,150,175]" pageId="7" pageNumber="538" pagination="304 - 1134" refId="ref5580" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae Latreille, 1802, In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2 / 1 - 2. Revised and Updated Edition. Hydrophiloidea-Staphylinoidea. Brill, Leiden, pp. 304 - 1134." type="book chapter" year="2015">Schülke &amp; Smetana 2015: 897</bibRefCitation>
</treatmentCitation>
; Li, L.-Z.
<emphasis id="B94BEAABFFE9857BFB83F2CDFBABFF17" box="[1074,1118,151,175]" italics="true" pageId="7" pageNumber="538">et al</emphasis>
. 2019: 342;
<treatmentCitation id="0A9E10A8FFE9857BFB6AF2CDFF05FF75" author="Jaloszynski, P." page="358" pageId="7" pageNumber="538" year="2021">
<bibRefCitation id="EFAE4B48FFE9857BFB6AF2CDFF05FF75" author="Jaloszynski, P." pageId="7" pageNumber="538" pagination="354 - 368" refId="ref5186" refString="Jaloszynski, P. (2021 b) Review of Scydmaenus species of continental China described by Herbert Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4952 (2), 354 - 368. https: // doi. org / 10.11646 / zootaxa. 4952.2.8" type="journal article" year="2021">Jałoszyński 2021b: 358</bibRefCitation>
</treatmentCitation>
.
<emphasis id="B94BEAABFFE9857BFF4BF2EFFEAEFF75" bold="true" box="[250,347,181,205]" pageId="7" pageNumber="538">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF406631FFE9857BFF26F429FA6CF916" pageId="7" pageNumber="538" startId="7.[151,250,1651,1675]" targetBox="[156,1436,230,1625]" targetPageId="7" targetType="figure">
<paragraph id="8B8036B9FFE9857BFF26F429FA6CF916" blockId="7.[151,1437,1651,1711]" pageId="7" pageNumber="538">
<emphasis id="B94BEAABFFE9857BFF26F429FEE6F933" bold="true" box="[151,275,1651,1675]" pageId="7" pageNumber="538">FIGURE 4.</emphasis>
<taxonomicName id="4C3F4D3AFFE9857BFEA8F429FDC2F933" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[281,567,1651,1675]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="B94BEAABFFE9857BFEA8F429FE63F933" box="[281,406,1651,1675]" italics="true" pageId="7" pageNumber="538">Scydmaenus</emphasis>
(
<emphasis id="B94BEAABFFE9857BFE14F42EFE5AF933" box="[421,431,1652,1675]" italics="true" pageId="7" pageNumber="538">s</emphasis>
.
<emphasis id="B94BEAABFFE9857BFE0DF42EFE22F933" box="[444,471,1652,1675]" italics="true" pageId="7" pageNumber="538">str</emphasis>
.)
<emphasis id="B94BEAABFFE9857BFE5AF42EFDC2F933" box="[491,567,1652,1675]" italics="true" pageId="7" pageNumber="538">vestitus</emphasis>
</taxonomicName>
(AE. Male. F. Female).
<emphasis id="B94BEAABFFE9857BFC88F429FCA4F933" bold="true" box="[825,849,1651,1675]" pageId="7" pageNumber="538">A.</emphasis>
Dorsal habitus.
<emphasis id="B94BEAABFFE9857BFC46F429FBFBF933" bold="true" box="[1015,1038,1651,1675]" pageId="7" pageNumber="538">B.</emphasis>
Meso- and metaventrite.
<emphasis id="B94BEAABFFE9857BFAA3F429FADFF933" bold="true" box="[1298,1322,1651,1675]" pageId="7" pageNumber="538">C.</emphasis>
Mesotibia.
<emphasis id="B94BEAABFFE9857BFF26F4CDFF38F917" bold="true" box="[151,205,1687,1711]" pageId="7" pageNumber="538">D, E.</emphasis>
Aedeagus, dorsal (D) and lateral (E).
<emphasis id="B94BEAABFFE9857BFDE1F4CDFD96F917" bold="true" box="[592,611,1687,1711]" pageId="7" pageNumber="538">F.</emphasis>
Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in CE; 0.05 mm in F.
</paragraph>
</caption>
<subSubSection id="C3256532FFE9857BFF26F55BFE63F8DB" pageId="7" pageNumber="538" type="materials_examined">
<paragraph id="8B8036B9FFE9857BFF26F55BFE63F8DB" blockId="7.[151,1437,1793,2036]" pageId="7" pageNumber="538">
<emphasis id="B94BEAABFFE9857BFF26F55BFE8AF8A3" bold="true" box="[151,383,1793,1819]" pageId="7" pageNumber="538">Material examined</emphasis>
(4 exx.).
<materialsCitation id="3B573CE4FFE9857BFE4FF55BFE67F8DB" collectingDate="2022-07-13" collectionCode="SNUC" collectorName="Wei" country="China" county="Zhaotong City" elevation="1950" latitude="28.24986" location="Xiaoyanfang Station" longLatPrecision="1" longitude="103.97566" municipality="Wumengshan Natural Reserve" pageId="7" pageNumber="538" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Yunnan">
<specimenCount id="9D39FD30FFE9857BFE4FF55BFDB0F8A4" box="[510,581,1793,1820]" count="2" pageId="7" pageNumber="538" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D39FD30FFE9857BFDE6F55BFD6BF8A4" box="[599,670,1793,1820]" count="2" pageId="7" pageNumber="538" type="female">2 ♀♀</specimenCount>
:
<collectingCountry id="F3287629FFE9857BFD04F55BFCF4F8A3" box="[693,769,1793,1819]" name="China" pageId="7" pageNumber="538">China</collectingCountry>
:
<collectingRegion id="49FBF85BFFE9857BFCA1F558FC9CF8A4" box="[784,873,1794,1820]" country="China" name="Yunnan" pageId="7" pageNumber="538">Yunnan</collectingRegion>
,
<collectingCounty id="62E14E35FFE9857BFCCBF55BFBD6F8A3" box="[890,1059,1793,1819]" pageId="7" pageNumber="538">Zhaotong City</collectingCounty>
,
<collectingMunicipality id="6BE4ACC3FFE9857BFB85F55BFA6DF8A4" box="[1076,1432,1793,1820]" pageId="7" pageNumber="538">Wumengshan Natural Reserve</collectingMunicipality>
,
<location id="8EE06062FFE9857BFF26F57FFE74F887" LSID="urn:lsid:plazi:treatment:039687AFFFE88574FF26F485FB34FC1D:8EE06062FFE9857BFF26F57FFE74F887" box="[151,385,1829,1855]" country="China" county="Zhaotong City" latitude="28.24986" longLatPrecision="1" longitude="103.97566" municipality="Wumengshan Natural Reserve" name="Xiaoyanfang Station" pageId="7" pageNumber="538" stateProvince="Yunnan">Xiaoyanfang Station</location>
,
<geoCoordinate id="EE0B507EFFE9857BFE3CF57FFDD2F887" box="[397,551,1829,1855]" degrees="28.249860" direction="north" orientation="latitude" pageId="7" pageNumber="538" precision="1" value="28.24986">28.249860°N</geoCoordinate>
,
<geoCoordinate id="EE0B507EFFE9857BFD83F57FFD23F8F8" box="[562,726,1829,1856]" degrees="103.975665" direction="east" orientation="longitude" pageId="7" pageNumber="538" precision="1" value="103.97566">103.975665°E</geoCoordinate>
,
<quantity id="4CC79B5CFFE9857BFD53F57FFC8AF8F8" box="[738,895,1829,1856]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.06" metricValueMin="1.8399999999999999" pageId="7" pageNumber="538" unit="m" value="1950.0" valueMax="2060.0" valueMin="1840.0">
<elevation id="0012D18AFFE9857BFD53F57FFC8AF8F8" box="[738,895,1829,1856]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.06" metricValueMin="1.8399999999999999" pageId="7" pageNumber="538" unit="m" value="1950.0" valueMax="2060.0" valueMin="1840.0">18402060 m</elevation>
</quantity>
,
<date id="FF811079FFE9857BFC3AF57FFBFBF887" box="[907,1038,1829,1855]" pageId="7" pageNumber="538" value="2022-07-13">
<collectingDate id="EFC5E991FFE9857BFC3AF57FFBFBF887" box="[907,1038,1829,1855]" pageId="7" pageNumber="538" value="2022-07-13">13.vii.2022</collectingDate>
</date>
,
<collectorName id="26CA536FFFE9857BFBA8F57FFBB0F887" box="[1049,1093,1829,1855]" pageId="7" pageNumber="538">Wei</collectorName>
leg.,
<emphasis id="B94BEAABFFE9857BFB35F57CFED6F8D9" bold="true" pageId="7" pageNumber="538">云南DZdz乌AE山小岩坊 ṄM国Dzẍ</emphasis>
(
<collectionCode id="ED2EAE7CFFE9857BFE88F513FE7FF8DB" box="[313,394,1865,1891]" pageId="7" pageNumber="538">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFE98574FF76F537FD13FE4D" lastPageId="8" lastPageNumber="539" pageId="7" pageNumber="538" type="diagnosis">
<paragraph id="8B8036B9FFE98574FF76F537FE30FEDD" blockId="7.[151,1437,1793,2036]" lastBlockId="8.[151,1437,151,933]" lastPageId="8" lastPageNumber="539" pageId="7" pageNumber="538">
<emphasis id="B94BEAABFFE9857BFF76F537FEB7F83F" bold="true" box="[199,322,1901,1927]" pageId="7" pageNumber="538">Diagnosis.</emphasis>
<emphasis id="B94BEAABFFE9857BFEF9F537FE7CF83F" box="[328,393,1901,1927]" italics="true" pageId="7" pageNumber="538">Male.</emphasis>
Body (
<figureCitation id="13042A3CFFE9857BFE6BF537FDC4F83F" box="[474,561,1901,1927]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (AE. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in CE; 0.05 mm in F." pageId="7" pageNumber="538">Fig. 4A</figureCitation>
) length over
<quantity id="4CC79B5CFFE9857BFD72F537FCDFF830" box="[707,810,1901,1928]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="538" unit="mm" value="2.5">2.50 mm</quantity>
. Head and pronotum finely and rather sparsely punctate. Head slightly transverse, suboval, tempora much longer than eyes; antenna with antennomere 5 much longer than 4 and 6, 8 transverse, 911 greatly enlarged and forming distinct club, 11 subconical. Pronotum bell-shaped, with four large antebasal impressions. Each elytron with round apex, well exposing pygidium. Metaventrite (
<figureCitation id="13042A3CFFE9857BFAB3F583FAAFF84B" box="[1282,1370,2009,2035]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (AE. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in CE; 0.05 mm in F." pageId="7" pageNumber="538">Fig. 4B</figureCitation>
) with large impression at middle, sides of impression forming short, setose ridges posteriorly. Protarsomere 1 broadened, tarsomeres successively narrowing from 1 to 5; mesotrochanter angulate on ventral margin, mesotibia (
<figureCitation id="13042A3CFFE68574FA8AF2E1FA61FF6D" box="[1339,1428,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (AE. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in CE; 0.05 mm in F." pageId="8" pageNumber="539">Fig. 4C</figureCitation>
) with long apical spine; metatibia angulate at inner-apical margin. Aedeagus (
<figureCitation id="13042A3CFFE68574FC4AF285FB86FF41" box="[1019,1139,223,249]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (AE. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in CE; 0.05 mm in F." pageId="8" pageNumber="539">Fig. 4D, E</figureCitation>
) elongate, dorso-ventrally symmetric, laterally greatly curved at middle; apical portion of endophallus armature composed of two sclerites pointed at apices, one markedly long and the other much shorter; area above foramen and below apex of median with short and dense setae.
</paragraph>
<paragraph id="8B8036B9FFE68574FF76F335FD13FE4D" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="539">
<emphasis id="B94BEAABFFE68574FF76F335FEEEFE31" box="[199,283,367,393]" italics="true" pageId="8" pageNumber="539">Female</emphasis>
. Body length slightly shorter than male. Antenna shorter. Metaventrite moderately convex at middle, lacking impression or ridges. Protarsus slender; ventral margin of mesotrochanter rounded; mesotibia lacking apical spine; metatibia lacking apical angulation. Spermatheca (
<figureCitation id="13042A3CFFE68574FCBEF3EDFC97FE69" box="[783,866,439,465]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1675]" captionTargetBox="[156,1436,230,1625]" captionTargetId="figure-45@7.[151,1436,230,1628]" captionTargetPageId="7" captionText="FIGURE 4. Scydmaenus (s. str.) vestitus (AE. Male. F. Female). A. Dorsal habitus. B. Meso- and metaventrite. C. Mesotibia. D, E. Aedeagus, dorsal (D) and lateral (E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A, B; 0.2 mm in CE; 0.05 mm in F." pageId="8" pageNumber="539">Fig. 4F</figureCitation>
) spherical, dome-shaped, with funnel-like structure located on same axis as insertion of accessory gland.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFE68574FF76F3A4FCD3FD11" pageId="8" pageNumber="539" type="description">
<paragraph id="8B8036B9FFE68574FF76F3A4FCD3FD11" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="539">
<emphasis id="B94BEAABFFE68574FF76F3A4FEACFDA0" bold="true" box="[199,345,510,536]" pageId="8" pageNumber="539">Description.</emphasis>
See
<bibRefCitation id="EFAE4B48FFE68574FE2BF3A5FD5AFDA1" author="Jaloszynski, P." box="[410,687,510,537]" pageId="8" pageNumber="539" pagination="354 - 368" refId="ref5186" refString="Jaloszynski, P. (2021 b) Review of Scydmaenus species of continental China described by Herbert Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4952 (2), 354 - 368. https: // doi. org / 10.11646 / zootaxa. 4952.2.8" type="journal article" year="2021">Jałoszyński 2021b: 358</bibRefCitation>
. Measurements for Wumengshan population: male body length
<quantity id="4CC79B5CFFE68574FF26F079FECAFD85" box="[151,319,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="2.69" metricValueMax="2.72" metricValueMin="2.66" pageId="8" pageNumber="539" unit="mm" value="2.69" valueMax="2.72" valueMin="2.66">2.662.72 mm</quantity>
, length/width of head 0.490.50/0.56
<quantity id="4CC79B5CFFE68574FD4BF078FC97FD85" box="[762,866,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="539" unit="mm" value="0.5">0.50 mm</quantity>
, pronotum 0.730.78/
<quantity id="4CC79B5CFFE68574FBEEF078FAF3FD85" box="[1119,1286,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="6.699999999999999" metricValueMax="6.8999999999999995" metricValueMin="6.5" pageId="8" pageNumber="539" unit="mm" value="0.6699999999999999" valueMax="0.69" valueMin="0.65">0.650.69 mm</quantity>
, elytra 1.33 1.40/
<quantity id="4CC79B5CFFE68574FF60F01DFE8DFDD9" box="[209,376,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.105" metricValueMax="1.13" metricValueMin="1.08" pageId="8" pageNumber="539" unit="mm" value="1.105" valueMax="1.13" valueMin="1.08">1.081.13 mm</quantity>
, length of antenna
<quantity id="4CC79B5CFFE68574FDE8F01CFCF7FDD9" box="[601,770,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" metricValueMax="1.35" metricValueMin="1.29" pageId="8" pageNumber="539" unit="mm" value="1.32" valueMax="1.35" valueMin="1.29">1.291.35 mm</quantity>
; female body length
<quantity id="4CC79B5CFFE68574FC49F01DFB55FDD9" box="[1016,1184,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.38" metricValueMin="2.22" pageId="8" pageNumber="539" unit="mm" value="2.3" valueMax="2.38" valueMin="2.22">2.222.38 mm</quantity>
, length/width of head 0.460.48/
<quantity id="4CC79B5CFFE68574FEA1F030FE40FD3D" box="[272,437,618,645]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="8" pageNumber="539" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.450.48 mm</quantity>
, pronotum 0.630.74/
<quantity id="4CC79B5CFFE68574FD01F030FCA3FD3D" box="[688,854,618,645]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.7" metricValueMin="5.6" pageId="8" pageNumber="539" unit="mm" value="0.615" valueMax="0.67" valueMin="0.56">0.560.67 mm</quantity>
, elytra 1.221.27/
<quantity id="4CC79B5CFFE68574FB95F031FB3FFD3D" box="[1060,1226,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.055" metricValueMax="1.07" metricValueMin="1.04" pageId="8" pageNumber="539" unit="mm" value="1.055" valueMax="1.07" valueMin="1.04">1.041.07 mm</quantity>
, length of antenna
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; longest diameter of spermatheca
<quantity id="4CC79B5CFFE68574FD0AF0D4FCD7FD11" box="[699,802,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="539" unit="mm" value="0.15">0.15 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFE68574FF76F0E8FAC5FD75" box="[199,1328,690,717]" pageId="8" pageNumber="539" type="distribution">
<paragraph id="8B8036B9FFE68574FF76F0E8FAC5FD75" blockId="8.[151,1437,151,933]" box="[199,1328,690,717]" pageId="8" pageNumber="539">
<emphasis id="B94BEAABFFE68574FF76F0E8FE94FD74" bold="true" box="[199,353,690,716]" pageId="8" pageNumber="539">Distribution.</emphasis>
<collectingCountry id="F3287629FFE68574FED6F0E9FE52FD75" box="[359,423,691,717]" name="Japan" pageId="8" pageNumber="539">Japan</collectingCountry>
(Honshu, Kyushu, Ryukyu Is.);
<collectingCountry id="F3287629FFE68574FCA5F0E9FC51FD75" box="[788,932,691,717]" name="South Korea" pageId="8" pageNumber="539">South Korea</collectingCountry>
;
<collectingCountry id="F3287629FFE68574FC1EF0E9FC07FD75" box="[943,1010,691,717]" name="China" pageId="8" pageNumber="539">China</collectingCountry>
(
<collectingRegion id="49FBF85BFFE68574FBB0F0E9FB95FD75" box="[1025,1120,691,717]" country="China" name="Sichuan" pageId="8" pageNumber="539">Sichuan</collectingRegion>
,
<collectingRegion id="49FBF85BFFE68574FBDBF0E9FB36FD75" box="[1130,1219,691,717]" country="China" name="Yunnan" pageId="8" pageNumber="539">Yunnan</collectingRegion>
,
<collectingRegion id="49FBF85BFFE68574FB7CF0E9FAD9FD75" box="[1229,1324,691,717]" country="China" name="Taiwan)" pageId="8" pageNumber="539">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFE68574FF76F08DFB34FC1D" pageId="8" pageNumber="539" type="discussion">
<paragraph id="8B8036B9FFE68574FF76F08DFB34FC1D" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="539">
<emphasis id="B94BEAABFFE68574FF76F08DFECEFD49" bold="true" box="[199,315,727,753]" pageId="8" pageNumber="539">Remarks.</emphasis>
This species is readily distinguishable and exhibits a widespread distribution ranging from
<collectingCountry id="F3287629FFE68574FAEDF08DFA69FD49" box="[1372,1436,727,753]" name="Japan" pageId="8" pageNumber="539">Japan</collectingCountry>
through
<collectingCountry id="F3287629FFE68574FF46F0A1FECFFCAD" box="[247,314,763,789]" name="China" pageId="8" pageNumber="539">China</collectingCountry>
(mainland and
<collectingRegion id="49FBF85BFFE68574FE5DF0A1FDBEFCAD" box="[492,587,763,789]" country="China" name="Taiwan)" pageId="8" pageNumber="539">Taiwan)</collectingRegion>
. The proposed synonymy of
<taxonomicName id="4C3F4D3AFFE68574FC27F0A1FBF2FCAC" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[918,1031,763,789]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="B94BEAABFFE68574FC27F0A1FC51FCAD" box="[918,932,763,789]" italics="true" pageId="8" pageNumber="539">S</emphasis>
.
<emphasis id="B94BEAABFFE68574FC02F0A1FBF2FCAC" box="[947,1031,763,788]" italics="true" pageId="8" pageNumber="539">vestitus</emphasis>
</taxonomicName>
with
<taxonomicName id="4C3F4D3AFFE68574FBFBF0A1FB0AFCAD" authorityName="Franz" authorityYear="1988" box="[1098,1279,763,789]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="kunmingensis">
<emphasis id="B94BEAABFFE68574FBFBF0A1FBADFCAD" box="[1098,1112,763,789]" italics="true" pageId="8" pageNumber="539">S</emphasis>
.
<emphasis id="B94BEAABFFE68574FBD6F0A1FB0AFCAD" box="[1127,1279,763,789]" italics="true" pageId="8" pageNumber="539">kunmingensis</emphasis>
</taxonomicName>
is based on a comparison of specimens from
<collectingRegion id="49FBF85BFFE68574FE4CF145FDA6FC81" box="[509,595,799,825]" country="China" name="Yunnan" pageId="8" pageNumber="539">Yunnan</collectingRegion>
with the original descriptions and aedeagal illustrations of both taxa. The large body size, male-specific tibial modifications, metaventrite characteristics, and the structural configuration of the aedeagal endophallus are distinctive and consistent across all known populations. The spermatheca of this species is illustrated for the first time. All individuals were collected together with
<taxonomicName id="4C3F4D3AFFE68574FB87F1D1FB4FFC1D" authorityName="Franz" authorityYear="1988" box="[1078,1210,907,933]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="539" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B94BEAABFFE68574FB87F1D1FBB1FC1D" box="[1078,1092,907,933]" italics="true" pageId="8" pageNumber="539">S</emphasis>
.
<emphasis id="B94BEAABFFE68574FBE3F1D1FB4FFC1D" box="[1106,1210,907,933]" italics="true" pageId="8" pageNumber="539">chinensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,260 @@
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<mods:title id="0258D7DCD2171E744ADE9AFF814BDD1A">A report of the Scydmaeninae species (Coleoptera: Staphylinidae) taken from Wumenshan National Nature Reserve, Yunnan, China</mods:title>
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<heading id="D0C881D5FFEB8579FF26F08CFD6AFD48" bold="true" box="[151,671,726,753]" fontSize="11" level="1" pageId="5" pageNumber="536" reason="1">
<taxonomicName id="4C3F4D3AFFEB8579FF26F08CFD6AFD48" ID-CoL="4W3ZD" authority="Franz, 1988" authorityName="Franz" authorityYear="1988" box="[151,671,726,753]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B94BEAABFFEB8579FF26F08CFD6AFD48" bold="true" box="[151,671,726,753]" pageId="5" pageNumber="536">
<emphasis id="B94BEAABFFEB8579FF26F08CFEDAFD49" bold="true" box="[151,303,726,753]" italics="true" pageId="5" pageNumber="536">Scydmaenus</emphasis>
(
<emphasis id="B94BEAABFFEB8579FEF1F08DFEBEFD49" bold="true" box="[320,331,727,753]" italics="true" pageId="5" pageNumber="536">s</emphasis>
.
<emphasis id="B94BEAABFFEB8579FEEBF08DFE8CFD49" bold="true" box="[346,377,727,753]" italics="true" pageId="5" pageNumber="536">str</emphasis>
.)
<emphasis id="B94BEAABFFEB8579FE20F08CFDF6FD49" bold="true" box="[401,515,726,753]" italics="true" pageId="5" pageNumber="536">chinensis</emphasis>
<bibRefCitation id="EFAE4B48FFEB8579FDBAF08DFD6AFD48" author="Franz, H." box="[523,671,726,753]" pageId="5" pageNumber="536" pagination="23 - 25" refId="ref4934" refString="Franz, H. (1988) Zwei neue Scydmaenus - Arten aus sud-China (Coleoptera, Scydmaenidae). Elytron, 2, 23 - 25." type="journal article" year="1988">Franz, 1988</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8036B9FFEB8579FF26F0A1FDE8FCAA" blockId="5.[151,671,726,825]" box="[151,541,763,789]" pageId="5" pageNumber="536">
Chinese common name:
<emphasis id="B94BEAABFFEB8579FE1CF0A1FDE8FCAA" bold="true" box="[429,541,763,786]" pageId="5" pageNumber="536">中华ẅm</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEB8579FF26F145FF1EFC81" box="[151,235,799,825]" pageId="5" pageNumber="536" type="description">
<paragraph id="8B8036B9FFEB8579FF26F145FF1EFC81" blockId="5.[151,671,726,825]" box="[151,235,799,825]" pageId="5" pageNumber="536">
(
<figureCitation id="13042A3CFFEB8579FF2EF145FF17FC81" box="[159,226,799,825]" captionStart="FIGURE 3" captionStartId="6.[151,250,1617,1641]" captionTargetBox="[154,1433,185,1587]" captionTargetId="figure-20@6.[151,1436,181,1593]" captionTargetPageId="6" captionText="FIGURE 3. Scydmaenus (s. str.) chinensis (AC. Male. D, E. Female). A. Dorsal habitus. B, C. Aedeagus, dorsal (B), and lateral (C). D, E. Spermatheca, dorsal (D) and lateral (E). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D, E." pageId="5" pageNumber="536">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEB8579FF26F139FF01FC00" pageId="5" pageNumber="536" type="reference_group">
<paragraph id="8B8036B9FFEB8579FF26F139FB47FCC3" blockId="5.[151,1437,867,952]" box="[151,1202,867,892]" pageId="5" pageNumber="536">
<treatmentCitationGroup id="AB2F1197FFEB8579FF26F139FB47FCC3" box="[151,1202,867,892]" pageId="5" pageNumber="536">
<treatmentCitation id="0A9E10A8FFEB8579FF26F139FDD7FCC3" author="Franz, H." box="[151,546,867,891]" page="24" pageId="5" pageNumber="536" year="1988">
<taxonomicName id="4C3F4D3AFFEB8579FF26F139FDD7FCC3" ID-CoL="4W3ZD" authority="Franz, 1988: 24" authorityName="Franz" authorityPageNumber="24" authorityYear="1988" box="[151,546,867,891]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B94BEAABFFEB8579FF26F139FE8CFCC3" box="[151,377,867,891]" italics="true" pageId="5" pageNumber="536">Scydmaenus chinensis</emphasis>
<materialsCitation id="3B573CE4FFEB8579FECEF13EFDD7FCC3" box="[383,546,867,891]" collectorName="O'Keefe &amp; Li, J. - K." location="Leshan" pageId="5" pageNumber="536" specimenCount="1" stateProvince="Sichuan">
<bibRefCitation id="EFAE4B48FFEB8579FECEF13EFDD7FCC3" author="Franz, H." box="[383,546,867,891]" pageId="5" pageNumber="536" pagination="23 - 25" refId="ref4934" refString="Franz, H. (1988) Zwei neue Scydmaenus - Arten aus sud-China (Coleoptera, Scydmaenidae). Elytron, 2, 23 - 25." type="journal article" year="1988">Franz, 1988: 24</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
;
<collectorName id="26CA536FFFEB8579FD9DF139FD76FCC3" box="[556,643,867,891]" pageId="5" pageNumber="536">OKeefe</collectorName>
&amp;
<collectorName id="26CA536FFFEB8579FD15F139FD0CFCC3" box="[676,761,867,891]" pageId="5" pageNumber="536">Li, J.-K.</collectorName>
1999: 160.
<typeStatus id="5484881BFFEB8579FCC0F13EFC56FCC4" box="[881,931,868,892]" pageId="5" pageNumber="536">Type</typeStatus>
locality:
<collectingRegion id="49FBF85BFFEB8579FBB2F139FBA2FCC3" box="[1027,1111,867,891]" country="China" name="Sichuan" pageId="5" pageNumber="536">Sichuan</collectingRegion>
,
<location id="8EE06062FFEB8579FBD0F139FB58FCC3" LSID="urn:lsid:plazi:treatment:039687AFFFEB8579FF26F08CFB0AF811:8EE06062FFEB8579FBD0F139FB58FCC3" box="[1121,1197,867,891]" name="Leshan" pageId="5" pageNumber="536" stateProvince="Sichuan">Leshan</location>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8036B9FFEB8579FF26F1D8FF01FC00" blockId="5.[151,1437,867,952]" pageId="5" pageNumber="536">
<treatmentCitationGroup id="AB2F1197FFEB8579FF26F1D8FF01FC00" pageId="5" pageNumber="536">
<taxonomicName id="4C3F4D3AFFEB8579FF26F1D8FE39FC22" ID-CoL="4W3ZD" authorityName="Franz" authorityYear="1988" box="[151,460,898,922]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B94BEAABFFEB8579FF26F1D8FEE1FC22" box="[151,276,898,922]" italics="true" pageId="5" pageNumber="536">Scydmaenus</emphasis>
(
<emphasis id="B94BEAABFFEB8579FE94F1D9FEDAFC22" box="[293,303,899,922]" italics="true" pageId="5" pageNumber="536">s</emphasis>
.
<emphasis id="B94BEAABFFEB8579FE8CF1D9FEADFC22" box="[317,344,899,922]" italics="true" pageId="5" pageNumber="536">str</emphasis>
.)
<emphasis id="B94BEAABFFEB8579FEDFF1D8FE39FC22" box="[366,460,898,922]" italics="true" pageId="5" pageNumber="536">chinensis</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A9E10A8FFEB8579FE6AF1D8FD9BFC22" author="Vit, S." box="[475,622,898,922]" page="226" pageId="5" pageNumber="536" year="2004">
<bibRefCitation id="EFAE4B48FFEB8579FE6AF1D8FD9BFC22" author="Vit, S." box="[475,622,898,922]" pageId="5" pageNumber="536" pagination="223 - 228" refId="ref5709" refString="Vit, S. (2004) Scydmaenidae [except Cyrtoscydmini, Cephenniini]. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 206, 223 - 228." type="journal article" year="2004">Vít 2004: 226</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9E10A8FFEB8579FDCBF1D8FC4CFC22" author="Schulke, M. &amp; Smetana, A." box="[634,953,898,922]" page="897" pageId="5" pageNumber="536" year="2015">
<bibRefCitation id="EFAE4B48FFEB8579FDCBF1D8FC4CFC22" author="Schulke, M. &amp; Smetana, A." box="[634,953,898,922]" pageId="5" pageNumber="536" pagination="304 - 1134" refId="ref5580" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae Latreille, 1802, In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2 / 1 - 2. Revised and Updated Edition. Hydrophiloidea-Staphylinoidea. Brill, Leiden, pp. 304 - 1134." type="book chapter" year="2015">Schülke &amp; Smetana 2015: 897</bibRefCitation>
</treatmentCitation>
; Li, L.-Z.
<emphasis id="B94BEAABFFEB8579FB97F1D9FBA1FC22" box="[1062,1108,898,922]" italics="true" pageId="5" pageNumber="536">et al</emphasis>
. 2019: 342;
<treatmentCitation id="0A9E10A8FFEB8579FB68F1D8FF05FC00" author="Jaloszynski, P." page="355" pageId="5" pageNumber="536" year="2021">
<bibRefCitation id="EFAE4B48FFEB8579FB68F1D8FF05FC00" author="Jaloszynski, P." pageId="5" pageNumber="536" pagination="354 - 368" refId="ref5186" refString="Jaloszynski, P. (2021 b) Review of Scydmaenus species of continental China described by Herbert Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4952 (2), 354 - 368. https: // doi. org / 10.11646 / zootaxa. 4952.2.8" type="journal article" year="2021">Jałoszyński 2021b: 355</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEB8579FF26F1BCFE8FFBF1" pageId="5" pageNumber="536" type="materials_examined">
<paragraph id="8B8036B9FFEB8579FF26F1BCFE8FFBF1" blockId="5.[151,1437,998,1961]" pageId="5" pageNumber="536">
<emphasis id="B94BEAABFFEB8579FF26F1BCFE8BFBB8" bold="true" box="[151,382,998,1024]" pageId="5" pageNumber="536">Material examined</emphasis>
(15 exx.).
<materialsCitation id="3B573CE4FFEB8579FDB9F1BDFE83FBF1" collectingDate="2022-07-13" collectionCode="SNUC" collectorName="Wei" country="China" county="Zhaotong City" elevation="1950" latitude="28.24986" location="Xiaoyanfang Station" longLatPrecision="1" longitude="103.97566" municipality="Wumengshan Natural Reserve" pageId="5" pageNumber="536" specimenCount="15" specimenCount-female="6" specimenCount-male="9" stateProvince="Yunnan">
<specimenCount id="9D39FD30FFEB8579FDB9F1BDFDBBFBB9" box="[520,590,999,1025]" count="9" pageId="5" pageNumber="536" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="9D39FD30FFEB8579FDEEF1BDFD50FBB9" box="[607,677,999,1025]" count="6" pageId="5" pageNumber="536" type="female">6 ♀♀</specimenCount>
:
<collectingCountry id="F3287629FFEB8579FD0AF1BDFCF2FBB9" box="[699,775,999,1025]" name="China" pageId="5" pageNumber="536">China</collectingCountry>
:
<collectingRegion id="49FBF85BFFEB8579FCA4F1BDFC9BFBB9" box="[789,878,999,1025]" country="China" name="Yunnan" pageId="5" pageNumber="536">Yunnan</collectingRegion>
,
<collectingCounty id="62E14E35FFEB8579FCCFF1BDFBD3FBB9" box="[894,1062,999,1025]" pageId="5" pageNumber="536">Zhaotong City</collectingCounty>
,
<collectingMunicipality id="6BE4ACC3FFEB8579FB87F1BDFA6DFBB9" box="[1078,1432,999,1025]" pageId="5" pageNumber="536">Wumengshan Natural Reserve</collectingMunicipality>
,
<location id="8EE06062FFEB8579FF26F651FE74FB9D" LSID="urn:lsid:plazi:treatment:039687AFFFEB8579FF26F08CFB0AF811:8EE06062FFEB8579FF26F651FE74FB9D" box="[151,385,1035,1061]" country="China" county="Zhaotong City" latitude="28.24986" longLatPrecision="1" longitude="103.97566" municipality="Wumengshan Natural Reserve" name="Xiaoyanfang Station" pageId="5" pageNumber="536" stateProvince="Yunnan">Xiaoyanfang Station</location>
,
<geoCoordinate id="EE0B507EFFEB8579FE3CF651FDD2FB9D" box="[397,551,1035,1061]" degrees="28.249860" direction="north" orientation="latitude" pageId="5" pageNumber="536" precision="1" value="28.24986">28.249860°N</geoCoordinate>
,
<geoCoordinate id="EE0B507EFFEB8579FD83F650FD23FB9D" box="[562,726,1034,1061]" degrees="103.975665" direction="east" orientation="longitude" pageId="5" pageNumber="536" precision="1" value="103.97566">103.975665°E</geoCoordinate>
,
<quantity id="4CC79B5CFFEB8579FD53F651FC8AFB9D" box="[738,895,1035,1061]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.06" metricValueMin="1.8399999999999999" pageId="5" pageNumber="536" unit="m" value="1950.0" valueMax="2060.0" valueMin="1840.0">
<elevation id="0012D18AFFEB8579FD53F651FC8AFB9D" box="[738,895,1035,1061]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.06" metricValueMin="1.8399999999999999" pageId="5" pageNumber="536" unit="m" value="1950.0" valueMax="2060.0" valueMin="1840.0">18402060 m</elevation>
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,
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<collectingDate id="EFC5E991FFEB8579FC3AF651FBFBFB9D" box="[907,1038,1035,1061]" pageId="5" pageNumber="536" value="2022-07-13">13.vii.2022</collectingDate>
</date>
,
<collectorName id="26CA536FFFEB8579FBA8F651FBB0FB9D" box="[1049,1093,1035,1061]" pageId="5" pageNumber="536">Wei</collectorName>
leg.,
<emphasis id="B94BEAABFFEB8579FB35F651FEF2FBFE" bold="true" pageId="5" pageNumber="536">云南DZdz乌AE山小岩坊 M国Dzẍ</emphasis>
(
<collectionCode id="ED2EAE7CFFEB8579FEACF675FE9BFBF1" box="[285,366,1071,1097]" pageId="5" pageNumber="536">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEB8579FF76F608FF28FA41" pageId="5" pageNumber="536" type="diagnosis">
<paragraph id="8B8036B9FFEB8579FF76F608FAD8FA35" blockId="5.[151,1437,998,1961]" pageId="5" pageNumber="536">
<emphasis id="B94BEAABFFEB8579FF76F608FEB7FBD4" bold="true" box="[199,322,1106,1132]" pageId="5" pageNumber="536">Diagnosis.</emphasis>
<emphasis id="B94BEAABFFEB8579FEFDF609FE78FBD5" box="[332,397,1107,1133]" italics="true" pageId="5" pageNumber="536">Male.</emphasis>
Body (
<figureCitation id="13042A3CFFEB8579FE57F609FDB7FBD5" box="[486,578,1107,1133]" captionStart="FIGURE 3" captionStartId="6.[151,250,1617,1641]" captionTargetBox="[154,1433,185,1587]" captionTargetId="figure-20@6.[151,1436,181,1593]" captionTargetPageId="6" captionText="FIGURE 3. Scydmaenus (s. str.) chinensis (AC. Male. D, E. Female). A. Dorsal habitus. B, C. Aedeagus, dorsal (B), and lateral (C). D, E. Spermatheca, dorsal (D) and lateral (E). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D, E." pageId="5" pageNumber="536">Fig. 3A</figureCitation>
) length approximately
<quantity id="4CC79B5CFFEB8579FCE0F609FC2BFBD5" box="[849,990,1107,1133]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="5" pageNumber="536" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.81.9 mm</quantity>
. Head and pronotum finely and rather sparsely punctate.Head slightly longer than wide, suboval, tempora much longer than eyes; antenna with antennomere 5 much longer than 4 and 6, 7 and 8 transversely subconical, 911 greatly enlarged and forming distinct club, 11 subconical. Pronotum bell-shaped, with four large antebasal impressions; lateral margins distinctly convergent posteriorly from broadest point to base. Each elytron with round apex, well exposing pygidium. Metaventrite and legs lacking modifications. Protarsomere 1 broadened, tarsomeres successively narrowing from 1 to 5. Aedeagus (
<figureCitation id="13042A3CFFEB8579FADEF75DFF29FAFD" captionStart="FIGURE 3" captionStartId="6.[151,250,1617,1641]" captionTargetBox="[154,1433,185,1587]" captionTargetId="figure-20@6.[151,1436,181,1593]" captionTargetPageId="6" captionText="FIGURE 3. Scydmaenus (s. str.) chinensis (AC. Male. D, E. Female). A. Dorsal habitus. B, C. Aedeagus, dorsal (B), and lateral (C). D, E. Spermatheca, dorsal (D) and lateral (E). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D, E." pageId="5" pageNumber="536">Fig. 3B, C</figureCitation>
) elongate, dorso-ventrally almost symmetric, laterally moderately curved at middle; endophallus armature composed of broad sclerite bifurcate in apical part, forming two lobes, left lobe shorter than right one, curved at apex, right one gradually narrowing toward apex; areas lateral to ostium with rows of long, curved setae.
</paragraph>
<paragraph id="8B8036B9FFEB8579FF76F7CDFF28FA41" blockId="5.[151,1437,998,1961]" pageId="5" pageNumber="536">
<emphasis id="B94BEAABFFEB8579FF76F7CDFEEEFA09" box="[199,283,1431,1457]" italics="true" pageId="5" pageNumber="536">Female</emphasis>
. Body length slightly shorter than male. Antenna as long as male. Protarsus slender. Spermatheca (
<figureCitation id="13042A3CFFEB8579FADEF7CDFF2FFA6D" captionStart="FIGURE 3" captionStartId="6.[151,250,1617,1641]" captionTargetBox="[154,1433,185,1587]" captionTargetId="figure-20@6.[151,1436,181,1593]" captionTargetPageId="6" captionText="FIGURE 3. Scydmaenus (s. str.) chinensis (AC. Male. D, E. Female). A. Dorsal habitus. B, C. Aedeagus, dorsal (B), and lateral (C). D, E. Spermatheca, dorsal (D) and lateral (E). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D, E." pageId="5" pageNumber="536">Fig. 3D, E</figureCitation>
) spherical, with broad opening and long funnel-like structure located on same axis as insertion of accessory gland.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEB8579FF76F458FC71F915" pageId="5" pageNumber="536" type="description">
<paragraph id="8B8036B9FFEB8579FF76F458FC71F915" blockId="5.[151,1437,998,1961]" pageId="5" pageNumber="536">
<emphasis id="B94BEAABFFEB8579FF76F458FEACF9A4" bold="true" box="[199,345,1538,1564]" pageId="5" pageNumber="536">Description.</emphasis>
See
<bibRefCitation id="EFAE4B48FFEB8579FE3FF459FD63F9A5" author="Jaloszynski, P." box="[398,662,1538,1565]" pageId="5" pageNumber="536" pagination="354 - 368" refId="ref5186" refString="Jaloszynski, P. (2021 b) Review of Scydmaenus species of continental China described by Herbert Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4952 (2), 354 - 368. https: // doi. org / 10.11646 / zootaxa. 4952.2.8" type="journal article" year="2021">Jałoszyński 2021b: 355</bibRefCitation>
. Measurements for Wumengshan population: male body length
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, length/width of head 0.380.40/
<quantity id="4CC79B5CFFEB8579FDD8F47CFCF9F9F9" box="[617,780,1574,1601]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="5" pageNumber="536" unit="mm" value="0.365" valueMax="0.38" valueMin="0.35">0.350.38 mm</quantity>
, pronotum 0.570.58/
<quantity id="4CC79B5CFFEB8579FBB0F47CFB51F9F9" box="[1025,1188,1574,1601]" metricMagnitude="-4" metricUnit="m" metricValue="5.05" metricValueMax="5.1" metricValueMin="5.0" pageId="5" pageNumber="536" unit="mm" value="0.505" valueMax="0.51" valueMin="0.5">0.500.51 mm</quantity>
, elytra 0.971.02/
<quantity id="4CC79B5CFFEB8579FADDF47DFF30F9DD" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="5" pageNumber="536" unit="mm" value="0.81">0.81 mm</quantity>
, length of antenna
<quantity id="4CC79B5CFFEB8579FE15F411FDB9F9DD" box="[420,588,1611,1637]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" metricValueMax="9.9" metricValueMin="9.3" pageId="5" pageNumber="536" unit="mm" value="0.96" valueMax="0.99" valueMin="0.93">0.930.99 mm</quantity>
; length of aedeagus
<quantity id="4CC79B5CFFEB8579FC8BF411FC51F9DD" box="[826,932,1611,1637]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="536" unit="mm" value="0.46">0.46 mm</quantity>
; female body length
<quantity id="4CC79B5CFFEB8579FB29F410FAF4F9DD" box="[1176,1281,1610,1637]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="5" pageNumber="536" unit="mm" value="1.85">1.85 mm</quantity>
, length/width of head 0.370.38/
<quantity id="4CC79B5CFFEB8579FEDCF435FDE6F931" box="[365,531,1647,1673]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="3.8" metricValueMin="3.7" pageId="5" pageNumber="536" unit="mm" value="0.375" valueMax="0.38" valueMin="0.37">0.370.38 mm</quantity>
, pronotum 0.560.59/
<quantity id="4CC79B5CFFEB8579FCBEF434FC40F931" box="[783,949,1646,1673]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.2" metricValueMin="5.1" pageId="5" pageNumber="536" unit="mm" value="0.515" valueMax="0.52" valueMin="0.51">0.510.52 mm</quantity>
, elytra 0.951.01/
<quantity id="4CC79B5CFFEB8579FB32F434FADCF931" box="[1155,1321,1646,1673]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" metricValueMax="1.83" metricValueMin="0.81" pageId="5" pageNumber="536" unit="mm" value="1.32" valueMax="1.83" valueMin="0.81">0.811.83 mm</quantity>
, length of antenna
<quantity id="4CC79B5CFFEB8579FF44F4C9FE6EF915" box="[245,411,1683,1709]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" metricValueMax="9.9" metricValueMin="9.3" pageId="5" pageNumber="536" unit="mm" value="0.96" valueMax="0.99" valueMin="0.93">0.930.99 mm</quantity>
; longest diameter of spermatheca
<quantity id="4CC79B5CFFEB8579FCA8F4C9FC75F915" box="[793,896,1683,1709]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="536" unit="mm" value="0.12">0.12 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEB8579FF76F4ECFD0CF969" box="[199,761,1718,1745]" pageId="5" pageNumber="536" type="distribution">
<paragraph id="8B8036B9FFEB8579FF76F4ECFD0CF969" blockId="5.[151,1437,998,1961]" box="[199,761,1718,1745]" pageId="5" pageNumber="536">
<emphasis id="B94BEAABFFEB8579FF76F4ECFE94F968" bold="true" box="[199,353,1718,1744]" pageId="5" pageNumber="536">Distribution.</emphasis>
Southwest
<collectingCountry id="F3287629FFEB8579FE52F4EDFDDFF969" box="[483,554,1719,1745]" name="China" pageId="5" pageNumber="536">China</collectingCountry>
:
<collectingRegion id="49FBF85BFFEB8579FD84F4EDFD67F969" box="[565,658,1719,1745]" country="China" name="Sichuan" pageId="5" pageNumber="536">Sichuan</collectingRegion>
,
<collectingRegion id="49FBF85BFFEB8579FD2DF4EDFD00F969" box="[668,757,1719,1745]" country="China" name="Yunnan" pageId="5" pageNumber="536">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEB8579FF76F481FB0AF811" pageId="5" pageNumber="536" type="discussion">
<paragraph id="8B8036B9FFEB8579FF76F481FE57F885" blockId="5.[151,1437,998,1961]" pageId="5" pageNumber="536">
<emphasis id="B94BEAABFFEB8579FF76F481FECEF94D" bold="true" box="[199,315,1755,1781]" pageId="5" pageNumber="536">Remarks.</emphasis>
This species is easily distinguishable by the aedeagal structures, particularly by the shape of the endophallus armature. The spermatheca is illustrated for the first time.All individuals were simultaneously collected with
<taxonomicName id="4C3F4D3AFFEB8579FF61F579FE6BF885" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[208,414,1827,1853]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="B94BEAABFFEB8579FF61F579FF2BF885" box="[208,222,1827,1853]" italics="true" pageId="5" pageNumber="536">S</emphasis>
.
<emphasis id="B94BEAABFFEB8579FF5DF579FEB5F884" box="[236,320,1827,1852]" italics="true" pageId="5" pageNumber="536">vestitus</emphasis>
(Sharp)
</taxonomicName>
.
</paragraph>
<paragraph id="8B8036B9FFEB8579FF76F51DFB0AF811" blockId="5.[151,1437,998,1961]" pageId="5" pageNumber="536">
In his redescription, Jałoszyński reported the aedeagus measurement as
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, which is comparable to the measurement obtained in the present study (
<quantity id="4CC79B5CFFEB8579FD72F531FCDAF83D" box="[707,815,1899,1925]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="536" unit="mm" value="0.46">0.46 mm</quantity>
). However, the scale bar in the corresponding figures (
<bibRefCitation id="EFAE4B48FFEB8579FF2EF5D5FE86F811" author="Jaloszynski, P." box="[159,371,1935,1961]" pageId="5" pageNumber="536" pagination="354 - 368" refId="ref5186" refString="Jaloszynski, P. (2021 b) Review of Scydmaenus species of continental China described by Herbert Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4952 (2), 354 - 368. https: // doi. org / 10.11646 / zootaxa. 4952.2.8" type="journal article" year="2021">Jałoszyński 2021b</bibRefCitation>
: figs 25) appears to suggest a larger size for the aedeagus, approaching 1.0 mm.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,445 @@
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<mods:title id="F3FF80503B51ECCA85AA2497C451A960">A report of the Scydmaeninae species (Coleoptera: Staphylinidae) taken from Wumenshan National Nature Reserve, Yunnan, China</mods:title>
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<mods:namePart id="A89750FBE23F9D4CC497886A7A3DF4A2">Yin, Zi-Wei</mods:namePart>
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<mods:namePart id="E9E5754311595523A410737E31C49F57">Zhou, De-Yao</mods:namePart>
<mods:affiliation id="332CDE0DE805CF50EA81A8B95086A35E">Jiading Agriculture Technology Extension Service Center, Jiading District, Shanghai 201403, China.</mods:affiliation>
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<mods:typeOfResource id="EAFDC58D0D2605578764E30ED4B282F7">text</mods:typeOfResource>
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<mods:titleInfo id="6293B3ADC8AD6FE5CFEB85D8C88DBFE1">
<mods:title id="294286E0C42D4C809667F2D71BE35F1C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EE06816CB0B35DEF8FD5671481C0E115">
<mods:date id="0C4C7B981F607846C18FDF410C0F0B16">2025</mods:date>
<mods:detail id="F67E6741D66AC77C2C28BDFA77CB2834" type="pubDate">
<mods:number id="F2D8CF61430D62DA678C144B44ED3E34">2025-01-15</mods:number>
</mods:detail>
<mods:detail id="F87514465D94F9084FDFF3DCEF352A40" type="volume">
<mods:number id="74A9C018C2B9D164BA45F6F69F7AB3CB">5569</mods:number>
</mods:detail>
<mods:detail id="F0F65AA6101D7CA91D08A9F95B1C5529" type="issue">
<mods:number id="8A33569BE3AC141F52D3A90AEDBF12CB">3</mods:number>
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<mods:start id="F96B43433A79313D148B71D44FBD52E8">531</mods:start>
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<mods:location id="CED30B52F86ACBD91C851319D2770B1E">
<mods:url id="6932FF19E11F8CFE1FDB06AA4EBC8C83">https://doi.org/10.11646/zootaxa.5569.3.8</mods:url>
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<mods:classification id="7BD0AC53A9CEF400E6A97F3D2A21A689">journal article</mods:classification>
<mods:identifier id="118A8DF24DEE7A0B7102075A46774E1A" type="DOI">10.11646/zootaxa.5569.3.8</mods:identifier>
<mods:identifier id="73441A6F662A4CD41144A5999878BC92" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039687AFFFED8579FF26F2CDFF05FD3D" LSID="urn:lsid:plazi:treatment:039687AFFFED8579FF26F2CDFF05FD3D" httpUri="http://treatment.plazi.org/id/039687AFFFED8579FF26F2CDFF05FD3D" lastPageId="5" lastPageNumber="536" pageId="3" pageNumber="534">
<subSubSection id="C3256532FFED857FFF26F2CDFDA0FF6B" pageId="3" pageNumber="534" type="nomenclature">
<paragraph id="8B8036B9FFED857FFF26F2CDFD58FF09" blockId="3.[151,685,151,249]" box="[151,685,151,178]" pageId="3" pageNumber="534">
<heading id="D0C881D5FFED857FFF26F2CDFD58FF09" bold="true" box="[151,685,151,178]" fontSize="11" level="1" pageId="3" pageNumber="534" reason="1">
<emphasis id="B94BEAABFFED857FFF26F2CDFD58FF09" bold="true" box="[151,685,151,178]" pageId="3" pageNumber="534">
<taxonomicName id="4C3F4D3AFFED857FFF26F2CDFDB6FF09" authority="Yin &amp; Zhou" authorityName="Yin &amp; Zhou" authorityYear="2025" box="[151,579,151,178]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="obesus" status="sp. nov.">
<emphasis id="B94BEAABFFED857FFF26F2CDFE56FF0A" bold="true" box="[151,419,151,178]" italics="true" pageId="3" pageNumber="534">Himaloconnus obesus</emphasis>
Yin &amp; Zhou
</taxonomicName>
,
<taxonomicNameLabel id="A27857D0FFED857FFDE1F2CDFD58FF09" box="[592,685,151,177]" pageId="3" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8036B9FFED857FFF26F2E1FDA0FF6B" blockId="3.[151,685,151,249]" box="[151,597,187,214]" pageId="3" pageNumber="534">
Chinese common name:
<emphasis id="B94BEAABFFED857FFE1CF2E6FDA0FF6B" bold="true" box="[429,597,188,211]" pageId="3" pageNumber="534">Ṁ体喜Ḣẅm</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFED857FFF26F285FF19FF41" box="[151,236,223,249]" pageId="3" pageNumber="534" type="description">
<paragraph id="8B8036B9FFED857FFF26F285FF19FF41" blockId="3.[151,685,151,249]" box="[151,236,223,249]" pageId="3" pageNumber="534">
(
<figureCitation id="13042A3CFFED857FFF2EF285FF16FF41" box="[159,227,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. FI. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in EI." pageId="3" pageNumber="534">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFED857FFF26F37DFE63FE31" pageId="3" pageNumber="534" type="materials_examined">
<paragraph id="8B8036B9FFED857FFF26F37DFE63FE31" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
<emphasis id="B94BEAABFFED857FFF26F37DFEC8FEF9" bold="true" box="[151,317,295,321]" pageId="3" pageNumber="534">Type material</emphasis>
(
<materialsCitation id="3B573CE4FFED857FFEFEF37DFE63FE31" collectingDate="2022-07-20" collectionCode="SNUC" collectorName="Wei" country="China" county="China" elevation="1830" latitude="27.846716" location="Wumengshan Natural Reserve" longLatPrecision="1" longitude="104.29242" municipality="Zhaotong City" pageId="3" pageNumber="534" specimenCount="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="9D39FD30FFED857FFEFEF37DFE73FEF9" box="[335,390,295,321]" count="1" pageId="3" pageNumber="534" type="generic">1 ex.</specimenCount>
).
<emphasis id="B94BEAABFFED857FFE2BF37DFD5AFEF9" bold="true" box="[410,687,295,321]" pageId="3" pageNumber="534">
<typeStatus id="5484881BFFED857FFE2BF37DFDC8FEF9" box="[410,573,295,321]" pageId="3" pageNumber="534" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="F3287629FFED857FFDF9F37DFD5FFEF9" box="[584,682,295,321]" name="China" pageId="3" pageNumber="534">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="9D39FD30FFED857FFD04F372FD38FEFA" box="[693,717,296,322]" count="1" pageId="3" pageNumber="534" type="male"></specimenCount>
:
<collectingCounty id="62E14E35FFED857FFD6CF37DFCDCFEF9" box="[733,809,295,321]" pageId="3" pageNumber="534">China</collectingCounty>
:
<collectingRegion id="49FBF85BFFED857FFC80F372FC7FFEFA" box="[817,906,296,322]" country="China" name="Yunnan" pageId="3" pageNumber="534">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BE4ACC3FFED857FFC25F37DFBC2FEF9" box="[916,1079,295,321]" pageId="3" pageNumber="534">Zhaotong City</collectingMunicipality>
,
<location id="8EE06062FFED857FFBF0F37DFA6DFEFA" LSID="urn:lsid:plazi:treatment:039687AFFFED8579FF26F2CDFF05FD3D:8EE06062FFED857FFBF0F37DFA6DFEFA" box="[1089,1432,295,322]" country="China" county="China" latitude="27.846716" longLatPrecision="1" longitude="104.29242" municipality="Zhaotong City" name="Wumengshan Natural Reserve" pageId="3" pageNumber="534" stateProvince="Yunnan">Wumengshan Natural Reserve</location>
,
<location id="8EE06062FFED857FFF26F311FEE7FEDD" LSID="urn:lsid:plazi:treatment:039687AFFFED8579FF26F2CDFF05FD3D:8EE06062FFED857FFF26F311FEE7FEDD" box="[151,274,331,357]" country="China" county="China" latitude="27.846716" longLatPrecision="1" longitude="104.29242" municipality="Zhaotong City" name="Xiaocaoba" pageId="3" pageNumber="534" stateProvince="Yunnan">Xiaocaoba</location>
,
<location id="8EE06062FFED857FFEACF311FE7BFEDD" LSID="urn:lsid:plazi:treatment:039687AFFFED8579FF26F2CDFF05FD3D:8EE06062FFED857FFEACF311FE7BFEDD" box="[285,398,331,357]" country="China" county="China" latitude="27.846716" longLatPrecision="1" longitude="104.29242" municipality="Zhaotong City" name="Miaoshan" pageId="3" pageNumber="534" stateProvince="Yunnan">Miaoshan</location>
,
<geoCoordinate id="EE0B507EFFED857FFE2BF311FDC1FEDD" box="[410,564,331,357]" degrees="27.846716" direction="north" orientation="latitude" pageId="3" pageNumber="534" precision="1" value="27.846716">27.846716°N</geoCoordinate>
,
<geoCoordinate id="EE0B507EFFED857FFD8EF311FD11FEDD" box="[575,740,331,357]" degrees="104.292419" direction="east" orientation="longitude" pageId="3" pageNumber="534" precision="1" value="104.29242">104.292419°E</geoCoordinate>
,
<quantity id="4CC79B5CFFED857FFD5EF311FCB2FEDE" box="[751,839,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="3" pageNumber="534" unit="m" value="1830.0">
<elevation id="0012D18AFFED857FFD5EF311FCB2FEDE" box="[751,839,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="3" pageNumber="534" unit="m" value="1830.0">1830 m</elevation>
</quantity>
,
<date id="FF811079FFED857FFCE2F311FC23FEDD" box="[851,982,331,357]" pageId="3" pageNumber="534" value="2022-07-20">
<collectingDate id="EFC5E991FFED857FFCE2F311FC23FEDD" box="[851,982,331,357]" pageId="3" pageNumber="534" value="2022-07-20">20.vii.2022</collectingDate>
</date>
,
<collectorName id="26CA536FFFED857FFC50F311FBF8FEDD" box="[993,1037,331,357]" pageId="3" pageNumber="534">Wei</collectorName>
leg.,
<emphasis id="B94BEAABFFED857FFBFDF316FED6FE3F" bold="true" pageId="3" pageNumber="534">云南DZdz乌AE山海小¤坝庙 山M国Dzẍ</emphasis>
(
<collectionCode id="ED2EAE7CFFED857FFE88F335FE7FFE31" box="[313,394,367,393]" pageId="3" pageNumber="534">SNUC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFED857FFF76F3C9FCCAFD11" pageId="3" pageNumber="534" type="diagnosis">
<paragraph id="8B8036B9FFED857FFF76F3C9FCCAFD11" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
<emphasis id="B94BEAABFFED857FFF76F3C9FEB7FE15" bold="true" box="[199,322,403,429]" pageId="3" pageNumber="534">Diagnosis.</emphasis>
<emphasis id="B94BEAABFFED857FFEF8F3C9FE7FFE15" box="[329,394,403,429]" italics="true" pageId="3" pageNumber="534">Male.</emphasis>
Body length approximately
<quantity id="4CC79B5CFFED857FFD7FF3C9FCDDFE16" box="[718,808,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="534" unit="mm" value="1.4">1.4 mm</quantity>
. Head dark brown to nearly black, pronotum and elytra reddish-brown. Dorsal surface of body appearing glabrous, almost devoid of setae, thick bristles present on tempora, posterior portion of vertex, and sides of pronotum and humeri of elytra. Antenna slender, antennomeres 1 and 2 each elongate, length 0.09 and
<quantity id="4CC79B5CFFED857FFE40F3A5FDA2FDA2" box="[497,599,511,538]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="534" unit="mm" value="0.08">0.08 mm</quantity>
, antennomere 9 and 11 of similar width, 10 broader than 9 and 11, together these three antennomeres form distinct club, antennomeres 37 gradually longer. Elytra oval, length/width 1.15. Aedeagus
<quantity id="4CC79B5CFFED857FFEBFF01DFE81FDDA" box="[270,372,583,610]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="534" unit="mm" value="0.26">0.26 mm</quantity>
long, apex of median lobe with acute projection at middle, endophallus armature symmetric, complex in structure; parameres almost reaching apex of median lobe, elongate and slender, each with one long apical and another shorter preapical seta.
<emphasis id="B94BEAABFFED857FFDD6F0D5FD4EFD11" box="[615,699,655,681]" italics="true" pageId="3" pageNumber="534">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFED8579FF76F0E9FC13FDA1" lastPageId="5" lastPageNumber="536" pageId="3" pageNumber="534" type="description">
<paragraph id="8B8036B9FFED857FFF76F0E9FA70FCAD" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
<emphasis id="B94BEAABFFED857FFF76F0E9FEACFD75" bold="true" box="[199,345,691,717]" pageId="3" pageNumber="534">Description.</emphasis>
<emphasis id="B94BEAABFFED857FFED3F0E9FE69FD75" box="[354,412,691,717]" italics="true" pageId="3" pageNumber="534">Male</emphasis>
. Body (
<figureCitation id="13042A3CFFED857FFE4AF0E9FDA3FD75" box="[507,598,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. FI. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in EI." pageId="3" pageNumber="534">Fig. 2A</figureCitation>
) length
<quantity id="4CC79B5CFFED857FFD06F0E9FCD4FD76" box="[695,801,691,718]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="3" pageNumber="534" unit="mm" value="1.38">1.38 mm</quantity>
; head dark brown, pronotum, elytra, antennae and legs reddish-brown, mouthparts and tarsi paler in color. Vertex, sides of head and pronotum and humeri of elytra with long bristles; body dorsum almost impunctate, glabrous, with short and sparse setae found only on disc of elytra.
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F145FF10FBC5" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
Head (
<figureCitation id="13042A3CFFED857FFEA4F145FE61FC81" box="[277,404,799,825]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. FI. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in EI." pageId="3" pageNumber="534">Fig. 2A, B</figureCitation>
) broadest at eyes, length
<quantity id="4CC79B5CFFED857FFD74F145FCDAFC82" box="[709,815,799,826]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="534" unit="mm" value="0.3">0.30 mm</quantity>
, width across eyes
<quantity id="4CC79B5CFFED857FFBA7F145FB74FC82" box="[1046,1153,799,826]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="534" unit="mm" value="0.28">0.28 mm</quantity>
; tempora in dorsal view approximately as long as eyes; vertex and frons confluent, together weakly convex, anterior portion of frons with pair of small, obtuse admesal tubercles; eyes large, strongly prominent laterally, in lateral view suboval, with distinct posteromedian emargination, each eye composed of approximately 20 ommatidia. Vertex with obscure, inconspicuous punctures, almost devoid of setae; tempora and posterior margin of vertex densely covered with long and thick bristles. Submentum with broad and deep central cavity. Hypopharynx with one pair of oval, membranous lateral lobes. Antenna slender, length
<quantity id="4CC79B5CFFED857FFD89F1ADFD68FBAA" box="[568,669,1015,1042]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="3" pageNumber="534" unit="mm" value="0.59">0.59 mm</quantity>
, club formed by apical three enlarged antennomeres; antennomeres 1 and 2 each strongly elongate, 37 gradually longer, 8 as long as and slightly broader than 7, 9 weakly transverse, 10 more distinctly transverse and much larger than 9, 11 as wide as 9, suboval, club (
<quantity id="4CC79B5CFFED857FFBA6F665FB89FBE1" box="[1047,1148,1087,1114]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="534" unit="mm" value="0.25">0.25 mm</quantity>
) less than half of antennal length.
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F6D2FE50FAB6" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
Pronotum roundly triangular, length
<quantity id="4CC79B5CFFED857FFDD8F6DDFD25FB1A" box="[617,720,1159,1186]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="3" pageNumber="534" unit="mm" value="0.36">0.36 mm</quantity>
, width
<quantity id="4CC79B5CFFED857FFC97F6DDFC7BFB1A" box="[806,910,1159,1186]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="534" unit="mm" value="0.42">0.42 mm</quantity>
, broadest near posterior 1/3, distinctly rapidly narrowing from broadest point toward apex.Anterior margin slightly arcuate, posterior margin indistinctly bisinuate and weakly convex posteriorly at middle, lacking antebasal pits. Pronotal disc almost impunctate, lateral portion with few suberect setae.
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F74DFAB5FAC1" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
Elytra together oval, broadest approximately at middle, length
<quantity id="4CC79B5CFFED857FFC29F74DFBF5FA8A" box="[920,1024,1303,1330]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="3" pageNumber="534" unit="mm" value="0.78">0.78 mm</quantity>
, width
<quantity id="4CC79B5CFFED857FFBE6F74DFB4AFA8A" box="[1111,1215,1303,1330]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="534" unit="mm" value="0.68">0.68 mm</quantity>
, length/width 1.15; humeral calli distinct, basal impressions short but well developed; apices separately rounded; disc almost impunctate, glabrous, only center with few scattered short setae. Metathoracic wings fully-developed, long, functional.
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F7D9FA7EFA79" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
Mesoventrite (
<figureCitation id="13042A3CFFED857FFEDAF7D9FE37FA25" box="[363,450,1411,1437]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. FI. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in EI." pageId="3" pageNumber="534">Fig. 2C</figureCitation>
) with broad intercoxal process; large, transversely oval area surrounding coxae greatly impressed and with setose fringes. Metaventrite with intercoxal process (
<figureCitation id="13042A3CFFED857FFC79F7FDFBD5FA79" box="[968,1056,1447,1473]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. FI. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in EI." pageId="3" pageNumber="534">Fig. 2D</figureCitation>
) formed by pair of acute spines.
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F796FAD0FA5D" blockId="3.[151,1437,295,2014]" box="[199,1317,1483,1510]" pageId="3" pageNumber="534">
Legs moderately long and slender, unmodified; with small plantulae (
<figureCitation id="13042A3CFFED857FFC63F791FBD3FA5D" box="[978,1062,1483,1509]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[170,1417,181,1918]" captionTargetId="figure-20@4.[170,1417,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Himaloconnus obesus sp. nov., male.A. Dorsal habitus. B. Head venter. C. Mesoventrite. D. Metaventral intercoxal process. E. Metatarsus. FI. Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in EI." pageId="3" pageNumber="534">Fig. 2E</figureCitation>
) between tarsal claws.
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F7B5FECEF906" blockId="3.[151,1437,295,2014]" pageId="3" pageNumber="534">
Aedeagus (Figs FI) elongate, length
<quantity id="4CC79B5CFFED857FFDD7F7B5FD3EF9B2" box="[614,715,1519,1546]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="534" unit="mm" value="0.26">0.26 mm</quantity>
, dorso-ventrally symmetric; in ventral view broadest in sub-basal region, weakly narrowing distad; apical region broadly subtriangular with blunt apex, with acute apical projection at middle; endophallus armature symmetric and complex, composed of longitudinal tubular component flanked by variously distinct, sclerotized plates of various shapes, in submedian region tubular structure distinctly broadened and forming large vesicle well-visible in lateral view; parameres slender, each with one long apical and one short subapical seta.
</paragraph>
<paragraph id="8B8036B9FFED857FFF76F49DFE6AF959" blockId="3.[151,1437,295,2014]" box="[199,415,1735,1762]" pageId="3" pageNumber="534">
<emphasis id="B94BEAABFFED857FFF76F49DFEEEF959" box="[199,283,1735,1761]" italics="true" pageId="3" pageNumber="534">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B8036B9FFED8579FF76F4B1FC13FDA1" blockId="3.[151,1437,295,2014]" lastBlockId="5.[151,1437,151,645]" lastPageId="5" lastPageNumber="536" pageId="3" pageNumber="534">
<emphasis id="B94BEAABFFED857FFF76F4B1FE47F8BD" bold="true" box="[199,434,1771,1797]" pageId="3" pageNumber="534">Comparative notes.</emphasis>
Nine species of
<taxonomicName id="4C3F4D3AFFED857FFDC2F4B1FCAAF8BE" authority="Franz" authorityName="Franz" authorityYear="1979" box="[627,863,1771,1798]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABFFED857FFDC2F4B1FCE3F8BD" box="[627,790,1771,1797]" italics="true" pageId="3" pageNumber="534">Himaloconnus</emphasis>
Franz
</taxonomicName>
have been described from
<collectingCountry id="F3287629FFED857FFB27F4B1FB2CF8BD" box="[1174,1241,1771,1797]" name="China" pageId="3" pageNumber="534">China</collectingCountry>
(
<collectingRegion id="49FBF85BFFED857FFB58F4B1FABCF8BD" box="[1257,1353,1771,1797]" country="China" name="Taiwan)" pageId="3" pageNumber="534">Taiwan)</collectingRegion>
,
<collectingCountry id="F3287629FFED857FFAE4F4B6FA6DF8BE" box="[1365,1432,1772,1798]" name="Japan" pageId="3" pageNumber="534">Japan</collectingCountry>
,
<collectingCountry id="F3287629FFED857FFF26F555FF08F891" box="[151,253,1807,1833]" name="Thailand" pageId="3" pageNumber="534">Thailand</collectingCountry>
,
<collectingCountry id="F3287629FFED857FFEB6F555FEB8F891" box="[263,333,1807,1833]" name="Nepal" pageId="3" pageNumber="534">Nepal</collectingCountry>
, and northern
<collectingCountry id="F3287629FFED857FFE58F555FDD6F891" box="[489,547,1807,1833]" name="India" pageId="3" pageNumber="534">India</collectingCountry>
(
<bibRefCitation id="EFAE4B48FFED857FFD9EF555FD00F891" author="Jaloszynski, P." box="[559,757,1807,1833]" pageId="3" pageNumber="534" pagination="334 - 360" refId="ref5078" refString="Jaloszynski, P. (2020) Himaloconnus Franz and Nogunius gen. n. of Japan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4822 (3), 334 - 360. https: // doi. org / 10.11646 / zootaxa. 4822.3.2" type="journal article" year="2020">Jałoszyński 2020</bibRefCitation>
). The aedeagi of most species, where known, exhibit a similar morphology. Only two species,
<taxonomicName id="4C3F4D3AFFED857FFDBBF56EFD0EF8F5" authority="(Franz)" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[522,763,1843,1870]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="B94BEAABFFED857FFDBBF56EFDEBF8F5" box="[522,542,1844,1869]" italics="true" pageId="3" pageNumber="534">H</emphasis>
.
<emphasis id="B94BEAABFFED857FFD81F569FD6BF8F5" box="[560,670,1843,1869]" italics="true" pageId="3" pageNumber="534">shutjensis</emphasis>
(Franz)
</taxonomicName>
from
<collectingCountry id="F3287629FFED857FFCF6F569FC7FF8F5" box="[839,906,1843,1869]" name="Nepal" pageId="3" pageNumber="534">Nepal</collectingCountry>
and
<taxonomicName id="4C3F4D3AFFED857FFC79F56EFB0AF8F5" authority="(Franz)" baseAuthorityName="Franz" baseAuthorityYear="1985" box="[968,1279,1843,1870]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="klapperichianus">
<emphasis id="B94BEAABFFED857FFC79F56EFC29F8F5" box="[968,988,1844,1869]" italics="true" pageId="3" pageNumber="534">H</emphasis>
.
<emphasis id="B94BEAABFFED857FFC5CF569FB54F8F5" box="[1005,1185,1843,1869]" italics="true" pageId="3" pageNumber="534">klapperichianus</emphasis>
(Franz)
</taxonomicName>
from
<collectingCountry id="F3287629FFED857FFAFBF569FF2EF8C9" name="Taiwan Island" pageId="3" pageNumber="534">Taiwan Island</collectingCountry>
and
<collectingCountry id="F3287629FFED857FFEBEF502FEA7F8CA" box="[271,338,1880,1906]" name="Japan" pageId="3" pageNumber="534">Japan</collectingCountry>
, have been adequately characterized.
<taxonomicName id="4C3F4D3AFFED857FFD43F50DFC12F8C9" authorityName="Yin &amp; Zhou" authorityYear="2025" box="[754,999,1879,1905]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="obesus" status="sp. nov.">
<emphasis id="B94BEAABFFED857FFD43F50DFC12F8C9" box="[754,999,1879,1905]" italics="true" pageId="3" pageNumber="534">Himaloconnus obesus</emphasis>
</taxonomicName>
<emphasis id="B94BEAABFFED857FFC5CF50DFBB0F8C9" bold="true" box="[1005,1093,1879,1905]" pageId="3" pageNumber="534">
<taxonomicNameLabel id="A27857D0FFED857FFC5CF50DFBB0F8C9" box="[1005,1093,1879,1905]" pageId="3" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is readily distinguishable from both species by its significantly larger body size, and a bicolored and nearly glabrous dorsal surface of the body. In contrast,
<taxonomicName id="4C3F4D3AFFED857FFF4CF5FAFE78F801" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[253,397,1951,1978]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="shutjensis">
<emphasis id="B94BEAABFFED857FFF4CF5FAFEE4F801" box="[253,273,1952,1977]" italics="true" pageId="3" pageNumber="534">H</emphasis>
.
<emphasis id="B94BEAABFFED857FFEAEF5C5FE78F801" box="[287,397,1951,1977]" italics="true" pageId="3" pageNumber="534">shutjensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D3AFFED857FFE73F5FAFD6DF801" baseAuthorityName="Franz" baseAuthorityYear="1985" box="[450,664,1951,1978]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="klapperichianus">
<emphasis id="B94BEAABFFED857FFE73F5FAFE23F801" box="[450,470,1952,1977]" italics="true" pageId="3" pageNumber="534">H</emphasis>
.
<emphasis id="B94BEAABFFED857FFE55F5C5FD6DF801" box="[484,664,1951,1977]" italics="true" pageId="3" pageNumber="534">klapperichianus</emphasis>
</taxonomicName>
are considerably smaller (≤ 1.0 mm), have uniformly colored bodies, and exhibit evenly distributed setae on the pronotum and elytra. The body size of
<taxonomicName id="4C3F4D3AFFED857FFB87F59EFB50F865" authorityName="Yin &amp; Zhou" authorityYear="2025" box="[1078,1189,1987,2014]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="obesus" status="sp. nov.">
<emphasis id="B94BEAABFFED857FFB87F59EFBBFF865" box="[1078,1098,1988,2013]" italics="true" pageId="3" pageNumber="534">H</emphasis>
.
<emphasis id="B94BEAABFFED857FFBE8F599FB50F865" box="[1113,1189,1987,2013]" italics="true" pageId="3" pageNumber="534">obesus</emphasis>
</taxonomicName>
<emphasis id="B94BEAABFFED857FFB1CF599FAFDF865" bold="true" box="[1197,1288,1987,2013]" pageId="3" pageNumber="534">
<taxonomicNameLabel id="A27857D0FFED857FFB1CF599FAFDF865" box="[1197,1288,1987,2013]" pageId="3" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
reaches
<quantity id="4CC79B5CFFED8579FADAF599FF30FF09" lastPageId="5" lastPageNumber="536" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="3" pageNumber="534" unit="mm" value="1.38">1.38 mm</quantity>
, making it one of the largest known species within the genus. Among other species, only
<taxonomicName id="4C3F4D3AFFEB8579FB28F2CDFA69FF09" authority="(Franz)" baseAuthorityName="Franz" baseAuthorityYear="1985" box="[1177,1436,151,177]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="alishanensis">
<emphasis id="B94BEAABFFEB8579FB28F2CDFB58FF08" box="[1177,1197,151,176]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FB0BF2CDFAB1FF09" box="[1210,1348,151,177]" italics="true" pageId="5" pageNumber="536">alishanensis</emphasis>
(Franz)
</taxonomicName>
(
<quantity id="4CC79B5CFFEB8579FF2EF2E0FEF1FF6D" box="[159,260,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="536" unit="mm" value="1.5">1.50 mm</quantity>
) from
<collectingCountry id="F3287629FFEB8579FEFCF2E1FE1CFF6D" box="[333,489,187,213]" name="Taiwan Island" pageId="5" pageNumber="536">Taiwan Island</collectingCountry>
and
<taxonomicName id="4C3F4D3AFFEB8579FDACF2E1FD30FF6C" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[541,709,187,213]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="ramamensis">
<emphasis id="B94BEAABFFEB8579FDACF2E1FDC4FF6C" box="[541,561,187,212]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FD8FF2E1FD30FF6C" box="[574,709,187,212]" italics="true" pageId="5" pageNumber="536">ramamensis</emphasis>
</taxonomicName>
(
<quantity id="4CC79B5CFFEB8579FD62F2E1FCCDFF6D" box="[723,824,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="536" unit="mm" value="1.4">1.40 mm</quantity>
) from
<collectingCountry id="F3287629FFEB8579FC33F2E1FC30FF6D" box="[898,965,187,213]" name="Nepal" pageId="5" pageNumber="536">Nepal</collectingCountry>
have comparative body sizes (
<bibRefCitation id="EFAE4B48FFEB8579FAA7F2E1FA6DFF6D" author="Franz, H." box="[1302,1432,187,213]" pageId="5" pageNumber="536" pagination="235 - 274" refId="ref4833" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz 1979</bibRefCitation>
,
<bibRefCitation id="EFAE4B48FFEB8579FF26F284FF26FF41" author="Franz, H." box="[151,211,222,249]" pageId="5" pageNumber="536" pagination="91 - 128" refId="ref4896" refString="Franz, H. (1985) Neue und ungenugend bekannte Scydmaeniden (Coleoptera) aus Taiwan, Fukien und Thailand. Mitteilungen der Munchner Entomologischen Gesellschaft, 74, 91 - 128. n [1984]" type="journal article" year="1985">1985</bibRefCitation>
). The new species can be separated from
<taxonomicName id="4C3F4D3AFFEB8579FD14F285FCA4FF41" baseAuthorityName="Franz" baseAuthorityYear="1985" box="[677,849,223,249]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="alishanensis">
<emphasis id="B94BEAABFFEB8579FD14F285FD4CFF40" box="[677,697,223,248]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FD76F285FCA4FF41" box="[711,849,223,249]" italics="true" pageId="5" pageNumber="536">alishanensis</emphasis>
</taxonomicName>
by the black head and reddish-brown pronotum and elytra, in contrast to the black head and pronotum and reddish-brown elytra of the latter species. Additionally,
<taxonomicName id="4C3F4D3AFFEB8579FA30F359FF16FEF9" authorityName="Yin &amp; Zhou" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="534" phylum="Arthropoda" rank="species" species="obesus" status="sp. nov.">
H.
<emphasis id="B94BEAABFFEB8579FF26F37DFF16FEF9" box="[151,227,295,321]" italics="true" pageId="5" pageNumber="536">obesus</emphasis>
</taxonomicName>
<emphasis id="B94BEAABFFEB8579FF5CF37DFEBCFEF9" bold="true" box="[237,329,295,321]" pageId="5" pageNumber="536">
<taxonomicNameLabel id="A27857D0FFEB8579FF5CF37DFEBCFEF9" box="[237,329,295,321]" pageId="5" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lacks antebasal pits on the pronotal disc, whereas
<taxonomicName id="4C3F4D3AFFEB8579FC24F37DFBB1FEF9" baseAuthorityName="Franz" baseAuthorityYear="1985" box="[917,1092,295,321]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="alishanensis">
<emphasis id="B94BEAABFFEB8579FC24F37DFC5CFEF8" box="[917,937,295,320]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FC0BF37DFBB1FEF9" box="[954,1092,295,321]" italics="true" pageId="5" pageNumber="536">alishanensis</emphasis>
</taxonomicName>
possesses six small antebasal pits. From
<taxonomicName id="4C3F4D3AFFEB8579FEA8F311FE33FEDC" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[281,454,331,357]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="ramamensis">
<emphasis id="B94BEAABFFEB8579FEA8F311FED8FEDC" box="[281,301,331,356]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FE8EF311FE33FEDC" box="[319,454,331,356]" italics="true" pageId="5" pageNumber="536">ramamensis</emphasis>
</taxonomicName>
the new species differs by the bicolored body (uniformly dark reddish-brown in
<taxonomicName id="4C3F4D3AFFEB8579FA30F311FEEBFE30" baseAuthorityName="Franz" baseAuthorityYear="1979" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="ramamensis">
<emphasis id="B94BEAABFFEB8579FA30F311FA60FEDC" box="[1409,1429,331,356]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FF26F335FEEBFE30" box="[151,286,367,392]" italics="true" pageId="5" pageNumber="536">ramamensis</emphasis>
</taxonomicName>
), and the lack of antebasal pits on the pronotal disc (with one pair of antebasal pits on the pronotal disc of
<taxonomicName id="4C3F4D3AFFEB8579FF06F3C9FE94FE14" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[183,353,403,429]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="ramamensis">
<emphasis id="B94BEAABFFEB8579FF06F3C9FF3EFE14" box="[183,203,403,428]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FF6BF3C9FE94FE14" box="[218,353,403,428]" italics="true" pageId="5" pageNumber="536">ramamensis</emphasis>
</taxonomicName>
). The remaining species,
<taxonomicName id="4C3F4D3AFFEB8579FD36F3C9FC37FE15" authority="Newton" authorityName="Newton" authorityYear="2017" box="[647,962,403,429]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="reductipenisides">
<emphasis id="B94BEAABFFEB8579FD36F3C9FC94FE15" box="[647,865,403,429]" italics="true" pageId="5" pageNumber="536">H. reductipenisides</emphasis>
Newton
</taxonomicName>
(
<quantity id="4CC79B5CFFEB8579FC60F3C8FB98FE15" box="[977,1133,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="5" pageNumber="536" unit="mm" value="0.925" valueMax="0.95" valueMin="0.9">0.90.95 mm</quantity>
),
<taxonomicName id="4C3F4D3AFFEB8579FB30F3C9FA68FE15" authority="(Franz)" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[1153,1437,403,429]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="nagarkotensis">
<emphasis id="B94BEAABFFEB8579FB30F3C9FB60FE14" box="[1153,1173,403,428]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FB14F3C9FAB7FE15" box="[1189,1346,403,429]" italics="true" pageId="5" pageNumber="536">nagarkotensis</emphasis>
(Franz)
</taxonomicName>
(
<quantity id="4CC79B5CFFEB8579FF2EF3ECFEFCFE69" box="[159,265,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="536" unit="mm" value="1.05">1.05 mm</quantity>
),
<taxonomicName id="4C3F4D3AFFEB8579FEACF3EDFE29FE69" authority="(Franz)" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[285,476,439,465]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="topali">
<emphasis id="B94BEAABFFEB8579FEACF3EDFEC4FE68" box="[285,305,439,464]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FEF1F3EDFE74FE69" box="[320,385,439,465]" italics="true" pageId="5" pageNumber="536">topali</emphasis>
(Franz)
</taxonomicName>
(
<quantity id="4CC79B5CFFEB8579FE5AF3EDFD61FE69" box="[491,660,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="5" pageNumber="536" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.800.90 mm</quantity>
),
<taxonomicName id="4C3F4D3AFFEB8579FD19F3EDFC5FFE69" authority="(Franz)" baseAuthorityName="Franz" baseAuthorityYear="1981" box="[680,938,439,465]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="pokharanus">
<emphasis id="B94BEAABFFEB8579FD19F3EDFD49FE68" box="[680,700,439,464]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FD7AF3EDFCBAFE69" box="[715,847,439,465]" italics="true" pageId="5" pageNumber="536">pokharanus</emphasis>
(Franz)
</taxonomicName>
(
<quantity id="4CC79B5CFFEB8579FC08F3ECFBD1FE69" box="[953,1060,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="536" unit="mm" value="0.95">0.95 mm</quantity>
),
<taxonomicName id="4C3F4D3AFFEB8579FB86F3EDFB01FE69" authority="(Franz)" baseAuthorityName="Franz" baseAuthorityYear="1981" box="[1079,1268,439,465]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="hindu">
<emphasis id="B94BEAABFFEB8579FB86F3EDFBBEFE68" box="[1079,1099,439,464]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FBEBF3EDFB6FFE69" box="[1114,1178,439,465]" italics="true" pageId="5" pageNumber="536">hindu</emphasis>
(Franz)
</taxonomicName>
(
<quantity id="4CC79B5CFFEB8579FAB5F3EDFA95FE69" box="[1284,1376,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="536" unit="mm" value="0.8">0.8 mm</quantity>
), and
<taxonomicName id="4C3F4D3AFFEB8579FF26F381FEA9FE4D" authority="(Franz)" baseAuthorityName="Franz" baseAuthorityYear="1979" box="[151,348,475,501]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="topali">
<emphasis id="B94BEAABFFEB8579FF26F381FF5EFE4C" box="[151,171,475,500]" italics="true" pageId="5" pageNumber="536">H</emphasis>
.
<emphasis id="B94BEAABFFEB8579FF0CF381FF0BFE4D" box="[189,254,475,501]" italics="true" pageId="5" pageNumber="536">topali</emphasis>
(Franz)
</taxonomicName>
(
<quantity id="4CC79B5CFFEB8579FEDEF381FE3BFE4D" box="[367,462,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="536" unit="mm" value="0.8">0.8 mm</quantity>
), are all much smaller (
<bibRefCitation id="EFAE4B48FFEB8579FD56F381FC9BFE4D" author="Franz, H." box="[743,878,475,501]" pageId="5" pageNumber="536" pagination="235 - 274" refId="ref4833" refString="Franz, H. (1979) Weitere Beitrage zur Kenntnis der Scydmaenidenfauna des Himalaja und seiner Grenzgebiete (Coleoptera: Scydmaenidae). Entomologica Basiliensia, 4, 235 - 274." type="journal article" year="1979">Franz 1979</bibRefCitation>
,
<bibRefCitation id="EFAE4B48FFEB8579FCCCF381FC42FE4D" author="Franz, H." box="[893,951,475,501]" pageId="5" pageNumber="536" pagination="295 - 316" refId="ref4866" refString="Franz, H. (1981) 10. Beitrag zur Kenntnis der Scydmaenidenfauna des Himalaja (Col.). Entomologica Basiliensia, 6, 295 - 316." type="journal article" year="1981">1981</bibRefCitation>
,
<bibRefCitation id="EFAE4B48FFEB8579FC76F381FBF6FE4D" author="Franz, H." box="[967,1027,475,501]" pageId="5" pageNumber="536" pagination="33 - 80" refId="ref4962" refString="Franz, H. (1989) 4. Beitrag zur Scydmaenidenfauna von Thailand (Die Scydmaenidenausbeute von D. H. Burckhardt u. I. Lobl). Revue suisse de zoologie, 96 (1), 33 - 80. https: // doi. org / 10.5962 / bhl. part. 82027" type="journal article" year="1989">1989</bibRefCitation>
).
<taxonomicName id="4C3F4D3AFFEB8579FBABF381FA8AFE4D" authorityName="Newton" authorityYear="2017" box="[1050,1407,475,501]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="536" phylum="Arthropoda" rank="species" species="reductipenisides">
<emphasis id="B94BEAABFFEB8579FBABF381FA8AFE4D" box="[1050,1407,475,501]" italics="true" pageId="5" pageNumber="536">Himaloconnus reductipenisides</emphasis>
</taxonomicName>
is distributed in
<collectingCountry id="F3287629FFEB8579FE83F3A5FE6DFDA1" box="[306,408,511,537]" name="Thailand" pageId="5" pageNumber="536">Thailand</collectingCountry>
, while the others are found in the Himalayan region.
</paragraph>
</subSubSection>
<caption id="DF406631FFEA8578FF26F5CDFF36F84F" pageId="4" pageNumber="535" startId="4.[151,250,1943,1967]" targetBox="[170,1417,181,1918]" targetPageId="4" targetType="figure">
<paragraph id="8B8036B9FFEA8578FF26F5CDFF36F84F" blockId="4.[151,1436,1943,2039]" pageId="4" pageNumber="535">
<emphasis id="B94BEAABFFEA8578FF26F5CDFEE7F817" bold="true" box="[151,274,1943,1967]" pageId="4" pageNumber="535">FIGURE 2.</emphasis>
<taxonomicName id="4C3F4D3AFFEA8578FEA6F5CDFE00F817" authorityName="Yin &amp; Zhou" authorityYear="2025" box="[279,501,1943,1967]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="535" phylum="Arthropoda" rank="species" species="obesus" status="sp. nov.">
<emphasis id="B94BEAABFFEA8578FEA6F5CDFE00F817" box="[279,501,1943,1967]" italics="true" pageId="4" pageNumber="535">Himaloconnus obesus</emphasis>
</taxonomicName>
<emphasis id="B94BEAABFFEA8578FE4BF5C2FDBCF808" bold="true" box="[506,585,1944,1968]" pageId="4" pageNumber="535">
<taxonomicNameLabel id="A27857D0FFEA8578FE4BF5C2FDBCF808" box="[506,585,1944,1968]" pageId="4" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B94BEAABFFEA8578FD21F5CDFD5CF817" bold="true" box="[656,681,1943,1967]" pageId="4" pageNumber="535">A.</emphasis>
Dorsal habitus.
<emphasis id="B94BEAABFFEA8578FCFAF5C2FC97F808" bold="true" box="[843,866,1944,1968]" pageId="4" pageNumber="535">B.</emphasis>
Head venter.
<emphasis id="B94BEAABFFEA8578FC5AF5CDFBF1F817" bold="true" box="[1003,1028,1943,1967]" pageId="4" pageNumber="535">C.</emphasis>
Mesoventrite.
<emphasis id="B94BEAABFFEA8578FB2BF5C2FB47F808" bold="true" box="[1178,1202,1944,1968]" pageId="4" pageNumber="535">D.</emphasis>
Metaventral intercoxal process.
<emphasis id="B94BEAABFFEA8578FF40F5E6FEFDF86C" bold="true" box="[241,264,1980,2004]" pageId="4" pageNumber="535">E.</emphasis>
Metatarsus.
<emphasis id="B94BEAABFFEA8578FE3DF5E6FE4CF86C" bold="true" box="[396,441,1980,2004]" pageId="4" pageNumber="535">FI.</emphasis>
Aedeagus, lateral (F, H) and ventral (G, I). Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in EI.
</paragraph>
</caption>
<subSubSection id="C3256532FFEB8579FF76F078FD64FD85" box="[199,657,546,573]" pageId="5" pageNumber="536" type="distribution">
<paragraph id="8B8036B9FFEB8579FF76F078FD64FD85" blockId="5.[151,1437,151,645]" box="[199,657,546,573]" pageId="5" pageNumber="536">
<emphasis id="B94BEAABFFEB8579FF76F078FE94FD84" bold="true" box="[199,353,546,572]" pageId="5" pageNumber="536">Distribution.</emphasis>
Southwest
<collectingCountry id="F3287629FFEB8579FE52F079FDDFFD85" box="[483,554,547,573]" name="China" pageId="5" pageNumber="536">China</collectingCountry>
:
<collectingRegion id="49FBF85BFFEB8579FD85F079FD78FD85" box="[564,653,547,573]" country="China" name="Yunnan" pageId="5" pageNumber="536">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEB8579FF76F01DFF05FD3D" pageId="5" pageNumber="536" type="etymology">
<paragraph id="8B8036B9FFEB8579FF76F01DFF05FD3D" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="536">
<emphasis id="B94BEAABFFEB8579FF76F01DFEBBFDD9" bold="true" box="[199,334,583,609]" pageId="5" pageNumber="536">Etymology.</emphasis>
The Latin participle
<emphasis id="B94BEAABFFEB8579FDF3F01DFD7BFDD9" box="[578,654,583,609]" italics="true" pageId="5" pageNumber="536">obçsus</emphasis>
(-
<emphasis id="B94BEAABFFEB8579FD1BF01DFD4DFDD8" box="[682,696,583,608]" italics="true" pageId="5" pageNumber="536">a</emphasis>
, -
<emphasis id="B94BEAABFFEB8579FD63F01DFD01FDD8" box="[722,756,583,608]" italics="true" pageId="5" pageNumber="536">um</emphasis>
) means fat, stout, referring to the stout body form of this species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,353 @@
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<mods:title id="D3675B5DB447F3923B6AEABD02517E6D">A report of the Scydmaeninae species (Coleoptera: Staphylinidae) taken from Wumenshan National Nature Reserve, Yunnan, China</mods:title>
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<mods:namePart id="BC35C85A72A392CBE69AA346B667B107">Yin, Zi-Wei</mods:namePart>
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<mods:namePart id="3AEC412D994E7CF61C8002FA4423B7D0">Zhou, De-Yao</mods:namePart>
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<treatment id="039687AFFFEF857EFF26F3D2FDE0F850" LSID="urn:lsid:plazi:treatment:039687AFFFEF857EFF26F3D2FDE0F850" httpUri="http://treatment.plazi.org/id/039687AFFFEF857EFF26F3D2FDE0F850" lastPageId="2" lastPageNumber="533" pageId="1" pageNumber="532">
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<paragraph id="8B8036B9FFEF857DFF26F3D2FCC8FE1A" blockId="1.[151,829,391,490]" box="[151,829,391,418]" pageId="1" pageNumber="532">
<heading id="D0C881D5FFEF857DFF26F3D2FCC8FE1A" bold="true" box="[151,829,391,418]" fontSize="11" level="1" pageId="1" pageNumber="532" reason="1">
<emphasis id="B94BEAABFFEF857DFF26F3D2FCC8FE1A" bold="true" box="[151,829,391,418]" pageId="1" pageNumber="532">
<taxonomicName id="4C3F4D3AFFEF857DFF26F3D2FD26FE1A" authority="Yin &amp; Zhou" authorityName="Yin &amp; Zhou" authorityYear="2025" box="[151,723,391,418]" class="Insecta" family="Staphylinidae" genus="Euconnus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="wumengshanus" status="sp. nov.">
<emphasis id="B94BEAABFFEF857DFF26F3D2FEE6FE1A" bold="true" box="[151,275,392,418]" italics="true" pageId="1" pageNumber="532">Euconnus</emphasis>
(
<emphasis id="B94BEAABFFEF857DFE95F3D2FEDAFE1A" bold="true" box="[292,303,392,418]" italics="true" pageId="1" pageNumber="532">s</emphasis>
.
<emphasis id="B94BEAABFFEF857DFE8FF3D2FEA8FE1A" bold="true" box="[318,349,392,418]" italics="true" pageId="1" pageNumber="532">str</emphasis>
.)
<emphasis id="B94BEAABFFEF857DFEC7F3DDFDC6FE1A" bold="true" box="[374,563,391,418]" italics="true" pageId="1" pageNumber="532">wumengshanus</emphasis>
Yin &amp; Zhou
</taxonomicName>
,
<taxonomicNameLabel id="A27857D0FFEF857DFD6EF3D2FCC8FE1A" box="[735,829,392,418]" pageId="1" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8036B9FFEF857DFF26F3F6FD85FE7C" blockId="1.[151,829,391,490]" box="[151,624,428,455]" pageId="1" pageNumber="532">
Chinese common name:
<emphasis id="B94BEAABFFEF857DFE1CF3F7FD85FE7C" bold="true" box="[429,624,429,452]" pageId="1" pageNumber="532">乌AE山ṀẆẅm</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEF857DFF26F38AFF1EFE52" box="[151,235,464,490]" pageId="1" pageNumber="532" type="description">
<paragraph id="8B8036B9FFEF857DFF26F38AFF1EFE52" blockId="1.[151,829,391,490]" box="[151,235,464,490]" pageId="1" pageNumber="532">
(
<figureCitation id="13042A3CFFEF857DFF2EF38AFF15FE52" box="[159,224,464,490]" captionStart="FIGURE 1" captionStartId="2.[151,250,1316,1340]" captionTargetBox="[180,1409,189,1287]" captionTargetId="figure-20@2.[177,1411,179,1289]" captionTargetPageId="2" captionText="FIGURE 1. Euconnus (s. str.) wumengshanus sp. nov. (AE. Male. F. Female). A. Dorsal habitus. BE. Aedeagus, lateral (B, D) and ventral (C, E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A; 0.1 mm in BE; 0.05 mm in F." pageId="1" pageNumber="532">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEF857DFF26F042FE63FD5E" pageId="1" pageNumber="532" type="materials_examined">
<paragraph id="8B8036B9FFEF857DFF26F042FE63FDC2" blockId="1.[151,1437,536,2039]" pageId="1" pageNumber="532">
<emphasis id="B94BEAABFFEF857DFF26F042FEC8FD8A" bold="true" box="[151,317,536,562]" pageId="1" pageNumber="532">Type material</emphasis>
(
<materialsCitation id="3B573CE4FFEF857DFEFEF042FE63FDC2" collectingDate="2022-07-15" collectionCode="SNUC" collectorName="Wei" country="China" county="China" elevation="1980" latitude="28.207336" location="Wumengshan Natural Reserve" longLatPrecision="1" longitude="103.91411" municipality="Zhaotong City" pageId="1" pageNumber="532" specimenCount="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="9D39FD30FFEF857DFEFEF042FE71FD8A" box="[335,388,536,562]" count="1" pageId="1" pageNumber="532" type="generic">1 ex.</specimenCount>
).
<emphasis id="B94BEAABFFEF857DFE2BF042FD5AFD8A" bold="true" box="[410,687,536,562]" pageId="1" pageNumber="532">
<typeStatus id="5484881BFFEF857DFE2BF042FDC8FD8A" box="[410,573,536,562]" pageId="1" pageNumber="532" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="F3287629FFEF857DFDF9F042FD5FFD8A" box="[584,682,536,562]" name="China" pageId="1" pageNumber="532">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="9D39FD30FFEF857DFD04F043FD38FD8B" box="[693,717,537,563]" count="1" pageId="1" pageNumber="532" type="male"></specimenCount>
:
<collectingCounty id="62E14E35FFEF857DFD6CF042FCDCFD8A" box="[733,809,536,562]" pageId="1" pageNumber="532">China</collectingCounty>
:
<collectingRegion id="49FBF85BFFEF857DFC80F043FC7FFD8B" box="[817,906,537,563]" country="China" name="Yunnan" pageId="1" pageNumber="532">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BE4ACC3FFEF857DFC25F042FBC2FD8A" box="[916,1079,536,562]" pageId="1" pageNumber="532">Zhaotong City</collectingMunicipality>
,
<location id="8EE06062FFEF857DFBF0F042FA6DFD8B" LSID="urn:lsid:plazi:treatment:039687AFFFEF857EFF26F3D2FDE0F850:8EE06062FFEF857DFBF0F042FA6DFD8B" box="[1089,1432,536,563]" country="China" county="China" latitude="28.207336" longLatPrecision="1" longitude="103.91411" municipality="Zhaotong City" name="Wumengshan Natural Reserve" pageId="1" pageNumber="532" stateProvince="Yunnan">Wumengshan Natural Reserve</location>
,
<location id="8EE06062FFEF857DFF26F066FED3FDEE" LSID="urn:lsid:plazi:treatment:039687AFFFEF857EFF26F3D2FDE0F850:8EE06062FFEF857DFF26F066FED3FDEE" box="[151,294,572,598]" country="China" county="China" latitude="28.207336" longLatPrecision="1" longitude="103.91411" municipality="Zhaotong City" name="Sanjiangkou" pageId="1" pageNumber="532" stateProvince="Yunnan">Sanjiangkou</location>
,
<location id="8EE06062FFEF857DFE80F066FE62FDEE" LSID="urn:lsid:plazi:treatment:039687AFFFEF857EFF26F3D2FDE0F850:8EE06062FFEF857DFE80F066FE62FDEE" box="[305,407,572,598]" country="China" county="China" latitude="28.207336" longLatPrecision="1" longitude="103.91411" municipality="Zhaotong City" name="Laziping" pageId="1" pageNumber="532" stateProvince="Yunnan">Laziping</location>
,
<geoCoordinate id="EE0B507EFFEF857DFE13F066FDCEFDEE" box="[418,571,572,598]" degrees="28.207336" direction="north" orientation="latitude" pageId="1" pageNumber="532" precision="1" value="28.207336">28.207336°N</geoCoordinate>
,
<geoCoordinate id="EE0B507EFFEF857DFDF7F066FD1FFDEE" box="[582,746,572,598]" degrees="103.914108" direction="east" orientation="longitude" pageId="1" pageNumber="532" precision="1" value="103.91411">103.914108°E</geoCoordinate>
,
<quantity id="4CC79B5CFFEF857DFD45F066FCBEFDEF" box="[756,843,572,599]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="1" pageNumber="532" unit="m" value="1980.0">
<elevation id="0012D18AFFEF857DFD45F066FCBEFDEF" box="[756,843,572,599]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="1" pageNumber="532" unit="m" value="1980.0">1980 m</elevation>
</quantity>
,
<date id="FF811079FFEF857DFCE7F066FC2DFDEF" box="[854,984,572,599]" pageId="1" pageNumber="532" value="2022-07-15">
<collectingDate id="EFC5E991FFEF857DFCE7F066FC2DFDEF" box="[854,984,572,599]" pageId="1" pageNumber="532" value="2022-07-15">15.vii.2022</collectingDate>
</date>
,
<collectorName id="26CA536FFFEF857DFC52F066FBFAFDEE" box="[995,1039,572,598]" pageId="1" pageNumber="532">Wei</collectorName>
leg.,
<emphasis id="B94BEAABFFEF857DFBFDF067FED6FDC0" bold="true" pageId="1" pageNumber="532">云南DZdz乌AE山三江口ḆŦ 坪M国Dzẍ</emphasis>
(
<collectionCode id="ED2EAE7CFFEF857DFE88F03AFE7FFDC2" box="[313,394,608,634]" pageId="1" pageNumber="532">SNUC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8036B9FFEF857DFF76F0DEFE63FD5E" blockId="1.[151,1437,536,2039]" pageId="1" pageNumber="532">
<emphasis id="B94BEAABFFEF857DFF76F0DEFDF5FD26" bold="true" box="[199,512,644,670]" pageId="1" pageNumber="532">Other material examined</emphasis>
(
<specimenCount id="9D39FD30FFEF857DFDA6F0DEFDA6FD26" box="[535,595,644,670]" count="1" pageId="1" pageNumber="532" type="generic">1 ex.</specimenCount>
).
<materialsCitation id="3B573CE4FFEF857DFDDEF0DEFE67FD5E" collectingDate="2022-07-20" collectionCode="SNUC" collectorName="Wei" country="China" county="Zhaotong City" elevation="1830" latitude="27.846716" location="Xiaocaoba" longLatPrecision="1" longitude="104.29242" municipality="Wumengshan Natural Reserve" pageId="1" pageNumber="532" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D39FD30FFEF857DFDDEF0DEFD55FD27" box="[623,672,644,671]" count="1" pageId="1" pageNumber="532" type="female">1 ♀</specimenCount>
:
<collectingCountry id="F3287629FFEF857DFD06F0DEFCF6FD26" box="[695,771,644,670]" name="China" pageId="1" pageNumber="532">China</collectingCountry>
:
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,
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,
<collectingMunicipality id="6BE4ACC3FFEF857DFB84F0DEFA6DFD27" box="[1077,1432,644,671]" pageId="1" pageNumber="532">Wumengshan Natural Reserve</collectingMunicipality>
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<collectingDate id="EFC5E991FFEF857DFCE2F0F2FC23FD7A" box="[851,982,680,706]" pageId="1" pageNumber="532" value="2022-07-20">20.vii.2022</collectingDate>
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leg.,
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(
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.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEF857DFF76F0AAFE6EFBBE" pageId="1" pageNumber="532" type="diagnosis">
<paragraph id="8B8036B9FFEF857DFF76F0AAFE6EFBBE" blockId="1.[151,1437,536,2039]" pageId="1" pageNumber="532">
<emphasis id="B94BEAABFFEF857DFF76F0AAFEB7FCB2" bold="true" box="[199,322,752,778]" pageId="1" pageNumber="532">Diagnosis.</emphasis>
<emphasis id="B94BEAABFFEF857DFEF7F0AAFE7DFCB2" box="[326,392,752,778]" italics="true" pageId="1" pageNumber="532">Male.</emphasis>
Body length approximately
<quantity id="4CC79B5CFFEF857DFD71F0AAFCE2FCB3" box="[704,791,752,779]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="532" unit="mm" value="1.4">1.4 mm</quantity>
. Terminal four antennomeres greatly enlarged and forming distinct club, occupying approximately 6.2/10 of antennal length. Aedeagus with compressor plate broad and plate-like in ventral view, obliquely connected to median lobe in lateral view; apical projection of median lobe broad and protruding in ventral view, curved dorsally in lateral view; median lobe with pair of membranous, lateral projections apically rounded, and transversely rhomboidal plate on ventral wall; broad and elongate parameres narrowing from bases toward apices, each with two long setae at apex and two similar long setae at approximately apical 1/3.
<emphasis id="B94BEAABFFEF857DFF26F192FF1EFC5A" box="[151,235,968,994]" italics="true" pageId="1" pageNumber="532">Female</emphasis>
. External morphology similar to male, with antennomeres of clubs slightly shorter; spermatheca in lateral view transversely oval.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEF857EFF76F64AFBD7F970" lastPageId="2" lastPageNumber="533" pageId="1" pageNumber="532" type="description">
<paragraph id="8B8036B9FFEF857DFF76F64AFCC8FBCB" blockId="1.[151,1437,536,2039]" pageId="1" pageNumber="532">
<emphasis id="B94BEAABFFEF857DFF76F64AFEACFB92" bold="true" box="[199,345,1040,1066]" pageId="1" pageNumber="532">Description.</emphasis>
<emphasis id="B94BEAABFFEF857DFEEEF64AFE6CFB92" box="[351,409,1040,1066]" italics="true" pageId="1" pageNumber="532">Male</emphasis>
. Body (
<figureCitation id="13042A3CFFEF857DFE40F64AFDB2FB92" box="[497,583,1040,1066]" captionStart="FIGURE 1" captionStartId="2.[151,250,1316,1340]" captionTargetBox="[180,1409,189,1287]" captionTargetId="figure-20@2.[177,1411,179,1289]" captionTargetPageId="2" captionText="FIGURE 1. Euconnus (s. str.) wumengshanus sp. nov. (AE. Male. F. Female). A. Dorsal habitus. BE. Aedeagus, lateral (B, D) and ventral (C, E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A; 0.1 mm in BE; 0.05 mm in F." pageId="1" pageNumber="532">Fig. 1A</figureCitation>
) length
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; body uniformly reddish-brown, mouthparts and tarsi paler in color. Posterior margin of vertex, sides of head, and sides and disc of pronotum with scattered thick bristles, with long setae on body surface. Dorsum of body finely punctate.
</paragraph>
<paragraph id="8B8036B9FFEF857DFF76F626FD13FAF3" blockId="1.[151,1437,536,2039]" pageId="1" pageNumber="532">
Head roundly rhomboidal, broadest at eyes, length
<quantity id="4CC79B5CFFEF857DFC93F626FC79FB2F" box="[802,908,1148,1175]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="532" unit="mm" value="0.32">0.32 mm</quantity>
, width
<quantity id="4CC79B5CFFEF857DFC56F626FBA7FB2F" box="[999,1106,1148,1175]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="532" unit="mm" value="0.3">0.30 mm</quantity>
; vertex and frons confluent, convex; supraantennal tubercles barely prominent; eyes large, strongly convex and coarsely faceted. Punctures on vertex and frons fine; setae long, dense, suberect to erect, additionally tempora and vertex with long bristles directed posteriorly. Antenna moderately short, length
<quantity id="4CC79B5CFFEF857DFD47F6B2FCAEFABB" box="[758,859,1256,1283]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="1" pageNumber="532" unit="mm" value="0.63">0.63 mm</quantity>
, club
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; antennomeres 1 and 2 elongate, 37 compact, each short and distinctly transverse, 810 greatly enlarged, conical, 11 roundly conical, distinctly longer than 10, with sharp constriction at point of basal 2/5.
</paragraph>
<paragraph id="8B8036B9FFEF857DFF76F70FFAB0FA0E" blockId="1.[151,1437,536,2039]" pageId="1" pageNumber="532">
Pronotum in dorsal view subtrapezoidal, broadest at base and strongly narrowing anteriorly, length
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, width
<quantity id="4CC79B5CFFEF857DFF6FF722FEB0FA2B" box="[222,325,1400,1427]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="532" unit="mm" value="0.43">0.43 mm</quantity>
; lateral antebasal pits small but distinct, asetose, transverse antebasal groove shallow. Punctures on pronotal disc fine; setae moderately fine and obscured by dense, long and thick bristles especially on sides.
</paragraph>
<paragraph id="8B8036B9FFEF857DFF76F79AFEE5F9FE" blockId="1.[151,1437,536,2039]" pageId="1" pageNumber="532">
Elytra suboval and slightly flattened, broadest approximately at middle, length
<quantity id="4CC79B5CFFEF857DFB9CF79AFB64FA63" box="[1069,1169,1472,1499]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="532" unit="mm" value="0.82">0.82 mm</quantity>
, width
<quantity id="4CC79B5CFFEF857DFB51F79AFAB0FA63" box="[1248,1349,1472,1499]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="1" pageNumber="532" unit="mm" value="0.66">0.66 mm</quantity>
, length/ width 1.24; basal impressions shallow but distinct, humeral calli elongate; apices of elytra separately rounded. Punctures on elytral disc fine and shallow; setae long, sparse and strongly erect. Metathoracic wings fully developed, functional.
</paragraph>
<paragraph id="8B8036B9FFEF857DFF76F40BFDA0F9D2" blockId="1.[151,1437,536,2039]" box="[199,597,1616,1643]" pageId="1" pageNumber="532">Legs long and slender; unmodified.</paragraph>
<paragraph id="8B8036B9FFEF857DFF76F42EFC80F8DE" blockId="1.[151,1437,536,2039]" pageId="1" pageNumber="532">
Aedeagus (
<figureCitation id="13042A3CFFEF857DFEF6F42EFE4AF936" box="[327,447,1652,1678]" captionStart="FIGURE 1" captionStartId="2.[151,250,1316,1340]" captionTargetBox="[180,1409,189,1287]" captionTargetId="figure-20@2.[177,1411,179,1289]" captionTargetPageId="2" captionText="FIGURE 1. Euconnus (s. str.) wumengshanus sp. nov. (AE. Male. F. Female). A. Dorsal habitus. BE. Aedeagus, lateral (B, D) and ventral (C, E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A; 0.1 mm in BE; 0.05 mm in F." pageId="1" pageNumber="532">Fig. 1DE</figureCitation>
) moderately stout, dorso-ventrally almost symmetric, length
<quantity id="4CC79B5CFFEF857DFBCAF42EFB16F937" box="[1147,1251,1652,1679]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="532" unit="mm" value="0.22">0.22 mm</quantity>
, in ventral view median lobe with abruptly delimited and long, broad apical projection greatly curved dorsally; compressor plate broad and plate-like in dorso-ventral view, subquadrate with narrowed apical margin, obliquely connected to median lobe in lateral view; lateral projections broad and membranous, curved dorsally, with round apices; ventral wall of median lobe with large, transversely rhomboidal plate, its apical margin with two admesal acute projection; parameres broad and elongate, narrowing from bases toward apices, each with two long setae at apex, two similar long setae at apical 2/5 and several short and thin setae at middle.
</paragraph>
<paragraph id="8B8036B9FFEF857DFF76F52AFD5BF84F" blockId="1.[151,1437,536,2039]" pageId="1" pageNumber="532">
<emphasis id="B94BEAABFFEF857DFF76F52AFEEEF832" box="[199,283,1904,1930]" italics="true" pageId="1" pageNumber="532">Female</emphasis>
. Similar to male in habitus and external morphological features. Antennal clubs (
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) slightly shorter than male. Spermatheca (
<figureCitation id="13042A3CFFEF857DFDB5F5CEFDA0F816" box="[516,597,1940,1966]" captionStart="FIGURE 1" captionStartId="2.[151,250,1316,1340]" captionTargetBox="[180,1409,189,1287]" captionTargetId="figure-20@2.[177,1411,179,1289]" captionTargetPageId="2" captionText="FIGURE 1. Euconnus (s. str.) wumengshanus sp. nov. (AE. Male. F. Female). A. Dorsal habitus. BE. Aedeagus, lateral (B, D) and ventral (C, E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A; 0.1 mm in BE; 0.05 mm in F." pageId="1" pageNumber="532">Fig. 1F</figureCitation>
) in lateral view transversely oval. Measurements (as for male): body length
<quantity id="4CC79B5CFFEF857DFF26F5E2FF09F86B" box="[151,252,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="532" unit="mm" value="1.4">1.40 mm</quantity>
, length/width of head 0.34/
<quantity id="4CC79B5CFFEF857DFD9DF5E2FD64F86B" box="[556,657,1976,2003]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="532" unit="mm" value="0.32">0.32 mm</quantity>
, pronotum 0.41/
<quantity id="4CC79B5CFFEF857DFCF6F5E2FC59F86B" box="[839,940,1976,2003]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="532" unit="mm" value="0.43">0.43 mm</quantity>
, elytra 0.81/
<quantity id="4CC79B5CFFEF857DFB84F5E2FB6FF86B" box="[1077,1178,1976,2003]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="532" unit="mm" value="0.67">0.67 mm</quantity>
, length of antenna
<quantity id="4CC79B5CFFEF857DFADAF5E2FF30F84F" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="1" pageNumber="532" unit="mm" value="0.61">0.61 mm</quantity>
, longest diameter of spermatheca
<quantity id="4CC79B5CFFEF857DFDF2F586FD5FF84F" box="[579,682,2012,2039]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="532" unit="mm" value="0.09">0.09 mm</quantity>
.
</paragraph>
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<paragraph id="8B8036B9FFEC857EFF26F77EFB50FAE7" blockId="2.[151,1437,1316,1376]" pageId="2" pageNumber="533">
<emphasis id="B94BEAABFFEC857EFF26F77EFEE6FA84" bold="true" box="[151,275,1316,1340]" pageId="2" pageNumber="533">FIGURE 1.</emphasis>
<taxonomicName id="4C3F4D3AFFEC857EFEABF77FFD9AFA84" authorityName="Yin &amp; Zhou" authorityYear="2025" box="[282,623,1316,1340]" class="Insecta" family="Staphylinidae" genus="Euconnus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="wumengshanus" status="sp. nov.">
<emphasis id="B94BEAABFFEC857EFEABF77FFE8BFA84" box="[282,382,1317,1340]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
(
<emphasis id="B94BEAABFFEC857EFE3CF77FFE62FA84" box="[397,407,1317,1340]" italics="true" pageId="2" pageNumber="533">s</emphasis>
.
<emphasis id="B94BEAABFFEC857EFE15F77FFE4AFA84" box="[420,447,1317,1340]" italics="true" pageId="2" pageNumber="533">str</emphasis>
.)
<emphasis id="B94BEAABFFEC857EFE65F77EFD9AFA84" box="[468,623,1316,1340]" italics="true" pageId="2" pageNumber="533">wumengshanus</emphasis>
</taxonomicName>
<emphasis id="B94BEAABFFEC857EFDC4F77EFD33FA84" bold="true" box="[629,710,1316,1340]" pageId="2" pageNumber="533">
<taxonomicNameLabel id="A27857D0FFEC857EFDC4F77EFD33FA84" box="[629,710,1316,1340]" pageId="2" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(AE. Male. F. Female).
<emphasis id="B94BEAABFFEC857EFC7AF77EFC11FA84" bold="true" box="[971,996,1316,1340]" pageId="2" pageNumber="533">A.</emphasis>
Dorsal habitus.
<emphasis id="B94BEAABFFEC857EFB3BF77EFB35FA84" bold="true" box="[1162,1216,1316,1340]" pageId="2" pageNumber="533">BE.</emphasis>
Aedeagus, lateral (B, D) and ventral (C, E). F. Spermatheca, lateral. Scale bars: 0.5 mm in A; 0.1 mm in BE; 0.05 mm in F.
</paragraph>
</caption>
<paragraph id="8B8036B9FFEC857EFF76F7E8FBD7F970" blockId="2.[151,1437,1458,2025]" pageId="2" pageNumber="533">
<emphasis id="B94BEAABFFEC857EFF76F7E8FE45FA74" bold="true" box="[199,432,1458,1484]" pageId="2" pageNumber="533">Comparative notes.</emphasis>
This is a typical member of the former subgenus
<taxonomicName id="4C3F4D3AFFEC857EFC6DF7E8FB35FA74" authority="Thomson" authorityName="Thomson" box="[988,1216,1458,1484]" pageId="2" pageNumber="533" rank="subGenus" subGenus="Napochus">
<emphasis id="B94BEAABFFEC857EFC6DF7E8FBB9FA74" box="[988,1100,1458,1484]" italics="true" pageId="2" pageNumber="533">Napochus</emphasis>
Thomson
</taxonomicName>
, which has recently been treated as a junior synonym of the nominotypical subgenus (
<bibRefCitation id="EFAE4B48FFEC857EFCC6F78CFBBFFA48" author="Jaloszynski, P." box="[887,1098,1494,1520]" pageId="2" pageNumber="533" pagination="255 - 270" refId="ref5126" refString="Jaloszynski, P. (2021 a) Taxonomy of ' Euconnus complex'. Part XXIII. Status of Napochus Thomson, Pycnophus Casey, and Filonapochus Franz revisited (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5026 (2), 255 - 270. https: // doi. org / 10.11646 / zootaxa. 5026.2.6" type="journal article" year="2021">Jałoszyński 2021a</bibRefCitation>
).
<taxonomicName id="4C3F4D3AFFEC857EFBECF78CFB38FA48" box="[1117,1229,1494,1520]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="540" phylum="Arthropoda" rank="subGenus" subGenus="Napochus">
<emphasis id="B94BEAABFFEC857EFBECF78CFB38FA48" box="[1117,1229,1494,1520]" italics="true" pageId="2" pageNumber="533">Napochus</emphasis>
</taxonomicName>
represents a fairly diverse group within
<taxonomicName id="4C3F4D3AFFEC857EFE35F7A1FE07F9AC" box="[388,498,1531,1556]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABFFEC857EFE35F7A1FE07F9AC" box="[388,498,1531,1556]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
</taxonomicName>
, and has been the subject of recent taxonomic efforts for the Australian (
<bibRefCitation id="EFAE4B48FFEC857EFAA8F7A0FF26F981" author="Jaloszynski, P." pageId="2" pageNumber="533" pagination="1 - 24" refId="ref5020" refString="Jaloszynski, P. (2015) Taxonomy of ' Euconnus complex'. Part III. Morphology of Euconnus subgenus Napochus and revision of the Australian species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3925 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 3925.1.1" type="journal article" year="2015">Jałoszyński 2015</bibRefCitation>
), European (
<bibRefCitation id="EFAE4B48FFEC857EFEEFF444FDF3F980" author="Orousset, J." box="[350,518,1566,1592]" pageId="2" pageNumber="533" pagination="176 - 201" refId="ref5421" refString="Orousset, J. (2021) Les especes du genre Euconnus Thomson, 1859, sous-genre Napochus Thomson, 1859, de la faune de France (Coleoptera, Staphylinidae, Scydmaeninae). Le Coleopteriste, 23 (3), 176 - 201. [2020]" type="journal article" year="2021">Orousset 2021</bibRefCitation>
), and North American (
<bibRefCitation id="EFAE4B48FFEC857EFCB8F444FC14F980" author="Stephan, K. &amp; Chandler, D. S. &amp; O'Keefe, S. T. &amp; Riley, E. G." box="[777,993,1566,1593]" pageId="2" pageNumber="533" pagination="1 - 319" refId="ref5643" refString="Stephan, K., Chandler, D. S., O'Keefe, S. T. &amp; Riley, E. G. (2021) A revision of the North American species of Euconnus (Napochus) Thomson, north of Mexico (Coleoptera: Staphylinidae: Scydmaeninae). Memoirs of the American Entomological Society, 51, 1 - 319." type="journal article" year="2021">
Stephan
<emphasis id="B94BEAABFFEC857EFCDAF445FC6EF980" box="[875,923,1566,1592]" italics="true" pageId="2" pageNumber="533">et al</emphasis>
. 2021
</bibRefCitation>
) faunas. In
<collectingCountry id="F3287629FFEC857EFBD2F444FB5CF980" box="[1123,1193,1566,1592]" name="China" pageId="2" pageNumber="533">China</collectingCountry>
, only a few
<taxonomicName id="4C3F4D3AFFEC857EFA9FF445FF1DF9E4" pageId="2" pageNumber="533">
<emphasis id="B94BEAABFFEC857EFA9FF445FA69F980" box="[1326,1436,1567,1592]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
species
</taxonomicName>
have been properly assigned to subgenera (Li, L.-Z.
<emphasis id="B94BEAABFFEC857EFC80F419FC94F9E4" box="[817,865,1602,1628]" italics="true" pageId="2" pageNumber="533">et al</emphasis>
. 2019;
<bibRefCitation id="EFAE4B48FFEC857EFC03F418FB8EF9E4" author="Yin, Z. - W. &amp; Zhou, D. - Y." box="[946,1147,1602,1628]" pageId="2" pageNumber="533" pagination="582 - 592" refId="ref5768" refString="Yin, Z. - W. &amp; Zhou, D. - Y. (2024) Chenzhilinus, a new subgenus of Euconnus Thomson (Coleoptera: Staphylinidae: Scydmaeninae) from southern China. Zootaxa, 5541 (4), 582 - 592. https: // doi. org / 10.11646 / zootaxa. 5541.4.9" type="journal article" year="2024">Yin &amp; Zhou 2024</bibRefCitation>
). Nevertheless, no species is known to possess an aedeagus that bears even a slight resemblance to that of
<taxonomicName id="4C3F4D3AFFEC857EFBA4F43DFB2AF938" authorityName="Yin &amp; Zhou" authorityYear="2025" box="[1045,1247,1638,1665]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="wumengshanus" status="sp. nov.">
<emphasis id="B94BEAABFFEC857EFBA4F43DFBD3F938" box="[1045,1062,1639,1664]" italics="true" pageId="2" pageNumber="533">E</emphasis>
.
<emphasis id="B94BEAABFFEC857EFB85F43CFB2AF938" box="[1076,1247,1638,1664]" italics="true" pageId="2" pageNumber="533">wumengshanus</emphasis>
</taxonomicName>
<emphasis id="B94BEAABFFEC857EFB57F43CFAB5F938" bold="true" box="[1254,1344,1638,1664]" pageId="2" pageNumber="533">
<taxonomicNameLabel id="A27857D0FFEC857EFB57F43CFAB5F938" box="[1254,1344,1638,1664]" pageId="2" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Prior to this study, only one
<taxonomicName id="4C3F4D3AFFEC857EFEC8F4D1FDB5F91C" box="[377,576,1674,1700]" pageId="2" pageNumber="533">
<emphasis id="B94BEAABFFEC857EFEC8F4D1FE12F91C" box="[377,487,1675,1700]" italics="true" pageId="2" pageNumber="533">Euconnus</emphasis>
species
</taxonomicName>
had been documented from
<collectingRegion id="49FBF85BFFEC857EFC31F4D1FC23F91D" box="[896,982,1675,1701]" country="China" name="Yunnan" pageId="2" pageNumber="533">Yunnan</collectingRegion>
(Mount Tangli) (Li, Q.-Q. &amp; Yin 2021), which is characterized by a modified male head and a stout, transverse aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEC857EFF76F488FD64F955" box="[199,657,1746,1773]" pageId="2" pageNumber="533" type="distribution">
<paragraph id="8B8036B9FFEC857EFF76F488FD64F955" blockId="2.[151,1437,1458,2025]" box="[199,657,1746,1773]" pageId="2" pageNumber="533">
<emphasis id="B94BEAABFFEC857EFF76F488FE94F954" bold="true" box="[199,353,1746,1772]" pageId="2" pageNumber="533">Distribution.</emphasis>
Southwest
<collectingCountry id="F3287629FFEC857EFE52F488FDDFF954" box="[483,554,1746,1772]" name="China" pageId="2" pageNumber="533">China</collectingCountry>
:
<collectingRegion id="49FBF85BFFEC857EFD85F489FD78F955" box="[564,653,1747,1773]" country="China" name="Yunnan" pageId="2" pageNumber="533">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEC857EFF76F4ACFF0CF88D" pageId="2" pageNumber="533" type="etymology">
<paragraph id="8B8036B9FFEC857EFF76F4ACFF0CF88D" blockId="2.[151,1437,1458,2025]" pageId="2" pageNumber="533">
<emphasis id="B94BEAABFFEC857EFF76F4ACFEBBF8A8" bold="true" box="[199,334,1782,1808]" pageId="2" pageNumber="533">Etymology.</emphasis>
The species name is a toponym derived from its
<typeStatus id="5484881BFFEC857EFC1DF4ADFC29F8A9" box="[940,988,1783,1809]" pageId="2" pageNumber="533">type</typeStatus>
locality, Wumenshan National Nature Reserve.
</paragraph>
</subSubSection>
<subSubSection id="C3256532FFEC857EFF76F564FDE0F850" pageId="2" pageNumber="533" type="discussion">
<paragraph id="8B8036B9FFEC857EFF76F564FDE0F850" blockId="2.[151,1437,1458,2025]" pageId="2" pageNumber="533">
<emphasis id="B94BEAABFFEC857EFF76F564FECEF8E0" bold="true" box="[199,315,1854,1880]" pageId="2" pageNumber="533">Remarks.</emphasis>
The association of the female specimen with the male
<typeStatus id="5484881BFFEC857EFC7DF564FBDBF8E0" box="[972,1070,1854,1880]" pageId="2" pageNumber="533" type="holotype">holotype</typeStatus>
was based on the similarity in morphological features and proportions of body segments. However, the collection site of the female specimen is situated approximately
<quantity id="4CC79B5CFFEC857EFE73F5DCFDF8F818" box="[450,525,1926,1953]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="533" unit="km" value="50.0">50 km</quantity>
southeast of Laziping, where the male was collected. Given this geographical separation, the possibility that the female represents a separate species cannot be excluded. Thus, this female has been omitted from the type series.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,333 @@
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<treatment id="039D5914FF91FFA7FF4DFDC1FC73FD7F" LSID="urn:lsid:plazi:treatment:039D5914FF91FFA7FF4DFDC1FC73FD7F" httpUri="http://treatment.plazi.org/id/039D5914FF91FFA7FF4DFDC1FC73FD7F" lastPageId="8" lastPageNumber="108" pageId="6" pageNumber="106">
<subSubSection id="C32EBB89FF91FFA9FF4DFDC1FE2DFDB9" box="[151,501,552,579]" pageId="6" pageNumber="106" type="nomenclature">
<paragraph id="8B8BE802FF91FFA9FF4DFDC1FE2DFDB9" blockId="6.[151,532,552,616]" box="[151,501,552,579]" pageId="6" pageNumber="106">
<heading id="D0C35F6EFF91FFA9FF4DFDC1FE2DFDB9" bold="true" box="[151,501,552,579]" fontSize="11" level="1" pageId="6" pageNumber="106" reason="1">
<emphasis id="B9403410FF91FFA9FF4DFDC1FE2DFDB9" bold="true" box="[151,501,552,579]" pageId="6" pageNumber="106">
<taxonomicName id="4C349381FF91FFA9FF4DFDC1FE48FDB9" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[151,400,552,579]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="106" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF91FFA9FF4DFDC1FE48FDB9" bold="true" box="[151,400,552,579]" italics="true" pageId="6" pageNumber="106">Mallinella moncheri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273896BFF91FFA9FE4DFDC0FE2DFDB9" box="[407,501,553,579]" pageId="6" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF91FFA9FF4DFDA4FDCCFD9D" box="[151,532,589,616]" pageId="6" pageNumber="106" type="description">
<paragraph id="8B8BE802FF91FFA9FF4DFDA4FDCCFD9D" blockId="6.[151,532,552,616]" box="[151,532,589,616]" pageId="6" pageNumber="106">
<figureCitation id="130FF487FF91FFA9FF4DFDA4FF05FD9D" box="[151,221,589,615]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Figs 3</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FF30FDA4FEE5FD9D" box="[234,317,589,615]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="6" pageNumber="106">6C, 6H</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FE93FDA4FE44FD92" box="[329,412,589,616]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">7C, 7H</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FE72FDA4FE13FD9D" box="[424,459,589,615]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="6" pageNumber="106">8C</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FE0DFDA4FDCCFD9D" box="[471,532,589,615]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="6" pageNumber="106">9EF</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF91FFA9FF4DFD7FFB4AFCB8" pageId="6" pageNumber="106" type="materials_examined">
<paragraph id="8B8BE802FF91FFA9FF4DFD7FFD2EFD03" blockId="6.[151,1437,662,2003]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF4DFD7FFE91FD4A" bold="true" box="[151,329,662,688]" pageId="6" pageNumber="106">Type material.</emphasis>
<materialsCitation id="3B5CE25FFF91FFA9FE8FFD7FFBFAFD2F" collectingDate="2023-08-23" country="Sri Lanka" elevation="152" latitude="6.0944448" location="Kottawa Forest" longLatPrecision="21" longitude="80.32361" municipality="Galle District" pageId="6" pageNumber="106" specimenCount="1" specimenCount-male="1" stateProvince="Southern" typeStatus="holotype">
<emphasis id="B9403410FF91FFA9FE8FFD7FFC4AFD4A" bold="true" box="[341,914,662,692]" pageId="6" pageNumber="106">
<typeStatus id="548F56A0FF91FFA9FE8FFD7FFE1AFD4A" box="[341,450,662,688]" pageId="6" pageNumber="106" type="holotype">Holotype</typeStatus>
<specimenCount id="9D32238BFF91FFA9FE17FD71FE3CFD4E" box="[461,484,664,692]" count="1" pageId="6" pageNumber="106" type="male"></specimenCount>
:
<collectingCountry id="F323A892FF91FFA9FE2DFD7FFD42FD4A" box="[503,666,662,688]" name="Sri Lanka" pageId="6" pageNumber="106">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF91FFA9FD71FD7FFC4AFD4A" bold="true" box="[683,914,662,689]" italics="true" pageId="6" pageNumber="106">
<collectingRegion id="49F026E0FF91FFA9FD71FD7FFC53FD4A" box="[683,907,662,689]" country="Sri Lanka" name="Southern" pageId="6" pageNumber="106">Southern Province</collectingRegion>
:
</emphasis>
</emphasis>
<collectingMunicipality id="6BEF7278FF91FFA9FC47FD7FFBE5FD4A" box="[925,1085,662,688]" pageId="6" pageNumber="106">Galle District</collectingMunicipality>
,
<location id="8EEBBED9FF91FFA9FB96FD7EFB27FD4B" LSID="urn:lsid:plazi:treatment:039D5914FF91FFA7FF4DFDC1FC73FD7F:8EEBBED9FF91FFA9FB96FD7EFB27FD4B" box="[1100,1279,663,689]" country="Sri Lanka" latitude="6.0944448" longLatPrecision="21" longitude="80.32361" municipality="Galle District" name="Kottawa Forest" pageId="6" pageNumber="106" stateProvince="Southern">Kottawa Forest</location>
,
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<geoCoordinate id="EE008EC5FF91FFA9FF4DFD53FEFBFD2F" box="[151,291,698,725]" degrees="80" direction="east" minutes="19" orientation="longitude" pageId="6" pageNumber="106" precision="15" seconds="25" value="80.32361">80°1925E</geoCoordinate>
,
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<elevation id="00190F31FF91FFA9FEF4FD53FEAEFD2F" box="[302,374,698,725]" metricMagnitude="2" metricUnit="m" metricValue="1.52" pageId="6" pageNumber="106" unit="m" value="152.0">152 m</elevation>
</quantity>
,
<date id="FF8ACEC2FF91FFA9FE5BFD52FDECFD2F" box="[385,564,699,725]" pageId="6" pageNumber="106" value="2023-08-23">
<collectingDate id="EFCE372AFF91FFA9FE5BFD52FDECFD2F" box="[385,564,699,725]" pageId="6" pageNumber="106" value="2023-08-23">23 August 2023</collectingDate>
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, leg. NG. Dayananda
<emphasis id="B9403410FF91FFA9FCFCFD52FC8EFD2F" box="[806,854,699,725]" italics="true" pageId="6" pageNumber="106">et al</emphasis>
. (IFS_ZOD_231)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF91FFA9FBF1FD52FD2BFD03" collectingDate="2023-08-23" country="Sri Lanka" elevation="152" latitude="6.0944448" location="Benjamin" longLatPrecision="21" longitude="80.32361" municipality="Galle District" pageId="6" pageNumber="106" specimenCount="1" specimenCount-female="1" stateProvince="Paratype" typeStatus="paratype">
<emphasis id="B9403410FF91FFA9FBF1FD52FB79FD2F" bold="true" box="[1067,1185,699,725]" pageId="6" pageNumber="106">
<typeStatus id="548F56A0FF91FFA9FBF1FD52FB44FD2F" box="[1067,1180,699,725]" pageId="6" pageNumber="106" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D32238BFF91FFA9FB7DFD52FB12FD2F" box="[1191,1226,699,725]" count="1" pageId="6" pageNumber="106" type="female">1♀</specimenCount>
, leg. SP.
<location id="8EEBBED9FF91FFA9FAF5FD52FA44FD2F" LSID="urn:lsid:plazi:treatment:039D5914FF91FFA7FF4DFDC1FC73FD7F:8EEBBED9FF91FFA9FAF5FD52FA44FD2F" box="[1327,1436,699,725]" country="Sri Lanka" name="Benjamin" pageId="6" pageNumber="106" stateProvince="Paratype">Benjamin</location>
<emphasis id="B9403410FF91FFA9FF4DFD09FF10FD03" box="[151,200,735,761]" italics="true" pageId="6" pageNumber="106">et al</emphasis>
. with same data as for holotype (IFS_ZOD_103)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BE802FF91FFA9FF1DFCEAFB4AFCB8" blockId="6.[151,1437,662,2003]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFCEAFDDCFCE7" bold="true" box="[199,516,771,797]" pageId="6" pageNumber="106">Other material examined.</emphasis>
<materialsCitation id="3B5CE25FFF91FFA9FDD5FCEAFB56FCB8" collectingDate="2023-08-23" country="Sri Lanka" elevation="121" latitude="6.058889" location="Hiyare Forest" longLatPrecision="21" longitude="80.32083" municipality="Galle District" pageId="6" pageNumber="106" specimenCount="2" specimenCount-female="2" stateProvince="Southern">
<emphasis id="B9403410FF91FFA9FDD5FCEAFC7EFCE4" bold="true" box="[527,934,771,798]" pageId="6" pageNumber="106">
<collectingCountry id="F323A892FF91FFA9FDD5FCEAFD6AFCE7" box="[527,690,771,797]" name="Sri Lanka" pageId="6" pageNumber="106">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF91FFA9FD19FCEAFC7EFCE4" bold="true" box="[707,934,771,798]" italics="true" pageId="6" pageNumber="106">
<collectingRegion id="49F026E0FF91FFA9FD19FCEAFC47FCE4" box="[707,927,771,798]" country="Sri Lanka" name="Southern" pageId="6" pageNumber="106">Southern province</collectingRegion>
:
</emphasis>
</emphasis>
<collectingMunicipality id="6BEF7278FF91FFA9FC6BFCEDFB88FCE4" box="[945,1104,772,798]" pageId="6" pageNumber="106">Galle District</collectingMunicipality>
,
<location id="8EEBBED9FF91FFA9FB85FCEDFAD8FCE4" LSID="urn:lsid:plazi:treatment:039D5914FF91FFA7FF4DFDC1FC73FD7F:8EEBBED9FF91FFA9FB85FCEDFAD8FCE4" box="[1119,1280,772,798]" country="Sri Lanka" latitude="6.058889" longLatPrecision="21" longitude="80.32083" municipality="Galle District" name="Hiyare Forest" pageId="6" pageNumber="106" stateProvince="Southern">Hiyare Forest</location>
,
<geoCoordinate id="EE008EC5FF91FFA9FAD5FCEDFA44FCE4" box="[1295,1436,772,798]" degrees="06" direction="north" minutes="03" orientation="latitude" pageId="6" pageNumber="106" precision="15" seconds="32" value="6.058889">06°0332N</geoCoordinate>
<geoCoordinate id="EE008EC5FF91FFA9FF4DFCC1FEFCFCB9" box="[151,292,808,835]" degrees="80" direction="east" minutes="19" orientation="longitude" pageId="6" pageNumber="106" precision="15" seconds="15" value="80.32083">80°1915E</geoCoordinate>
,
<quantity id="4CCC45E7FF91FFA9FEF5FCC1FEAAFCB8" box="[303,370,808,834]" metricMagnitude="2" metricUnit="m" metricValue="1.21" pageId="6" pageNumber="106" unit="m" value="121.0">
<elevation id="00190F31FF91FFA9FEF5FCC1FEAAFCB8" box="[303,370,808,834]" metricMagnitude="2" metricUnit="m" metricValue="1.21" pageId="6" pageNumber="106" unit="m" value="121.0">121m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF91FFA9FEA7FCC1FE7AFCB8" box="[381,418,808,834]" count="2" pageId="6" pageNumber="106" type="female">2♀</specimenCount>
,
<date id="FF8ACEC2FF91FFA9FE74FCC1FDBAFCB8" box="[430,610,808,834]" pageId="6" pageNumber="106" value="2023-08-23">
<collectingDate id="EFCE372AFF91FFA9FE74FCC1FDBAFCB8" box="[430,610,808,834]" pageId="6" pageNumber="106" value="2023-08-23">23 August 2023</collectingDate>
</date>
, leg. NG. Dayananda
<emphasis id="B9403410FF91FFA9FC83FCC0FC52FCB8" box="[857,906,808,834]" italics="true" pageId="6" pageNumber="106">et al</emphasis>
. (IFS_ZOD_232, 233)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF91FFA9FF1DFCA4FAE3FC9D" box="[199,1339,845,871]" pageId="6" pageNumber="106" type="etymology">
<paragraph id="8B8BE802FF91FFA9FF1DFCA4FAE3FC9D" blockId="6.[151,1437,662,2003]" box="[199,1339,845,871]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFCA4FE96FC9D" bold="true" box="[199,334,845,871]" pageId="6" pageNumber="106">Etymology.</emphasis>
The new species is named after the chocolate Mon Cheri. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF91FFA9FF1DFC98FEFFFB4A" pageId="6" pageNumber="106" type="diagnosis">
<paragraph id="8B8BE802FF91FFA9FF1DFC98FEFFFB4A" blockId="6.[151,1437,662,2003]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFC98FE9AFC71" bold="true" box="[199,322,881,907]" pageId="6" pageNumber="106">Diagnosis.</emphasis>
<taxonomicName id="4C349381FF91FFA9FE9DFC98FDF2FC71" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[327,554,881,907]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="106" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF91FFA9FE9DFC98FDF2FC71" box="[327,554,881,907]" italics="true" pageId="6" pageNumber="106">Mallinella moncheri</emphasis>
</taxonomicName>
<emphasis id="B9403410FF91FFA9FDF5FC98FD4FFC71" bold="true" box="[559,663,881,907]" pageId="6" pageNumber="106">
<taxonomicNameLabel id="A273896BFF91FFA9FDF5FC98FD5FFC71" box="[559,647,881,907]" pageId="6" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
s
</emphasis>
hare features such as triangular lateral border of the epigyne, V-shaped epigynal plate with the members of the annulipes-group. Within the annulipes-group, males of
<taxonomicName id="4C349381FF91FFA9FB6AFC7EFAE6FC4A" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[1200,1342,918,944]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="106" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF91FFA9FB6AFC7EFAE6FC4A" box="[1200,1342,918,944]" italics="true" pageId="6" pageNumber="106">M. moncheri</emphasis>
</taxonomicName>
<emphasis id="B9403410FF91FFA9FA9EFC7FFA44FC4A" bold="true" box="[1348,1436,918,944]" pageId="6" pageNumber="106">
<taxonomicNameLabel id="A273896BFF91FFA9FA9EFC7FFA44FC4A" box="[1348,1436,918,944]" pageId="6" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are most similar to the males of
<taxonomicName id="4C349381FF91FFA9FDD2FC52FD74FC2E" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[520,684,954,980]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="106" phylum="Arthropoda" rank="species" species="dhanahami" status="sp. nov.">
<emphasis id="B9403410FF91FFA9FDD2FC52FD74FC2E" box="[520,684,954,980]" italics="true" pageId="6" pageNumber="106">M. dhanahami</emphasis>
</taxonomicName>
<emphasis id="B9403410FF91FFA9FD6FFC53FCD7FC2E" bold="true" box="[693,783,954,980]" pageId="6" pageNumber="106">
<taxonomicNameLabel id="A273896BFF91FFA9FD6FFC53FCD7FC2E" box="[693,783,954,980]" pageId="6" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as both share digitiform RTA, and black colour bands on either side of the carapace of both male and female but can be separated from it by slender, unbranched embolus (bifurcated embolus in
<taxonomicName id="4C349381FF91FFA9FE41FBEDFDE6FBE7" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[411,574,1027,1053]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="106" phylum="Arthropoda" rank="species" species="dhanahami" status="sp. nov.">
<emphasis id="B9403410FF91FFA9FE41FBEDFDE6FBE7" box="[411,574,1027,1053]" italics="true" pageId="6" pageNumber="106">M. dhanahami</emphasis>
</taxonomicName>
<emphasis id="B9403410FF91FFA9FD9FFBEDFD46FBE4" bold="true" box="[581,670,1028,1054]" pageId="6" pageNumber="106">
<taxonomicNameLabel id="A273896BFF91FFA9FD9FFBEDFD46FBE4" box="[581,670,1028,1054]" pageId="6" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and diagonally aligned emobolic base (aligned longitudinally in
<taxonomicName id="4C349381FF91FFA9FAA4FBEDFECDFBB8" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="106" phylum="Arthropoda" rank="species" species="dhanahami" status="sp. nov.">
<emphasis id="B9403410FF91FFA9FAA4FBEDFECDFBB8" italics="true" pageId="6" pageNumber="106">M. dhanahami</emphasis>
</taxonomicName>
<emphasis id="B9403410FF91FFA9FEC1FBC1FEACFBB8" bold="true" box="[283,372,1064,1090]" pageId="6" pageNumber="106">
<taxonomicNameLabel id="A273896BFF91FFA9FEC1FBC1FEACFBB8" box="[283,372,1064,1090]" pageId="6" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) (
<figureCitation id="130FF487FF91FFA9FE56FBC1FE3FFBB8" box="[396,487,1064,1090]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="6" pageNumber="106">Figs 1G</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FE29FBC1FDC1FBB8" box="[499,537,1064,1090]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="6" pageNumber="106">8A</figureCitation>
;
<figureCitation id="130FF487FF91FFA9FDFEFBC1FD90FBB8" box="[548,584,1064,1090]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">3G</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FD89FBC1FDAFFBB8" box="[595,631,1064,1090]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="6" pageNumber="106">8C</figureCitation>
). Females are also most similar to the females of
<taxonomicName id="4C349381FF91FFA9FB40FBC0FAE5FBB8" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[1178,1341,1064,1090]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="106" phylum="Arthropoda" rank="species" species="dhanahami" status="sp. nov.">
<emphasis id="B9403410FF91FFA9FB40FBC0FAE5FBB8" box="[1178,1341,1064,1090]" italics="true" pageId="6" pageNumber="106">M. dhanahami</emphasis>
</taxonomicName>
<emphasis id="B9403410FF91FFA9FA99FBC1FA44FBB8" bold="true" box="[1347,1436,1064,1090]" pageId="6" pageNumber="106">
<taxonomicNameLabel id="A273896BFF91FFA9FA99FBC1FA44FBB8" box="[1347,1436,1064,1090]" pageId="6" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as both share V-shaped epigynal plate but can be separated from it by deeper anterior depression of the epigyne and triangular lateral borders of the epigyne (digitiform lateral borders in
<taxonomicName id="4C349381FF91FFA9FC77FB9BFB89FB71" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[941,1105,1137,1163]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="106" phylum="Arthropoda" rank="species" species="dhanahami" status="sp. nov.">
<emphasis id="B9403410FF91FFA9FC77FB9BFB89FB71" box="[941,1105,1137,1163]" italics="true" pageId="6" pageNumber="106">M. dhanahami</emphasis>
</taxonomicName>
<emphasis id="B9403410FF91FFA9FB80FB98FB6CFB71" bold="true" box="[1114,1204,1137,1163]" pageId="6" pageNumber="106">
<taxonomicNameLabel id="A273896BFF91FFA9FB80FB98FB6CFB71" box="[1114,1204,1137,1163]" pageId="6" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). (
<figureCitation id="130FF487FF91FFA9FB0FFB98FAEFFB71" box="[1237,1335,1137,1163]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="6" pageNumber="106">Figs. 1C</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FA9FFB98FA4FFB76" box="[1349,1431,1137,1164]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">7A, 7F</figureCitation>
;
<figureCitation id="130FF487FF91FFA9FF4DFB7FFF62FB4A" box="[151,186,1174,1200]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">3C</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FF1CFB7FFEC2FB4A" box="[198,282,1174,1200]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">7C, 7H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF91FFA7FF1DFB53FAF3FD9B" lastPageId="8" lastPageNumber="108" pageId="6" pageNumber="106" type="description">
<paragraph id="8B8BE802FF91FFA9FF1DFB53FE8FF92E" blockId="6.[151,1437,662,2003]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFB53FE81FB2E" bold="true" box="[199,345,1210,1236]" pageId="6" pageNumber="106">Description.</emphasis>
<emphasis id="B9403410FF91FFA9FE87FB53FE4FFB2E" box="[349,407,1210,1236]" italics="true" pageId="6" pageNumber="106">Male</emphasis>
in alcohol (
<typeStatus id="548F56A0FF91FFA9FDCEFB53FDA3FB2E" box="[532,635,1210,1236]" pageId="6" pageNumber="106" type="holotype">holotype</typeStatus>
;
<figureCitation id="130FF487FF91FFA9FD5EFB53FD01FB2E" box="[644,729,1210,1236]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Fig. 3A</figureCitation>
)). Body length 3.57; carapace 2.16 long, 1.49 wide; opisthosoma 1.41 long, 1.08 wide. Habitus as in
<figureCitation id="130FF487FF91FFA9FDC1FB36FDABFB03" box="[539,627,1246,1273]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="6" pageNumber="106">Fig. 5A</figureCitation>
. Carapace ovoid, smooth and shiny, orange-brown, with two longitudinal lateral dark bands (
<figureCitation id="130FF487FF91FFA9FEB4FAEAFE1FFAE7" box="[366,455,1283,1309]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Fig. 3A</figureCitation>
); fovea red-brown. Clypeus light brown, 0.44 high. Eye sizes and inter-distances: AME 0.17, ALE 0.10, PME 0.11, PLE 0.11; AMEAME 0.04, AMEPME 0.09, AMEALE 0.08, ALEALE 0.45, PME PME 0.08, PMEPLE 0.20, PLEPLE 0.60, ALEPLE 0.03. MOA 0.39 long, front width 0.30, back width 0.35. Chilum unipartite, trapezoid. Chelicerae 0.79 long, light brown, without teeth. Endites yellow. Labium triangular, yellow-brown, 0.20 long, 0.35 wide. Sternum yellow, furnished with sparse black setae, 0.89 long, 0.74 wide. Lateral margins of sternum with small semi-circular pits in front of each coxa of third and fourth pair of legs. Legs yellow-brown, except pale yellowish coxae, proximal tubercle on anterior femora. Measurement of palp and legs: pedipalp (right) 1.93 (0.70 + 0.33 + 0.35+0.55), I 4.62 (1. 25 + 0.52 + 1.20 + 0.75+ 0.90), II 4.35 (1.20 + 0.50 + 0.80 + 0.95 + 0.90), III 4.60 (1.20 + 0.50 + 0.90 + 1.15 + 0.85), IV 6.21 (1.55 + 0.51+ 1.35 + 1.85 + 0.95). Leg formula: 4132. Opisthosoma oval, covered with black short setae, Dorsum of opisthosoma dark sepia, with lanceolate scutum. Pattern on dorsum of opisthosoma with anterior paired patch followed by medially connected large transverse patches (
<figureCitation id="130FF487FF91FFA9FE68F97CFDD4F955" box="[434,524,1685,1711]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Fig. 3A</figureCitation>
). Posterior ventral spines cylindrical, with blunt apices (
<figureCitation id="130FF487FF91FFA9FB5FF97CFB04F955" box="[1157,1244,1685,1711]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Fig. 3E</figureCitation>
). Spinnerets pale yellow (
<figureCitation id="130FF487FF91FFA9FF29F950FE92F92E" box="[243,330,1721,1748]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF91FFA9FF1DF937FF00F829" blockId="6.[151,1437,662,2003]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DF937FF24F902" box="[199,252,1758,1784]" italics="true" pageId="6" pageNumber="106">Palp</emphasis>
(
<figureCitation id="130FF487FF91FFA9FED0F937FE5CF902" box="[266,388,1758,1784]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Figs 3FH</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FE57F937FE69F902" box="[397,433,1758,1784]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="6" pageNumber="106">8C</figureCitation>
). VTA moderately developed, subtriangular, with curved apex (
<figureCitation id="130FF487FF91FFA9FBB7F937FB1CF902" box="[1133,1220,1758,1784]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="6" pageNumber="106">Fig. 8C</figureCitation>
). RTA digitiform in retrolateral view, broad at base, gradually tapered towards acute apex (
<figureCitation id="130FF487FF91FFA9FC77F8EAFBDFF8E7" box="[941,1031,1795,1821]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Fig. 3H</figureCitation>
). Cymbial fold deep, more than half the length of the cymbium. Apicoprolateral process of TA with, sharply pointed, elongated, slender flange, directed posteromesad (
<figureCitation id="130FF487FF91FFA9FE9AF8A5FE45F89C" box="[320,413,1868,1894]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Figs 3G</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FE73F8A5FE15F89C" box="[425,461,1868,1894]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="6" pageNumber="106">8C</figureCitation>
). Tegulum with large, well developed tegular tubercle (
<figureCitation id="130FF487FF91FFA9FBE0F8A5FB4AF89C" box="[1082,1170,1868,1894]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="6" pageNumber="106">Fig. 8C</figureCitation>
). Basoprolateral fold of TA, subtriangular (
<figureCitation id="130FF487FF91FFA9FEB6F899FE1CF870" box="[364,452,1904,1930]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="6" pageNumber="106">Fig. 8C</figureCitation>
). Embolic base aligned diagonally; membranous area of embolic base narrow. Embolus slender, unbranched, elongated, with blunt apex (
<figureCitation id="130FF487FF91FFA9FD11F87CFCF2F855" box="[715,810,1941,1967]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="6" pageNumber="106">Figs 3G</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FCE2F87CFC84F855" box="[824,860,1941,1967]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="6" pageNumber="106">8C</figureCitation>
). Tegular spine absent. Conductor triangular, apex blunt.
</paragraph>
<caption id="DF4BB88AFF90FFA8FF4DF938FBFBF81F" ID-DOI="http://doi.org/10.5281/zenodo.14734666" ID-Zenodo-Dep="14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="7" pageNumber="107" startId="7.[151,250,1745,1769]" targetBox="[253,1335,186,1717]" targetPageId="7" targetType="figure">
<paragraph id="8B8BE802FF90FFA8FF4DF938FBFBF81F" blockId="7.[151,1437,1745,2021]" pageId="7" pageNumber="107">
<emphasis id="B9403410FF90FFA8FF4DF938FECBF913" bold="true" box="[151,275,1745,1769]" pageId="7" pageNumber="107">FIGURE 3.</emphasis>
<taxonomicName id="4C349381FF90FFA8FEC0F938FE33F913" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[282,491,1745,1769]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="107" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF90FFA8FEC0F938FE33F913" box="[282,491,1745,1769]" italics="true" pageId="7" pageNumber="107">Mallinella moncheri</emphasis>
</taxonomicName>
<emphasis id="B9403410FF90FFA8FE2BF938FD9AF913" bold="true" box="[497,578,1745,1769]" pageId="7" pageNumber="107">
<taxonomicNameLabel id="A273896BFF90FFA8FE2BF938FD9AF913" box="[497,578,1745,1769]" pageId="7" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8BE802FF9FFFA7FF1DFF7EFDA6FE0F" blockId="8.[151,1437,150,645]" pageId="8" pageNumber="108">
<emphasis id="B9403410FF9FFFA7FF1DFF7EFEC3FF4B" box="[199,283,151,177]" italics="true" pageId="8" pageNumber="108">Female</emphasis>
in alcohol (
<typeStatus id="548F56A0FF9FFFA7FE73FF7EFDD7FF4B" box="[425,527,151,177]" pageId="8" pageNumber="108" type="paratype">paratype</typeStatus>
;
<figureCitation id="130FF487FF9FFFA7FDC7FF7EFDAEFF4B" box="[541,630,151,177]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="8" pageNumber="108">Fig. 3B</figureCitation>
). Body length 4.51; carapace 2.41 long, 1.70 wide; opisthosoma 2.10 long, 1.62 wide. Habitus and details as in male except for the following (
<figureCitation id="130FF487FF9FFFA7FC3AFF52FBE3FF2F" box="[992,1083,186,213]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="8" pageNumber="108">Fig. 5B</figureCitation>
). Clypeus 0.46 high. Eye sizes and inter-distances: AME 0.15, ALE 0.10, PME 0.11, PLE 0.10; AMEAME 0.07, AMEPME 0.14, AMEALE 0.08, ALEALE 0.53, PMEPME 0.14, PMEPLE 0.18, PLEPLE 0.68, ALEPLE 0.06. MOA 0.42 long, front width 0.32, back width 0.35. Chelicerae 0.81 long, Labium 0.30 long, 0.44 wide. Sternum: 1.00 long, 0.96 wide. Measurements of palp and legs: pedipalp (right) 1.68 (0.63 + 0.27 + 0.29 + 0.49), I 6.00 (1.50 + 0.50 + 1.45 + 1.30 + 1.25), II 5.75 (1.35 + 0.45 + 1.30 + 1.45 + 1.20), III 5.80 (1.35 + 0.50+ 1.20 + 1.55 + 1.20), IV 7.85 (1.75 + 0.50 + 1.85 + 2.25 + 1.50). Dorsum of pear-shaped opisthosoma, covered with dense black setae. Dorsal scutum on opisthosoma absent (
<figureCitation id="130FF487FF9FFFA7FE72FE5EFDD9FE2B" box="[424,513,439,465]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="8" pageNumber="108">Fig. 3B</figureCitation>
). Pattern on dorsum of opisthosoma with a pair of anterior small oblique patches, followed by numerous pale spots (
<figureCitation id="130FF487FF9FFFA7FDC3FE32FDA8FE0F" box="[537,624,475,501]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="8" pageNumber="108">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF9FFFA7FF1DFE16FAF3FD9B" blockId="8.[151,1437,150,645]" pageId="8" pageNumber="108">
<emphasis id="B9403410FF9FFFA7FF1DFE16FEE9FDE3" box="[199,305,511,537]" italics="true" pageId="8" pageNumber="108">Genitalia</emphasis>
(
<figureCitation id="130FF487FF9FFFA7FE9EFE16FE1EFDE3" box="[324,454,511,537]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="8" pageNumber="108">Figs 3CD</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FE0DFE16FDF6FDE3" box="[471,558,511,537]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="8" pageNumber="108">7C, 7H</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FDE5FE16FD58FDE3" box="[575,640,511,537]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="8" pageNumber="108">9EF</figureCitation>
). Epigynal plate, V-shaped, with deep anterior median invagination. Lateral borders triangular, terminally blunt. Epigynal furrow highly sclerotized (
<figureCitation id="130FF487FF9FFFA7FBE8FDCAFB49FDC7" box="[1074,1169,547,573]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="8" pageNumber="108">Figs 3C</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FB45FDCAFB1AFDC7" box="[1183,1218,547,573]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="8" pageNumber="108">7C</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FB0BFDCAFB2CFDC7" box="[1233,1268,547,573]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="8" pageNumber="108">9E</figureCitation>
). Insemination ducts longer than spermathecal length. Spermathecae globular in shape. Fertilization ducts as in
<figureCitation id="130FF487FF9FFFA7FB14FDAEFAFFFD9B" box="[1230,1319,583,609]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="8" pageNumber="108">Fig. 7H</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9FFFA7FF1DFD83FC73FD7F" box="[199,939,618,645]" pageId="8" pageNumber="108" type="distribution">
<paragraph id="8B8BE802FF9FFFA7FF1DFD83FC73FD7F" blockId="8.[151,1437,150,645]" box="[199,939,618,645]" pageId="8" pageNumber="108">
<emphasis id="B9403410FF9FFFA7FF1DFD83FEB9FD7E" bold="true" box="[199,353,618,644]" pageId="8" pageNumber="108">Distribution.</emphasis>
Known only from the collecting localities (
<figureCitation id="130FF487FF9FFFA7FC90FD82FC46FD7F" box="[842,926,619,645]" captionStart="FIGURE 12" captionStartId="17.[151,250,1133,1157]" captionText="FIGURE 12. Distribution map of known Mallinella species of Sri Lanka." pageId="8" pageNumber="108">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,527 @@
<document id="2BEE3A38379606407CFBD0E6C335E40D" ID-DOI="10.11646/zootaxa.5570.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734655" ID-ZooBank="1995C18B-1993-4DC4-B8AB-E4891F148A1E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737736390657" checkinUser="plazi" docAuthor="Dayananda, Naruwan &amp; Benjamin, Suresh P." docDate="2025" docId="039D5914FF94FFA9FF4DF8E7FCB7FE2F" docLanguage="en" docName="zootaxa.5570.1.4.pdf" docOrigin="Zootaxa 5570 (1)" docSource="https://doi.org/10.11646/zootaxa.5570.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mallinella milkyway Dayananda &amp; Benjamin, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="106" masterDocId="FFA4216CFF97FFAFFFDAFFE9FFD8FFFA" masterDocTitle="Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka" masterLastPageNumber="118" masterPageNumber="100" pageNumber="103" updateTime="1737736825942" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="884E17C329252DD627397D3A32592139">Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka</mods:title>
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<mods:name id="64D735145C372EEBDDDBE38EFC018B17" type="personal">
<mods:role id="4775772C44524D362D53C89643D55434">
<mods:roleTerm id="7CDFF6DBE095BBA6A417E3A685D73645">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7E8605E16511F558C4C1F27C2636E10B">Dayananda, Naruwan</mods:namePart>
<mods:affiliation id="C19D651C5A46C8B309869D3C03216C29">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka.</mods:affiliation>
</mods:name>
<mods:name id="2B02A2AEBF7736CC68DA7DB1373EB1DA" type="personal">
<mods:role id="A5C8DE76FEF3E492A44CC4D06C066DA2">
<mods:roleTerm id="8280D69785BE508116B090DA0179835E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3590E8EADA29DCCB6B22458ABF00443A">Benjamin, Suresh P.</mods:namePart>
<mods:affiliation id="C326A98C1F580340A2831F3BB770E080">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="A79DC9EE0DB8B55A1169C8F1887B763E">text</mods:typeOfResource>
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<mods:titleInfo id="61C980EC358289A6CF43D452E02329A1">
<mods:title id="8A018FE66C0D5FCB3F4ED995A0E29CAE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EAA4A93B8FA4E4159EAE50DF8CD312C0">
<mods:date id="B745158086454CF8DCE54490E431B2E1">2025</mods:date>
<mods:detail id="80BC088ACA0B592E36A0937C4025CBEE" type="pubDate">
<mods:number id="489CFCA9A56953770A354DA7B64B2964">2025-01-16</mods:number>
</mods:detail>
<mods:detail id="50A7CEC2966573514D3239441AA972DB" type="volume">
<mods:number id="1F5753E8354CFC765B18D939E0A71999">5570</mods:number>
</mods:detail>
<mods:detail id="82CFCC55657C3C02E9C06DC2E9BDA8C9" type="issue">
<mods:number id="73BD6AF9091CA7954C142EE7E6E78634">1</mods:number>
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<mods:extent id="9A082407EAF3EFCCD0196FE0D00F529E" unit="page">
<mods:start id="DD5F182513B5CCEFF16A71B35B082212">100</mods:start>
<mods:end id="DBE63D2B4A69A9AF4DAF6A8249BDB5C4">118</mods:end>
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<mods:location id="59B4589F11D3153CC4A5AE65DDB5BA60">
<mods:url id="BBD4FB500EF9132EF1C6F5DCBD18CED9">https://doi.org/10.11646/zootaxa.5570.1.4</mods:url>
</mods:location>
<mods:classification id="69F2F0C86729F4FB39AD750F1BC0C11E">journal article</mods:classification>
<mods:identifier id="81F880674E2865E7F9B1D67E0915FF3E" type="DOI">10.11646/zootaxa.5570.1.4</mods:identifier>
<mods:identifier id="A132F170AF5DB7C0046932D759BDB4EE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A6D1FE56972F4BC57A0ACF0CC2546E6F" type="Zenodo-Dep">14734655</mods:identifier>
<mods:identifier id="3BD14704C3E5288937D3C0E85EC26340" type="ZooBank">1995C18B-1993-4DC4-B8AB-E4891F148A1E</mods:identifier>
</mods:mods>
<treatment id="039D5914FF94FFA9FF4DF8E7FCB7FE2F" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F" httpUri="http://treatment.plazi.org/id/039D5914FF94FFA9FF4DF8E7FCB7FE2F" lastPageId="6" lastPageNumber="106" pageId="3" pageNumber="103">
<subSubSection id="C32EBB89FF94FFACFF4DF8E7FE2BF8D3" box="[151,499,1806,1833]" pageId="3" pageNumber="103" type="nomenclature">
<paragraph id="8B8BE802FF94FFACFF4DF8E7FE2BF8D3" blockId="3.[151,537,1806,1869]" box="[151,499,1806,1833]" pageId="3" pageNumber="103">
<heading id="D0C35F6EFF94FFACFF4DF8E7FE2BF8D3" bold="true" box="[151,499,1806,1833]" fontSize="11" level="1" pageId="3" pageNumber="103" reason="1">
<emphasis id="B9403410FF94FFACFF4DF8E7FE2BF8D3" bold="true" box="[151,499,1806,1833]" pageId="3" pageNumber="103">
<taxonomicName id="4C349381FF94FFACFF4DF8E7FE56F8D3" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[151,398,1806,1833]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="milkyway" status="sp. nov.">
<emphasis id="B9403410FF94FFACFF4DF8E7FE56F8D3" bold="true" box="[151,398,1806,1833]" italics="true" pageId="3" pageNumber="103">Mallinella milkyway</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273896BFF94FFACFE4FF8E6FE2BF8D3" box="[405,499,1807,1833]" pageId="3" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF94FFACFF4DF8DAFDC1F8B7" box="[151,537,1843,1869]" pageId="3" pageNumber="103" type="description">
<paragraph id="8B8BE802FF94FFACFF4DF8DAFDC1F8B7" blockId="3.[151,537,1806,1869]" box="[151,537,1843,1869]" pageId="3" pageNumber="103">
<figureCitation id="130FF487FF94FFACFF4DF8DAFF06F8B7" box="[151,222,1843,1869]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="3" pageNumber="103">Figs 2</figureCitation>
,
<figureCitation id="130FF487FF94FFACFF30F8DAFEE5F8B7" box="[234,317,1843,1869]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="3" pageNumber="103">6B, 6G</figureCitation>
,
<figureCitation id="130FF487FF94FFACFE93F8DAFE44F8B7" box="[329,412,1843,1869]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="3" pageNumber="103">7B, 7G</figureCitation>
,
<figureCitation id="130FF487FF94FFACFE72F8DAFE13F8B7" box="[424,459,1843,1869]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="3" pageNumber="103">8B</figureCitation>
,
<figureCitation id="130FF487FF94FFACFE0DF8DAFDC1F8B7" box="[471,537,1843,1869]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="3" pageNumber="103">9CD</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF94FFAAFF4DF892FC03FD7F" lastPageId="5" lastPageNumber="105" pageId="3" pageNumber="103" type="materials_examined">
<paragraph id="8B8BE802FF94FFACFF4DF892FD25F827" blockId="3.[151,1436,1914,2013]" pageId="3" pageNumber="103">
<emphasis id="B9403410FF94FFACFF4DF892FE9FF86E" bold="true" box="[151,327,1914,1941]" pageId="3" pageNumber="103">Type material.</emphasis>
<materialsCitation id="3B5CE25FFF94FFACFE8AF892FB68F843" collectingDate="2010-02-02" collectorName="S. Batuwita" country="Sri Lanka" county="Kandy District" elevation="1360" latitude="7.361111" location="Knuckles Range" longLatPrecision="21" longitude="80.833336" municipality="Corbett's Gap" pageId="3" pageNumber="103" specimenCount="1" specimenCount-male="1" stateProvince="Central" typeStatus="holotype">
<emphasis id="B9403410FF94FFACFE8AF892FCB4F86F" bold="true" box="[336,876,1914,1944]" pageId="3" pageNumber="103">
<typeStatus id="548F56A0FF94FFACFE8AF892FE65F86F" box="[336,445,1915,1941]" pageId="3" pageNumber="103" type="holotype">Holotype</typeStatus>
<specimenCount id="9D32238BFF94FFACFE1CF895FE05F862" box="[454,477,1916,1944]" count="1" pageId="3" pageNumber="103" type="male"></specimenCount>
:
<collectingCountry id="F323A892FF94FFACFE37F893FD56F86E" box="[493,654,1914,1940]" name="Sri Lanka" pageId="3" pageNumber="103">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF94FFACFD46F893FCB4F86F" bold="true" box="[668,876,1914,1941]" italics="true" pageId="3" pageNumber="103">
<collectingRegion id="49F026E0FF94FFACFD46F893FCBDF86F" box="[668,869,1914,1941]" country="Sri Lanka" name="Central" pageId="3" pageNumber="103">Central Province</collectingRegion>
:
</emphasis>
</emphasis>
<collectingCounty id="62EA908EFF94FFACFCAFF892FBF8F86F" box="[885,1056,1915,1941]" pageId="3" pageNumber="103">Kandy District</collectingCounty>
,
<collectingMunicipality id="6BEF7278FF94FFACFBF7F892FB15F86F" box="[1069,1229,1915,1941]" pageId="3" pageNumber="103">Corbetts Gap</collectingMunicipality>
,
<location id="8EEBBED9FF94FFACFB01F892FA40F86F" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF94FFACFB01F892FA40F86F" box="[1243,1432,1915,1941]" country="Sri Lanka" county="Kandy District" latitude="7.361111" longLatPrecision="21" longitude="80.833336" municipality="Corbett's Gap" name="Knuckles Range" pageId="3" pageNumber="103" stateProvince="Central">Knuckles Range</location>
,
<geoCoordinate id="EE008EC5FF94FFACFF4DF876FEFCF843" box="[151,292,1951,1977]" degrees="07" direction="north" minutes="21" orientation="latitude" pageId="3" pageNumber="103" precision="15" seconds="40" value="7.361111">07°2140N</geoCoordinate>
<geoCoordinate id="EE008EC5FF94FFACFEF4F877FE62F843" box="[302,442,1950,1977]" degrees="80" direction="east" minutes="50" orientation="longitude" pageId="3" pageNumber="103" precision="15" seconds="00" value="80.833336">80°5000E</geoCoordinate>
,
<quantity id="4CCC45E7FF94FFACFE12F876FDF9F843" box="[456,545,1951,1977]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="3" pageNumber="103" unit="m" value="1360.0">
<elevation id="00190F31FF94FFACFE12F876FDF9F843" box="[456,545,1951,1977]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="3" pageNumber="103" unit="m" value="1360.0">1360 m</elevation>
</quantity>
,
<date id="FF8ACEC2FF94FFACFDF5F876FD37F843" box="[559,751,1951,1977]" pageId="3" pageNumber="103" value="2010-02-02">
<collectingDate id="EFCE372AFF94FFACFDF5F876FD37F843" box="[559,751,1951,1977]" pageId="3" pageNumber="103" value="2010-02-02">2 February 2010</collectingDate>
</date>
, leg.
<collectorName id="26C18DD4FF94FFACFCF4F876FC6BF843" box="[814,947,1951,1977]" pageId="3" pageNumber="103">S. Batuwita</collectorName>
<emphasis id="B9403410FF94FFACFC66F876FC37F843" box="[956,1007,1951,1977]" italics="true" pageId="3" pageNumber="103">et al</emphasis>
. (IFS_ZOD_36)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF94FFACFB66F876FD21F827" collectingDate="2010-02-02" collectorName="S. Batuwita" country="Sri Lanka" county="Kandy District" elevation="1360" latitude="7.361111" location="Knuckles Range" longLatPrecision="21" longitude="80.833336" municipality="Corbett's Gap" pageId="3" pageNumber="103" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Paratypes" typeStatus="paratype">
<emphasis id="B9403410FF94FFACFB66F876FAE5F843" bold="true" box="[1212,1341,1951,1977]" pageId="3" pageNumber="103">
<typeStatus id="548F56A0FF94FFACFB66F876FAE0F843" box="[1212,1336,1951,1977]" pageId="3" pageNumber="103" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D32238BFF94FFACFA9CF876FAB1F843" box="[1350,1385,1951,1977]" count="2" pageId="3" pageNumber="103" type="male">2♂</specimenCount>
<specimenCount id="9D32238BFF94FFACFAA8F876FA4DF843" box="[1394,1429,1951,1977]" count="1" pageId="3" pageNumber="103" type="female">1♀</specimenCount>
, with same data as for holotype (IFS_ZOD_34, 35, 37)
</materialsCitation>
.
</paragraph>
<caption id="DF4BB88AFF93FFABFF4DF938FC55F81F" ID-DOI="http://doi.org/10.5281/zenodo.14734664" ID-Zenodo-Dep="14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="4" pageNumber="104" startId="4.[151,250,1745,1769]" targetBox="[256,1330,187,1717]" targetPageId="4" targetType="figure">
<paragraph id="8B8BE802FF93FFABFF4DF938FC55F81F" blockId="4.[151,1437,1745,2021]" pageId="4" pageNumber="104">
<emphasis id="B9403410FF93FFABFF4DF938FECDF913" bold="true" box="[151,277,1745,1769]" pageId="4" pageNumber="104">FIGURE 2.</emphasis>
<taxonomicName id="4C349381FF93FFABFEC4F938FE28F913" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[286,496,1745,1769]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="104" phylum="Arthropoda" rank="species" species="milkyway" status="sp. nov.">
<emphasis id="B9403410FF93FFABFEC4F938FE28F913" box="[286,496,1745,1769]" italics="true" pageId="4" pageNumber="104">Mallinella milkyway</emphasis>
</taxonomicName>
<emphasis id="B9403410FF93FFABFE22F938FD93F913" bold="true" box="[504,587,1745,1769]" pageId="4" pageNumber="104">
<taxonomicNameLabel id="A273896BFF93FFABFE22F938FD93F913" box="[504,587,1745,1769]" pageId="4" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8BE802FF92FFAAFF1DFF7FFC03FD7F" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DFF7FFE27FF4A" bold="true" box="[199,511,150,176]" pageId="5" pageNumber="105">Other material examined.</emphasis>
<materialsCitation id="3B5CE25FFF92FFAAFDD2FF7FFE40FEBB" collectingDate="2010-08-24" collectingDateMax="2019-02-16" collectingDateMin="2010-08-24" collectorName="Benjamin &amp; Athukorala" country="Sri Lanka" elevation="180" latitude="7.859444" location="Matale District" longLatPrecision="21" longitude="80.67444" pageId="5" pageNumber="105" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Central">
<emphasis id="B9403410FF92FFAAFDD2FF7FFC5BFF4B" bold="true" box="[520,899,150,177]" pageId="5" pageNumber="105">
<collectingCountry id="F323A892FF92FFAAFDD2FF7FFD70FF4A" box="[520,680,150,176]" name="Sri Lanka" pageId="5" pageNumber="105">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF92FFAAFD6DFF7FFC5BFF4B" bold="true" box="[695,899,150,177]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFD6DFF7FFCA5FF4B" box="[695,893,150,177]" country="Sri Lanka" name="Central" pageId="5" pageNumber="105">Central province</collectingRegion>
:
</emphasis>
</emphasis>
<location id="8EEBBED9FF92FFAAFC56FF7EFBE2FF4B" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFC56FF7EFBE2FF4B" box="[908,1082,151,177]" country="Sri Lanka" latitude="7.859444" longLatPrecision="21" longitude="80.67444" name="Matale District" pageId="5" pageNumber="105" stateProvince="Central">Matale District</location>
, NIFS arboretum,
<geoCoordinate id="EE008EC5FF92FFAAFAD5FF7FFA45FF4B" box="[1295,1437,150,177]" degrees="07" direction="north" minutes="51" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="34" value="7.859444">07°5134N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFF4DFF52FEFCFF2F" box="[151,292,187,213]" degrees="80" direction="east" minutes="40" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="28" value="80.67444">80°4028E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFEF5FF52FEAFFF2F" box="[303,375,187,213]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="105" unit="m" value="180.0">
<elevation id="00190F31FF92FFAAFEF5FF52FEAFFF2F" box="[303,375,187,213]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="105" unit="m" value="180.0">180 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFE58FF52FE7EFF2F" box="[386,422,187,213]" count="1" pageId="5" pageNumber="105" type="male">1♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFE68FF52FDEEFF2F" box="[434,566,187,213]" pageId="5" pageNumber="105" value="2013-07-07">
<collectingDate id="EFCE372AFF92FFAAFE68FF52FDEEFF2F" box="[434,566,187,213]" pageId="5" pageNumber="105" value="2013-07-07">7 July 2013</collectingDate>
</date>
, leg. SP.
<collectorName id="26C18DD4FF92FFAAFD43FF52FCDEFF2F" box="[665,774,187,213]" pageId="5" pageNumber="105">Benjamin</collectorName>
<emphasis id="B9403410FF92FFAAFCD6FF52FCE5FF2F" box="[780,829,187,213]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_25);
<specimenCount id="9D32238BFF92FFAAFBDFFF52FBF2FF2F" box="[1029,1066,187,213]" count="3" pageId="5" pageNumber="105" type="male">3♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFBEFFF52FB30FF2F" box="[1077,1256,187,213]" pageId="5" pageNumber="105" value="2010-08-24">
<collectingDate id="EFCE372AFF92FFAAFBEFFF52FB30FF2F" box="[1077,1256,187,213]" pageId="5" pageNumber="105" value="2010-08-24">24 August 2010</collectingDate>
</date>
, NP.
<collectorName id="26C18DD4FF92FFAAFAC4FF52FA44FF2F" box="[1310,1436,187,213]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFF4DFF36FF13FF03" box="[151,203,223,249]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_41, 210, 211);
<specimenCount id="9D32238BFF92FFAAFDD6FF36FDE8FF03" box="[524,560,223,249]" count="3" pageId="5" pageNumber="105" type="female">3♀</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFDE5FF36FD27FF03" box="[575,767,223,249]" pageId="5" pageNumber="105" value="2016-01-20">
<collectingDate id="EFCE372AFF92FFAAFDE5FF36FD27FF03" box="[575,767,223,249]" pageId="5" pageNumber="105" value="2016-01-20">20 January 2016</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFCB4FF36FC34FF03" box="[878,1004,223,249]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFC2CFF36FBF1FF03" box="[1014,1065,223,249]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_132, 133, 134);
<specimenCount id="9D32238BFF92FFAAFAA3FF36FA44FF03" box="[1401,1436,223,249]" count="1" pageId="5" pageNumber="105" type="male">1♂</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFF1CFEEAFF31FEE7" box="[198,233,259,285]" count="1" pageId="5" pageNumber="105" type="female">1♀</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFF2DFEEAFE19FEE7" box="[247,449,259,285]" pageId="5" pageNumber="105" value="2019-02-16">
<collectingDate id="EFCE372AFF92FFAAFF2DFEEAFE19FEE7" box="[247,449,259,285]" pageId="5" pageNumber="105" value="2019-02-16">16 February 2019</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFDF3FEEAFD7FFEE7" box="[553,679,259,285]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFD74FEEAFD07FEE7" box="[686,735,259,285]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_182, 183);
<specimenCount id="9D32238BFF92FFAAFC34FEEAFBCAFEE7" box="[1006,1042,259,285]" count="1" pageId="5" pageNumber="105" type="male">1♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFBC4FEEAFB6EFEE7" box="[1054,1206,258,285]" pageId="5" pageNumber="105" value="2015-05-28">
<collectingDate id="EFCE372AFF92FFAAFBC4FEEAFB6EFEE7" box="[1054,1206,258,285]" pageId="5" pageNumber="105" value="2015-05-28">28 May 2015</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFAC4FEEAFA44FEE7" box="[1310,1436,259,285]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFF4DFECEFF12FEBB" box="[151,202,295,321]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_209)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF92FFAAFE7EFECFFBAEFE9F" collectingDate="2010-05-24" collectorName="Athukorala" country="Sri Lanka" elevation="30" latitude="8.170834" location="Wanatha Villu" longLatPrecision="21" longitude="79.875" municipality="Puttalam District" pageId="5" pageNumber="105" specimenCount="6" specimenCount-female="6" stateProvince="North Western">
<emphasis id="B9403410FF92FFAAFE7EFECFFD1FFEBB" bold="true" box="[420,711,294,321]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFE7EFECFFD18FEBB" box="[420,704,294,321]" country="Sri Lanka" name="North Western" pageId="5" pageNumber="105">North Western Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BEF7278FF92FFAAFD0AFECEFC4BFEBB" box="[720,915,295,321]" pageId="5" pageNumber="105">Puttalam District</collectingMunicipality>
,
<location id="8EEBBED9FF92FFAAFC45FECEFB9CFEBB" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFC45FECEFB9CFEBB" box="[927,1092,295,321]" country="Sri Lanka" latitude="8.170834" longLatPrecision="21" longitude="79.875" municipality="Puttalam District" name="Wanatha Villu" pageId="5" pageNumber="105" stateProvince="North Western">Wanatha Villu</location>
,
<geoCoordinate id="EE008EC5FF92FFAAFB8AFECFFB05FEBB" box="[1104,1245,294,321]" degrees="08" direction="north" minutes="10" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="15" value="8.170834">08°1015N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFB3CFECFFAABFEBB" box="[1254,1395,294,321]" degrees="79" direction="east" minutes="52" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="30" value="79.875">79°5230E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFA5AFECEFF77FE9F" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="105" unit="m" value="30.0">
<elevation id="00190F31FF92FFAAFA5AFECEFF77FE9F" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="105" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFF66FEA2FF38FE9F" box="[188,224,331,357]" count="6" pageId="5" pageNumber="105" type="female">6♀</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFF37FEA2FE5FFE9F" box="[237,391,331,357]" pageId="5" pageNumber="105" value="2010-05-24">
<collectingDate id="EFCE372AFF92FFAAFF37FEA2FE5FFE9F" box="[237,391,331,357]" pageId="5" pageNumber="105" value="2010-05-24">24 May 2010</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFE28FEA2FDA8FE9F" box="[498,624,331,357]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFDA2FEA2FD72FE9F" box="[632,682,331,357]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_23, 53, 142, 143, 144, 145)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF92FFAAFB5BFEA3FDDCFE57" collectingDate="2012-01-10" collectorName="Athukorala" country="Sri Lanka" elevation="56" latitude="8.8" location="Padaviya" longLatPrecision="21" longitude="80.75" municipality="Anuradhapura District" pageId="5" pageNumber="105" specimenCount="3" specimenCount-female="3" stateProvince="North Central">
<emphasis id="B9403410FF92FFAAFB5BFEA3FA44FE9F" bold="true" box="[1153,1436,330,357]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFB5BFEA3FA4DFE9F" box="[1153,1429,330,357]" country="Sri Lanka" name="North Central" pageId="5" pageNumber="105">North Central Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BEF7278FF92FFAAFF4DFE86FE42FE73" box="[151,410,367,393]" pageId="5" pageNumber="105">Anuradhapura District</collectingMunicipality>
,
<location id="8EEBBED9FF92FFAAFE70FE86FDCCFE73" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFE70FE86FDCCFE73" box="[426,532,367,393]" country="Sri Lanka" latitude="8.8" longLatPrecision="21" longitude="80.75" municipality="Anuradhapura District" name="Padaviya" pageId="5" pageNumber="105" stateProvince="North Central">Padaviya</location>
,
<geoCoordinate id="EE008EC5FF92FFAAFDFEFE86FD69FE73" box="[548,689,367,393]" degrees="08" direction="north" minutes="48" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="00" value="8.8">08°4800N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFD67FE87FC92FE73" box="[701,842,366,393]" degrees="80" direction="east" minutes="45" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="00" value="80.75">80°4500E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFC80FE87FC42FE73" box="[858,922,366,393]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="105" unit="m" value="56.0">
<elevation id="00190F31FF92FFAAFC80FE87FC42FE73" box="[858,922,366,393]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="105" unit="m" value="56.0">56 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFC71FE86FC17FE73" box="[939,975,367,393]" count="3" pageId="5" pageNumber="105" type="female">3♀</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFC3BFE86FB7FFE73" box="[993,1191,367,393]" pageId="5" pageNumber="105" value="2012-01-10">
<collectingDate id="EFCE372AFF92FFAAFC3BFE86FB7FFE73" box="[993,1191,367,393]" pageId="5" pageNumber="105" value="2012-01-10">10 January 2012</collectingDate>
</date>
, leg. NP.
<collectorName id="26C18DD4FF92FFAAFAC4FE86FA44FE73" box="[1310,1436,367,393]" pageId="5" pageNumber="105">Athukorala</collectorName>
<emphasis id="B9403410FF92FFAAFF4DFE7AFF17FE57" box="[151,207,403,429]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_158,159,160)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF92FFAAFDCEFE7AFA40FE2B" collectingDate="2023-09-26" collectorName="Dayananda" country="Sri Lanka" elevation="9" latitude="8.9366665" location="Beach Side" longLatPrecision="21" longitude="81.006386" municipality="Trincomalee District" pageId="5" pageNumber="105" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Eastern">
<emphasis id="B9403410FF92FFAAFDCEFE7AFD34FE57" bold="true" box="[532,748,403,429]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFDCEFE7AFD3DFE57" box="[532,741,403,429]" country="Sri Lanka" name="Eastern" pageId="5" pageNumber="105">Eastern Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BEF7278FF92FFAAFD23FE7AFC32FE57" box="[761,1002,403,429]" pageId="5" pageNumber="105">Trincomalee District</collectingMunicipality>
, Arisimale
<location id="8EEBBED9FF92FFAAFBAFFE7AFB25FE57" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFBAFFE7AFB25FE57" box="[1141,1277,403,429]" country="Sri Lanka" latitude="8.9366665" longLatPrecision="21" longitude="81.006386" municipality="Trincomalee District" name="Beach Side" pageId="5" pageNumber="105" stateProvince="Eastern">Beach Side</location>
,
<geoCoordinate id="EE008EC5FF92FFAAFAD5FE7BFA44FE57" box="[1295,1436,402,429]" degrees="08" direction="north" minutes="56" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="12" value="8.9366665">08°5612N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFF4DFE5EFEFBFE2B" box="[151,291,439,465]" degrees="81" direction="east" minutes="00" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="23" value="81.006386">81°0023E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFEE8FE5EFEBAFE2B" box="[306,354,439,465]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="105" unit="m" value="9.0">
<elevation id="00190F31FF92FFAAFEE8FE5EFEBAFE2B" box="[306,354,439,465]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="105" unit="m" value="9.0">9 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFEAAFE5EFE4BFE2B" box="[368,403,439,465]" count="2" pageId="5" pageNumber="105" type="female">2♀</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFE15FE5EFE2DFE2B" box="[463,501,439,465]" count="2" pageId="5" pageNumber="105" type="male">2♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFDD9FE5EFD3DFE2B" box="[515,741,439,465]" pageId="5" pageNumber="105" value="2023-09-26">
<collectingDate id="EFCE372AFF92FFAAFDD9FE5EFD3DFE2B" box="[515,741,439,465]" pageId="5" pageNumber="105" value="2023-09-26">26 September 2023</collectingDate>
</date>
, leg. NG.
<collectorName id="26C18DD4FF92FFAAFC85FE5EFC05FE2B" box="[863,989,439,465]" pageId="5" pageNumber="105">Dayananda</collectorName>
<emphasis id="B9403410FF92FFAAFC3CFE5EFBC2FE2B" box="[998,1050,439,465]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_205, 206, 207, 208)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF92FFAAFF4DFE33FC0EFD7F" collectingDate="2016-09-20" collectingDateMax="2023-10-04" collectingDateMin="2016-09-20" collectorName="Dayananda" country="Sri Lanka" elevation="4" latitude="9.62" location="Mandaitivu" longLatPrecision="21" longitude="79.97806" municipality="Jaffna District" pageId="5" pageNumber="105" specimenCount="16" specimenCount-female="5" specimenCount-male="11" stateProvince="Nothern Province">
<emphasis id="B9403410FF92FFAAFF4DFE33FEABFE0F" bold="true" box="[151,371,474,501]" italics="true" pageId="5" pageNumber="105">
<collectingRegion id="49F026E0FF92FFAAFF4DFE33FEB4FE0F" box="[151,364,474,501]" country="Sri Lanka" name="Nothern Province" pageId="5" pageNumber="105">Nothern Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BEF7278FF92FFAAFEA4FE32FDFCFE0F" box="[382,548,475,501]" pageId="5" pageNumber="105">Jaffna District</collectingMunicipality>
,
<location id="8EEBBED9FF92FFAAFDE9FE32FD6FFE0F" LSID="urn:lsid:plazi:treatment:039D5914FF94FFA9FF4DF8E7FCB7FE2F:8EEBBED9FF92FFAAFDE9FE32FD6FFE0F" box="[563,695,475,501]" country="Sri Lanka" latitude="9.62" longLatPrecision="21" longitude="79.97806" municipality="Jaffna District" name="Mandaitivu" pageId="5" pageNumber="105" stateProvince="Nothern Province">Mandaitivu</location>
,
<geoCoordinate id="EE008EC5FF92FFAAFD1CFE32FC8CFE0F" box="[710,852,475,501]" degrees="09" direction="north" minutes="37" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="30" value="9.625">09°3730N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFC85FE33FC34FE0F" box="[863,1004,474,501]" degrees="79" direction="east" minutes="59" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="50" value="79.997215">79°5950E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFC21FE32FBF4FE0F" box="[1019,1068,475,501]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="105" unit="m" value="4.0">
<elevation id="00190F31FF92FFAAFC21FE32FBF4FE0F" box="[1019,1068,475,501]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="105" unit="m" value="4.0">4 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFBE6FE32FB87FE0F" box="[1084,1119,475,501]" count="1" pageId="5" pageNumber="105" type="female">1♀</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFB44FE32FB1BFE0F" box="[1182,1219,475,501]" count="1" pageId="5" pageNumber="105" type="male">1♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFB09FE32FF0AFDE3" pageId="5" pageNumber="105" value="2016-09-20" valueMax="2016-09-22" valueMin="2016-09-20">
<collectingDate id="EFCE372AFF92FFAAFB09FE32FF0AFDE3" pageId="5" pageNumber="105" value="2016-09-20" valueMax="2016-09-22" valueMin="2016-09-20">2022 September 2016</collectingDate>
</date>
, leg. SP. Benjamin
<emphasis id="B9403410FF92FFAAFE76FE16FE05FDE3" box="[428,477,511,537]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_180, 181);
<specimenCount id="9D32238BFF92FFAAFD37FE16FCCAFDE3" box="[749,786,511,537]" count="4" pageId="5" pageNumber="105" type="male">4♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFCC4FE16FC06FDE3" box="[798,990,511,537]" pageId="5" pageNumber="105" value="2023-10-03">
<collectingDate id="EFCE372AFF92FFAAFCC4FE16FC06FDE3" box="[798,990,511,537]" pageId="5" pageNumber="105" value="2023-10-03">03 October 2023</collectingDate>
</date>
, NG. Dayananda
<emphasis id="B9403410FF92FFAAFB7FFE16FB0EFDE3" box="[1189,1238,511,537]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_224, 225, 226, 227); Chettypulam Road side,
<geoCoordinate id="EE008EC5FF92FFAAFDB1FDCAFD20FDC7" box="[619,760,547,573]" degrees="09" direction="north" minutes="37" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="48" value="9.63">09°3748N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFCD8FDCBFC56FDC7" box="[770,910,546,573]" degrees="79" direction="east" minutes="54" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="45" value="79.9125">79°5445E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFC46FDCAFC13FDC7" box="[924,971,547,573]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="105" unit="m" value="6.0">
<elevation id="00190F31FF92FFAAFC46FDCAFC13FDC7" box="[924,971,547,573]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="105" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFC03FDCAFC24FDC7" box="[985,1020,547,573]" count="1" pageId="5" pageNumber="105" type="female">1♀</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFBEDFDCAFB84FDC7" box="[1079,1116,547,573]" count="3" pageId="5" pageNumber="105" type="male">3♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFBB0FDCAFAF5FDC7" box="[1130,1325,547,573]" pageId="5" pageNumber="105" value="2023-10-04">
<collectingDate id="EFCE372AFF92FFAAFBB0FDCAFAF5FDC7" box="[1130,1325,547,573]" pageId="5" pageNumber="105" value="2023-10-04">04 October 2023</collectingDate>
</date>
, leg. NG. Dayananda
<emphasis id="B9403410FF92FFAAFEC1FDAEFE94FD9B" box="[283,332,583,609]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS_ZOD_220,221,222,223); Allaipiddy,
<geoCoordinate id="EE008EC5FF92FFAAFCECFDAEFC1CFD9B" box="[822,964,583,609]" degrees="09" direction="north" minutes="37" orientation="latitude" pageId="5" pageNumber="105" precision="15" seconds="12" value="9.62">09°3712N</geoCoordinate>
<geoCoordinate id="EE008EC5FF92FFAAFC13FDAFFB8EFD9B" box="[969,1110,582,609]" degrees="79" direction="east" minutes="58" orientation="longitude" pageId="5" pageNumber="105" precision="15" seconds="41" value="79.97806">79°5841E</geoCoordinate>
,
<quantity id="4CCC45E7FF92FFAAFBBBFDAEFB55FD9B" box="[1121,1165,583,609]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="105" unit="m" value="6.0">
<elevation id="00190F31FF92FFAAFBBBFDAEFB55FD9B" box="[1121,1165,583,609]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="105" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<specimenCount id="9D32238BFF92FFAAFB42FDAEFB63FD9B" box="[1176,1211,583,609]" count="3" pageId="5" pageNumber="105" type="female">3♀</specimenCount>
and
<specimenCount id="9D32238BFF92FFAAFB2AFDAEFACDFD9B" box="[1264,1301,583,609]" count="3" pageId="5" pageNumber="105" type="male">3♂</specimenCount>
,
<date id="FF8ACEC2FF92FFAAFAFAFDAEFF0AFD7F" pageId="5" pageNumber="105" value="2023-10-04">
<collectingDate id="EFCE372AFF92FFAAFAFAFDAEFF0AFD7F" pageId="5" pageNumber="105" value="2023-10-04">04 October 2023</collectingDate>
</date>
, leg. NG.
<collectorName id="26C18DD4FF92FFAAFE9EFD82FE1AFD7F" box="[324,450,619,645]" pageId="5" pageNumber="105">Dayananda</collectorName>
<emphasis id="B9403410FF92FFAAFE13FD82FE22FD7F" box="[457,506,619,645]" italics="true" pageId="5" pageNumber="105">et al</emphasis>
. (IFS ZOD_214, 215, 219, 216, 217, 218)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF92FFAAFF1DFD66FAE0FD53" box="[199,1336,655,681]" pageId="5" pageNumber="105" type="etymology">
<paragraph id="8B8BE802FF92FFAAFF1DFD66FAE0FD53" blockId="5.[151,1437,150,2013]" box="[199,1336,655,681]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DFD66FE96FD53" bold="true" box="[199,334,655,681]" pageId="5" pageNumber="105">Etymology.</emphasis>
The new species is named after the chocolate MilkyWay. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF92FFAAFF1DFD5BFCDFFC17" pageId="5" pageNumber="105" type="diagnosis">
<paragraph id="8B8BE802FF92FFAAFF1DFD5BFCDFFC17" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DFD5BFE9AFD36" bold="true" box="[199,322,690,716]" pageId="5" pageNumber="105">Diagnosis.</emphasis>
<taxonomicName id="4C349381FF92FFAAFE92FD5AFDF6FD37" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[328,558,691,717]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="milkyway" status="sp. nov.">
<emphasis id="B9403410FF92FFAAFE92FD5AFDF6FD37" box="[328,558,691,717]" italics="true" pageId="5" pageNumber="105">Mallinella milkyway</emphasis>
</taxonomicName>
<emphasis id="B9403410FF92FFAAFDEEFD5AFD47FD37" bold="true" box="[564,671,691,717]" pageId="5" pageNumber="105">
<taxonomicNameLabel id="A273896BFF92FFAAFDEEFD5AFD55FD37" box="[564,653,691,717]" pageId="5" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
s
</emphasis>
hare features such as U-shaped epigynal plate, tear-shaped PVS, and tremendously lengthened insemination ducts with the members of the redimita-group. Within the redimita-group, males of
<taxonomicName id="4C349381FF92FFAAFF21FD12FE52FCEF" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[251,394,763,789]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="milkyway" status="sp. nov.">
<emphasis id="B9403410FF92FFAAFF21FD12FE52FCEF" box="[251,394,763,789]" italics="true" pageId="5" pageNumber="105">M. milkyway</emphasis>
</taxonomicName>
<emphasis id="B9403410FF92FFAAFE55FD12FE30FCEF" bold="true" box="[399,488,763,789]" pageId="5" pageNumber="105">
<taxonomicNameLabel id="A273896BFF92FFAAFE55FD12FE30FCEF" box="[399,488,763,789]" pageId="5" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are most similar to the males of
<taxonomicName id="4C349381FF92FFAAFC97FD12FC17FCEF" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[845,975,763,789]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="redimita">
<emphasis id="B9403410FF92FFAAFC97FD12FC17FCEF" box="[845,975,763,789]" italics="true" pageId="5" pageNumber="105">M. redimita</emphasis>
</taxonomicName>
as both share flange-like RTA, bifurcated embolus, sharp apical ridge directed posteromesad and a triangular apico-retrolateral fold with a blunt apex but can be separated from it by the rami of the embolus with blunt apices (sharply pointed in
<taxonomicName id="4C349381FF92FFAAFBBEFCAAFB31FCA7" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[1124,1257,835,861]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="redimita">
<emphasis id="B9403410FF92FFAAFBBEFCAAFB31FCA7" box="[1124,1257,835,861]" italics="true" pageId="5" pageNumber="105">M. redimita</emphasis>
</taxonomicName>
) and lanceolate dorsal scutum (cf.
<bibRefCitation id="EFA595F3FF92FFAAFEBDFC8EFD5AFC7B" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[359,642,871,897]" pageId="5" pageNumber="105" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF92FFAAFDD2FC8EFD98FC7B" box="[520,576,871,897]" italics="true" pageId="5" pageNumber="105">et al.</emphasis>
2012
</bibRefCitation>
: figs. 706707 and
<figureCitation id="130FF487FF92FFAAFC87FC8EFBD5FC7B" box="[861,1037,871,897]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs. 2A, 2FH</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFBC2FC8EFBE4FC7B" box="[1048,1084,871,897]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">8B</figureCitation>
). Females are also most similar to the females of
<taxonomicName id="4C349381FF92FFAAFE86FC62FE3EFC5F" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[348,486,907,933]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="redimata">
<emphasis id="B9403410FF92FFAAFE86FC62FE3EFC5F" box="[348,486,907,933]" italics="true" pageId="5" pageNumber="105">M. redimata</emphasis>
</taxonomicName>
as both share U-shaped epigynal plate, and elongated insemination ducts but can be separated from it by globular spermathecae (truncated spermathecae in
<taxonomicName id="4C349381FF92FFAAFC60FC46FBE6FC33" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[954,1086,943,969]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="redimita">
<emphasis id="B9403410FF92FFAAFC60FC46FBE6FC33" box="[954,1086,943,969]" italics="true" pageId="5" pageNumber="105">M. redimita</emphasis>
</taxonomicName>
) (cf.
<bibRefCitation id="EFA595F3FF92FFAAFBA6FC46FA40FC33" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[1148,1432,943,969]" pageId="5" pageNumber="105" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF92FFAAFAC7FC46FA8DFC33" box="[1309,1365,943,969]" italics="true" pageId="5" pageNumber="105">et al.</emphasis>
2012
</bibRefCitation>
: figs. 694699, 701-703 and
<figureCitation id="130FF487FF92FFAAFE09FC3AFD8FFC17" box="[467,599,979,1005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs. 2CD</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFDB8FC3AFD70FC17" box="[610,680,979,1005]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="5" pageNumber="105">7BG</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFD69FC3AFD23FC17" box="[691,763,979,1005]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="5" pageNumber="105">9CD</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF92FFA9FF1DFC1FFB7CFE77" lastPageId="6" lastPageNumber="106" pageId="5" pageNumber="105" type="description">
<paragraph id="8B8BE802FF92FFAAFF1DFC1FFE70F9F3" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DFC1FFE81FBEA" bold="true" box="[199,345,1014,1040]" pageId="5" pageNumber="105">Description.</emphasis>
<emphasis id="B9403410FF92FFAAFE84FC1EFE40FBEB" box="[350,408,1015,1041]" italics="true" pageId="5" pageNumber="105">Male</emphasis>
in alcohol (
<typeStatus id="548F56A0FF92FFAAFDCCFC1EFDA5FBEB" box="[534,637,1015,1041]" pageId="5" pageNumber="105" type="holotype">holotype</typeStatus>
;
<figureCitation id="130FF487FF92FFAAFD5DFC1EFD06FBEB" box="[647,734,1015,1041]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2A</figureCitation>
). Body length 3.96; carapace 2.25 long, 1.76 wide; opisthosoma 1.71 long, 1.36 wide. Habitus as in
<figureCitation id="130FF487FF92FFAAFDF0FBF2FD5EFBCF" box="[554,646,1051,1077]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2A</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFD49FBF2FD3EFBCF" box="[659,742,1051,1077]" captionStart="FIGURE 11" captionStartId="16.[151,250,771,795]" captionTargetBox="[151,1436,181,747]" captionTargetId="figure-17@16.[151,1436,181,747]" captionTargetPageId="16" captionText="FIGURE 11. Photographs of live Mallinella milkyway sp. nov. from Sri Lanka, paratype male from Arisimale Beach Side (IFS_ZOD_207). Photographs by SP. Benjamin" figureDoi="http://doi.org/10.5281/zenodo.14734687" httpUri="https://zenodo.org/record/14734687/files/figure.png" pageId="5" pageNumber="105">11AB</figureCitation>
. Carapace ovoid, granulate, brown, slightly darker anteriorly; fovea red-brown (
<figureCitation id="130FF487FF92FFAAFEBEFBD6FE18FBA3" box="[356,448,1087,1113]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2A</figureCitation>
). Clypeus light brown, 0.42 high. Eye sizes and inter-distances: AME 0.13, ALE 0.12, PME 0.12, PLE 0.12; AMEAME 0.08, AMEPME 0.15, AMEALE 0.12, ALEALE 0.55, PMEPME 0.12, PMEPLE 0.20, PLEPLE 0.68, ALEPLE 0.06. MOA 0.41 long, front width 0.34, back width 0.38. Chilum unipartite, trapezoid. Chelicerae 0.78 long, brown, without teeth. Endites yellow-brown. Labium triangular, yellow-brown, 0.23 long, 0.38 wide. Sternum brown, with sparse black setae, 1.01 long, 0.93 wide. Lateral margins of sternum with small semi-circular pits in front of each coxa of third and fourth pair of legs (
<figureCitation id="130FF487FF92FFAAFB50FB1AFB3CFAF7" box="[1162,1252,1267,1293]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="5" pageNumber="105">Fig. 6G</figureCitation>
). Legs generally yellowish, except for brown femora, proximal tubercle on anterior femora (
<figureCitation id="130FF487FF92FFAAFC21FAFEFB8FFACB" box="[1019,1111,1303,1329]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="5" pageNumber="105">Fig. 6B</figureCitation>
). Measurements of palp and legs: pedipalp (right) 2.78 (0.97 + 0.45 + 0.47 + 0.89) I 6.70 (1.60 + 0.50 + 1.75 + 1.50+ 1.35), II 5.90 (1.45 + 0.45 + 1.30 + 1.40 + 1.30), III 6.25 (1.40 + 0.35 + 1.25 + 1.35 + 1.90), IV 8.17 (1.95 + 0.50+ 1.85 + 2.15 + 1.72). Leg formula: 4132. Opisthosoma oval, covered with black short setae, with lanceolate dorsal scutum. Dorsum of opisthosoma dark sepia. Pattern on dorsum of opisthosoma represented by a paired of large anterior patches followed by irregular arrangement of pale spots. Posterior ventral spines short, tear-shaped (
<figureCitation id="130FF487FF92FFAAFB6DFA22FAC9FA1F" box="[1207,1297,1483,1509]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2E</figureCitation>
). Spinnerets yellow-brown (
<figureCitation id="130FF487FF92FFAAFE9EFA06FE43F9F3" box="[324,411,1519,1545]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF92FFAAFF1DF9FAFAB2F91B" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DF9FAFF24F9D7" box="[199,252,1555,1581]" italics="true" pageId="5" pageNumber="105">Palp</emphasis>
(
<figureCitation id="130FF487FF92FFAAFED7F9FAFE52F9D7" box="[269,394,1555,1581]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2FH</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFE4DF9FAFE62F9D7" box="[407,442,1555,1581]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">8B</figureCitation>
). RTA flange-like in retrolateral view, broad at base, gradually tapered towards sharply pointed apex (
<figureCitation id="130FF487FF92FFAAFEEDF9DEFE4DF9AB" box="[311,405,1591,1617]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2G</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFE78F9DEFE1DF9AB" box="[418,453,1591,1617]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">8B</figureCitation>
). VTA subtriangular blunt apex. Tegular spine minute (
<figureCitation id="130FF487FF92FFAAFBEFF9DEFB55F9AB" box="[1077,1165,1591,1617]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">Fig. 8B</figureCitation>
). Cymbial fold shallow, less than half the length of cymbium (
<figureCitation id="130FF487FF92FFAAFD91F9B2FD7FF98F" box="[587,679,1627,1653]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2G</figureCitation>
). TA with sharply pointed apical ridge directed posteromesad with triangular apicoprotrolateral process having blunt apex, directed mesad (
<figureCitation id="130FF487FF92FFAAFC75F996FBDCF963" box="[943,1028,1663,1689]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">Fig. 8B</figureCitation>
). Embolic base aligned in retrolateral direction, anterior membranous part broad, triangular (
<figureCitation id="130FF487FF92FFAAFCDEF94AFCBBF947" box="[772,867,1699,1725]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2G</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFCAAF94AFC4BF947" box="[880,915,1699,1725]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">8B</figureCitation>
). Embolus blade-like, broadest medially, subterminally divided, both rami slender, with blunt tips (
<figureCitation id="130FF487FF92FFAAFD2FF92EFC95F91B" box="[757,845,1735,1761]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="5" pageNumber="105">Fig. 8B</figureCitation>
). Conductor beak-shaped, sharply pointed apex.
</paragraph>
<paragraph id="8B8BE802FF92FFA9FF1DF902FC75FF2F" blockId="5.[151,1437,150,2013]" lastBlockId="6.[151,1437,150,469]" lastPageId="6" lastPageNumber="106" pageId="5" pageNumber="105">
<emphasis id="B9403410FF92FFAAFF1DF902FEC3F8FF" box="[199,283,1771,1797]" italics="true" pageId="5" pageNumber="105">Female</emphasis>
(
<typeStatus id="548F56A0FF92FFAAFEF2F902FE56F8FF" box="[296,398,1771,1797]" pageId="5" pageNumber="105" type="paratype">paratype</typeStatus>
;
<figureCitation id="130FF487FF92FFAAFE4DF902FE35F8FF" box="[407,493,1771,1797]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Fig. 2B</figureCitation>
). Body length 4.98; carapace 2.24 long, 1.54 wide; opisthosoma 2.74 long, 1.91 wide. Habitus and details as in male except for the following (
<figureCitation id="130FF487FF92FFAAFCDAF8E6FC83F8D3" box="[768,859,1807,1833]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="5" pageNumber="105">Figs 2B</figureCitation>
,
<figureCitation id="130FF487FF92FFAAFCBCF8E6FC63F8D3" box="[870,955,1807,1833]" captionStart="FIGURE 10" captionStartId="15.[151,250,1923,1947]" captionTargetBox="[151,1435,181,1900]" captionTargetId="figure-25@15.[151,1435,181,1900]" captionTargetPageId="15" captionText="FIGURE 10. Photographs of live Mallinella spp. described from Sri Lanka. AB Mallinella milkyway sp. nov., paratype female from Arisimale Beach Side (IFS_ZOD_205). CE Mallinella dhanahami sp. nov., paratype female from Gilimale Forest Reserve (IFS_ZOD_235). Photographs by SP. Benjamin." figureDoi="http://doi.org/10.5281/zenodo.14734683" httpUri="https://zenodo.org/record/14734683/files/figure.png" pageId="5" pageNumber="105">10AB</figureCitation>
). Carapace with smooth tegument. Clypeus 0.50 high. Eye sizes and inter-distances: AME 0.10, ALE 0.09, PME 0.09, PLE 0.09; AMEAME 0.08, AMEPME 0.14, AMEALE 0.12, ALEALE 0.41, PMEPME 0.12, PMEPLE 0.16, PLEPLE 0.58, ALEPLE 0.05. MOA 0.36 long, front width 0.27, back width 0.31. Chelicerae 0.75 long. Labium 0.25 long, 0.39 wide. Sternum with shallow semi-circular pits, 0.94 long, 0.89 wide Legs with proximal tubercle on anterior femora. Measurements of palp and legs: palp (right) 1.94 (0.64 + 0.36 + 0.39 + 0.55), I 4.85 (1.25 + 0.50 + 1.10 + 1.15 + 0.85), II 4.75 (1.15 + 0.45 + 1.05 + 1.15 + 0.95), III 5.00 (1.25 + 0.45+ 1.00 + 1.35 + 0.95), IV 6.25 (1.50 + 0.55 + 1.80 + 1.25 + 1.15). Leg formula: 4132. Dorsum of opisthosoma without scutum (
<figureCitation id="130FF487FF91FFA9FC92FF52FC78FF2F" box="[840,928,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="6" pageNumber="106">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF91FFA9FF1DFF09FB5FFE9D" blockId="6.[151,1437,150,469]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFF09FEE9FF00" box="[199,305,224,250]" italics="true" pageId="6" pageNumber="106">Genitalia</emphasis>
(
<figureCitation id="130FF487FF91FFA9FEE4FF09FE62FF00" box="[318,442,224,250]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="6" pageNumber="106">Figs 2CD</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FE19FF09FDCBFF00" box="[451,531,224,250]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">7B, 7G</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FDC7FF09FDBDFF00" box="[541,613,224,250]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="6" pageNumber="106">9CD</figureCitation>
). Epigynal plate; narrow, U-shaped, prominent anterior median depression, weakly sclerotized. Lateral borders retracted (
<figureCitation id="130FF487FF91FFA9FD70FEEDFCD2FEE4" box="[682,778,260,286]" captionStart="FIGURE 2" captionStartId="4.[151,250,1745,1769]" captionTargetBox="[256,1330,187,1717]" captionTargetId="figure-17@4.[252,1336,181,1721]" captionTargetPageId="4" captionText="FIGURE 2. Mallinella milkyway sp. nov. A, EH holotype male (IFS_ZOD_36) and BE paratype female (IFS_ZOD_37). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734664" httpUri="https://zenodo.org/record/14734664/files/figure.png" pageId="6" pageNumber="106">Figs 2C</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FCC3FEECFCE4FEE5" box="[793,828,261,287]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">7B</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FC91FEEDFCB7FEE4" box="[843,879,260,286]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="6" pageNumber="106">9C</figureCitation>
). Insemination ducts elongated, slightly curving inwardly (
<figureCitation id="130FF487FF91FFA9FED1FEC0FEB2FEB9" box="[267,362,297,323]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">Figs 7G</figureCitation>
,
<figureCitation id="130FF487FF91FFA9FEA2FEC0FE46FEB9" box="[376,414,297,323]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="6" pageNumber="106">9D</figureCitation>
). Spermathecae globular in shape, slightly widened distally. Insemination ducts short and straight extend posteriorly from the copulatory openings. Fertilization ducts as in
<figureCitation id="130FF487FF91FFA9FBF0FEA4FB5BFE9D" box="[1066,1155,333,359]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="6" pageNumber="106">Fig. 7G</figureCitation>
.
</paragraph>
<paragraph id="8B8BE802FF91FFA9FF1DFE9BFB7CFE77" blockId="6.[151,1437,150,469]" box="[199,1188,370,397]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFE9BFEE7FE76" bold="true" box="[199,319,370,396]" pageId="6" pageNumber="106">Variation.</emphasis>
Male (n=22): body length 3.904.02. Female (n=21): body length 5.005.19.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF91FFA9FF1DFE7FFCB7FE2F" pageId="6" pageNumber="106" type="distribution">
<paragraph id="8B8BE802FF91FFA9FF1DFE7FFCB7FE2F" blockId="6.[151,1437,150,469]" pageId="6" pageNumber="106">
<emphasis id="B9403410FF91FFA9FF1DFE7FFEB9FE4A" bold="true" box="[199,353,406,432]" pageId="6" pageNumber="106">Distribution.</emphasis>
<collectingCountry id="F323A892FF91FFA9FEBCFE7FFE0BFE4A" box="[358,467,406,432]" name="Sri Lanka" pageId="6" pageNumber="106">Sri Lanka</collectingCountry>
(Matale District, Jaffna District, Anuradhapura District, Trincomalee District, Puttalam District, Kandy District) (
<figureCitation id="130FF487FF91FFA9FE6DFE52FDD3FE2F" box="[439,523,443,469]" captionStart="FIGURE 12" captionStartId="17.[151,250,1133,1157]" captionText="FIGURE 12. Distribution map of known Mallinella species of Sri Lanka." pageId="6" pageNumber="106">Fig. 12</figureCitation>
). Possibly also present in
<collectingCountry id="F323A892FF91FFA9FCF4FE52FCB3FE2F" box="[814,875,443,469]" name="India" pageId="6" pageNumber="106">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,338 @@
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<mods:title id="CA3B10AB4A7A86F4FA5F5CE2F2C43B93">Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka</mods:title>
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<subSubSection id="C32EBB89FF96FFAEFF4DF934FDD3F90D" box="[151,523,1757,1784]" pageId="1" pageNumber="101" type="nomenclature">
<paragraph id="8B8BE802FF96FFAEFF4DF934FDD3F90D" blockId="1.[151,621,1757,1820]" box="[151,523,1757,1784]" pageId="1" pageNumber="101">
<heading id="D0C35F6EFF96FFAEFF4DF934FDD3F90D" bold="true" box="[151,523,1757,1784]" fontSize="11" level="1" pageId="1" pageNumber="101" reason="1">
<emphasis id="B9403410FF96FFAEFF4DF934FDD3F90D" bold="true" box="[151,523,1757,1784]" pageId="1" pageNumber="101">
<taxonomicName id="4C349381FF96FFAEFF4DF934FE7FF902" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[151,423,1757,1784]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="101" phylum="Arthropoda" rank="species" species="dhanahami" status="sp. nov.">
<emphasis id="B9403410FF96FFAEFF4DF934FE7FF902" bold="true" box="[151,423,1757,1784]" italics="true" pageId="1" pageNumber="101">Mallinella dhanahami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273896BFF96FFAEFE74F934FDD3F90D" box="[430,523,1757,1783]" pageId="1" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF96FFAEFF4DF8E8FDB5F8E1" box="[151,621,1793,1820]" pageId="1" pageNumber="101" type="description">
<paragraph id="8B8BE802FF96FFAEFF4DF8E8FDB5F8E1" blockId="1.[151,621,1757,1820]" box="[151,621,1793,1820]" pageId="1" pageNumber="101">
<figureCitation id="130FF487FF96FFAEFF4DF8E8FF04F8E1" box="[151,220,1793,1819]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="1" pageNumber="101">Figs 1</figureCitation>
,
<figureCitation id="130FF487FF96FFAEFF30F8E8FEE1F8E1" box="[234,313,1793,1819]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="1" pageNumber="101">6A, 6F</figureCitation>
,
<figureCitation id="130FF487FF96FFAEFE9EF8EBFE4BF8E6" box="[324,403,1794,1820]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="1" pageNumber="101">7A, 7F</figureCitation>
,
<figureCitation id="130FF487FF96FFAEFE44F8E8FE1AF8E1" box="[414,450,1793,1819]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="1" pageNumber="101">8A</figureCitation>
,
<figureCitation id="130FF487FF96FFAEFE14F8E8FDCBF8E1" box="[462,531,1793,1819]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="1" pageNumber="101">9AB</figureCitation>
,
<figureCitation id="130FF487FF96FFAEFDC5F8E8FDB5F8E1" box="[543,621,1793,1819]" captionStart="FIGURE 10" captionStartId="15.[151,250,1923,1947]" captionTargetBox="[151,1435,181,1900]" captionTargetId="figure-25@15.[151,1435,181,1900]" captionTargetPageId="15" captionText="FIGURE 10. Photographs of live Mallinella spp. described from Sri Lanka. AB Mallinella milkyway sp. nov., paratype female from Arisimale Beach Side (IFS_ZOD_205). CE Mallinella dhanahami sp. nov., paratype female from Gilimale Forest Reserve (IFS_ZOD_235). Photographs by SP. Benjamin." figureDoi="http://doi.org/10.5281/zenodo.14734683" httpUri="https://zenodo.org/record/14734683/files/figure.png" pageId="1" pageNumber="101">10CE</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF96FFAEFF4DF8A0FD30F856" pageId="1" pageNumber="101" type="materials_examined">
<paragraph id="8B8BE802FF96FFAEFF4DF8A0FD30F856" blockId="1.[151,1436,1865,2035]" pageId="1" pageNumber="101">
<emphasis id="B9403410FF96FFAEFF4DF8A0FE93F899" bold="true" box="[151,331,1865,1891]" pageId="1" pageNumber="101">Type material.</emphasis>
<materialsCitation id="3B5CE25FFF96FFAEFE82F8A0FAC2F87D" collectingDate="2024-03-25" country="Sri Lanka" elevation="126" latitude="0.76416665" location="Gilimale Forest Reserve" longLatPrecision="21" longitude="0.42777777" municipality="Rathnapura District" pageId="1" pageNumber="101" specimenCount="1" specimenCount-male="1" stateProvince="Sabaragamuwa" typeStatus="holotype">
<emphasis id="B9403410FF96FFAEFE82F8A0FC3DF899" bold="true" box="[344,997,1865,1895]" pageId="1" pageNumber="101">
<typeStatus id="548F56A0FF96FFAEFE82F8A0FE1DF899" box="[344,453,1865,1891]" pageId="1" pageNumber="101" type="holotype">Holotype</typeStatus>
<specimenCount id="9D32238BFF96FFAEFE0BF8A2FE30F89D" box="[465,488,1867,1895]" count="1" pageId="1" pageNumber="101" type="male"></specimenCount>
:
<collectingCountry id="F323A892FF96FFAEFE27F8A0FD79F899" box="[509,673,1865,1891]" name="Sri Lanka" pageId="1" pageNumber="101">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF96FFAEFD69F8A0FC3DF899" bold="true" box="[691,997,1865,1892]" italics="true" pageId="1" pageNumber="101">
<collectingRegion id="49F026E0FF96FFAEFD69F8A0FC06F899" box="[691,990,1865,1892]" country="Sri Lanka" name="Sabaragamuwa" pageId="1" pageNumber="101">Sabaragamuwa Province</collectingRegion>
:
</emphasis>
</emphasis>
<collectingMunicipality id="6BEF7278FF96FFAEFC28F8A0FB0FF899" box="[1010,1239,1865,1891]" pageId="1" pageNumber="101">Rathnapura District</collectingMunicipality>
,
<location id="8EEBBED9FF96FFAEFB3DF8A0FF2CF872" LSID="urn:lsid:plazi:treatment:039D5914FF96FFACFF4DF934FC84F947:8EEBBED9FF96FFAEFB3DF8A0FF2CF872" country="Sri Lanka" latitude="0.76416665" longLatPrecision="21" longitude="0.42777777" municipality="Rathnapura District" name="Gilimale Forest Reserve" pageId="1" pageNumber="101" stateProvince="Sabaragamuwa">Gilimale Forest Reserve</location>
, 06
<geoCoordinate id="EE008EC5FF96FFAEFEFAF886FE54F870" box="[288,396,1903,1930]" degrees="0" direction="north" minutes="45" orientation="latitude" pageId="1" pageNumber="101" precision="15" seconds="51" value="0.76416665">⁰4551N</geoCoordinate>
80
<geoCoordinate id="EE008EC5FF96FFAEFE69F886FDC6F870" box="[435,542,1903,1930]" degrees="0" direction="east" minutes="25" orientation="longitude" pageId="1" pageNumber="101" precision="15" seconds="40" value="0.42777777">⁰2540E</geoCoordinate>
,
<quantity id="4CCC45E7FF96FFAEFDF6F884FDAFF872" box="[556,631,1901,1928]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="101" unit="m" value="126.0">
<elevation id="00190F31FF96FFAEFDF6F884FDAFF872" box="[556,631,1901,1928]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="101" unit="m" value="126.0">126 m</elevation>
</quantity>
,
<date id="FF8ACEC2FF96FFAEFD5FF884FCEFF87D" box="[645,823,1901,1928]" pageId="1" pageNumber="101" value="2024-03-25">
<collectingDate id="EFCE372AFF96FFAEFD5FF884FCEFF87D" box="[645,823,1901,1928]" pageId="1" pageNumber="101" value="2024-03-25">25 March 2024</collectingDate>
</date>
, leg. SP. Benjamin
<emphasis id="B9403410FF96FFAEFBC2F887FB8BF87D" box="[1048,1107,1901,1927]" italics="true" pageId="1" pageNumber="101">et al.</emphasis>
(IFS_ZOD_234)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF96FFAEFAFCF884FD3DF856" collectingDate="2024-03-25" country="Sri Lanka" elevation="126" latitude="0.76416665" location="Gilimale Forest Reserve" longLatPrecision="21" longitude="0.42777777" municipality="Rathnapura District" pageId="1" pageNumber="101" specimenCount="1" specimenCount-female="1" stateProvince="Paratype" typeStatus="paratype">
<emphasis id="B9403410FF96FFAEFAFCF884FA44F87D" bold="true" box="[1318,1436,1901,1927]" pageId="1" pageNumber="101">
<typeStatus id="548F56A0FF96FFAEFAFCF884FA4FF87D" box="[1318,1431,1901,1927]" pageId="1" pageNumber="101" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D32238BFF96FFAEFF4DF878FF62F851" box="[151,186,1937,1963]" count="1" pageId="1" pageNumber="101" type="female">1♀</specimenCount>
, with same data as for holotype (IFS_ZOD_235)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF96FFAEFF1DF85CFECCF809" pageId="1" pageNumber="101" type="etymology">
<paragraph id="8B8BE802FF96FFAEFF1DF85CFECCF809" blockId="1.[151,1436,1865,2035]" pageId="1" pageNumber="101">
<emphasis id="B9403410FF96FFAEFF1DF85CFE96F835" bold="true" box="[199,334,1973,1999]" pageId="1" pageNumber="101">Etymology.</emphasis>
This new species is named after the first authors grandmother, Dhanahami. Used as a noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF4BB88AFF95FFADFF4DF938FC55F81F" ID-DOI="http://doi.org/10.5281/zenodo.14734662" ID-Zenodo-Dep="14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="2" pageNumber="102" startId="2.[151,250,1745,1769]" targetBox="[255,1334,184,1715]" targetPageId="2" targetType="figure">
<paragraph id="8B8BE802FF95FFADFF4DF938FC55F81F" blockId="2.[151,1437,1745,2021]" pageId="2" pageNumber="102">
<emphasis id="B9403410FF95FFADFF4DF938FECAF913" bold="true" box="[151,274,1745,1769]" pageId="2" pageNumber="102">FIGURE 1.</emphasis>
<taxonomicName id="4C349381FF95FFADFEC2F938FE21F913" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[280,505,1745,1769]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="102" phylum="Arthropoda" rank="species" species="dhanahami" status="sp. nov.">
<emphasis id="B9403410FF95FFADFEC2F938FE21F913" box="[280,505,1745,1769]" italics="true" pageId="2" pageNumber="102">Mallinella dhanahami</emphasis>
</taxonomicName>
<emphasis id="B9403410FF95FFADFE25F938FD96F913" bold="true" box="[511,590,1745,1769]" pageId="2" pageNumber="102">
<taxonomicNameLabel id="A273896BFF95FFADFE25F938FD96F913" box="[511,590,1745,1769]" pageId="2" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C32EBB89FF94FFACFF1DFF7FFABBFE2B" pageId="3" pageNumber="103" type="diagnosis">
<paragraph id="8B8BE802FF94FFACFF1DFF7FFABBFE2B" blockId="3.[151,1437,150,1725]" pageId="3" pageNumber="103">
<emphasis id="B9403410FF94FFACFF1DFF7FFE9AFF4A" bold="true" box="[199,322,150,176]" pageId="3" pageNumber="103">Diagnosis.</emphasis>
<taxonomicName id="4C349381FF94FFACFE92FF7EFD99FF4B" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[328,577,151,177]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="dhanahami" status="sp. nov.">
<emphasis id="B9403410FF94FFACFE92FF7EFD99FF4B" box="[328,577,151,177]" italics="true" pageId="3" pageNumber="103">Mallinella dhanahami</emphasis>
</taxonomicName>
<emphasis id="B9403410FF94FFACFD92FF7EFD79FF4B" bold="true" box="[584,673,151,177]" pageId="3" pageNumber="103">
<taxonomicNameLabel id="A273896BFF94FFACFD92FF7EFD79FF4B" box="[584,673,151,177]" pageId="3" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share features such as inwardly projecting digitiform lateral borders of the epigyne, round proximal tubercle on anterior femora and pale oblong spots on the dorsum of opisthosoma with the members of the annulipes-group. Within the annulipes-group, males of
<taxonomicName id="4C349381FF94FFACFC04FF36FB59FF03" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[990,1153,223,249]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="dhanahami" status="sp. nov.">
<emphasis id="B9403410FF94FFACFC04FF36FB59FF03" box="[990,1153,223,249]" italics="true" pageId="3" pageNumber="103">M. dhanahami</emphasis>
</taxonomicName>
<emphasis id="B9403410FF94FFACFB52FF36FB3AFF03" bold="true" box="[1160,1250,223,249]" pageId="3" pageNumber="103">
<taxonomicNameLabel id="A273896BFF94FFACFB52FF36FB3AFF03" box="[1160,1250,223,249]" pageId="3" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are most similar to the males of
<taxonomicName id="4C349381FF94FFACFE93FEEAFE3DFEE7" baseAuthorityName="Barrion &amp; Litsinger" baseAuthorityYear="1992" box="[329,485,259,285]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="calilungae">
<emphasis id="B9403410FF94FFACFE93FEEAFE3DFEE7" box="[329,485,259,285]" italics="true" pageId="3" pageNumber="103">M. calilungae</emphasis>
</taxonomicName>
as both share bifurcated embolus and elongated TA but can be separated from it by beak shaped VTA (triangular shaped in
<taxonomicName id="4C349381FF94FFACFD96FECEFD3FFEBB" baseAuthorityName="Barrion &amp; Litsinger" baseAuthorityYear="1992" box="[588,743,295,321]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="calilungae">
<emphasis id="B9403410FF94FFACFD96FECEFD3FFEBB" box="[588,743,295,321]" italics="true" pageId="3" pageNumber="103">M. calilungae</emphasis>
</taxonomicName>
), and digitiform RTA (broad triangular ridge in
<taxonomicName id="4C349381FF94FFACFB22FECEFA4BFEBB" baseAuthorityName="Barrion &amp; Litsinger" baseAuthorityYear="1992" box="[1272,1427,295,321]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="calilungae">
<emphasis id="B9403410FF94FFACFB22FECEFA4BFEBB" box="[1272,1427,295,321]" italics="true" pageId="3" pageNumber="103">M. calilungae</emphasis>
</taxonomicName>
) (cf.
<bibRefCitation id="EFA595F3FF94FFACFF1CFEA2FE3FFE9F" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[198,487,331,357]" pageId="3" pageNumber="103" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF94FFACFEB3FEA2FE7BFE9F" box="[361,419,331,357]" italics="true" pageId="3" pageNumber="103">et al.</emphasis>
2012
</bibRefCitation>
: figs. 757759 and
<figureCitation id="130FF487FF94FFACFD10FEA2FC9DFE9F" box="[714,837,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Figs 1FH</figureCitation>
,
<figureCitation id="130FF487FF94FFACFC88FEA2FCA0FE9F" box="[850,888,331,357]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="3" pageNumber="103">8A</figureCitation>
). Females are most similar to the females of
<taxonomicName id="4C349381FF94FFACFAA4FEA2FEDCFE73" baseAuthorityName="Thorell" baseAuthorityYear="1892" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="B9403410FF94FFACFAA4FEA2FEDCFE73" italics="true" pageId="3" pageNumber="103">M. annulipes</emphasis>
</taxonomicName>
as both share V-shaped epigynal plate and the digitiform lateral border of the epigyne but can be separated from it by the anterior median depression (absent in
<taxonomicName id="4C349381FF94FFACFD07FE7AFCB6FE57" baseAuthorityName="Thorell" baseAuthorityYear="1892" box="[733,878,403,429]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="B9403410FF94FFACFD07FE7AFCB6FE57" box="[733,878,403,429]" italics="true" pageId="3" pageNumber="103">M. annulipes</emphasis>
</taxonomicName>
) and the highly sclerotized epigynal furrow of the epigyne (weakly sclerotized in
<taxonomicName id="4C349381FF94FFACFE2FFE5EFD5FFE2B" baseAuthorityName="Thorell" baseAuthorityYear="1892" box="[501,647,439,465]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="B9403410FF94FFACFE2FFE5EFD5FFE2B" box="[501,647,439,465]" italics="true" pageId="3" pageNumber="103">M. annulipes</emphasis>
</taxonomicName>
). (cf.
<bibRefCitation id="EFA595F3FF94FFACFD16FE5EFC30FE2B" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[716,1000,439,465]" pageId="3" pageNumber="103" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF94FFACFCB7FE5EFC7DFE2B" box="[877,933,439,465]" italics="true" pageId="3" pageNumber="103">et al.</emphasis>
2012
</bibRefCitation>
: figs. 733734 and
<figureCitation id="130FF487FF94FFACFB19FE5EFAFBFE2B" box="[1219,1315,439,465]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Figs. 1C</figureCitation>
,
<figureCitation id="130FF487FF94FFACFAEAFE5EFA8DFE2B" box="[1328,1365,439,465]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="3" pageNumber="103">9A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF94FFACFF1DFE33FA81F963" pageId="3" pageNumber="103" type="description">
<paragraph id="8B8BE802FF94FFACFF1DFE33FC13FBEB" blockId="3.[151,1437,150,1725]" pageId="3" pageNumber="103">
<emphasis id="B9403410FF94FFACFF1DFE33FE81FE0E" bold="true" box="[199,345,474,500]" pageId="3" pageNumber="103">Description.</emphasis>
<emphasis id="B9403410FF94FFACFE84FE32FE40FE0F" box="[350,408,475,501]" italics="true" pageId="3" pageNumber="103">Male</emphasis>
in alcohol (
<typeStatus id="548F56A0FF94FFACFDCCFE32FDA5FE0F" box="[534,637,475,501]" pageId="3" pageNumber="103" type="holotype">holotype</typeStatus>
;
<figureCitation id="130FF487FF94FFACFD5DFE32FD05FE0F" box="[647,733,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Fig. 1A</figureCitation>
). Body length 4.92; carapace 2.70 long, 1.93 wide; opisthosoma 2.22 long, 1.67 wide. Habitus as in
<figureCitation id="130FF487FF94FFACFDC7FE16FDA0FDE3" box="[541,632,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Figs 1A</figureCitation>
,
<figureCitation id="130FF487FF94FFACFD58FE16FD0CFDE3" box="[642,724,511,537]" captionStart="FIGURE 10" captionStartId="15.[151,250,1923,1947]" captionTargetBox="[151,1435,181,1900]" captionTargetId="figure-25@15.[151,1435,181,1900]" captionTargetPageId="15" captionText="FIGURE 10. Photographs of live Mallinella spp. described from Sri Lanka. AB Mallinella milkyway sp. nov., paratype female from Arisimale Beach Side (IFS_ZOD_205). CE Mallinella dhanahami sp. nov., paratype female from Gilimale Forest Reserve (IFS_ZOD_235). Photographs by SP. Benjamin." figureDoi="http://doi.org/10.5281/zenodo.14734683" httpUri="https://zenodo.org/record/14734683/files/figure.png" pageId="3" pageNumber="103">10CE</figureCitation>
. Carapace ovoid, smooth and shiny; fovea straight, longitudinal, red-brown. Clypeus brown, 0.45 high. Eye sizes and inter-distances: AME 0.16, ALE 0.09, PME 0.12, PLE 0.09; AMEAME 0.11, AMEPME 0.17, AMEALE 0.10, ALEALE 0.60, PMEPME 0.15, PMEPLE 0.20, PLEPLE 0.77, ALEPLE 0.08. MOA 0.45 long, front width 0.39, back width 0.43. Chilum unipartite, trapezoid. Chelicerae 0.92 long, light brown, without teeth. Endites yellow. Labium triangular, yellow-brown, 0.19 long, 0.38 wide. Sternum lateral margins with small semi-circular pits in front of each coxa of third and fourth pair of legs, furnished heavily with sparse black setae, 1.17 long, 0.95 wide (
<figureCitation id="130FF487FF94FFACFD2AFD3EFC9DFD0B" box="[752,837,727,753]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="3" pageNumber="103">Fig. 6F</figureCitation>
). Legs yellowish, round proximal tubercle on anterior femora (
<figureCitation id="130FF487FF94FFACFF2CFD12FE89FCEF" box="[246,337,763,789]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="3" pageNumber="103">Fig. 6A</figureCitation>
). Measurements of palp and legs: pedipalp (right) 3.45 (1.15 + 0.59 + 0.62 + 1.09), I 8.50 (2.10 + 0.70 + 1.90 + 1.95+ 1.85), II 7.40 (1.85 + 0.65 + 1.55 + 1.75 + 1.60), III 8.20 (2.05 + 0.70 + 1.60 + 2.15 + 1.70), IV 10.15 (2.30 + 0.60+ 2.25 + 3.25 + 1.75). Leg formula: 4132. Opisthosoma pear-shaped with lanceolate dorsal scutum (
<figureCitation id="130FF487FF94FFACFF20FC8EFE8DFC7B" box="[250,341,871,897]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Fig. 1A</figureCitation>
). Pattern on dorsum of opisthosoma with five pairs of spots: first pair is represented by minute pale spots; second pair is represented by oval spots obliquely arranged; third pair by transverse pale spots; fourth pair by paired spots medially connected; fifth pairs of the pattern on opisthosoma represented by minute pale spots distinctly smaller than anterior second and third pairs (
<figureCitation id="130FF487FF94FFACFCE6FC3AFC4DFC17" box="[828,917,979,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Fig. 1A</figureCitation>
). Posterior ventral spines elongated, gradually tapering with smoothly rounded apices. Spinnerets pale yellow (
<figureCitation id="130FF487FF94FFACFCBDFC1EFC64FBEB" box="[871,956,1015,1041]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF94FFACFF1DFBF2FD6DFB13" blockId="3.[151,1437,150,1725]" pageId="3" pageNumber="103">
<emphasis id="B9403410FF94FFACFF1DFBF2FF24FBCF" box="[199,252,1051,1077]" italics="true" pageId="3" pageNumber="103">Palp</emphasis>
(
<figureCitation id="130FF487FF94FFACFED3FBF2FE5AFBCF" box="[265,386,1051,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Figs 1FH</figureCitation>
,
<figureCitation id="130FF487FF94FFACFE51FBF2FE69FBCF" box="[395,433,1051,1077]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="3" pageNumber="103">8A</figureCitation>
). RTA digitiform, tapered towards blunt apex; VTA beak shaped (
<figureCitation id="130FF487FF94FFACFBA1FBF2FB0DFBCF" box="[1147,1237,1051,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Figs 1G</figureCitation>
,
<figureCitation id="130FF487FF94FFACFB3AFBF2FADDFBCF" box="[1248,1285,1051,1077]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="3" pageNumber="103">8A</figureCitation>
); Retrolateral cymbial fold broad, slightly more than half the length of cymbium (
<figureCitation id="130FF487FF94FFACFC76FBD6FBD0FBA3" box="[940,1032,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Fig. 1G</figureCitation>
). TA hook-shaped; apico-prolateral process of TA rod-shaped, directed posteromesad (
<figureCitation id="130FF487FF94FFACFD3FFB8AFC9EFB87" box="[741,838,1123,1149]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Figs 1G</figureCitation>
,
<figureCitation id="130FF487FF94FFACFC8CFB8AFCA4FB87" box="[854,892,1123,1149]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="3" pageNumber="103">8A</figureCitation>
). TS almost indistinct. TT rounded protrusion. Embolic base aligned in longitudinal direction, with narrow membranous area. Embolus medially bifurcated; mesal ramus slender, elongated, lateral ramus broad, blade-like (
<figureCitation id="130FF487FF94FFACFCE6FB42FC45FB3F" box="[828,925,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Figs 1G</figureCitation>
,
<figureCitation id="130FF487FF94FFACFC74FB42FC0BFB3F" box="[942,979,1195,1221]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="3" pageNumber="103">8A</figureCitation>
). Conductor an elevated mound with a groove to accommodate embolus (
<figureCitation id="130FF487FF94FFACFDC3FB26FDADFB13" box="[537,629,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Figs 1G</figureCitation>
,
<figureCitation id="130FF487FF94FFACFD58FB26FD70FB13" box="[642,680,1231,1257]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="3" pageNumber="103">8A</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF94FFACFF1DFB1AFE40F9D7" blockId="3.[151,1437,150,1725]" pageId="3" pageNumber="103">
<emphasis id="B9403410FF94FFACFF1DFB1AFEC3FAF7" box="[199,283,1267,1293]" italics="true" pageId="3" pageNumber="103">Female</emphasis>
<materialsCitation id="3B5CE25FFF94FFACFEFFFB1AFE4CF9D7" collectionCode="AME, ALE, PME, PLE, MOA" pageId="3" pageNumber="103" specimenCount="1" typeStatus="paratype">
in alcohol (
<typeStatus id="548F56A0FF94FFACFE73FB1AFDD7FAF7" box="[425,527,1267,1293]" pageId="3" pageNumber="103" type="paratype">paratype</typeStatus>
;
<figureCitation id="130FF487FF94FFACFDC7FB1AFDADFAF7" box="[541,629,1267,1293]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Fig. 1B</figureCitation>
). Body length 5.46; carapace 2.70 long, 1.96 wide; opisthosoma 2.76 long, 2.11 wide. Habitus and details as in male except for the following (
<figureCitation id="130FF487FF94FFACFC1AFAFEFBCEFACB" box="[960,1046,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Fig. 1B</figureCitation>
). Clypeus 0.58 high. Eye sizes and inter-distances:
<collectionCode id="ED2570C7FF94FFACFE92FAD2FE5EFAAF" box="[328,390,1339,1365]" pageId="3" pageNumber="103">AME</collectionCode>
0.16,
<collectionCode id="ED2570C7FF94FFACFE11FAD2FDD9FAAF" box="[459,513,1339,1365]" pageId="3" pageNumber="103">ALE</collectionCode>
0.10,
<collectionCode id="ED2570C7FF94FFACFD92FAD2FD5AFAAF" box="[584,642,1339,1365]" pageId="3" pageNumber="103">PME</collectionCode>
0.13,
<collectionCode id="ED2570C7FF94FFACFD13FAD2FD23FAAF" box="[713,763,1339,1365]" pageId="3" pageNumber="103">PLE</collectionCode>
0.11;
<collectionCode id="ED2570C7FF94FFACFCE5FAD2FCA5FAAF" box="[831,893,1339,1365]" pageId="3" pageNumber="103">AME</collectionCode>
<collectionCode id="ED2570C7FF94FFACFC56FAD2FC12FAAF" box="[908,970,1339,1365]" pageId="3" pageNumber="103">AME</collectionCode>
0.09,
<collectionCode id="ED2570C7FF94FFACFBD5FAD2FB95FAAF" box="[1039,1101,1339,1365]" pageId="3" pageNumber="103">AME</collectionCode>
<collectionCode id="ED2570C7FF94FFACFB81FAD2FB4DFAAF" box="[1115,1173,1339,1365]" pageId="3" pageNumber="103">PME</collectionCode>
0.14,
<collectionCode id="ED2570C7FF94FFACFB00FAD2FAC0FAAF" box="[1242,1304,1339,1365]" pageId="3" pageNumber="103">AME</collectionCode>
<collectionCode id="ED2570C7FF94FFACFAFCFAD2FA84FAAF" box="[1318,1372,1339,1365]" pageId="3" pageNumber="103">ALE</collectionCode>
0.13,
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<collectionCode id="ED2570C7FF94FFACFF06FAB6FECAFA83" box="[220,274,1375,1401]" pageId="3" pageNumber="103">ALE</collectionCode>
0.62,
<collectionCode id="ED2570C7FF94FFACFE80FAB6FE4CFA83" box="[346,404,1375,1401]" pageId="3" pageNumber="103">PME</collectionCode>
<collectionCode id="ED2570C7FF94FFACFE78FAB6FE04FA83" box="[418,476,1375,1401]" pageId="3" pageNumber="103">PME</collectionCode>
0.16,
<collectionCode id="ED2570C7FF94FFACFDFEFAB6FD85FA83" box="[548,605,1375,1401]" pageId="3" pageNumber="103">PME</collectionCode>
<collectionCode id="ED2570C7FF94FFACFDB1FAB6FD45FA83" box="[619,669,1375,1401]" pageId="3" pageNumber="103">PLE</collectionCode>
0.24,
<collectionCode id="ED2570C7FF94FFACFD3FFAB6FCCFFA83" box="[741,791,1375,1401]" pageId="3" pageNumber="103">PLE</collectionCode>
<collectionCode id="ED2570C7FF94FFACFCFFFAB6FC8FFA83" box="[805,855,1375,1401]" pageId="3" pageNumber="103">PLE</collectionCode>
0.80,
<collectionCode id="ED2570C7FF94FFACFC44FAB6FC0CFA83" box="[926,980,1375,1401]" pageId="3" pageNumber="103">ALE</collectionCode>
<collectionCode id="ED2570C7FF94FFACFC38FAB6FBCCFA83" box="[994,1044,1375,1401]" pageId="3" pageNumber="103">PLE</collectionCode>
0.08.
<collectionCode id="ED2570C7FF94FFACFB86FAB6FB44FA83" box="[1116,1180,1375,1401]" pageId="3" pageNumber="103">MOA</collectionCode>
0.46 long, front width 0.40, back width 0.42. Chelicerae 0.94 long, light brown. Labium 0.29 long, 0.45 wide. Sternum; 1.20 long, 1.06 wide. Measurements of palp and legs: pedipalp (right) 2.72 (0.91 + 0.47 + 0.49 + 0.85), I 7.40 (1.90 + 0.60 + 1.75 + 1.40 + 1.75), II 6.50 (1.60 + 0.55 + 1.55 + 1.30 + 1.50), III 7.35 (1.85 + 0.60+ 1.70 + 1.65 + 1.55), IV 9.05 (2.10 + 0.60 + 1.85 + 2.80 + 1.70). Leg formula: 4132. Opisthosoma oval-shaped; dorsum covered with dense black hairs; without dorsal scutum
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BE802FF94FFACFF1DF9DEFA81F963" blockId="3.[151,1437,150,1725]" pageId="3" pageNumber="103">
<emphasis id="B9403410FF94FFACFF1DF9DEFEE9F9AB" box="[199,305,1591,1617]" italics="true" pageId="3" pageNumber="103">Genitalia</emphasis>
(
<figureCitation id="130FF487FF94FFACFE98F9DEFE1BF9AB" box="[322,451,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1745,1769]" captionTargetBox="[255,1334,184,1715]" captionTargetId="figure-17@2.[249,1338,179,1719]" captionTargetPageId="2" captionText="FIGURE 1. Mallinella dhanahami sp. nov. A, EH holotype male (IFS_ZOD_234) and BE paratype female (IFS_ZOD_235). AB habitus, dorsal; C epigyne intact, ventral; D vulva, ventral; E posterior ventral spines, ventral; F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP = apicoprolateral process of tegular apophysis; C = conductor; Cy = cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts, LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TA, tegular apophysis; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734662" httpUri="https://zenodo.org/record/14734662/files/figure.png" pageId="3" pageNumber="103">Figs 1CD</figureCitation>
,
<figureCitation id="130FF487FF94FFACFE0BF9DEFDFAF9AB" box="[465,546,1591,1617]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="3" pageNumber="103">7A, 7F</figureCitation>
,
<figureCitation id="130FF487FF94FFACFDEAF9DEFDA0F9AB" box="[560,632,1591,1617]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="3" pageNumber="103">9AB</figureCitation>
). Epigynal plate narrowed, highly sclerotized, V-shaped, with anterior median depression. Lateral borders narrow, digitiform, terminally blunt. Epigynal furrow sclerotized. Spermathecae short, truncated. Insemination ducts shorter than the length of spermathecae. Fertilization ducts as in
<figureCitation id="130FF487FF94FFACFAD9F996FA8DF963" box="[1283,1365,1663,1689]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="3" pageNumber="103">
Fig.
<specimenCount id="9D32238BFF94FFACFAECF996FA8DF963" box="[1334,1365,1663,1689]" count="7" pageId="3" pageNumber="103" type="generic">7F</specimenCount>
</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF94FFACFF1DF94BFC84F947" box="[199,860,1698,1725]" pageId="3" pageNumber="103" type="distribution">
<paragraph id="8B8BE802FF94FFACFF1DF94BFC84F947" blockId="3.[151,1437,150,1725]" box="[199,860,1698,1725]" pageId="3" pageNumber="103">
<emphasis id="B9403410FF94FFACFF1DF94BFEB9F946" bold="true" box="[199,353,1698,1724]" pageId="3" pageNumber="103">Distribution.</emphasis>
Known only from the
<typeStatus id="548F56A0FF94FFACFDBAF94AFD48F947" box="[608,656,1699,1725]" pageId="3" pageNumber="103">type</typeStatus>
locality (
<figureCitation id="130FF487FF94FFACFD21F94AFC96F947" box="[763,846,1699,1725]" captionStart="FIGURE 12" captionStartId="17.[151,250,1133,1157]" captionText="FIGURE 12. Distribution map of known Mallinella species of Sri Lanka." pageId="3" pageNumber="103">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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Genus
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<emphasis id="B9403410FF96FFAEFF34FE26FEB4FE10" bold="true" box="[238,364,463,490]" italics="true" pageId="1" pageNumber="101">Mallinella</emphasis>
<bibRefCitation id="EFA595F3FF96FFAEFEA9FE39FDCBFE10" author="Strand, E." box="[371,531,464,490]" pageId="1" pageNumber="101" pagination="604 - 637" refId="ref12067" refString="Strand, E. (1906) Diagnosen nordafrikanischer, hauptsachlich von Carlo Freiherr von Erlanger gesammelter Spinnen. Zoologischer Anzeiger, 30, 604 - 637 + 655 - 690." type="journal article" year="1906">Strand, 1906</bibRefCitation>
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<emphasis id="B9403410FF96FFAEFF4DFDF1FECAFDC8" bold="true" box="[151,274,536,562]" pageId="1" pageNumber="101">Diagnosis.</emphasis>
The genus is characterised by the presence of a single row of ventral spines situated in front of the spinnerets, the palpal tibia with one or three apophyses, the conductor consisting of several parts, the embolic base connected to a tegulum via a thin membrane, the epigyne with a deep anterior median incision and the short and diverging copulatory ducts (
<bibRefCitation id="EFA595F3FF96FFAEFE0BFD6DFD28FD64" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[465,752,644,670]" pageId="1" pageNumber="101" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
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2012
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).
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<typeStatus id="548F56A0FF96FFAEFF1DFD41FEDAFD38" box="[199,258,680,706]" pageId="1" pageNumber="101">Type</typeStatus>
species.
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, by monotypy.
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<emphasis id="B9403410FF96FFAEFF1DFD25FEB9FD1C" bold="true" box="[199,353,716,742]" pageId="1" pageNumber="101">Distribution.</emphasis>
Africa, Asia, Central and South America
<emphasis id="B9403410FF96FFAEFCEEFD25FCE5FD1C" bold="true" box="[820,829,716,742]" pageId="1" pageNumber="101">(</emphasis>
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)
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<emphasis id="B9403410FF9DFFA5FF4DFC8FFEABFC7B" bold="true" box="[151,371,870,897]" italics="true" pageId="10" pageNumber="110">Mallinella truffles</emphasis>
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<figureCitation id="130FF487FF9DFFA5FF4DFC62FF06FC5F" box="[151,222,906,933]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">
Figs
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<figureCitation id="130FF487FF9DFFA5FF30FC62FEEAFC5F" box="[234,306,907,933]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="10" pageNumber="110">6E, 6J</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FEE4FC62FE58FC5F" box="[318,384,907,933]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="10" pageNumber="110">7E,7J</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FE51FC62FE75FC5F" box="[395,429,907,933]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">8E</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FE63FC62FE31FC5F" box="[441,489,907,933]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="10" pageNumber="110">9IJ</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9DFFA5FF4DFC3AFE1AFBCF" pageId="10" pageNumber="110" type="materials_examined">
<paragraph id="8B8BE802FF9DFFA5FF4DFC3AFE1AFBCF" blockId="10.[151,1437,978,2013]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF4DFC3AFE9DFC16" bold="true" box="[151,325,978,1005]" pageId="10" pageNumber="110">Type material.</emphasis>
<materialsCitation id="3B5CE25FFF9DFFA5FE96FC3AFC34FBEB" collectingDate="2010-07-13" country="Sri Lanka" elevation="1480" latitude="7.145833" location="Loolecondera Estate" longLatPrecision="21" longitude="80.69806" municipality="Kandy District" pageId="10" pageNumber="110" specimenCount="1" specimenCount-male="1" stateProvince="Central" typeStatus="holotype">
<emphasis id="B9403410FF9DFFA5FE96FC3AFCB8FC17" bold="true" box="[332,864,978,1008]" pageId="10" pageNumber="110">
<typeStatus id="548F56A0FF9DFFA5FE96FC3AFE61FC17" box="[332,441,979,1005]" pageId="10" pageNumber="110" type="holotype">Holotype</typeStatus>
<specimenCount id="9D32238BFF9DFFA5FE1AFC3DFE0FFC0A" box="[448,471,980,1008]" count="1" pageId="10" pageNumber="110" type="male"></specimenCount>
:
<collectingCountry id="F323A892FF9DFFA5FE3CFC3BFD5DFC16" box="[486,645,978,1004]" name="Sri Lanka" pageId="10" pageNumber="110">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF9DFFA5FD48FC3BFCB8FC17" bold="true" box="[658,864,978,1005]" italics="true" pageId="10" pageNumber="110">
<collectingRegion id="49F026E0FF9DFFA5FD48FC3BFC81FC17" box="[658,857,978,1005]" country="Sri Lanka" name="Central" pageId="10" pageNumber="110">Central Province</collectingRegion>
:
</emphasis>
</emphasis>
<collectingMunicipality id="6BEF7278FF9DFFA5FCBDFC3AFBC8FC17" box="[871,1040,979,1005]" pageId="10" pageNumber="110">Kandy District</collectingMunicipality>
,
<location id="8EEBBED9FF9DFFA5FBC1FC3AFADBFC17" LSID="urn:lsid:plazi:treatment:039D5914FF9DFFA3FF4DFC8FFC8DFD7F:8EEBBED9FF9DFFA5FBC1FC3AFADBFC17" box="[1051,1283,979,1005]" country="Sri Lanka" latitude="7.145833" longLatPrecision="21" longitude="80.69806" municipality="Kandy District" name="Loolecondera Estate" pageId="10" pageNumber="110" stateProvince="Central">Loolecondera Estate</location>
,
<geoCoordinate id="EE008EC5FF9DFFA5FAD5FC3BFA44FC17" box="[1295,1436,978,1005]" degrees="07" direction="north" minutes="08" orientation="latitude" pageId="10" pageNumber="110" precision="15" seconds="45" value="7.145833">07°0845N</geoCoordinate>
<geoCoordinate id="EE008EC5FF9DFFA5FF4DFC1FFEFBFBEB" box="[151,291,1014,1041]" degrees="80" direction="east" minutes="41" orientation="longitude" pageId="10" pageNumber="110" precision="15" seconds="53" value="80.69806">80°4153E</geoCoordinate>
,
<quantity id="4CCC45E7FF9DFFA5FEF5FC1EFE5EFBEB" box="[303,390,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="10" pageNumber="110" unit="m" value="1480.0">
<elevation id="00190F31FF9DFFA5FEF5FC1EFE5EFBEB" box="[303,390,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="10" pageNumber="110" unit="m" value="1480.0">1480 m</elevation>
</quantity>
,
<date id="FF8ACEC2FF9DFFA5FE48FC1EFDFDFBEB" box="[402,549,1015,1041]" pageId="10" pageNumber="110" value="2010-07-13">
<collectingDate id="EFCE372AFF9DFFA5FE48FC1EFDFDFBEB" box="[402,549,1015,1041]" pageId="10" pageNumber="110" value="2010-07-13">13 July 2010</collectingDate>
</date>
, leg. SP. Benjamin
<emphasis id="B9403410FF9DFFA5FD27FC1EFCF6FBEB" box="[765,814,1015,1041]" italics="true" pageId="10" pageNumber="110">et al</emphasis>
. (IFS_ZOD_43)
</materialsCitation>
.
<materialsCitation id="3B5CE25FFF9DFFA5FC2CFC1EFE67FBCF" collectingDate="2010-07-13" country="Sri Lanka" elevation="1480" latitude="7.145833" location="Loolecondera Estate" longLatPrecision="21" longitude="80.69806" municipality="Kandy District" pageId="10" pageNumber="110" specimenCount="1" specimenCount-female="1" stateProvince="Paratypes" typeStatus="paratype">
<emphasis id="B9403410FF9DFFA5FC2CFC1EFBAFFBEB" bold="true" box="[1014,1143,1015,1041]" pageId="10" pageNumber="110">
<typeStatus id="548F56A0FF9DFFA5FC2CFC1EFBAAFBEB" box="[1014,1138,1015,1041]" pageId="10" pageNumber="110" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D32238BFF9DFFA5FBA4FC1EFB79FBEB" box="[1150,1185,1015,1041]" count="1" pageId="10" pageNumber="110" type="female">1♀</specimenCount>
, with same data as for holotype (IFS_ZOD_230)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9DFFA5FF1DFBD6FAC2FBA3" box="[199,1306,1087,1113]" pageId="10" pageNumber="110" type="etymology">
<paragraph id="8B8BE802FF9DFFA5FF1DFBD6FAC2FBA3" blockId="10.[151,1437,978,2013]" box="[199,1306,1087,1113]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DFBD6FE96FBA3" bold="true" box="[199,334,1087,1113]" pageId="10" pageNumber="110">Etymology.</emphasis>
The new species is named after the chocolate Truffles. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9DFFA5FF1DFB8BFE11FA83" pageId="10" pageNumber="110" type="diagnosis">
<paragraph id="8B8BE802FF9DFFA5FF1DFB8BFE11FA83" blockId="10.[151,1437,978,2013]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DFB8BFE9AFB86" bold="true" box="[199,322,1122,1148]" pageId="10" pageNumber="110">Diagnosis.</emphasis>
<taxonomicName id="4C349381FF9DFFA5FE94FB8AFDC7FB87" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[334,543,1123,1149]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="110" phylum="Arthropoda" rank="species" species="truffles" status="sp. nov.">
<emphasis id="B9403410FF9DFFA5FE94FB8AFDC7FB87" box="[334,543,1123,1149]" italics="true" pageId="10" pageNumber="110">Mallinella truffles</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9DFFA5FDF6FB8AFD53FB87" bold="true" box="[556,651,1123,1149]" pageId="10" pageNumber="110">
<taxonomicNameLabel id="A273896BFF9DFFA5FDF6FB8AFD53FB87" box="[556,651,1123,1149]" pageId="10" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share features such as unbranched embolus, simple rostrated TA, longitudinal furrow of the epigyne with the members of the tuberculata group. Within tuberculata group, males of
<taxonomicName id="4C349381FF9DFFA5FF4DFB42FED3FB3F" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[151,267,1195,1221]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="110" phylum="Arthropoda" rank="species" species="truffles" status="sp. nov.">
<emphasis id="B9403410FF9DFFA5FF4DFB42FED3FB3F" box="[151,267,1195,1221]" italics="true" pageId="10" pageNumber="110">M. truffles</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9DFFA5FEC8FB42FEB4FB3F" bold="true" box="[274,364,1195,1221]" pageId="10" pageNumber="110">
<taxonomicNameLabel id="A273896BFF9DFFA5FEC8FB42FEB4FB3F" box="[274,364,1195,1221]" pageId="10" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are most similar to the males of
<taxonomicName id="4C349381FF9DFFA5FD02FB42FCB2FB3F" authorityName="Dankittipakul, Jocque &amp; Singtripop" authorityYear="2012" box="[728,874,1195,1221]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="110" phylum="Arthropoda" rank="species" species="brachiata">
<emphasis id="B9403410FF9DFFA5FD02FB42FCB2FB3F" box="[728,874,1195,1221]" italics="true" pageId="10" pageNumber="110">M. brachiata</emphasis>
</taxonomicName>
as both share digitiform RTA, cymbial fold which is less than half the length of the cymbium, triangular apico prolateral process but can be separated from it by long narrow unbranched embolus. (cf.
<bibRefCitation id="EFA595F3FF9DFFA5FDC0FB1AFCE5FAF7" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[538,829,1267,1293]" pageId="10" pageNumber="110" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF9DFFA5FD64FB1AFD29FAF7" box="[702,753,1267,1293]" italics="true" pageId="10" pageNumber="110">et al</emphasis>
. 2012
</bibRefCitation>
: figs 147151 and
<figureCitation id="130FF487FF9DFFA5FBC1FB1AFB4EFAF7" box="[1051,1174,1266,1293]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs 5FH</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FB7EFB1AFB1FFAF7" box="[1188,1223,1267,1293]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">8E</figureCitation>
). Females are also most similar to the females of
<taxonomicName id="4C349381FF9DFFA5FE3FFAFEFD52FACB" authorityName="Dankittipakul, Jocque &amp; Singtripop" authorityYear="2012" box="[485,650,1303,1329]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="110" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B9403410FF9DFFA5FE3FFAFEFD52FACB" box="[485,650,1303,1329]" italics="true" pageId="10" pageNumber="110">M. tuberculata</emphasis>
</taxonomicName>
as both share sub-rectangular epigynal plate, and deep anterior median depression, but can be separated from it by globular spermathecae (cf.
<bibRefCitation id="EFA595F3FF9DFFA5FC68FAD2FB16FAAF" author="Dankittipakul, P. &amp; Jocque, R. &amp; Singtripop, T." box="[946,1230,1339,1365]" pageId="10" pageNumber="110" pagination="1 - 327" refId="ref11715" refString="Dankittipakul, P., Jocque, R. &amp; Singtripop, T. (2012) Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from south East Asia (Araneae, Zodariidae). Zootaxa, 3369 (1), 1 - 327. https: // doi. org / 10.11646 / zootaxa. 3369.1.1" type="journal article" year="2012">
Dankittipakul
<emphasis id="B9403410FF9DFFA5FB89FAD2FB5CFAAF" box="[1107,1156,1339,1365]" italics="true" pageId="10" pageNumber="110">et al</emphasis>
. 2012
</bibRefCitation>
: figs 152155 and
<figureCitation id="130FF487FF9DFFA5FF4DFAB6FEC3FA83" box="[151,283,1374,1401]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs. 5CD</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FEFCFAB6FEB6FA83" box="[294,366,1375,1401]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="10" pageNumber="110">7E, 7J</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FEA0FAB6FE65FA83" box="[378,445,1375,1401]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="10" pageNumber="110">9I9J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9DFFA3FF1DFA6BFA9EFD9B" lastPageId="12" lastPageNumber="112" pageId="10" pageNumber="110" type="description">
<paragraph id="8B8BE802FF9DFFA5FF1DFA6BFB45F88B" blockId="10.[151,1437,978,2013]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DFA6BFE81FA66" bold="true" box="[199,345,1410,1436]" pageId="10" pageNumber="110">Description.</emphasis>
<emphasis id="B9403410FF9DFFA5FE84FA6AFE40FA67" box="[350,408,1411,1437]" italics="true" pageId="10" pageNumber="110">Male</emphasis>
in alcohol (
<typeStatus id="548F56A0FF9DFFA5FDCCFA6AFDA5FA67" box="[534,637,1411,1437]" pageId="10" pageNumber="110" type="holotype">holotype</typeStatus>
;
<figureCitation id="130FF487FF9DFFA5FD5DFA6AFD06FA67" box="[647,734,1410,1437]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5A</figureCitation>
). Body length 4.96; carapace 2.66 long, 2.20 wide; opisthosoma 2.30 long, 1.60 wide. Habitus as in
<figureCitation id="130FF487FF9DFFA5FDF8FA4EFDA2FA3B" box="[546,634,1447,1473]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="10" pageNumber="110">Fig. 7A</figureCitation>
. Carapace ovoid, smooth, red-brown; fovea red-brown. Clypeus brown, 0.57 high. Eye sizes and inter-distances: AME 0.16, ALE 0.10, PME 0.12, PLE 0.10; AMEAME 0.10, AMEPME 0.13, AMEALE 0.12, ALEALE 0.55, PMEPME 0.12, PMEPLE 0.25, PLEPLE 0.76, ALEPLE 0.06. MOA 0.44 long, front width 0.38, back width 0.41. Chelicerae 0.89 long, light brown, without teeth. Endites yellow. Labium triangular, yellow-brown, 0.20 long, 0.35 wide. Sternum yellowish, furnished with sparse black setae, 0.89 long, 0.74 wide. Lateral margins of sternum with small semi-circular pits in front of each coxa of third and fourth pair of legs. Legs yellowish, with proximal tubercle. Measurements of palp and legs: pedipalp (right) 2.36 (0.83 + 0.40 + 0.44 + 0.69), I 6.10 (1.60 + 0.65 + 1.40 + 1.45+ 1.00), II 5.45 (1.45 + 0.65 + 1.10 + 1.25 + 1.00), III 6.00 (1.60 + 0.65 + 1.25 + 1.50 + 1.00), IV 8.00 (2.00 + 0.70+ 1.60 + 2.35 + 1.35). Leg formula: 4132. Opisthosoma oval; pattern on dorsum of opisthosoma provided large pale patches, roughly oval first pair of patches connected vertically with nearly round second pair, third and fourth pairs of nearly round patches connected medially and vertically with each other (
<figureCitation id="130FF487FF9DFFA5FE0BF8DAFDF7F8B7" box="[465,559,1842,1869]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5A</figureCitation>
). Dorsum of opisthosoma with large ovoid scutum. Posterior ventral spines elongated, gradually tapering, with acute apices (
<figureCitation id="130FF487FF9DFFA5FD67F8BEFCCFF88B" box="[701,791,1878,1905]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5A</figureCitation>
). Spinnerets pale yellow (
<figureCitation id="130FF487FF9DFFA5FBE3F8BEFB48F88B" box="[1081,1168,1878,1905]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5E</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF9DFFA4FF1DF892FD71FF2F" blockId="10.[151,1437,978,2013]" lastBlockId="11.[151,1436,150,213]" lastPageId="11" lastPageNumber="111" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DF892FF24F86F" box="[199,252,1915,1941]" italics="true" pageId="10" pageNumber="110">Palp</emphasis>
(
<figureCitation id="130FF487FF9DFFA5FED5F892FE57F86F" box="[271,399,1914,1941]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs 5FH</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FE45F892FE1AF86F" box="[415,450,1915,1941]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">8E</figureCitation>
). VTA elevated subtriangular hump (
<figureCitation id="130FF487FF9DFFA5FCACF892FC00F86F" box="[886,984,1914,1941]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs 5G</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FC32F892FBD3F86F" box="[1000,1035,1915,1941]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">8E</figureCitation>
); RTA digitiform, tapered towards blunt apex (
<figureCitation id="130FF487FF9DFFA5FEFBF876FE74F843" box="[289,428,1950,1977]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Figs. 5GH</figureCitation>
); Cymbium with deep retrolateral fold, less than half the length of cymbium (
<figureCitation id="130FF487FF9DFFA5FAE9F876FA57F843" box="[1331,1423,1950,1977]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="10" pageNumber="110">Fig. 5G</figureCitation>
), apicoprolateral process triangular with blunt apex, TT large triangular with pointed apex. Embolic base aligned in diagonally, directed posteriorly, with narrow membranous area (
<figureCitation id="130FF487FF9CFFA4FCB8FF7EFC64FF4B" box="[866,956,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="11" pageNumber="111">Fig. 5G</figureCitation>
). Embolus, long, narrow, unbranched, with blunt apex. Conductor triangular, pointed apex.
</paragraph>
<caption id="DF4BB88AFF9CFFA4FF4DF922FD8DF825" ID-DOI="http://doi.org/10.5281/zenodo.14734672" ID-Zenodo-Dep="14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="11" pageNumber="111" startId="11.[151,250,1739,1763]" targetBox="[275,1312,241,1710]" targetPageId="11" targetType="figure">
<paragraph id="8B8BE802FF9CFFA4FF4DF922FD8DF825" blockId="11.[151,1437,1739,2016]" pageId="11" pageNumber="111">
<emphasis id="B9403410FF9CFFA4FF4DF922FECAF91E" bold="true" box="[151,274,1739,1764]" pageId="11" pageNumber="111">FIGURE 5.</emphasis>
<taxonomicName id="4C349381FF9CFFA4FEC2F922FE08F919" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[280,464,1739,1763]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="111" phylum="Arthropoda" rank="species" species="truffles" status="sp. nov.">
<emphasis id="B9403410FF9CFFA4FEC2F922FE08F919" box="[280,464,1739,1763]" italics="true" pageId="11" pageNumber="111">Mallinella truffles</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9CFFA4FE0FF925FDFDF91E" bold="true" box="[469,549,1740,1764]" pageId="11" pageNumber="111">
<taxonomicNameLabel id="A273896BFF9CFFA4FE0FF925FDFDF91E" box="[469,549,1740,1764]" pageId="11" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8BE802FF9BFFA3FF1DFF7EFEDDFE2B" blockId="12.[151,1437,150,645]" pageId="12" pageNumber="112">
<emphasis id="B9403410FF9BFFA3FF1DFF7EFEC3FF4B" box="[199,283,151,177]" italics="true" pageId="12" pageNumber="112">Female</emphasis>
<materialsCitation id="3B5CE25FFF9BFFA3FEFFFF7EFED9FE2B" collectionCode="AME, ALE, PME, PLE, MOA" pageId="12" pageNumber="112" specimenCount="1" typeStatus="paratype">
in alcohol (
<typeStatus id="548F56A0FF9BFFA3FE73FF7EFDD7FF4B" box="[425,527,151,177]" pageId="12" pageNumber="112" type="paratype">paratype</typeStatus>
;
<figureCitation id="130FF487FF9BFFA3FDC6FF7EFDAEFF4B" box="[540,630,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Fig. 5B</figureCitation>
); Body length 6.46; carapace 2.97 long, 2.01 wide; opisthosoma 3.49 long, 2.46 wide. Habitus and details as in male except for the following (
<figureCitation id="130FF487FF9BFFA3FC1AFF52FBC0FF2F" box="[960,1048,186,213]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Fig. 5B</figureCitation>
). Clypeus 0.34 high. Eye sizes and inter-distances:
<collectionCode id="ED2570C7FF9BFFA3FE92FF36FE5EFF03" box="[328,390,223,249]" pageId="12" pageNumber="112">AME</collectionCode>
0.13,
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0.11,
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0.12,
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0.11;
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<collectionCode id="ED2570C7FF9BFFA3FC51FF36FC11FF03" box="[907,969,223,249]" pageId="12" pageNumber="112">AME</collectionCode>
0.09,
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<collectionCode id="ED2570C7FF9BFFA3FB81FF36FB4DFF03" box="[1115,1173,223,249]" pageId="12" pageNumber="112">PME</collectionCode>
0.18,
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<collectionCode id="ED2570C7FF9BFFA3FAFCFF36FA84FF03" box="[1318,1372,223,249]" pageId="12" pageNumber="112">ALE</collectionCode>
0.13,
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<collectionCode id="ED2570C7FF9BFFA3FF06FEEAFECAFEE7" box="[220,274,259,285]" pageId="12" pageNumber="112">ALE</collectionCode>
0.60,
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<collectionCode id="ED2570C7FF9BFFA3FE78FEEAFE04FEE7" box="[418,476,259,285]" pageId="12" pageNumber="112">PME</collectionCode>
0.13,
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<collectionCode id="ED2570C7FF9BFFA3FDB1FEEAFD45FEE7" box="[619,669,259,285]" pageId="12" pageNumber="112">PLE</collectionCode>
0.26,
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0.78,
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0.04.
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0.45 long, front width 0.36, back width 0.38. Labium 0.34 long, 0.47 wide. Sternum with shallow semi-circular pits, 1.25 long, 1.08 wide. Measurements of palp and legs: pedipalp (right) 2.25 (0.81 + 0.37 + 0.40 + 0.67) I 6.65 (1.80 + 0.75 + 1.60 + 1.55 + 0.95), II 6.60 (1.65 + 0.75 + 1.40 + 1.55 + 1.25), III 6.70 (1.70 + 0.75+ 1.50 + 1.75 + 1.00), IV 8.05 (1.95 + 0.75 + 1.75 + 2.25 + 1.35). Dorsum pattern of opisthosoma, indistinct, covered with thick black and white pubescence (
<figureCitation id="130FF487FF9BFFA3FF45FE5EFF20FE2B" box="[159,248,438,465]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Fig. 5B</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BE802FF9BFFA3FF1DFE32FA9EFD9B" blockId="12.[151,1437,150,645]" pageId="12" pageNumber="112">
<emphasis id="B9403410FF9BFFA3FF1DFE32FEE9FE0F" box="[199,305,475,501]" italics="true" pageId="12" pageNumber="112">Genitalia</emphasis>
(
<figureCitation id="130FF487FF9BFFA3FE9BFE32FE19FE0F" box="[321,449,474,501]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Figs 5CD</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FE14FE32FDC0FE0F" box="[462,536,475,501]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="12" pageNumber="112">7E, 7J</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FDFFFE32FDB0FE0F" box="[549,616,475,501]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">9I9J</figureCitation>
). Epigynal plate wide, sub rectangular, with curved sclerotized epigynal furrows directed towards posteriorly (
<figureCitation id="130FF487FF9BFFA3FD8EFE16FD6EFDE3" box="[596,694,510,537]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Figs 5C</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FD1DFE16FD3AFDE3" box="[711,738,511,537]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">9I</figureCitation>
). Lateral borders sub-triangular, terminally acute (
<figureCitation id="130FF487FF9BFFA3FAECFE16FA40FDE3" box="[1334,1432,510,537]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Figs 5C</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FF4DFDCAFF6AFDC7" box="[151,178,547,573]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">9I</figureCitation>
). Insemination ducts short, thick, distally diverging (
<figureCitation id="130FF487FF9BFFA3FCC5FDCAFC59FDC7" box="[799,897,546,573]" captionStart="FIGURE 5" captionStartId="11.[151,250,1739,1763]" captionTargetBox="[275,1312,241,1710]" captionTargetId="figure-45@11.[271,1316,237,1716]" captionTargetPageId="11" captionText="FIGURE 5. Mallinella truffles sp. nov. A, EH holotype male (IFS_ZOD_43) and BE paratype female (IFS_ZOD_230).AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734672" httpUri="https://zenodo.org/record/14734672/files/figure.png" pageId="12" pageNumber="112">Figs 5D</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FC4AFDCAFC05FDC7" box="[912,989,547,573]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="12" pageNumber="112">7E, 7J</figureCitation>
,
<figureCitation id="130FF487FF9BFFA3FC37FDCAFBD1FDC7" box="[1005,1033,547,573]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">9J</figureCitation>
), longer than spermathecal length. Spermathecae, globular in shape distally with coiled internal ducts (
<figureCitation id="130FF487FF9BFFA3FC55FDAEFC07FD9B" box="[911,991,583,609]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="12" pageNumber="112">Fig. 9J</figureCitation>
). Fertilization ducts as in fig 7J.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9BFFA3FF1DFD83FC8DFD7F" box="[199,853,618,645]" pageId="12" pageNumber="112" type="distribution">
<paragraph id="8B8BE802FF9BFFA3FF1DFD83FC8DFD7F" blockId="12.[151,1437,150,645]" box="[199,853,618,645]" pageId="12" pageNumber="112">
<emphasis id="B9403410FF9BFFA3FF1DFD83FEB9FD7E" bold="true" box="[199,353,618,644]" pageId="12" pageNumber="112">Distribution.</emphasis>
Known only from the
<typeStatus id="548F56A0FF9BFFA3FDBAFD82FD48FD7F" box="[608,656,619,645]" pageId="12" pageNumber="112">type</typeStatus>
locality (
<figureCitation id="130FF487FF9BFFA3FD21FD82FC90FD7F" box="[763,840,619,645]" captionStart="FIGURE 12" captionStartId="17.[151,250,1133,1157]" captionText="FIGURE 12. Distribution map of known Mallinella species of Sri Lanka." pageId="12" pageNumber="112">Fig.12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="6A1D9B25EE8E5B5072192F724F6D6D7B" ID-DOI="10.11646/zootaxa.5570.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734655" ID-ZooBank="1995C18B-1993-4DC4-B8AB-E4891F148A1E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737736390657" checkinUser="plazi" docAuthor="Dayananda, Naruwan &amp; Benjamin, Suresh P." docDate="2025" docId="039D5914FF9FFFA5FF4DFD3FFC84FCEF" docLanguage="en" docName="zootaxa.5570.1.4.pdf" docOrigin="Zootaxa 5570 (1)" docSource="https://doi.org/10.11646/zootaxa.5570.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mallinella oreo Dayananda &amp; Benjamin, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="110" masterDocId="FFA4216CFF97FFAFFFDAFFE9FFD8FFFA" masterDocTitle="Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka" masterLastPageNumber="118" masterPageNumber="100" pageNumber="108" updateTime="1737736825942" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2688BEE0124AFBBB277B83473457FF3A">Five new species of the genus Mallinella Strand, 1906 (Araneae: Zodariidae) from Sri Lanka</mods:title>
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<mods:namePart id="B296B5A0B56CF159D1C495EB872C8DB3">Dayananda, Naruwan</mods:namePart>
<mods:affiliation id="4F12D7B658D7720741859248B3A6AC62">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka.</mods:affiliation>
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<mods:namePart id="74996770D2B7D1934F7ADDB8CAC0FA69">Benjamin, Suresh P.</mods:namePart>
<mods:affiliation id="30992E20B16E574BF09A7ABC940C6B58">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka.</mods:affiliation>
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<mods:date id="518B957861D3B22EF4C1F9D961DCF77C">2025</mods:date>
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<heading id="D0C35F6EFF9FFFA7FF4DFD3FFE6FFD0B" bold="true" box="[151,439,726,753]" fontSize="11" level="1" pageId="8" pageNumber="108" reason="1">
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<taxonomicName id="4C349381FF9FFFA7FF4DFD3FFE8AFD0B" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[151,338,726,753]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="oreo" status="sp. nov.">
<emphasis id="B9403410FF9FFFA7FF4DFD3FFE8AFD0B" bold="true" box="[151,338,726,753]" italics="true" pageId="8" pageNumber="108">Mallinella oreo</emphasis>
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<subSubSection id="C32EBB89FF9FFFA7FF4DFD12FDD2FCEF" box="[151,522,763,789]" pageId="8" pageNumber="108" type="description">
<paragraph id="8B8BE802FF9FFFA7FF4DFD12FDD2FCEF" blockId="8.[151,522,726,789]" box="[151,522,763,789]" pageId="8" pageNumber="108">
<figureCitation id="130FF487FF9FFFA7FF4DFD12FF06FCEF" box="[151,222,763,789]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Figs 4</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FF30FD12FEECFCEF" box="[234,308,763,789]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="8" pageNumber="108">6D, 6I</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FE9AFD12FE51FCEF" box="[320,393,763,789]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="8" pageNumber="108">7D, 7I</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FE4CFD12FE62FCEF" box="[406,442,763,789]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="8" pageNumber="108">8D</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FE1CFD12FDD2FCEF" box="[454,522,763,789]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="8" pageNumber="108">9GH</figureCitation>
</paragraph>
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<paragraph id="8B8BE802FF9FFFA7FF4DFCAAFD6FFC5F" blockId="8.[151,1437,834,2013]" pageId="8" pageNumber="108">
<emphasis id="B9403410FF9FFFA7FF4DFCAAFE9EFCA6" bold="true" box="[151,326,834,861]" pageId="8" pageNumber="108">Type material.</emphasis>
<materialsCitation id="3B5CE25FFF9FFFA7FE94FCAAFB34FC7B" collectingDate="2014-01-23" country="Sri Lanka" county="Badulla District" elevation="2000" latitude="0.8666667" location="Passara - Ella Road" longLatPrecision="1308" longitude="0.11666667" municipality="Namunukula Peak" pageId="8" pageNumber="108" specimenCount="1" specimenCount-male="1" stateProvince="Uva" typeStatus="holotype">
<emphasis id="B9403410FF9FFFA7FE94FCAAFCE5FCA7" bold="true" box="[334,829,834,864]" pageId="8" pageNumber="108">
<typeStatus id="548F56A0FF9FFFA7FE94FCAAFE63FCA7" box="[334,443,835,861]" pageId="8" pageNumber="108" type="holotype">Holotype</typeStatus>
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:
<collectingCountry id="F323A892FF9FFFA7FE33FCABFD51FCA6" box="[489,649,834,860]" name="Sri Lanka" pageId="8" pageNumber="108">SRI LANKA</collectingCountry>
:
<emphasis id="B9403410FF9FFFA7FD4DFCAAFCE5FCA7" bold="true" box="[663,829,835,861]" italics="true" pageId="8" pageNumber="108">
<collectingRegion id="49F026E0FF9FFFA7FD4DFCAAFCEEFCA7" box="[663,822,835,861]" country="Sri Lanka" name="Uva" pageId="8" pageNumber="108">Uva Province</collectingRegion>
:
</emphasis>
</emphasis>
<collectingCounty id="62EA908EFF9FFFA7FC9FFCAAFC24FCA7" box="[837,1020,835,861]" pageId="8" pageNumber="108">Badulla District</collectingCounty>
,
<collectingMunicipality id="6BEF7278FF9FFFA7FBD2FCAAFB01FCA7" box="[1032,1241,835,861]" pageId="8" pageNumber="108">Namunukula Peak</collectingMunicipality>
, along
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, 06
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, 81
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, leg. SP. Benjamin
<emphasis id="B9403410FF9FFFA7FC2AFC8EFBF8FC7B" box="[1008,1056,871,897]" italics="true" pageId="8" pageNumber="108">et al</emphasis>
. (IFS_ZOD_137)
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.
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<emphasis id="B9403410FF9FFFA7FB2CFC8EFAB4FC7B" bold="true" box="[1270,1388,871,897]" pageId="8" pageNumber="108">
<typeStatus id="548F56A0FF9FFFA7FB2CFC8EFABFFC7B" box="[1270,1383,871,897]" pageId="8" pageNumber="108" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D32238BFF9FFFA7FAA8FC8EFA4DFC7B" box="[1394,1429,871,897]" count="1" pageId="8" pageNumber="108" type="female">1♀</specimenCount>
, with same data as for holotype (IFS_ZOD_138)
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.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9FFFA7FF1DFC46FB21FC33" box="[199,1273,943,969]" pageId="8" pageNumber="108" type="etymology">
<paragraph id="8B8BE802FF9FFFA7FF1DFC46FB21FC33" blockId="8.[151,1437,834,2013]" box="[199,1273,943,969]" pageId="8" pageNumber="108">
<emphasis id="B9403410FF9FFFA7FF1DFC46FE96FC33" bold="true" box="[199,334,943,969]" pageId="8" pageNumber="108">Etymology.</emphasis>
The new species is named after the chocolate Oreo. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9FFFA7FF1DFC3BFF21FACB" pageId="8" pageNumber="108" type="diagnosis">
<paragraph id="8B8BE802FF9FFFA7FF1DFC3BFF21FACB" blockId="8.[151,1437,834,2013]" pageId="8" pageNumber="108">
<emphasis id="B9403410FF9FFFA7FF1DFC3BFE9AFC16" bold="true" box="[199,322,978,1004]" pageId="8" pageNumber="108">Diagnosis.</emphasis>
<taxonomicName id="4C349381FF9FFFA7FE90FC3AFE22FC16" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[330,506,979,1005]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="oreo" status="sp. nov.">
<emphasis id="B9403410FF9FFFA7FE90FC3AFE22FC16" box="[330,506,979,1005]" italics="true" pageId="8" pageNumber="108">Mallinella oreo</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9FFFA7FDD9FC3AFDAAFC17" bold="true" box="[515,626,979,1005]" pageId="8" pageNumber="108">
<taxonomicNameLabel id="A273896BFF9FFFA7FDD9FC3AFD86FC17" box="[515,606,979,1005]" pageId="8" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
s
</emphasis>
hares the feature of inwardly projected digitiform lateral borders of the epigyne with the members of the annulipes-group. Within the annulipes-group, males of
<taxonomicName id="4C349381FF9FFFA7FBA6FC1EFB0CFBEA" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[1148,1236,1015,1040]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="oreo" status="sp. nov.">
<emphasis id="B9403410FF9FFFA7FBA6FC1EFB0CFBEA" box="[1148,1236,1015,1040]" italics="true" pageId="8" pageNumber="108">M. oreo</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9FFFA7FB01FC1EFAEDFBEB" bold="true" box="[1243,1333,1015,1041]" pageId="8" pageNumber="108">
<taxonomicNameLabel id="A273896BFF9FFFA7FB01FC1EFAEDFBEB" box="[1243,1333,1015,1041]" pageId="8" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are most similar to the males of
<taxonomicName id="4C349381FF9FFFA7FE7AFBF2FDE8FBCF" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[416,560,1051,1077]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF9FFFA7FE7AFBF2FDE8FBCF" box="[416,560,1051,1077]" italics="true" pageId="8" pageNumber="108">M. moncheri</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9FFFA7FDE3FBF2FD4BFBCF" bold="true" box="[569,659,1051,1077]" pageId="8" pageNumber="108">
<taxonomicNameLabel id="A273896BFF9FFFA7FDE3FBF2FD4BFBCF" box="[569,659,1051,1077]" pageId="8" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as both share digitiform RTA, less prominent cymbial fold which is less than half the length of the cymbium, and diagonally aligned embolic base but can be separated from it by the triangular hump like VTA (moderately developed with curved apex in
<taxonomicName id="4C349381FF9FFFA7FC1BFB8AFB8BFB87" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[961,1107,1123,1149]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF9FFFA7FC1BFB8AFB8BFB87" box="[961,1107,1123,1149]" italics="true" pageId="8" pageNumber="108">M. moncheri</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9FFFA7FB86FB8AFB6FFB87" bold="true" box="[1116,1207,1123,1149]" pageId="8" pageNumber="108">
<taxonomicNameLabel id="A273896BFF9FFFA7FB86FB8AFB6FFB87" box="[1116,1207,1123,1149]" pageId="8" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), and subterminally bifurcated embolus (slender unbranched in
<taxonomicName id="4C349381FF9FFFA7FD52FB6EFCC1FB5B" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[648,793,1159,1185]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF9FFFA7FD52FB6EFCC1FB5B" box="[648,793,1159,1185]" italics="true" pageId="8" pageNumber="108">M. moncheri</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9FFFA7FCF8FB6EFCA6FB5B" bold="true" box="[802,894,1159,1185]" pageId="8" pageNumber="108">
<taxonomicNameLabel id="A273896BFF9FFFA7FCF8FB6EFCA6FB5B" box="[802,894,1159,1185]" pageId="8" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) (
<figureCitation id="130FF487FF9FFFA7FC42FB6EFC2FFB5B" box="[920,1015,1159,1185]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="8" pageNumber="108">Figs 3G</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FBDFFB6EFBF1FB5B" box="[1029,1065,1159,1185]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="8" pageNumber="108">8C</figureCitation>
;
<figureCitation id="130FF487FF9FFFA7FBEDFB6EFB84FB5B" box="[1079,1116,1159,1185]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">4G</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FBB3FB6EFB54FB5B" box="[1129,1164,1159,1185]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="8" pageNumber="108">8D</figureCitation>
) Females are also most similar to the females of
<taxonomicName id="4C349381FF9FFFA7FE1BFB42FD8CFB3F" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[449,596,1195,1221]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF9FFFA7FE1BFB42FD8CFB3F" box="[449,596,1195,1221]" italics="true" pageId="8" pageNumber="108">M. moncheri</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9FFFA7FD84FB42FD64FB3F" bold="true" box="[606,700,1195,1221]" pageId="8" pageNumber="108">
<taxonomicNameLabel id="A273896BFF9FFFA7FD84FB42FD64FB3F" box="[606,700,1195,1221]" pageId="8" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as both share terminally blunt digitiform lateral borders of the epigyne but can be separated from it by semi-circular shaped epigynal plate (
<emphasis id="B9403410FF9FFFA7FC37FB26FB8DFB13" box="[1005,1109,1231,1257]" italics="true" pageId="8" pageNumber="108">V-shaped</emphasis>
in
<taxonomicName id="4C349381FF9FFFA7FBADFB26FADEFB13" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[1143,1286,1231,1257]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF9FFFA7FBADFB26FADEFB13" box="[1143,1286,1231,1257]" italics="true" pageId="8" pageNumber="108">M. moncheri</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9FFFA7FAD6FB26FABCFB13" bold="true" box="[1292,1380,1231,1257]" pageId="8" pageNumber="108">
<taxonomicNameLabel id="A273896BFF9FFFA7FAD6FB26FABCFB13" box="[1292,1380,1231,1257]" pageId="8" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and small anterior median depression of the epigyne (deep median depression in
<taxonomicName id="4C349381FF9FFFA7FC25FB1AFB48FAF7" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[1023,1168,1267,1293]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="moncheri" status="sp. nov.">
<emphasis id="B9403410FF9FFFA7FC25FB1AFB48FAF7" box="[1023,1168,1267,1293]" italics="true" pageId="8" pageNumber="108">M. moncheri</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9FFFA7FB42FB1AFB2AFAF7" bold="true" box="[1176,1266,1267,1293]" pageId="8" pageNumber="108">
<taxonomicNameLabel id="A273896BFF9FFFA7FB42FB1AFB2AFAF7" box="[1176,1266,1267,1293]" pageId="8" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) (
<figureCitation id="130FF487FF9FFFA7FAD6FB1AFAB0FAF7" box="[1292,1384,1267,1293]" captionStart="FIGURE 3" captionStartId="7.[151,250,1745,1769]" captionTargetBox="[253,1335,186,1717]" captionTargetId="figure-25@7.[249,1339,181,1721]" captionTargetPageId="7" captionText="FIGURE 3. Mallinella moncheri sp. nov. A, EH holotype male (IFS_ZOD_231) and BE paratype female (IFS_ZOD_232). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734666" httpUri="https://zenodo.org/record/14734666/files/figure.png" pageId="8" pageNumber="108">Figs 3C</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FAAFFB1AFA40FAF7" box="[1397,1432,1267,1293]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="8" pageNumber="108">9E</figureCitation>
;
<figureCitation id="130FF487FF9FFFA7FF4DFAFEFF62FACB" box="[151,186,1303,1329]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">4C</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FF1CFAFEFF34FACB" box="[198,236,1303,1329]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="8" pageNumber="108">9G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9FFFA5FF1DFAD3FCF1FD0B" lastPageId="10" lastPageNumber="110" pageId="8" pageNumber="108" type="description">
<paragraph id="8B8BE802FF9FFFA7FF1DFAD3FCD5F8B7" blockId="8.[151,1437,834,2013]" pageId="8" pageNumber="108">
<emphasis id="B9403410FF9FFFA7FF1DFAD3FE81FAAE" bold="true" box="[199,345,1338,1364]" pageId="8" pageNumber="108">Description.</emphasis>
<emphasis id="B9403410FF9FFFA7FE84FAD2FE40FAAF" box="[350,408,1339,1365]" italics="true" pageId="8" pageNumber="108">Male</emphasis>
in alcohol (
<typeStatus id="548F56A0FF9FFFA7FDCCFAD2FDA5FAAF" box="[534,637,1339,1365]" pageId="8" pageNumber="108" type="holotype">holotype</typeStatus>
;
<figureCitation id="130FF487FF9FFFA7FD5DFAD2FD07FAAF" box="[647,735,1339,1365]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Fig. 4A</figureCitation>
). Body length 4.41; carapace 2.42 long, 1.83 wide; opisthosoma 1.99 long, 1.45 wide. Habitus as in
<figureCitation id="130FF487FF9FFFA7FDC7FAB6FDADFA83" box="[541,629,1375,1401]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Fig. 4A</figureCitation>
. Carapace ovoid, smooth, shiny, dark brown anteriorly; fovea red brown. Clypeus brown, 0.38 high. Eye sizes and inter-distances: AME 0.16, ALE 0.11, PME 0.13, PLE 0.11; AMEAME 0.07, AMEPME 0.13, AMEALE 0.08, ALEALE 0.50, PMEPME 0.10, PMEPLE 0.18, PLEPLE 0.68, ALE PLE 0.04. MOA 0.42 long, front width 0.36, back width 0.39. Chilum unipartite, trapezoid. Chelicerae 0.86 long, light brown, without teeth. Endites yellow. Labium triangular, yellow-brown, 0.20 long, 0.35 wide. Sternum yellow, furnished with sparse black setae, 1.05 long, 0.96 wide. Lateral margins of sternum with small semi-circular pits in front of each coxa of third and fourth pair of legs. Legs yellowish, with proximal tubercle on anterior femora. Measurements of palp and legs: pedipalp (right) 2.20 (0.88 + 0.40 + 0.42 + 0.50) I 5.51 (1. 51 + 0.55 + 1.15 + 1.35+ 0.95), II 5.30 (1.50 + 0.55 + 1.05 + 1.25 + 0.95), III 5.39 (1.35 + 0.54 + 1.10 + 1.50 + 0.90), IV 6.95 (1.60 + 0.50+ 1.65 + 2.10 + 1.10). Leg formula: 4132. Opisthosoma oval; dorsum dark sepia (
<figureCitation id="130FF487FF9FFFA7FBBCF94AFB62F947" box="[1126,1210,1699,1725]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Fig.4A</figureCitation>
). Pattern on dorsum of opisthosoma provided with large pale patches, first and second pairs of patches fused to form reniform patches, followed by third and fourth pairs of oblong patches, fifth pair of patches connected vertically with fourth pair. Dorsum of opisthosoma with lanceolate scutum. Posterior ventral spines moderate in length, gradually tapering, with blunt apices (
<figureCitation id="130FF487FF9FFFA7FEBCF8DAFE66F8B7" box="[358,446,1843,1869]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Fig. 4E</figureCitation>
). Spinnerets yellow (
<figureCitation id="130FF487FF9FFFA7FD73F8DAFCD9F8B7" box="[681,769,1843,1869]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="8B8BE802FF9FFFA5FF1DF8BEFF0CFEE7" blockId="8.[151,1437,834,2013]" lastBlockId="10.[151,1437,151,789]" lastPageId="10" lastPageNumber="110" pageId="8" pageNumber="108">
<emphasis id="B9403410FF9FFFA7FF1DF8BEFF24F88B" box="[199,252,1879,1905]" italics="true" pageId="8" pageNumber="108">Palp</emphasis>
(
<figureCitation id="130FF487FF9FFFA7FED1F8BEFE5FF88B" box="[267,391,1879,1905]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Figs 4FH</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FE48F8BEFE60F88B" box="[402,440,1879,1905]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="8" pageNumber="108">8D</figureCitation>
). VTA triangular hump (
<figureCitation id="130FF487FF9FFFA7FD16F8BEFCFFF88B" box="[716,807,1879,1905]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Fig. 4G</figureCitation>
); RTA digitiform, tapered towards blunt apex (
<figureCitation id="130FF487FF9FFFA7FAE9F8BEFF76F86F" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Figs 4G H</figureCitation>
,
<figureCitation id="130FF487FF9FFFA7FF63F892FF05F86F" box="[185,221,1915,1941]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="8" pageNumber="108">8D</figureCitation>
), Cymbial fold less prominent, less than half the length of cymbium (
<figureCitation id="130FF487FF9FFFA7FC3AF892FBE2F86F" box="[992,1082,1915,1941]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="8" pageNumber="108">Fig. 4G</figureCitation>
). TA elongated, Apicoprolateral fold of TA triangular, apicoprolateral process of TA triangular, broad at base, with blunt apex, directed posteromesad (
<figureCitation id="130FF487FF9FFFA7FF45F82AFF25F827" box="[159,253,1987,2013]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="8" pageNumber="108">Fig. 8D</figureCitation>
). Apicoretrolateral fold of TA triangular, with blunt apex. Mesoprolateral fold of TA, short, triangular (
<figureCitation id="130FF487FF9DFFA5FF45FF7EFF22FF4B" box="[159,250,151,177]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">Fig. 8D</figureCitation>
). Baso-prolateral fold of TA large, directed posteriad. Tegular spine large, prolaterally directed (
<figureCitation id="130FF487FF9DFFA5FAEFFF7EFA57FF4B" box="[1333,1423,151,177]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">Fig. 8D</figureCitation>
). Embolic base aligned diagonally, directed posteriorly, with broad membranous area (
<figureCitation id="130FF487FF9DFFA5FB83FF52FB6EFF2F" box="[1113,1206,187,213]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="10" pageNumber="110">Fig. 4G</figureCitation>
). Embolus branched sub-terminally, with long lateral ramus. Both rami with blunt apices. (
<figureCitation id="130FF487FF9DFFA5FC70FF36FBDCFF03" box="[938,1028,223,249]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="10" pageNumber="110">Fig. 4G</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FBCAFF36FBEEFF03" box="[1040,1078,223,249]" captionStart="FIGURE 8" captionStartId="13.[151,250,1711,1735]" captionTargetBox="[156,1430,186,1683]" captionTargetId="figure-25@13.[151,1435,181,1688]" captionTargetPageId="13" captionText="FIGURE 8. AE. Mallinella spp., male left palps, ventral. A Mallinella dhanahami sp. nov. (IFS_ZOD_234). B Mallinella milkyway sp. nov. (IFS_ZOD_36). C Mallinella moncheri sp. nov. (IFS_ZOD_231). D Mallinella oreo sp. nov. (IFS_ZOD_ 137). E Mallinella truffles sp. nov. (IFS_ZOD_43). Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; E, embolus; EB, embolic base; LR, lateral ramus; MPF, meso prolateral fold; MR, mesal ramus; pC, prolateral extension of conductor; RTA, retrolateral tibial apophysis; T, tegulum; TA, tegular apophysis; TS, tegular spine; TT, tegular tubercle; VTA, ventral tibial apophysis. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734678" httpUri="https://zenodo.org/record/14734678/files/figure.png" pageId="10" pageNumber="110">8D</figureCitation>
). Embolus elongated with blunt apex.
</paragraph>
<caption id="DF4BB88AFF9EFFA6FF4DF95BFF34F810" ID-DOI="http://doi.org/10.5281/zenodo.14734668" ID-Zenodo-Dep="14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="9" pageNumber="109" startId="9.[151,250,1714,1738]" targetBox="[266,1321,185,1686]" targetPageId="9" targetType="figure">
<paragraph id="8B8BE802FF9EFFA6FF4DF95BFF34F810" blockId="9.[151,1437,1714,2026]" pageId="9" pageNumber="109">
<emphasis id="B9403410FF9EFFA6FF4DF95BFECBF930" bold="true" box="[151,275,1714,1738]" pageId="9" pageNumber="109">FIGURE 4.</emphasis>
<taxonomicName id="4C349381FF9EFFA6FEC0F95BFE6FF930" authorityName="Dayananda &amp; Benjamin" authorityYear="2025" box="[282,439,1714,1738]" class="Arachnida" family="Zodariidae" genus="Mallinella" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="109" phylum="Arthropoda" rank="species" species="oreo" status="sp. nov.">
<emphasis id="B9403410FF9EFFA6FEC0F95BFE6FF930" box="[282,439,1714,1738]" italics="true" pageId="9" pageNumber="109">Mallinella oreo</emphasis>
</taxonomicName>
<emphasis id="B9403410FF9EFFA6FE64F95BFDD7F930" bold="true" box="[446,527,1714,1738]" pageId="9" pageNumber="109">
<taxonomicNameLabel id="A273896BFF9EFFA6FE64F95BFDD7F930" box="[446,527,1714,1738]" pageId="9" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8BE802FF9DFFA5FF1DFECEFEA9FD9B" blockId="10.[151,1437,151,789]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DFECEFEC3FEBB" box="[199,283,295,321]" italics="true" pageId="10" pageNumber="110">Female</emphasis>
<materialsCitation id="3B5CE25FFF9DFFA5FEFFFECEFEB5FD9B" collectionCode="AME, ALE, PME, PLE, MOA" pageId="10" pageNumber="110" specimenCount="1" typeStatus="paratype">
in alcohol (
<typeStatus id="548F56A0FF9DFFA5FE73FECEFDD7FEBB" box="[425,527,295,321]" pageId="10" pageNumber="110" type="paratype">paratype</typeStatus>
;
<figureCitation id="130FF487FF9DFFA5FDC7FECEFDAFFEBB" box="[541,631,295,321]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="10" pageNumber="110">Fig. 4B</figureCitation>
). Body length 5.22; carapace 2.46 long, 1.67 wide; opisthosoma 2.76 long, 2.03 wide. Habitus and details as in male except for the following (
<figureCitation id="130FF487FF9DFFA5FC3AFEA2FBE3FE9F" box="[992,1083,331,357]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="10" pageNumber="110">Fig. 4B</figureCitation>
). Clypeus 0.34 high. Eye sizes and inter-distances:
<collectionCode id="ED2570C7FF9DFFA5FEA0FE86FE60FE73" box="[378,440,367,393]" pageId="10" pageNumber="110">AME</collectionCode>
0.11,
<collectionCode id="ED2570C7FF9DFFA5FE25FE86FDEDFE73" box="[511,565,367,393]" pageId="10" pageNumber="110">ALE</collectionCode>
0.09,
<collectionCode id="ED2570C7FF9DFFA5FDA4FE86FD60FE73" box="[638,696,367,393]" pageId="10" pageNumber="110">PME</collectionCode>
0.11,
<collectionCode id="ED2570C7FF9DFFA5FCDAFE86FCEAFE73" box="[768,818,367,393]" pageId="10" pageNumber="110">PLE</collectionCode>
0.09;
<collectionCode id="ED2570C7FF9DFFA5FCA0FE86FC60FE73" box="[890,952,367,393]" pageId="10" pageNumber="110">AME</collectionCode>
<collectionCode id="ED2570C7FF9DFFA5FC1CFE86FBDCFE73" box="[966,1028,367,393]" pageId="10" pageNumber="110">AME</collectionCode>
0.06,
<collectionCode id="ED2570C7FF9DFFA5FB96FE86FB52FE73" box="[1100,1162,367,393]" pageId="10" pageNumber="110">AME</collectionCode>
<collectionCode id="ED2570C7FF9DFFA5FB42FE86FB0AFE73" box="[1176,1234,367,393]" pageId="10" pageNumber="110">PME</collectionCode>
0.15,
<collectionCode id="ED2570C7FF9DFFA5FAC0FE86FA80FE73" box="[1306,1368,367,393]" pageId="10" pageNumber="110">AME</collectionCode>
<collectionCode id="ED2570C7FF9DFFA5FABCFE86FA44FE73" box="[1382,1436,367,393]" pageId="10" pageNumber="110">ALE</collectionCode>
0.11,
<collectionCode id="ED2570C7FF9DFFA5FF0FFE7AFED3FE57" box="[213,267,403,429]" pageId="10" pageNumber="110">ALE</collectionCode>
<collectionCode id="ED2570C7FF9DFFA5FEC0FE7AFE88FE57" box="[282,336,403,429]" pageId="10" pageNumber="110">ALE</collectionCode>
0.50,
<collectionCode id="ED2570C7FF9DFFA5FE40FE7AFE0CFE57" box="[410,468,403,429]" pageId="10" pageNumber="110">PME</collectionCode>
<collectionCode id="ED2570C7FF9DFFA5FE38FE7AFDC4FE57" box="[482,540,403,429]" pageId="10" pageNumber="110">PME</collectionCode>
0.10,
<collectionCode id="ED2570C7FF9DFFA5FDBFFE7AFD46FE57" box="[613,670,403,429]" pageId="10" pageNumber="110">PME</collectionCode>
<collectionCode id="ED2570C7FF9DFFA5FD76FE7AFD06FE57" box="[684,734,403,429]" pageId="10" pageNumber="110">PLE</collectionCode>
0.19,
<collectionCode id="ED2570C7FF9DFFA5FCF2FE7AFC82FE57" box="[808,858,403,429]" pageId="10" pageNumber="110">PLE</collectionCode>
<collectionCode id="ED2570C7FF9DFFA5FCB2FE7AFC42FE57" box="[872,922,403,429]" pageId="10" pageNumber="110">PLE</collectionCode>
0.64,
<collectionCode id="ED2570C7FF9DFFA5FC38FE7AFBC0FE57" box="[994,1048,403,429]" pageId="10" pageNumber="110">ALE</collectionCode>
<collectionCode id="ED2570C7FF9DFFA5FBFCFE7AFB80FE57" box="[1062,1112,403,429]" pageId="10" pageNumber="110">PLE</collectionCode>
0.06.
<collectionCode id="ED2570C7FF9DFFA5FB78FE7AFB3AFE57" box="[1186,1250,403,429]" pageId="10" pageNumber="110">MOA</collectionCode>
0.39 long, front width 0.28, back width 0.32. Chelicerae 0.83 long Labium 0.30 long, 0.44 wide. Sternum yellow with shallow semi-circular pits, 1.06 long, 1.04 wide. Legs with proximal tubercle on anterior femora (
<figureCitation id="130FF487FF9DFFA5FBE2FE32FB4AFE0F" box="[1080,1170,475,501]" captionStart="FIGURE 6" captionStartId="12.[151,250,1167,1191]" captionTargetBox="[152,1435,705,1143]" captionTargetId="figure-244@12.[151,1435,705,1143]" captionTargetPageId="12" captionText="FIGURE 6. Mallinella spp., Legs:A, B, C, D, E, anterior femora; F, G, H, I, J, sternum, ventral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: CP, semicircular pits on sternum; PT, posterior tubercle. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734674" httpUri="https://zenodo.org/record/14734674/files/figure.png" pageId="10" pageNumber="110">Fig. 6D</figureCitation>
). Measurements of palp and legs: pedipalp (right) 1.98 (0.77 + 0.37 + 0.38 + 0.46), I 5.00 (1.40 + 0.45 + 1.20 + 1.00 + 0.95), II 4.85 (1.30 + 0.45 + 1.25 + 0.95 + 0.90), III 4.95 (1.35 + 0.45+ 1.15 + 1.10 + 0.90), IV 6.75 (1.65 + 0.50 + 1.50 + 2.00 + 1.10). Leg formula: 4132
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BE802FF9DFFA5FF1DFD82FCF1FD0B" blockId="10.[151,1437,151,789]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DFD82FEE9FD7F" box="[199,305,619,645]" italics="true" pageId="10" pageNumber="110">Genitalia</emphasis>
(
<figureCitation id="130FF487FF9DFFA5FE9AFD82FE67FD7F" box="[320,447,619,645]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="10" pageNumber="110">Figs 4CD</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FE10FD82FDCCFD7F" box="[458,532,619,645]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="10" pageNumber="110">7D, 7I</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FDFAFD82FDB1FD7F" box="[544,617,619,645]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="10" pageNumber="110">9GH</figureCitation>
). Epigynal plate, semi-circular, small median depression. Lateral borders digitiform, terminally blunt (
<figureCitation id="130FF487FF9DFFA5FE0EFD66FDF5FD53" box="[468,557,655,681]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="10" pageNumber="110">Fig. 9G</figureCitation>
). Both epigynal plate and furrow highly sclerotized (
<figureCitation id="130FF487FF9DFFA5FBB4FD66FB10FD53" box="[1134,1224,655,681]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="10" pageNumber="110">Figs 4C</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FB08FD66FB20FD53" box="[1234,1272,655,681]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="10" pageNumber="110">9G</figureCitation>
). Insemination ducts long, thick, sclerotized, distally diverging (
<figureCitation id="130FF487FF9DFFA5FD69FD5AFCD7FD37" box="[691,783,691,717]" captionStart="FIGURE 4" captionStartId="9.[151,250,1714,1738]" captionTargetBox="[266,1321,185,1686]" captionTargetId="figure-25@9.[262,1325,181,1690]" captionTargetPageId="9" captionText="FIGURE 4. Mallinella oreo sp. nov. A, EH holotype male (IFS_ZOD_137) and BE paratype female (IFS_ZOD_138). AB habitus, dorsal. C epigyne intact, ventral. D vulva, ventral; E posterior ventral spines, ventral. F left palp, prolateral; G idem, ventral; H idem, retrolateral. Abbreviations: APF, apicoprolateral fold of tegular apophysis; APP, apicoprolateral process of tegular apophysis; BPF, basoprolateral fold of tegular apophysis; C, conductor; Cy, cymbium; CF, cymbial fold; DB, dark bands; DS, dorsal scutum; E, embolus; EB, embolic base; EF, epigynal furrow; EP, epigynal plate; F, fovea; ID, insemination ducts; LB, lateral border of the epigyne; LR, lateral ramus of embolus; MR, mesal ramus of embolus; mEB, membranous area of the embolic base; pC, prolateral extension of conductor; PVS, posterior ventral spines; RTA, retrolateral tibial apophysis; S, spermatheca; Sp, spinnerets; T, tegulum; TS, Tegular spine; VTA, ventral tibial apophysis. Scale lines: AB = 0.5 mm; CH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734668" httpUri="https://zenodo.org/record/14734668/files/figure.png" pageId="10" pageNumber="110">Figs 4D</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FCC3FD5AFCB9FD37" box="[793,865,691,717]" captionStart="FIGURE 7" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[155,1435,1326,1857]" captionTargetId="figure-305@12.[151,1435,1323,1861]" captionTargetPageId="12" captionText="FIGURE 7. Mallinella spp., female genitalia: A, B, C, D, E, vulva ventral; F, G, H, I, J, vulva, retrolateral AF Mallinella dhanahami sp. nov. BG Mallinella milkyway sp. nov. CH Mallinella moncheri sp. nov. DI Mallinella oreo sp. nov. EJ Mallinella truffles sp. nov. Abbreviations: FD, fertilization duct; ID, insemination ducts; S, spermathecae. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734676" httpUri="https://zenodo.org/record/14734676/files/figure.png" pageId="10" pageNumber="110">7D, 7I</figureCitation>
,
<figureCitation id="130FF487FF9DFFA5FCB7FD5AFC4DFD37" box="[877,917,691,717]" captionStart="FIGURE 9" captionStartId="14.[151,250,1874,1898]" captionTargetBox="[228,1359,186,1846]" captionTargetId="figure-17@14.[223,1364,181,1850]" captionTargetPageId="14" captionText="FIGURE 9. Mallinella spp., female genitalia: A, C, E, G, I, epigyne, ventral; B, D, F, H, J, vulvae, ventral. AB Mallinella dhanahami sp. nov. (IFS_ZOD_235). CD Mallinella milkyway sp. nov. (IFS_ZOD_37). EF Mallinella moncheri sp. nov. (IFS_ZOD_232).GH Mallinellaoreo sp. nov. (IFS_ZOD_138).IJ Mallinellatruffles sp.nov. (IFS_ZOD_230).Abbreviations: EF, epigynal furrow; EP, epigynal plate; ID, insemination duct; LB, lateral border; S, spermatheca. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14734681" httpUri="https://zenodo.org/record/14734681/files/figure.png" pageId="10" pageNumber="110">9H</figureCitation>
.). Insemination ducts longer than spermathecal length. Spermathecae round, fertilization ducts as in Fig. I.
</paragraph>
</subSubSection>
<subSubSection id="C32EBB89FF9DFFA5FF1DFD13FC84FCEF" box="[199,860,762,789]" pageId="10" pageNumber="110" type="distribution">
<paragraph id="8B8BE802FF9DFFA5FF1DFD13FC84FCEF" blockId="10.[151,1437,151,789]" box="[199,860,762,789]" pageId="10" pageNumber="110">
<emphasis id="B9403410FF9DFFA5FF1DFD13FEB9FCEE" bold="true" box="[199,353,762,788]" pageId="10" pageNumber="110">Distribution.</emphasis>
Known only from the
<typeStatus id="548F56A0FF9DFFA5FDBAFD12FD48FCEF" box="[608,656,763,789]" pageId="10" pageNumber="110">type</typeStatus>
locality (
<figureCitation id="130FF487FF9DFFA5FD21FD12FC96FCEF" box="[763,846,763,789]" captionStart="FIGURE 12" captionStartId="17.[151,250,1133,1157]" captionText="FIGURE 12. Distribution map of known Mallinella species of Sri Lanka." pageId="10" pageNumber="110">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,166 @@
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<mods:title id="6B36BABDD495D14D11B7C6272E8B49AB">A new species of the genus Puto Signoret (Hemiptera: Coccomorpha: Putoidae) from China and redescription of Puto huangshanensis Wu</mods:title>
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<mods:namePart id="0B677A933EA0B932B4834BB3AADFEA74">Zheng, Xinyi</mods:namePart>
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<mods:namePart id="6283A56C07D1AD0CA5E2B60531E8D9F2">Li, Yuang</mods:namePart>
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<mods:namePart id="DC594EBDCBFC6BD890AC58BBDC935F3D">Tan, Zhixiang</mods:namePart>
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<mods:namePart id="954E0D32AC06C44A449FEEC11115D028">Wu, SanAn</mods:namePart>
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<taxonomicName id="4C644EA2232DFF84FF5C578DFE50FA82" authority="Zheng &amp; Wu" authorityName="Zheng &amp; Wu" authorityYear="2025" box="[151,485,1266,1293]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="sinensis" status="sp. nov.">
<emphasis id="B910E933232DFF84FF5C578DFE8DFA82" bold="true" box="[151,312,1267,1293]" italics="true" pageId="8" pageNumber="127">Puto sinensis</emphasis>
Zheng &amp; Wu
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,
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<emphasis id="B910E933232DFF84FF5C5644FDB1FADA" bold="true" box="[151,516,1338,1365]" pageId="8" pageNumber="127">
Material examined.
<typeStatus id="54DF8B83232DFF84FE465645FE4BFADA" box="[397,510,1339,1365]" pageId="8" pageNumber="127" type="holotype">Holotype</typeStatus>
:
</emphasis>
Adult
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;
<collectingCountry id="F37375B1232DFF84FDB55645FD6EFADA" box="[638,731,1339,1365]" name="China" pageId="8" pageNumber="127">CHINA</collectingCountry>
:
<collectingRegion id="49A0FBC3232DFF84FD235645FC25FADA" box="[744,912,1339,1365]" country="China" name="Guizhou" pageId="8" pageNumber="127">Guizhou Prov.</collectingRegion>
, / Guiyang City, Kaiyang County, / Nanjiang Grand Canyon, 26°9556″N, 106º9874″E, / on
<taxonomicName id="4C644EA2232DFF84FD645621FC37FAF7" box="[687,898,1375,1401]" class="Magnoliopsida" family="Lauraceae" genus="Lindera" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="127" phylum="Tracheophyta" rank="species" species="communis">
<emphasis id="B910E933232DFF84FD645621FC37FAF7" box="[687,898,1375,1401]" italics="true" pageId="8" pageNumber="127">Lindera communis</emphasis>
</taxonomicName>
(
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), /
<date id="FFDA13E1232DFF84FBE65621FB0FFAF6" box="[1069,1210,1375,1401]" pageId="8" pageNumber="127" value="2023-05-01">1 May 2023</date>
, / Zhixiang Tan leg.
<typeStatus id="54DF8B83232DFF84FF5C56FDFF4AFA12" box="[151,255,1411,1437]" pageId="8" pageNumber="127" type="holotype">Holotype</typeStatus>
mounted together with
<specimenCount id="9D62FEA8232DFF84FDC456FDFD17FA12" box="[527,674,1411,1437]" count="1" pageId="8" pageNumber="127" type="female" typeStatus="paratype">1 paratype ♀</specimenCount>
on 1 slide (
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);
<typeStatus id="54DF8B83232DFF84FC4656FDFC5AFA12" box="[909,1007,1411,1437]" pageId="8" pageNumber="127" type="holotype">holotype</typeStatus>
specimen is indicated with a circle in permanent marker on the coverslip.
</paragraph>
<paragraph id="8BDB3521232DFF84FF0C56B5FA2FF9FA" blockId="8.[151,1437,1338,1725]" pageId="8" pageNumber="127">
<emphasis id="B910E933232DFF84FF0C56B5FEFDFA6A" bold="true" box="[199,328,1483,1509]" pageId="8" pageNumber="127">
<typeStatus id="54DF8B83232DFF84FF0C56B5FEF6FA6A" box="[199,323,1483,1509]" pageId="8" pageNumber="127" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="F37375B1232DFF84FE9A56B5FE1BFA6A" box="[337,430,1483,1509]" name="China" pageId="8" pageNumber="127">CHINA</collectingCountry>
: same data as
<typeStatus id="54DF8B83232DFF84FD9F56B5FD0CFA6A" box="[596,697,1483,1509]" pageId="8" pageNumber="127" type="holotype">holotype</typeStatus>
,
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: 1 mounted together with
<typeStatus id="54DF8B83232DFF84FBF356B5FB2FFA6A" box="[1080,1178,1483,1509]" pageId="8" pageNumber="127" type="holotype">holotype</typeStatus>
on 1 slide, 1 mounted singly on 1 slide, and 2 slides each containing
<specimenCount id="9D62FEA8232DFF84FD605691FD63F986" box="[683,726,1519,1545]" count="2" pageId="8" pageNumber="127" type="female">2 ♀</specimenCount>
mounted together (
<collectionCode id="ED75ADE4232DFF84FC7F5691FBB6F986" box="[948,1027,1519,1545]" pageId="8" pageNumber="127">BFUC</collectionCode>
); also
<specimenCount id="9D62FEA8232DFF84FB865691FB24F986" box="[1101,1169,1519,1545]" count="3" pageId="8" pageNumber="127" type="male">3 ♂♂</specimenCount>
: 2 mounted singly on 2 slides and 1 preserved in glycerine; 4 forewings mounted together on 1 slide, and 2 forewings mounted together on 1 slide.
<collectingCountry id="F37375B1232DFF84FF3A5549FEFBF9DE" box="[241,334,1591,1617]" name="China" pageId="8" pageNumber="127">CHINA</collectingCountry>
:
<collectingRegion id="49A0FBC3232DFF84FE915549FD97F9DE" box="[346,546,1591,1617]" country="China" name="Guangdong" pageId="8" pageNumber="127">Guangdong Prov.</collectingRegion>
, Shaoguan City, Yao Autonomous County of Ruyuan, Nanling National Forest Park, on
<taxonomicName id="4C644EA2232DFF84FF375525FE54F9FA" box="[252,481,1627,1653]" class="Magnoliopsida" family="Rosaceae" genus="Photinia" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="127" phylum="Tracheophyta" rank="species" species="serratifolia">
<emphasis id="B910E933232DFF84FF375525FE54F9FA" box="[252,481,1627,1653]" italics="true" pageId="8" pageNumber="127">Photinia serratifolia</emphasis>
</taxonomicName>
(
<taxonomicName id="4C644EA2232DFF84FE3B5525FDD5F9FA" box="[496,608,1627,1653]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="127" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
), San-an Wu leg.,
<date id="FFDA13E1232DFF84FCE75525FC12F9FA" box="[812,935,1627,1653]" pageId="8" pageNumber="127" value="2021-04-20">20.iv.2021</date>
:
<specimenCount id="9D62FEA8232DFF84FC7A5525FC45F9FA" box="[945,1008,1627,1653]" count="2" pageId="8" pageNumber="127" type="female">2 ♀♀</specimenCount>
mounted together on 1 slide (
<collectionCode id="ED75ADE4232DFF84FAF55525FA38F9FA" box="[1342,1421,1627,1653]" pageId="8" pageNumber="127">BFUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E66AA232DFF84FF0C5501FC21F932" pageId="8" pageNumber="127" type="etymology">
<paragraph id="8BDB3521232DFF84FF0C5501FC21F932" blockId="8.[151,1437,1338,1725]" pageId="8" pageNumber="127">
<emphasis id="B910E933232DFF84FF0C5501FEFBF916" bold="true" box="[199,334,1663,1689]" pageId="8" pageNumber="127">Etymology.</emphasis>
The specific name is formed by combination of the Latin prefix
<emphasis id="B910E933232DFF84FB9E5501FBC3F917" box="[1109,1142,1663,1688]" italics="true" pageId="8" pageNumber="127">sin</emphasis>
-, referring to
<collectingCountry id="F37375B1232DFF84FAED5501FAD9F916" box="[1318,1388,1663,1689]" name="China" pageId="8" pageNumber="127">China</collectingCountry>
, the country of collection, with the Latin suffix -
<emphasis id="B910E933232DFF84FD4655DDFD70F933" box="[653,709,1699,1724]" italics="true" pageId="8" pageNumber="127">ensis</emphasis>
, meaning from.
</paragraph>
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</treatment>
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@ -1,69 +1,70 @@
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@ -81,7 +82,7 @@
<subSubSection id="C34105D0FFC2E243FF392081FECBF884" box="[151,309,1771,1797]" pageId="24" pageNumber="25" type="description">
<paragraph id="8BE4565BFFC2E243FF392081FECBF884" blockId="24.[151,562,1734,1797]" box="[151,309,1771,1797]" pageId="24" pageNumber="25">
(
<figureCitation id="13604ADEFFC2E243FF312081FED3F884" box="[159,301,1771,1797]" captionStart-0="FIGURES 99101" captionStart-1="FIGURES 102106" captionStart-2="FIGURES 107116" captionStart-3="FIGURES 117122" captionStartId-0="25.[151,264,1922,1946]" captionStartId-1="26.[151,264,1885,1909]" captionStartId-2="27.[151,264,1866,1890]" captionStartId-3="28.[151,264,1074,1098]" captionTargetBox-0="[166,1424,250,1891]" captionTargetBox-1="[172,1417,209,1854]" captionTargetBox-2="[158,1429,209,1834]" captionTargetBox-3="[159,1423,212,1042]" captionTargetId-0="figure-58@25.[153,1434,237,1899]" captionTargetId-1="figure-27@26.[158,1430,199,1862]" captionTargetId-2="figure-18@27.[151,1436,199,1842]" captionTargetId-3="figure-27@28.[151,1436,199,1051]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionText-0="FIGURES 99101. Calomera bordonii Wiesner (all from Maluku, Buru Island, Waedea). 99100—habitus: 99—♂, 14.2 mm, HT (JWCM); 100—♀, 14.5 mm, PT (JWCM); 101—labels of the HT. Bar = 1 mm (photography P. Schüle)." captionText-1="FIGURES 102106. Calomera bordonii Wiesner (all from Buru Island, Waedea). 102—♀, habitus, 15.1 mm, PT (ABCF); 103—104—pronotum: 103—♀, PT (ABCF); 104—♂, HT (JWCM); 105—♂ aedeagus, HT; 106—♂, elytron, HT. Bars = 1 mm (photography P. Schüle)" captionText-2="FIGURES 107116. Calomera bordonii Wiesner (all from Buru Island). 107—head, ♂, Remals Mts., Ilat (CCJM); 108111— labrum: 108109—♂, ibid. (CCJM); 110—♀, ibid. (CCJM); 111—♀, Waedea, PT (JWCM); 112113—pronotum: 112—♂, Remals Mts., Ilat (CCJM); 113—♀, Waedea, PT (JWCM); 114116—elytron: 114—♂, Remals Mts., Ilat (CCJM); 115—♀, ibid. (CCJM); 116—♀, Waedea, PT (JWCM). Bars= 1 mm." captionText-3="FIGURES 117122. Calomera bordonii Wiesner, male aedeagus or its apex (all from Maluku, Buru Island, Remals Mts., Ilat, CCJM). 117—left lateral view; 118—dorsal view; 119—ventral view; 120—left lateral view; 121122 —internal sac in left and right lateral view. Bar = 1. mm." pageId="24" pageNumber="25">Figs 99122</figureCitation>
<figureCitation id="13604ADEFFC2E243FF312081FED3F884" box="[159,301,1771,1797]" captionStart-0="FIGURES 99101" captionStart-1="FIGURES 102106" captionStart-2="FIGURES 107116" captionStart-3="FIGURES 117122" captionStartId-0="25.[151,264,1922,1946]" captionStartId-1="26.[151,264,1885,1909]" captionStartId-2="27.[151,264,1866,1890]" captionStartId-3="28.[151,264,1074,1098]" captionTargetBox-0="[166,1424,250,1891]" captionTargetBox-1="[172,1417,209,1854]" captionTargetBox-2="[158,1429,209,1834]" captionTargetBox-3="[159,1423,212,1042]" captionTargetId-0="figure-58@25.[153,1434,237,1899]" captionTargetId-1="figure-27@26.[158,1430,199,1862]" captionTargetId-2="figure-18@27.[151,1436,199,1842]" captionTargetId-3="figure-27@28.[151,1436,199,1051]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionText-0="FIGURES 99101. Calomera bordonii Wiesner (all from Maluku, Buru Island, Waedea). 99100—habitus: 99—♂, 14.2 mm, HT (JWCM); 100—♀, 14.5 mm, PT (JWCM); 101—labels of the HT. Bar = 1 mm (photography P. Schüle)." captionText-1="FIGURES 102106. Calomera bordonii Wiesner (all from Buru Island, Waedea). 102—♀, habitus, 15.1 mm, PT (ABCF); 103—104—pronotum: 103—♀, PT (ABCF); 104—♂, HT (JWCM); 105—♂ aedeagus, HT; 106—♂, elytron, HT. Bars = 1 mm (photography P. Schüle)" captionText-2="FIGURES 107116. Calomera bordonii Wiesner (all from Buru Island). 107—head, ♂, Remals Mts., Ilat (CCJM); 108111— labrum: 108109—♂, ibid. (CCJM); 110—♀, ibid. (CCJM); 111—♀, Waedea, PT (JWCM); 112113—pronotum: 112—♂, Remals Mts., Ilat (CCJM); 113—♀, Waedea, PT (JWCM); 114116—elytron: 114—♂, Remals Mts., Ilat (CCJM); 115—♀, ibid. (CCJM); 116—♀, Waedea, PT (JWCM). Bars= 1 mm." captionText-3="FIGURES 117122. Calomera bordonii Wiesner, male aedeagus or its apex (all from Maluku, Buru Island, Remals Mts., Ilat, CCJM). 117—left lateral view; 118—dorsal view; 119—ventral view; 120—left lateral view; 121122 —internal sac in left and right lateral view. Bar = 1. mm." figureDoi-0="http://doi.org/10.5281/zenodo.14734518" figureDoi-1="http://doi.org/10.5281/zenodo.14734520" figureDoi-2="http://doi.org/10.5281/zenodo.14734522" figureDoi-3="http://doi.org/10.5281/zenodo.14734524" httpUri-0="https://zenodo.org/record/14734518/files/figure.png" httpUri-1="https://zenodo.org/record/14734520/files/figure.png" httpUri-2="https://zenodo.org/record/14734522/files/figure.png" httpUri-3="https://zenodo.org/record/14734524/files/figure.png" pageId="24" pageNumber="25">Figs 99122</figureCitation>
)
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@ -199,7 +200,7 @@ labelled: “
</materialsCitation>
.
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<caption id="DF2406D3FFC3E242FF3921E8FB1FF83F" ID-DOI="http://doi.org/10.5281/zenodo.14734518" ID-Zenodo-Dep="14734518" httpUri="https://zenodo.org/record/14734518/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,264,1922,1946]" targetBox="[166,1424,250,1891]" targetPageId="25" targetType="figure">
<paragraph id="8BE4565BFFC3E242FF3921E8FB1FF83F" blockId="25.[151,1436,1922,1983]" pageId="25" pageNumber="26">
<emphasis id="B92F8A49FFC3E242FF3921E8FE9FF81B" bold="true" box="[151,353,1922,1946]" pageId="25" pageNumber="26">FIGURES 99101.</emphasis>
<taxonomicName id="4C5B2DD8FFC3E242FEC921E8FD7DF81B" authority="Wiesner" authorityName="Wiesner" authorityYear="2018" box="[359,643,1922,1946]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="bordonii">
@ -209,7 +210,7 @@ Wiesner
(all from Maluku, Buru Island, Waedea). 99100—habitus: 99—♂, 14.2 mm, HT (JWCM); 100—♀, 14.5 mm, PT (JWCM); 101—labels of the HT. Bar = 1 mm (photography P. Schüle).
</paragraph>
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<paragraph id="8BE4565BFFC0E241FF392137FE75F83C" blockId="26.[151,1436,1885,1981]" pageId="26" pageNumber="27">
<emphasis id="B92F8A49FFC0E241FF392137FE8EF8F4" bold="true" box="[151,368,1885,1909]" pageId="26" pageNumber="27">FIGURES 102106.</emphasis>
<taxonomicName id="4C5B2DD8FFC0E241FED72137FD67F8F4" authority="Wiesner" authorityName="Wiesner" authorityYear="2018" box="[377,665,1885,1909]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bordonii">
@ -219,7 +220,7 @@ Wiesner
(all from Buru Island, Waedea). 102—♀, habitus, 15.1 mm, PT (ABCF); 103—104—pronotum: 103—♀, PT (ABCF); 104—♂, HT (JWCM); 105—♂ aedeagus, HT; 106—♂, elytron, HT. Bars = 1 mm (photography P. Schüle)
</paragraph>
</caption>
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<caption id="DF2406D3FFC1E240FF392120FD0AF84C" ID-DOI="http://doi.org/10.5281/zenodo.14734522" ID-Zenodo-Dep="14734522" httpUri="https://zenodo.org/record/14734522/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,264,1866,1890]" targetBox="[158,1429,209,1834]" targetPageId="27" targetType="figure">
<paragraph id="8BE4565BFFC1E240FF392120FD0AF84C" blockId="27.[151,1436,1866,1998]" pageId="27" pageNumber="28">
<emphasis id="B92F8A49FFC1E240FF392120FE92F8E3" bold="true" box="[151,364,1866,1890]" pageId="27" pageNumber="28">FIGURES 107116.</emphasis>
<taxonomicName id="4C5B2DD8FFC1E240FEDF2120FD72F8E3" authority="Wiesner" authorityName="Wiesner" authorityYear="2018" box="[369,652,1866,1890]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bordonii">
@ -229,7 +230,7 @@ Wiesner
(all from Buru Island). 107—head, ♂, Remals Mts., Ilat (CCJM); 108111— labrum: 108109—♂, ibid. (CCJM); 110—♀, ibid. (CCJM); 111—♀, Waedea, PT (JWCM); 112113—pronotum: 112—♂, Remals Mts., Ilat (CCJM); 113—♀, Waedea, PT (JWCM); 114116—elytron: 114—♂, Remals Mts., Ilat (CCJM); 115—♀, ibid. (CCJM); 116—♀, Waedea, PT (JWCM). Bars= 1 mm.
</paragraph>
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<caption id="DF2406D3FFC6E247FF392258FE29FB13" ID-DOI="http://doi.org/10.5281/zenodo.14734524" ID-Zenodo-Dep="14734524" httpUri="https://zenodo.org/record/14734524/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,264,1074,1098]" targetBox="[159,1423,212,1042]" targetPageId="28" targetType="figure">
<paragraph id="8BE4565BFFC6E247FF392258FE29FB13" blockId="28.[151,1437,1074,1170]" pageId="28" pageNumber="29">
<emphasis id="B92F8A49FFC6E247FF392258FE92FBCB" bold="true" box="[151,364,1074,1098]" pageId="28" pageNumber="29">FIGURES 117122.</emphasis>
<taxonomicName id="4C5B2DD8FFC6E247FEDD2258FD6EFBCB" authority="Wiesner" authorityName="Wiesner" authorityYear="2018" box="[371,656,1074,1098]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bordonii">
@ -618,19 +619,19 @@ possesses a similar pattern of its elytral maculation with
<emphasis id="B92F8A49FFC6E247FDFD21B3FD2AF873" box="[595,724,2008,2035]" italics="true" pageId="28" pageNumber="29">C. bordonii</emphasis>
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is rounded or at least always shorter, and female upper-discal catoptric area indistinct (
<figureCitation id="13604ADEFFC7E246FE0426FDFDF3FF30" box="[426,525,151,177]" captionStart="FIGURES 99101" captionStartId="25.[151,264,1922,1946]" captionTargetBox="[166,1424,250,1891]" captionTargetId="figure-58@25.[153,1434,237,1899]" captionTargetPageId="25" captionText="FIGURES 99101. Calomera bordonii Wiesner (all from Maluku, Buru Island, Waedea). 99100—habitus: 99—♂, 14.2 mm, HT (JWCM); 100—♀, 14.5 mm, PT (JWCM); 101—labels of the HT. Bar = 1 mm (photography P. Schüle)." pageId="29" pageNumber="30">Figs 100</figureCitation>
<figureCitation id="13604ADEFFC7E246FE0426FDFDF3FF30" box="[426,525,151,177]" captionStart="FIGURES 99101" captionStartId="25.[151,264,1922,1946]" captionTargetBox="[166,1424,250,1891]" captionTargetId="figure-58@25.[153,1434,237,1899]" captionTargetPageId="25" captionText="FIGURES 99101. Calomera bordonii Wiesner (all from Maluku, Buru Island, Waedea). 99100—habitus: 99—♂, 14.2 mm, HT (JWCM); 100—♀, 14.5 mm, PT (JWCM); 101—labels of the HT. Bar = 1 mm (photography P. Schüle)." figureDoi="http://doi.org/10.5281/zenodo.14734518" httpUri="https://zenodo.org/record/14734518/files/figure.png" pageId="29" pageNumber="30">Figs 100</figureCitation>
,
<figureCitation id="13604ADEFFC7E246FDB626FDFDBAFF30" box="[536,580,151,177]" captionStart="FIGURES 102106" captionStartId="26.[151,264,1885,1909]" captionTargetBox="[172,1417,209,1854]" captionTargetId="figure-27@26.[158,1430,199,1862]" captionTargetPageId="26" captionText="FIGURES 102106. Calomera bordonii Wiesner (all from Buru Island, Waedea). 102—♀, habitus, 15.1 mm, PT (ABCF); 103—104—pronotum: 103—♀, PT (ABCF); 104—♂, HT (JWCM); 105—♂ aedeagus, HT; 106—♂, elytron, HT. Bars = 1 mm (photography P. Schüle)" pageId="29" pageNumber="30">102</figureCitation>
<figureCitation id="13604ADEFFC7E246FDB626FDFDBAFF30" box="[536,580,151,177]" captionStart="FIGURES 102106" captionStartId="26.[151,264,1885,1909]" captionTargetBox="[172,1417,209,1854]" captionTargetId="figure-27@26.[158,1430,199,1862]" captionTargetPageId="26" captionText="FIGURES 102106. Calomera bordonii Wiesner (all from Buru Island, Waedea). 102—♀, habitus, 15.1 mm, PT (ABCF); 103—104—pronotum: 103—♀, PT (ABCF); 104—♂, HT (JWCM); 105—♂ aedeagus, HT; 106—♂, elytron, HT. Bars = 1 mm (photography P. Schüle)" figureDoi="http://doi.org/10.5281/zenodo.14734520" httpUri="https://zenodo.org/record/14734520/files/figure.png" pageId="29" pageNumber="30">102</figureCitation>
,
<figureCitation id="13604ADEFFC7E246FDE126FCFD4AFF30" box="[591,692,150,177]" captionStart="FIGURES 107116" captionStartId="27.[151,264,1866,1890]" captionTargetBox="[158,1429,209,1834]" captionTargetId="figure-18@27.[151,1436,199,1842]" captionTargetPageId="27" captionText="FIGURES 107116. Calomera bordonii Wiesner (all from Buru Island). 107—head, ♂, Remals Mts., Ilat (CCJM); 108111— labrum: 108109—♂, ibid. (CCJM); 110—♀, ibid. (CCJM); 111—♀, Waedea, PT (JWCM); 112113—pronotum: 112—♂, Remals Mts., Ilat (CCJM); 113—♀, Waedea, PT (JWCM); 114116—elytron: 114—♂, Remals Mts., Ilat (CCJM); 115—♀, ibid. (CCJM); 116—♀, Waedea, PT (JWCM). Bars= 1 mm." pageId="29" pageNumber="30">115116</figureCitation>
<figureCitation id="13604ADEFFC7E246FDE126FCFD4AFF30" box="[591,692,150,177]" captionStart="FIGURES 107116" captionStartId="27.[151,264,1866,1890]" captionTargetBox="[158,1429,209,1834]" captionTargetId="figure-18@27.[151,1436,199,1842]" captionTargetPageId="27" captionText="FIGURES 107116. Calomera bordonii Wiesner (all from Buru Island). 107—head, ♂, Remals Mts., Ilat (CCJM); 108111— labrum: 108109—♂, ibid. (CCJM); 110—♀, ibid. (CCJM); 111—♀, Waedea, PT (JWCM); 112113—pronotum: 112—♂, Remals Mts., Ilat (CCJM); 113—♀, Waedea, PT (JWCM); 114116—elytron: 114—♂, Remals Mts., Ilat (CCJM); 115—♀, ibid. (CCJM); 116—♀, Waedea, PT (JWCM). Bars= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734522" httpUri="https://zenodo.org/record/14734522/files/figure.png" pageId="29" pageNumber="30">115116</figureCitation>
). Basic elytral coloration velvety black; furthermore,
<taxonomicName id="4C5B2DD8FFC7E246FAAA26FDFA7DFF30" authorityName="Wiesner" authorityYear="2018" box="[1284,1411,151,177]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bordonii">
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is immediately distinguished by the almost subglobose shape of the pronotal disc in males, olivaceous-green pronotal surface with faint or sometimes bright green, green-blue and violet reflections and with rather coarser surface sculpture, yet the rugae on sublateral areas are usually more spaced (
<figureCitation id="13604ADEFFC7E246FC012769FBAEFE9C" box="[943,1104,259,285]" captionStart="FIGURES 102106" captionStartId="26.[151,264,1885,1909]" captionTargetBox="[172,1417,209,1854]" captionTargetId="figure-27@26.[158,1430,199,1862]" captionTargetPageId="26" captionText="FIGURES 102106. Calomera bordonii Wiesner (all from Buru Island, Waedea). 102—♀, habitus, 15.1 mm, PT (ABCF); 103—104—pronotum: 103—♀, PT (ABCF); 104—♂, HT (JWCM); 105—♂ aedeagus, HT; 106—♂, elytron, HT. Bars = 1 mm (photography P. Schüle)" pageId="29" pageNumber="30">Figs 103104</figureCitation>
<figureCitation id="13604ADEFFC7E246FC012769FBAEFE9C" box="[943,1104,259,285]" captionStart="FIGURES 102106" captionStartId="26.[151,264,1885,1909]" captionTargetBox="[172,1417,209,1854]" captionTargetId="figure-27@26.[158,1430,199,1862]" captionTargetPageId="26" captionText="FIGURES 102106. Calomera bordonii Wiesner (all from Buru Island, Waedea). 102—♀, habitus, 15.1 mm, PT (ABCF); 103—104—pronotum: 103—♀, PT (ABCF); 104—♂, HT (JWCM); 105—♂ aedeagus, HT; 106—♂, elytron, HT. Bars = 1 mm (photography P. Schüle)" figureDoi="http://doi.org/10.5281/zenodo.14734520" httpUri="https://zenodo.org/record/14734520/files/figure.png" pageId="29" pageNumber="30">Figs 103104</figureCitation>
). In addition, male labrum is generally longer and the surface of female labrum has deeper sublateral impression (
<figureCitation id="13604ADEFFC7E246FB92274DFB29FEC0" box="[1084,1239,295,321]" captionStart="FIGURES 107116" captionStartId="27.[151,264,1866,1890]" captionTargetBox="[158,1429,209,1834]" captionTargetId="figure-18@27.[151,1436,199,1842]" captionTargetPageId="27" captionText="FIGURES 107116. Calomera bordonii Wiesner (all from Buru Island). 107—head, ♂, Remals Mts., Ilat (CCJM); 108111— labrum: 108109—♂, ibid. (CCJM); 110—♀, ibid. (CCJM); 111—♀, Waedea, PT (JWCM); 112113—pronotum: 112—♂, Remals Mts., Ilat (CCJM); 113—♀, Waedea, PT (JWCM); 114116—elytron: 114—♂, Remals Mts., Ilat (CCJM); 115—♀, ibid. (CCJM); 116—♀, Waedea, PT (JWCM). Bars= 1 mm." pageId="29" pageNumber="30">Figs 108111</figureCitation>
<figureCitation id="13604ADEFFC7E246FB92274DFB29FEC0" box="[1084,1239,295,321]" captionStart="FIGURES 107116" captionStartId="27.[151,264,1866,1890]" captionTargetBox="[158,1429,209,1834]" captionTargetId="figure-18@27.[151,1436,199,1842]" captionTargetPageId="27" captionText="FIGURES 107116. Calomera bordonii Wiesner (all from Buru Island). 107—head, ♂, Remals Mts., Ilat (CCJM); 108111— labrum: 108109—♂, ibid. (CCJM); 110—♀, ibid. (CCJM); 111—♀, Waedea, PT (JWCM); 112113—pronotum: 112—♂, Remals Mts., Ilat (CCJM); 113—♀, Waedea, PT (JWCM); 114116—elytron: 114—♂, Remals Mts., Ilat (CCJM); 115—♀, ibid. (CCJM); 116—♀, Waedea, PT (JWCM). Bars= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734522" httpUri="https://zenodo.org/record/14734522/files/figure.png" pageId="29" pageNumber="30">Figs 108111</figureCitation>
). Body
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long.

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@ -1,71 +1,72 @@
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<paragraph id="8BE4565BFFC7E246FF392448FE08FDBD" blockId="29.[151,502,546,609]" box="[151,502,546,573]" pageId="29" pageNumber="30">
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@ -82,7 +83,7 @@
<subSubSection id="C34105D0FFC7E246FF39242DFEBDFDE0" box="[151,323,583,609]" pageId="29" pageNumber="30" type="description">
<paragraph id="8BE4565BFFC7E246FF39242DFEBDFDE0" blockId="29.[151,502,546,609]" box="[151,323,583,609]" pageId="29" pageNumber="30">
(
<figureCitation id="13604ADEFFC7E246FF31242DFEC5FDE0" box="[159,315,583,609]" captionStart-0="FIGURES 123125" captionStart-1="FIGURES 126132" captionStart-2="FIGURES 133138" captionStartId-0="30.[151,264,1861,1885]" captionStartId-1="31.[151,264,1591,1615]" captionStartId-2="32.[151,264,1867,1891]" captionTargetBox-0="[164,1425,194,1824]" captionTargetBox-1="[163,1422,191,1554]" captionTargetBox-2="[162,1427,191,1832]" captionTargetId-0="figure-27@30.[151,1435,181,1837]" captionTargetId-1="figure-18@31.[153,1434,181,1567]" captionTargetId-2="figure-27@32.[151,1436,181,1844]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="32" captionText-0="FIGURES 123125. Calomera jakli Schüle, from the type locality in the Obi Island. 123124—habitus: 123—♂, 13.1 mm, HT (PSCH) (photography P. Schüle); 124—♀, 13.7 mm, PT (CCJM); 125—labels of the HT. Bar = 1 mm." captionText-1="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." captionText-2="FIGURES 133138. Calomera jakli Schüle, from the type locality in the Obi Island. 133—habitus, ♂, 13.1 mm, PT (PSCH); 134135—pronotum:134—♂, PT(PSCH); 135—♂, HT (PSCH); 136—elytron, ♂, PT(PSCH); 137—aedeagus, ♂, PT (PSCH); 138—internal sac, PT (PSCH). Bars = 1 mm (photography P. Schüle)." pageId="29" pageNumber="30">Figs 123138</figureCitation>
<figureCitation id="13604ADEFFC7E246FF31242DFEC5FDE0" box="[159,315,583,609]" captionStart-0="FIGURES 123125" captionStart-1="FIGURES 126132" captionStart-2="FIGURES 133138" captionStartId-0="30.[151,264,1861,1885]" captionStartId-1="31.[151,264,1591,1615]" captionStartId-2="32.[151,264,1867,1891]" captionTargetBox-0="[164,1425,194,1824]" captionTargetBox-1="[163,1422,191,1554]" captionTargetBox-2="[162,1427,191,1832]" captionTargetId-0="figure-27@30.[151,1435,181,1837]" captionTargetId-1="figure-18@31.[153,1434,181,1567]" captionTargetId-2="figure-27@32.[151,1436,181,1844]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="32" captionText-0="FIGURES 123125. Calomera jakli Schüle, from the type locality in the Obi Island. 123124—habitus: 123—♂, 13.1 mm, HT (PSCH) (photography P. Schüle); 124—♀, 13.7 mm, PT (CCJM); 125—labels of the HT. Bar = 1 mm." captionText-1="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." captionText-2="FIGURES 133138. Calomera jakli Schüle, from the type locality in the Obi Island. 133—habitus, ♂, 13.1 mm, PT (PSCH); 134135—pronotum:134—♂, PT(PSCH); 135—♂, HT (PSCH); 136—elytron, ♂, PT(PSCH); 137—aedeagus, ♂, PT (PSCH); 138—internal sac, PT (PSCH). Bars = 1 mm (photography P. Schüle)." figureDoi-0="http://doi.org/10.5281/zenodo.14734526" figureDoi-1="http://doi.org/10.5281/zenodo.14734528" figureDoi-2="http://doi.org/10.5281/zenodo.14734530" httpUri-0="https://zenodo.org/record/14734526/files/figure.png" httpUri-1="https://zenodo.org/record/14734528/files/figure.png" httpUri-2="https://zenodo.org/record/14734530/files/figure.png" pageId="29" pageNumber="30">Figs 123138</figureCitation>
)
</paragraph>
</subSubSection>
@ -259,13 +260,13 @@ shares glabrous genae with the three preceding species and
<emphasis id="B92F8A49FFC7E246FC682239FBE3FBEC" box="[966,1053,1107,1133]" italics="true" pageId="29" pageNumber="30">C. jakli</emphasis>
</taxonomicName>
is immediately distinguished by subtrapezoid shape of lateral margins of its pronotal disc (
<figureCitation id="13604ADEFFC7E246FCFE221DFC0AFB10" box="[848,1012,1143,1169]" captionStart="FIGURES 126132" captionStartId="31.[151,264,1591,1615]" captionTargetBox="[163,1422,191,1554]" captionTargetId="figure-18@31.[153,1434,181,1567]" captionTargetPageId="31" captionText="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." pageId="29" pageNumber="30">Figs 131132</figureCitation>
<figureCitation id="13604ADEFFC7E246FCFE221DFC0AFB10" box="[848,1012,1143,1169]" captionStart="FIGURES 126132" captionStartId="31.[151,264,1591,1615]" captionTargetBox="[163,1422,191,1554]" captionTargetId="figure-18@31.[153,1434,181,1567]" captionTargetPageId="31" captionText="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734528" httpUri="https://zenodo.org/record/14734528/files/figure.png" pageId="29" pageNumber="30">Figs 131132</figureCitation>
). Elytral maculation (
<figureCitation id="13604ADEFFC7E246FB5B221DFA66FB10" box="[1269,1432,1143,1169]" captionStart="FIGURES 123125" captionStartId="30.[151,264,1861,1885]" captionTargetBox="[164,1425,194,1824]" captionTargetId="figure-27@30.[151,1435,181,1837]" captionTargetPageId="30" captionText="FIGURES 123125. Calomera jakli Schüle, from the type locality in the Obi Island. 123124—habitus: 123—♂, 13.1 mm, HT (PSCH) (photography P. Schüle); 124—♀, 13.7 mm, PT (CCJM); 125—labels of the HT. Bar = 1 mm." pageId="29" pageNumber="30">Figs 123124</figureCitation>
<figureCitation id="13604ADEFFC7E246FB5B221DFA66FB10" box="[1269,1432,1143,1169]" captionStart="FIGURES 123125" captionStartId="30.[151,264,1861,1885]" captionTargetBox="[164,1425,194,1824]" captionTargetId="figure-27@30.[151,1435,181,1837]" captionTargetPageId="30" captionText="FIGURES 123125. Calomera jakli Schüle, from the type locality in the Obi Island. 123124—habitus: 123—♂, 13.1 mm, HT (PSCH) (photography P. Schüle); 124—♀, 13.7 mm, PT (CCJM); 125—labels of the HT. Bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734526" httpUri="https://zenodo.org/record/14734526/files/figure.png" pageId="29" pageNumber="30">Figs 123124</figureCitation>
,
<figureCitation id="13604ADEFFC7E246FF3922F1FF02FB34" box="[151,252,1179,1205]" captionStart="FIGURES 126132" captionStartId="31.[151,264,1591,1615]" captionTargetBox="[163,1422,191,1554]" captionTargetId="figure-18@31.[153,1434,181,1567]" captionTargetPageId="31" captionText="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." pageId="29" pageNumber="30">129130</figureCitation>
<figureCitation id="13604ADEFFC7E246FF3922F1FF02FB34" box="[151,252,1179,1205]" captionStart="FIGURES 126132" captionStartId="31.[151,264,1591,1615]" captionTargetBox="[163,1422,191,1554]" captionTargetId="figure-18@31.[153,1434,181,1567]" captionTargetPageId="31" captionText="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734528" httpUri="https://zenodo.org/record/14734528/files/figure.png" pageId="29" pageNumber="30">129130</figureCitation>
,
<figureCitation id="13604ADEFFC7E246FEA622F1FE92FB34" box="[264,364,1179,1205]" captionStart="FIGURES 133138" captionStartId="32.[151,264,1867,1891]" captionTargetBox="[162,1427,191,1832]" captionTargetId="figure-27@32.[151,1436,181,1844]" captionTargetPageId="32" captionText="FIGURES 133138. Calomera jakli Schüle, from the type locality in the Obi Island. 133—habitus, ♂, 13.1 mm, PT (PSCH); 134135—pronotum:134—♂, PT(PSCH); 135—♂, HT (PSCH); 136—elytron, ♂, PT(PSCH); 137—aedeagus, ♂, PT (PSCH); 138—internal sac, PT (PSCH). Bars = 1 mm (photography P. Schüle)." pageId="29" pageNumber="30">133, 136</figureCitation>
<figureCitation id="13604ADEFFC7E246FEA622F1FE92FB34" box="[264,364,1179,1205]" captionStart="FIGURES 133138" captionStartId="32.[151,264,1867,1891]" captionTargetBox="[162,1427,191,1832]" captionTargetId="figure-27@32.[151,1436,181,1844]" captionTargetPageId="32" captionText="FIGURES 133138. Calomera jakli Schüle, from the type locality in the Obi Island. 133—habitus, ♂, 13.1 mm, PT (PSCH); 134135—pronotum:134—♂, PT(PSCH); 135—♂, HT (PSCH); 136—elytron, ♂, PT(PSCH); 137—aedeagus, ♂, PT (PSCH); 138—internal sac, PT (PSCH). Bars = 1 mm (photography P. Schüle)." figureDoi="http://doi.org/10.5281/zenodo.14734530" httpUri="https://zenodo.org/record/14734530/files/figure.png" pageId="29" pageNumber="30">133, 136</figureCitation>
) generally large and less spaced, with humeral macula separated from rather large, rounded or subquadrate subhumeral macula; lateromedian macula large, almost adjacent to outer elytral margin and closer to central macula or connected with it with a narrow stripe but not forming entire lateromedian band; anteapical-apical lunula wide but rather short;
<specimenCount id="9D5D9DD2FFC7E246FE42236DFD62FAA0" box="[492,668,1287,1313]" count="3" pageId="29" pageNumber="30" type="generic" typeStatus="paratype">three paratypes</specimenCount>
,
@ -279,9 +280,9 @@ and
(
<collectionCode id="ED4ACE9EFFC7E246FAAC236DFAB3FAA0" box="[1282,1357,1287,1313]" pageId="29" pageNumber="30">CDCL</collectionCode>
), have unusually an additional, small but clear basal spot (
<figureCitation id="13604ADEFFC7E246FD4F2341FCBAFAC4" box="[737,836,1323,1349]" captionStart="FIGURES 126132" captionStartId="31.[151,264,1591,1615]" captionTargetBox="[163,1422,191,1554]" captionTargetId="figure-18@31.[153,1434,181,1567]" captionTargetPageId="31" captionText="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." pageId="29" pageNumber="30">Fig. 130</figureCitation>
<figureCitation id="13604ADEFFC7E246FD4F2341FCBAFAC4" box="[737,836,1323,1349]" captionStart="FIGURES 126132" captionStartId="31.[151,264,1591,1615]" captionTargetBox="[163,1422,191,1554]" captionTargetId="figure-18@31.[153,1434,181,1567]" captionTargetPageId="31" captionText="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734528" httpUri="https://zenodo.org/record/14734528/files/figure.png" pageId="29" pageNumber="30">Fig. 130</figureCitation>
). Labrum with almost effaced anterolateral teeth and with only shallow or indistinct excision on either side of median tooth (
<figureCitation id="13604ADEFFC7E246FC132325FBA3FAE8" box="[957,1117,1359,1385]" captionStart="FIGURES 126132" captionStartId="31.[151,264,1591,1615]" captionTargetBox="[163,1422,191,1554]" captionTargetId="figure-18@31.[153,1434,181,1567]" captionTargetPageId="31" captionText="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." pageId="29" pageNumber="30">Figs 127128</figureCitation>
<figureCitation id="13604ADEFFC7E246FC132325FBA3FAE8" box="[957,1117,1359,1385]" captionStart="FIGURES 126132" captionStartId="31.[151,264,1591,1615]" captionTargetBox="[163,1422,191,1554]" captionTargetId="figure-18@31.[153,1434,181,1567]" captionTargetPageId="31" captionText="FIGURES 126132. Calomera jakli Schüle, from the type locality in the Obi Island. 126—head, ♀, PT (CDCL); 127128— labrum: 127—♂, HT (PSCH); 128—♀, PT (CDCL); 129130—elytron: 129—♂, PT (PSCH); 130—♀, PT (CDCL); 131 132—pronotum: 131—♀, PT (CDCL); 132—♂, PT (PSCH). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734528" httpUri="https://zenodo.org/record/14734528/files/figure.png" pageId="29" pageNumber="30">Figs 127128</figureCitation>
). Body 12.513.9 (
<specimenCode id="DBFDFE20FFC7E246FA9A2325FA6AFAE8" box="[1332,1428,1359,1385]" collectionCode="HT" pageId="29" pageNumber="30">HT 13.1</specimenCode>
) mm long, 4.254.90 (

View file

@ -1,71 +1,72 @@
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<treatment id="03F2E74DFFC7E278FF392020FDC5F871" ID-DOI="http://doi.org/10.5281/zenodo.14734498" ID-Zenodo-Dep="14734498" LSID="urn:lsid:plazi:treatment:03F2E74DFFC7E278FF392020FDC5F871" httpUri="http://treatment.plazi.org/id/03F2E74DFFC7E278FF392020FDC5F871" lastPageId="35" lastPageNumber="36" pageId="29" pageNumber="30">
<subSubSection id="C34105D0FFC7E246FF392020FDD6F9E5" box="[151,552,1610,1637]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8BE4565BFFC7E246FF392020FDD6F9E5" blockId="29.[151,552,1610,1673]" box="[151,552,1610,1637]" pageId="29" pageNumber="30">
<heading id="D0ACE137FFC7E246FF392020FDD6F9E5" bold="true" box="[151,552,1610,1637]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
@ -81,7 +82,7 @@
<subSubSection id="C34105D0FFC7E246FF392005FEBDF908" box="[151,323,1647,1673]" pageId="29" pageNumber="30" type="description">
<paragraph id="8BE4565BFFC7E246FF392005FEBDF908" blockId="29.[151,552,1610,1673]" box="[151,323,1647,1673]" pageId="29" pageNumber="30">
(
<figureCitation id="13604ADEFFC7E246FF312005FEC5F908" box="[159,315,1647,1673]" captionStart-0="FIGURES 139145" captionStart-1="FIGURES 146151" captionStart-2="FIGURES 152157" captionStart-3="FIGURES 158166" captionStartId-0="33.[151,264,1867,1891]" captionStartId-1="34.[151,264,1859,1883]" captionStartId-2="35.[151,264,1395,1419]" captionStartId-3="36.[151,264,1368,1392]" captionTargetBox-0="[159,1430,191,1837]" captionTargetBox-1="[165,1425,191,1822]" captionTargetBox-2="[161,1423,190,1359]" captionTargetBox-3="[166,1422,192,1333]" captionTargetId-0="figure-18@33.[151,1436,181,1844]" captionTargetId-1="figure-27@34.[152,1435,181,1835]" captionTargetId-2="figure-18@35.[152,1436,181,1371]" captionTargetId-3="figure-27@36.[153,1434,181,1345]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="35" captionTargetPageId-3="36" captionText-0="FIGURES 139145. Calomera cabigasi Cassola, Philippines, Gingoog, male holotype (ZSM). 139—habitus, 11.7 mm; 140— labrum; 141—elytron; 142—pronotum; 143—labels; 144—aedeagus; 145—ditto, shape of slightly turned apex. Bars = 1 mm (image credit Katja Neven (ZSM), partly modified)." captionText-1="FIGURES 146151. Calomera cabigasi Cassola, from Philippines. 146147—habitus, Trento, Agusan del Sur (CCJM): 146— ♂, 11.7 mm; 147—♀, 12.7 mm; 148—head, ibid. (CCJM); 149—labrum, ♂, ibid. (CCJM); 150—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 151—♀, Trento, Agusan del Sur (CCJM). Bars = 1 mm." captionText-2="FIGURES 152157. Calomera cabigasi Cassola, from Philippines. 152—habitus, ♀, 12.7 mm, Luzon Nueva Viscaya, Kayapa (PSCH); 153—labrum, ♀, ibid. (PSCH); 154—pronotum, ♀, ibid. (PSCH); 155—elytron, ♀, ibid. (PSCH); 156157—aedeagus showing internal sac in its left and right lateral view. Bars = 1 mm (photography P. Schüle)." captionText-3="FIGURES 158166. Calomera cabigasi Cassola, from Philippines. 158160—pronotum: 158—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 159—♂, Trento, Agusan del Sur (CCJM); 160—♀, ibid (CCJM); 161163—elytron: 161—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 162—♂, Trento, Agusan del Sur (CCJM); 163—♀, Trento, Agusan del Sur (CCJM); 164 166—male aedeagus: 164—Mindanao, Bukidnon, PT (FCCR-MCZR) 165—ditto, dorsal view; 166—Trento, Agusan del Sur (CCJM). Bars = 1 mm." pageId="29" pageNumber="30">Figs 139166</figureCitation>
<figureCitation id="13604ADEFFC7E246FF312005FEC5F908" box="[159,315,1647,1673]" captionStart-0="FIGURES 139145" captionStart-1="FIGURES 146151" captionStart-2="FIGURES 152157" captionStart-3="FIGURES 158166" captionStartId-0="33.[151,264,1867,1891]" captionStartId-1="34.[151,264,1859,1883]" captionStartId-2="35.[151,264,1395,1419]" captionStartId-3="36.[151,264,1368,1392]" captionTargetBox-0="[159,1430,191,1837]" captionTargetBox-1="[165,1425,191,1822]" captionTargetBox-2="[161,1423,190,1359]" captionTargetBox-3="[166,1422,192,1333]" captionTargetId-0="figure-18@33.[151,1436,181,1844]" captionTargetId-1="figure-27@34.[152,1435,181,1835]" captionTargetId-2="figure-18@35.[152,1436,181,1371]" captionTargetId-3="figure-27@36.[153,1434,181,1345]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="35" captionTargetPageId-3="36" captionText-0="FIGURES 139145. Calomera cabigasi Cassola, Philippines, Gingoog, male holotype (ZSM). 139—habitus, 11.7 mm; 140— labrum; 141—elytron; 142—pronotum; 143—labels; 144—aedeagus; 145—ditto, shape of slightly turned apex. Bars = 1 mm (image credit Katja Neven (ZSM), partly modified)." captionText-1="FIGURES 146151. Calomera cabigasi Cassola, from Philippines. 146147—habitus, Trento, Agusan del Sur (CCJM): 146— ♂, 11.7 mm; 147—♀, 12.7 mm; 148—head, ibid. (CCJM); 149—labrum, ♂, ibid. (CCJM); 150—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 151—♀, Trento, Agusan del Sur (CCJM). Bars = 1 mm." captionText-2="FIGURES 152157. Calomera cabigasi Cassola, from Philippines. 152—habitus, ♀, 12.7 mm, Luzon Nueva Viscaya, Kayapa (PSCH); 153—labrum, ♀, ibid. (PSCH); 154—pronotum, ♀, ibid. (PSCH); 155—elytron, ♀, ibid. (PSCH); 156157—aedeagus showing internal sac in its left and right lateral view. Bars = 1 mm (photography P. Schüle)." captionText-3="FIGURES 158166. Calomera cabigasi Cassola, from Philippines. 158160—pronotum: 158—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 159—♂, Trento, Agusan del Sur (CCJM); 160—♀, ibid (CCJM); 161163—elytron: 161—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 162—♂, Trento, Agusan del Sur (CCJM); 163—♀, Trento, Agusan del Sur (CCJM); 164 166—male aedeagus: 164—Mindanao, Bukidnon, PT (FCCR-MCZR) 165—ditto, dorsal view; 166—Trento, Agusan del Sur (CCJM). Bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14734532" figureDoi-1="http://doi.org/10.5281/zenodo.14734534" figureDoi-2="http://doi.org/10.5281/zenodo.14734536" figureDoi-3="http://doi.org/10.5281/zenodo.14734538" httpUri-0="https://zenodo.org/record/14734532/files/figure.png" httpUri-1="https://zenodo.org/record/14734534/files/figure.png" httpUri-2="https://zenodo.org/record/14734536/files/figure.png" httpUri-3="https://zenodo.org/record/14734538/files/figure.png" pageId="29" pageNumber="30">Figs 139166</figureCitation>
)
</paragraph>
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@ -162,7 +163,7 @@ dedit 2010” [red, printed]
</materialsCitation>
.
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<caption id="DF2406D3FFC4E245FF39212FFB56F800" pageId="30" pageNumber="31" startId="30.[151,264,1861,1885]" targetBox="[164,1425,194,1824]" targetPageId="30" targetType="figure">
<caption id="DF2406D3FFC4E245FF39212FFB56F800" ID-DOI="http://doi.org/10.5281/zenodo.14734526" ID-Zenodo-Dep="14734526" httpUri="https://zenodo.org/record/14734526/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,264,1861,1885]" targetBox="[164,1425,194,1824]" targetPageId="30" targetType="figure">
<paragraph id="8BE4565BFFC4E245FF39212FFB56F800" blockId="30.[151,1437,1861,1922]" pageId="30" pageNumber="31">
<emphasis id="B92F8A49FFC4E245FF39212FFE93F8DC" bold="true" box="[151,365,1861,1885]" pageId="30" pageNumber="31">FIGURES 123125.</emphasis>
<taxonomicName id="4C5B2DD8FFC4E245FEDC212FFDA8F8DC" authority="Schule" authorityName="Schule" authorityYear="2010" box="[370,598,1861,1885]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="jakli">
@ -172,7 +173,7 @@ Schüle
, from the type locality in the Obi Island. 123124—habitus: 123—♂, 13.1 mm, HT (PSCH) (photography P. Schüle); 124—♀, 13.7 mm, PT (CCJM); 125—labels of the HT. Bar = 1 mm.
</paragraph>
</caption>
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<caption id="DF2406D3FFC5E244FF39205DFC6BF916" ID-DOI="http://doi.org/10.5281/zenodo.14734528" ID-Zenodo-Dep="14734528" httpUri="https://zenodo.org/record/14734528/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,264,1591,1615]" targetBox="[163,1422,191,1554]" targetPageId="31" targetType="figure">
<paragraph id="8BE4565BFFC5E244FF39205DFC6BF916" blockId="31.[151,1436,1591,1687]" pageId="31" pageNumber="32">
<emphasis id="B92F8A49FFC5E244FF39205DFE90F9CE" bold="true" box="[151,366,1591,1615]" pageId="31" pageNumber="32">FIGURES 126132.</emphasis>
<taxonomicName id="4C5B2DD8FFC5E244FEDB205DFDA3F9CE" authority="Schule" authorityName="Schule" authorityYear="2010" box="[373,605,1591,1615]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="jakli">
@ -305,7 +306,7 @@ local collector”
.
</paragraph>
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<caption id="DF2406D3FFFAE27BFF392121FCA3F82D" ID-DOI="http://doi.org/10.5281/zenodo.14734530" ID-Zenodo-Dep="14734530" httpUri="https://zenodo.org/record/14734530/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,264,1867,1891]" targetBox="[162,1427,191,1832]" targetPageId="32" targetType="figure">
<paragraph id="8BE4565BFFFAE27BFF392121FCA3F82D" blockId="32.[151,1436,1867,1964]" pageId="32" pageNumber="33">
<emphasis id="B92F8A49FFFAE27BFF392121FE90F8E2" bold="true" box="[151,366,1867,1891]" pageId="32" pageNumber="33">FIGURES 133138.</emphasis>
<taxonomicName id="4C5B2DD8FFFAE27BFEDB2126FDA2F8E5" authority="Schule" authorityName="Schule" authorityYear="2010" box="[373,604,1868,1892]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="jakli">
@ -315,7 +316,7 @@ Schüle
, from the type locality in the Obi Island. 133—habitus, ♂, 13.1 mm, PT (PSCH); 134135—pronotum: 134—♂, PT (PSCH); 135—♂, HT (PSCH); 136—elytron, ♂, PT (PSCH); 137—aedeagus, ♂, PT (PSCH); 138—internal sac, PT (PSCH). Bars = 1 mm (photography P. Schüle).
</paragraph>
</caption>
<caption id="DF2406D3FFFBE27AFF392121FD56F82D" pageId="33" pageNumber="34" startId="33.[151,264,1867,1891]" targetBox="[159,1430,191,1837]" targetPageId="33" targetType="figure">
<caption id="DF2406D3FFFBE27AFF392121FD56F82D" ID-DOI="http://doi.org/10.5281/zenodo.14734532" ID-Zenodo-Dep="14734532" httpUri="https://zenodo.org/record/14734532/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,264,1867,1891]" targetBox="[159,1430,191,1837]" targetPageId="33" targetType="figure">
<paragraph id="8BE4565BFFFBE27AFF392121FD56F82D" blockId="33.[151,1437,1867,1964]" pageId="33" pageNumber="34">
<emphasis id="B92F8A49FFFBE27AFF392121FE93F8E2" bold="true" box="[151,365,1867,1891]" pageId="33" pageNumber="34">FIGURES 139145.</emphasis>
<taxonomicName id="4C5B2DD8FFFBE27AFEDD2126FD72F8E5" authority="Cassola" authorityName="Cassola" authorityYear="2011" box="[371,652,1868,1892]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cabigasi">
@ -325,7 +326,7 @@ Cassola
, Philippines, Gingoog, male holotype (ZSM). 139—habitus, 11.7 mm; 140— labrum; 141—elytron; 142—pronotum; 143—labels; 144—aedeagus; 145—ditto, shape of slightly turned apex. Bars = 1 mm (image credit Katja Neven (ZSM), partly modified).
</paragraph>
</caption>
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<caption id="DF2406D3FFF8E279FF392129FC7EF822" ID-DOI="http://doi.org/10.5281/zenodo.14734534" ID-Zenodo-Dep="14734534" httpUri="https://zenodo.org/record/14734534/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,264,1859,1883]" targetBox="[165,1425,191,1822]" targetPageId="34" targetType="figure">
<paragraph id="8BE4565BFFF8E279FF392129FC7EF822" blockId="34.[151,1437,1859,1956]" pageId="34" pageNumber="35">
<emphasis id="B92F8A49FFF8E279FF392129FE93F8DA" bold="true" box="[151,365,1859,1883]" pageId="34" pageNumber="35">FIGURES 146151.</emphasis>
<taxonomicName id="4C5B2DD8FFF8E279FEDC2129FD74F8DA" authority="Cassola" authorityName="Cassola" authorityYear="2011" box="[370,650,1859,1883]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cabigasi">
@ -335,7 +336,7 @@ Cassola
, from Philippines. 146147—habitus, Trento, Agusan del Sur (CCJM): 146— ♂, 11.7 mm; 147—♀, 12.7 mm; 148—head, ibid. (CCJM); 149—labrum, ♂, ibid. (CCJM); 150—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 151—♀, Trento, Agusan del Sur (CCJM). Bars = 1 mm.
</paragraph>
</caption>
<caption id="DF2406D3FFF9E278FF392319FBC9FA52" pageId="35" pageNumber="36" startId="35.[151,264,1395,1419]" targetBox="[161,1423,190,1359]" targetPageId="35" targetType="figure">
<caption id="DF2406D3FFF9E278FF392319FBC9FA52" ID-DOI="http://doi.org/10.5281/zenodo.14734536" ID-Zenodo-Dep="14734536" httpUri="https://zenodo.org/record/14734536/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,264,1395,1419]" targetBox="[161,1423,190,1359]" targetPageId="35" targetType="figure">
<paragraph id="8BE4565BFFF9E278FF392319FBC9FA52" blockId="35.[151,1436,1395,1492]" pageId="35" pageNumber="36">
<emphasis id="B92F8A49FFF9E278FF392319FE93FA0A" bold="true" box="[151,365,1395,1419]" pageId="35" pageNumber="36">FIGURES 152157.</emphasis>
<taxonomicName id="4C5B2DD8FFF9E278FEDD2319FD75FA0A" authority="Cassola" authorityName="Cassola" authorityYear="2011" box="[371,651,1395,1419]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="cabigasi">
@ -353,25 +354,25 @@ Cassola
<emphasis id="B92F8A49FFF9E278FE73204CFD49F9C1" box="[477,695,1574,1600]" italics="true" pageId="35" pageNumber="36">Calomera cabigasi</emphasis>
</taxonomicName>
is rather clearly distinguished from all preceding species, with which it shares glabrous genae, by its elytral maculae which are isolated and notably spaced (central macula rarely indistinctly thinly connected with lateromedian one), lateromedian macula placed more dorsally, while small, elongate lateromedian ventral-most spot is perfectly or almost at the same level with central macula (
<figureCitation id="13604ADEFFF9E278FB5520F8FA69F92D" box="[1275,1431,1682,1708]" captionStart="FIGURES 139145" captionStartId="33.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1837]" captionTargetId="figure-18@33.[151,1436,181,1844]" captionTargetPageId="33" captionText="FIGURES 139145. Calomera cabigasi Cassola, Philippines, Gingoog, male holotype (ZSM). 139—habitus, 11.7 mm; 140— labrum; 141—elytron; 142—pronotum; 143—labels; 144—aedeagus; 145—ditto, shape of slightly turned apex. Bars = 1 mm (image credit Katja Neven (ZSM), partly modified)." pageId="35" pageNumber="36">Figs 139, 141</figureCitation>
<figureCitation id="13604ADEFFF9E278FB5520F8FA69F92D" box="[1275,1431,1682,1708]" captionStart="FIGURES 139145" captionStartId="33.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1837]" captionTargetId="figure-18@33.[151,1436,181,1844]" captionTargetPageId="33" captionText="FIGURES 139145. Calomera cabigasi Cassola, Philippines, Gingoog, male holotype (ZSM). 139—habitus, 11.7 mm; 140— labrum; 141—elytron; 142—pronotum; 143—labels; 144—aedeagus; 145—ditto, shape of slightly turned apex. Bars = 1 mm (image credit Katja Neven (ZSM), partly modified)." figureDoi="http://doi.org/10.5281/zenodo.14734532" httpUri="https://zenodo.org/record/14734532/files/figure.png" pageId="35" pageNumber="36">Figs 139, 141</figureCitation>
,
<figureCitation id="13604ADEFFF9E278FF3920DCFF02F951" box="[151,252,1718,1744]" captionStart="FIGURES 146151" captionStartId="34.[151,264,1859,1883]" captionTargetBox="[165,1425,191,1822]" captionTargetId="figure-27@34.[152,1435,181,1835]" captionTargetPageId="34" captionText="FIGURES 146151. Calomera cabigasi Cassola, from Philippines. 146147—habitus, Trento, Agusan del Sur (CCJM): 146— ♂, 11.7 mm; 147—♀, 12.7 mm; 148—head, ibid. (CCJM); 149—labrum, ♂, ibid. (CCJM); 150—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 151—♀, Trento, Agusan del Sur (CCJM). Bars = 1 mm." pageId="35" pageNumber="36">146147</figureCitation>
<figureCitation id="13604ADEFFF9E278FF3920DCFF02F951" box="[151,252,1718,1744]" captionStart="FIGURES 146151" captionStartId="34.[151,264,1859,1883]" captionTargetBox="[165,1425,191,1822]" captionTargetId="figure-27@34.[152,1435,181,1835]" captionTargetPageId="34" captionText="FIGURES 146151. Calomera cabigasi Cassola, from Philippines. 146147—habitus, Trento, Agusan del Sur (CCJM): 146— ♂, 11.7 mm; 147—♀, 12.7 mm; 148—head, ibid. (CCJM); 149—labrum, ♂, ibid. (CCJM); 150—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 151—♀, Trento, Agusan del Sur (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734534" httpUri="https://zenodo.org/record/14734534/files/figure.png" pageId="35" pageNumber="36">146147</figureCitation>
,
<figureCitation id="13604ADEFFF9E278FEA920DFFE94F951" box="[263,362,1717,1744]" captionStart="FIGURES 152157" captionStartId="35.[151,264,1395,1419]" captionTargetBox="[161,1423,190,1359]" captionTargetId="figure-18@35.[152,1436,181,1371]" captionTargetPageId="35" captionText="FIGURES 152157. Calomera cabigasi Cassola, from Philippines. 152—habitus, ♀, 12.7 mm, Luzon Nueva Viscaya, Kayapa (PSCH); 153—labrum, ♀, ibid. (PSCH); 154—pronotum, ♀, ibid. (PSCH); 155—elytron, ♀, ibid. (PSCH); 156157—aedeagus showing internal sac in its left and right lateral view. Bars = 1 mm (photography P. Schüle)." pageId="35" pageNumber="36">152, 155</figureCitation>
<figureCitation id="13604ADEFFF9E278FEA920DFFE94F951" box="[263,362,1717,1744]" captionStart="FIGURES 152157" captionStartId="35.[151,264,1395,1419]" captionTargetBox="[161,1423,190,1359]" captionTargetId="figure-18@35.[152,1436,181,1371]" captionTargetPageId="35" captionText="FIGURES 152157. Calomera cabigasi Cassola, from Philippines. 152—habitus, ♀, 12.7 mm, Luzon Nueva Viscaya, Kayapa (PSCH); 153—labrum, ♀, ibid. (PSCH); 154—pronotum, ♀, ibid. (PSCH); 155—elytron, ♀, ibid. (PSCH); 156157—aedeagus showing internal sac in its left and right lateral view. Bars = 1 mm (photography P. Schüle)." figureDoi="http://doi.org/10.5281/zenodo.14734536" httpUri="https://zenodo.org/record/14734536/files/figure.png" pageId="35" pageNumber="36">152, 155</figureCitation>
,
<figureCitation id="13604ADEFFF9E278FED820DCFE23F951" box="[374,477,1718,1744]" captionStart="FIGURES 158166" captionStartId="36.[151,264,1368,1392]" captionTargetBox="[166,1422,192,1333]" captionTargetId="figure-27@36.[153,1434,181,1345]" captionTargetPageId="36" captionText="FIGURES 158166. Calomera cabigasi Cassola, from Philippines. 158160—pronotum: 158—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 159—♂, Trento, Agusan del Sur (CCJM); 160—♀, ibid (CCJM); 161163—elytron: 161—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 162—♂, Trento, Agusan del Sur (CCJM); 163—♀, Trento, Agusan del Sur (CCJM); 164 166—male aedeagus: 164—Mindanao, Bukidnon, PT (FCCR-MCZR) 165—ditto, dorsal view; 166—Trento, Agusan del Sur (CCJM). Bars = 1 mm." pageId="35" pageNumber="36">161163</figureCitation>
<figureCitation id="13604ADEFFF9E278FED820DCFE23F951" box="[374,477,1718,1744]" captionStart="FIGURES 158166" captionStartId="36.[151,264,1368,1392]" captionTargetBox="[166,1422,192,1333]" captionTargetId="figure-27@36.[153,1434,181,1345]" captionTargetPageId="36" captionText="FIGURES 158166. Calomera cabigasi Cassola, from Philippines. 158160—pronotum: 158—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 159—♂, Trento, Agusan del Sur (CCJM); 160—♀, ibid (CCJM); 161163—elytron: 161—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 162—♂, Trento, Agusan del Sur (CCJM); 163—♀, Trento, Agusan del Sur (CCJM); 164 166—male aedeagus: 164—Mindanao, Bukidnon, PT (FCCR-MCZR) 165—ditto, dorsal view; 166—Trento, Agusan del Sur (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734538" httpUri="https://zenodo.org/record/14734538/files/figure.png" pageId="35" pageNumber="36">161163</figureCitation>
). Lateral areas of pronotal surface mostly glabrous or with occasional setae only (
<figureCitation id="13604ADEFFF9E278FAC220DCFF3DF975" captionStart="FIGURES 139145" captionStartId="33.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1837]" captionTargetId="figure-18@33.[151,1436,181,1844]" captionTargetPageId="33" captionText="FIGURES 139145. Calomera cabigasi Cassola, Philippines, Gingoog, male holotype (ZSM). 139—habitus, 11.7 mm; 140— labrum; 141—elytron; 142—pronotum; 143—labels; 144—aedeagus; 145—ditto, shape of slightly turned apex. Bars = 1 mm (image credit Katja Neven (ZSM), partly modified)." pageId="35" pageNumber="36">Figs 142</figureCitation>
<figureCitation id="13604ADEFFF9E278FAC220DCFF3DF975" captionStart="FIGURES 139145" captionStartId="33.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1837]" captionTargetId="figure-18@33.[151,1436,181,1844]" captionTargetPageId="33" captionText="FIGURES 139145. Calomera cabigasi Cassola, Philippines, Gingoog, male holotype (ZSM). 139—habitus, 11.7 mm; 140— labrum; 141—elytron; 142—pronotum; 143—labels; 144—aedeagus; 145—ditto, shape of slightly turned apex. Bars = 1 mm (image credit Katja Neven (ZSM), partly modified)." figureDoi="http://doi.org/10.5281/zenodo.14734532" httpUri="https://zenodo.org/record/14734532/files/figure.png" pageId="35" pageNumber="36">Figs 142</figureCitation>
,
<figureCitation id="13604ADEFFF9E278FF6120B3FF05F975" box="[207,251,1753,1780]" captionStart="FIGURES 152157" captionStartId="35.[151,264,1395,1419]" captionTargetBox="[161,1423,190,1359]" captionTargetId="figure-18@35.[152,1436,181,1371]" captionTargetPageId="35" captionText="FIGURES 152157. Calomera cabigasi Cassola, from Philippines. 152—habitus, ♀, 12.7 mm, Luzon Nueva Viscaya, Kayapa (PSCH); 153—labrum, ♀, ibid. (PSCH); 154—pronotum, ♀, ibid. (PSCH); 155—elytron, ♀, ibid. (PSCH); 156157—aedeagus showing internal sac in its left and right lateral view. Bars = 1 mm (photography P. Schüle)." pageId="35" pageNumber="36">154</figureCitation>
<figureCitation id="13604ADEFFF9E278FF6120B3FF05F975" box="[207,251,1753,1780]" captionStart="FIGURES 152157" captionStartId="35.[151,264,1395,1419]" captionTargetBox="[161,1423,190,1359]" captionTargetId="figure-18@35.[152,1436,181,1371]" captionTargetPageId="35" captionText="FIGURES 152157. Calomera cabigasi Cassola, from Philippines. 152—habitus, ♀, 12.7 mm, Luzon Nueva Viscaya, Kayapa (PSCH); 153—labrum, ♀, ibid. (PSCH); 154—pronotum, ♀, ibid. (PSCH); 155—elytron, ♀, ibid. (PSCH); 156157—aedeagus showing internal sac in its left and right lateral view. Bars = 1 mm (photography P. Schüle)." figureDoi="http://doi.org/10.5281/zenodo.14734536" httpUri="https://zenodo.org/record/14734536/files/figure.png" pageId="35" pageNumber="36">154</figureCitation>
,
<figureCitation id="13604ADEFFF9E278FEA920B3FE90F975" box="[263,366,1753,1780]" captionStart="FIGURES 158166" captionStartId="36.[151,264,1368,1392]" captionTargetBox="[166,1422,192,1333]" captionTargetId="figure-27@36.[153,1434,181,1345]" captionTargetPageId="36" captionText="FIGURES 158166. Calomera cabigasi Cassola, from Philippines. 158160—pronotum: 158—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 159—♂, Trento, Agusan del Sur (CCJM); 160—♀, ibid (CCJM); 161163—elytron: 161—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 162—♂, Trento, Agusan del Sur (CCJM); 163—♀, Trento, Agusan del Sur (CCJM); 164 166—male aedeagus: 164—Mindanao, Bukidnon, PT (FCCR-MCZR) 165—ditto, dorsal view; 166—Trento, Agusan del Sur (CCJM). Bars = 1 mm." pageId="35" pageNumber="36">158160</figureCitation>
<figureCitation id="13604ADEFFF9E278FEA920B3FE90F975" box="[263,366,1753,1780]" captionStart="FIGURES 158166" captionStartId="36.[151,264,1368,1392]" captionTargetBox="[166,1422,192,1333]" captionTargetId="figure-27@36.[153,1434,181,1345]" captionTargetPageId="36" captionText="FIGURES 158166. Calomera cabigasi Cassola, from Philippines. 158160—pronotum: 158—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 159—♂, Trento, Agusan del Sur (CCJM); 160—♀, ibid (CCJM); 161163—elytron: 161—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 162—♂, Trento, Agusan del Sur (CCJM); 163—♀, Trento, Agusan del Sur (CCJM); 164 166—male aedeagus: 164—Mindanao, Bukidnon, PT (FCCR-MCZR) 165—ditto, dorsal view; 166—Trento, Agusan del Sur (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734538" httpUri="https://zenodo.org/record/14734538/files/figure.png" pageId="35" pageNumber="36">158160</figureCitation>
). Labrum with mostly markedly blackened anterior area (
<figureCitation id="13604ADEFFF9E278FC5C20B0FBA8F975" box="[1010,1110,1754,1780]" captionStart="FIGURES 139145" captionStartId="33.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1837]" captionTargetId="figure-18@33.[151,1436,181,1844]" captionTargetPageId="33" captionText="FIGURES 139145. Calomera cabigasi Cassola, Philippines, Gingoog, male holotype (ZSM). 139—habitus, 11.7 mm; 140— labrum; 141—elytron; 142—pronotum; 143—labels; 144—aedeagus; 145—ditto, shape of slightly turned apex. Bars = 1 mm (image credit Katja Neven (ZSM), partly modified)." pageId="35" pageNumber="36">Figs 140</figureCitation>
<figureCitation id="13604ADEFFF9E278FC5C20B0FBA8F975" box="[1010,1110,1754,1780]" captionStart="FIGURES 139145" captionStartId="33.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1837]" captionTargetId="figure-18@33.[151,1436,181,1844]" captionTargetPageId="33" captionText="FIGURES 139145. Calomera cabigasi Cassola, Philippines, Gingoog, male holotype (ZSM). 139—habitus, 11.7 mm; 140— labrum; 141—elytron; 142—pronotum; 143—labels; 144—aedeagus; 145—ditto, shape of slightly turned apex. Bars = 1 mm (image credit Katja Neven (ZSM), partly modified)." figureDoi="http://doi.org/10.5281/zenodo.14734532" httpUri="https://zenodo.org/record/14734532/files/figure.png" pageId="35" pageNumber="36">Figs 140</figureCitation>
,
<figureCitation id="13604ADEFFF9E278FBCC20B3FB38F975" box="[1122,1222,1753,1780]" captionStart="FIGURES 146151" captionStartId="34.[151,264,1859,1883]" captionTargetBox="[165,1425,191,1822]" captionTargetId="figure-27@34.[152,1435,181,1835]" captionTargetPageId="34" captionText="FIGURES 146151. Calomera cabigasi Cassola, from Philippines. 146147—habitus, Trento, Agusan del Sur (CCJM): 146— ♂, 11.7 mm; 147—♀, 12.7 mm; 148—head, ibid. (CCJM); 149—labrum, ♂, ibid. (CCJM); 150—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 151—♀, Trento, Agusan del Sur (CCJM). Bars = 1 mm." pageId="35" pageNumber="36">149151</figureCitation>
<figureCitation id="13604ADEFFF9E278FBCC20B3FB38F975" box="[1122,1222,1753,1780]" captionStart="FIGURES 146151" captionStartId="34.[151,264,1859,1883]" captionTargetBox="[165,1425,191,1822]" captionTargetId="figure-27@34.[152,1435,181,1835]" captionTargetPageId="34" captionText="FIGURES 146151. Calomera cabigasi Cassola, from Philippines. 146147—habitus, Trento, Agusan del Sur (CCJM): 146— ♂, 11.7 mm; 147—♀, 12.7 mm; 148—head, ibid. (CCJM); 149—labrum, ♂, ibid. (CCJM); 150—♂, Mindanao, Bukidnon, PT (FCCR-MCZR); 151—♀, Trento, Agusan del Sur (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734534" httpUri="https://zenodo.org/record/14734534/files/figure.png" pageId="35" pageNumber="36">149151</figureCitation>
,
<figureCitation id="13604ADEFFF9E278FB7C20B3FAFEF975" box="[1234,1280,1753,1780]" captionStart="FIGURES 152157" captionStartId="35.[151,264,1395,1419]" captionTargetBox="[161,1423,190,1359]" captionTargetId="figure-18@35.[152,1436,181,1371]" captionTargetPageId="35" captionText="FIGURES 152157. Calomera cabigasi Cassola, from Philippines. 152—habitus, ♀, 12.7 mm, Luzon Nueva Viscaya, Kayapa (PSCH); 153—labrum, ♀, ibid. (PSCH); 154—pronotum, ♀, ibid. (PSCH); 155—elytron, ♀, ibid. (PSCH); 156157—aedeagus showing internal sac in its left and right lateral view. Bars = 1 mm (photography P. Schüle)." pageId="35" pageNumber="36">153</figureCitation>
<figureCitation id="13604ADEFFF9E278FB7C20B3FAFEF975" box="[1234,1280,1753,1780]" captionStart="FIGURES 152157" captionStartId="35.[151,264,1395,1419]" captionTargetBox="[161,1423,190,1359]" captionTargetId="figure-18@35.[152,1436,181,1371]" captionTargetPageId="35" captionText="FIGURES 152157. Calomera cabigasi Cassola, from Philippines. 152—habitus, ♀, 12.7 mm, Luzon Nueva Viscaya, Kayapa (PSCH); 153—labrum, ♀, ibid. (PSCH); 154—pronotum, ♀, ibid. (PSCH); 155—elytron, ♀, ibid. (PSCH); 156157—aedeagus showing internal sac in its left and right lateral view. Bars = 1 mm (photography P. Schüle)." figureDoi="http://doi.org/10.5281/zenodo.14734536" httpUri="https://zenodo.org/record/14734536/files/figure.png" pageId="35" pageNumber="36">153</figureCitation>
).
</paragraph>
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@ -1,71 +1,72 @@
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(
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<figureCitation id="13604ADEFFFEE27FFF31201DFEC5F910" box="[159,315,1655,1681]" captionStart-0="FIGURES 167171" captionStart-1="FIGURES 172178" captionStart-2="FIGURES 179181" captionStart-3="FIGURES 182186" captionStart-4="FIGURES 187199" captionStartId-0="37.[151,264,1867,1891]" captionStartId-1="38.[151,264,1864,1888]" captionStartId-2="39.[151,264,1885,1909]" captionStartId-3="40.[151,264,1224,1248]" captionStartId-4="41.[151,264,1822,1846]" captionTargetBox-0="[151,1425,181,1834]" captionTargetBox-1="[164,1427,194,1831]" captionTargetBox-2="[160,1428,210,1852]" captionTargetBox-3="[159,1429,189,1188]" captionTargetBox-4="[162,1428,161,1807]" captionTargetId-0="figure-18@37.[151,1436,181,1844]" captionTargetId-1="figure-27@38.[151,1435,181,1841]" captionTargetId-2="figure-18@39.[151,1436,199,1862]" captionTargetId-3="figure-27@40.[151,1435,181,1200]" captionTargetId-4="figure-18@41.[151,1436,151,1814]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionTargetPageId-3="40" captionTargetPageId-4="41" captionText-0="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." captionText-1="FIGURES 172178. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 172—habitus, ♂, 14.5 mm, Malabo, Mamasa Mamuju road, PT (FCCR-MCZR); 173—head, ♂, ibid., PT (FCCR-MCZR); 174178—labrum: 174—♂, ibid., PT (FCCR- MCZR); 175—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 176—♂, Barat, Mamasa (PSCH); 177—♂, Sabbang (JWCM); 178—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR). Bars = 1 mm." captionText-2="FIGURES 179181. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 179180—habitus: 179—♂, 14.3 mm, Sabbang (JWCM); 180—♂, 14.9 mm, Tengah / Gunung Sampuraga (AOCW); 181—male aedeagus showing internal sac, Barat, Mamasa (PSCH). Bars = 1 mm." captionText-3="FIGURES 182186. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi, pronotum. 182—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 183— ♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 184—♂, Sabbang (JWCM); 185—♂, Barat, Mamasa (PSCH). 186—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR). Bars = 1 mm." captionText-4="FIGURES 187199. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 187191—elytron: 187—♂, Barat, Polewali— Mamasa road, PT (FCCR-MCZR);188—♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 189—♂, Barat, Mamasa (PSCH); 190—♂, Sabbang (JWCM); 191—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 192199—male aedeagus or its apex: 192—PolewaliMamasa road, PT (CCJM); 193— Malabo, MamasaMamuju road, PT (FCCR-MCZR); 194— ditto, dorsal view; 195—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 196—♂, Barat, Mamasa (PSCH); 197—ditto., dorsal view; 198—Sabbang (JWCM); 199—ditto, dorsal view. Bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14734540" figureDoi-1="http://doi.org/10.5281/zenodo.14734542" figureDoi-2="http://doi.org/10.5281/zenodo.14734544" figureDoi-3="http://doi.org/10.5281/zenodo.14734546" figureDoi-4="http://doi.org/10.5281/zenodo.14734548" httpUri-0="https://zenodo.org/record/14734540/files/figure.png" httpUri-1="https://zenodo.org/record/14734542/files/figure.png" httpUri-2="https://zenodo.org/record/14734544/files/figure.png" httpUri-3="https://zenodo.org/record/14734546/files/figure.png" httpUri-4="https://zenodo.org/record/14734548/files/figure.png" pageId="36" pageNumber="37">Figs 167199</figureCitation>
)
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@ -136,7 +137,7 @@ northeast of
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<paragraph id="8BE4565BFFFFE27EFF392121FC97F82D" blockId="37.[151,1437,1867,1964]" pageId="37" pageNumber="38">
<emphasis id="B92F8A49FFFFE27EFF392121FE8AF8E2" bold="true" box="[151,372,1867,1891]" pageId="37" pageNumber="38">FIGURES 167171.</emphasis>
<taxonomicName id="4C5B2DD8FFFFE27EFE2F2126FCD6F8E5" authority="Cassola &amp; Brzoska" authorityName="Cassola &amp; Brzoska" authorityYear="2008" box="[385,808,1868,1892]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mamasa">
@ -308,7 +309,7 @@ Cassola &amp; Brzoska
, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG).
</paragraph>
</caption>
<caption id="DF2406D3FFFCE27DFF392122FBF8F84D" pageId="38" pageNumber="39" startId="38.[151,264,1864,1888]" targetBox="[164,1427,194,1831]" targetPageId="38" targetType="figure">
<caption id="DF2406D3FFFCE27DFF392122FBF8F84D" ID-DOI="http://doi.org/10.5281/zenodo.14734542" ID-Zenodo-Dep="14734542" httpUri="https://zenodo.org/record/14734542/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,264,1864,1888]" targetBox="[164,1427,194,1831]" targetPageId="38" targetType="figure">
<paragraph id="8BE4565BFFFCE27DFF392122FBF8F84D" blockId="38.[151,1436,1864,1997]" pageId="38" pageNumber="39">
<emphasis id="B92F8A49FFFCE27DFF392122FE93F8E1" bold="true" box="[151,365,1864,1888]" pageId="38" pageNumber="39">FIGURES 172178.</emphasis>
<taxonomicName id="4C5B2DD8FFFCE27DFEDD2122FD03F8E1" authority="Cassola &amp; Brzoska" authorityName="Cassola &amp; Brzoska" authorityYear="2008" box="[371,765,1864,1889]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mamasa">
@ -320,7 +321,7 @@ Cassola &amp; Brzoska
; 177—♂, Sabbang (JWCM); 178—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR). Bars = 1 mm.
</paragraph>
</caption>
<caption id="DF2406D3FFFDE27CFF392137FE81F83C" pageId="39" pageNumber="40" startId="39.[151,264,1885,1909]" targetBox="[160,1428,210,1852]" targetPageId="39" targetType="figure">
<caption id="DF2406D3FFFDE27CFF392137FE81F83C" ID-DOI="http://doi.org/10.5281/zenodo.14734544" ID-Zenodo-Dep="14734544" httpUri="https://zenodo.org/record/14734544/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,264,1885,1909]" targetBox="[160,1428,210,1852]" targetPageId="39" targetType="figure">
<paragraph id="8BE4565BFFFDE27CFF392137FE81F83C" blockId="39.[151,1436,1885,1981]" pageId="39" pageNumber="40">
<emphasis id="B92F8A49FFFDE27CFF392137FE92F8F4" bold="true" box="[151,364,1885,1909]" pageId="39" pageNumber="40">FIGURES 179181.</emphasis>
<taxonomicName id="4C5B2DD8FFFDE27CFEDF2137FD06F8F4" authority="Cassola &amp; Brzoska" authorityName="Cassola &amp; Brzoska" authorityYear="2008" box="[369,760,1885,1909]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mamasa">
@ -332,7 +333,7 @@ Cassola &amp; Brzoska
. Bars = 1 mm.
</paragraph>
</caption>
<caption id="DF2406D3FFF2E273FF3922A2FB65FAA9" pageId="40" pageNumber="41" startId="40.[151,264,1224,1248]" targetBox="[159,1429,189,1188]" targetPageId="40" targetType="figure">
<caption id="DF2406D3FFF2E273FF3922A2FB65FAA9" ID-DOI="http://doi.org/10.5281/zenodo.14734546" ID-Zenodo-Dep="14734546" httpUri="https://zenodo.org/record/14734546/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,264,1224,1248]" targetBox="[159,1429,189,1188]" targetPageId="40" targetType="figure">
<paragraph id="8BE4565BFFF2E273FF3922A2FB65FAA9" blockId="40.[151,1436,1224,1321]" pageId="40" pageNumber="41">
<emphasis id="B92F8A49FFF2E273FF3922A2FE93FB61" bold="true" box="[151,365,1224,1248]" pageId="40" pageNumber="41">FIGURES 182186.</emphasis>
<taxonomicName id="4C5B2DD8FFF2E273FEDD22A2FD03FB61" authority="Cassola &amp; Brzoska" authorityName="Cassola &amp; Brzoska" authorityYear="2008" box="[371,765,1224,1249]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mamasa">
@ -562,7 +563,7 @@ leg.” // “ex col.
.
</paragraph>
</subSubSection>
<caption id="DF2406D3FFF3E272FF392174FC5AF86B" pageId="41" pageNumber="42" startId="41.[151,264,1822,1846]" targetBox="[162,1428,161,1807]" targetPageId="41" targetType="figure">
<caption id="DF2406D3FFF3E272FF392174FC5AF86B" ID-DOI="http://doi.org/10.5281/zenodo.14734548" ID-Zenodo-Dep="14734548" httpUri="https://zenodo.org/record/14734548/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,264,1822,1846]" targetBox="[162,1428,161,1807]" targetPageId="41" targetType="figure">
<paragraph id="8BE4565BFFF3E272FF392174FC5AF86B" blockId="41.[151,1436,1822,2026]" pageId="41" pageNumber="42">
<emphasis id="B92F8A49FFF3E272FF392174FE93F8B7" bold="true" box="[151,365,1822,1846]" pageId="41" pageNumber="42">FIGURES 187199.</emphasis>
<taxonomicName id="4C5B2DD8FFF3E272FEDC2174FD02F8B7" authority="Cassola &amp; Brzoska" authorityName="Cassola &amp; Brzoska" authorityYear="2008" box="[370,764,1822,1846]" class="Insecta" family="Carabidae" genus="Calomera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mamasa">
@ -584,31 +585,31 @@ Cassola &amp; Brzoska
<emphasis id="B92F8A49FFF0E271FE7B26FCFD57FF31" box="[469,681,150,176]" italics="true" pageId="42" pageNumber="43">Calomera mamasa</emphasis>
</taxonomicName>
belongs to the largest species of the complex. Body 13.515.5 (HT 14.0) mm long, 4.955.45 (HT 5.05) mm wide. It can be easily distinguished from other species of the species-complex by its remarkable dorsal coloration: head and pronotum deep coppery with reddish reflections, notably prevailingly reddish-cupreous on the pronotum, while elytra almost velvety black, usually with faintly diffusing greenish lustre on anteapical-lateral area and indistinct cupreous lustre on basal area. Setosity: genae glabrous (
<figureCitation id="13604ADEFFF0E271FAC1274CFF3AFEE5" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." pageId="42" pageNumber="43">Fig. 168</figureCitation>
<figureCitation id="13604ADEFFF0E271FAC1274CFF3AFEE5" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." figureDoi="http://doi.org/10.5281/zenodo.14734540" httpUri="https://zenodo.org/record/14734540/files/figure.png" pageId="42" pageNumber="43">Fig. 168</figureCitation>
), proepisterna with setae at their ventral margin, sometimes up to proepisternal half (as in HT,
<figureCitation id="13604ADEFFF0E271FB482720FAB9FEE5" box="[1254,1351,330,356]" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." pageId="42" pageNumber="43">Fig. 168</figureCitation>
<figureCitation id="13604ADEFFF0E271FB482720FAB9FEE5" box="[1254,1351,330,356]" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." figureDoi="http://doi.org/10.5281/zenodo.14734540" httpUri="https://zenodo.org/record/14734540/files/figure.png" pageId="42" pageNumber="43">Fig. 168</figureCitation>
); rather copious or very sparse setae along dorsolateral pronotal margins (
<figureCitation id="13604ADEFFF0E271FC362704FBC4FE09" box="[920,1082,366,392]" captionStart="FIGURES 182186" captionStartId="40.[151,264,1224,1248]" captionTargetBox="[159,1429,189,1188]" captionTargetId="figure-27@40.[151,1435,181,1200]" captionTargetPageId="40" captionText="FIGURES 182186. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi, pronotum. 182—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 183— ♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 184—♂, Sabbang (JWCM); 185—♂, Barat, Mamasa (PSCH). 186—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR). Bars = 1 mm." pageId="42" pageNumber="43">Figs 182186</figureCitation>
<figureCitation id="13604ADEFFF0E271FC362704FBC4FE09" box="[920,1082,366,392]" captionStart="FIGURES 182186" captionStartId="40.[151,264,1224,1248]" captionTargetBox="[159,1429,189,1188]" captionTargetId="figure-27@40.[151,1435,181,1200]" captionTargetPageId="40" captionText="FIGURES 182186. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi, pronotum. 182—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 183— ♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 184—♂, Sabbang (JWCM); 185—♂, Barat, Mamasa (PSCH). 186—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734546" httpUri="https://zenodo.org/record/14734546/files/figure.png" pageId="42" pageNumber="43">Figs 182186</figureCitation>
). Elytral maculation yellowish or pale orange-yellow, conspicuously wide, laterally adjacent to outer elytral margin (
<figureCitation id="13604ADEFFF0E271FBD127F8FB16FE2D" box="[1151,1256,402,428]" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." pageId="42" pageNumber="43">Figs 167</figureCitation>
<figureCitation id="13604ADEFFF0E271FBD127F8FB16FE2D" box="[1151,1256,402,428]" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." figureDoi="http://doi.org/10.5281/zenodo.14734540" httpUri="https://zenodo.org/record/14734540/files/figure.png" pageId="42" pageNumber="43">Figs 167</figureCitation>
,
<figureCitation id="13604ADEFFF0E271FB5927F8FADDFE2D" box="[1271,1315,402,428]" captionStart="FIGURES 172178" captionStartId="38.[151,264,1864,1888]" captionTargetBox="[164,1427,194,1831]" captionTargetId="figure-27@38.[151,1435,181,1841]" captionTargetPageId="38" captionText="FIGURES 172178. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 172—habitus, ♂, 14.5 mm, Malabo, Mamasa Mamuju road, PT (FCCR-MCZR); 173—head, ♂, ibid., PT (FCCR-MCZR); 174178—labrum: 174—♂, ibid., PT (FCCR- MCZR); 175—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 176—♂, Barat, Mamasa (PSCH); 177—♂, Sabbang (JWCM); 178—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR). Bars = 1 mm." pageId="42" pageNumber="43">172</figureCitation>
<figureCitation id="13604ADEFFF0E271FB5927F8FADDFE2D" box="[1271,1315,402,428]" captionStart="FIGURES 172178" captionStartId="38.[151,264,1864,1888]" captionTargetBox="[164,1427,194,1831]" captionTargetId="figure-27@38.[151,1435,181,1841]" captionTargetPageId="38" captionText="FIGURES 172178. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 172—habitus, ♂, 14.5 mm, Malabo, Mamasa Mamuju road, PT (FCCR-MCZR); 173—head, ♂, ibid., PT (FCCR-MCZR); 174178—labrum: 174—♂, ibid., PT (FCCR- MCZR); 175—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 176—♂, Barat, Mamasa (PSCH); 177—♂, Sabbang (JWCM); 178—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734542" httpUri="https://zenodo.org/record/14734542/files/figure.png" pageId="42" pageNumber="43">172</figureCitation>
,
<figureCitation id="13604ADEFFF0E271FA9D27F8FA66FE2D" box="[1331,1432,402,428]" captionStart="FIGURES 179181" captionStartId="39.[151,264,1885,1909]" captionTargetBox="[160,1428,210,1852]" captionTargetId="figure-18@39.[151,1436,199,1862]" captionTargetPageId="39" captionText="FIGURES 179181. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 179180—habitus: 179—♂, 14.3 mm, Sabbang (JWCM); 180—♂, 14.9 mm, Tengah / Gunung Sampuraga (AOCW); 181—male aedeagus showing internal sac, Barat, Mamasa (PSCH). Bars = 1 mm." pageId="42" pageNumber="43">179180</figureCitation>
<figureCitation id="13604ADEFFF0E271FA9D27F8FA66FE2D" box="[1331,1432,402,428]" captionStart="FIGURES 179181" captionStartId="39.[151,264,1885,1909]" captionTargetBox="[160,1428,210,1852]" captionTargetId="figure-18@39.[151,1436,199,1862]" captionTargetPageId="39" captionText="FIGURES 179181. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 179180—habitus: 179—♂, 14.3 mm, Sabbang (JWCM); 180—♂, 14.9 mm, Tengah / Gunung Sampuraga (AOCW); 181—male aedeagus showing internal sac, Barat, Mamasa (PSCH). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734544" httpUri="https://zenodo.org/record/14734544/files/figure.png" pageId="42" pageNumber="43">179180</figureCitation>
,
<figureCitation id="13604ADEFFF0E271FF3927DCFF03FE51" box="[151,253,438,464]" captionStart="FIGURES 187199" captionStartId="41.[151,264,1822,1846]" captionTargetBox="[162,1428,161,1807]" captionTargetId="figure-18@41.[151,1436,151,1814]" captionTargetPageId="41" captionText="FIGURES 187199. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 187191—elytron: 187—♂, Barat, Polewali— Mamasa road, PT (FCCR-MCZR);188—♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 189—♂, Barat, Mamasa (PSCH); 190—♂, Sabbang (JWCM); 191—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 192199—male aedeagus or its apex: 192—PolewaliMamasa road, PT (CCJM); 193— Malabo, MamasaMamuju road, PT (FCCR-MCZR); 194— ditto, dorsal view; 195—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 196—♂, Barat, Mamasa (PSCH); 197—ditto., dorsal view; 198—Sabbang (JWCM); 199—ditto, dorsal view. Bars = 1 mm." pageId="42" pageNumber="43">187191</figureCitation>
<figureCitation id="13604ADEFFF0E271FF3927DCFF03FE51" box="[151,253,438,464]" captionStart="FIGURES 187199" captionStartId="41.[151,264,1822,1846]" captionTargetBox="[162,1428,161,1807]" captionTargetId="figure-18@41.[151,1436,151,1814]" captionTargetPageId="41" captionText="FIGURES 187199. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 187191—elytron: 187—♂, Barat, Polewali— Mamasa road, PT (FCCR-MCZR);188—♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 189—♂, Barat, Mamasa (PSCH); 190—♂, Sabbang (JWCM); 191—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 192199—male aedeagus or its apex: 192—PolewaliMamasa road, PT (CCJM); 193— Malabo, MamasaMamuju road, PT (FCCR-MCZR); 194— ditto, dorsal view; 195—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 196—♂, Barat, Mamasa (PSCH); 197—ditto., dorsal view; 198—Sabbang (JWCM); 199—ditto, dorsal view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734548" httpUri="https://zenodo.org/record/14734548/files/figure.png" pageId="42" pageNumber="43">187191</figureCitation>
). Labrum (
<figureCitation id="13604ADEFFF0E271FED627DCFDECFE51" box="[376,530,438,464]" captionStart="FIGURES 172178" captionStartId="38.[151,264,1864,1888]" captionTargetBox="[164,1427,194,1831]" captionTargetId="figure-27@38.[151,1435,181,1841]" captionTargetPageId="38" captionText="FIGURES 172178. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 172—habitus, ♂, 14.5 mm, Malabo, Mamasa Mamuju road, PT (FCCR-MCZR); 173—head, ♂, ibid., PT (FCCR-MCZR); 174178—labrum: 174—♂, ibid., PT (FCCR- MCZR); 175—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 176—♂, Barat, Mamasa (PSCH); 177—♂, Sabbang (JWCM); 178—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR). Bars = 1 mm." pageId="42" pageNumber="43">Figs 174178</figureCitation>
<figureCitation id="13604ADEFFF0E271FED627DCFDECFE51" box="[376,530,438,464]" captionStart="FIGURES 172178" captionStartId="38.[151,264,1864,1888]" captionTargetBox="[164,1427,194,1831]" captionTargetId="figure-27@38.[151,1435,181,1841]" captionTargetPageId="38" captionText="FIGURES 172178. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 172—habitus, ♂, 14.5 mm, Malabo, Mamasa Mamuju road, PT (FCCR-MCZR); 173—head, ♂, ibid., PT (FCCR-MCZR); 174178—labrum: 174—♂, ibid., PT (FCCR- MCZR); 175—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 176—♂, Barat, Mamasa (PSCH); 177—♂, Sabbang (JWCM); 178—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734542" httpUri="https://zenodo.org/record/14734542/files/figure.png" pageId="42" pageNumber="43">Figs 174178</figureCitation>
) primarily 6setose as in other species, yet examined specimens possess also 78 labral setae (in male
<typeStatus id="54E0E8F9FFF0E271FE2E27B0FE1CFE75" box="[384,482,474,500]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
only five setae developed). Aedeagus similar to other species in its lateral view (
<figureCitation id="13604ADEFFF0E271FAC427B0FF3DFD99" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." pageId="42" pageNumber="43">Figs 170</figureCitation>
<figureCitation id="13604ADEFFF0E271FAC427B0FF3DFD99" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." figureDoi="http://doi.org/10.5281/zenodo.14734540" httpUri="https://zenodo.org/record/14734540/files/figure.png" pageId="42" pageNumber="43">Figs 170</figureCitation>
,
<figureCitation id="13604ADEFFF0E271FF612794FE22FD99" box="[207,476,510,537]" captionStart="FIGURES 187199" captionStartId="41.[151,264,1822,1846]" captionTargetBox="[162,1428,161,1807]" captionTargetId="figure-18@41.[151,1436,151,1814]" captionTargetPageId="41" captionText="FIGURES 187199. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 187191—elytron: 187—♂, Barat, Polewali— Mamasa road, PT (FCCR-MCZR);188—♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 189—♂, Barat, Mamasa (PSCH); 190—♂, Sabbang (JWCM); 191—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 192199—male aedeagus or its apex: 192—PolewaliMamasa road, PT (CCJM); 193— Malabo, MamasaMamuju road, PT (FCCR-MCZR); 194— ditto, dorsal view; 195—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 196—♂, Barat, Mamasa (PSCH); 197—ditto., dorsal view; 198—Sabbang (JWCM); 199—ditto, dorsal view. Bars = 1 mm." pageId="42" pageNumber="43">192193, 195196, 198</figureCitation>
<figureCitation id="13604ADEFFF0E271FF612794FE22FD99" box="[207,476,510,537]" captionStart="FIGURES 187199" captionStartId="41.[151,264,1822,1846]" captionTargetBox="[162,1428,161,1807]" captionTargetId="figure-18@41.[151,1436,151,1814]" captionTargetPageId="41" captionText="FIGURES 187199. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 187191—elytron: 187—♂, Barat, Polewali— Mamasa road, PT (FCCR-MCZR);188—♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 189—♂, Barat, Mamasa (PSCH); 190—♂, Sabbang (JWCM); 191—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 192199—male aedeagus or its apex: 192—PolewaliMamasa road, PT (CCJM); 193— Malabo, MamasaMamuju road, PT (FCCR-MCZR); 194— ditto, dorsal view; 195—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 196—♂, Barat, Mamasa (PSCH); 197—ditto., dorsal view; 198—Sabbang (JWCM); 199—ditto, dorsal view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734548" httpUri="https://zenodo.org/record/14734548/files/figure.png" pageId="42" pageNumber="43">192193, 195196, 198</figureCitation>
), lacking lateral indentations in its dorsal (or ventral) view (
<figureCitation id="13604ADEFFF0E271FBD52794FB1EFD99" box="[1147,1248,510,536]" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." pageId="42" pageNumber="43">Figs 171</figureCitation>
<figureCitation id="13604ADEFFF0E271FBD52794FB1EFD99" box="[1147,1248,510,536]" captionStart="FIGURES 167171" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[151,1425,181,1834]" captionTargetId="figure-18@37.[151,1436,181,1844]" captionTargetPageId="37" captionText="FIGURES 167171. Calomera mamasa Cassola &amp; Brzoska, Sulawesi TorajaMamasa. 167—habitus, ♂, 14 mm, HT (MSNG).168—lateral view, showing glabrous gena and setose proepisternum; 169—labels of the HT; 170—aedeagus; 171— ditto., dorsal view. Bars = 1 mm (image credit Roberto Poggi (MSNG)." figureDoi="http://doi.org/10.5281/zenodo.14734540" httpUri="https://zenodo.org/record/14734540/files/figure.png" pageId="42" pageNumber="43">Figs 171</figureCitation>
,
<figureCitation id="13604ADEFFF0E271FB422794FA74FD99" box="[1260,1418,510,536]" captionStart="FIGURES 187199" captionStartId="41.[151,264,1822,1846]" captionTargetBox="[162,1428,161,1807]" captionTargetId="figure-18@41.[151,1436,151,1814]" captionTargetPageId="41" captionText="FIGURES 187199. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 187191—elytron: 187—♂, Barat, Polewali— Mamasa road, PT (FCCR-MCZR);188—♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 189—♂, Barat, Mamasa (PSCH); 190—♂, Sabbang (JWCM); 191—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 192199—male aedeagus or its apex: 192—PolewaliMamasa road, PT (CCJM); 193— Malabo, MamasaMamuju road, PT (FCCR-MCZR); 194— ditto, dorsal view; 195—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 196—♂, Barat, Mamasa (PSCH); 197—ditto., dorsal view; 198—Sabbang (JWCM); 199—ditto, dorsal view. Bars = 1 mm." pageId="42" pageNumber="43">194, 197, 199</figureCitation>
<figureCitation id="13604ADEFFF0E271FB422794FA74FD99" box="[1260,1418,510,536]" captionStart="FIGURES 187199" captionStartId="41.[151,264,1822,1846]" captionTargetBox="[162,1428,161,1807]" captionTargetId="figure-18@41.[151,1436,151,1814]" captionTargetPageId="41" captionText="FIGURES 187199. Calomera mamasa Cassola &amp; Brzoska, all from Sulawesi. 187191—elytron: 187—♂, Barat, Polewali— Mamasa road, PT (FCCR-MCZR);188—♂, Malabo, MamasaMamuju road, PT (FCCR-MCZR); 189—♂, Barat, Mamasa (PSCH); 190—♂, Sabbang (JWCM); 191—♀, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 192199—male aedeagus or its apex: 192—PolewaliMamasa road, PT (CCJM); 193— Malabo, MamasaMamuju road, PT (FCCR-MCZR); 194— ditto, dorsal view; 195—♂, Barat, Polewali—Mamasa road, PT (FCCR-MCZR); 196—♂, Barat, Mamasa (PSCH); 197—ditto., dorsal view; 198—Sabbang (JWCM); 199—ditto, dorsal view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14734548" httpUri="https://zenodo.org/record/14734548/files/figure.png" pageId="42" pageNumber="43">194, 197, 199</figureCitation>
).
</paragraph>
<paragraph id="8BE4565BFFF0E271FF692449FD02FDBD" blockId="42.[151,1437,150,1041]" box="[199,764,546,573]" pageId="42" pageNumber="43">

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<mods:title id="576EDE68EB9A43F148B8356FF1B931DC">Description of two new species of Rhaphidophora (Orthoptera: Rhaphidophoridae: Rhaphidophorinae) from Yunnan, China</mods:title>
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<subSubSection id="C34ECED2FFD0B102FF6088A8FEA3FA17" pageId="3" pageNumber="397" type="nomenclature">
<paragraph id="8BEB9D59FFD0B102FF6088A8FDF7FA34" blockId="3.[151,595,1410,1509]" box="[151,595,1410,1437]" pageId="3" pageNumber="397">
<heading id="D0A32A35FFD0B102FF6088A8FDF7FA34" bold="true" box="[151,595,1410,1437]" fontSize="11" level="1" pageId="3" pageNumber="397" reason="1">
<emphasis id="B920414BFFD0B102FF6088A8FDF7FA34" bold="true" box="[151,595,1410,1437]" pageId="3" pageNumber="397">
<taxonomicName id="4C54E6DAFFD0B102FF6088A8FE4AFA34" authorityName="Yu &amp; Liu &amp; Bian &amp; Zhang" authorityYear="2025" box="[151,494,1410,1437]" class="Dothideomycetes" family="Leptosphaeriaceae" genus="Rhaphidophora" kingdom="Fungi" order="Pleosporales" pageId="3" pageNumber="397" phylum="Ascomycota" rank="species" species="minutispina" status="sp. nov.">
<emphasis id="B920414BFFD0B102FF6088A8FE4AFA34" bold="true" box="[151,494,1410,1437]" italics="true" pageId="3" pageNumber="397">Rhaphidophora minutispina</emphasis>
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<taxonomicNameLabel id="A213FC30FFD0B102FE0188A9FDF7FA34" box="[502,595,1411,1437]" pageId="3" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BEB9D59FFD0B102FF60888DFEA3FA17" blockId="3.[151,595,1410,1509]" box="[151,263,1447,1470]" pageId="3" pageNumber="397">
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<emphasis id="B920414BFFD0B102FF60888DFEA3FA17" bold="true" box="[151,263,1447,1470]" pageId="3" pageNumber="397">微ḦȆdz</emphasis>
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<subSubSection id="C34ECED2FFD0B105FF6088E1FE10F8B5" lastPageId="4" lastPageNumber="398" pageId="3" pageNumber="397" type="description">
<paragraph id="8BEB9D59FFD0B102FF6088E1FEB8FA4C" blockId="3.[151,595,1410,1509]" box="[151,284,1483,1509]" pageId="3" pageNumber="397">
<figureCitation id="136F81DCFFD0B102FF6088E1FEB8FA4C" box="[151,284,1483,1509]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1327,1351]" captionStartId-1="5.[151,250,1795,1819]" captionTargetBox-0="[151,1436,179,1300]" captionTargetBox-1="[151,1436,597,1772]" captionTargetId-0="figure-18@4.[151,1436,179,1300]" captionTargetId-1="figure-202@5.[151,1436,597,1772]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." captionText-1="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GH. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Figures 34</figureCitation>
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF608B38FB66F8AC" blockId="3.[151,1437,1554,2013]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF608B38FED7F984" bold="true" box="[151,371,1554,1581]" pageId="3" pageNumber="397">Description. Male.</emphasis>
Body medium.
<emphasis id="B920414BFFD0B102FDDF8B39FDC8F984" box="[552,620,1555,1581]" italics="true" pageId="3" pageNumber="397">Head.</emphasis>
Head and pronotum as other species of the genus.
<emphasis id="B920414BFFD0B102FB528B39FB45F985" box="[1189,1249,1555,1580]" italics="true" pageId="3" pageNumber="397">Legs.</emphasis>
Fore coxae with 1 spine; femora with 1 short ventral spine on internal margin; tibiae ventrally with 1 internal and 2 external spines, apices with 1 pair of spines on ventral surface. Middle femora with 1 pair of apical spines; tibiae dorsally with 2 pairs of spines, ventral surface with 13 pairs of spines, apices with 1 pair of spines on dorsal and ventral surfaces separately. Hind femora with 1 short internal spine (
<figureCitation id="136F81DCFFD0B102FD018B89FCF4F914" box="[758,848,1699,1725]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." pageId="3" pageNumber="397">Fig. 3E</figureCitation>
); tibiae dorsally with 1925 internal spines, 2123 external spines, and 1 pair of small subapical spines, apices with 1 pair of dorsal spines and 2 pairs of ventral spines; basitarsi with 46 small spines and 1 apical spine on dorsal surface along the midline (
<figureCitation id="136F81DCFFD0B102FB978BC1FB11F8AC" box="[1120,1205,1771,1797]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." pageId="3" pageNumber="397">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF308A25FAE8F874" blockId="3.[151,1437,1554,2013]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF308A25FE9DF880" box="[199,313,1807,1833]" italics="true" pageId="3" pageNumber="397">Abdomen.</emphasis>
Posterior margin of tenth abdominal tergite faintly concave in middle. Dorsal surface of epiproct with obviously raised lateral margins, between them concave (
<figureCitation id="136F81DCFFD0B102FCD28A19FCD9F8E4" box="[805,893,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." pageId="3" pageNumber="397">Fig. 3H</figureCitation>
), and then prolonged into a pair of wide spines (
<figureCitation id="136F81DCFFD0B102FF688A7DFF53F8D8" box="[159,247,1879,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." pageId="3" pageNumber="397">Fig. 3G</figureCitation>
), the internal margin of the spine with 1 minute spine about the middle area (
<figureCitation id="136F81DCFFD0B102FB918A7DFB19F8D8" box="[1126,1213,1879,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." pageId="3" pageNumber="397">Fig. 3H</figureCitation>
); ventral surface of epiproct transverse, posterolateral area with 1 thin spine on each side, its apices directing forward and acute (
<figureCitation id="136F81DCFFD0B102FA988A51FF15F810" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." pageId="3" pageNumber="397">Fig. 3I</figureCitation>
). Subgenital plate wider than long, lateral margins convex, posterior margin truncate; styli conical, about two times as long as subgenital plate, inserted on the posterolateral areas of the plate (
<figureCitation id="136F81DCFFD0B102FBDF8AE9FB26F874" box="[1064,1154,1987,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." pageId="3" pageNumber="397">Fig. 3K</figureCitation>
), apices subacute.
</paragraph>
<caption id="DF2BCDD1FFD7B105FF608805FC7CFA26" pageId="4" pageNumber="398" startId="4.[151,250,1327,1351]" targetBox="[151,1436,179,1300]" targetPageId="4" targetType="figure">
<paragraph id="8BEB9D59FFD7B105FF608805FC7CFA26" blockId="4.[151,1436,1326,1423]" pageId="4" pageNumber="398">
<emphasis id="B920414BFFD7B105FF608805FCA4FAEE" bold="true" box="[151,768,1326,1351]" pageId="4" pageNumber="398">
FIGURE 3.
<taxonomicName id="4C54E6DAFFD7B105FEEC8804FDE7FAEE" authorityName="Yu &amp; Liu &amp; Bian &amp; Zhang" authorityYear="2025" box="[283,579,1326,1351]" class="Dothideomycetes" family="Leptosphaeriaceae" genus="Rhaphidophora" kingdom="Fungi" order="Pleosporales" pageId="4" pageNumber="398" phylum="Ascomycota" rank="species" species="minutispina" status="sp. nov.">
<emphasis id="B920414BFFD7B105FEEC8804FDE7FAEE" bold="true" box="[283,579,1326,1351]" italics="true" pageId="4" pageNumber="398">Rhaphidophora minutispina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213FC30FFD7B105FDBD8805FD3FFAEE" box="[586,667,1327,1351]" pageId="4" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
Male: A.
</emphasis>
head in frontal view;
<emphasis id="B920414BFFD7B105FC158805FBBDFAEE" bold="true" box="[994,1049,1327,1351]" pageId="4" pageNumber="398">BC.</emphasis>
head and thoraces:
<emphasis id="B920414BFFD7B105FB138805FB5FFAEE" bold="true" box="[1252,1275,1327,1351]" pageId="4" pageNumber="398">B.</emphasis>
dorsal view,
<emphasis id="B920414BFFD7B105FA738805FA38FAEE" bold="true" box="[1412,1436,1327,1351]" pageId="4" pageNumber="398">C.</emphasis>
lateral view;
<emphasis id="B920414BFFD7B105FEE98879FEF0FAC2" bold="true" box="[286,340,1363,1387]" pageId="4" pageNumber="398">DE.</emphasis>
apex of hind femur:
<emphasis id="B920414BFFD7B105FDD98879FDE3FAC2" bold="true" box="[558,583,1363,1387]" pageId="4" pageNumber="398">D.</emphasis>
external view,
<emphasis id="B920414BFFD7B105FD128879FD58FAC2" bold="true" box="[741,764,1363,1387]" pageId="4" pageNumber="398">E.</emphasis>
internal view;
<emphasis id="B920414BFFD7B105FC6F8879FC0FFAC2" bold="true" box="[920,939,1363,1387]" pageId="4" pageNumber="398">F.</emphasis>
tarsus of hind leg in lateral view;
<emphasis id="B920414BFFD7B105FAF98879FAEDFAC2" bold="true" box="[1294,1353,1363,1387]" pageId="4" pageNumber="398">GK.</emphasis>
apex of abdomen:
<emphasis id="B920414BFFD7B105FEF6885DFEB8FA26" bold="true" box="[257,284,1399,1423]" pageId="4" pageNumber="398">G.</emphasis>
apical view,
<emphasis id="B920414BFFD7B105FE56885DFE1FFA26" bold="true" box="[417,443,1399,1423]" pageId="4" pageNumber="398">H.</emphasis>
apico-lateral view,
<emphasis id="B920414BFFD7B105FD75885DFD36FA26" bold="true" box="[642,658,1399,1423]" pageId="4" pageNumber="398">I.</emphasis>
lateral view,
<emphasis id="B920414BFFD7B105FCEC885DFCEBFA26" bold="true" box="[795,847,1399,1423]" pageId="4" pageNumber="398">JK.</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="8BEB9D59FFD7B105FF3088CBFD49F922" blockId="4.[151,1436,1505,2036]" pageId="4" pageNumber="398">
<emphasis id="B920414BFFD7B105FF3088CBFE81FA52" bold="true" box="[199,293,1505,1531]" pageId="4" pageNumber="398">Female.</emphasis>
Tenth abdominal tergite concave in middle (
<figureCitation id="136F81DCFFD7B105FCCE88CBFC37FA52" box="[825,915,1505,1531]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GH. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4G</figureCitation>
). Epiproct roughly triangular, dorsal surface concave along middle, posterior margin rounded (
<figureCitation id="136F81DCFFD7B105FD4B8B2FFCB5F9B6" box="[700,785,1541,1567]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GH. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4G</figureCitation>
). Cerci slender. Seventh abdominal sternite longer than the preceding sternites. Subgenital plate semi-circular, separated by a prolonged membranous zone from the preceding sternite (
<figureCitation id="136F81DCFFD7B105FF0B8B67FEE9F9CE" box="[252,333,1613,1639]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GH. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4I</figureCitation>
), posterior margin with 1 median concavity (
<figureCitation id="136F81DCFFD7B105FCAE8B67FC08F9CE" box="[857,940,1613,1639]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GH. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4J</figureCitation>
). Ovipositor about as long as hind femora, gradually upcurved behind the middle area (
<figureCitation id="136F81DCFFD7B105FD718B5BFD79F925" box="[646,733,1649,1676]" captionStart="FIGURE 4" captionStartId="5.[151,250,1795,1819]" captionTargetBox="[151,1436,597,1772]" captionTargetId="figure-202@5.[151,1436,597,1772]" captionTargetPageId="5" captionText="FIGURE 4. Rhaphidophora minutispina sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GH. apex of abdomen: G. dorsal view, H. lateral view; I. subgenital plate in ventral view; J. seventh abdominal sternite and subgenital plate in ventral view." pageId="4" pageNumber="398">Fig. 4H</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD7B105FF308BBFFA9BF906" blockId="4.[151,1436,1505,2036]" box="[199,1343,1685,1711]" pageId="4" pageNumber="398">
<emphasis id="B920414BFFD7B105FF308BBFFEEBF906" bold="true" box="[199,335,1685,1711]" pageId="4" pageNumber="398">Coloration.</emphasis>
Male body blackish brown, dorsal surface of head almost black. Apices of femora darker.
</paragraph>
<paragraph id="8BEB9D59FFD7B105FF308B93FE10F8B5" blockId="4.[151,1436,1505,2036]" pageId="4" pageNumber="398">
<emphasis id="B920414BFFD7B105FF308B93FE62F97A" bold="true" box="[199,454,1721,1747]" pageId="4" pageNumber="398">Measurements (mm).</emphasis>
BL:
<specimenCount id="9D5256D0FFD7B105FDF68B97FDB2F97A" box="[513,534,1725,1747]" count="1" pageId="4" pageNumber="398" type="male"></specimenCount>
17.417.8,
<specimenCount id="9D5256D0FFD7B105FD6C8B90FD14F97D" box="[667,688,1722,1748]" count="1" pageId="4" pageNumber="398" type="female"></specimenCount>
15.718.6; PL:
<specimenCount id="9D5256D0FFD7B105FC908B97FCD8F97A" box="[871,892,1725,1747]" count="1" pageId="4" pageNumber="398" type="male"></specimenCount>
5.05.2,
<specimenCount id="9D5256D0FFD7B105FC118B90FC5FF97D" box="[998,1019,1722,1748]" count="1" pageId="4" pageNumber="398" type="female"></specimenCount>
5.15.7; FFL:
<specimenCount id="9D5256D0FFD7B105FB528B97FB1EF97A" box="[1189,1210,1725,1747]" count="1" pageId="4" pageNumber="398" type="male"></specimenCount>
6.06.1,
<specimenCount id="9D5256D0FFD7B105FAD38B90FA9DF97D" box="[1316,1337,1722,1748]" count="1" pageId="4" pageNumber="398" type="female"></specimenCount>
5.77.7; MFL:
<specimenCount id="9D5256D0FFD7B105FF178BCBFF51F95E" box="[224,245,1761,1783]" count="1" pageId="4" pageNumber="398" type="male"></specimenCount>
6.06.2,
<specimenCount id="9D5256D0FFD7B105FE978BF4FED1F951" box="[352,373,1758,1784]" count="1" pageId="4" pageNumber="398" type="female"></specimenCount>
6.06.2; HFL:
<specimenCount id="9D5256D0FFD7B105FDD18BCBFD9FF95E" box="[550,571,1761,1783]" count="1" pageId="4" pageNumber="398" type="male"></specimenCount>
14.615.7,
<specimenCount id="9D5256D0FFD7B105FD368BF4FD72F951" box="[705,726,1758,1784]" count="1" pageId="4" pageNumber="398" type="female"></specimenCount>
14.716.0; HTL:
<specimenCount id="9D5256D0FFD7B105FC538BCBFC1DF95E" box="[932,953,1761,1783]" count="1" pageId="4" pageNumber="398" type="male"></specimenCount>
12.813.5,
<specimenCount id="9D5256D0FFD7B105FBB78BF4FBF1F951" box="[1088,1109,1758,1784]" count="1" pageId="4" pageNumber="398" type="female"></specimenCount>
12.813.5; HBL:
<specimenCount id="9D5256D0FFD7B105FAD38BCBFA9DF95E" box="[1316,1337,1761,1783]" count="1" pageId="4" pageNumber="398" type="male"></specimenCount>
2.83.0,
<specimenCount id="9D5256D0FFD7B105FF608A28FF08F8B5" box="[151,172,1794,1820]" count="1" pageId="4" pageNumber="398" type="female"></specimenCount>
2.43.0; OvL: 6.59.2.
</paragraph>
</subSubSection>
<subSubSection id="C34ECED2FFD7B104FF308A0FFA38FEE8" lastPageId="5" lastPageNumber="399" pageId="4" pageNumber="398" type="materials_examined">
<paragraph id="8BEB9D59FFD7B104FF308A0FFA38FEE8" blockId="4.[151,1436,1505,2036]" lastBlockId="5.[151,1437,150,537]" lastPageId="5" lastPageNumber="399" pageId="4" pageNumber="398">
<emphasis id="B920414BFFD7B105FF308A0FFE14F896" bold="true" box="[199,432,1829,1855]" pageId="4" pageNumber="398">Material examined.</emphasis>
<materialsCitation id="3B3C9704FFD7B105FE418A0FFE97F8CD" collectingDate="2024-08-20" location="Gongshan" municipality="Dulongjiang" pageId="4" pageNumber="398" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54EF23FBFFD7B105FE418A0FFD87F896" box="[438,547,1829,1855]" pageId="4" pageNumber="398" type="holotype">Holotype</typeStatus>
: male,
<collectingMunicipality id="6B8F0723FFD7B105FD878A0FFD5BF896" box="[624,767,1829,1855]" pageId="4" pageNumber="398">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD7B105FCFE8A0FFCD9F896" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD7B105FCFE8A0FFCD9F896" box="[777,893,1829,1855]" municipality="Dulongjiang" name="Gongshan" pageId="4" pageNumber="398" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD7B105FC708A0CFC44F8E9" box="[903,992,1830,1856]" country="China" name="Yunnan" pageId="4" pageNumber="398">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD7B105FC1F8A0FFB00F896" box="[1000,1188,1829,1855]" pageId="4" pageNumber="398" value="2024-08-20">
<collectingDate id="EFAE4271FFD7B105FC1F8A0FFB00F896" box="[1000,1188,1829,1855]" pageId="4" pageNumber="398" value="2024-08-20">August 20, 2024</collectingDate>
</date>
, coll. by Yangting Qin and Ting Luo
</materialsCitation>
.
<materialsCitation id="3B3C9704FFD7B105FEC98A60FEA7F821" collectingDate="2024-08-21" county="Dulongjiang" location="Yangting Qin" municipality="Gongshan" pageId="4" pageNumber="398" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54EF23FBFFD7B105FEC98A60FE0BF8CD" box="[318,431,1866,1892]" pageId="4" pageNumber="398" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5256D0FFD7B105FE4D8A63FD8DF8CA" box="[442,553,1865,1891]" count="2" pageId="4" pageNumber="398" type="female">2 females</specimenCount>
,
<collectingCounty id="628AE5D5FFD7B105FDC28A63FD60F8CA" box="[565,708,1865,1891]" pageId="4" pageNumber="398">Dulongjiang</collectingCounty>
,
<collectingMunicipality id="6B8F0723FFD7B105FD388A63FCE7F8CA" box="[719,835,1865,1891]" pageId="4" pageNumber="398">Gongshan</collectingMunicipality>
,
<collectingRegion id="499053BBFFD7B105FCB98A60FC03F8CD" box="[846,935,1866,1892]" country="China" name="Yunnan" pageId="4" pageNumber="398">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD7B105FC468A63FBCBF8CA" box="[945,1135,1865,1891]" pageId="4" pageNumber="398" value="2024-08-21">
<collectingDate id="EFAE4271FFD7B105FC468A63FBCBF8CA" box="[945,1135,1865,1891]" pageId="4" pageNumber="398" value="2024-08-21">August 21, 2024</collectingDate>
</date>
, coll. by
<location id="8E8BCB82FFD7B105FB228A63FAC8F8CA" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD7B105FB228A63FAC8F8CA" box="[1237,1388,1865,1891]" county="Dulongjiang" municipality="Gongshan" name="Yangting Qin" pageId="4" pageNumber="398" stateProvince="Yunnan">Yangting Qin</location>
and
<location id="8E8BCB82FFD7B105FF608A47FEA7F821" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD7B105FF608A47FEA7F821" box="[151,259,1901,1928]" county="Dulongjiang" municipality="Gongshan" name="Ting Luo" pageId="4" pageNumber="398" stateProvince="Yunnan">Ting Luo</location>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD7B105FEF98A47FA9BF821" box="[270,1343,1901,1928]" collectingDate="2024-08-23" location="Gongshan" municipality="Dulongjiang" pageId="4" pageNumber="398" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D5256D0FFD7B105FEF98A47FEF8F82E" box="[270,348,1901,1927]" count="1" pageId="4" pageNumber="398" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD7B105FE9F8A47FE53F82E" box="[360,503,1901,1927]" pageId="4" pageNumber="398">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD7B105FDF58A47FDD2F82E" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD7B105FDF58A47FDD2F82E" box="[514,630,1901,1927]" municipality="Dulongjiang" name="Gongshan" pageId="4" pageNumber="398" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD7B105FD768A44FD7EF821" box="[641,730,1902,1928]" country="China" name="Yunnan" pageId="4" pageNumber="398">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD7B105FD148A47FC04F82E" box="[739,928,1901,1927]" pageId="4" pageNumber="398" value="2024-08-23">
<collectingDate id="EFAE4271FFD7B105FD148A47FC04F82E" box="[739,928,1901,1927]" pageId="4" pageNumber="398" value="2024-08-23">August 23, 2024</collectingDate>
</date>
, coll. by Yangting Qin and Ting Luo
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD7B105FABD8A47FBD7F805" collectingDate="2024-08-25" location="Gongshan" municipality="Dulongjiang" pageId="4" pageNumber="398" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D5256D0FFD7B105FABD8A47FA3CF82E" box="[1354,1432,1901,1927]" count="1" pageId="4" pageNumber="398" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD7B105FF608ABBFE82F802" box="[151,294,1937,1963]" pageId="4" pageNumber="398">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD7B105FEC58ABBFE02F802" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD7B105FEC58ABBFE02F802" box="[306,422,1937,1963]" municipality="Dulongjiang" name="Gongshan" pageId="4" pageNumber="398" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD7B105FE468AB8FDAEF805" box="[433,522,1938,1964]" country="China" name="Yunnan" pageId="4" pageNumber="398">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD7B105FDE48ABBFD76F802" box="[531,722,1937,1964]" pageId="4" pageNumber="398" value="2024-08-25">
<collectingDate id="EFAE4271FFD7B105FDE48ABBFD76F802" box="[531,722,1937,1964]" pageId="4" pageNumber="398" value="2024-08-25">August 25, 2024</collectingDate>
</date>
, coll. by Yangting Qin and Ting Luo
</materialsCitation>
. Other specimens:
<materialsCitation id="3B3C9704FFD7B105FABE8ABBFC64F866" collectingDate="2024-08-11" collectorName="Mingyu Liu" location="Gongshan" municipality="Dulongjiang" pageId="4" pageNumber="398" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D5256D0FFD7B105FABE8ABBFA3CF802" box="[1353,1432,1937,1963]" count="1" pageId="4" pageNumber="398" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD7B105FF608A9FFE82F866" box="[151,294,1973,1999]" pageId="4" pageNumber="398">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD7B105FEC68A9FFE01F866" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD7B105FEC68A9FFE01F866" box="[305,421,1973,1999]" municipality="Dulongjiang" name="Gongshan" pageId="4" pageNumber="398" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD7B105FE478A9CFDADF879" box="[432,521,1974,2000]" country="China" name="Yunnan" pageId="4" pageNumber="398">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD7B105FDE48A9FFD6BF866" box="[531,719,1973,1999]" pageId="4" pageNumber="398" value="2024-08-11">
<collectingDate id="EFAE4271FFD7B105FDE48A9FFD6BF866" box="[531,719,1973,1999]" pageId="4" pageNumber="398" value="2024-08-11">August 11, 2024</collectingDate>
</date>
, coll. by
<collectorName id="26A1F88FFFD7B105FCC18A9FFC64F866" box="[822,960,1973,1999]" pageId="4" pageNumber="398">Mingyu Liu</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD7B105FC3B8A9FFD55F85D" collectingDate="2024-08-19" collectorName="Yangting Qin &amp; Ting Luo" location="Gongshan" municipality="Dulongjiang" pageId="4" pageNumber="398" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D5256D0FFD7B105FC3B8A9FFBBFF866" box="[972,1051,1973,1999]" count="1" pageId="4" pageNumber="398" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD7B105FBD18A9FFB11F866" box="[1062,1205,1973,1999]" pageId="4" pageNumber="398">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD7B105FB368A9FFA91F866" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD7B105FB368A9FFA91F866" box="[1217,1333,1973,1999]" municipality="Dulongjiang" name="Gongshan" pageId="4" pageNumber="398" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD7B105FAB78A9CFA3DF879" box="[1344,1433,1974,2000]" country="China" name="Yunnan" pageId="4" pageNumber="398">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD7B105FF608AF3FEF0F85A" box="[151,340,2009,2035]" pageId="4" pageNumber="398" value="2024-08-19">
<collectingDate id="EFAE4271FFD7B105FF608AF3FEF0F85A" box="[151,340,2009,2035]" pageId="4" pageNumber="398" value="2024-08-19">August 19, 2024</collectingDate>
</date>
, coll. by
<collectorName id="26A1F88FFFD7B105FE4E8AF3FDF4F85A" box="[441,592,2009,2035]" pageId="4" pageNumber="398">Yangting Qin</collectorName>
and
<collectorName id="26A1F88FFFD7B105FD718AF3FD55F85D" box="[646,753,2009,2036]" pageId="4" pageNumber="398">Ting Luo</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD7B104FD0B8AF3FD8EFF18" collectingDate="2024-08-20" collectorName="Yangting Qin &amp; Ting Luo" lastPageId="5" lastPageNumber="399" location="Gongshan" municipality="Dulongjiang" pageId="4" pageNumber="398" specimenCount="6" specimenCount-male="6" stateProvince="Yunnan">
<specimenCount id="9D5256D0FFD7B105FD0B8AF3FCF1F85A" box="[764,853,2009,2035]" count="6" pageId="4" pageNumber="398" type="male">6 males</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD7B105FC968AF3FC54F85A" box="[865,1008,2009,2035]" pageId="4" pageNumber="398">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD7B105FC0C8AF3FBCBF85A" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD7B105FC0C8AF3FBCBF85A" box="[1019,1135,2009,2035]" municipality="Dulongjiang" name="Gongshan" pageId="4" pageNumber="398" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD7B105FB8E8AF0FB76F85D" box="[1145,1234,2010,2036]" country="China" name="Yunnan" pageId="4" pageNumber="398">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD7B105FB2C8AF3FA3CF85A" box="[1243,1432,2009,2035]" pageId="4" pageNumber="398" value="2024-08-20">
<collectingDate id="EFAE4271FFD7B105FB2C8AF3FA3CF85A" box="[1243,1432,2009,2035]" pageId="4" pageNumber="398" value="2024-08-20">August 20, 2024</collectingDate>
</date>
, coll. by
<collectorName id="26A1F88FFFD6B104FF068DBDFE2CFF18" box="[241,392,151,177]" pageId="5" pageNumber="399">Yangting Qin</collectorName>
and
<collectorName id="26A1F88FFFD6B104FE498DBDFD8EFF18" box="[446,554,151,177]" pageId="5" pageNumber="399">Ting Luo</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD6B104FDC28DBCFEC7FF7C" collectingDate="2024-08-21" collectorName="Yangting Qin &amp; Ting Luo" location="Gongshan" municipality="Dulongjiang" pageId="5" pageNumber="399" specimenCount="5" specimenCount-male="5" stateProvince="Yunnan">
<specimenCount id="9D5256D0FFD6B104FDC28DBCFD2AFF18" box="[565,654,150,177]" count="5" pageId="5" pageNumber="399" type="male">5 males</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD6B104FD6D8DBDFC8DFF18" box="[666,809,151,177]" pageId="5" pageNumber="399">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD6B104FCC38DBDFC0CFF18" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD6B104FCC38DBDFC0CFF18" box="[820,936,151,177]" municipality="Dulongjiang" name="Gongshan" pageId="5" pageNumber="399" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD6B104FC458DBDFBAFFF18" box="[946,1035,151,177]" country="China" name="Yunnan" pageId="5" pageNumber="399">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD6B104FBE38DBDFB76FF18" box="[1044,1234,151,177]" pageId="5" pageNumber="399" value="2024-08-21">
<collectingDate id="EFAE4271FFD6B104FBE38DBDFB76FF18" box="[1044,1234,151,177]" pageId="5" pageNumber="399" value="2024-08-21">August 21, 2024</collectingDate>
</date>
, coll. by
<collectorName id="26A1F88FFFD6B104FAC08DBDFF65FF7C" pageId="5" pageNumber="399">Yangting Qin</collectorName>
and
<collectorName id="26A1F88FFFD6B104FF008D91FEC7FF7C" box="[247,355,187,213]" pageId="5" pageNumber="399">Ting Luo</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD6B104FE998D91FE96FF50" collectingDate="2024-08-22" collectorName="Yangting Qin &amp; Ting Luo" location="Gongshan" municipality="Dulongjiang" pageId="5" pageNumber="399" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D5256D0FFD6B104FE998D91FE1DFF7C" box="[366,441,187,213]" count="1" pageId="5" pageNumber="399" type="male">1 male</specimenCount>
and
<specimenCount id="9D5256D0FFD6B104FE078D91FDFBFF7C" box="[496,607,187,213]" count="3" pageId="5" pageNumber="399" type="female">3 females</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD6B104FD9D8D91FD5DFF7C" box="[618,761,187,213]" pageId="5" pageNumber="399">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD6B104FCF38D91FCDCFF7C" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD6B104FCF38D91FCDCFF7C" box="[772,888,187,213]" municipality="Dulongjiang" name="Gongshan" pageId="5" pageNumber="399" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD6B104FC758D91FC7FFF7C" box="[898,987,187,213]" country="China" name="Yunnan" pageId="5" pageNumber="399">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD6B104FC138D91FB05FF7C" box="[996,1185,187,213]" pageId="5" pageNumber="399" value="2024-08-22">
<collectingDate id="EFAE4271FFD6B104FC138D91FB05FF7C" box="[996,1185,187,213]" pageId="5" pageNumber="399" value="2024-08-22">August 22, 2024</collectingDate>
</date>
, coll. by
<collectorName id="26A1F88FFFD6B104FAF18D91FA38FF7C" box="[1286,1436,187,213]" pageId="5" pageNumber="399">Yangting Qin</collectorName>
and
<collectorName id="26A1F88FFFD6B104FF318DF5FE96FF50" box="[198,306,223,249]" pageId="5" pageNumber="399">Ting Luo</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD6B104FECA8DF5FEA6FEB4" collectingDate="2024-08-23" collectorName="Yangting Qin &amp; Ting Luo" location="Gongshan" municipality="Dulongjiang" pageId="5" pageNumber="399" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Yunnan">
<specimenCount id="9D5256D0FFD6B104FECA8DF5FE37FF50" box="[317,403,223,249]" count="3" pageId="5" pageNumber="399" type="male">3 males</specimenCount>
and
<specimenCount id="9D5256D0FFD6B104FE3D8DF5FD8AFF50" box="[458,558,223,249]" count="1" pageId="5" pageNumber="399" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD6B104FDCE8DF5FD6CFF50" box="[569,712,223,249]" pageId="5" pageNumber="399">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD6B104FD248DF5FCE3FF50" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD6B104FD248DF5FCE3FF50" box="[723,839,223,249]" municipality="Dulongjiang" name="Gongshan" pageId="5" pageNumber="399" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD6B104FCA58DF5FC0FFF50" box="[850,939,223,249]" country="China" name="Yunnan" pageId="5" pageNumber="399">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD6B104FC438DF5FBD5FF50" box="[948,1137,223,249]" pageId="5" pageNumber="399" value="2024-08-23">
<collectingDate id="EFAE4271FFD6B104FC438DF5FBD5FF50" box="[948,1137,223,249]" pageId="5" pageNumber="399" value="2024-08-23">August 23, 2024</collectingDate>
</date>
, coll. by
<collectorName id="26A1F88FFFD6B104FB218DF5FAC9FF50" box="[1238,1389,223,249]" pageId="5" pageNumber="399">Yangting Qin</collectorName>
and
<collectorName id="26A1F88FFFD6B104FF608C29FEA6FEB4" box="[151,258,259,285]" pageId="5" pageNumber="399">Ting Luo</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD6B104FEFB8C29FF6CFEE8" collectingDate="2024-08-24" collectorName="Yangting Qin &amp; Ting Luo" location="Gongshan" municipality="Dulongjiang" pageId="5" pageNumber="399" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Yunnan">
<specimenCount id="9D5256D0FFD6B104FEFB8C29FEC4FEB4" box="[268,352,259,285]" count="3" pageId="5" pageNumber="399" type="male">3 males</specimenCount>
and
<specimenCount id="9D5256D0FFD6B104FE628C29FDA7FEB4" box="[405,515,259,285]" count="2" pageId="5" pageNumber="399" type="female">2 females</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD6B104FDFA8C29FD38FEB4" box="[525,668,259,285]" pageId="5" pageNumber="399">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD6B104FD528C29FCBDFEB4" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD6B104FD528C29FCBDFEB4" box="[677,793,259,285]" municipality="Dulongjiang" name="Gongshan" pageId="5" pageNumber="399" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD6B104FCD48C29FCD8FEB4" box="[803,892,259,285]" country="China" name="Yunnan" pageId="5" pageNumber="399">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD6B104FC738C29FB9AFEB4" box="[900,1086,259,285]" pageId="5" pageNumber="399" value="2024-08-24">
<collectingDate id="EFAE4271FFD6B104FC738C29FB9AFEB4" box="[900,1086,259,285]" pageId="5" pageNumber="399" value="2024-08-24">August 24, 2024</collectingDate>
</date>
, coll. by
<collectorName id="26A1F88FFFD6B104FB688C29FA91FEB4" box="[1183,1333,259,285]" pageId="5" pageNumber="399">Yangting Qin</collectorName>
and
<collectorName id="26A1F88FFFD6B104FA9F8C29FF6CFEE8" pageId="5" pageNumber="399">Ting Luo</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD6B104FF248C0DFA3DFEE8" box="[211,1433,294,321]" collectingDate="2024-08-25" collectorName="Yangting Qin &amp; Ting Luo" location="Gongshan" municipality="Dulongjiang" pageId="5" pageNumber="399" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D5256D0FFD6B104FF248C0DFEBAFEE8" box="[211,286,295,321]" count="1" pageId="5" pageNumber="399" type="male">1 male</specimenCount>
and
<specimenCount id="9D5256D0FFD6B104FEA28C0DFE1DFEE8" box="[341,441,295,321]" count="1" pageId="5" pageNumber="399" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6B8F0723FFD6B104FE338C0DFDF7FEE8" box="[452,595,295,321]" pageId="5" pageNumber="399">Dulongjiang</collectingMunicipality>
,
<location id="8E8BCB82FFD6B104FDA98C0DFD76FEE8" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD0B104FF6088A8FEB8FDB0:8E8BCB82FFD6B104FDA98C0DFD76FEE8" box="[606,722,295,321]" municipality="Dulongjiang" name="Gongshan" pageId="5" pageNumber="399" stateProvince="Yunnan">Gongshan</location>
,
<collectingRegion id="499053BBFFD6B104FD2A8C0DFC92FEE8" box="[733,822,295,321]" country="China" name="Yunnan" pageId="5" pageNumber="399">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD6B104FCC88C0DFC5FFEE8" box="[831,1019,294,321]" pageId="5" pageNumber="399" value="2024-08-25">
<collectingDate id="EFAE4271FFD6B104FCC88C0DFC5FFEE8" box="[831,1019,294,321]" pageId="5" pageNumber="399" value="2024-08-25">August 25, 2024</collectingDate>
</date>
, coll. by
<collectorName id="26A1F88FFFD6B104FB978C0DFB53FEE8" box="[1120,1271,295,321]" pageId="5" pageNumber="399">Yangting Qin</collectorName>
and
<collectorName id="26A1F88FFFD6B104FADA8C0DFA3DFEE8" box="[1325,1433,295,321]" pageId="5" pageNumber="399">Ting Luo</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34ECED2FFD6B104FF398C60FDF2FECC" box="[206,598,330,357]" pageId="5" pageNumber="399" type="distribution">
<paragraph id="8BEB9D59FFD6B104FF398C60FDF2FECC" blockId="5.[151,1437,150,537]" box="[206,598,330,357]" pageId="5" pageNumber="399">
<emphasis id="B920414BFFD6B104FF398C60FECCFECD" bold="true" box="[206,360,330,356]" pageId="5" pageNumber="399">Distribution.</emphasis>
<collectingRegion id="499053BBFFD6B104FE9A8C61FE67FECC" box="[365,451,331,357]" country="China" name="Yunnan" pageId="5" pageNumber="399">Yunnan</collectingRegion>
(Gongshan).
</paragraph>
</subSubSection>
<subSubSection id="C34ECED2FFD6B104FF308C44FD2FFE78" pageId="5" pageNumber="399" type="discussion">
<paragraph id="8BEB9D59FFD6B104FF308C44FD2FFE78" blockId="5.[151,1437,150,537]" pageId="5" pageNumber="399">
<emphasis id="B920414BFFD6B104FF308C44FEE8FE21" bold="true" box="[199,332,366,392]" pageId="5" pageNumber="399">Discussion.</emphasis>
The new species differs from
<taxonomicName id="4C54E6DAFFD6B104FD6C8C45FB52FE20" authority="Shen, Wang &amp; He, 2024" authorityName="Shen, Wang &amp; He" authorityYear="2024" box="[667,1270,367,393]" class="Dothideomycetes" family="Leptosphaeriaceae" genus="Rhaphidophora" kingdom="Fungi" order="Pleosporales" pageId="5" pageNumber="399" phylum="Ascomycota" rank="species" species="imbricofurca">
<emphasis id="B920414BFFD6B104FD6C8C45FC46FE20" box="[667,994,367,393]" italics="true" pageId="5" pageNumber="399">Rhaphidophora imbricofurca</emphasis>
Shen, Wang &amp; He, 2024
</taxonomicName>
by the internal margin of male epiproct bearing a minuscule spine on the upper plate (
<figureCitation id="136F81DCFFD6B104FC4C8CB9FBB7FE04" box="[955,1043,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." pageId="5" pageNumber="399">Fig. 3G</figureCitation>
), and male subgenital plate with 1 pair of longer and subconical styli (
<figureCitation id="136F81DCFFD6B104FDD38C9DFDDAFE78" box="[548,638,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1327,1351]" captionTargetBox="[151,1436,179,1300]" captionTargetId="figure-18@4.[151,1436,179,1300]" captionTargetPageId="4" captionText="FIGURE 3. Rhaphidophora minutispina sp. nov. Male: A. head in frontal view; BC. head and thoraces: B. dorsal view, C. lateral view; DE. apex of hind femur: D. external view, E. internal view; F. tarsus of hind leg in lateral view; GK. apex of abdomen: G. apical view, H. apico-lateral view, I. lateral view, JK. ventral view." pageId="5" pageNumber="399">Fig. 3K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34ECED2FFD6B104FF308CF1FEB8FDB0" pageId="5" pageNumber="399" type="etymology">
<paragraph id="8BEB9D59FFD6B104FF308CF1FEB8FDB0" blockId="5.[151,1437,150,537]" pageId="5" pageNumber="399">
<emphasis id="B920414BFFD6B104FF308CF1FEEAFE5C" bold="true" box="[199,334,475,501]" pageId="5" pageNumber="399">Etymology.</emphasis>
The new name refers to the male epiproct with a pair of minuscule spines on internal margin of the upper plate.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,361 @@
<document id="C1D03743D868B98B924FA1A13FBC2A01" ID-DOI="10.11646/zootaxa.5570.2.11" ID-ISSN="1175-5326" ID-ZooBank="FEAAC872-3152-4FAC-BED7-5FFFB5457387" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737737037635" checkinUser="plazi" docAuthor="Yu, Tingting, Liu, Mingyu, Bian, Xun &amp; Zhang, Bin" docDate="2025" docId="03FD2C4FFFD3B102FF608ABFFB86FA98" docLanguage="en" docName="zootaxa.5570.2.11.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhaphidophora condoprocera Yu, Liu, Bian &amp; Zhang, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="397" masterDocId="FFC45437FFD3B101FFF78D2AFFA4FFA9" masterDocTitle="Description of two new species of Rhaphidophora (Orthoptera: Rhaphidophoridae: Rhaphidophorinae) from Yunnan, China" masterLastPageNumber="400" masterPageNumber="394" pageNumber="394" updateTime="1737737096432" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="ADD039A7BA77D17C47D5357BB5F8B330">Description of two new species of Rhaphidophora (Orthoptera: Rhaphidophoridae: Rhaphidophorinae) from Yunnan, China</mods:title>
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<mods:name id="F18E5D46082DD9A2121EEAC2BE218C72" type="personal">
<mods:role id="50908DDFC15BB8B048B2AF860930D86C">
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<mods:namePart id="16850E4EDF811C2BE56C3682B63D24B7">Yu, Tingting</mods:namePart>
<mods:nameIdentifier id="77FE8B15053335DF5C7F98AD3EAD9A46" type="ORCID">0009-0005-7392-1243</mods:nameIdentifier>
<mods:affiliation id="370E04312ADD43A102859542B9BC3C7B">College of Life Sciences &amp; Technology, Inner Mongolia Normal University, Hohhot 010022, China.</mods:affiliation>
<mods:nameIdentifier id="A9017A78893A3382907B09ADB9F0E3EF" type="email">yutingtingnmg@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name id="C26B5772A42B87CF90B1F5229F3F7449" type="personal">
<mods:role id="CE77B842F7BE1392ED3B56A8ECC04C8F">
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</mods:role>
<mods:namePart id="C81E1213C6AC192B55412B625D25AE7F">Liu, Mingyu</mods:namePart>
<mods:affiliation id="05EB9AB9C33F3518453E888858698067">College of Agricultural and Life Sciences, Kunming University, Kunming 650214, China.</mods:affiliation>
</mods:name>
<mods:name id="BB8E5A838DD46E8756901E455DB23B76" type="personal">
<mods:role id="D8F30E889167BCD29659073C8EBE3517">
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</mods:role>
<mods:namePart id="13CD59ED7CAD50BAA3208C1D571B40E4">Bian, Xun</mods:namePart>
<mods:affiliation id="55660E9E22E56506D6AA44EF509A8FD9">Key Laboratory of Ecology of Rare and Endangered Species and Environmental Protection (Guangxi Normal University), Ministry of Education, Guilin 541006, China.</mods:affiliation>
</mods:name>
<mods:name id="48B1BAC509E82766D7036F1EEA958CA0" type="personal">
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</mods:role>
<mods:namePart id="9502FE54606B28C88E0F166F7711C9DE">Zhang, Bin</mods:namePart>
<mods:affiliation id="41150C10612291294F09843E35637674">College of Life Sciences &amp; Technology, Inner Mongolia Normal University, Hohhot 010022, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="DD5B8769B549EC1CAFD446F5E1F0F066">text</mods:typeOfResource>
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<mods:titleInfo id="FA97834AD789F4F4D7137C7CC6EFDC95">
<mods:title id="A87AD991F783B312A46CCD7080146611">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B53D46823A31DB7A75809DA80FFAE75A">
<mods:date id="56D7317938CCEE1BAD5E61AE872BBA1A">2025</mods:date>
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<mods:number id="44BE4ABF5E58D904B5B0DA27B9B512E3">2025-01-17</mods:number>
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<mods:detail id="BCAF2C1878230089E3ACA68ECCAC5F14" type="volume">
<mods:number id="9A3ACAA53E7A8DBC4C3518F1E5AD9927">5570</mods:number>
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<mods:number id="B9CF678D75AFA70BAF906B40896D1720">2</mods:number>
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<mods:start id="8C1CEA91C5E6D0C7E7698B1A80BE09E6">394</mods:start>
<mods:end id="53E3E995AB422254CE97B61EFA3CF290">400</mods:end>
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<mods:location id="3942059023370394E3B1C8CB5187039F">
<mods:url id="4D1292E92782CC1FD52E0CCED44A7934">https://doi.org/10.11646/zootaxa.5570.2.11</mods:url>
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<mods:identifier id="128A9FB646C3D5559619A5B004E13EA3" type="DOI">10.11646/zootaxa.5570.2.11</mods:identifier>
<mods:identifier id="62E0922D2223A6A5E052C4D7DB6C68B4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FD2C4FFFD3B102FF608ABFFB86FA98" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD3B102FF608ABFFB86FA98" httpUri="http://treatment.plazi.org/id/03FD2C4FFFD3B102FF608ABFFB86FA98" lastPageId="3" lastPageNumber="397" pageId="0" pageNumber="394">
<subSubSection id="C34ECED2FFD3B101FF608ABFFEA3F87B" pageId="0" pageNumber="394" type="nomenclature">
<paragraph id="8BEB9D59FFD3B101FF608ABFFDCCF819" blockId="0.[151,616,1941,2040]" box="[151,616,1941,1968]" pageId="0" pageNumber="394">
<heading id="D0A32A35FFD3B101FF608ABFFDCCF819" bold="true" box="[151,616,1941,1968]" fontSize="11" level="1" pageId="0" pageNumber="394" reason="1">
<emphasis id="B920414BFFD3B101FF608ABFFDCCF819" bold="true" box="[151,616,1941,1968]" pageId="0" pageNumber="394">
<taxonomicName id="4C54E6DAFFD3B101FF608ABFFDA7F819" authorityName="Yu &amp; Liu &amp; Bian &amp; Zhang" authorityYear="2025" box="[151,515,1941,1968]" class="Dothideomycetes" family="Leptosphaeriaceae" genus="Rhaphidophora" kingdom="Fungi" order="Pleosporales" pageId="0" pageNumber="394" phylum="Ascomycota" rank="species" species="condoprocera" status="sp. nov.">
<emphasis id="B920414BFFD3B101FF608ABFFDA7F819" bold="true" box="[151,515,1941,1968]" italics="true" pageId="0" pageNumber="394">Rhaphidophora condoprocera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213FC30FFD3B101FDFC8ABCFDCCF819" box="[523,616,1942,1968]" pageId="0" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEB9D59FFD3B101FF608A91FEA3F87B" blockId="0.[151,616,1941,2040]" box="[151,263,1979,2002]" pageId="0" pageNumber="394">
<heading id="D0A32A35FFD3B101FF608A91FEA3F87B" bold="true" box="[151,263,1979,2002]" fontSize="11" level="1" pageId="0" pageNumber="394" reason="1">
<emphasis id="B920414BFFD3B101FF608A91FEA3F87B" bold="true" box="[151,263,1979,2002]" pageId="0" pageNumber="394">DȐȆdz</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34ECED2FFD3B102FF608AF4FE10FC0C" lastPageId="3" lastPageNumber="397" pageId="0" pageNumber="394" type="description">
<paragraph id="8BEB9D59FFD3B101FF608AF4FEB8F851" blockId="0.[151,616,1941,2040]" box="[151,284,2014,2040]" pageId="0" pageNumber="394">
<figureCitation id="136F81DCFFD3B101FF608AF4FEB8F851" box="[151,284,2014,2040]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1896,1920]" captionStartId-1="2.[151,250,1516,1540]" captionTargetBox-0="[158,1430,179,1870]" captionTargetBox-1="[151,1436,181,1492]" captionTargetId-0="figure-27@1.[158,1430,179,1870]" captionTargetId-1="figure-18@2.[151,1436,181,1492]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." captionText-1="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; FG. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="0" pageNumber="394">Figures 12</figureCitation>
</paragraph>
<caption id="DF2BCDD1FFD2B100FF608A42FD93F845" pageId="1" pageNumber="395" startId="1.[151,250,1896,1920]" targetBox="[158,1430,179,1870]" targetPageId="1" targetType="figure">
<paragraph id="8BEB9D59FFD2B100FF608A42FD93F845" blockId="1.[151,1436,1896,2029]" pageId="1" pageNumber="395">
<emphasis id="B920414BFFD2B100FF608A42FCB7F829" bold="true" box="[151,787,1896,1921]" pageId="1" pageNumber="395">
FIGURE 1.
<taxonomicName id="4C54E6DAFFD2B100FEEC8A42FDF2F829" authorityName="Yu &amp; Liu &amp; Bian &amp; Zhang" authorityYear="2025" box="[283,598,1896,1920]" class="Dothideomycetes" family="Leptosphaeriaceae" genus="Rhaphidophora" kingdom="Fungi" order="Pleosporales" pageId="1" pageNumber="395" phylum="Ascomycota" rank="species" species="condoprocera" status="sp. nov.">
<emphasis id="B920414BFFD2B100FEEC8A42FDF2F829" bold="true" box="[283,598,1896,1920]" italics="true" pageId="1" pageNumber="395">Rhaphidophora condoprocera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213FC30FFD2B100FDAA8A43FD0AF828" box="[605,686,1897,1921]" pageId="1" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
Male: A.
</emphasis>
head in frontal view;
<emphasis id="B920414BFFD2B100FC028A43FB88F828" bold="true" box="[1013,1068,1897,1921]" pageId="1" pageNumber="395">B, D.</emphasis>
head and thorax:
<emphasis id="B920414BFFD2B100FB138A43FB5FF828" bold="true" box="[1252,1275,1897,1921]" pageId="1" pageNumber="395">B.</emphasis>
dorsal view,
<emphasis id="B920414BFFD2B100FA738A43FA38F828" bold="true" box="[1412,1436,1897,1921]" pageId="1" pageNumber="395">D.</emphasis>
lateral view;
<emphasis id="B920414BFFD2B100FEE98AA6FE93F80D" bold="true" box="[286,311,1932,1956]" pageId="1" pageNumber="395">C.</emphasis>
head and pronotum in dorsal view;
<emphasis id="B920414BFFD2B100FD598AA7FD7BF80C" bold="true" box="[686,735,1933,1957]" pageId="1" pageNumber="395">EF.</emphasis>
apex of hind femur:
<emphasis id="B920414BFFD2B100FC4B8AA7FC77F80C" bold="true" box="[956,979,1933,1957]" pageId="1" pageNumber="395">E.</emphasis>
external view,
<emphasis id="B920414BFFD2B100FB858AA7FB21F80C" bold="true" box="[1138,1157,1933,1957]" pageId="1" pageNumber="395">F.</emphasis>
internal view;
<emphasis id="B920414BFFD2B100FAD58AA6FA98F80D" bold="true" box="[1314,1340,1932,1956]" pageId="1" pageNumber="395">G.</emphasis>
tarsus of hind leg in lateral view;
<emphasis id="B920414BFFD2B100FE648A9BFE6FF860" bold="true" box="[403,459,1969,1993]" pageId="1" pageNumber="395">HL.</emphasis>
apex of abdomen:
<emphasis id="B920414BFFD2B100FD678A9BFD0EF860" bold="true" box="[656,682,1969,1993]" pageId="1" pageNumber="395">H.</emphasis>
apical view,
<emphasis id="B920414BFFD2B100FCC58A9BFCE6F860" bold="true" box="[818,834,1969,1993]" pageId="1" pageNumber="395">I.</emphasis>
lateral view,
<emphasis id="B920414BFFD2B100FC3A8A9BFC44F860" bold="true" box="[973,992,1969,1993]" pageId="1" pageNumber="395">J.</emphasis>
apical and slightly ventral view,
<emphasis id="B920414BFFD2B100FAC28A9BFAEBF860" bold="true" box="[1333,1359,1969,1993]" pageId="1" pageNumber="395">K.</emphasis>
ventral and slightly lateral view,
<emphasis id="B920414BFFD2B100FE618AFFFE0AF844" bold="true" box="[406,430,2005,2029]" pageId="1" pageNumber="395">L.</emphasis>
ventral view.
</paragraph>
</caption>
<caption id="DF2BCDD1FFD1B103FF6088C6FBB9F9E5" pageId="2" pageNumber="396" startId="2.[151,250,1516,1540]" targetBox="[151,1436,181,1492]" targetPageId="2" targetType="figure">
<paragraph id="8BEB9D59FFD1B103FF6088C6FBB9F9E5" blockId="2.[151,1436,1515,1612]" pageId="2" pageNumber="396">
<emphasis id="B920414BFFD1B103FF6088C6FC87F9AD" bold="true" box="[151,803,1515,1540]" pageId="2" pageNumber="396">
FIGURE 2.
<taxonomicName id="4C54E6DAFFD1B103FEEF88C1FDF6F9AA" authorityName="Yu &amp; Liu &amp; Bian &amp; Zhang" authorityYear="2025" box="[280,594,1515,1539]" class="Dothideomycetes" family="Leptosphaeriaceae" genus="Rhaphidophora" kingdom="Fungi" order="Pleosporales" pageId="2" pageNumber="396" phylum="Ascomycota" rank="species" species="condoprocera" status="sp. nov.">
<emphasis id="B920414BFFD1B103FEEF88C1FDF6F9AA" bold="true" box="[280,594,1515,1539]" italics="true" pageId="2" pageNumber="396">Rhaphidophora condoprocera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213FC30FFD1B103FDAF88C6FD03F9AD" box="[600,679,1516,1540]" pageId="2" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
Female: A.
</emphasis>
head in frontal view;
<emphasis id="B920414BFFD1B103FC0888C6FB91F9AD" bold="true" box="[1023,1077,1516,1540]" pageId="2" pageNumber="396">BC.</emphasis>
head and thorax:
<emphasis id="B920414BFFD1B103FB1F88C6FB5BF9AD" bold="true" box="[1256,1279,1516,1540]" pageId="2" pageNumber="396">B.</emphasis>
dorsal view,
<emphasis id="B920414BFFD1B103FA7388C6FA38F9AD" bold="true" box="[1412,1436,1516,1540]" pageId="2" pageNumber="396">C.</emphasis>
lateral view;
<emphasis id="B920414BFFD1B103FEEA8B3AFE92F981" bold="true" box="[285,310,1552,1576]" pageId="2" pageNumber="396">D.</emphasis>
apices of hind femora in internal view;
<emphasis id="B920414BFFD1B103FD268B3AFD4CF981" bold="true" box="[721,744,1552,1576]" pageId="2" pageNumber="396">E.</emphasis>
tarsus of hind leg in lateral view;
<emphasis id="B920414BFFD1B103FBBF8B3AFBDBF981" bold="true" box="[1096,1151,1552,1576]" pageId="2" pageNumber="396">FG.</emphasis>
apex of abdomen:
<emphasis id="B920414BFFD1B103FAB38B3AFAF3F981" bold="true" box="[1348,1367,1552,1576]" pageId="2" pageNumber="396">F.</emphasis>
dorsal view,
<emphasis id="B920414BFFD1B103FF248B1EFF4AF9E5" bold="true" box="[211,238,1588,1612]" pageId="2" pageNumber="396">G.</emphasis>
lateral view;
<emphasis id="B920414BFFD1B103FE8F8B1EFE36F9E5" bold="true" box="[376,402,1588,1612]" pageId="2" pageNumber="396">H.</emphasis>
seventh abdominal sternite and subgenital plate in ventral view.
</paragraph>
</caption>
<paragraph id="8BEB9D59FFD1B103FF608BB4FF6CF8C5" blockId="2.[151,1437,1694,2009]" pageId="2" pageNumber="396">
<emphasis id="B920414BFFD1B103FF608BB4FED1F911" bold="true" box="[151,373,1694,1720]" pageId="2" pageNumber="396">Description. Male.</emphasis>
Body medium.
<emphasis id="B920414BFFD1B103FDDA8BB4FDCEF911" box="[557,618,1694,1720]" italics="true" pageId="2" pageNumber="396">Head</emphasis>
. Face smooth (
<figureCitation id="136F81DCFFD1B103FCEE8BB4FCD0F911" box="[793,884,1694,1720]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1A</figureCitation>
). Fastigium verticis projecting forward in lateral view (
<figureCitation id="136F81DCFFD1B103FF178BE8FE9EF975" box="[224,314,1730,1756]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1B</figureCitation>
), dorsal surface with a longitudinal median furrow, apices which divide fastigium verticis into 2 distinct rostral tubercles and slightly separated from each other (
<figureCitation id="136F81DCFFD1B103FC9C8BCCFC66F8A9" box="[875,962,1766,1792]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1B</figureCitation>
). Eyes ovoid, protruding forward; median ocellus slightly larger than lateral ocelli (
<figureCitation id="136F81DCFFD1B103FD9C8A20FD62F88D" box="[619,710,1802,1828]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1A</figureCitation>
); lateral ocelli large, nearly occupying all the lateral margins of rostral tubercles (
<figureCitation id="136F81DCFFD1B103FE968A04FE1EF8E1" box="[353,442,1838,1864]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1B</figureCitation>
). Apical segments of maxillary palpi about two times as long as subapical ones (
<figureCitation id="136F81DCFFD1B103FA988A04FF1DF8C5" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD1B102FF308A5CFC80FECC" blockId="2.[151,1437,1694,2009]" lastBlockId="3.[151,1437,151,1329]" lastPageId="3" lastPageNumber="397" pageId="2" pageNumber="396">
<emphasis id="B920414BFFD1B103FF308A5CFE84F839" box="[199,288,1910,1936]" italics="true" pageId="2" pageNumber="396">Thorax.</emphasis>
Anterior margin of pronotum slightly projecting, posterior margin widely rounded, protruding backward (
<figureCitation id="136F81DCFFD1B103FF688AB0FF57F81D" box="[159,243,1946,1972]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1C</figureCitation>
); lateral lobes longer than high, ventral margin undulate (
<figureCitation id="136F81DCFFD1B103FCA88AB0FC12F81D" box="[863,950,1946,1972]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="2" pageNumber="396">Fig. 1D</figureCitation>
). Posterior margin of mesonotum protruding backward, posterior margin of metanotum faintly protruding.
<emphasis id="B920414BFFD1B103FCB88A95FC2FF871" box="[847,907,1983,2008]" italics="true" pageId="2" pageNumber="396">Legs.</emphasis>
Fore coxae swollen with 1 stout spine; femora with 1 internal spine; tibiae ventrally with 1 internal and 12 external spines, apices with 1 pair of spines on ventral surface. Middle femora with 1 pair of apical spines; tibiae dorsally with 2 pairs of spines, ventral surface with 13 pairs of spines, apices with 1 pair of spines on dorsal and ventral surfaces separately. Apices of hind femora with 1 short internal spine (
<figureCitation id="136F81DCFFD0B102FE888C29FE70FEB4" box="[383,468,259,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1E</figureCitation>
); tibiae dorsally with 2123 internal spines, 1925 external spines and 1 pair of small subapical spines, apices with 1 pair of dorsal and 2 pairs of ventral spines; basitarsi with 46 small spines and 1 large apical spine on dorsal surface along the midline (
<figureCitation id="136F81DCFFD0B102FD4A8C61FCB7FECC" box="[701,787,331,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF308C45FCA9FDC8" blockId="3.[151,1437,151,1329]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF308C45FE9DFE20" box="[199,313,367,393]" italics="true" pageId="3" pageNumber="397">Abdomen.</emphasis>
Apex of abdomen curving downwards. Posterior margin of ninth abdominal tergite concave, tenth abdominal tergite narrow. Epiproct prolonged and gradually curving downward, basal half with lateral margins convex and converging, apical area slightly widened, bilobate in dorsal view (
<figureCitation id="136F81DCFFD0B102FBE68C9DFBCDFE78" box="[1041,1129,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1H</figureCitation>
), its ventral surface with 1 depressed process (
<figureCitation id="136F81DCFFD0B102FE838CF1FE6FFE5C" box="[372,459,475,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1L</figureCitation>
); ventral surface of epiproct with 1 pair of stout spines, its apices directing outwards and faintly curved forward (
<figureCitation id="136F81DCFFD0B102FE228CD5FD8BFDB0" box="[469,559,511,537]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1K</figureCitation>
). Paraprocts prolonged into a spine, the apical half incurved. Subgenital plate wider than long, posterior margin almost straight; styli gradually upcurved before the middle (
<figureCitation id="136F81DCFFD0B102FB4B8F09FAACFD94" box="[1212,1288,547,573]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1I</figureCitation>
), inserted on posterolateral areas of the plate (
<figureCitation id="136F81DCFFD0B102FDF38F6DFDFDFDC8" box="[516,601,583,609]" captionStart="FIGURE 1" captionStartId="1.[151,250,1896,1920]" captionTargetBox="[158,1430,179,1870]" captionTargetId="figure-27@1.[158,1430,179,1870]" captionTargetPageId="1" captionText="FIGURE 1. Rhaphidophora condoprocera sp. nov. Male: A. head in frontal view; B, D. head and thorax: B. dorsal view, D. lateral view; C. head and pronotum in dorsal view; EF. apex of hind femur: E. external view, F. internal view; G. tarsus of hind leg in lateral view; HL. apex of abdomen: H. apical view, I. lateral view, J. apical and slightly ventral view, K. ventral and slightly lateral view, L. ventral view." pageId="3" pageNumber="397">Fig. 1L</figureCitation>
), apices obtuse.
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF308F41FCB7FD58" blockId="3.[151,1437,151,1329]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF308F41FE81FD2C" bold="true" box="[199,293,619,645]" pageId="3" pageNumber="397">Female.</emphasis>
Tenth abdominal tergite with posterior margin concave in middle (
<figureCitation id="136F81DCFFD0B102FBDF8F41FBDBFD2C" box="[1064,1151,619,645]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; FG. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2F</figureCitation>
). Epiproct semielliptical, dorsal surface furrowed along the midline, posterior margin obtuse (
<figureCitation id="136F81DCFFD0B102FC358FA5FBB9FD00" box="[962,1053,655,681]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; FG. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2F</figureCitation>
). Cerci slender. Subgenital plate transverse, posterior margin undulating and faintly projecting in middle (
<figureCitation id="136F81DCFFD0B102FC3E8F99FBBBFD64" box="[969,1055,691,717]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; FG. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2H</figureCitation>
). Ovipositor about half as long as hind femora, gradually upcurved, apices acute (
<figureCitation id="136F81DCFFD0B102FD5B8FFDFCA6FD58" box="[684,770,727,753]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; FG. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2G</figureCitation>
).
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF308FD0FBB1FC90" blockId="3.[151,1437,151,1329]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF308FD0FEEBFCBD" bold="true" box="[199,335,762,788]" pageId="3" pageNumber="397">Coloration.</emphasis>
Male body brown, Face and eyes black ocelli pale yellow, disc of abdomen nearly black, apices of femora and apices of tibiae darker. Female abdomen with irregularly pale spots.
</paragraph>
<paragraph id="8BEB9D59FFD0B102FF308E69FE10FC0C" blockId="3.[151,1437,151,1329]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF308E69FE62FCF5" bold="true" box="[199,454,834,861]" pageId="3" pageNumber="397">Measurements (mm).</emphasis>
BL:
<specimenCount id="9D5256D0FFD0B102FDF68E6CFDB2FCF5" box="[513,534,838,860]" count="1" pageId="3" pageNumber="397" type="male"></specimenCount>
17.417.8,
<specimenCount id="9D5256D0FFD0B102FD6C8E69FD14FCF4" box="[667,688,835,861]" count="1" pageId="3" pageNumber="397" type="female"></specimenCount>
15.718.6; PL:
<specimenCount id="9D5256D0FFD0B102FC908E6CFCD8FCF5" box="[871,892,838,860]" count="1" pageId="3" pageNumber="397" type="male"></specimenCount>
5.05.2,
<specimenCount id="9D5256D0FFD0B102FC118E69FC5FFCF4" box="[998,1019,835,861]" count="1" pageId="3" pageNumber="397" type="female"></specimenCount>
5.15.7; FFL:
<specimenCount id="9D5256D0FFD0B102FB528E6CFB1EFCF5" box="[1189,1210,838,860]" count="1" pageId="3" pageNumber="397" type="male"></specimenCount>
6.06.1,
<specimenCount id="9D5256D0FFD0B102FAD38E69FA9DFCF4" box="[1316,1337,835,861]" count="1" pageId="3" pageNumber="397" type="female"></specimenCount>
5.77.7; MFL:
<specimenCount id="9D5256D0FFD0B102FF178E40FF51FC29" box="[224,245,874,896]" count="1" pageId="3" pageNumber="397" type="male"></specimenCount>
5.96.0,
<specimenCount id="9D5256D0FFD0B102FE978E4DFED1FC28" box="[352,373,871,897]" count="1" pageId="3" pageNumber="397" type="female"></specimenCount>
6.06.4; HFL:
<specimenCount id="9D5256D0FFD0B102FDD18E40FD9FFC29" box="[550,571,874,896]" count="1" pageId="3" pageNumber="397" type="male"></specimenCount>
14.615.7,
<specimenCount id="9D5256D0FFD0B102FD368E4DFD72FC28" box="[705,726,871,897]" count="1" pageId="3" pageNumber="397" type="female"></specimenCount>
14.716.0; HTL:
<specimenCount id="9D5256D0FFD0B102FC538E40FC1DFC29" box="[932,953,874,896]" count="1" pageId="3" pageNumber="397" type="male"></specimenCount>
12.813.5,
<specimenCount id="9D5256D0FFD0B102FBB78E4DFBF1FC28" box="[1088,1109,871,897]" count="1" pageId="3" pageNumber="397" type="female"></specimenCount>
12.813.5; HBL:
<specimenCount id="9D5256D0FFD0B102FAD38E40FA9DFC29" box="[1316,1337,874,896]" count="1" pageId="3" pageNumber="397" type="male"></specimenCount>
2.82.9,
<specimenCount id="9D5256D0FFD0B102FF608EA1FF08FC0C" box="[151,172,907,933]" count="1" pageId="3" pageNumber="397" type="female"></specimenCount>
2.43.0; OvL: 6.59.2.
</paragraph>
</subSubSection>
<subSubSection id="C34ECED2FFD0B102FF308E84FCA1FB9C" pageId="3" pageNumber="397" type="materials_examined">
<paragraph id="8BEB9D59FFD0B102FF308E84FCA1FB9C" blockId="3.[151,1437,151,1329]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF308E84FE14FC61" bold="true" box="[199,432,942,968]" pageId="3" pageNumber="397">Material examined.</emphasis>
<materialsCitation id="3B3C9704FFD0B102FE418E85FF61FC44" collectingDate="2024-08-17" location="Lushui" municipality="Pianma" pageId="3" pageNumber="397" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54EF23FBFFD0B102FE418E85FD87FC60" box="[438,547,943,969]" pageId="3" pageNumber="397" type="holotype">Holotype</typeStatus>
: male,
<collectingMunicipality id="6B8F0723FFD0B102FD988E85FD62FC60" box="[623,710,943,969]" pageId="3" pageNumber="397">Pianma</collectingMunicipality>
,
<location id="8E8BCB82FFD0B102FD268E85FC86FC60" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD3B102FF608ABFFB86FA98:8E8BCB82FFD0B102FD268E85FC86FC60" box="[721,802,943,969]" municipality="Pianma" name="Lushui" pageId="3" pageNumber="397" stateProvince="Yunnan">Lushui</location>
,
<collectingRegion id="499053BBFFD0B102FCDC8E85FC20FC60" box="[811,900,943,969]" country="China" name="Yunnan" pageId="3" pageNumber="397">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD0B102FC7B8E85FBE3FC60" box="[908,1095,943,969]" pageId="3" pageNumber="397" value="2024-08-17">
<collectingDate id="EFAE4271FFD0B102FC7B8E85FBE3FC60" box="[908,1095,943,969]" pageId="3" pageNumber="397" value="2024-08-17">August 17, 2024</collectingDate>
</date>
, coll. by Ting Luo and Yanting Qin
</materialsCitation>
.
<materialsCitation id="3B3C9704FFD0B102FF248EF9FE87FBB8" collectingDate="2024-08-17" county="Pianma" location="Ting Luo" municipality="Lushui" pageId="3" pageNumber="397" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54EF23FBFFD0B102FF248EF9FEE0FC44" box="[211,324,979,1005]" pageId="3" pageNumber="397" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5256D0FFD0B102FEA58EF9FE05FC44" box="[338,417,979,1005]" count="1" pageId="3" pageNumber="397" type="male">1 male</specimenCount>
and
<specimenCount id="9D5256D0FFD0B102FE288EF9FDE3FC44" box="[479,583,979,1005]" count="1" pageId="3" pageNumber="397" type="female">1 female</specimenCount>
, male,
<collectingCounty id="628AE5D5FFD0B102FD698EF9FD51FC44" box="[670,757,979,1005]" pageId="3" pageNumber="397">Pianma</collectingCounty>
,
<collectingMunicipality id="6B8F0723FFD0B102FCF38EF9FCF1FC44" box="[772,853,979,1005]" pageId="3" pageNumber="397">Lushui</collectingMunicipality>
,
<collectingRegion id="499053BBFFD0B102FC958EF9FC1FFC44" box="[866,955,979,1005]" country="China" name="Yunnan" pageId="3" pageNumber="397">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD0B102FC3F8EF9FB29FC44" box="[968,1165,979,1005]" pageId="3" pageNumber="397" value="2024-08-17">
<collectingDate id="EFAE4271FFD0B102FC3F8EF9FB29FC44" box="[968,1165,979,1005]" pageId="3" pageNumber="397" value="2024-08-17">August 17, 2024</collectingDate>
</date>
, coll. by
<location id="8E8BCB82FFD0B102FB0A8EF9FACDFC44" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD3B102FF608ABFFB86FA98:8E8BCB82FFD0B102FB0A8EF9FACDFC44" box="[1277,1385,979,1005]" county="Pianma" municipality="Lushui" name="Ting Luo" pageId="3" pageNumber="397" stateProvince="Yunnan">Ting Luo</location>
and
<location id="8E8BCB82FFD0B102FF608EDDFE87FBB8" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD3B102FF608ABFFB86FA98:8E8BCB82FFD0B102FF608EDDFE87FBB8" box="[151,291,1015,1041]" county="Pianma" municipality="Lushui" name="Yanting Qin" pageId="3" pageNumber="397" stateProvince="Yunnan">Yanting Qin</location>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD0B102FEC78EDDFB7DFBB8" box="[304,1241,1015,1041]" county="Pianma" location="August" municipality="Lushui" pageId="3" pageNumber="397" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D5256D0FFD0B102FEC78EDDFE04FBB8" box="[304,416,1015,1041]" count="3" pageId="3" pageNumber="397" type="female">3 females</specimenCount>
,
<collectingCounty id="628AE5D5FFD0B102FE5B8EDDFDA7FBB8" box="[428,515,1015,1041]" pageId="3" pageNumber="397">Pianma</collectingCounty>
,
<collectingMunicipality id="6B8F0723FFD0B102FDF88EDDFDC4FBB8" box="[527,608,1015,1041]" pageId="3" pageNumber="397">Lushui</collectingMunicipality>
,
<collectingRegion id="499053BBFFD0B102FD9D8EDDFD67FBB8" box="[618,707,1015,1041]" country="China" name="Yunnan" pageId="3" pageNumber="397">Yunnan</collectingRegion>
,
<location id="8E8BCB82FFD0B102FD3A8EDDFCBAFBB8" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD3B102FF608ABFFB86FA98:8E8BCB82FFD0B102FD3A8EDDFCBAFBB8" box="[717,798,1015,1041]" county="Pianma" municipality="Lushui" name="August" pageId="3" pageNumber="397" stateProvince="Yunnan">August</location>
17, coll. by
<location id="8E8BCB82FFD0B102FC5A8EDDFBB2FBB8" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD3B102FF608ABFFB86FA98:8E8BCB82FFD0B102FC5A8EDDFBB2FBB8" box="[941,1046,1015,1041]" county="Pianma" municipality="Lushui" name="Ting Luo" pageId="3" pageNumber="397" stateProvince="Yunnan">Ting Luo</location>
and
<location id="8E8BCB82FFD0B102FBBA8EDDFB7DFBB8" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD3B102FF608ABFFB86FA98:8E8BCB82FFD0B102FBBA8EDDFB7DFBB8" box="[1101,1241,1015,1041]" county="Pianma" municipality="Lushui" name="Yanting Qin" pageId="3" pageNumber="397" stateProvince="Yunnan">Yanting Qin</location>
</materialsCitation>
;
<materialsCitation id="3B3C9704FFD0B102FB118EDDFCA4FB9C" collectingDate="2024-08-21" county="Pianma" location="Mingyu Liu" municipality="Lushui" pageId="3" pageNumber="397" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D5256D0FFD0B102FB118EDDFA91FBB8" box="[1254,1333,1015,1041]" count="1" pageId="3" pageNumber="397" type="male">1 male</specimenCount>
,
<collectingCounty id="628AE5D5FFD0B102FAB68EDDFA3CFBB8" box="[1345,1432,1015,1041]" pageId="3" pageNumber="397">Pianma</collectingCounty>
,
<collectingMunicipality id="6B8F0723FFD0B102FF608931FF4CFB9C" box="[151,232,1051,1077]" pageId="3" pageNumber="397">Lushui</collectingMunicipality>
,
<collectingRegion id="499053BBFFD0B102FF058931FEEFFB9C" box="[242,331,1051,1077]" country="China" name="Yunnan" pageId="3" pageNumber="397">Yunnan</collectingRegion>
,
<date id="FFEABB99FFD0B102FEA38931FDB5FB9C" box="[340,529,1051,1077]" pageId="3" pageNumber="397" value="2024-08-21">
<collectingDate id="EFAE4271FFD0B102FEA38931FDB5FB9C" box="[340,529,1051,1077]" pageId="3" pageNumber="397" value="2024-08-21">August 21, 2024</collectingDate>
</date>
, coll. by
<location id="8E8BCB82FFD0B102FD808931FCA4FB9C" LSID="urn:lsid:plazi:treatment:03FD2C4FFFD3B102FF608ABFFB86FA98:8E8BCB82FFD0B102FD808931FCA4FB9C" box="[631,768,1051,1077]" county="Pianma" municipality="Lushui" name="Mingyu Liu" pageId="3" pageNumber="397" stateProvince="Yunnan">Mingyu Liu</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34ECED2FFD0B102FF398914FD96FBF0" box="[206,562,1086,1113]" pageId="3" pageNumber="397" type="distribution">
<paragraph id="8BEB9D59FFD0B102FF398914FD96FBF0" blockId="3.[151,1437,151,1329]" box="[206,562,1086,1113]" pageId="3" pageNumber="397">
<emphasis id="B920414BFFD0B102FF398914FECCFBF1" bold="true" box="[206,360,1086,1112]" pageId="3" pageNumber="397">Distribution.</emphasis>
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(Lushui).
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<emphasis id="B920414BFFD0B102FF308948FEE8FBD5" bold="true" box="[199,332,1122,1148]" pageId="3" pageNumber="397">Discussion.</emphasis>
The new species differs from other species of
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<emphasis id="B920414BFFD0B102FC628949FBE1FBD4" box="[917,1093,1123,1149]" italics="true" pageId="3" pageNumber="397">Rhaphidophora</emphasis>
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by male epiproct prolonged downwards and backwards apical area with three swellings: two dorsad and one ventrad, apical margin of each swelling undulant (
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), ventral surface of epiproct with 1 pair of stout and widely separated spines; posterior margin of female subgenital plate faintly projecting in middle (
<figureCitation id="136F81DCFFD0B102FCAE89E5FC0BFB40" box="[857,943,1231,1257]" captionStart="FIGURE 2" captionStartId="2.[151,250,1516,1540]" captionTargetBox="[151,1436,181,1492]" captionTargetId="figure-18@2.[151,1436,181,1492]" captionTargetPageId="2" captionText="FIGURE 2. Rhaphidophora condoprocera sp. nov. Female: A. head in frontal view; BC. head and thorax: B. dorsal view, C. lateral view; D. apices of hind femora in internal view; E. tarsus of hind leg in lateral view; FG. apex of abdomen: F. dorsal view, G. lateral view; H. seventh abdominal sternite and subgenital plate in ventral view." pageId="3" pageNumber="397">Fig. 2H</figureCitation>
).
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<emphasis id="B920414BFFD0B102FF3089D9FEEAFAA4" bold="true" box="[199,334,1267,1293]" pageId="3" pageNumber="397">Etymology.</emphasis>
The name of the new species derives from the Latin words “
<emphasis id="B920414BFFD0B102FBFD89D9FBE4FAA4" box="[1034,1088,1267,1293]" italics="true" pageId="3" pageNumber="397">cond</emphasis>
” and “
<emphasis id="B920414BFFD0B102FB6489D9FB79FAA5" box="[1171,1245,1267,1292]" italics="true" pageId="3" pageNumber="397">procer</emphasis>
” referring to the male epiproct with 1 subcylindrical process on ventral surface of the upper plate.
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Diagnosis. (modified from
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. Fronto-orbital plates not enlarged; eyes densely covered with short hairs; ocellar setae present; fronto-orbital plate narrower than frontal vitta; proboscis short, stout and bulbous, dusted; facial ridges with few setulae near vibrissa; prosternum bare; antenna dark brown; arista nearly bare; palpus grey, slightly dilated apically, flattened, spoon-shaped; dorsocentrals 2:3, the anterior presutural one very weak and short; anepimeron bare; presutural intra-alar seta present; katepisternals 0:1. Fore tibia with one posteroventral seta. Hind tibia with a weak posterodorsal seta, one anterodorsal and one anterovental. Wings brownish, halter and calypter yellow.
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Notes. Described from male and female from
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<emphasis id="B3B98961FF81207D6BD540A3FE770EB5" bold="true" box="[151,436,474,501]" italics="true" pageId="19" pageNumber="279">Xenomyia stuckenbergi</emphasis>
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Diagnosis. (modified from
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) Length:
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. Fronto-orbital plate narrower than frontal vitta; profile of head at antennal base much longer than at mouth margin, forming an acute angle. Eyes densely covered with short hairs; ocellar setae present; fronto-orbital plate narrower than frontal vitta; proboscis short, stout and bulbous, dusted; palpus brown; antenna brown; arista bare; dorsocentrals 2:3; katepisternals 1:1; prosternum bare; presutural intra-alar seta present. Fore tibia with 12 posterior seta; the last two tarsomeres dorsally dilated; hind tibia with only one short anterodorsal seta at middle; claws not enlarged. Calypter brownish; halter yellow.
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Notes. Known from
<typeStatus id="5E76EBD1FF81207D6AEF4383FDCC0C55" box="[429,527,763,789]" pageId="19" pageNumber="279" type="holotype">holotype</typeStatus>
male from
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,
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. (
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.
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<taxonomicName id="46CD2EF0FF82207E6BD547DBFD9309FC" ID-CoL="7FVV9" authority="Emden, 1951" authorityName="Emden" authorityYear="1951" box="[151,592,1698,1725]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="276" phylum="Arthropoda" rank="species" species="pseudolispe">
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<emphasis id="B3B98961FF82207E6BD547DBFE6609FD" bold="true" box="[151,421,1698,1725]" italics="true" pageId="16" pageNumber="276">Xenomyia pseudolispe</emphasis>
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Diagnosis. (modified from
<bibRefCitation id="E55C2882FF82207E6A854793FD9A0845" author="Emden, F. I. van" box="[455,601,1770,1797]" pageId="16" pageNumber="276" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
). Length: body
<quantity id="4635F896FF82207E684B4793FCA20845" box="[777,865,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="16" pageNumber="276" unit="mm" value="4.9">4.9 mm</quantity>
; wing
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. Ocellar setae present; fronto-orbital plate narrower than frontal vitta; proboscis stout and bulbous, dusted; facial ridges with few setulae near vibrissa; claws long and strong; presutural intra-alar seta absent; anepimeron setulose (
<figureCitation id="19F649F6FF82207E6F55464BFBAF080D" box="[1047,1132,1842,1869]" captionStart="FIGURES 2126" captionStartId="10.[151,264,1378,1402]" captionTargetBox="[154,1432,184,1351]" captionTargetId="figure-18@10.[151,1435,181,1354]" captionTargetPageId="10" captionText="FIGURES 2126. 21. Xenomyia calyptrata Emden, wing and calypters of male; a: upper calypter; b: lower calypter (modified from Fig. 53 of Emden 1951); 22. Xenomyia edwardsi Emden, wing of male (modified from Fig. 4, plate 1 of Emden 1951); 23. Xenomyia hirtibasis (Bigot), wing of male; 24. Xenomyia osculata Pont &amp; Werner, wing of male (modified from Fig. 5 of Pont &amp; Werner 2003); 25. Xenomyia pseudolispe Emden, thorax of male, lateral view (modified from Fig. 47 of Emden 1951); 26. Xenomyia bispina Emden, abdomen of male, lateral view (modified from Fig. 7, plate X of Emden 1951). Scale bars: 1 mm. Figs 21, 22 and 24 not to scale." pageId="16" pageNumber="276">Fig. 25</figureCitation>
, modified from
<figureCitation id="19F649F6FF82207E6E6B464BFAB8080D" box="[1321,1403,1843,1869]" captionStart="FIGURES 4651" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[250,1338,183,1913]" captionTargetId="figure-33@17.[244,1344,181,1919]" captionTargetPageId="17" captionText="FIGURES 4651. 46. Xenomyia hirtibasis (Bigot), male, lateral habitus; 47. Xenomyia hirtibasis (Bigot), male, dorsal habitus; 48. Xenomyia hirtibasis (Bigot), male, head, frontal view; 49. Xenomyia hirtibasis (Bigot), female, lateral habitus; 50. Xenomyia hirtibasis (Bigot), female, dorsal habitus; 51. Xenomyia hirtibasis (Bigot), female, head, frontal view. Scale bars: 1 mm." pageId="16" pageNumber="276">Fig. 47</figureCitation>
of
<bibRefCitation id="E55C2882FF82207E6BD5462FFEEA0831" author="Emden, F. I. van" box="[151,297,1878,1905]" pageId="16" pageNumber="276" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
); katepisternals 0:1; postpedicel sharply angular at dorsal apex, fore tibia with a posterior seta beyond middle; hind tibia with a fine median posterodorsal seta; hind metatarsus modified, very slender in basal two-thirds and strongly thickened and with some setae on apical third (
<figureCitation id="19F649F6FF82207E687A46E7FC4808F9" box="[824,907,1951,1977]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1909,1933]" captionTargetBox="[216,1371,184,1882]" captionTargetId="figure-18@8.[213,1374,181,1884]" captionTargetPageId="8" captionText="FIGURES 1620. 16. Xenomyia bispina Emden, hind leg of male (modified from Fig. 50 of Emden 1951); 17. Xenomyia hirtibasis Emden, hind leg of male; 18. Xenomyia osculata Pont &amp; Werner, fore femur of male (modified from Fig. 4 of Pont &amp; Werner 2003); 19. Xenomyia perplexa Emden, hind leg of male (modified from Fig. 49 of Emden 1951); 20. Xenomyia pseudolispe Emden, hind leg of male (modified from Fig. 48 of Emden 1951). Scale bars: 1 mm. Fig. 18, not to scale." pageId="16" pageNumber="276">Fig. 20</figureCitation>
, modified from
<figureCitation id="19F649F6FF82207E6F0246E7FB4C08F9" box="[1088,1167,1951,1977]" captionStart="FIGURES 4651" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[250,1338,183,1913]" captionTargetId="figure-33@17.[244,1344,181,1919]" captionTargetPageId="17" captionText="FIGURES 4651. 46. Xenomyia hirtibasis (Bigot), male, lateral habitus; 47. Xenomyia hirtibasis (Bigot), male, dorsal habitus; 48. Xenomyia hirtibasis (Bigot), male, head, frontal view; 49. Xenomyia hirtibasis (Bigot), female, lateral habitus; 50. Xenomyia hirtibasis (Bigot), female, dorsal habitus; 51. Xenomyia hirtibasis (Bigot), female, head, frontal view. Scale bars: 1 mm." pageId="16" pageNumber="276">Fig. 48</figureCitation>
of
<bibRefCitation id="E55C2882FF82207E6FF646E7FA8408F9" author="Emden, F. I. van" box="[1204,1351,1950,1977]" pageId="16" pageNumber="276" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
).
</paragraph>
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<paragraph id="81725573FF83207F6BD546EFFA9B08B7" blockId="17.[151,1436,1943,2040]" pageId="17" pageNumber="277">
<emphasis id="B3B98961FF83207F6BD546EFFE9008EF" bold="true" box="[151,339,1943,1967]" pageId="17" pageNumber="277">FIGURES 4651.</emphasis>
46.
<taxonomicName id="46CD2EF0FF83207F6A3D46E0FD5808EF" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[383,667,1943,1967]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="277" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF83207F6A3D46E0FD8908EF" box="[383,586,1943,1967]" italics="true" pageId="17" pageNumber="277">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, male, lateral habitus; 47.
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<emphasis id="B3B98961FF83207F68E746E0FBB308EF" box="[933,1136,1943,1967]" italics="true" pageId="17" pageNumber="277">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, male, dorsal habitus; 48.
<taxonomicName id="46CD2EF0FF83207F6BFF46C4FE1B0893" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[189,472,1979,2003]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="277" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF83207F6BFF46C4FE440893" box="[189,391,1979,2003]" italics="true" pageId="17" pageNumber="277">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, male, head, frontal view; 49.
<taxonomicName id="46CD2EF0FF83207F684646C4FBDC0893" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[772,1055,1979,2003]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="277" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF83207F684646C4FC0D0893" box="[772,974,1979,2003]" italics="true" pageId="17" pageNumber="277">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, female, lateral habitus; 50.
<taxonomicName id="46CD2EF0FF83207F6E7546C4FE8B08B7" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="277" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF83207F6E7546C4FF3408B7" italics="true" pageId="17" pageNumber="277">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, female, dorsal habitus; 51.
<taxonomicName id="46CD2EF0FF83207F69274698FC4108B7" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[613,898,2015,2039]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="277" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF83207F69274698FCF208B7" box="[613,817,2015,2039]" italics="true" pageId="17" pageNumber="277">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, female, head, frontal view. Scale bars: 1 mm.
</paragraph>
</caption>
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<emphasis id="B3B98961FF80207C6BD5448DFEE3094D" bold="true" box="[151,288,1525,1549]" pageId="18" pageNumber="278">FIGURE 52.</emphasis>
Map with the known distribution of the 16 species of
<taxonomicName id="46CD2EF0FF80207C6807448EFBC6094D" authority="Malloch" authorityName="Malloch" authorityYear="1921" box="[837,1029,1525,1549]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B3B98961FF80207C6807448EFC69094D" box="[837,938,1526,1549]" italics="true" pageId="18" pageNumber="278">Xenomyia</emphasis>
Malloch
</taxonomicName>
(
<taxonomicName id="46CD2EF0FF80207C6F51448DFBA0094D" box="[1043,1123,1525,1549]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="18" pageNumber="278" phylum="Tracheophyta" rank="order">Diptera</taxonomicName>
,
<taxonomicName id="46CD2EF0FF80207C6F2F448DFB17094D" box="[1133,1236,1525,1549]" class="Insecta" family="Muscidae" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="278" phylum="Arthropoda" rank="family">Muscidae</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="81725573FF80207C6B854721FE400867" blockId="18.[151,1436,1625,1831]" pageId="18" pageNumber="278">
Note. Described from male from
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, Aberdares, Thompsons Falls.
<typeStatus id="5E76EBD1FF80207C68B04721FB990933" box="[1010,1114,1625,1651]" pageId="18" pageNumber="278" type="holotype">Holotype</typeStatus>
deposited at NHMUK. This is the only
<taxonomicName id="46CD2EF0FF80207C6A514706FE1F09D7" box="[275,476,1661,1687]" pageId="18" pageNumber="278">
<emphasis id="B3B98961FF80207C6A514706FE4009D7" box="[275,387,1662,1687]" italics="true" pageId="18" pageNumber="278">Xenomyia</emphasis>
species
</taxonomicName>
with setulae on anepimeron, but in a different position as in
<taxonomicName id="46CD2EF0FF80207C6FC44706FB0009D7" authorityName="Latreille" authorityYear="1797" box="[1158,1219,1662,1687]" class="Insecta" family="Muscidae" genus="Lispe" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B3B98961FF80207C6FC44706FB0009D7" box="[1158,1219,1662,1687]" italics="true" pageId="18" pageNumber="278">Lispe</emphasis>
</taxonomicName>
(the former having the setulae in front of the anterior end of the infrasquamal ridge, and not like the latter where the setulae are in the middle of anepimeron between lower anterior end of infrasquamal ridge and the suture with katepisternum. Also, the parafacialia of this species is bare.
<bibRefCitation id="E55C2882FF80207C69194791FCF80843" author="Emden, F. I. van" box="[603,827,1769,1796]" pageId="18" pageNumber="278" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden (1951: 496)</bibRefCitation>
made some considerations about the position of this species in
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<emphasis id="B3B98961FF80207C6A4E4676FEBF0867" box="[268,380,1806,1831]" italics="true" pageId="18" pageNumber="278">Xenomyia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,115 @@
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<mods:title id="D8538410D4296C9FAC7D3F0B63AAEE76">The puzzling genus Xenomyia Malloch (Diptera, Muscidae): taxonomic notes, key to species and redescription of Xenomyia hirtibasis (Bigot)</mods:title>
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<taxonomicName id="46CD2EF0FF82207E6BD5405FFDE10E00" ID-CoL="7G7V8" authority="Emden, 1948" authorityName="Emden" authorityYear="1948" box="[151,546,294,321]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="276" phylum="Arthropoda" rank="species" species="palpalis">
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<emphasis id="B3B98961FF82207E6BD5405FFEBB0E01" bold="true" box="[151,376,294,321]" italics="true" pageId="16" pageNumber="276">Xenomyia palpalis</emphasis>
<bibRefCitation id="E55C2882FF82207E6A3D405FFDE10E00" author="Emden, F. I. van" box="[383,546,294,321]" pageId="16" pageNumber="276" pagination="460 - 484" refId="ref11097" refString="Emden, F. I. van (1948) Entomological Expedition to Abyssinia, 1926 - 27. Diptera Cyclorrhapha: Muscidae, II. Annals and Magazine of Natural History, Series 11, 14, 460 - 484, 6 figs. https: // doi. org / 10.1080 / 03745481.1943.9727997" type="journal article" year="1948">Emden, 1948</bibRefCitation>
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Diagnosis. (modified from the
<bibRefCitation id="E55C2882FF82207E6AB34017FD400EC9" author="Emden, F. I. van" box="[497,643,367,393]" pageId="16" pageNumber="276" pagination="460 - 484" refId="ref11097" refString="Emden, F. I. van (1948) Entomological Expedition to Abyssinia, 1926 - 27. Diptera Cyclorrhapha: Muscidae, II. Annals and Magazine of Natural History, Series 11, 14, 460 - 484, 6 figs. https: // doi. org / 10.1080 / 03745481.1943.9727997" type="journal article" year="1948">Emden 1948</bibRefCitation>
). Female. (
<figureCitation id="19F649F6FF82207E69BD4017FC8C0EC9" box="[767,847,366,393]" captionStart="FIGURES 1015" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[199,1388,185,1847]" captionTargetId="figure-33@7.[196,1391,181,1850]" captionTargetPageId="7" captionText="FIGURES 1015. 10. Xenomyia hirtibasis (Bigot), head of male, lateral view; 11. Xenomyia hirtibasis (Bigot), head of female, lateral view; 12. Xenomyia osculata Pont &amp; Werner, head of male, frontal view (modified from Fig. 1 of Pont &amp; Werner 2003); 13. Xenomyia osculata Pont &amp; Werner, head of female, frontal view (modified from Fig. 2 of Pont &amp; Werner 2003); 14. Xenomyia oxycera Emden, head of male, lateral view (modified from Fig. 55 of Emden 1951); 15. Xenomyia palpalis Emden, head and thorax of female, lateral view (modified from Fig. 4 of Emden 1948). Scale bars: 1 mm. Figs 12, 13 and 15 not to scale." pageId="16" pageNumber="276">Fig. 15</figureCitation>
modified from
<figureCitation id="19F649F6FF82207E68BD4017FB830EC9" box="[1023,1088,367,393]" captionStart="FIGURES 49" captionStartId="3.[151,264,1822,1846]" captionTargetBox="[221,1369,185,1795]" captionTargetId="figure-33@3.[218,1369,181,1798]" captionTargetPageId="3" captionText="FIGURES 49. 4. Xenomyia acuticeps Emden, head of male, lateral view (modified from Fig. 54 of Emden 1951); 5. Xenomyia atra Malloch, head of female, lateral view (modified from Fig. 53 of Emden 1951); 6. Xenomyia atra Malloch, head of male, lateral view (modified from Fig. 52 of Emden 1951, after Jeanneliotis notabilis Séguy); 7. Xenomyia calyptrata Emden, head and thorax of male, lateral view (modified from Fig. 56 of Emden 1951); 8. Xenomyia edwardsi Emden, head of male, lateral view (modified from Fig. 57 of Emden 1951); 9. Xenomyia edwardsi Emden, head of female, lateral view (modified from Fig. 58 of Emden 1951). Scale bars: 1 mm. Fig. 6 not to scale." pageId="16" pageNumber="276">Fig. 4</figureCitation>
of
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). Body length:
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; wing length:
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. General ground-color fuscous-brown. Ocellar setae present, fine and short; antennal insertion produced; arista damaged; eye a little oblique with dense and short velvety-like hairs; fronto-orbital plate narrower than frontal vitta and without setae, except for 12 setulae near lunula; proboscis stout and bulbous, dusted; palpus very large; facial ridges with few setulae near vibrissa. Mesonotum without acrostichal setae, dorsocentrals 1:4, only the last one strong; presutural intra-alar seta absent; anepimeron bare; katepisternals 0:1. Legs very long and slender; fore tibia without submedian setae; hind tibia with 1 anterodorsal submedian seta; claws long and strong. Wing brownish along the veins; calypter brown; halter yellow.
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Notes. Known only from
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,
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,
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, collected in house (
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). Figure of head and thorax in lateral view in
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. Male unknown
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View file

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Diagnosis. (modified from
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. Eyes bare. Fronto-orbital plates enlarged, completely suppressing frontal vitta and covered with short dense setulae; ocellar setae absent in male and present in female; proboscis slender, elongate and glossy dark brown; prosternum bare; anepimeron bare; posthumeral seta present. Dorsocentrals 2:3, katepisternals 1:2. Fore tibia without a posterior seta; hind tibia with only one median anterodorsal seta. Claws not elongated. Calypter yellow with brownish margins; halter yellow.
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Notes. Described from male and female from
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<bibRefCitation id="E55C2882FF82207E6ACA45B7FDEF0BA8" author="Emden, F. I. van" box="[392,556,1230,1257]" pageId="16" pageNumber="276" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden, 1951</bibRefCitation>
</emphasis>
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</paragraph>
</subSubSection>
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Diagnosis. (modified from
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). Length: 7.0 mm. Ocellar setae present; fronto-orbital plate narrower than frontal vitta; proboscis short, stout and bulbous, dusted; palpus narrowly dilated apically, flattened, somewhat spoon-shaped; facial ridges with few setulae near vibrissa; several rows of occipital hairs; anepimeron bare; presutural intra-alar seta present in male and absent in female; dorsocentrals 1:3; katepisternals 0:1; coxae with slender setae, more numerous on mid coxa; claws very long and strong almost same size as the length of pedicel plus postpedicel; puvilli small; fore tibia with a posterior seta; apex of hind tibia in male with long posteroventral fine setae and very long anteroventral setae; hind tarsi compressed and very short (
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, modified from
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of
<bibRefCitation id="E55C2882FF82207E6F9B4497FAA80949" author="Emden, F. I. van" box="[1241,1387,1518,1545]" pageId="16" pageNumber="276" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
).
</paragraph>
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Note. Described from male and female from
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,
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,
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deposited at
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View file

@ -0,0 +1,112 @@
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<taxonomicName id="46CD2EF0FF9420686BD542ABFDD80CAC" ID-CoL="7FW77" authority="Emden, 1951" authorityName="Emden" authorityYear="1951" box="[151,539,978,1005]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="bispina">
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<emphasis id="B3B98961FF9420686BD542ABFEB20CAD" bold="true" box="[151,369,978,1005]" italics="true" pageId="6" pageNumber="266">Xenomyia bispina</emphasis>
<bibRefCitation id="E55C2882FF9420686A3A42ABFDD80CAC" author="Emden, F. I. van" box="[376,539,978,1005]" pageId="6" pageNumber="266" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden, 1951</bibRefCitation>
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Diagnosis. (modified from
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). Length:
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. Ocellar setae present; fronto-orbital plate narrower than frontal vitta; proboscis stout and bulbous, dusted; facial ridges with few setulae near vibrissa; profile of head at antennal base not longer than at mouth margin, forming an obtuse angle. Several rows of occipital hairs. Eyes oblique. Anepimeron bare. Hind tibia without anterodorsal seta. Claws moderately long, not longer than antenna. Dorsocentrals 2:3. Katepisternals 1:1. Presutural intra-alar absent or small. Fore tibia with posterior seta. Hind femur with two short spines in basal two-fifth (
<figureCitation id="19F649F6FF942068691645B7FD600BA9" box="[596,675,1231,1257]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1909,1933]" captionTargetBox="[216,1371,184,1882]" captionTargetId="figure-18@8.[213,1374,181,1884]" captionTargetPageId="8" captionText="FIGURES 1620. 16. Xenomyia bispina Emden, hind leg of male (modified from Fig. 50 of Emden 1951); 17. Xenomyia hirtibasis Emden, hind leg of male; 18. Xenomyia osculata Pont &amp; Werner, fore femur of male (modified from Fig. 4 of Pont &amp; Werner 2003); 19. Xenomyia perplexa Emden, hind leg of male (modified from Fig. 49 of Emden 1951); 20. Xenomyia pseudolispe Emden, hind leg of male (modified from Fig. 48 of Emden 1951). Scale bars: 1 mm. Fig. 18, not to scale." pageId="6" pageNumber="266">Fig. 16</figureCitation>
(modified from
<figureCitation id="19F649F6FF942068681A45B7FC650BA9" box="[856,934,1230,1257]" captionStart="FIGURES 4651" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[250,1338,183,1913]" captionTargetId="figure-33@17.[244,1344,181,1919]" captionTargetPageId="17" captionText="FIGURES 4651. 46. Xenomyia hirtibasis (Bigot), male, lateral habitus; 47. Xenomyia hirtibasis (Bigot), male, dorsal habitus; 48. Xenomyia hirtibasis (Bigot), male, head, frontal view; 49. Xenomyia hirtibasis (Bigot), female, lateral habitus; 50. Xenomyia hirtibasis (Bigot), female, dorsal habitus; 51. Xenomyia hirtibasis (Bigot), female, head, frontal view. Scale bars: 1 mm." pageId="6" pageNumber="266">Fig. 50</figureCitation>
of
<bibRefCitation id="E55C2882FF942068688B45B7FB990BA9" author="Emden, F. I. van" box="[969,1114,1230,1257]" pageId="6" pageNumber="266" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
). Hind tibia anterodorsal seta absent in male and present in female.Abdomen subcylindrical with apex truncated in lateral view (
<figureCitation id="19F649F6FF9420686F9B458BFAEB0A4D" box="[1241,1320,1267,1293]" captionStart="FIGURES 2126" captionStartId="10.[151,264,1378,1402]" captionTargetBox="[154,1432,184,1351]" captionTargetId="figure-18@10.[151,1435,181,1354]" captionTargetPageId="10" captionText="FIGURES 2126. 21. Xenomyia calyptrata Emden, wing and calypters of male; a: upper calypter; b: lower calypter (modified from Fig. 53 of Emden 1951); 22. Xenomyia edwardsi Emden, wing of male (modified from Fig. 4, plate 1 of Emden 1951); 23. Xenomyia hirtibasis (Bigot), wing of male; 24. Xenomyia osculata Pont &amp; Werner, wing of male (modified from Fig. 5 of Pont &amp; Werner 2003); 25. Xenomyia pseudolispe Emden, thorax of male, lateral view (modified from Fig. 47 of Emden 1951); 26. Xenomyia bispina Emden, abdomen of male, lateral view (modified from Fig. 7, plate X of Emden 1951). Scale bars: 1 mm. Figs 21, 22 and 24 not to scale." pageId="6" pageNumber="266">Fig. 26</figureCitation>
(modified from
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of
<bibRefCitation id="E55C2882FF9420686A0B446FFE1F0A71" author="Emden, F. I. van" box="[329,476,1302,1329]" pageId="6" pageNumber="266" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
).
</paragraph>
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Note. Described from male and female from
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, Namwamba Valley at
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(
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Diagnosis. (modified from
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. Ocellar setae absent; fronto-orbital plate much broader than frontal vitta, densely covered by short setulae; postpedicel round at apex; proboscis slender and glossy black; antenna and palpus black, long and slender; prosternum bare; superior frontal seta absent; acrostichals present; dorsocentrals 2:3; katepisternals 1:2; presutural intra-alar seta present; anterior intralar seta strong, posterior weak. Anepimeron bare. Fore tibia without median seta. Hind tibia with an anterodorsal and an anteroventral setae, without posterodorsal. Claws moderately long, not longer than antenna.
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Note. Described from
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<emphasis id="B3B98961FF9420686BD544B3FE510AA5" bold="true" box="[151,402,1482,1509]" italics="true" pageId="6" pageNumber="266">Xenomyia calyptrata</emphasis>
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Diagnosis. (modified from
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). Length: 5.0
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. Ocellar absent; fronto-orbital plate much broader than frontal vitta, densely covered by short setulae; postpedicel round at apex; proboscis slender and glossy black; antenna and palpus black, long and slender; arista very short (
<figureCitation id="19F649F6FF94206868324723FC7B0935" box="[880,952,1627,1653]" captionStart="FIGURES 49" captionStartId="3.[151,264,1822,1846]" captionTargetBox="[221,1369,185,1795]" captionTargetId="figure-33@3.[218,1369,181,1798]" captionTargetPageId="3" captionText="FIGURES 49. 4. Xenomyia acuticeps Emden, head of male, lateral view (modified from Fig. 54 of Emden 1951); 5. Xenomyia atra Malloch, head of female, lateral view (modified from Fig. 53 of Emden 1951); 6. Xenomyia atra Malloch, head of male, lateral view (modified from Fig. 52 of Emden 1951, after Jeanneliotis notabilis Séguy); 7. Xenomyia calyptrata Emden, head and thorax of male, lateral view (modified from Fig. 56 of Emden 1951); 8. Xenomyia edwardsi Emden, head of male, lateral view (modified from Fig. 57 of Emden 1951); 9. Xenomyia edwardsi Emden, head of female, lateral view (modified from Fig. 58 of Emden 1951). Scale bars: 1 mm. Fig. 6 not to scale." pageId="6" pageNumber="266">Fig. 7</figureCitation>
, modified from Fig. 56 of
<bibRefCitation id="E55C2882FF9420686FBB4723FA4C0935" author="Emden, F. I. van" box="[1273,1423,1626,1653]" pageId="6" pageNumber="266" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
); prosternum bare; superior frontal seta absent; acrostichals present; dorsocentrals 1:2; katepisternals 1+2, the posterior seta very strong; presutural intra-alar seta absent; two intralar fine. Anepimeron bare. Fore tibia without median seta. Hind tibia with an anterodorsal seta, without posterodorsal. Wings spotted Lower calypter elongate, more than twice as long as wide (
<figureCitation id="19F649F6FF94206869454793FD9B0845" box="[519,600,1771,1797]" captionStart="FIGURES 2126" captionStartId="10.[151,264,1378,1402]" captionTargetBox="[154,1432,184,1351]" captionTargetId="figure-18@10.[151,1435,181,1354]" captionTargetPageId="10" captionText="FIGURES 2126. 21. Xenomyia calyptrata Emden, wing and calypters of male; a: upper calypter; b: lower calypter (modified from Fig. 53 of Emden 1951); 22. Xenomyia edwardsi Emden, wing of male (modified from Fig. 4, plate 1 of Emden 1951); 23. Xenomyia hirtibasis (Bigot), wing of male; 24. Xenomyia osculata Pont &amp; Werner, wing of male (modified from Fig. 5 of Pont &amp; Werner 2003); 25. Xenomyia pseudolispe Emden, thorax of male, lateral view (modified from Fig. 47 of Emden 1951); 26. Xenomyia bispina Emden, abdomen of male, lateral view (modified from Fig. 7, plate X of Emden 1951). Scale bars: 1 mm. Figs 21, 22 and 24 not to scale." pageId="6" pageNumber="266">Fig. 21</figureCitation>
, modified from
<figureCitation id="19F649F6FF94206868484793FC8E0845" box="[778,845,1771,1797]" captionStart="FIGURES 49" captionStartId="3.[151,264,1822,1846]" captionTargetBox="[221,1369,185,1795]" captionTargetId="figure-33@3.[218,1369,181,1798]" captionTargetPageId="3" captionText="FIGURES 49. 4. Xenomyia acuticeps Emden, head of male, lateral view (modified from Fig. 54 of Emden 1951); 5. Xenomyia atra Malloch, head of female, lateral view (modified from Fig. 53 of Emden 1951); 6. Xenomyia atra Malloch, head of male, lateral view (modified from Fig. 52 of Emden 1951, after Jeanneliotis notabilis Séguy); 7. Xenomyia calyptrata Emden, head and thorax of male, lateral view (modified from Fig. 56 of Emden 1951); 8. Xenomyia edwardsi Emden, head of male, lateral view (modified from Fig. 57 of Emden 1951); 9. Xenomyia edwardsi Emden, head of female, lateral view (modified from Fig. 58 of Emden 1951). Scale bars: 1 mm. Fig. 6 not to scale." pageId="6" pageNumber="266">Fig. 7</figureCitation>
, plate X of
<bibRefCitation id="E55C2882FF942068688E4793FB9E0845" author="Emden, F. I. van" box="[972,1117,1770,1797]" pageId="6" pageNumber="266" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
). Claws a little long.
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R
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bare. Head and thorax in lateral view as in
<figureCitation id="19F649F6FF94206869784677FDB80869" box="[570,635,1807,1833]" captionStart="FIGURES 49" captionStartId="3.[151,264,1822,1846]" captionTargetBox="[221,1369,185,1795]" captionTargetId="figure-33@3.[218,1369,181,1798]" captionTargetPageId="3" captionText="FIGURES 49. 4. Xenomyia acuticeps Emden, head of male, lateral view (modified from Fig. 54 of Emden 1951); 5. Xenomyia atra Malloch, head of female, lateral view (modified from Fig. 53 of Emden 1951); 6. Xenomyia atra Malloch, head of male, lateral view (modified from Fig. 52 of Emden 1951, after Jeanneliotis notabilis Séguy); 7. Xenomyia calyptrata Emden, head and thorax of male, lateral view (modified from Fig. 56 of Emden 1951); 8. Xenomyia edwardsi Emden, head of male, lateral view (modified from Fig. 57 of Emden 1951); 9. Xenomyia edwardsi Emden, head of female, lateral view (modified from Fig. 58 of Emden 1951). Scale bars: 1 mm. Fig. 6 not to scale." pageId="6" pageNumber="266">Fig. 7</figureCitation>
(modified from Fig. 56 of
<bibRefCitation id="E55C2882FF94206868EB4677FBFF0869" author="Emden, F. I. van" box="[937,1084,1806,1833]" pageId="6" pageNumber="266" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
).
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Note. Described from male and female from
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at
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at
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, on rocks) (
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,
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,
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View file

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<taxonomicName id="46CD2EF0FF97206B6BD5476EFDC20970" ID-CoL="7FVV7" authority="Malloch, 1921" authorityName="Malloch" authorityYear="1921" box="[151,513,1558,1585]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="265" phylum="Arthropoda" rank="species" species="atra">
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<emphasis id="B3B98961FF97206B6BD5476EFE890971" bold="true" box="[151,330,1558,1585]" italics="true" pageId="5" pageNumber="265">Xenomyia atra</emphasis>
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Diagnosis. (modified from
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). Length:
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. Black, shining with bluish-grey pruinescence. Head of female as in
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. (modified from Fig. 53 of
<bibRefCitation id="E55C2882FF97206B699547FAFCAF09DD" author="Emden, F. I. van" box="[727,876,1666,1693]" pageId="5" pageNumber="265" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
); black, frons velvety black; face, parafacial and gena with whitish pollinosity. Antenna black; palpus yellow. Dorsum of scutum velvety brownish black, most of pleura grey pruinescent. Legs black. Wings slightly smoky, calypter brown, margins fuscous. Halteres bright yellow. Frons a little over one-third of head-width; orbits not clearly differentiated; ocellar setae present; fronto-orbital plate narrower than frontal vitta; proboscis stout and bulbous, dusted; anepimeron bare; postpedicel about three times as long as pedicel, tapered from middle to apex; arista long and slender; gena about as high as width of postpedicel. Postsutural dorsocentrals 3, katerpisternals 1:1. Legs long and slender. Fore and mid tibia, each with one anterodorsal and one posterodorsal seta; hind femur with very short weak anteroventral setae; hind tibia with 1 anteroventral, 1 anterodorsal and 1 posterodorsal setae. crossvein rm at middle of discal cell. Lower calypter much longer than upper.
</paragraph>
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Note: Described from male and female from
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(NHMUK and MNHN) at
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, forest area.
<bibRefCitation id="E55C2882FF9420686E0F41EFFF210F95" author="Emden, F. I. van" pageId="6" pageNumber="266" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden (1951)</bibRefCitation>
considered
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<emphasis id="B3B98961FF9420686A2A41C3FD9B0F95" box="[360,600,187,213]" italics="true" pageId="6" pageNumber="266">Jeanneliotis notabilis</emphasis>
Séguy
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as a junior synonym of
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<emphasis id="B3B98961FF94206868F241C3FC3E0F94" box="[944,1021,187,212]" italics="true" pageId="6" pageNumber="266">X. atra</emphasis>
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. According to him, the description
<emphasis id="B3B98961FF9420686EC441C3FED00FB9" italics="true" pageId="6" pageNumber="266">
of
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</emphasis>
by
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gives enough details to consider them the same species. Moreover, they were collected in the same mountain, and the male of
<taxonomicName id="46CD2EF0FF9420686921407BFD770E5C" box="[611,692,259,284]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="atra">
<emphasis id="B3B98961FF9420686921407BFD770E5C" box="[611,692,259,284]" italics="true" pageId="6" pageNumber="266">X. atra</emphasis>
</taxonomicName>
and the female of
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<emphasis id="B3B98961FF94206868D6407BFBD70E5D" box="[916,1044,259,285]" italics="true" pageId="6" pageNumber="266">J. notabilis</emphasis>
</taxonomicName>
are unknown. The morphological differences between them, especially in the shape on the head (
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, modified from
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of
<bibRefCitation id="E55C2882FF9420686F9B405FFAA90E01" author="Emden, F. I. van" box="[1241,1386,294,321]" pageId="6" pageNumber="266" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
) are probably sexual differences as occuring in other species of the genus. As commented in the introduction,
<bibRefCitation id="E55C2882FF9420686E084033FF270EC9" author="Hennig, W." pageId="6" pageNumber="266" pagination="1 - 100" refId="ref11267" refString="Hennig, W. (1965) Vorarbeiten zu einem phylogenetischen System der Muscidae (Diptera: Cyclorrhapha). Stuttgater Beitrage zur Naturkunde, 141, 1 - 100, 53 figs." type="journal article" year="1965">Hennig (1965)</bibRefCitation>
, who observed the
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of
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<emphasis id="B3B98961FF942068694F4017FD570EC9" box="[525,660,367,393]" italics="true" pageId="6" pageNumber="266">Jeanneliotis</emphasis>
</taxonomicName>
in Paris, considered this synonym poorly substantiated and according to him it is not even possible to determine whether this genus belong to the
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. At the time of the original description,
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constituted an independent subfamily to it.
</paragraph>
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View file

@ -0,0 +1,111 @@
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<taxonomicName id="46CD2EF0FF97206B6BD5453AFDF70B1C" ID-CoL="7G8DS" authority="Emden, 1951" authorityName="Emden" authorityYear="1951" box="[151,564,1090,1117]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="265" phylum="Arthropoda" rank="species" species="acuticeps">
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<emphasis id="B3B98961FF97206B6BD5453AFE4A0B1C" bold="true" box="[151,393,1090,1116]" italics="true" pageId="5" pageNumber="265">Xenomyia acuticeps</emphasis>
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Diagnosis. (modified from
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). Length:
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. Ocellar setae present; fronto-orbital plate narrower than frontal vitta; profile of head at antennal base much longer than at mouth margin, forming an acute angle (
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, modified from Fig. 54 of
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); proboscis stout and bulbous, dusted; facial ridges with few setulae near vibrissa; claws small; dorsocentral presutural 1; katepisternals 1:1; presutural intra-alar seta absent; posterior intralar seta absent. Anepimeron bare. Mid tarsus of male compressed and slightly dilated, the anterior surface with dense pale golden yellow dust, which is interrupted by a longitudinal row of setae. Hind tibia without posterodorsal seta. Claws small.
</paragraph>
<paragraph id="81725573FF97206B6B8544FFFB610A85" blockId="5.[151,1437,1162,1477]" pageId="5" pageNumber="265">
<materialsCitation id="31A55F2EFF97206B6B8544FFFB5C0A85" collectionCode="NHMUK" country="Uganda" elevation="0" location="Namwamba Valley" pageId="5" pageNumber="265" specimenCount="1" specimenCount-male="1">
Note. Described from male and female from
<collectingCountry id="F9DA15E3FF97206B69FB44FEFCD00AE0" box="[697,787,1414,1440]" name="Uganda" pageId="5" pageNumber="265">Uganda</collectingCountry>
, collected in
<location id="841203A8FF97206B68E744FEFBB80AE0" LSID="urn:lsid:plazi:treatment:0964E465FF97206B6BD5453AFB610A85:841203A8FF97206B68E744FEFBB80AE0" box="[933,1147,1414,1440]" country="Uganda" name="Namwamba Valley" pageId="5" pageNumber="265">Namwamba Valley</location>
, at
<elevation id="0AE0B240FF97206B6FDD44FEFA5E0AE0" box="[1183,1437,1414,1441]" isEnumeration="true" pageId="5" pageNumber="265">
<quantity id="4635F896FF97206B6FDD44FEFB3E0AE0" box="[1183,1277,1414,1441]" metricMagnitude="3" metricUnit="m" metricValue="1.9812" pageId="5" pageNumber="265" unit="ft" value="6500.0">6,500 ft.</quantity>
and
<quantity id="4635F896FF97206B6E7344FFFA5E0AE0" box="[1329,1437,1414,1441]" metricMagnitude="3" metricUnit="m" metricValue="3.10896" pageId="5" pageNumber="265" unit="ft" value="10200.0">10,200 ft.</quantity>
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is labelled: on stones in small rapid stream (
<bibRefCitation id="E55C2882FF97206B69B744D2FC490A85" author="Emden, F. I. van" box="[757,906,1450,1477]" pageId="5" pageNumber="265" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
). Deposited at
<collectionCode id="E7DCCDB6FF97206B6F7344D3FB5C0A85" box="[1073,1183,1451,1477]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" pageNumber="265" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,439 @@
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<mods:title id="F406B97B6BBC115A1BC733107D15604C">The puzzling genus Xenomyia Malloch (Diptera, Muscidae): taxonomic notes, key to species and redescription of Xenomyia hirtibasis (Bigot)</mods:title>
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<taxonomicName id="46CD2EF0FF9B20676BD543CBFDF20D8D" ID-CoL="7G877" authority="(Bigot), 1885" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[151,561,691,718]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="269" phylum="Arthropoda" rank="species" species="hirtibasis">
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<emphasis id="B3B98961FF9B20676BD543CBFE4A0D8E" bold="true" box="[151,393,691,718]" italics="true" pageId="9" pageNumber="269">Xenomyia hirtibasis</emphasis>
(Bigot), 1885
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<emphasis id="B3B98961FF9B20676BD54383FE860C55" bold="true" box="[151,325,763,789]" pageId="9" pageNumber="269">Redescription</emphasis>
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</paragraph>
<paragraph id="81725573FF9B20676BD54245FCAC0C17" blockId="9.[151,1437,829,2043]" box="[151,879,829,856]" pageId="9" pageNumber="269">
Male. Length: body:
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; wing: 4.05.0 mm. (
<figureCitation id="19F649F6FF9B206769A54245FCA40C17" box="[743,871,829,855]" captionStart="FIGURES 4651" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[250,1338,183,1913]" captionTargetId="figure-33@17.[244,1344,181,1919]" captionTargetPageId="17" captionText="FIGURES 4651. 46. Xenomyia hirtibasis (Bigot), male, lateral habitus; 47. Xenomyia hirtibasis (Bigot), male, dorsal habitus; 48. Xenomyia hirtibasis (Bigot), male, head, frontal view; 49. Xenomyia hirtibasis (Bigot), female, lateral habitus; 50. Xenomyia hirtibasis (Bigot), female, dorsal habitus; 51. Xenomyia hirtibasis (Bigot), female, head, frontal view. Scale bars: 1 mm." pageId="9" pageNumber="269">Figs 4648</figureCitation>
)
</paragraph>
<paragraph id="81725573FF9B20676B854219FAD20B4B" blockId="9.[151,1437,829,2043]" pageId="9" pageNumber="269">Color: General color brown, with greyish-blue pollinosity. Frontal vitta and fronto-orbital plate velvety dark brown. Face and gena grey-blue pollinose. Antennae, arista and palpus dark brown. Scutum and scutellum dark brown, subshining, with postpronotum and notopleuron greyish-blue pollinose. Pleura greyish-blue pollinose, upper half of anepisternum dark brown. Calypters creamy-white. Halter yellow. Wing smoky brown. Legs brown with greyish-blue pollinosity. Abdomen dark brown, with lateral greyish-blue pollinose areas in all tergites.</paragraph>
<paragraph id="81725573FF9B20676B85456DFDAA0BFF" blockId="9.[151,1437,829,2043]" pageId="9" pageNumber="269">
Head (
<figureCitation id="19F649F6FF9B20676A50456DFE030B6F" box="[274,448,1045,1071]" captionStart="FIGURES 1015" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[199,1388,185,1847]" captionTargetId="figure-33@7.[196,1391,181,1850]" captionTargetPageId="7" captionText="FIGURES 1015. 10. Xenomyia hirtibasis (Bigot), head of male, lateral view; 11. Xenomyia hirtibasis (Bigot), head of female, lateral view; 12. Xenomyia osculata Pont &amp; Werner, head of male, frontal view (modified from Fig. 1 of Pont &amp; Werner 2003); 13. Xenomyia osculata Pont &amp; Werner, head of female, frontal view (modified from Fig. 2 of Pont &amp; Werner 2003); 14. Xenomyia oxycera Emden, head of male, lateral view (modified from Fig. 55 of Emden 1951); 15. Xenomyia palpalis Emden, head and thorax of female, lateral view (modified from Fig. 4 of Emden 1948). Scale bars: 1 mm. Figs 12, 13 and 15 not to scale." pageId="9" pageNumber="269">Figs 10 and 11</figureCitation>
): Frontal vitta broad; fronto-orbital plate narrow. Eye bare. Ocellar setae present, well-developed. Inner and outer vertical setae strong. Postpedicel short, about 1.5 times the length of the pedicel. Arista long, pubescens, enlarged on basal third. Vibrissae long, with vibrissal setulae. Gena tall, about one third of eye height. Proboscis stout and bulbous, dusted, labellum with pairs of stout teeth (
<figureCitation id="19F649F6FF9B20676F5B45F9FBAC0BDB" box="[1049,1135,1153,1179]" captionStart="FIGURES 4651" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[250,1338,183,1913]" captionTargetId="figure-33@17.[244,1344,181,1919]" captionTargetPageId="17" captionText="FIGURES 4651. 46. Xenomyia hirtibasis (Bigot), male, lateral habitus; 47. Xenomyia hirtibasis (Bigot), male, dorsal habitus; 48. Xenomyia hirtibasis (Bigot), male, head, frontal view; 49. Xenomyia hirtibasis (Bigot), female, lateral habitus; 50. Xenomyia hirtibasis (Bigot), female, dorsal habitus; 51. Xenomyia hirtibasis (Bigot), female, head, frontal view. Scale bars: 1 mm." pageId="9" pageNumber="269">Fig. 48</figureCitation>
). Palpus somewhat dilated apically, flattened, slightly spoon-shaped.
</paragraph>
<paragraph id="81725573FF9B20676B8545B1FF0C0AFB" blockId="9.[151,1437,829,2043]" pageId="9" pageNumber="269">Thorax: Acrostichal absent. Dorsocentrals 2:3. Intra-alars 1+0. Supra-alars 1, prealar absent. 1 postpronotal (the outer), the lobe covered with short dense setulae. Prosternum bare. Postpronotal 1, long. Notopleuron with 2 setae, the posterior seta one-third longer than anterior seta, without setulae. Anepisternum without a short seta in upper anterior corner. Katepisternals 1+1, the upper posterior seta strong. Anepimeron, katepimeron, meron, katatergite and anatergite bare. Posterior spiracle without setae on margins. Scutellum with a pair of sub-basal lateral and apical setae, the apical seta one-third longer than the basal seta; disc sparsely setulose, lateral margins and ventral surface bare.</paragraph>
<paragraph id="81725573FF9B20676B8544BDFE8E092F" blockId="9.[151,1437,829,2043]" pageId="9" pageNumber="269">
Legs: (
<figureCitation id="19F649F6FF9B20676A5A44BDFEB30A9F" box="[280,368,1477,1503]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1909,1933]" captionTargetBox="[216,1371,184,1882]" captionTargetId="figure-18@8.[213,1374,181,1884]" captionTargetPageId="8" captionText="FIGURES 1620. 16. Xenomyia bispina Emden, hind leg of male (modified from Fig. 50 of Emden 1951); 17. Xenomyia hirtibasis Emden, hind leg of male; 18. Xenomyia osculata Pont &amp; Werner, fore femur of male (modified from Fig. 4 of Pont &amp; Werner 2003); 19. Xenomyia perplexa Emden, hind leg of male (modified from Fig. 49 of Emden 1951); 20. Xenomyia pseudolispe Emden, hind leg of male (modified from Fig. 48 of Emden 1951). Scale bars: 1 mm. Fig. 18, not to scale." pageId="9" pageNumber="269">Figs 17</figureCitation>
and
<figureCitation id="19F649F6FF9B20676AEF44BDFE090A9F" box="[429,458,1477,1503]" captionStart="FIGURES 4651" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[250,1338,183,1913]" captionTargetId="figure-33@17.[244,1344,181,1919]" captionTargetPageId="17" captionText="FIGURES 4651. 46. Xenomyia hirtibasis (Bigot), male, lateral habitus; 47. Xenomyia hirtibasis (Bigot), male, dorsal habitus; 48. Xenomyia hirtibasis (Bigot), male, head, frontal view; 49. Xenomyia hirtibasis (Bigot), female, lateral habitus; 50. Xenomyia hirtibasis (Bigot), female, dorsal habitus; 51. Xenomyia hirtibasis (Bigot), female, head, frontal view. Scale bars: 1 mm." pageId="9" pageNumber="269">46</figureCitation>
) Fore coxa greatly developed and with a dense tuft of numerous long fine setae on anterior surface; mid coxa with a row of long seta, mid femur with 46 fine and long posteroventral setae near base followed by a row of 78 pointed setae until two-thirds of the length. Hind femur with 35 long anteroventral setae on apical two-thirds. Hind tibia with a very long anterodorsal seta on basal third (
<figureCitation id="19F649F6FF9B20676F664749FBB4090B" box="[1060,1143,1585,1611]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1909,1933]" captionTargetBox="[216,1371,184,1882]" captionTargetId="figure-18@8.[213,1374,181,1884]" captionTargetPageId="8" captionText="FIGURES 1620. 16. Xenomyia bispina Emden, hind leg of male (modified from Fig. 50 of Emden 1951); 17. Xenomyia hirtibasis Emden, hind leg of male; 18. Xenomyia osculata Pont &amp; Werner, fore femur of male (modified from Fig. 4 of Pont &amp; Werner 2003); 19. Xenomyia perplexa Emden, hind leg of male (modified from Fig. 49 of Emden 1951); 20. Xenomyia pseudolispe Emden, hind leg of male (modified from Fig. 48 of Emden 1951). Scale bars: 1 mm. Fig. 18, not to scale." pageId="9" pageNumber="269">Fig. 17</figureCitation>
). All tarsi long with claws long and strong.
</paragraph>
<paragraph id="81725573FF9B20676B854701FB3A09F7" blockId="9.[151,1437,829,2043]" pageId="9" pageNumber="269">
Wing (
<figureCitation id="19F649F6FF9B20676A574701FEA809D3" box="[277,363,1657,1683]" captionStart="FIGURES 2126" captionStartId="10.[151,264,1378,1402]" captionTargetBox="[154,1432,184,1351]" captionTargetId="figure-18@10.[151,1435,181,1354]" captionTargetPageId="10" captionText="FIGURES 2126. 21. Xenomyia calyptrata Emden, wing and calypters of male; a: upper calypter; b: lower calypter (modified from Fig. 53 of Emden 1951); 22. Xenomyia edwardsi Emden, wing of male (modified from Fig. 4, plate 1 of Emden 1951); 23. Xenomyia hirtibasis (Bigot), wing of male; 24. Xenomyia osculata Pont &amp; Werner, wing of male (modified from Fig. 5 of Pont &amp; Werner 2003); 25. Xenomyia pseudolispe Emden, thorax of male, lateral view (modified from Fig. 47 of Emden 1951); 26. Xenomyia bispina Emden, abdomen of male, lateral view (modified from Fig. 7, plate X of Emden 1951). Scale bars: 1 mm. Figs 21, 22 and 24 not to scale." pageId="9" pageNumber="269">Figs 23</figureCitation>
and
<figureCitation id="19F649F6FF9B20676AE74701FE0509D3" box="[421,454,1657,1683]" captionStart="FIGURES 4651" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[250,1338,183,1913]" captionTargetId="figure-33@17.[244,1344,181,1919]" captionTargetPageId="17" captionText="FIGURES 4651. 46. Xenomyia hirtibasis (Bigot), male, lateral habitus; 47. Xenomyia hirtibasis (Bigot), male, dorsal habitus; 48. Xenomyia hirtibasis (Bigot), male, head, frontal view; 49. Xenomyia hirtibasis (Bigot), female, lateral habitus; 50. Xenomyia hirtibasis (Bigot), female, dorsal habitus; 51. Xenomyia hirtibasis (Bigot), female, head, frontal view. Scale bars: 1 mm." pageId="9" pageNumber="269">46</figureCitation>
): Membrane completely covered with microtrichia. Greater ampula dark brown. Veins bare except
<collectingRegion id="43099B91FF9B20676A5C47E5FEA209F7" box="[286,353,1693,1719]" country="Tanzania" name="Pwani" pageId="9" pageNumber="269">Costa</collectingRegion>
. Both calypters glossiform, lower one about 1.4 times the length of the upper one.
</paragraph>
<paragraph id="81725573FF9B20676B8547B9FC76099B" blockId="9.[151,1437,829,2043]" box="[199,949,1729,1755]" pageId="9" pageNumber="269">Abdomen. Tergites with row of fine setae at margins of all tergites.</paragraph>
<paragraph id="81725573FF9B20676B85479DFAF70863" blockId="9.[151,1437,829,2043]" pageId="9" pageNumber="269">
Genitalia (
<figureCitation id="19F649F6FF9B20676A02479DFE0409BF" box="[320,455,1765,1791]" captionStart="FIGURES 2732" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[224,1366,182,1881]" captionTargetId="figure-33@11.[218,1369,181,1884]" captionTargetPageId="11" captionText="FIGURES 2732. 27. Xenomyia hirtibasis (Bigot), sternite 5 of male; 28. Xenomyia hirtibasis (Bigot), sternite 6 of male; 29. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, dorsal view; 30. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, lateral view; 31. Xenomyia hirtibasis (Bigot), aedeagus, dorsal view; 32. Xenomyia hirtibasis (Bigot), aedeagus, lateral. Scale bars: 0.1 mm." pageId="9" pageNumber="269">Figs 2732</figureCitation>
): Sternite 5 a little taller than wide, with a deep anterior margin (
<figureCitation id="19F649F6FF9B20676F82479DFAD409BF" box="[1216,1303,1765,1791]" captionStart="FIGURES 2732" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[224,1366,182,1881]" captionTargetId="figure-33@11.[218,1369,181,1884]" captionTargetPageId="11" captionText="FIGURES 2732. 27. Xenomyia hirtibasis (Bigot), sternite 5 of male; 28. Xenomyia hirtibasis (Bigot), sternite 6 of male; 29. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, dorsal view; 30. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, lateral view; 31. Xenomyia hirtibasis (Bigot), aedeagus, dorsal view; 32. Xenomyia hirtibasis (Bigot), aedeagus, lateral. Scale bars: 0.1 mm." pageId="9" pageNumber="269">Fig. 27</figureCitation>
). Sternite 6 asymmetric (
<figureCitation id="19F649F6FF9B20676A6B4671FEBD0863" box="[297,382,1801,1827]" captionStart="FIGURES 2732" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[224,1366,182,1881]" captionTargetId="figure-33@11.[218,1369,181,1884]" captionTargetPageId="11" captionText="FIGURES 2732. 27. Xenomyia hirtibasis (Bigot), sternite 5 of male; 28. Xenomyia hirtibasis (Bigot), sternite 6 of male; 29. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, dorsal view; 30. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, lateral view; 31. Xenomyia hirtibasis (Bigot), aedeagus, dorsal view; 32. Xenomyia hirtibasis (Bigot), aedeagus, lateral. Scale bars: 0.1 mm." pageId="9" pageNumber="269">Fig. 28</figureCitation>
). Cercal plate partially divided and surstylus as long as cercal plate (
<figureCitation id="19F649F6FF9B20676F3E4671FAE40863" box="[1148,1319,1801,1827]" captionStart="FIGURES 2732" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[224,1366,182,1881]" captionTargetId="figure-33@11.[218,1369,181,1884]" captionTargetPageId="11" captionText="FIGURES 2732. 27. Xenomyia hirtibasis (Bigot), sternite 5 of male; 28. Xenomyia hirtibasis (Bigot), sternite 6 of male; 29. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, dorsal view; 30. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, lateral view; 31. Xenomyia hirtibasis (Bigot), aedeagus, dorsal view; 32. Xenomyia hirtibasis (Bigot), aedeagus, lateral. Scale bars: 0.1 mm." pageId="9" pageNumber="269">Figs 29 and 30</figureCitation>
).
</paragraph>
<paragraph id="81725573FF9B20676B854655FCBE082B" blockId="9.[151,1437,829,2043]" pageId="9" pageNumber="269">
Aedeagus with the epiphallus well-developed; phallapodeme long, narrow; pregonite long; postgonite shorter than pregonite; distiphallus membranous at apex (
<figureCitation id="19F649F6FF9B206769844629FCB2082B" box="[710,881,1873,1899]" captionStart="FIGURES 2732" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[224,1366,182,1881]" captionTargetId="figure-33@11.[218,1369,181,1884]" captionTargetPageId="11" captionText="FIGURES 2732. 27. Xenomyia hirtibasis (Bigot), sternite 5 of male; 28. Xenomyia hirtibasis (Bigot), sternite 6 of male; 29. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, dorsal view; 30. Xenomyia hirtibasis (Bigot), cercal plate and surstyli, lateral view; 31. Xenomyia hirtibasis (Bigot), aedeagus, dorsal view; 32. Xenomyia hirtibasis (Bigot), aedeagus, lateral. Scale bars: 0.1 mm." pageId="9" pageNumber="269">Figs 31 and 32</figureCitation>
).
</paragraph>
<paragraph id="81725573FF9B20676B85460DFBC008D0" blockId="9.[151,1437,829,2043]" box="[199,1027,1909,1936]" pageId="9" pageNumber="269">
Female: Similar to male, mainly differing in leg chaetotaxy. (
<figureCitation id="19F649F6FF9B20676831460DFC3408D0" box="[883,1015,1909,1936]" captionStart="FIGURES 4651" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[250,1338,183,1913]" captionTargetId="figure-33@17.[244,1344,181,1919]" captionTargetPageId="17" captionText="FIGURES 4651. 46. Xenomyia hirtibasis (Bigot), male, lateral habitus; 47. Xenomyia hirtibasis (Bigot), male, dorsal habitus; 48. Xenomyia hirtibasis (Bigot), male, head, frontal view; 49. Xenomyia hirtibasis (Bigot), female, lateral habitus; 50. Xenomyia hirtibasis (Bigot), female, dorsal habitus; 51. Xenomyia hirtibasis (Bigot), female, head, frontal view. Scale bars: 1 mm." pageId="9" pageNumber="269">Figs 4951</figureCitation>
).
</paragraph>
<paragraph id="81725573FF9B20676B8546E1FD5F08BB" blockId="9.[151,1437,829,2043]" pageId="9" pageNumber="269">Legs: Fore coxa without a dense tuft of numerous long fine setae on anterior surface. Mid and hind femora practically without differentiated setae, except for a fine preapical dorsal seta on both femora. Hind tibia with the anterodorsal seta on basal third short and fine.</paragraph>
<caption id="D5B205FBFF9820646BD5441AFE17096E" pageId="10" pageNumber="270" startId="10.[151,264,1378,1402]" targetBox="[154,1432,184,1351]" targetPageId="10" targetType="figure">
<paragraph id="81725573FF9820646BD5441AFE17096E" blockId="10.[151,1437,1378,1582]" pageId="10" pageNumber="270">
<emphasis id="B3B98961FF9820646BD5441AFE970A3A" bold="true" box="[151,340,1378,1402]" pageId="10" pageNumber="270">FIGURES 2126.</emphasis>
21.
<taxonomicName id="46CD2EF0FF9820646AC2441BFD670A3A" authority="Emden" authorityName="Emden" box="[384,676,1378,1402]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="calyptrata">
<emphasis id="B3B98961FF9820646AC2441BFD970A3A" box="[384,596,1378,1402]" italics="true" pageId="10" pageNumber="270">Xenomyia calyptrata</emphasis>
Emden
</taxonomicName>
, wing and calypters of male; a: upper calypter; b: lower calypter (modified from Fig. 53 of
<bibRefCitation id="E55C2882FF9820646A7C44FEFE000ADE" author="Emden, F. I. van" box="[318,451,1414,1438]" pageId="10" pageNumber="270" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
); 22.
<taxonomicName id="46CD2EF0FF9820646ABF44FFFCD50ADE" authority="Emden" authorityName="Emden" box="[509,790,1414,1438]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="B3B98961FF9820646ABF44FFFD050ADE" box="[509,710,1414,1438]" italics="true" pageId="10" pageNumber="270">Xenomyia edwardsi</emphasis>
Emden
</taxonomicName>
, wing of male (modified from Fig. 4, plate 1 of
<bibRefCitation id="E55C2882FF9820646E4944FEFA530ADE" author="Emden, F. I. van" box="[1291,1424,1414,1438]" pageId="10" pageNumber="270" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
); 23.
<taxonomicName id="46CD2EF0FF9820646BFF44D3FE180A82" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[189,475,1450,1474]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF9820646BFF44D3FE490A82" box="[189,394,1450,1474]" italics="true" pageId="10" pageNumber="270">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, wing of male; 24.
<taxonomicName id="46CD2EF0FF98206469E044D3FBC40A82" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" box="[674,1031,1450,1474]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF98206469E044D3FCA60A82" box="[674,869,1450,1474]" italics="true" pageId="10" pageNumber="270">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, wing of male (modified from Fig. 5 of
<bibRefCitation id="E55C2882FF9820646BD544B7FEAE0AA6" author="Werner, D. &amp; Pont, A. C." box="[151,365,1486,1510]" pageId="10" pageNumber="270" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner 2003</bibRefCitation>
); 25.
<taxonomicName id="46CD2EF0FF9820646AE644B7FD1B0AA6" authority="Emden" authorityName="Emden" box="[420,728,1486,1510]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="pseudolispe">
<emphasis id="B3B98961FF9820646AE644B7FD4B0AA6" box="[420,648,1486,1510]" italics="true" pageId="10" pageNumber="270">Xenomyia pseudolispe</emphasis>
Emden
</taxonomicName>
, thorax of male, lateral view (modified from Fig. 47 of
<bibRefCitation id="E55C2882FF9820646E4E44B6FA530AA6" author="Emden, F. I. van" box="[1292,1424,1486,1510]" pageId="10" pageNumber="270" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
); 26.
<taxonomicName id="46CD2EF0FF9820646BFF448BFE00094A" authority="Emden" authorityName="Emden" box="[189,451,1522,1546]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="bispina">
<emphasis id="B3B98961FF9820646BFF448BFEB0094A" box="[189,371,1522,1546]" italics="true" pageId="10" pageNumber="270">Xenomyia bispina</emphasis>
Emden
</taxonomicName>
, abdomen of male, lateral view (modified from Fig. 7, plate X of
<bibRefCitation id="E55C2882FF9820646F16448AFB14094A" author="Emden, F. I. van" box="[1108,1239,1522,1546]" pageId="10" pageNumber="270" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
). Scale bars: 1 mm. Figs 21, 22 and 24 not to scale.
</paragraph>
</caption>
<paragraph id="81725573FF9820646B8547F8FB4009A2" blockId="10.[151,1437,1664,1979]" pageId="10" pageNumber="270">
Ovipositor (
<figureCitation id="19F649F6FF9820646A0C47F8FE3809DA" box="[334,507,1664,1690]" captionStart="FIGURES 3941" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[155,1431,316,1909]" captionTargetId="figure-78@14.[151,1435,309,1913]" captionTargetPageId="14" captionText="FIGURES 3941. 39. Xenomyia hirtibasis (Bigot), ovipositor, dorsal view; 40. Xenomyia hirtibasis (Bigot), ovipositor, lateral view; 41. Xenomyia osculata Pont &amp; Werner, ovipositor, dorsal view (modified from Fig. 11 of Pont &amp; Werner 2003). Scale bars: 0.2 mm. Fig. 41, not to scale." pageId="10" pageNumber="270">Figs 39 and 40</figureCitation>
): Long, all tergites broad, well-developed, cercus about twice longer as long as the epiproct, with long setae on outer margin. Sternite 6 elongate-rectangular; sternite 7 a small plate; sternite 8 absent. The membrane between sternites 7 and 8 with microtrichia. Three rounded spermathecae.
</paragraph>
<paragraph id="81725573FF9820646B854794FAB108FA" blockId="10.[151,1437,1664,1979]" pageId="10" pageNumber="270">
Material examined.
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, Stellenbosch, Jonkershoek NR, t. Tweede Waterval, Eerste R.,
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201812022,
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201812021,
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1053.7”
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E19
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0720.9”
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,
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<collectingDate id="E5378A5BFF9820646E564620FA5B0833" box="[1300,1432,1880,1907]" pageId="10" pageNumber="270" value="2018-12-05">05/xii/2018</collectingDate>
</date>
,
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&amp;
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,
<collectionCode id="E7DCCDB6FF9820646A114604FE6008D7" box="[339,419,1916,1943]" pageId="10" pageNumber="270">LHGA</collectionCode>
</materialsCitation>
201812051,
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<specimenCount id="97CB9EFAFF982064696F4604FDB808D6" box="[557,635,1916,1942]" count="1" pageId="10" pageNumber="270" type="male">1 male</specimenCount>
,
<specimenCount id="97CB9EFAFF98206469C44604FD2408D6" box="[646,743,1916,1942]" count="1" pageId="10" pageNumber="270" type="female">1 female</specimenCount>
(
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)
</materialsCitation>
,
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(NMB). All in good conditions. Although adults were not observed preying, they were seen flying over areas where immature
<taxonomicName id="46CD2EF0FF9820646F0546D8FB0308FA" authorityName="Newman" authorityYear="1834" box="[1095,1216,1952,1978]" class="Insecta" family="Simuliidae" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="270" phylum="Arthropoda" rank="family">Simuliidae</taxonomicName>
were collected.
</paragraph>
<caption id="D5B205FBFF9920656BD5460DFEE008B8" pageId="11" pageNumber="271" startId="11.[151,264,1909,1933]" targetBox="[224,1366,182,1881]" targetPageId="11" targetType="figure">
<paragraph id="81725573FF9920656BD5460DFEE008B8" blockId="11.[151,1436,1909,2041]" pageId="11" pageNumber="271">
<emphasis id="B3B98961FF9920656BD5460DFE9B08CD" bold="true" box="[151,344,1909,1933]" pageId="11" pageNumber="271">FIGURES 2732.</emphasis>
27.
<taxonomicName id="46CD2EF0FF9920656ACA460EFD6B08CD" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[392,680,1909,1933]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="271" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF9920656ACA460EFD9408CD" box="[392,599,1909,1933]" italics="true" pageId="11" pageNumber="271">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, sternite 5 of male; 28.
<taxonomicName id="46CD2EF0FF99206568E8460EFB0808CD" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[938,1227,1909,1933]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="271" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF99206568E8460EFBBA08CD" box="[938,1145,1909,1933]" italics="true" pageId="11" pageNumber="271">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, sternite 6 of male; 29.
<taxonomicName id="46CD2EF0FF9920656BFF46E2FE1A08F1" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[189,473,1945,1969]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="271" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF9920656BFF46E2FE4B08F1" box="[189,392,1945,1969]" italics="true" pageId="11" pageNumber="271">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, cercal plate and surstyli, dorsal view; 30.
<taxonomicName id="46CD2EF0FF99206568C146E2FB5C08F1" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[899,1183,1945,1969]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="271" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF99206568C146E2FB8D08F1" box="[899,1102,1945,1969]" italics="true" pageId="11" pageNumber="271">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, cercal plate and surstyli, lateral view; 31.
<taxonomicName id="46CD2EF0FF9920656A0646C6FDA10895" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[324,610,1981,2005]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="271" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF9920656A0646C6FDD20895" box="[324,529,1981,2005]" italics="true" pageId="11" pageNumber="271">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, aedeagus, dorsal view; 32.
<taxonomicName id="46CD2EF0FF99206568C646C6FB610895" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[900,1186,1981,2005]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="271" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF99206568C646C6FB930895" box="[900,1104,1981,2005]" italics="true" pageId="11" pageNumber="271">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, aedeagus, lateral. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="D5B205FBFF9E20626BD5462AFB1F08BA" pageId="12" pageNumber="272" startId="12.[151,264,1874,1898]" targetBox="[206,1382,184,1847]" targetPageId="12" targetType="figure">
<paragraph id="81725573FF9E20626BD5462AFB1F08BA" blockId="12.[151,1437,1874,2042]" pageId="12" pageNumber="272">
<emphasis id="B3B98961FF9E20626BD5462AFE90082A" bold="true" box="[151,339,1874,1898]" pageId="12" pageNumber="272">FIGURES 3338.</emphasis>
33.
<taxonomicName id="46CD2EF0FF9E20626A3C462BFD5B082A" authority="Zielke" authorityName="Zielke" authorityYear="1970" box="[382,664,1874,1898]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="272" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="B3B98961FF9E20626A3C462BFD8C082A" box="[382,591,1874,1898]" italics="true" pageId="12" pageNumber="272">Xenomyia natalensis</emphasis>
Zielke
</taxonomicName>
, cercal plate and surstyli of male, dorsal view (modified from Fig. 3 of
<bibRefCitation id="E55C2882FF9E20626E1E462AFF0D08CE" author="Zielke, E." pageId="12" pageNumber="272" pagination="1 - 14" refId="ref11648" refString="Zielke, E. (1970) Description of new Muscidae (Diptera) from South Africa. Novos Taxa entomologicos, 74, 1 - 14." type="journal article" year="1970">Zielke 1970</bibRefCitation>
); 34.
<taxonomicName id="46CD2EF0FF9E20626A44460FFDAB08CE" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" box="[262,616,1910,1934]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="272" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF9E20626A44460FFE0408CE" box="[262,455,1910,1934]" italics="true" pageId="12" pageNumber="272">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, sternite 5 of male; 35.
<taxonomicName id="46CD2EF0FF9E20626814460FFB7B08CE" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" box="[854,1208,1910,1934]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="272" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF9E20626814460FFBDB08CE" box="[854,1048,1910,1934]" italics="true" pageId="12" pageNumber="272">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, epandrium and cercal plate, dorsal view; 36.
<taxonomicName id="46CD2EF0FF9E20626A3D46E3FD2008F2" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" box="[383,739,1946,1970]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="272" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF9E20626A3D46E3FD8208F2" box="[383,577,1946,1970]" italics="true" pageId="12" pageNumber="272">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, epandrium, cercal plate and surstyli; 37.
<taxonomicName id="46CD2EF0FF9E20626FC946E3FF200896" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="272" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF9E20626FC946E3FA8E08F2" box="[1163,1357,1946,1970]" italics="true" pageId="12" pageNumber="272">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, hypandrium and aedeagus, dorsal view; 38.
<taxonomicName id="46CD2EF0FF9E206269EA46C7FBCA0896" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" box="[680,1033,1982,2006]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="272" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF9E206269EA46C7FCAA0896" box="[680,873,1982,2006]" italics="true" pageId="12" pageNumber="272">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, hypandrium and aedeagus, lateral view (Figs 3438 modified from Figs 610 of
<bibRefCitation id="E55C2882FF9E20626975469BFCCE08BA" author="Werner, D. &amp; Pont, A. C." box="[567,781,2018,2042]" pageId="12" pageNumber="272" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner 2003</bibRefCitation>
); Scale bar: 0.5 mm. Figs 3438, not to scale.
</paragraph>
</caption>
<paragraph id="81725573FF9F20636B8541EFFCC20EED" blockId="13.[151,1437,151,429]" pageId="13" pageNumber="273">
Note. Described from male and female from
<collectingCountry id="F9DA15E3FF9F206369F941EFFC880FF1" box="[699,843,151,177]" name="South Africa" pageId="13" pageNumber="273">South Africa</collectingCountry>
,
<collectingRegion id="43099B91FF9F2063681441EFFC510FF1" box="[854,914,151,177]" country="South Africa" name="Western Cape" pageId="13" pageNumber="273">Cape</collectingRegion>
,
<collectingRegion id="43099B91FF9F206368DF41EFFC140FF1" box="[925,983,151,177]" country="South Africa" name="Western Cape" pageId="13" pageNumber="273">Cape</collectingRegion>
of Good Hope. (OUMNH). The original description, in the genus
<taxonomicName id="46CD2EF0FF9F20636AF141C3FE330F94" authorityName="Latreille" authorityYear="1797" box="[435,496,187,212]" class="Insecta" family="Muscidae" genus="Lispe" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B3B98961FF9F20636AF141C3FE330F94" box="[435,496,187,212]" italics="true" pageId="13" pageNumber="273">Lispe</emphasis>
</taxonomicName>
, is very brief.
<bibRefCitation id="E55C2882FF9F206369D441C3FCE10F95" author="Stein, P." box="[662,802,187,213]" pageId="13" pageNumber="273" pagination="273 - 293" refId="ref11555" refString="Stein, P. (1907) Revision der Bigot'schen und einiger von Macquart beschriebenen aussereuropaischen Anthomyiden (Dipt.). [Concl.]. Zeitschrift fur systematische hymenopterologie und dipterologie, 7, 4, 273 - 293." type="journal article" year="1907">Stein (1907)</bibRefCitation>
redescribed the species with more details mainly on the leg chaetotaxy and mentioned that the absence of setae on anepimeron should better positioned it in the genus
<taxonomicName id="46CD2EF0FF9F20636BD5407BFE540E5D" authority="Rondani" authorityName="Rondani" authorityYear="1866" box="[151,407,259,285]" class="Insecta" family="Muscidae" genus="Melanochelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B3B98961FF9F20636BD5407BFEF20E5D" box="[151,305,259,285]" italics="true" pageId="13" pageNumber="273">Melanochelia</emphasis>
Rondani
</taxonomicName>
(=
<taxonomicName id="46CD2EF0FF9F20636AFD407BFCF40E5D" authority="Robineau-Desvoidy" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[447,823,259,285]" class="Insecta" family="Muscidae" genus="Limnophora" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B3B98961FF9F20636AFD407BFD8A0E5D" box="[447,585,259,285]" italics="true" pageId="13" pageNumber="273">Limnophora</emphasis>
Robineau-Desvoidy
</taxonomicName>
). Later,
<bibRefCitation id="E55C2882FF9F206368D2407BFBF20E5D" author="Emden, F. I. van" box="[912,1073,258,285]" pageId="13" pageNumber="273" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden (1951)</bibRefCitation>
saw the same specimen seen by
<bibRefCitation id="E55C2882FF9F20636BD5405FFEE10E01" author="Stein, P." box="[151,290,295,321]" pageId="13" pageNumber="273" pagination="273 - 293" refId="ref11555" refString="Stein, P. (1907) Revision der Bigot'schen und einiger von Macquart beschriebenen aussereuropaischen Anthomyiden (Dipt.). [Concl.]. Zeitschrift fur systematische hymenopterologie und dipterologie, 7, 4, 273 - 293." type="journal article" year="1907">Stein (1907)</bibRefCitation>
(designated by inference as
<typeStatus id="5E76EBD1FF9F2063691C405FFD060E01" box="[606,709,295,321]" pageId="13" pageNumber="273" type="lectotype">lectotype</typeStatus>
by
<bibRefCitation id="E55C2882FF9F206369AF405FFC9C0E01" author="Pont, A. C." box="[749,863,295,321]" pageId="13" pageNumber="273" pagination="721 - 761" refId="ref11404" refString="Pont, A. C. (1980) Family Muscidae. In: Crosskey, R. W. (Ed.), Catalog of the Diptera of the Afrotropical Region. London, British Museum (Natural History), pp. 721 - 761." type="book chapter" year="1980">Pont 1980</bibRefCitation>
) and included the species in his key. More recently,
<bibRefCitation id="E55C2882FF9F20636BD54033FE500E25" author="Werner, D. &amp; Pont, A. C." box="[151,403,331,357]" pageId="13" pageNumber="273" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner (2003)</bibRefCitation>
also included this species in their key for the
<taxonomicName id="46CD2EF0FF9F206368D84033FBC90E24" authorityName="Malloch" authorityYear="1921" box="[922,1034,331,356]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B3B98961FF9F206368D84033FBC90E24" box="[922,1034,331,356]" italics="true" pageId="13" pageNumber="273">Xenomyia</emphasis>
</taxonomicName>
of Southern Africa.
<specimenCount id="97CB9EFAFF9F20636FB04033FA9D0E25" box="[1266,1374,331,357]" count="1" pageId="13" pageNumber="273" type="male">One male</specimenCount>
from
<collectingRegion id="43099B91FF9F20636BD54017FEFA0EC9" box="[151,313,367,393]" country="South Africa" name="Western Cape" pageId="13" pageNumber="273">Western Cape</collectingRegion>
, Buffelsbos forest,
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,
<geoCoordinate id="E4F933B4FF9F206369F94017FC8F0EC9" box="[699,844,367,393]" degrees="23" direction="east" minutes="38.329" orientation="longitude" pageId="13" pageNumber="273" precision="1" value="23.638817">23°38.329E</geoCoordinate>
,
<quantity id="4635F896FF9F206368194017FC640EC9" box="[859,935,367,393]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="273" unit="m" value="400.0">400 m</quantity>
, indigenous montane forest (stream), Kirk-Spriggs, A.H. (BMSA(D)08463 is deposited at BMSA.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,123 @@
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<mods:title id="0300C428691824ADCE618A8217E3DDD5">The puzzling genus Xenomyia Malloch (Diptera, Muscidae): taxonomic notes, key to species and redescription of Xenomyia hirtibasis (Bigot)</mods:title>
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<emphasis id="B3B98961FF9B20676BD541EFFE460FF2" bold="true" box="[151,389,151,178]" italics="true" pageId="9" pageNumber="269">Xenomyia edwardsi</emphasis>
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Diagnosis. (modified from
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). Length:
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. Head of male and female, respectively in
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(modified from Figs 57 and 58 of
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). Ocellar absent; fronto-orbital plate much broader than frontal vitta, densely covered by short setulae; postpedicel round at apex; proboscis slender and glossy black; antenna and palpus black, long and slender; arista short (
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, modified, respectively from Figs 57 and 58 of
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); prosternum setulose; superior frontal seta absent; acrostichals present; dorsocentrals, only the last strong; katepisternals 0:2; presutural intra-alar seta absent; two intralar weak. Anepimeron bare. Fore tibia without median seta. Hind tibia with an anterodorsal seta, without posterodorsal. Wings spotted (
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, modified from
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, plate 1 of
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). Lower calypter fuscous. Halter yellow. Claws a little long.
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R
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setulose.
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Note. Described from male and female from
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, Mobuku River, at circa
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, and from
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in Swam River at
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, on rocks “wetted by spray in the both parts of the two rivers” (
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) (NHMUK). Known geographical distribution,
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and
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.
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<taxonomicName id="46CD2EF0FF9F20636BD54087FDF10D58" ID-CoL="7FVV8" authority="Zielke, 1970" authorityName="Zielke" authorityYear="1970" box="[151,562,510,537]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="273" phylum="Arthropoda" rank="species" species="natalensis">
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<emphasis id="B3B98961FF9F20636BD54087FE500D59" bold="true" box="[151,403,510,537]" italics="true" pageId="13" pageNumber="273">Xenomyia natalensis</emphasis>
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Diagnosis. (modified from
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). Length: body
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. Male with eyes bare. Fronto-orbital plates not enlarged; ocellar setae present; fronto-orbital plate narrower than frontal vitta; proboscis short, stout and bulbous, dusted; facial ridges with few setulae near vibrissa; antenna brown, arista long and bare; palpus light greyish dusted; dorsocentrals 1:3; katepisternals 0:1; presutural intra-alar seta present; fore tibia without a posterior seta, hind tibia with one median anterodorsal seta and a row of short and strong posterodorsals in the apical third; female with a ventral seta. Metatarsus of hind leg with a comb of short and strong setae, becoming stronger in the apical half. Claws long. Wing brownish; calypter yellowish; halter yellow. Cercal plate very broad (
<figureCitation id="19F649F6FF9F20636F284267FB7F0C79" box="[1130,1212,799,825]" captionStart="FIGURES 3338" captionStartId="12.[151,264,1874,1898]" captionTargetBox="[206,1382,184,1847]" captionTargetId="figure-18@12.[203,1385,181,1850]" captionTargetPageId="12" captionText="FIGURES 3338. 33. Xenomyia natalensis Zielke, cercal plate and surstyli of male, dorsal view (modified from Fig. 3 of Zielke 1970); 34. Xenomyia osculata Pont &amp; Werner, sternite 5 of male; 35. Xenomyia osculata Pont &amp; Werner, epandrium and cercal plate, dorsal view; 36. Xenomyia osculata Pont &amp; Werner, epandrium, cercal plate and surstyli; 37. Xenomyia osculata Pont &amp; Werner, hypandrium and aedeagus, dorsal view; 38. Xenomyia osculata Pont &amp; Werner, hypandrium and aedeagus, lateral view (Figs 3438 modified from Figs 610 of Pont &amp; Werner 2003); Scale bar: 0.5 mm. Figs 3438, not to scale." pageId="13" pageNumber="273">Fig. 33</figureCitation>
, modified from
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of
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).
</paragraph>
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Note. Known from male and female
<collectingCountry id="F9DA15E3FF9F20636921421FFD300CC1" box="[611,755,871,897]" name="South Africa" pageId="13" pageNumber="273">South Africa</collectingCountry>
,
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, Giants Castle. deposited in
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.
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View file

@ -0,0 +1,156 @@
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Diagnosis. (modified from
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). (
<figureCitation id="19F649F6FF9F206369964563FCE60B75" box="[724,805,1051,1077]" captionStart="FIGURES 4245" captionStartId="15.[151,264,1908,1932]" captionTargetBox="[179,1408,189,1878]" captionTargetId="figure-33@15.[172,1415,181,1885]" captionTargetPageId="15" captionText="FIGURES 4245. 42. Xenomyia osculata Pont &amp;Werner,male head, lateral view (from Pont &amp;Werner 2003, with the permission of the publisher); 43. Xenomyia osculata Pont &amp; Werner, male, lateral habitus; 44. Xenomyia osculata Pont &amp; Werner, male, dorsal habitus; 45. Xenomyia osculata Pont &amp; Werner, male, head, frontal view. Figs 4345 modified from Couri &amp; Pont 2000). Scale bars: 2 mm. Fig. 42, not to scale." pageId="13" pageNumber="273">Fig. 42</figureCitation>
reproduced from
<bibRefCitation id="E55C2882FF9F206368B24563FB1F0B75" author="Werner, D. &amp; Pont, A. C." box="[1008,1244,1051,1077]" pageId="13" pageNumber="273" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner 2003</bibRefCitation>
, with permission of the publisher, see Material and Methods);
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, modified from
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). Length: 4.55.0 mm. Shape of head very distinct with small eye, arched frons, fronto-orbital plates enlarged, with margins touching and covered with dense small setulae (
<figureCitation id="19F649F6FF9F2063695C45FFFD0A0BE1" box="[542,713,1159,1185]" captionStart="FIGURES 1015" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[199,1388,185,1847]" captionTargetId="figure-33@7.[196,1391,181,1850]" captionTargetPageId="7" captionText="FIGURES 1015. 10. Xenomyia hirtibasis (Bigot), head of male, lateral view; 11. Xenomyia hirtibasis (Bigot), head of female, lateral view; 12. Xenomyia osculata Pont &amp; Werner, head of male, frontal view (modified from Fig. 1 of Pont &amp; Werner 2003); 13. Xenomyia osculata Pont &amp; Werner, head of female, frontal view (modified from Fig. 2 of Pont &amp; Werner 2003); 14. Xenomyia oxycera Emden, head of male, lateral view (modified from Fig. 55 of Emden 1951); 15. Xenomyia palpalis Emden, head and thorax of female, lateral view (modified from Fig. 4 of Emden 1948). Scale bars: 1 mm. Figs 12, 13 and 15 not to scale." pageId="13" pageNumber="273">Figs 12 and 13</figureCitation>
modified from
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and
<figureCitation id="19F649F6FF9F20636F5945FFFBAF0BE1" box="[1051,1132,1159,1185]" captionStart="FIGURES 4245" captionStartId="15.[151,264,1908,1932]" captionTargetBox="[179,1408,189,1878]" captionTargetId="figure-33@15.[172,1415,181,1885]" captionTargetPageId="15" captionText="FIGURES 4245. 42. Xenomyia osculata Pont &amp;Werner,male head, lateral view (from Pont &amp;Werner 2003, with the permission of the publisher); 43. Xenomyia osculata Pont &amp; Werner, male, lateral habitus; 44. Xenomyia osculata Pont &amp; Werner, male, dorsal habitus; 45. Xenomyia osculata Pont &amp; Werner, male, head, frontal view. Figs 4345 modified from Couri &amp; Pont 2000). Scale bars: 2 mm. Fig. 42, not to scale." pageId="13" pageNumber="273">Fig. 42</figureCitation>
reproduced from
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of
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). Ocellar setae absent; antennal insertion not produced; arista very short, short pubescent; eye small with short sparse hairs; proboscis elongate, dark brown; palpus long and fine; facial ridges with few setulae near vibrissa. Mesonotum with acrostichal setulae in 23 rows before suture and 34 irregular rows behind suture, dorsocentrals 2:3; presutural intra-alar seta present; anepimeron bare; katepisternals 1+2, only the upper posterior seta strong; prosternum bare. Legs with fore femur with a spine-like row of 57 posteroventral setae (
<figureCitation id="19F649F6FF9F20636E034443FA540A15" box="[1345,1431,1339,1365]" captionStart="FIGURES 1620" captionStartId="8.[151,264,1909,1933]" captionTargetBox="[216,1371,184,1882]" captionTargetId="figure-18@8.[213,1374,181,1884]" captionTargetPageId="8" captionText="FIGURES 1620. 16. Xenomyia bispina Emden, hind leg of male (modified from Fig. 50 of Emden 1951); 17. Xenomyia hirtibasis Emden, hind leg of male; 18. Xenomyia osculata Pont &amp; Werner, fore femur of male (modified from Fig. 4 of Pont &amp; Werner 2003); 19. Xenomyia perplexa Emden, hind leg of male (modified from Fig. 49 of Emden 1951); 20. Xenomyia pseudolispe Emden, hind leg of male (modified from Fig. 48 of Emden 1951). Scale bars: 1 mm. Fig. 18, not to scale." pageId="13" pageNumber="273">Fig. 18</figureCitation>
, modified from
<figureCitation id="19F649F6FF9F20636A7D4427FEBC0A39" box="[319,383,1375,1401]" captionStart="FIGURES 49" captionStartId="3.[151,264,1822,1846]" captionTargetBox="[221,1369,185,1795]" captionTargetId="figure-33@3.[218,1369,181,1798]" captionTargetPageId="3" captionText="FIGURES 49. 4. Xenomyia acuticeps Emden, head of male, lateral view (modified from Fig. 54 of Emden 1951); 5. Xenomyia atra Malloch, head of female, lateral view (modified from Fig. 53 of Emden 1951); 6. Xenomyia atra Malloch, head of male, lateral view (modified from Fig. 52 of Emden 1951, after Jeanneliotis notabilis Séguy); 7. Xenomyia calyptrata Emden, head and thorax of male, lateral view (modified from Fig. 56 of Emden 1951); 8. Xenomyia edwardsi Emden, head of male, lateral view (modified from Fig. 57 of Emden 1951); 9. Xenomyia edwardsi Emden, head of female, lateral view (modified from Fig. 58 of Emden 1951). Scale bars: 1 mm. Fig. 6 not to scale." pageId="13" pageNumber="273">Fig. 4</figureCitation>
of
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); fore tibia without submedian setae; hind tibia with 1 anterodorsal and 1 posterodorsal submedian seta; claws not enlarged. Wing completely covered with microtrichia, denser in apical area between
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R
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and
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, resembling a cloud (
<figureCitation id="19F649F6FF9F2063698E44DFFCDC0A81" box="[716,799,1447,1473]" captionStart="FIGURES 2126" captionStartId="10.[151,264,1378,1402]" captionTargetBox="[154,1432,184,1351]" captionTargetId="figure-18@10.[151,1435,181,1354]" captionTargetPageId="10" captionText="FIGURES 2126. 21. Xenomyia calyptrata Emden, wing and calypters of male; a: upper calypter; b: lower calypter (modified from Fig. 53 of Emden 1951); 22. Xenomyia edwardsi Emden, wing of male (modified from Fig. 4, plate 1 of Emden 1951); 23. Xenomyia hirtibasis (Bigot), wing of male; 24. Xenomyia osculata Pont &amp; Werner, wing of male (modified from Fig. 5 of Pont &amp; Werner 2003); 25. Xenomyia pseudolispe Emden, thorax of male, lateral view (modified from Fig. 47 of Emden 1951); 26. Xenomyia bispina Emden, abdomen of male, lateral view (modified from Fig. 7, plate X of Emden 1951). Scale bars: 1 mm. Figs 21, 22 and 24 not to scale." pageId="13" pageNumber="273">Fig. 24</figureCitation>
, modified from
<figureCitation id="19F649F6FF9F2063689544DFFBDA0A81" box="[983,1049,1446,1473]" captionStart="FIGURES 49" captionStartId="3.[151,264,1822,1846]" captionTargetBox="[221,1369,185,1795]" captionTargetId="figure-33@3.[218,1369,181,1798]" captionTargetPageId="3" captionText="FIGURES 49. 4. Xenomyia acuticeps Emden, head of male, lateral view (modified from Fig. 54 of Emden 1951); 5. Xenomyia atra Malloch, head of female, lateral view (modified from Fig. 53 of Emden 1951); 6. Xenomyia atra Malloch, head of male, lateral view (modified from Fig. 52 of Emden 1951, after Jeanneliotis notabilis Séguy); 7. Xenomyia calyptrata Emden, head and thorax of male, lateral view (modified from Fig. 56 of Emden 1951); 8. Xenomyia edwardsi Emden, head of male, lateral view (modified from Fig. 57 of Emden 1951); 9. Xenomyia edwardsi Emden, head of female, lateral view (modified from Fig. 58 of Emden 1951). Scale bars: 1 mm. Fig. 6 not to scale." pageId="13" pageNumber="273">Fig. 5</figureCitation>
of
<bibRefCitation id="E55C2882FF9F20636F0244DFFAED0A81" author="Werner, D. &amp; Pont, A. C." box="[1088,1326,1447,1473]" pageId="13" pageNumber="273" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner 2003</bibRefCitation>
); calypter creamy-white, lower one projecting slightly beyond upper one; halter with the stalk creamy and the knob white. Sternite 5 and terminalia as in
<figureCitation id="19F649F6FF9F20636AAF4497FDA80949" box="[493,619,1519,1545]" captionStart="FIGURES 3338" captionStartId="12.[151,264,1874,1898]" captionTargetBox="[206,1382,184,1847]" captionTargetId="figure-18@12.[203,1385,181,1850]" captionTargetPageId="12" captionText="FIGURES 3338. 33. Xenomyia natalensis Zielke, cercal plate and surstyli of male, dorsal view (modified from Fig. 3 of Zielke 1970); 34. Xenomyia osculata Pont &amp; Werner, sternite 5 of male; 35. Xenomyia osculata Pont &amp; Werner, epandrium and cercal plate, dorsal view; 36. Xenomyia osculata Pont &amp; Werner, epandrium, cercal plate and surstyli; 37. Xenomyia osculata Pont &amp; Werner, hypandrium and aedeagus, dorsal view; 38. Xenomyia osculata Pont &amp; Werner, hypandrium and aedeagus, lateral view (Figs 3438 modified from Figs 610 of Pont &amp; Werner 2003); Scale bar: 0.5 mm. Figs 3438, not to scale." pageId="13" pageNumber="273">Figs 3438</figureCitation>
(modified from
<figureCitation id="19F649F6FF9F206368664497FC570949" box="[804,916,1519,1545]" captionStart-0="FIGURES 49" captionStart-1="FIGURES 1015" captionStartId-0="3.[151,264,1822,1846]" captionStartId-1="7.[151,264,1874,1898]" captionTargetBox-0="[221,1369,185,1795]" captionTargetBox-1="[199,1388,185,1847]" captionTargetId-0="figure-33@3.[218,1369,181,1798]" captionTargetId-1="figure-33@7.[196,1391,181,1850]" captionTargetPageId-0="3" captionTargetPageId-1="7" captionText-0="FIGURES 49. 4. Xenomyia acuticeps Emden, head of male, lateral view (modified from Fig. 54 of Emden 1951); 5. Xenomyia atra Malloch, head of female, lateral view (modified from Fig. 53 of Emden 1951); 6. Xenomyia atra Malloch, head of male, lateral view (modified from Fig. 52 of Emden 1951, after Jeanneliotis notabilis Séguy); 7. Xenomyia calyptrata Emden, head and thorax of male, lateral view (modified from Fig. 56 of Emden 1951); 8. Xenomyia edwardsi Emden, head of male, lateral view (modified from Fig. 57 of Emden 1951); 9. Xenomyia edwardsi Emden, head of female, lateral view (modified from Fig. 58 of Emden 1951). Scale bars: 1 mm. Fig. 6 not to scale." captionText-1="FIGURES 1015. 10. Xenomyia hirtibasis (Bigot), head of male, lateral view; 11. Xenomyia hirtibasis (Bigot), head of female, lateral view; 12. Xenomyia osculata Pont &amp; Werner, head of male, frontal view (modified from Fig. 1 of Pont &amp; Werner 2003); 13. Xenomyia osculata Pont &amp; Werner, head of female, frontal view (modified from Fig. 2 of Pont &amp; Werner 2003); 14. Xenomyia oxycera Emden, head of male, lateral view (modified from Fig. 55 of Emden 1951); 15. Xenomyia palpalis Emden, head and thorax of female, lateral view (modified from Fig. 4 of Emden 1948). Scale bars: 1 mm. Figs 12, 13 and 15 not to scale." pageId="13" pageNumber="273">Figs 610</figureCitation>
of
<bibRefCitation id="E55C2882FF9F206368FB4497FB660949" author="Werner, D. &amp; Pont, A. C." box="[953,1189,1519,1545]" pageId="13" pageNumber="273" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner 2003</bibRefCitation>
).
</paragraph>
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Female. Head very different from the male with broad frons; fronto-orbital plate broad, covered with small setulae and with 34 pairs of frontal setae; 2 pairs of orbital setae; eyes bare; arista longer and narrower. Wing smoky, but without the area with denser microtrichia. Ovipositor as in
<figureCitation id="19F649F6FF9F206368804723FBD50935" box="[962,1046,1627,1653]" captionStart="FIGURES 3941" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[155,1431,316,1909]" captionTargetId="figure-78@14.[151,1435,309,1913]" captionTargetPageId="14" captionText="FIGURES 3941. 39. Xenomyia hirtibasis (Bigot), ovipositor, dorsal view; 40. Xenomyia hirtibasis (Bigot), ovipositor, lateral view; 41. Xenomyia osculata Pont &amp; Werner, ovipositor, dorsal view (modified from Fig. 11 of Pont &amp; Werner 2003). Scale bars: 0.2 mm. Fig. 41, not to scale." pageId="13" pageNumber="273">Fig. 41</figureCitation>
, modified from
<figureCitation id="19F649F6FF9F20636F934723FAE20935" box="[1233,1313,1627,1653]" captionStart="FIGURES 1015" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[199,1388,185,1847]" captionTargetId="figure-33@7.[196,1391,181,1850]" captionTargetPageId="7" captionText="FIGURES 1015. 10. Xenomyia hirtibasis (Bigot), head of male, lateral view; 11. Xenomyia hirtibasis (Bigot), head of female, lateral view; 12. Xenomyia osculata Pont &amp; Werner, head of male, frontal view (modified from Fig. 1 of Pont &amp; Werner 2003); 13. Xenomyia osculata Pont &amp; Werner, head of female, frontal view (modified from Fig. 2 of Pont &amp; Werner 2003); 14. Xenomyia oxycera Emden, head of male, lateral view (modified from Fig. 55 of Emden 1951); 15. Xenomyia palpalis Emden, head and thorax of female, lateral view (modified from Fig. 4 of Emden 1948). Scale bars: 1 mm. Figs 12, 13 and 15 not to scale." pageId="13" pageNumber="273">Fig. 11</figureCitation>
of
<bibRefCitation id="E55C2882FF9F20636E084723FEEF09D9" author="Werner, D. &amp; Pont, A. C." pageId="13" pageNumber="273" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner 2003</bibRefCitation>
.
</paragraph>
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Notes. Described from male and female from
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.
<collectingMunicipality id="6116CF09FF9F2063682447DBFC1F09FD" box="[870,988,1699,1725]" pageId="13" pageNumber="273">According</collectingMunicipality>
to
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, the species is known only from the middle
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, and is a predator of black-flies.
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deposited in
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View file

@ -0,0 +1,161 @@
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<emphasis id="B3B98961FF9F20636BD5462FFDE10830" bold="true" box="[151,546,1878,1905]" pageId="13" pageNumber="273">
<emphasis id="B3B98961FF9F20636BD5462FFEB40831" bold="true" box="[151,375,1879,1905]" italics="true" pageId="13" pageNumber="273">Xenomyia oxycera</emphasis>
<bibRefCitation id="E55C2882FF9F20636A3D462FFDE10830" author="Emden, F. I. van" box="[383,546,1878,1905]" pageId="13" pageNumber="273" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden, 1951</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9D706F8FF9F207E6BD546E7FC2F0F95" lastPageId="16" lastPageNumber="276" pageId="13" pageNumber="273" type="diagnosis">
<paragraph id="81725573FF9F20606BD546E7FAAF0E5D" blockId="13.[151,1437,1950,2013]" lastBlockId="14.[151,1436,151,285]" lastPageId="14" lastPageNumber="274" pageId="13" pageNumber="273">
Diagnosis. (modified from
<bibRefCitation id="E55C2882FF9F20636A8F46E7FDA308F9" author="Emden, F. I. van" box="[461,608,1950,1977]" pageId="13" pageNumber="273" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
). Length:
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. (
<figureCitation id="19F649F6FF9F2063683746E7FC0A08F9" box="[885,969,1951,1977]" captionStart="FIGURES 1015" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[199,1388,185,1847]" captionTargetId="figure-33@7.[196,1391,181,1850]" captionTargetPageId="7" captionText="FIGURES 1015. 10. Xenomyia hirtibasis (Bigot), head of male, lateral view; 11. Xenomyia hirtibasis (Bigot), head of female, lateral view; 12. Xenomyia osculata Pont &amp; Werner, head of male, frontal view (modified from Fig. 1 of Pont &amp; Werner 2003); 13. Xenomyia osculata Pont &amp; Werner, head of female, frontal view (modified from Fig. 2 of Pont &amp; Werner 2003); 14. Xenomyia oxycera Emden, head of male, lateral view (modified from Fig. 55 of Emden 1951); 15. Xenomyia palpalis Emden, head and thorax of female, lateral view (modified from Fig. 4 of Emden 1948). Scale bars: 1 mm. Figs 12, 13 and 15 not to scale." pageId="13" pageNumber="273">Fig. 14</figureCitation>
, modified from Fig. 55 of
<bibRefCitation id="E55C2882FF9F20636FBE46E7FA5308F9" author="Emden, F. I. van" box="[1276,1424,1950,1977]" pageId="13" pageNumber="273" pagination="325 - 710" refId="ref11152" refString="Emden, F. I. van (1951) n. k. In: Muscidae: C. - Scatophaginae, Anthomyiinae, Lispinae, Fanniinae and Phaoniinae. Ruwenzori Expedition 1934 - 5. 2 (6). British Museum (Natural History), London, pp. 325 - 710, 106 figs, 3 tables." type="book chapter" year="1951">Emden 1951</bibRefCitation>
). Ocellar absent; fronto-orbital plate much broader than frontal vitta, densely covered by short setulae; postpedicel produced and acuminate; proboscis slender and glossy black; antenna brown; palpus dark yellow, long and slender; prosternum bare; superior frontal seta absent; acrostichals present; dorsocentrals 1:2; katepisternals 1:2; presutural intra-alar seta present. Anepimeron bare. Fore tibia without median seta. Hind tibia with an anterodorsal and posterodorsal setae. Claws moderately long, not longer than antenna. Wing smoky. Calypter and halter yellow.
</paragraph>
<caption id="D5B205FBFF9C20606BD546E8FE3B08B0" pageId="14" pageNumber="274" startId="14.[151,264,1936,1960]" targetBox="[155,1431,316,1909]" targetPageId="14" targetType="figure">
<paragraph id="81725573FF9C20606BD546E8FE3B08B0" blockId="14.[151,1436,1936,2032]" pageId="14" pageNumber="274">
<emphasis id="B3B98961FF9C20606BD546E8FE9708E8" bold="true" box="[151,340,1936,1960]" pageId="14" pageNumber="274">FIGURES 3941.</emphasis>
39.
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<emphasis id="B3B98961FF9C20606AC246E9FD8F08E8" box="[384,588,1936,1960]" italics="true" pageId="14" pageNumber="274">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, ovipositor, dorsal view; 40.
<taxonomicName id="46CD2EF0FF9C2060688146E9FB2308E8" authority="(Bigot)" authorityName="" authorityYear="1885" baseAuthorityName="Bigot" baseAuthorityYear="1885" box="[963,1248,1936,1960]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="274" phylum="Arthropoda" rank="species" species="hirtibasis">
<emphasis id="B3B98961FF9C2060688146E9FB4C08E8" box="[963,1167,1936,1960]" italics="true" pageId="14" pageNumber="274">Xenomyia hirtibasis</emphasis>
(Bigot)
</taxonomicName>
, ovipositor, lateral view; 41.
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<emphasis id="B3B98961FF9C20606BBC46CDFE02088C" box="[254,449,1972,1996]" italics="true" pageId="14" pageNumber="274">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, ovipositor, dorsal view (modified from Fig. 11 of
<bibRefCitation id="E55C2882FF9C20606F3A46CCFA90088C" author="Werner, D. &amp; Pont, A. C." box="[1144,1363,1972,1996]" pageId="14" pageNumber="274" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner 2003</bibRefCitation>
). Scale bars: 0.2 mm. Fig. 41, not to scale.
</paragraph>
</caption>
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<paragraph id="81725573FF9D20616BD5460CFDE108B9" blockId="15.[151,1437,1908,2041]" pageId="15" pageNumber="275">
<emphasis id="B3B98961FF9D20616BD5460CFE9108CD" bold="true" box="[151,338,1908,1933]" pageId="15" pageNumber="275">FIGURES 4245.</emphasis>
42.
<taxonomicName id="46CD2EF0FF9D20616A3E460DFD1408CC" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" box="[380,727,1909,1933]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="275" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF9D20616A3E460DFDF808CD" box="[380,571,1909,1933]" italics="true" pageId="15" pageNumber="275">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, male head, lateral view (from
<bibRefCitation id="E55C2882FF9D20616F46460DFB1208CD" author="Werner, D. &amp; Pont, A. C." box="[1028,1233,1909,1933]" pageId="15" pageNumber="275" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner 2003</bibRefCitation>
, with the permission of the publisher); 43.
<taxonomicName id="46CD2EF0FF9D20616A3A46E1FD2308F0" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" box="[376,736,1945,1969]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="275" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF9D20616A3A46E1FDF808F1" box="[376,571,1945,1969]" italics="true" pageId="15" pageNumber="275">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, male, lateral habitus; 44.
<taxonomicName id="46CD2EF0FF9D206168B346E1FA9A08F0" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" box="[1009,1369,1945,1969]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="275" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF9D206168B346E1FB7608F1" box="[1009,1205,1945,1969]" italics="true" pageId="15" pageNumber="275">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, male, dorsal habitus; 45.
<taxonomicName id="46CD2EF0FF9D20616A1546C5FD740894" authority="Pont &amp; Werner" authorityName="Pont &amp; Werner" authorityYear="2003" box="[343,695,1981,2005]" class="Insecta" family="Muscidae" genus="Xenomyia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="275" phylum="Arthropoda" rank="species" species="osculata">
<emphasis id="B3B98961FF9D20616A1546C5FDDB0895" box="[343,536,1981,2005]" italics="true" pageId="15" pageNumber="275">Xenomyia osculata</emphasis>
Pont &amp; Werner
</taxonomicName>
, male, head, frontal view. Figs 4345 modified from Couri &amp; Pont 2000). Scale bars: 2 mm. Fig. 42, not to scale.
</paragraph>
</caption>
<paragraph id="81725573FF82207E6B8541EFFC2F0F95" blockId="16.[151,1437,151,213]" pageId="16" pageNumber="276">
<materialsCitation id="31A55F2EFF82207E6B8541EFFC2F0F95" collectionCode="NHMUK" country="Uganda" location="West Africa" municipality="This" pageId="16" pageNumber="276" specimenCount="1" stateProvince="Jinja" typeStatus="holotype">
Note. Described from male and female from
<collectingCountry id="F9DA15E3FF82207E698741EFFCDC0FF1" box="[709,799,151,177]" name="Uganda" pageId="16" pageNumber="276">Uganda</collectingCountry>
,
<collectingRegion id="43099B91FF82207E686941EFFCA70FF1" box="[811,868,151,177]" country="Uganda" name="Jinja" pageId="16" pageNumber="276">Jinja</collectingRegion>
.
<collectingMunicipality id="6116CF09FF82207E682C41EFFC630FF1" box="[878,928,151,177]" pageId="16" pageNumber="276">This</collectingMunicipality>
species is well known as a predator of black flies in
<location id="841203A8FF82207E6BA941C3FEAC0F95" LSID="urn:lsid:plazi:treatment:0964E465FF9F207E6BD5462FFC2F0F95:841203A8FF82207E6BA941C3FEAC0F95" box="[235,367,187,213]" country="Uganda" municipality="This" name="West Africa" pageId="16" pageNumber="276" stateProvince="Jinja">West Africa</location>
(
<bibRefCitation id="E55C2882FF82207E6A3F41C3FDAF0F95" author="Werner, D. &amp; Pont, A. C." box="[381,620,187,213]" pageId="16" pageNumber="276" pagination="115 - 132" refId="ref11597" refString="Werner, D. &amp; Pont, A. C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 1, 115 - 132. https: // doi. org / 10.1046 / j. 1365 - 2915.2003.00431. x" type="journal article" year="2003">Pont &amp; Werner 2003</bibRefCitation>
).
<typeStatus id="5E76EBD1FF82207E693C41C3FD250F95" box="[638,742,187,213]" pageId="16" pageNumber="276" type="holotype">Holotype</typeStatus>
deposited at
<collectionCode id="E7DCCDB6FF82207E683941C3FC2A0F95" box="[891,1001,187,213]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="276" type="Museum">NHMUK</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,109 @@
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2.
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<emphasis id="879AFD306A74FFB3FF41EC98FE76FF17" bold="true" box="[181,504,151,178]" italics="true" pageId="9" pageNumber="188">Robertsmyia pallidoventris</emphasis>
(
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)
</taxonomicName>
</emphasis>
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<taxonomicName id="72EE5AA16A74FFB3FF63ECD3FDD7FF56" authority="Roberts, 1928: 452" authorityName="Roberts" authorityPageNumber="452" authorityYear="1928" box="[151,601,219,244]" class="Insecta" family="Bombyliidae" genus="Dischistus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="188" phylum="Arthropoda" rank="species" species="pallidoventer">
<emphasis id="879AFD306A74FFB3FF63ECD3FE00FF51" box="[151,398,220,244]" italics="true" pageId="9" pageNumber="188">Dischistus pallidoventer</emphasis>
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<bibRefCitation id="D17F5CD36A74FFB3FE60ECD3FDD7FF56" author="Roberts, F. H. S." box="[404,601,219,244]" pageId="9" pageNumber="188" pagination="413 - 455" refId="ref4624" refString="Roberts, F. H. S. (1928) A revision of the Australian Bombyliidae (Diptera). Part II. The Proceedings of the Linnean Society of New South Wales, 53, 413 - 455." type="journal article" year="1928">Roberts, 1928: 452</bibRefCitation>
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locality:
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(Tas.).
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<emphasis id="879AFD306A74FFB3FF63ED2EFE9CFE9E" bold="true" box="[151,274,289,315]" pageId="9" pageNumber="188">Diagnosis.</emphasis>
Medium-sized species, wing length
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. Pedicel elongate, 2.5× longer than wide, with dorsal apical spine. Wing dark brown on costal third. Cell
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~3.5× longer than wide.
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(NSW, Tas., Vic.) (
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<emphasis id="879AFD306A74FFB3FF33ED81FEB5FE0D" bold="true" box="[199,315,398,424]" pageId="9" pageNumber="188">Remarks.</emphasis>
This species was redescribed by
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: pp. 387388, figs 4850).
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View file

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Key to species of
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1. Wing dark brown on costal third; pedicel elongate, 2.5× as long as wide, with small dorsal apical spine........................................................................................
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Wing membrane almost uniformly subhyaline, only infuscation along the anterior costal margin; pedicel 1.7× as long as wide, without dorsal apical spine.......................................................
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(
<figureCitation id="2DD53DA76A7FFFB8FF6BE81EFE8CFB8E" box="[159,258,1041,1067]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="3.[151,250,1253,1277]" captionStartId-1="4.[151,250,1597,1621]" captionStartId-2="5.[151,250,1430,1454]" captionStartId-3="6.[151,250,1935,1959]" captionStartId-4="7.[151,250,1071,1095]" captionStartId-5="7.[151,250,1982,2006]" captionStartId-6="8.[151,250,1143,1167]" captionStartId-7="8.[151,250,1974,1998]" captionTargetBox-0="[346,1241,379,1223]" captionTargetBox-1="[167,1423,181,1562]" captionTargetBox-2="[151,1435,453,1406]" captionTargetBox-3="[191,1376,196,1911]" captionTargetBox-4="[371,1216,191,1037]" captionTargetBox-5="[476,1112,1156,1959]" captionTargetBox-6="[393,1194,181,1119]" captionTargetBox-7="[363,1226,1228,1948]" captionTargetId-0="figure-93@3.[340,1247,375,1229]" captionTargetId-1="figure-18@4.[165,1423,181,1573]" captionTargetId-2="figure-140@5.[151,1435,453,1406]" captionTargetId-3="figure-18@6.[191,1397,181,1911]" captionTargetId-4="figure-17@7.[361,1226,181,1047]" captionTargetId-5="figure-48@7.[476,1112,1156,1959]" captionTargetId-6="figure-18@8.[393,1194,181,1119]" captionTargetId-7="figure-53@8.[358,1228,1228,1951]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="8" captionText-0="FIGURE 1. Head of Robertsmyia aureola sp. nov., lateral view, measurements. Abbreviations: AL—antennal length; HH— head height; HL—head length; PL—proboscis length." captionText-1="FIGURE 2. Robertsmyia aureola sp. nov. Holotype, male, ANIC 29-020518 (a) habitus, lateral; (b) head and thorax, profile; (c) head, frontal. ♀ (d) mouthparts. Scale bars as shown." captionText-2="FIGURE 3. Robertsmyia aureola sp. nov. Female paratype, ANIC 29-020519 (a) habitus, dorsal; (b) head and thorax, frontal; (c) habitus, lateral; (d) head, profile. Scale bars as shown." captionText-3="FIGURE 4. Robertsmyia aureola sp. nov. Female genitalia. (a) showing sternite 8 (internal, faint), ventral; (b) showing tergite 8 and tergite 9 with acanthophorite spines, dorsal; (c) tergite 9, dorsal; (d) tergite 9, lateral; (e) spermathecal bulbs showing apically invaginated cone (inset), ducting and sperm pumps; (f) furca, dorsal. Scale bars as shown." captionText-4="FIGURE 5. Robertsmyia aureola sp. nov. Male genitalia. Holotype, ANIC 29-020518 (a) genital capsule, dorsal, (b) ventral; (c) epandrium, dorsal; (d) genital capsule, lateral. Scale bars as shown." captionText-5="FIGURE 6. Robertsmyia aureola sp. nov. (a) labellae showing pseudotracheae; (b) antenna, lateral; (c) wing, showing main area of infuscation. Scale bars as shown." captionText-6="FIGURE 7. Wings. (a) Laurella formosa [from Li &amp; Yeates (2018)]; (b) Robertsmyia pallidoventris [from Li &amp; Yeates (2018)]; (c) Robertsmyia aureola sp. nov. Holotype, male, ANIC 29-020518. Scale bars as shown." captionText-7="FIGURE 8. The known distribution records of Robertsmyia species in Australia: R. aureola sp. nov. (red triangle) and R. pallidoventris (blue stars)." figureDoi-0="http://doi.org/10.5281/zenodo.14734783" figureDoi-1="http://doi.org/10.5281/zenodo.14734785" figureDoi-2="http://doi.org/10.5281/zenodo.14734787" figureDoi-3="http://doi.org/10.5281/zenodo.14734791" figureDoi-4="http://doi.org/10.5281/zenodo.14734793" figureDoi-5="http://doi.org/10.5281/zenodo.14734797" figureDoi-6="http://doi.org/10.5281/zenodo.14734799" figureDoi-7="http://doi.org/10.5281/zenodo.14734804" httpUri-0="https://zenodo.org/record/14734783/files/figure.png" httpUri-1="https://zenodo.org/record/14734785/files/figure.png" httpUri-2="https://zenodo.org/record/14734787/files/figure.png" httpUri-3="https://zenodo.org/record/14734791/files/figure.png" httpUri-4="https://zenodo.org/record/14734793/files/figure.png" httpUri-5="https://zenodo.org/record/14734797/files/figure.png" httpUri-6="https://zenodo.org/record/14734799/files/figure.png" httpUri-7="https://zenodo.org/record/14734804/files/figure.png" pageId="2" pageNumber="181">Figs 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FDF472A96A7FFFB8FF63E856FF72FB45" pageId="2" pageNumber="181" type="materials_examined">
<paragraph id="B55121226A7FFFB8FF63E856FF72FB45" blockId="2.[151,1437,1113,2039]" pageId="2" pageNumber="181">
<emphasis id="879AFD306A7FFFB8FF63E856FED4FBD6" bold="true" box="[151,346,1113,1139]" pageId="2" pageNumber="181">Type specimens.</emphasis>
<materialsCitation id="05862B7F6A7FFFB8FE96E856FD83FB3D" collectingDate="2024-04-29" collectionCode="SA, ANIC" collectorName="MA Newton" country="Australia" latitude="-34.06389" location="Cadell" longLatPrecision="19" longitude="139.74333" municipality="MacKintosh Rd." pageId="2" pageNumber="181" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="879AFD306A7FFFB8FE96E856FE41FBD6" bold="true" box="[354,463,1113,1139]" pageId="2" pageNumber="181">
<typeStatus id="6A559F806A7FFFB8FE96E856FE41FBD6" box="[354,463,1113,1139]" pageId="2" pageNumber="181" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="A3E8EAAB6A7FFFB8FE23E855FE60FBD1" box="[471,494,1114,1140]" count="1" pageId="2" pageNumber="181" type="male"></specimenCount>
,
<collectingCountry id="CDF961B26A7FFFB8FE0EE856FDEAFBD6" box="[506,612,1113,1139]" name="Australia" pageId="2" pageNumber="181">Australia</collectingCountry>
,
<emphasis id="879AFD306A7FFFB8FD9BE856FD1DFBD6" bold="true" box="[623,659,1113,1139]" pageId="2" pageNumber="181">
<collectionCode id="D3FFB9E76A7FFFB8FD9BE856FD1DFBD6" box="[623,659,1113,1139]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="2" pageNumber="181">SA</collectionCode>
</emphasis>
,
<location id="B03177F96A7FFFB8FD56E856FD60FBD6" LSID="urn:lsid:plazi:treatment:3D4790346A7FFFBFFF63EFC6FC34F80C:B03177F96A7FFFB8FD56E856FD60FBD6" box="[674,750,1113,1139]" country="Australia" latitude="-34.06389" longLatPrecision="19" longitude="139.74333" municipality="MacKintosh Rd." name="Cadell" pageId="2" pageNumber="181">Cadell</location>
,
<collectingMunicipality id="5535BB586A7FFFB8FD0FE856FC39FBD6" box="[763,951,1113,1139]" pageId="2" pageNumber="181">MacKintosh Rd.</collectingMunicipality>
,
<geoCoordinate id="D0DA47E56A7FFFB8FC30E854FBCAFBD3" box="[964,1092,1115,1142]" degrees="34" direction="south" minutes="03" orientation="latitude" pageId="2" pageNumber="181" precision="15" seconds="50" value="-34.06389">34°0350″S</geoCoordinate>
<geoCoordinate id="D0DA47E56A7FFFB8FBB8E854FB51FBD3" box="[1100,1247,1115,1142]" degrees="139" direction="east" minutes="44" orientation="longitude" pageId="2" pageNumber="181" precision="15" seconds="36" value="139.74333">139°4436″E</geoCoordinate>
,
<date id="C15007E26A7FFFB8FB18E856FAEAFBD6" box="[1260,1380,1113,1139]" pageId="2" pageNumber="181" value="2024-04-29">
<collectingDate id="D114FE0A6A7FFFB8FB18E856FAEAFBD6" box="[1260,1380,1113,1139]" pageId="2" pageNumber="181" value="2024-04-29">29.iv.2024</collectingDate>
</date>
,
<collectorName id="181B44F46A7FFFB8FA85E855FF7EFB3D" pageId="2" pageNumber="181">MA Newton</collectorName>
(
<collectionCode id="D3FFB9E76A7FFFB8FEF6E872FEC6FB32" box="[258,328,1149,1175]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="2" pageNumber="181">ANIC</collectionCode>
No. 29-020518)
</materialsCitation>
.
<materialsCitation id="05862B7F6A7FFFB8FDEFE872FEBFFB1E" collectingDate="2024-04-29" collectionCode="ANIC, SA" collectorName="MA Newton" country="Australia" latitude="-34.06389" location="Cadell" longLatPrecision="19" longitude="139.74333" municipality="MacKintosh Rd." pageId="2" pageNumber="181" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="879AFD306A7FFFB8FDEFE872FD1DFB32" bold="true" box="[539,659,1149,1175]" pageId="2" pageNumber="181">
<typeStatus id="6A559F806A7FFFB8FDEFE872FD1DFB32" box="[539,659,1149,1175]" pageId="2" pageNumber="181" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="A3E8EAAB6A7FFFB8FD69E871FD3CFB3D" box="[669,690,1150,1176]" count="1" pageId="2" pageNumber="181" type="female"></specimenCount>
(5), Same data as holotype (
<collectionCode id="D3FFB9E76A7FFFB8FBF3E872FBC3FB32" box="[1031,1101,1149,1175]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="2" pageNumber="181">ANIC</collectionCode>
No. 29-02051929-020523).
<collectingCountry id="CDF961B26A7FFFB8FF63E8AEFE8FFB1E" box="[151,257,1185,1211]" name="Australia" pageId="2" pageNumber="181">Australia</collectingCountry>
,
<emphasis id="879AFD306A7FFFB8FEF9E8AEFEBFFB1E" bold="true" box="[269,305,1185,1211]" pageId="2" pageNumber="181">
<collectionCode id="D3FFB9E76A7FFFB8FEF9E8AEFEBFFB1E" box="[269,305,1185,1211]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="2" pageNumber="181">SA</collectionCode>
</emphasis>
</materialsCitation>
,
<materialsCitation id="05862B7F6A7FFFB8FEB5E8ADFF76FB45" collectingDate="2024-05-08" collectionCode="ANIC" collectorName="MA Newton." country="Australia" latitude="-34.12111" location="Pooginook" longLatPrecision="19" longitude="140.05917" municipality="Goyder" pageId="2" pageNumber="181" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A3E8EAAB6A7FFFB8FEB5E8ADFED8FB19" box="[321,342,1186,1212]" count="1" pageId="2" pageNumber="181" type="male"></specimenCount>
(1),
<location id="B03177F96A7FFFB8FE7BE8AEFD85FB1E" LSID="urn:lsid:plazi:treatment:3D4790346A7FFFBFFF63EFC6FC34F80C:B03177F96A7FFFB8FE7BE8AEFD85FB1E" box="[399,523,1185,1211]" country="Australia" latitude="-34.12111" longLatPrecision="19" longitude="140.05917" municipality="Goyder" name="Pooginook" pageId="2" pageNumber="181">Pooginook</location>
,
<collectingMunicipality id="5535BB586A7FFFB8FDECE8AEFDE2FB1E" box="[536,620,1185,1211]" pageId="2" pageNumber="181">Goyder</collectingMunicipality>
hwy.,
<geoCoordinate id="D0DA47E56A7FFFB8FD4DE8ACFCB4FB1B" box="[697,826,1187,1214]" degrees="34" direction="south" minutes="07" orientation="latitude" pageId="2" pageNumber="181" precision="15" seconds="16" value="-34.12111">34°0716″S</geoCoordinate>
<geoCoordinate id="D0DA47E56A7FFFB8FCB7E8ACFC58FB1B" box="[835,982,1187,1214]" degrees="140" direction="east" minutes="03" orientation="longitude" pageId="2" pageNumber="181" precision="15" seconds="33" value="140.05917">140°0333″E</geoCoordinate>
,
<date id="C15007E26A7FFFB8FC16E8AEFBC8FB1E" box="[994,1094,1185,1211]" pageId="2" pageNumber="181" value="2024-05-08">
<collectingDate id="D114FE0A6A7FFFB8FC16E8AEFBC8FB1E" box="[994,1094,1185,1211]" pageId="2" pageNumber="181" value="2024-05-08">8.v.2024</collectingDate>
</date>
.
<collectorName id="181B44F46A7FFFB8FBA6E8ADFB69FB19" box="[1106,1255,1186,1212]" pageId="2" pageNumber="181">MA Newton.</collectorName>
(
<collectionCode id="D3FFB9E76A7FFFB8FB02E8AEFAB2FB1E" box="[1270,1340,1185,1211]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="2" pageNumber="181">ANIC</collectionCode>
No. 29- 020524)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FDF472A96A7FFFB8FF33E8E6FD52FAEE" pageId="2" pageNumber="181" type="diagnosis">
<paragraph id="B55121226A7FFFB8FF33E8E6FD52FAEE" blockId="2.[151,1437,1113,2039]" pageId="2" pageNumber="181">
<emphasis id="879AFD306A7FFFB8FF33E8E6FECCFAA6" bold="true" box="[199,322,1257,1283]" pageId="2" pageNumber="181">Diagnosis.</emphasis>
Small-sized species, wing length
<quantity id="72168CC76A7FFFB8FD4BE8E6FCC7FAA1" box="[703,841,1257,1284]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="6.5" metricValueMin="3.3" pageId="2" pageNumber="181" unit="mm" value="4.9" valueMax="6.5" valueMin="3.3">3.36.5 mm</quantity>
. Pedicel moderately elongate, 1.7× longer than wide, without dorsal apical spine. Wing membrane almost uniformly subhyaline, only infuscation along the anterior costal margin. Cell
<emphasis id="879AFD306A7FFFB8FEDEE93EFEC2FAEE" box="[298,332,1329,1355]" italics="true" pageId="2" pageNumber="181">dm</emphasis>
greater than 3.5× longer than wide.
</paragraph>
</subSubSection>
<subSubSection id="FDF472A96A7FFFBEFF33E95AFE11F83C" lastPageId="4" lastPageNumber="183" pageId="2" pageNumber="181" type="description">
<paragraph id="B55121226A7FFFB8FF33E95AFC64FAD5" blockId="2.[151,1437,1113,2039]" box="[199,1002,1365,1392]" pageId="2" pageNumber="181">
<emphasis id="879AFD306A7FFFB8FF33E95AFE2DFACA" bold="true" box="[199,419,1365,1391]" pageId="2" pageNumber="181">Description. Male.</emphasis>
Body length
<quantity id="72168CC76A7FFFB8FDCFE95AFD4AFAD5" box="[571,708,1365,1392]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.9" metricValueMin="3.8" pageId="2" pageNumber="181" unit="mm" value="4.85" valueMax="5.9" valueMin="3.8">3.85.9 mm</quantity>
, wing length
<quantity id="72168CC76A7FFFB8FCAFE95AFC6BFAD5" box="[859,997,1365,1392]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.7" metricValueMin="3.3" pageId="2" pageNumber="181" unit="mm" value="4.5" valueMax="5.7" valueMin="3.3">3.35.7 mm</quantity>
.
</paragraph>
<paragraph id="B55121226A7FFFB8FF33E976FD36F80A" blockId="2.[151,1437,1113,2039]" pageId="2" pageNumber="181">
<emphasis id="879AFD306A7FFFB8FF33E976FE89FA36" bold="true" box="[199,263,1401,1427]" pageId="2" pageNumber="181">Head</emphasis>
. Head ~1.6× wider than long; clothed with dense mid-grey pruinescence, long white and gold hairs, the latter more concentrated dorsally with some adpressed gold scales. Eyes broadly connected by 2.0× length of ocellar tubercle (
<figureCitation id="2DD53DA76A7FFFB8FEF7E9CEFEDDFA7E" box="[259,339,1473,1499]" captionStart="FIGURE 2" captionStartId="4.[151,250,1597,1621]" captionTargetBox="[167,1423,181,1562]" captionTargetId="figure-18@4.[165,1423,181,1573]" captionTargetPageId="4" captionText="FIGURE 2. Robertsmyia aureola sp. nov. Holotype, male, ANIC 29-020518 (a) habitus, lateral; (b) head and thorax, profile; (c) head, frontal. ♀ (d) mouthparts. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734785" httpUri="https://zenodo.org/record/14734785/files/figure.png" pageId="2" pageNumber="181">
Fig. 2
<emphasis id="879AFD306A7FFFB8FEB3E9CDFEDDFA7E" box="[327,339,1474,1499]" italics="true" pageId="2" pageNumber="181">c</emphasis>
</figureCitation>
). Lower frons small and triangular with pale greyish yellow pruinescence, devoid of hairs; mid frons absent; upper frons bordering ocellar triangle (
<figureCitation id="2DD53DA76A7FFFB8FD14E9EAFCBEFA5A" box="[736,816,1509,1535]" captionStart="FIGURE 2" captionStartId="4.[151,250,1597,1621]" captionTargetBox="[167,1423,181,1562]" captionTargetId="figure-18@4.[165,1423,181,1573]" captionTargetPageId="4" captionText="FIGURE 2. Robertsmyia aureola sp. nov. Holotype, male, ANIC 29-020518 (a) habitus, lateral; (b) head and thorax, profile; (c) head, frontal. ♀ (d) mouthparts. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734785" httpUri="https://zenodo.org/record/14734785/files/figure.png" pageId="2" pageNumber="181">
Fig. 2
<emphasis id="879AFD306A7FFFB8FCD6E9EAFCBEFA5A" box="[802,816,1509,1535]" italics="true" pageId="2" pageNumber="181">b</emphasis>
</figureCitation>
). Ocellar tubercle slightly raised, its posterior edge just behind that of the eye margin; black with dark grey pruinescence and numerous blackish setae, variously directed. Face conically produced, 0.8× the length of the scape (
<figureCitation id="2DD53DA76A7FFFB8FD01EA22FCB8F9E2" box="[757,822,1581,1607]" captionStart="FIGURE 1" captionStartId="3.[151,250,1253,1277]" captionTargetBox="[346,1241,379,1223]" captionTargetId="figure-93@3.[340,1247,375,1229]" captionTargetPageId="3" captionText="FIGURE 1. Head of Robertsmyia aureola sp. nov., lateral view, measurements. Abbreviations: AL—antennal length; HH— head height; HL—head length; PL—proboscis length." figureDoi="http://doi.org/10.5281/zenodo.14734783" httpUri="https://zenodo.org/record/14734783/files/figure.png" pageId="2" pageNumber="181">Fig. 1</figureCitation>
); pale greyish-yellow pruinescence; moderately dense, long, whitish-gold hairs; gena vestiture the same and continuous with the face. Occiput with long, gold, reflective hairs and short, gold, reflective, adpressed scales. Posterior eye margin slightly sinuate. Antennae with dense grey pruinescence on all segments (
<figureCitation id="2DD53DA76A7FFFB8FE11EA96FDDFF916" box="[485,593,1689,1716]" captionStart="FIGURE 2" captionStartId="4.[151,250,1597,1621]" captionTargetBox="[167,1423,181,1562]" captionTargetId="figure-18@4.[165,1423,181,1573]" captionTargetPageId="4" captionText="FIGURE 2. Robertsmyia aureola sp. nov. Holotype, male, ANIC 29-020518 (a) habitus, lateral; (b) head and thorax, profile; (c) head, frontal. ♀ (d) mouthparts. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734785" httpUri="https://zenodo.org/record/14734785/files/figure.png" pageId="2" pageNumber="181">
Figs 2
<emphasis id="879AFD306A7FFFB8FDDDEA95FDB9F916" box="[553,567,1690,1715]" italics="true" pageId="2" pageNumber="181">a</emphasis>
,
<emphasis id="879AFD306A7FFFB8FDB7EA96FDDFF916" box="[579,593,1689,1715]" italics="true" pageId="2" pageNumber="181">b</emphasis>
</figureCitation>
). Scape and pedicel both with anteriorly directed, long black hairs; flagellum bare. Scape short, 2.7× longer than wide, slightly wider near apex, 0.3× length of entire antenna. Pedicel 1.7× longer than wide, widest distally, 0.2× length of entire antenna. Flagellum the longest segment, one-segmented, 5.2× longer than wide, widest 0.4× of its length from base, with apical ventral projection and distinctive dorsal spine-like style (
<figureCitation id="2DD53DA76A7FFFB8FF6BEB26FF61F8E6" box="[159,239,1833,1859]" captionStart="FIGURE 6" captionStartId="7.[151,250,1982,2006]" captionTargetBox="[476,1112,1156,1959]" captionTargetId="figure-48@7.[476,1112,1156,1959]" captionTargetPageId="7" captionText="FIGURE 6. Robertsmyia aureola sp. nov. (a) labellae showing pseudotracheae; (b) antenna, lateral; (c) wing, showing main area of infuscation. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734797" httpUri="https://zenodo.org/record/14734797/files/figure.png" pageId="2" pageNumber="181">
Fig. 6
<emphasis id="879AFD306A7FFFB8FF15EB26FF61F8E6" box="[225,239,1833,1859]" italics="true" pageId="2" pageNumber="181">b</emphasis>
</figureCitation>
), 0.4× length of pedicel; pinned specimens show the flagellum to be curved with some lateral compression. Palp one-segmented, slender and elongate, blackish grey, concolourous with antennae (
<figureCitation id="2DD53DA76A7FFFB8FB82EB42FB46F8C2" box="[1142,1224,1869,1895]" captionStart="FIGURE 2" captionStartId="4.[151,250,1597,1621]" captionTargetBox="[167,1423,181,1562]" captionTargetId="figure-18@4.[165,1423,181,1573]" captionTargetPageId="4" captionText="FIGURE 2. Robertsmyia aureola sp. nov. Holotype, male, ANIC 29-020518 (a) habitus, lateral; (b) head and thorax, profile; (c) head, frontal. ♀ (d) mouthparts. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734785" httpUri="https://zenodo.org/record/14734785/files/figure.png" pageId="2" pageNumber="181">
Fig. 2
<emphasis id="879AFD306A7FFFB8FB4EEB42FB46F8C2" box="[1210,1224,1869,1895]" italics="true" pageId="2" pageNumber="181">d</emphasis>
</figureCitation>
). Mouthparts long (extending well beyond oral cavity), robust, 2.0× head length, 2.5× antennae length (
<figureCitation id="2DD53DA76A7FFFB8FB85EB7EFB3BF82E" box="[1137,1205,1905,1931]" captionStart="FIGURE 1" captionStartId="3.[151,250,1253,1277]" captionTargetBox="[346,1241,379,1223]" captionTargetId="figure-93@3.[340,1247,375,1229]" captionTargetPageId="3" captionText="FIGURE 1. Head of Robertsmyia aureola sp. nov., lateral view, measurements. Abbreviations: AL—antennal length; HH— head height; HL—head length; PL—proboscis length." figureDoi="http://doi.org/10.5281/zenodo.14734783" httpUri="https://zenodo.org/record/14734783/files/figure.png" pageId="2" pageNumber="181">Fig. 1</figureCitation>
); labellae large and fleshy with five robust pseudotracheae (
<figureCitation id="2DD53DA76A7FFFB8FDA3EB9AFD29F80A" box="[599,679,1941,1967]" captionStart="FIGURE 6" captionStartId="7.[151,250,1982,2006]" captionTargetBox="[476,1112,1156,1959]" captionTargetId="figure-48@7.[476,1112,1156,1959]" captionTargetPageId="7" captionText="FIGURE 6. Robertsmyia aureola sp. nov. (a) labellae showing pseudotracheae; (b) antenna, lateral; (c) wing, showing main area of infuscation. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734797" httpUri="https://zenodo.org/record/14734797/files/figure.png" pageId="2" pageNumber="181">
Fig. 6
<emphasis id="879AFD306A7FFFB8FD6DEB99FD29F80A" box="[665,679,1942,1967]" italics="true" pageId="2" pageNumber="181">a</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="B55121226A7FFFB9FF33EBB6FEA1FEC0" blockId="2.[151,1437,1113,2039]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="182" pageId="2" pageNumber="181">
<emphasis id="879AFD306A7FFFB8FF33EBB6FEAEF876" bold="true" box="[199,288,1977,2003]" pageId="2" pageNumber="181">Thorax</emphasis>
. Integumental colour of scutum dark bluish black with dull shine and sparse pale brownish grey pruinescence; anterior third with moderately dense long gold hairs and admixed short reflective gold scales. Hairs and scales most prominent peripherally, integument mostly glabrous dorsally. Scutellum integument concolourous with scutum, short curved adpressed golden scales along lateral and posterior margin, mostly glabrous dorsally. Pleura densely ashy pruinose, integument blackish, mostly with sparse long whitish hairs, except anepimeron, meron, laterotergite and mediotergite bare. Leg I coxae mostly cinereous pruinose; leg II coxae distally yellow, basally cinereous pruinose, both with sparse long whitish hairs; leg III coxae integument yellow with sparse long whitish hairs.
</paragraph>
<caption id="E19171AA6A7EFFB9FF63E8EAFD33FA84" ID-DOI="http://doi.org/10.5281/zenodo.14734783" ID-Zenodo-Dep="14734783" httpUri="https://zenodo.org/record/14734783/files/figure.png" pageId="3" pageNumber="182" startId="3.[151,250,1253,1277]" targetBox="[346,1241,379,1223]" targetPageId="3" targetType="figure">
<paragraph id="B55121226A7EFFB9FF63E8EAFD33FA84" blockId="3.[151,1436,1253,1313]" pageId="3" pageNumber="182">
<emphasis id="879AFD306A7EFFB9FF63E8EAFE9BFB58" bold="true" box="[151,277,1253,1277]" pageId="3" pageNumber="182">FIGURE 1.</emphasis>
Head of
<taxonomicName id="72EE5AA16A7EFFB9FE83E8EAFDC0FB58" authorityName="Li" authorityYear="2025" box="[375,590,1253,1277]" class="Insecta" family="Bombyliidae" genus="Robertsmyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="aureola" status="sp. nov.">
<emphasis id="879AFD306A7EFFB9FE83E8EAFDC0FB58" box="[375,590,1253,1277]" italics="true" pageId="3" pageNumber="182">Robertsmyia aureola</emphasis>
</taxonomicName>
<emphasis id="879AFD306A7EFFB9FDA1E8EAFD26FB58" bold="true" box="[597,680,1253,1277]" pageId="3" pageNumber="182">
<taxonomicNameLabel id="9CA9404B6A7EFFB9FDA1E8EAFD26FB58" box="[597,680,1253,1277]" pageId="3" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, lateral view, measurements. Abbreviations: AL—antennal length; HH— head height; HL—head length; PL—proboscis length.
</paragraph>
</caption>
<paragraph id="B55121226A7EFFB9FF33E97CFB1BF9B8" blockId="3.[151,1437,1395,2034]" pageId="3" pageNumber="182">
<emphasis id="879AFD306A7EFFB9FF33E97CFF71FA28" bold="true" box="[199,255,1395,1421]" pageId="3" pageNumber="182">Legs</emphasis>
. Legs mostly yellow, hind femur and tibia darkening distally, tarsi mostly yellow, darkening distally to brown-black, trochanters with ventral black mark. Leg II, III tibiae distinctly spinose, leg I minimally. Leg I, II femora with fine, moderately dense white hairs. Legs with whitish yellow scales, most distinct on femora. Leg bristles mostly short and black, some pale bristles present, especially leg I. Fore tibia 1.6× longer than fore basitarsus, mid tibia 2.3× longer than mid basitarsus, hind tibia 2.0× longer than basitarsus.
</paragraph>
<paragraph id="B55121226A7EFFB9FF33EA28FAC0F908" blockId="3.[151,1437,1395,2034]" pageId="3" pageNumber="182">
<emphasis id="879AFD306A7EFFB9FF33EA28FE9DF9E4" bold="true" box="[199,275,1575,1601]" pageId="3" pageNumber="182">Wings</emphasis>
. Wing membrane uniformly subhyaline with iridescence except for cell
<emphasis id="879AFD306A7EFFB9FB95EA27FBF6F9E4" box="[1121,1144,1576,1601]" italics="true" pageId="3" pageNumber="182">sc</emphasis>
and wing base with thin brownish yellow infuscation (
<figureCitation id="2DD53DA76A7EFFB9FE16EA44FDBEF9C0" box="[482,560,1611,1638]" captionStart="FIGURE 7" captionStartId="8.[151,250,1143,1167]" captionTargetBox="[393,1194,181,1119]" captionTargetId="figure-18@8.[393,1194,181,1119]" captionTargetPageId="8" captionText="FIGURE 7. Wings. (a) Laurella formosa [from Li &amp; Yeates (2018)]; (b) Robertsmyia pallidoventris [from Li &amp; Yeates (2018)]; (c) Robertsmyia aureola sp. nov. Holotype, male, ANIC 29-020518. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734799" httpUri="https://zenodo.org/record/14734799/files/figure.png" pageId="3" pageNumber="182">
Fig. 7
<emphasis id="879AFD306A7EFFB9FDD0EA43FDBEF9C0" box="[548,560,1612,1637]" italics="true" pageId="3" pageNumber="182">c</emphasis>
</figureCitation>
). Cell
<emphasis id="879AFD306A7EFFB9FD8FEA43FD00F9C9" box="[635,654,1612,1644]" italics="true" pageId="3" pageNumber="182">
r
<subScript id="296A23676A7EFFB9FD72EA51FD00F9C9" attach="left" box="[646,654,1630,1644]" fontSize="6" pageId="3" pageNumber="182">5</subScript>
</emphasis>
open; cell
<emphasis id="879AFD306A7EFFB9FCF3EA44FCAEF9C0" box="[775,800,1611,1637]" italics="true" pageId="3" pageNumber="182">br</emphasis>
much longer than cell
<emphasis id="879AFD306A7EFFB9FBE9EA44FBB1F9C0" box="[1053,1087,1611,1637]" italics="true" pageId="3" pageNumber="182">bm</emphasis>
; crossvein
<emphasis id="879AFD306A7EFFB9FB49EA43FB6BF9C0" box="[1213,1253,1612,1637]" italics="true" pageId="3" pageNumber="182">r-m</emphasis>
arising 0.7 from the base of cell
<emphasis id="879AFD306A7EFFB9FEB9EA60FEE1F92C" box="[333,367,1647,1673]" italics="true" pageId="3" pageNumber="182">dm</emphasis>
; crossvein
<emphasis id="879AFD306A7EFFB9FE05EA7FFDADF92C" box="[497,547,1648,1673]" italics="true" pageId="3" pageNumber="182">m-m</emphasis>
long, nearly as long as crossvein
<emphasis id="879AFD306A7EFFB9FC53EA7FFC41F92C" box="[935,975,1648,1673]" italics="true" pageId="3" pageNumber="182">r-m</emphasis>
; cell
<emphasis id="879AFD306A7EFFB9FBE4EA7FFBB6F92C" box="[1040,1080,1648,1673]" italics="true" pageId="3" pageNumber="182">cup</emphasis>
open (
<figureCitation id="2DD53DA76A7EFFB9FB7CEA60FB56F92C" box="[1160,1240,1647,1674]" captionStart="FIGURE 7" captionStartId="8.[151,250,1143,1167]" captionTargetBox="[393,1194,181,1119]" captionTargetId="figure-18@8.[393,1194,181,1119]" captionTargetPageId="8" captionText="FIGURE 7. Wings. (a) Laurella formosa [from Li &amp; Yeates (2018)]; (b) Robertsmyia pallidoventris [from Li &amp; Yeates (2018)]; (c) Robertsmyia aureola sp. nov. Holotype, male, ANIC 29-020518. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734799" httpUri="https://zenodo.org/record/14734799/files/figure.png" pageId="3" pageNumber="182">
Fig. 7
<emphasis id="879AFD306A7EFFB9FB38EA7FFB56F92C" box="[1228,1240,1648,1673]" italics="true" pageId="3" pageNumber="182">c</emphasis>
</figureCitation>
). Alula vestigial, small and narrow. Haltere stem yellowish, darkening brown distally into brown area within yellowish knob.
</paragraph>
<paragraph id="B55121226A7EFFB9FF33EAB8FCADF854" blockId="3.[151,1437,1395,2034]" pageId="3" pageNumber="182">
<emphasis id="879AFD306A7EFFB9FF33EAB8FEB2F974" bold="true" box="[199,316,1719,1745]" pageId="3" pageNumber="182">Abdomen</emphasis>
. Long and narrow, only tapering apically. Integumental colour of tergites black; tergites IIV with lateral border distinctly yellowish orange, diminished or absent on tergites VVIII. Tergite I with moderately dense long gold hairs and some gold scales, similar to anterior scutum. Tergites IIVIII with sparse long fine pale hairs and sparse adpressed gold scales. Sternites with yellowish brown pruinescence and moderately dense whitish-yellow hairs, cream colour distinctly contrasting with tergites.
<emphasis id="879AFD306A7EFFB9FCFFEB48FCFBF8C4" box="[779,885,1863,1889]" italics="true" pageId="3" pageNumber="182">Genitalia</emphasis>
. Epandrium and gonocoxites nearly rectangular, sides slightly concave in distal half; length ~1.2× as long as wide (
<figureCitation id="2DD53DA76A7EFFB9FC9EEB64FC5BF820" box="[874,981,1899,1926]" captionStart="FIGURE 5" captionStartId="7.[151,250,1071,1095]" captionTargetBox="[371,1216,191,1037]" captionTargetId="figure-17@7.[361,1226,181,1047]" captionTargetPageId="7" captionText="FIGURE 5. Robertsmyia aureola sp. nov. Male genitalia. Holotype, ANIC 29-020518 (a) genital capsule, dorsal, (b) ventral; (c) epandrium, dorsal; (d) genital capsule, lateral. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734793" httpUri="https://zenodo.org/record/14734793/files/figure.png" pageId="3" pageNumber="182">
Figs 5
<emphasis id="879AFD306A7EFFB9FC5BEB63FC5BF820" box="[943,981,1900,1925]" italics="true" pageId="3" pageNumber="182">a, c</emphasis>
</figureCitation>
). Hypandrium membranous. Ejaculatory apodeme strong; gonocoxal apodeme short and weak with apex pointed; anterior arm of phallic sheath absent; inner and outer apex of gonocoxite rounded; phallus extends just beyond apex of gonocoxite; gonocoxal fusion seam medium; gonostylus pointed dorsolaterally (
<figureCitation id="2DD53DA76A7EFFB9FD72EBD8FC9DF854" box="[646,787,2007,2034]" captionStart="FIGURE 5" captionStartId="7.[151,250,1071,1095]" captionTargetBox="[371,1216,191,1037]" captionTargetId="figure-17@7.[361,1226,181,1047]" captionTargetPageId="7" captionText="FIGURE 5. Robertsmyia aureola sp. nov. Male genitalia. Holotype, ANIC 29-020518 (a) genital capsule, dorsal, (b) ventral; (c) epandrium, dorsal; (d) genital capsule, lateral. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734793" httpUri="https://zenodo.org/record/14734793/files/figure.png" pageId="3" pageNumber="182">
Figs 5
<emphasis id="879AFD306A7EFFB9FD39EBD7FC9DF854" box="[717,787,2007,2033]" italics="true" pageId="3" pageNumber="182">a, b, d</emphasis>
</figureCitation>
).
</paragraph>
<caption id="E19171AA6A79FFBEFF63EA32FD5BF9DC" ID-DOI="http://doi.org/10.5281/zenodo.14734785" ID-Zenodo-Dep="14734785" httpUri="https://zenodo.org/record/14734785/files/figure.png" pageId="4" pageNumber="183" startId="4.[151,250,1597,1621]" targetBox="[167,1423,181,1562]" targetPageId="4" targetType="figure">
<paragraph id="B55121226A79FFBEFF63EA32FD5BF9DC" blockId="4.[151,1436,1597,1657]" pageId="4" pageNumber="183">
<emphasis id="879AFD306A79FFBEFF63EA32FE9AF9F0" bold="true" box="[151,276,1597,1621]" pageId="4" pageNumber="183">FIGURE 2.</emphasis>
<taxonomicName id="72EE5AA16A79FFBEFEEFEA32FE7EF9F0" authorityName="Li" authorityYear="2025" box="[283,496,1597,1621]" class="Insecta" family="Bombyliidae" genus="Robertsmyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="aureola" status="sp. nov.">
<emphasis id="879AFD306A79FFBEFEEFEA32FE7EF9F0" box="[283,496,1597,1621]" italics="true" pageId="4" pageNumber="183">Robertsmyia aureola</emphasis>
</taxonomicName>
<emphasis id="879AFD306A79FFBEFE0CEA32FDC7F9F0" bold="true" box="[504,585,1597,1621]" pageId="4" pageNumber="183">
<taxonomicNameLabel id="9CA9404B6A79FFBEFE0CEA32FDC7F9F0" box="[504,585,1597,1621]" pageId="4" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, male, ANIC 29-020518 (a) habitus, lateral; (b) head and thorax, profile; (c) head, frontal. ♀ (d) mouthparts. Scale bars as shown.
</paragraph>
</caption>
<paragraph id="B55121226A79FFBEFF33EAC4FE11F83C" blockId="4.[151,1436,1739,2018]" pageId="4" pageNumber="183">
<emphasis id="879AFD306A79FFBEFF33EAC4FEABF940" bold="true" box="[199,293,1739,1765]" pageId="4" pageNumber="183">Female.</emphasis>
Body length
<quantity id="72168CC76A79FFBEFE34EAC4FDC4F943" box="[448,586,1739,1766]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="5.5" metricValueMin="3.3" pageId="4" pageNumber="183" unit="mm" value="4.4" valueMax="5.5" valueMin="3.3">3.35.5 mm</quantity>
, wing length
<quantity id="72168CC76A79FFBEFD10EAC4FCE1F943" box="[740,879,1739,1766]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="6.5" metricValueMin="4.1" pageId="4" pageNumber="183" unit="mm" value="5.3" valueMax="6.5" valueMin="4.1">4.16.5 mm</quantity>
. Female habitus very similar to male (
<figureCitation id="2DD53DA76A79FFBEFAD6EAC4FA1CF940" box="[1314,1426,1739,1766]" captionStart="FIGURE 3" captionStartId="5.[151,250,1430,1454]" captionTargetBox="[151,1435,453,1406]" captionTargetId="figure-140@5.[151,1435,453,1406]" captionTargetPageId="5" captionText="FIGURE 3. Robertsmyia aureola sp. nov. Female paratype, ANIC 29-020519 (a) habitus, dorsal; (b) head and thorax, frontal; (c) habitus, lateral; (d) head, profile. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734787" httpUri="https://zenodo.org/record/14734787/files/figure.png" pageId="4" pageNumber="183">
Figs 3
<emphasis id="879AFD306A79FFBEFA9DEAC3FAF9F940" box="[1385,1399,1740,1765]" italics="true" pageId="4" pageNumber="183">a</emphasis>
,
<emphasis id="879AFD306A79FFBEFA72EAC3FA1CF940" box="[1414,1426,1740,1765]" italics="true" pageId="4" pageNumber="183">c</emphasis>
</figureCitation>
) except frons complete with eyes well separated, ~2× width of ocellar triangle mid frons (
<figureCitation id="2DD53DA76A79FFBEFB8DEAE0FB67F8AC" box="[1145,1257,1775,1802]" captionStart="FIGURE 3" captionStartId="5.[151,250,1430,1454]" captionTargetBox="[151,1435,453,1406]" captionTargetId="figure-140@5.[151,1435,453,1406]" captionTargetPageId="5" captionText="FIGURE 3. Robertsmyia aureola sp. nov. Female paratype, ANIC 29-020519 (a) habitus, dorsal; (b) head and thorax, frontal; (c) habitus, lateral; (d) head, profile. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734787" httpUri="https://zenodo.org/record/14734787/files/figure.png" pageId="4" pageNumber="183">
Figs 3
<emphasis id="879AFD306A79FFBEFB4BEAE0FB43F8AC" box="[1215,1229,1775,1801]" italics="true" pageId="4" pageNumber="183">b</emphasis>
,
<emphasis id="879AFD306A79FFBEFB2FEAE0FB67F8AC" box="[1243,1257,1775,1801]" italics="true" pageId="4" pageNumber="183">d</emphasis>
</figureCitation>
). Tergite 8 with thick, long, fine hairs (
<figureCitation id="2DD53DA76A79FFBEFE6EEB1CFD83F888" box="[410,525,1811,1838]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[191,1376,196,1911]" captionTargetId="figure-18@6.[191,1397,181,1911]" captionTargetPageId="6" captionText="FIGURE 4. Robertsmyia aureola sp. nov. Female genitalia. (a) showing sternite 8 (internal, faint), ventral; (b) showing tergite 8 and tergite 9 with acanthophorite spines, dorsal; (c) tergite 9, dorsal; (d) tergite 9, lateral; (e) spermathecal bulbs showing apically invaginated cone (inset), ducting and sperm pumps; (f) furca, dorsal. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734791" httpUri="https://zenodo.org/record/14734791/files/figure.png" pageId="4" pageNumber="183">
Figs 4
<emphasis id="879AFD306A79FFBEFE16EB1BFE7EF888" box="[482,496,1812,1837]" italics="true" pageId="4" pageNumber="183">a</emphasis>
,
<emphasis id="879AFD306A79FFBEFE0BEB1CFD83F888" box="[511,525,1811,1837]" italics="true" pageId="4" pageNumber="183">b</emphasis>
</figureCitation>
);&gt; 30 acanthophorite spines present in a dense row (
<figureCitation id="2DD53DA76A79FFBEFB9AEB1CFB6EF888" box="[1134,1248,1811,1838]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[191,1376,196,1911]" captionTargetId="figure-18@6.[191,1397,181,1911]" captionTargetPageId="6" captionText="FIGURE 4. Robertsmyia aureola sp. nov. Female genitalia. (a) showing sternite 8 (internal, faint), ventral; (b) showing tergite 8 and tergite 9 with acanthophorite spines, dorsal; (c) tergite 9, dorsal; (d) tergite 9, lateral; (e) spermathecal bulbs showing apically invaginated cone (inset), ducting and sperm pumps; (f) furca, dorsal. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734791" httpUri="https://zenodo.org/record/14734791/files/figure.png" pageId="4" pageNumber="183">
Figs 4
<emphasis id="879AFD306A79FFBEFB42EB1BFB4CF888" box="[1206,1218,1812,1837]" italics="true" pageId="4" pageNumber="183">c</emphasis>
,
<emphasis id="879AFD306A79FFBEFB26EB1CFB6EF888" box="[1234,1248,1811,1837]" italics="true" pageId="4" pageNumber="183">d</emphasis>
</figureCitation>
). Furca sclerites slender, largely parallel, slightly concave, unjointed (
<figureCitation id="2DD53DA76A79FFBEFD00EB38FCCEF8F4" box="[756,832,1847,1873]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[191,1376,196,1911]" captionTargetId="figure-18@6.[191,1397,181,1911]" captionTargetPageId="6" captionText="FIGURE 4. Robertsmyia aureola sp. nov. Female genitalia. (a) showing sternite 8 (internal, faint), ventral; (b) showing tergite 8 and tergite 9 with acanthophorite spines, dorsal; (c) tergite 9, dorsal; (d) tergite 9, lateral; (e) spermathecal bulbs showing apically invaginated cone (inset), ducting and sperm pumps; (f) furca, dorsal. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734791" httpUri="https://zenodo.org/record/14734791/files/figure.png" pageId="4" pageNumber="183">
Fig. 4
<emphasis id="879AFD306A79FFBEFCCCEB38FCCEF8F4" box="[824,832,1847,1873]" italics="true" pageId="4" pageNumber="183">f</emphasis>
</figureCitation>
). Spermathecal bulb large, sclerotized and spherical, with small invaginated apical cone (
<figureCitation id="2DD53DA76A79FFBEFDC7EB54FD0DF8D0" box="[563,643,1883,1909]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[191,1376,196,1911]" captionTargetId="figure-18@6.[191,1397,181,1911]" captionTargetPageId="6" captionText="FIGURE 4. Robertsmyia aureola sp. nov. Female genitalia. (a) showing sternite 8 (internal, faint), ventral; (b) showing tergite 8 and tergite 9 with acanthophorite spines, dorsal; (c) tergite 9, dorsal; (d) tergite 9, lateral; (e) spermathecal bulbs showing apically invaginated cone (inset), ducting and sperm pumps; (f) furca, dorsal. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734791" httpUri="https://zenodo.org/record/14734791/files/figure.png" pageId="4" pageNumber="183">
Fig. 4
<emphasis id="879AFD306A79FFBEFD83EB53FD0DF8D0" box="[631,643,1884,1909]" italics="true" pageId="4" pageNumber="183">e</emphasis>
</figureCitation>
inset). Distal spermathecal duct of normal length; sperm pump short, collars absent (
<figureCitation id="2DD53DA76A79FFBEFEB4EB70FE00F83C" box="[320,398,1919,1945]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[191,1376,196,1911]" captionTargetId="figure-18@6.[191,1397,181,1911]" captionTargetPageId="6" captionText="FIGURE 4. Robertsmyia aureola sp. nov. Female genitalia. (a) showing sternite 8 (internal, faint), ventral; (b) showing tergite 8 and tergite 9 with acanthophorite spines, dorsal; (c) tergite 9, dorsal; (d) tergite 9, lateral; (e) spermathecal bulbs showing apically invaginated cone (inset), ducting and sperm pumps; (f) furca, dorsal. Scale bars as shown." figureDoi="http://doi.org/10.5281/zenodo.14734791" httpUri="https://zenodo.org/record/14734791/files/figure.png" pageId="4" pageNumber="183">
Fig. 4
<emphasis id="879AFD306A79FFBEFE76EB8FFE00F83C" box="[386,398,1920,1945]" italics="true" pageId="4" pageNumber="183">e</emphasis>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FDF472A96A79FFBFFF33EBACFD6EFE2C" lastPageId="5" lastPageNumber="184" pageId="4" pageNumber="183" type="discussion">
<paragraph id="B55121226A79FFBFFF33EBACFD6EFE2C" blockId="4.[151,1436,1739,2018]" lastBlockId="5.[151,1436,151,393]" lastPageId="5" lastPageNumber="184" pageId="4" pageNumber="183">
<emphasis id="879AFD306A79FFBEFF33EBACFEB5F818" bold="true" box="[199,315,1955,1981]" pageId="4" pageNumber="183">Remarks.</emphasis>
<taxonomicName id="72EE5AA16A79FFBEFEB0EBACFDBEF818" authorityName="Li" authorityYear="2025" box="[324,560,1955,1981]" class="Insecta" family="Bombyliidae" genus="Robertsmyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="aureola" status="sp. nov.">
<emphasis id="879AFD306A79FFBEFEB0EBACFDBEF818" box="[324,560,1955,1981]" italics="true" pageId="4" pageNumber="183">Robertsmyia aureola</emphasis>
</taxonomicName>
<emphasis id="879AFD306A79FFBEFDCDEBACFD1AF818" bold="true" box="[569,660,1955,1981]" pageId="4" pageNumber="183">
<taxonomicNameLabel id="9CA9404B6A79FFBEFDCDEBACFD1AF818" box="[569,660,1955,1981]" pageId="4" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be readily distinguished from its congener,
<taxonomicName id="72EE5AA16A79FFBEFB4FEBABFAF6F818" baseAuthorityName="Roberts" baseAuthorityYear="1928" box="[1211,1400,1955,1981]" class="Insecta" family="Bombyliidae" genus="Robertsmyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="pallidoventris">
<emphasis id="879AFD306A79FFBEFB4FEBABFAF6F818" box="[1211,1400,1955,1981]" italics="true" pageId="4" pageNumber="183">R. pallidoventris</emphasis>
</taxonomicName>
by the antennal pedicel being shorter and lacking a dorsal apical spine; wings subhyaline with cell
<emphasis id="879AFD306A79FFBEFB2EEBC7FB7FF844" box="[1242,1265,1992,2017]" italics="true" pageId="4" pageNumber="183">sc</emphasis>
and wing base lightly infuscated brownish. The invaginated cone (in
<taxonomicName id="72EE5AA16A78FFBFFCFCEC98FC0EFF14" authorityName="Li" authorityYear="2025" box="[776,896,151,177]" class="Insecta" family="Bombyliidae" genus="Robertsmyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="aureola" status="sp. nov.">
<emphasis id="879AFD306A78FFBFFCFCEC98FC0EFF14" box="[776,896,151,177]" italics="true" pageId="5" pageNumber="184">R. aureola</emphasis>
</taxonomicName>
<emphasis id="879AFD306A78FFBFFC7DEC98FC68FF14" bold="true" box="[905,998,151,177]" pageId="5" pageNumber="184">
<taxonomicNameLabel id="9CA9404B6A78FFBFFC7DEC98FC68FF14" box="[905,998,151,177]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and apical small conical point (in
<taxonomicName id="72EE5AA16A78FFBFFA70EC98FEBDFF70" baseAuthorityName="Roberts" baseAuthorityYear="1928" class="Insecta" family="Bombyliidae" genus="Robertsmyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="pallidoventris">
<emphasis id="879AFD306A78FFBFFA70EC98FEBDFF70" italics="true" pageId="5" pageNumber="184">R. pallidoventris</emphasis>
</taxonomicName>
) on the sperm bulb might be the same character. It depends on the pressure differences between the inside and the outside of the sperm bulb during dissection. Vesture variations on the dorsal surface of the scutum are observed in different specimens. These variations most likely indicate the age of the specimen as hairs and scales on the dorsal surface are easier to drop off. Some specimens (often males) of
<taxonomicName id="72EE5AA16A78FFBFFBBFED28FB4EFEE4" authorityName="Li" authorityYear="2025" box="[1099,1216,295,321]" class="Insecta" family="Bombyliidae" genus="Robertsmyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="aureola" status="sp. nov.">
<emphasis id="879AFD306A78FFBFFBBFED28FB4EFEE4" box="[1099,1216,295,321]" italics="true" pageId="5" pageNumber="184">R. aureola</emphasis>
</taxonomicName>
<emphasis id="879AFD306A78FFBFFB3CED28FAACFEE4" bold="true" box="[1224,1314,295,321]" pageId="5" pageNumber="184">
<taxonomicNameLabel id="9CA9404B6A78FFBFFB3CED28FAACFEE4" box="[1224,1314,295,321]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
display an unusual-shaped abdomen, which is typically long and narrow, only tapering apically that resembles
<taxonomicName id="72EE5AA16A78FFBFFB11ED44FACBFEC0" authorityName="Hall" authorityYear="1969" box="[1253,1349,331,357]" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="tribe" tribe="Eclimini">Eclimini</taxonomicName>
species according to the key in
<bibRefCitation id="D17F5CD36A78FFBFFE6BED60FDFEFE2C" author="Li, X. &amp; Yeates, D. K." box="[415,624,367,393]" pageId="5" pageNumber="184" pagination="319 - 399" refId="ref4309" refString="Li, X. &amp; Yeates, D. K. (2018) Morphological phylogeny of the Australian genera of the bee fly subfamily Bombyliinae (Diptera: Bombyliidae) with description of four new genera. Invertebrate Systematics, 32 (2), 319 - 399. https: // doi. org / 10.1071 / IS 17039" type="journal article" year="2018">Li &amp; Yeates (2018</bibRefCitation>
: pp. 326).
</paragraph>
</subSubSection>
<caption id="E19171AA6A78FFBFFF63E999FD51FA77" ID-DOI="http://doi.org/10.5281/zenodo.14734787" ID-Zenodo-Dep="14734787" httpUri="https://zenodo.org/record/14734787/files/figure.png" pageId="5" pageNumber="184" startId="5.[151,250,1430,1454]" targetBox="[151,1435,453,1406]" targetPageId="5" targetType="figure">
<paragraph id="B55121226A78FFBFFF63E999FD51FA77" blockId="5.[151,1437,1430,1490]" pageId="5" pageNumber="184">
<emphasis id="879AFD306A78FFBFFF63E999FE9DFA0B" bold="true" box="[151,275,1430,1454]" pageId="5" pageNumber="184">FIGURE 3.</emphasis>
<taxonomicName id="72EE5AA16A78FFBFFEEEE999FE61FA0B" authorityName="Li" authorityYear="2025" box="[282,495,1430,1454]" class="Insecta" family="Bombyliidae" genus="Robertsmyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="aureola" status="sp. nov.">
<emphasis id="879AFD306A78FFBFFEEEE999FE61FA0B" box="[282,495,1430,1454]" italics="true" pageId="5" pageNumber="184">Robertsmyia aureola</emphasis>
</taxonomicName>
<emphasis id="879AFD306A78FFBFFE01E999FDC8FA0B" bold="true" box="[501,582,1430,1454]" pageId="5" pageNumber="184">
<taxonomicNameLabel id="9CA9404B6A78FFBFFE01E999FDC8FA0B" box="[501,582,1430,1454]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female paratype, ANIC 29-020519 (a) habitus, dorsal; (b) head and thorax, frontal; (c) habitus, lateral; (d) head, profile. Scale bars as shown.
</paragraph>
</caption>
<subSubSection id="FDF472A96A78FFBFFF33EA2BFD7DF99B" box="[199,755,1572,1598]" pageId="5" pageNumber="184" type="distribution">
<paragraph id="B55121226A78FFBFFF33EA2BFD7DF99B" blockId="5.[151,1437,1572,1961]" box="[199,755,1572,1598]" pageId="5" pageNumber="184">
<emphasis id="879AFD306A78FFBFFF33EA2BFEEFF99B" bold="true" box="[199,353,1572,1598]" pageId="5" pageNumber="184">Distribution.</emphasis>
Riverland,
<collectingRegion id="772AEFC06A78FFBFFE10EA2BFD1CF99B" box="[484,658,1572,1598]" country="Australia" name="South Australia" pageId="5" pageNumber="184">South Australia</collectingRegion>
(
<figureCitation id="2DD53DA76A78FFBFFD55EA2BFD6BF99B" box="[673,741,1572,1598]" captionStart="FIGURE 8" captionStartId="8.[151,250,1974,1998]" captionTargetBox="[363,1226,1228,1948]" captionTargetId="figure-53@8.[358,1228,1228,1951]" captionTargetPageId="8" captionText="FIGURE 8. The known distribution records of Robertsmyia species in Australia: R. aureola sp. nov. (red triangle) and R. pallidoventris (blue stars)." figureDoi="http://doi.org/10.5281/zenodo.14734804" httpUri="https://zenodo.org/record/14734804/files/figure.png" pageId="5" pageNumber="184">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FDF472A96A78FFBFFF33EA47FC34F80C" pageId="5" pageNumber="184" type="etymology">
<paragraph id="B55121226A78FFBFFF33EA47FD56F923" blockId="5.[151,1437,1572,1961]" pageId="5" pageNumber="184">
<emphasis id="879AFD306A78FFBFFF33EA47FEC0F9C7" bold="true" box="[199,334,1608,1634]" pageId="5" pageNumber="184">Etymology.</emphasis>
The specific epithet is derived from the Latin,
<emphasis id="879AFD306A78FFBFFCA0EA47FC3BF9C7" box="[852,949,1608,1634]" italics="true" pageId="5" pageNumber="184">aureolus</emphasis>
, meaning glittering gold, in reference to the reflective gold scales on the dorsum of this species.
</paragraph>
<paragraph id="B55121226A78FFBFFF33EA9FFC34F80C" blockId="5.[151,1437,1572,1961]" pageId="5" pageNumber="184">
<emphasis id="879AFD306A78FFBFFF33EA9FFE11F90F" bold="true" box="[199,415,1680,1706]" pageId="5" pageNumber="184">Collection details.</emphasis>
During April of 2024, manual sweep-netting along a roadside nature strip (MacKintosh Rd) near Cadell,
<collectingRegion id="772AEFC06A78FFBFFEADEABBFD86F96B" box="[345,520,1716,1742]" country="Australia" name="South Australia" pageId="5" pageNumber="184">South Australia</collectingRegion>
(
<geoCoordinate id="D0DA47E56A78FFBFFDECEAB9FD14F974" box="[536,666,1718,1745]" degrees="34" direction="south" minutes="03" orientation="latitude" pageId="5" pageNumber="184" precision="15" seconds="50" value="-34.06389">34°0350″S</geoCoordinate>
<geoCoordinate id="D0DA47E56A78FFBFFD56EAB9FCBDF974" box="[674,819,1718,1745]" degrees="139" direction="east" minutes="44" orientation="longitude" pageId="5" pageNumber="184" precision="15" seconds="36" value="139.74333">139°4436″E</geoCoordinate>
) of mallee bushes including
<taxonomicName id="72EE5AA16A78FFBFFB8FEABBFA84F96A" box="[1147,1290,1716,1743]" class="Magnoliopsida" family="Sapindaceae" genus="Dodonea" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="879AFD306A78FFBFFB8FEABBFB6CF96B" box="[1147,1250,1716,1742]" italics="true" pageId="5" pageNumber="184">Dodonea</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="72EE5AA16A78FFBFFAEEEABBFA17F96B" box="[1306,1433,1716,1742]" class="Magnoliopsida" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="order">Sapindales</taxonomicName>
:
<taxonomicName id="72EE5AA16A78FFBFFF63EAD7FEA6F957" box="[151,296,1752,1778]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
),
<taxonomicName id="72EE5AA16A78FFBFFECEEAD6FE3AF956" box="[314,436,1753,1779]" class="Magnoliopsida" family="Fabaceae" genus="Senna" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="879AFD306A78FFBFFECEEAD6FEF0F956" box="[314,382,1753,1779]" italics="true" pageId="5" pageNumber="184">Senna</emphasis>
spp.
</taxonomicName>
(
<taxonomicName id="72EE5AA16A78FFBFFE36EAD7FD93F957" box="[450,541,1752,1778]" class="Magnoliopsida" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="order">Fabales</taxonomicName>
:
<taxonomicName id="72EE5AA16A78FFBFFDDCEAD7FD1BF957" box="[552,661,1752,1778]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="184" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
), revealed
<specimenCount id="A3E8EAAB6A78FFBFFCF8EAD7FC18F957" box="[780,918,1752,1778]" count="5" pageId="5" pageNumber="184" type="female">five females</specimenCount>
and
<specimenCount id="A3E8EAAB6A78FFBFFC38EAD6FBBFF957" box="[972,1073,1752,1779]" count="1" pageId="5" pageNumber="184" type="male">one male</specimenCount>
specimen of
<taxonomicName id="72EE5AA16A78FFBFFB33EAD6FAB2F957" authorityName="Li" authorityYear="2025" box="[1223,1340,1752,1778]" class="Insecta" family="Bombyliidae" genus="Robertsmyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="aureola" status="sp. nov.">
<emphasis id="879AFD306A78FFBFFB33EAD6FAB2F957" box="[1223,1340,1752,1778]" italics="true" pageId="5" pageNumber="184">R. aureola</emphasis>
</taxonomicName>
<emphasis id="879AFD306A78FFBFFAB7EAD7FA12F957" bold="true" box="[1347,1436,1752,1778]" pageId="5" pageNumber="184">
<taxonomicNameLabel id="9CA9404B6A78FFBFFAB7EAD7FA12F957" box="[1347,1436,1752,1778]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Capture frequency was low, with only one or
<specimenCount id="A3E8EAAB6A78FFBFFD6AEAF2FCCBF8B3" box="[670,837,1788,1815]" count="2" pageId="5" pageNumber="184" type="generic">two specimens</specimenCount>
being collected on any given day within an hour. As autumn was well underway, very few plants were in flower and insect activity was generally quite low. Further netting on warmer days in
<date id="C15007E26A78FFBFFE3DEB4AFDB3F8FB" box="[457,573,1860,1887]" pageId="5" pageNumber="184" value="2024-05">May 2024</date>
revealed more females with no males. No visual observation of these small flies was ever made before netting. In
<date id="C15007E26A78FFBFFDA7EB66FD45F827" box="[595,715,1896,1923]" pageId="5" pageNumber="184" value="2024-05">May 2024</date>
,
<specimenCount id="A3E8EAAB6A78FFBFFD2CEB66FCB1F827" box="[728,831,1896,1923]" count="1" pageId="5" pageNumber="184" type="male">one male</specimenCount>
specimen was collected in a roadside nature strip of sandy mallee at Pooginook,
<collectingRegion id="772AEFC06A78FFBFFE20EB83FD0FF803" box="[468,641,1932,1958]" country="Australia" name="South Australia" pageId="5" pageNumber="184">South Australia</collectingRegion>
(
<geoCoordinate id="D0DA47E56A78FFBFFD64EB81FC9CF80C" box="[656,786,1934,1961]" degrees="34" direction="south" minutes="07" orientation="latitude" pageId="5" pageNumber="184" precision="15" seconds="16" value="-34.12111">34°0716″S</geoCoordinate>
<geoCoordinate id="D0DA47E56A78FFBFFCEDEB81FC20F80C" box="[793,942,1934,1961]" degrees="140" direction="east" minutes="03" orientation="longitude" pageId="5" pageNumber="184" precision="15" seconds="33" value="140.05917">140°0333″E</geoCoordinate>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,271 @@
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<heading id="796C2065FF80FFDBC5D278B0A227FA81" bold="true" box="[151,582,1238,1265]" fontSize="11" level="1" pageId="7" pageNumber="158" reason="1">
<taxonomicName id="E59BEC8AFF80FFDBC5D278B0A227FA81" authority="Viets, 1940" authorityName="Viets" authorityYear="1940" box="[151,582,1238,1265]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="10EF4B1BFF80FFDBC5D278B0A227FA81" bold="true" box="[151,582,1238,1265]" pageId="7" pageNumber="158">
<emphasis id="10EF4B1BFF80FFDBC5D278B0A1D6FA80" bold="true" box="[151,439,1238,1265]" italics="true" pageId="7" pageNumber="158">Simognathus adriaticus</emphasis>
<bibRefCitation id="460AEAF8FF80FFDBC4FB78B0A227FA81" author="Viets, K." box="[446,582,1238,1264]" pageId="7" pageNumber="158" pagination="1 - 135" refId="ref9235" refString="Viets, K. (1940) Meeresmilben aus der Adria (Halacaridae und Hydrachnellae, Acari). Archiv fur Naturgeschichte, Neue Folge, 9, 1 - 135." type="journal article" year="1940">Viets, 1940</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="6A81C482FF80FFDBC5D2789DA1BDFB64" box="[151,476,1274,1301]" pageId="7" pageNumber="158" type="description">
<paragraph id="22249709FF80FFDBC5D2789DA1BDFB64" blockId="7.[151,582,1238,1301]" box="[151,476,1274,1301]" pageId="7" pageNumber="158">
<figureCitation id="BAA08B8CFF80FFDBC5D2789DA14FFB64" box="[151,302,1274,1301]" captionStart="FIGURE 5" captionStartId="8.[151,250,1797,1821]" captionTargetBox="[167,1427,195,1759]" captionTargetId="figure-27@8.[151,1435,181,1775]" captionTargetPageId="8" captionText="FIGURE 5. Simognathus adriaticus Viets, 1940: Line art illustrations of deutonymph (AG) (FDHAL-24/110) and protonymph (H, I) (FDHAL-24/112) A Idiosoma, dorsal view B Idiosoma, ventral view C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734735" httpUri="https://zenodo.org/record/14734735/files/figure.png" pageId="7" pageNumber="158">Figures 5AI</figureCitation>
,
<figureCitation id="BAA08B8CFF80FFDBC47F789DA11FFB64" box="[314,382,1275,1301]" captionStart="FIGURE 6" captionStartId="9.[151,250,1512,1536]" captionTargetBox="[264,1322,181,1487]" captionTargetId="figure-16@9.[264,1323,181,1488]" captionTargetPageId="9" captionText="FIGURE 6. Acaromantis monnioti Morselli, 1970 (AC) (FDHAL-24/107) &amp; Simognathus adriaticus Viets, 1940 (D, E) (FDHAL-24/110,111): Light-microscope images of habitus dorsal deutonymph (A) habitus dorsal larva (B) and ventral view of quiescent pupal stage larva (C) habitus dorsal deutonymph (D) habitus dorsal protonymph (E) Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14734737" httpUri="https://zenodo.org/record/14734737/files/figure.png" pageId="7" pageNumber="158">6D, E</figureCitation>
;
<tableCitation id="6F19A2B2FF80FFDBC4CC789DA1BDFB64" box="[393,476,1275,1301]" captionStart="TABLE 3" captionStartId="10.[152,244,151,175]" captionText="TABLE 3. Measurements for Simognathus adriaticus (in micrometers)." pageId="7" pageNumber="158">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="6A81C482FF80FFDBC5D27923A29CFB2C" box="[151,765,1349,1373]" pageId="7" pageNumber="158" type="reference_group">
<paragraph id="22249709FF80FFDBC5D27923A29CFB2C" blockId="7.[151,765,1349,1373]" box="[151,765,1349,1373]" pageId="7" pageNumber="158">
<treatmentCitationGroup id="028BB027FF80FFDBC5D27923A29CFB2C" box="[151,765,1349,1373]" pageId="7" pageNumber="158">
[
<treatmentCitation id="A33AB118FF80FFDBC5E57923A238FB2C" author="Viets, K." box="[160,601,1349,1373]" page="97" pageId="7" pageNumber="158" year="1940">
<taxonomicName id="E59BEC8AFF80FFDBC5E57923A238FB2C" authority="Viets, 1940: 95 - 97" authorityName="Viets" authorityPageNumber="95 - 97" authorityYear="1940" box="[160,601,1349,1373]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="10EF4B1BFF80FFDBC5E57923A1F1FB2C" box="[160,400,1349,1373]" italics="true" pageId="7" pageNumber="158">Simognathus adriaticus</emphasis>
<bibRefCitation id="460AEAF8FF80FFDBC4D37923A238FB2C" author="Viets, K." box="[406,601,1349,1373]" pageId="7" pageNumber="158" pagination="1 - 135" refId="ref9235" refString="Viets, K. (1940) Meeresmilben aus der Adria (Halacaridae und Hydrachnellae, Acari). Archiv fur Naturgeschichte, Neue Folge, 9, 1 - 135." type="journal article" year="1940">Viets, 1940: 9597</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; Figs. 143149]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6A81C482FF80FFDBC5D279EDA2DFF99A" pageId="7" pageNumber="158" type="materials_examined">
<paragraph id="22249709FF80FFDBC5D279EDA0B6F862" blockId="7.[151,1437,1419,2027]" pageId="7" pageNumber="158">
<emphasis id="10EF4B1BFF80FFDBC5D279EDA1E0FBD4" bold="true" box="[151,385,1419,1445]" pageId="7" pageNumber="158">Material examined.</emphasis>
<materialsCitation id="92F39D54FF80FFDBC4C279EAA3E7FBBB" collectingDate="2019-09-30" elevation="5" latitude="39.34206" location="Ayvalik" longLatPrecision="1" longitude="26.71007" municipality="Protonymph" pageId="7" pageNumber="158" specimenCount="1" stateProvince="Balikesir">
<collectingMunicipality id="C2400D73FF80FFDBC4C279EAA270FBD7" box="[391,529,1420,1446]" pageId="7" pageNumber="158">Protonymph</collectingMunicipality>
(FDHAL-24/112), undissected and slide mounted, St.1 (
<location id="2744C1D2FF80FFDBC1C879EAA488FBD7" LSID="urn:lsid:plazi:treatment:AA32261FFF80FFD5C5D278B0A2BFF9A5:2744C1D2FF80FFDBC1C879EAA488FBD7" box="[1165,1257,1420,1446]" latitude="39.34206" longLatPrecision="1" longitude="26.71007" municipality="Protonymph" name="Ayvalik" pageId="7" pageNumber="158" stateProvince="Balikesir">Ayvalik</location>
,
<collectingRegion id="E05F59EBFF80FFDBC1B179EAA53FFBD7" box="[1268,1374,1420,1446]" country="Turkey" name="Balikesir" pageId="7" pageNumber="158">Balikesir</collectingRegion>
), fine sand with
<emphasis id="10EF4B1BFF80FFDBC44879D7A1EFFBBB" box="[269,398,1457,1482]" italics="true" pageId="7" pageNumber="158">P. oceanica</emphasis>
,
<quantity id="E5633AECFF80FFDBC4DB79D6A1ADFBBA" box="[414,460,1456,1483]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="158" unit="m" value="5.0">
<elevation id="A9B6703AFF80FFDBC4DB79D6A1ADFBBA" box="[414,460,1456,1483]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="158" unit="m" value="5.0">5 m</elevation>
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,
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,
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,
<date id="5625B1C9FF80FFDBC7A079D6A3E7FBBB" box="[741,902,1456,1482]" pageId="7" pageNumber="158" value="2019-09-30">
<collectingDate id="46614821FF80FFDBC7A079D6A3E7FBBB" box="[741,902,1456,1482]" pageId="7" pageNumber="158" value="2019-09-30">30 Sept. 2019</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="92F39D54FF80FFDBC6D479D7A0B2F862" collectingDate="2019-06-22" elevation="2" latitude="36.84936" location="Kundu" longLatPrecision="1" longitude="30.839" municipality="Two" pageId="7" pageNumber="158" specimenCount="1" stateProvince="Antalya">
<collectingMunicipality id="C2400D73FF80FFDBC6D479D7A3A3FBBA" box="[913,962,1457,1483]" pageId="7" pageNumber="158">Two</collectingMunicipality>
deutonymphs (FDHAL-24/110, FDHAL-24/111) undissected and slide mounted, St. 5 (
<location id="2744C1D2FF80FFDBC7EA79B2A29FFB9F" LSID="urn:lsid:plazi:treatment:AA32261FFF80FFD5C5D278B0A2BFF9A5:2744C1D2FF80FFDBC7EA79B2A29FFB9F" box="[687,766,1492,1518]" latitude="36.84936" longLatPrecision="1" longitude="30.839" municipality="Two" name="Kundu" pageId="7" pageNumber="158" stateProvince="Antalya">Kundu</location>
,
<collectingRegion id="E05F59EBFF80FFDBC64F79B2A306FB9F" box="[778,871,1492,1518]" country="Turkey" name="Antalya" pageId="7" pageNumber="158">Antalya</collectingRegion>
), fine sand,
<quantity id="E5633AECFF80FFDBC6B379B2A444FB9E" box="[1014,1061,1492,1519]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="158" unit="m" value="2.0">
<elevation id="A9B6703AFF80FFDBC6B379B2A444FB9E" box="[1014,1061,1492,1519]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="158" unit="m" value="2.0">2 m</elevation>
</quantity>
,
<geoCoordinate id="47AFF1CEFF80FFDBC17179B2A4CFFB9F" box="[1076,1198,1492,1518]" degrees="36.849361" direction="north" orientation="latitude" pageId="7" pageNumber="158" precision="1" value="36.84936">36.849361</geoCoordinate>
,
<geoCoordinate id="47AFF1CEFF80FFDBC1F979B2A557FB9F" box="[1212,1334,1492,1518]" degrees="30.839000" direction="east" orientation="longitude" pageId="7" pageNumber="158" precision="1" value="30.839">30.839000</geoCoordinate>
,
<date id="5625B1C9FF80FFDBC00679B2A0B2F862" pageId="7" pageNumber="158" value="2019-06-22">
<collectingDate id="46614821FF80FFDBC00679B2A0B2F862" pageId="7" pageNumber="158" value="2019-06-22">22 June 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="22249709FF80FFDBC5827A7BA1E2F928" blockId="7.[151,1437,1419,2027]" pageId="7" pageNumber="158">
<emphasis id="10EF4B1BFF80FFDBC5827A7BA2E5F846" bold="true" box="[199,644,1564,1591]" pageId="7" pageNumber="158">
Morphology and notes.
<emphasis id="10EF4B1BFF80FFDBC4987A7AA21CF846" bold="true" box="[477,637,1564,1591]" italics="true" pageId="7" pageNumber="158">Deutonymphs</emphasis>
.
</emphasis>
Dorsal plates uniformly foveate (
<figureCitation id="BAA08B8CFF80FFDBC6B17A7BA42CF849" box="[1012,1101,1565,1592]" captionStart="FIGURE 5" captionStartId="8.[151,250,1797,1821]" captionTargetBox="[167,1427,195,1759]" captionTargetId="figure-27@8.[151,1435,181,1775]" captionTargetPageId="8" captionText="FIGURE 5. Simognathus adriaticus Viets, 1940: Line art illustrations of deutonymph (AG) (FDHAL-24/110) and protonymph (H, I) (FDHAL-24/112) A Idiosoma, dorsal view B Idiosoma, ventral view C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734735" httpUri="https://zenodo.org/record/14734735/files/figure.png" pageId="7" pageNumber="158">Fig. 5A</figureCitation>
).AD with a transverse hyaline lens at anterior end. Posterior to hyaline lens, area of AD foveate. Dorsal setae small. AD anteriorly truncated, posteriorly rounded., bearing ds-1. OC triangular, narrowing posteriorly. Pair of ds-2 to ds-4 striated in integument. Pair of ds-5 and ds-6 on
<collectionCode id="448A0FCCFF80FFDBC4F67AECA1BBF8D5" box="[435,474,1674,1700]" country="Netherlands" name="Dutch Plant Protection Service, Culture Collection of Plant Pathogenic Bacteria" pageId="7" pageNumber="158">PD</collectionCode>
. All ventral plates separated and foveated (
<figureCitation id="BAA08B8CFF80FFDBC6807AEFA47CF8D5" box="[965,1053,1673,1700]" captionStart="FIGURE 5" captionStartId="8.[151,250,1797,1821]" captionTargetBox="[167,1427,195,1759]" captionTargetId="figure-27@8.[151,1435,181,1775]" captionTargetPageId="8" captionText="FIGURE 5. Simognathus adriaticus Viets, 1940: Line art illustrations of deutonymph (AG) (FDHAL-24/110) and protonymph (H, I) (FDHAL-24/112) A Idiosoma, dorsal view B Idiosoma, ventral view C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734735" httpUri="https://zenodo.org/record/14734735/files/figure.png" pageId="7" pageNumber="158">Fig. 5B</figureCitation>
) AE with three pairs of setae and a pair of large epimeral vesicles. Posterior part of AE truncate. PE with a dorsal and three ventral setae. Anterior part of GA truncate. GA with two pairs of gac and one pair of pgs, no sgs (
<figureCitation id="BAA08B8CFF80FFDBC6BE7AB4A45EF89C" box="[1019,1087,1746,1773]" captionStart="FIGURE 5" captionStartId="8.[151,250,1797,1821]" captionTargetBox="[167,1427,195,1759]" captionTargetId="figure-27@8.[151,1435,181,1775]" captionTargetPageId="8" captionText="FIGURE 5. Simognathus adriaticus Viets, 1940: Line art illustrations of deutonymph (AG) (FDHAL-24/110) and protonymph (H, I) (FDHAL-24/112) A Idiosoma, dorsal view B Idiosoma, ventral view C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734735" httpUri="https://zenodo.org/record/14734735/files/figure.png" pageId="7" pageNumber="158">Fig. 5</figureCitation>
B') Gnathosomal base foveate. Palps three segmented (
<figureCitation id="BAA08B8CFF80FFDBC4E77A90A19AF960" box="[418,507,1782,1809]" captionStart="FIGURE 5" captionStartId="8.[151,250,1797,1821]" captionTargetBox="[167,1427,195,1759]" captionTargetId="figure-27@8.[151,1435,181,1775]" captionTargetPageId="8" captionText="FIGURE 5. Simognathus adriaticus Viets, 1940: Line art illustrations of deutonymph (AG) (FDHAL-24/110) and protonymph (H, I) (FDHAL-24/112) A Idiosoma, dorsal view B Idiosoma, ventral view C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734735" httpUri="https://zenodo.org/record/14734735/files/figure.png" pageId="7" pageNumber="158">Fig. 5C</figureCitation>
). Surface of all telofemora foveated. Leg chaetotaxy as follows (bipectinated setae between parentheses) (
<figureCitation id="BAA08B8CFF80FFDBC4D27B7DA241F944" box="[407,544,1818,1845]" captionStart="FIGURE 5" captionStartId="8.[151,250,1797,1821]" captionTargetBox="[167,1427,195,1759]" captionTargetId="figure-27@8.[151,1435,181,1775]" captionTargetPageId="8" captionText="FIGURE 5. Simognathus adriaticus Viets, 1940: Line art illustrations of deutonymph (AG) (FDHAL-24/110) and protonymph (H, I) (FDHAL-24/112) A Idiosoma, dorsal view B Idiosoma, ventral view C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734735" httpUri="https://zenodo.org/record/14734735/files/figure.png" pageId="7" pageNumber="158">Figs. 5DG</figureCitation>
); Leg I: 1, 2, 1, 4, 5, 6; Leg
<collectionCode id="448A0FCCFF80FFDBC61E7B7DA310F944" box="[859,881,1819,1845]" pageId="7" pageNumber="158">II</collectionCode>
: 1, 1, 2, 4, 5(2), 6; Leg
<collectionCode id="448A0FCCFF80FFDBC13E7B7DA4FAF944" box="[1147,1179,1819,1845]" pageId="7" pageNumber="158">III</collectionCode>
: 1, 1, 2, 3, 5(2), 6; Leg
<collectionCode id="448A0FCCFF80FFDBC5D27B59A0D6F928" box="[151,183,1855,1881]" pageId="7" pageNumber="158">IV</collectionCode>
: 1, 1, 1, 3, 5(2), 6.
</paragraph>
<paragraph id="22249709FF80FFDBC5827B05A2DFF99A" blockId="7.[151,1437,1419,2027]" pageId="7" pageNumber="158">
<emphasis id="10EF4B1BFF80FFDBC5827B05A13CF90F" bold="true" box="[199,349,1891,1918]" italics="true" pageId="7" pageNumber="158">Protonymph.</emphasis>
All plates are smaller than deutonymphs (
<figureCitation id="BAA08B8CFF80FFDBC6247B05A380F90C" box="[865,993,1891,1918]" captionStart="FIGURE 5" captionStartId="8.[151,250,1797,1821]" captionTargetBox="[167,1427,195,1759]" captionTargetId="figure-27@8.[151,1435,181,1775]" captionTargetPageId="8" captionText="FIGURE 5. Simognathus adriaticus Viets, 1940: Line art illustrations of deutonymph (AG) (FDHAL-24/110) and protonymph (H, I) (FDHAL-24/112) A Idiosoma, dorsal view B Idiosoma, ventral view C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734735" httpUri="https://zenodo.org/record/14734735/files/figure.png" pageId="7" pageNumber="158">Figs 5H, I</figureCitation>
). Number and arrangement of dorsal idiosomatic setae same as in deutonymph. Epimeral vesicles clearly seen (
<figureCitation id="BAA08B8CFF80FFDBC1457BE1A433F9D3" box="[1024,1106,1927,1954]" captionStart="FIGURE 5" captionStartId="8.[151,250,1797,1821]" captionTargetBox="[167,1427,195,1759]" captionTargetId="figure-27@8.[151,1435,181,1775]" captionTargetPageId="8" captionText="FIGURE 5. Simognathus adriaticus Viets, 1940: Line art illustrations of deutonymph (AG) (FDHAL-24/110) and protonymph (H, I) (FDHAL-24/112) A Idiosoma, dorsal view B Idiosoma, ventral view C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734735" httpUri="https://zenodo.org/record/14734735/files/figure.png" pageId="7" pageNumber="158">Fig. 5I</figureCitation>
). Leg chaetotaxy as follows (bipectinated setae between parentheses); Leg I: 1, 2, 1, 4, 5, 6; Leg
<collectionCode id="448A0FCCFF80FFDBC6D67BCAA3C8F9B7" box="[915,937,1964,1990]" pageId="7" pageNumber="158">II</collectionCode>
: 1, 1, 2, 4, 5(2), 6; Leg
<collectionCode id="448A0FCCFF80FFDBC1F57BCAA4B1F9B7" box="[1200,1232,1964,1990]" pageId="7" pageNumber="158">III</collectionCode>
: 1, 1, 2, 3, 5(2), 6; Leg
<collectionCode id="448A0FCCFF80FFDBC58F7BB7A08BF99A" box="[202,234,2001,2027]" pageId="7" pageNumber="158">IV</collectionCode>
: 0, 1+2 (basifemur+telofemur), 2, 5(2), 5.
</paragraph>
</subSubSection>
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<paragraph id="22249709FF8FFFD4C5D27B63A179F9FB" blockId="8.[151,1436,1797,1930]" pageId="8" pageNumber="159">
<emphasis id="10EF4B1BFF8FFFD4C5D27B63A173F96C" bold="true" box="[151,274,1797,1821]" pageId="8" pageNumber="159">FIGURE 5.</emphasis>
<taxonomicName id="E59BEC8AFF8FFFD4C4527B63A2E1F96C" authority="Viets, 1940" authorityName="Viets" authorityYear="1940" box="[279,640,1797,1821]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="10EF4B1BFF8FFFD4C4527B63A267F96C" box="[279,518,1797,1821]" italics="true" pageId="8" pageNumber="159">Simognathus adriaticus</emphasis>
<bibRefCitation id="460AEAF8FF8FFFD4C74F7B63A2E1F96C" author="Viets, K." box="[522,640,1797,1821]" pageId="8" pageNumber="159" pagination="1 - 135" refId="ref9235" refString="Viets, K. (1940) Meeresmilben aus der Adria (Halacaridae und Hydrachnellae, Acari). Archiv fur Naturgeschichte, Neue Folge, 9, 1 - 135." type="journal article" year="1940">Viets, 1940</bibRefCitation>
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: Line art illustrations of deutonymph
<emphasis id="10EF4B1BFF8FFFD4C6B37B63A458F96C" bold="true" box="[1014,1081,1797,1821]" pageId="8" pageNumber="159">(AG)</emphasis>
(FDHAL-24/110) and protonymph
<emphasis id="10EF4B1BFF8FFFD4C5D27B4FA0B2F930" bold="true" box="[151,211,1833,1857]" pageId="8" pageNumber="159">(H, I)</emphasis>
(FDHAL-24/112)
<emphasis id="10EF4B1BFF8FFFD4C4D17B4FA1C4F930" bold="true" box="[404,421,1833,1857]" pageId="8" pageNumber="159">A</emphasis>
Idiosoma, dorsal view
<emphasis id="10EF4B1BFF8FFFD4C7D17B4FA2C4F930" bold="true" box="[660,677,1833,1857]" pageId="8" pageNumber="159">B</emphasis>
Idiosoma, ventral view
<emphasis id="10EF4B1BFF8FFFD4C6D87B4FA3CEF930" bold="true" box="[925,943,1833,1857]" pageId="8" pageNumber="159">C</emphasis>
Gnathosoma, lateral view
<emphasis id="10EF4B1BFF8FFFD4C1877B4FA4B5F930" bold="true" box="[1218,1236,1833,1857]" pageId="8" pageNumber="159">D</emphasis>
Leg I, medial view
<emphasis id="10EF4B1BFF8FFFD4C5D27B2BA0C9F914" bold="true" box="[151,168,1869,1893]" pageId="8" pageNumber="159">E</emphasis>
Leg II, medial view
<emphasis id="10EF4B1BFF8FFFD4C43B7B2BA1ECF914" bold="true" box="[382,397,1869,1893]" pageId="8" pageNumber="159">F</emphasis>
Leg III, medial view
<emphasis id="10EF4B1BFF8FFFD4C7287B2BA2E0F914" bold="true" box="[621,641,1869,1893]" pageId="8" pageNumber="159">G</emphasis>
Leg IV, medial view
<emphasis id="10EF4B1BFF8FFFD4C6187B2BA310F914" bold="true" box="[861,881,1869,1893]" pageId="8" pageNumber="159">H</emphasis>
Idiosoma, dorsal view
<emphasis id="10EF4B1BFF8FFFD4C1257B2BA40BF914" bold="true" box="[1120,1130,1869,1893]" pageId="8" pageNumber="159">I</emphasis>
Idiosoma, ventral view. Scale bars: 50 µm.
</paragraph>
</caption>
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<paragraph id="22249709FF8EFFD5C5D2798EA542F839" blockId="9.[151,1436,1512,1608]" pageId="9" pageNumber="160">
<emphasis id="10EF4B1BFF8EFFD5C5D2798EA177F871" bold="true" box="[151,278,1512,1536]" pageId="9" pageNumber="160">FIGURE 6.</emphasis>
<taxonomicName id="E59BEC8AFF8EFFD5C464798EA2C9F871" authority="Morselli, 1970" authorityName="Morselli" authorityYear="1970" box="[289,680,1512,1536]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="monnioti">
<emphasis id="10EF4B1BFF8EFFD5C464798EA264F871" box="[289,517,1512,1536]" italics="true" pageId="9" pageNumber="160">Acaromantis monnioti</emphasis>
<bibRefCitation id="460AEAF8FF8EFFD5C74A798EA2C9F871" author="Morselli, I." box="[527,680,1512,1536]" pageId="9" pageNumber="160" pagination="109 - 119" refId="ref8592" refString="Morselli, I. (1970) Ricerche sugli Alacaridi delle coste livornesi. II. - Acaromantis monnioti, una nuova specie raccoltasui fondi sabbiosi delle Secche della 206 Meloria (Livorno). Pubblicazioni della Stazione Zoologica di Napoli, 38, 109 - 119." type="journal article" year="1970">Morselli, 1970</bibRefCitation>
</taxonomicName>
<emphasis id="10EF4B1BFF8EFFD5C7F7798EA295F871" bold="true" box="[690,756,1512,1536]" pageId="9" pageNumber="160">(AC)</emphasis>
(FDHAL-24/107) &amp;
<taxonomicName id="E59BEC8AFF8EFFD5C69F798EA52FF871" authority="Viets, 1940" authorityName="Viets" authorityYear="1940" box="[986,1358,1512,1536]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="10EF4B1BFF8EFFD5C69F798EA4ACF871" box="[986,1229,1512,1536]" italics="true" pageId="9" pageNumber="160">Simognathus adriaticus</emphasis>
<bibRefCitation id="460AEAF8FF8EFFD5C192798EA52FF871" author="Viets, K." box="[1239,1358,1512,1536]" pageId="9" pageNumber="160" pagination="1 - 135" refId="ref9235" refString="Viets, K. (1940) Meeresmilben aus der Adria (Halacaridae und Hydrachnellae, Acari). Archiv fur Naturgeschichte, Neue Folge, 9, 1 - 135." type="journal article" year="1940">Viets, 1940</bibRefCitation>
</taxonomicName>
<emphasis id="10EF4B1BFF8EFFD5C01D798EA5FDF871" bold="true" box="[1368,1436,1512,1536]" pageId="9" pageNumber="160">(D, E)</emphasis>
(FDHAL-24/110,111): Light-microscope images of habitus dorsal deutonymph
<emphasis id="10EF4B1BFF8EFFD5C6F97A6AA381F855" bold="true" box="[956,992,1548,1572]" pageId="9" pageNumber="160">(A)</emphasis>
habitus dorsal larva
<emphasis id="10EF4B1BFF8EFFD5C1F47A6AA4B2F855" bold="true" box="[1201,1235,1548,1572]" pageId="9" pageNumber="160">(B)</emphasis>
and ventral view of quiescent pupal stage larva
<emphasis id="10EF4B1BFF8EFFD5C4EA7A56A1B3F839" bold="true" box="[431,466,1584,1608]" pageId="9" pageNumber="160">(C)</emphasis>
habitus dorsal deutonymph
<emphasis id="10EF4B1BFF8EFFD5C7B47A56A374F839" bold="true" box="[753,789,1584,1608]" pageId="9" pageNumber="160">(D)</emphasis>
habitus dorsal protonymph
<emphasis id="10EF4B1BFF8EFFD5C1747A56A432F839" bold="true" box="[1073,1107,1584,1608]" pageId="9" pageNumber="160">(E)</emphasis>
Scale bars: 100 µm.
</paragraph>
</caption>
<subSubSection id="6A81C482FF8EFFD5C5827A10A2BFF9A5" pageId="9" pageNumber="160" type="discussion">
<paragraph id="22249709FF8EFFD5C5827A10A2BFF9A5" blockId="9.[151,1437,1654,2005]" pageId="9" pageNumber="160">
<emphasis id="10EF4B1BFF8EFFD5C5827A10A15AF8E1" bold="true" box="[199,315,1654,1680]" pageId="9" pageNumber="160">Remarks.</emphasis>
The genus
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<emphasis id="10EF4B1BFF8EFFD5C4847A10A233F8E1" box="[449,594,1654,1680]" italics="true" pageId="9" pageNumber="160">Simognathus</emphasis>
Trouessart, 1889
</taxonomicName>
comprises 44 named species of which only two species,
<taxonomicName id="E59BEC8AFF8EFFD5C5D27AFCA1CFF8C5" authority="Viets, 1940" authorityName="Viets" authorityYear="1940" box="[151,430,1690,1716]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="10EF4B1BFF8EFFD5C5D27AFCA147F8C5" box="[151,294,1690,1716]" italics="true" pageId="9" pageNumber="160">S. adriaticus</emphasis>
<bibRefCitation id="460AEAF8FF8EFFD5C46B7AFCA1CFF8C5" author="Viets, K." box="[302,430,1690,1716]" pageId="9" pageNumber="160" pagination="1 - 135" refId="ref9235" refString="Viets, K. (1940) Meeresmilben aus der Adria (Halacaridae und Hydrachnellae, Acari). Archiv fur Naturgeschichte, Neue Folge, 9, 1 - 135." type="journal article" year="1940">Viets, 1940</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="E59BEC8AFF8EFFD5C4A37AFCA358F8C5" authority="Trouessart, 1894" authorityName="Trouessart" authorityYear="1894" box="[486,825,1690,1717]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF8EFFD5C4A37AFCA210F8C5" box="[486,625,1690,1716]" italics="true" pageId="9" pageNumber="160">S. leiomerus</emphasis>
Trouessart, 1894
</taxonomicName>
, are known from the Mediterranean Sea.
<taxonomicName id="E59BEC8AFF8EFFD5C04B7AFCA5FCF8C5" authorityName="Viets" authorityYear="1940" box="[1294,1437,1690,1716]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="10EF4B1BFF8EFFD5C04B7AFCA5FCF8C5" box="[1294,1437,1690,1716]" italics="true" pageId="9" pageNumber="160">S. adriaticus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="460AEAF8FF8EFFD5C4277AD8A18FF8A9" author="Viets, K." box="[354,494,1726,1752]" pageId="9" pageNumber="160" pagination="1 - 135" refId="ref9235" refString="Viets, K. (1940) Meeresmilben aus der Adria (Halacaridae und Hydrachnellae, Acari). Archiv fur Naturgeschichte, Neue Folge, 9, 1 - 135." type="journal article" year="1940">Viets (1940)</bibRefCitation>
from
<collectingCountry id="5A8CD799FF8EFFD5C7717AD8A2E7F8A9" box="[564,646,1726,1752]" name="Croatia" pageId="9" pageNumber="160">Croatia</collectingCountry>
(Rovigno) among various macroalgae and other substrates. Later, the species was reported from sublittoral sand habitat from Antalya (in
<bibRefCitation id="460AEAF8FF8EFFD5C6D07A84A4E1F88D" author="Stamouli, C. &amp; Akel, E. &amp; Azzurro, E. &amp; Bakiu, R. &amp; Bas, A. A. &amp; Bitar, G. &amp; Boyaci, Y. O. &amp; Cakalli, M. &amp; Corsini-Foka, M. &amp; Crocetta, F. &amp; Dragicevic, B. &amp; Dulcic, J. &amp; Durucan, F. &amp; El Zrelli, R. &amp; Erguden, D. &amp; Filiz, H. &amp; Giardina, F. &amp; Giovos, I. &amp; Gonulal, O. &amp; Hemida, F. &amp; Kassar, A. &amp; Kondylatos, G. &amp; Macali, A. &amp; Mancini, E. &amp; Ovalis, P. &amp; Paladini De Mendoza, F. &amp; Pavicic, M. &amp; Rabaoui, L. &amp; Rizkalla, S. I. &amp; Tiralongo, F. &amp; Turan, C. &amp; Vrdoljak, D. &amp; Yapici, S. &amp; Zenetos, A" box="[917,1152,1762,1789]" pageId="9" pageNumber="160" pagination="534 - 556" refId="ref8978" refString="Stamouli, C., Akel, E. H Kh., Azzurro, E., Bakiu, R., Bas, A. A., Bitar, G., Boyaci, Y. O., Cakalli, M., Corsini-Foka, M., Crocetta, F., Dragicevic, B., Dulcic, J., Durucan, F., El Zrelli, R., Erguden, D., Filiz, H., Giardina, F., Giovos, I., Gonulal, O., Hemida, F., Kassar, A., Kondylatos, G., Macali, A., Mancini, E., Ovalis, P., Paladini De Mendoza, F., Pavicic, M., Rabaoui, L., Rizkalla, S. I., Tiralongo, F., Turan, C., Vrdoljak, D., Yapici, S. &amp; Zenetos, A (2017) New Mediterranean Biodiversity Records (December 2017) Mediterranean Marine Science, 18 (3), 534 - 556. https: // doi. org / 10.12681 / mms. 15823" type="book chapter" year="2017">
Stamouli
<emphasis id="10EF4B1BFF8EFFD5C1477A85A454F88D" box="[1026,1077,1762,1788]" italics="true" pageId="9" pageNumber="160">et al</emphasis>
. 2017
</bibRefCitation>
). The present author also reported this species on the north of Aegean Sea (Çanakkale, Bozcaada) (
<bibRefCitation id="460AEAF8FF8EFFD5C6987B60A4E5F951" author="Durucan, F." box="[989,1156,1798,1824]" pageId="9" pageNumber="160" pagination="341 - 346" refId="ref7848" refString="Durucan, F. (2021) Halacaridae (Acari: Prostigmata) of the Aegean Sea of Turkey (Canakkale and Izmir). Persian Journal of Acarology, 10 (3), 341 - 346. https: // doi. org / 10.22073 / pja. v 10 i 3.67699" type="journal article" year="2021">Durucan 2021</bibRefCitation>
). The species belongs to
<emphasis id="10EF4B1BFF8EFFD5C5D27B4DA091F935" box="[151,240,1835,1860]" italics="true" pageId="9" pageNumber="160">minutus</emphasis>
group. This species group may be recognised by having large, triangular or rounded OC and generally with cornea (
<bibRefCitation id="460AEAF8FF8EFFD5C4777B28A1AFF919" author="Bartsch, I." box="[306,462,1870,1896]" pageId="9" pageNumber="160" pagination="135 - 152" refId="ref6910" refString="Bartsch, I. (1994) The genus Simognathus (Acari: Halacaridae), description of six new species from southern Australia and a tabular key to all species. Acarologia, 35, 135 - 152." type="journal article" year="1994">Bartsch 1994</bibRefCitation>
) and additionally
<taxonomicName id="E59BEC8AFF8EFFD5C7ED7B28A35AF919" authorityName="Viets" authorityYear="1940" box="[680,827,1870,1896]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="10EF4B1BFF8EFFD5C7ED7B28A35AF919" box="[680,827,1870,1896]" italics="true" pageId="9" pageNumber="160">S. adriaticus</emphasis>
</taxonomicName>
may recognised by: (1) body uniformly with lightbrownish pigmented, (2) median plate punctate in AE, (3) OC large, triangular, narrowing posteriorly in adults and deutonymphs, but the corners more rounded in protonymphs, (4) knob present on second palpal segment in adults but not observed in deutonymphs and protonmyphs.
</paragraph>
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<taxonomicName id="E59BEC8AFF85FFDEC5D27D09A201FFF8" authority="Morselli, 1970" authorityName="Morselli" authorityYear="1970" box="[151,608,367,393]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="monnioti">
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<emphasis id="10EF4B1BFF85FFDEC5D27D09A1C7FFF8" bold="true" box="[151,422,367,393]" italics="true" pageId="2" pageNumber="153">Acaromantis monnioti</emphasis>
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<paragraph id="22249709FF85FFDEC5D27DF5A188FFDC" blockId="2.[151,608,367,430]" box="[151,489,403,430]" pageId="2" pageNumber="153">
<figureCitation id="BAA08B8CFF85FFDEC5D27DF5A157FFDF" box="[151,310,403,430]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Figures 2AK</figureCitation>
,
<figureCitation id="BAA08B8CFF85FFDEC4007DF5A1EAFFDC" box="[325,395,403,429]" captionStart="FIGURE 6" captionStartId="9.[151,250,1512,1536]" captionTargetBox="[264,1322,181,1487]" captionTargetId="figure-16@9.[264,1323,181,1488]" captionTargetPageId="9" captionText="FIGURE 6. Acaromantis monnioti Morselli, 1970 (AC) (FDHAL-24/107) &amp; Simognathus adriaticus Viets, 1940 (D, E) (FDHAL-24/110,111): Light-microscope images of habitus dorsal deutonymph (A) habitus dorsal larva (B) and ventral view of quiescent pupal stage larva (C) habitus dorsal deutonymph (D) habitus dorsal protonymph (E) Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14734737" httpUri="https://zenodo.org/record/14734737/files/figure.png" pageId="2" pageNumber="153">6AC</figureCitation>
;
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[
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<taxonomicName id="E59BEC8AFF85FFDEC5E57DBEA2E5FF81" authority="Morselli, 1970: 109 - 119" authorityName="Morselli" authorityPageNumber="109 - 119" authorityYear="1970" box="[160,644,472,496]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="monnioti">
<emphasis id="10EF4B1BFF85FFDEC5E57DBEA1E1FF9E" box="[160,384,472,495]" italics="true" pageId="2" pageNumber="153">Acaromantis monnioti</emphasis>
<bibRefCitation id="460AEAF8FF85FFDEC4C27DBEA2E5FF81" author="Morselli, I." box="[391,644,472,496]" pageId="2" pageNumber="153" pagination="109 - 119" refId="ref8592" refString="Morselli, I. (1970) Ricerche sugli Alacaridi delle coste livornesi. II. - Acaromantis monnioti, una nuova specie raccoltasui fondi sabbiosi delle Secche della 206 Meloria (Livorno). Pubblicazioni della Stazione Zoologica di Napoli, 38, 109 - 119." type="journal article" year="1970">Morselli, 1970: 109119</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<figureCitation id="BAA08B8CFF85FFDEC7CA7DBEA2B5FF81" box="[655,724,472,496]" captionStart="FIGURE 1" captionStartId="1.[151,250,1962,1986]" captionTargetBox="[157,1436,955,1930]" captionTargetId="figure-369@1.[151,1436,955,1936]" captionTargetPageId="1" captionText="FIGURE 1. Map showing sampling localities for halacarid mite species inAntalya and Balikesir, Türkiye.Upper right: overview map of Türkiye indicates location of detailed map (Map-Source: Ocean Data View)." figureDoi="http://doi.org/10.5281/zenodo.14734725" httpUri="https://zenodo.org/record/14734725/files/figure.png" pageId="2" pageNumber="153">Figs. 1</figureCitation>
,
<figureCitation id="BAA08B8CFF85FFDEC7A57DBEA28CFF81" box="[736,749,472,496]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">2</figureCitation>
&amp; Tav.1]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6A81C482FF85FFDEC5D27E7BA4A2F896" pageId="2" pageNumber="153" type="materials_examined">
<paragraph id="22249709FF85FFDEC5D27E7BA3E0FCD5" blockId="2.[151,1436,541,676]" pageId="2" pageNumber="153">
<emphasis id="10EF4B1BFF85FFDEC5D27E7BA1E0FC46" bold="true" box="[151,385,541,567]" pageId="2" pageNumber="153">Material examined.</emphasis>
<materialsCitation id="92F39D54FF85FFDEC4CD7E78A31CFCD5" collectingDate="2018-10-18" latitude="36.84936" location="Kundu" longLatPrecision="1" longitude="30.839" municipality="Mites" pageId="2" pageNumber="153" specimenCount="2" specimenCount-larva="2" stateProvince="Antalya">
<collectingMunicipality id="C2400D73FF85FFDEC4CD7E78A1A9FC49" box="[392,456,542,568]" pageId="2" pageNumber="153">Mites</collectingMunicipality>
obtained at St. 5 (
<location id="2744C1D2FF85FFDEC7DD7E78A286FC49" LSID="urn:lsid:plazi:treatment:AA32261FFF85FFD8C5D27D09A088FFF8:2744C1D2FF85FFDEC7DD7E78A286FC49" box="[664,743,542,568]" latitude="36.84936" longLatPrecision="1" longitude="30.839" municipality="Mites" name="Kundu" pageId="2" pageNumber="153" stateProvince="Antalya">Kundu</location>
,
<collectingRegion id="E05F59EBFF85FFDEC7B47E78A32FFC49" box="[753,846,542,568]" country="Turkey" name="Antalya" pageId="2" pageNumber="153">Antalya</collectingRegion>
), well sorted medium grained sand, interstitial water at
<quantity id="E5633AECFF85FFDEC5F67E24A098FC2D" box="[179,249,578,604]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="153" unit="cm" value="20.0">20 cm</quantity>
depth on the seashore near the water,
<geoCoordinate id="47AFF1CEFF85FFDEC7E87E24A346FC2D" box="[685,807,578,604]" degrees="36.849361" direction="north" orientation="latitude" pageId="2" pageNumber="153" precision="1" value="36.84936">36.849361</geoCoordinate>
,
<geoCoordinate id="47AFF1CEFF85FFDEC6767E24A3CCFC2D" box="[819,941,578,604]" degrees="30.839000" direction="east" orientation="longitude" pageId="2" pageNumber="153" precision="1" value="30.839">30.839000</geoCoordinate>
,
<date id="5625B1C9FF85FFDEC6FC7E24A431FC2D" box="[953,1104,578,604]" pageId="2" pageNumber="153" value="2018-10-18">
<collectingDate id="46614821FF85FFDEC6FC7E24A431FC2D" box="[953,1104,578,604]" pageId="2" pageNumber="153" value="2018-10-18">18 Oct. 2018</collectingDate>
</date>
. Deutonymph and
<specimenCount id="349D5C80FF85FFDEC0637E24A5FDFC2D" box="[1318,1436,578,604]" count="2" pageId="2" pageNumber="153" type="larva">two larvae</specimenCount>
(FDHAL-24/107), undissected and slide mounted; Deutonymph (FDHAL-24/108), undissected and slide mounted; Quiescent larva (FDHAL-24/109), undissected and slide mounted
</materialsCitation>
.
</paragraph>
<caption id="76E4C781FF85FFDEC5DD7EB5A3EAFC9D" box="[152,907,723,748]" pageId="2" pageNumber="153" startId="2.[152,244,723,747]" targetType="table">
<paragraph id="22249709FF85FFDEC5DD7EB5A3EAFC9D" blockId="2.[152,907,723,748]" box="[152,907,723,748]" pageId="2" pageNumber="153">
<emphasis id="10EF4B1BFF85FFDEC5DD7EB5A16EFC9A" bold="true" box="[152,271,723,747]" pageId="2" pageNumber="153">TABLE 1.</emphasis>
Measurements for
<taxonomicName id="E59BEC8AFF85FFDEC49B7EB2A2AAFC9A" authorityName="Morselli" authorityYear="1970" box="[478,715,724,747]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="monnioti">
<emphasis id="10EF4B1BFF85FFDEC49B7EB2A2AAFC9A" box="[478,715,724,747]" italics="true" pageId="2" pageNumber="153">Acaromantis monnioti</emphasis>
</taxonomicName>
(in micrometers).
</paragraph>
</caption>
<paragraph id="22249709FF85FFDEC4497F76A537FAD7" pageId="2" pageNumber="153">
<table id="509B65A9FF850023C4497E9AA517FAD7" box="[268,1398,764,1190]" gridcols="5" gridrows="10" pageId="2" pageNumber="153">
<tr id="9CAB954BFF850023C4497E9AA517FD4D" box="[268,1398,764,828]" gridrow="0" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C4497E9AA2E8FD4D" box="[268,649,764,828]" gridcol="0" gridrow="0" pageId="2" pageNumber="153">
<taxonomicName id="E59BEC8AFF85FFDEC4497F76A2E8FD59" authority="Morselli, 1970 Deutonymphs Length Width Larvae Length Width Idiosoma" authorityName="Morselli" authorityYear="1970" box="[268,649,784,808]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="monnioti">
<emphasis id="10EF4B1BFF85FFDEC4497F76A18CFD56" box="[268,493,784,807]" italics="true" pageId="2" pageNumber="153">Acaromantis monnioti</emphasis>
<bibRefCitation id="460AEAF8FF85FFDEC4B67F76A2E8FD59" author="Morselli, I." box="[499,649,784,808]" pageId="2" pageNumber="153" pagination="109 - 119" refId="ref8592" refString="Morselli, I. (1970) Ricerche sugli Alacaridi delle coste livornesi. II. - Acaromantis monnioti, una nuova specie raccoltasui fondi sabbiosi delle Secche della 206 Meloria (Livorno). Pubblicazioni della Stazione Zoologica di Napoli, 38, 109 - 119." type="journal article" year="1970">Morselli, 1970</bibRefCitation>
</taxonomicName>
</th>
<th id="DF7AFC37FF850023C6617E9AA444FD4D" box="[804,1061,764,828]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="2" pageNumber="153">
<emphasis id="10EF4B1BFF85FFDEC61F7E9AA390FD65" bold="true" box="[858,1009,764,788]" pageId="2" pageNumber="153">Deutonymphs</emphasis>
Length Width
</th>
<th id="DF7AFC37FF850023C1307E9AA517FD4D" box="[1141,1398,764,828]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="2" pageNumber="153">
<emphasis id="10EF4B1BFF85FFDEC18A7E9AA57DFD65" bold="true" box="[1231,1308,764,788]" pageId="2" pageNumber="153">Larvae</emphasis>
Length Width
</th>
</tr>
<tr id="9CAB954BFF850023C4497F2AA517FD15" box="[268,1398,844,868]" gridrow="1" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C4497F2AA2E8FD15" box="[268,649,844,868]" gridcol="0" gridrow="1" pageId="2" pageNumber="153">Idiosoma</th>
<td id="DF7AFC37FF850023C6617F2AA31CFD15" box="[804,893,844,868]" gridcol="1" gridrow="1" pageId="2" pageNumber="153">250275</td>
<td id="DF7AFC37FF850023C6887F2AA444FD15" box="[973,1061,844,868]" gridcol="2" gridrow="1" pageId="2" pageNumber="153">125150</td>
<td id="DF7AFC37FF850023C1307F2AA4AFFD15" box="[1141,1230,844,868]" gridcol="3" gridrow="1" pageId="2" pageNumber="153">187207</td>
<td id="DF7AFC37FF850023C0587F2AA517FD15" box="[1309,1398,844,868]" gridcol="4" gridrow="1" pageId="2" pageNumber="153">105125</td>
</tr>
<tr id="9CAB954BFF850023C4497F12A517FDFD" box="[268,1398,884,908]" gridrow="2" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C4497F12A2E8FDFD" box="[268,649,884,908]" gridcol="0" gridrow="2" pageId="2" pageNumber="153">Gnathosoma</th>
<td id="DF7AFC37FF850023C6617F12A31CFDFD" box="[804,893,884,908]" gridcol="1" gridrow="2" pageId="2" pageNumber="153">68</td>
<td id="DF7AFC37FF850023C6887F12A444FDFD" box="[973,1061,884,908]" gridcol="2" gridrow="2" pageId="2" pageNumber="153">68</td>
<td id="DF7AFC37FF850023C1307F12A4AFFDFD" box="[1141,1230,884,908]" gridcol="3" gridrow="2" pageId="2" pageNumber="153">65</td>
<td id="DF7AFC37FF850023C0587F12A517FDFD" box="[1309,1398,884,908]" gridcol="4" gridrow="2" pageId="2" pageNumber="153">47</td>
</tr>
<tr id="9CAB954BFF850023C4497FFAA517FDC4" box="[268,1398,924,949]" gridrow="3" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C4497FFAA2E8FDC4" box="[268,649,924,949]" gridcol="0" gridrow="3" pageId="2" pageNumber="153">AD</th>
<td id="DF7AFC37FF850023C6617FFAA31CFDC4" box="[804,893,924,949]" gridcol="1" gridrow="3" pageId="2" pageNumber="153">100</td>
<td id="DF7AFC37FF850023C6887FFAA444FDC4" box="[973,1061,924,949]" gridcol="2" gridrow="3" pageId="2" pageNumber="153">88</td>
<td id="DF7AFC37FF850023C1307FFAA4AFFDC4" box="[1141,1230,924,949]" gridcol="3" gridrow="3" pageId="2" pageNumber="153">66</td>
<td id="DF7AFC37FF850023C0587FFAA517FDC4" box="[1309,1398,924,949]" gridcol="4" gridrow="3" pageId="2" pageNumber="153">56</td>
</tr>
<tr id="9CAB954BFF850023C4497FA3A517FDAC" box="[268,1398,965,989]" gridrow="4" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C4497FA3A2E8FDAC" box="[268,649,965,989]" gridcol="0" gridrow="4" pageId="2" pageNumber="153">OC</th>
<td id="DF7AFC37FF850023C6617FA3A31CFDAC" box="[804,893,965,989]" gridcol="1" gridrow="4" pageId="2" pageNumber="153">17</td>
<td id="DF7AFC37FF850023C6887FA3A444FDAC" box="[973,1061,965,989]" gridcol="2" gridrow="4" pageId="2" pageNumber="153">7</td>
<td id="DF7AFC37FF850023C1307FA3A4AFFDAC" box="[1141,1230,965,989]" gridcol="3" gridrow="4" pageId="2" pageNumber="153">13</td>
<td id="DF7AFC37FF850023C0587FA3A517FDAC" box="[1309,1398,965,989]" gridcol="4" gridrow="4" pageId="2" pageNumber="153">-</td>
</tr>
<tr id="9CAB954BFF850023C4497F8BA517FA74" box="[268,1398,1005,1029]" gridrow="5" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C4497F8BA2E8FA74" box="[268,649,1005,1029]" gridcol="0" gridrow="5" pageId="2" pageNumber="153">PD</th>
<td id="DF7AFC37FF850023C6617F8BA31CFA74" box="[804,893,1005,1029]" gridcol="1" gridrow="5" pageId="2" pageNumber="153">125</td>
<td id="DF7AFC37FF850023C6887F8BA444FA74" box="[973,1061,1005,1029]" gridcol="2" gridrow="5" pageId="2" pageNumber="153">100</td>
<td id="DF7AFC37FF850023C1307F8BA4AFFA74" box="[1141,1230,1005,1029]" gridcol="3" gridrow="5" pageId="2" pageNumber="153">38</td>
<td id="DF7AFC37FF850023C0587F8BA517FA74" box="[1309,1398,1005,1029]" gridcol="4" gridrow="5" pageId="2" pageNumber="153">62</td>
</tr>
<tr id="9CAB954BFF850023C4497873A517FA5C" box="[268,1398,1045,1069]" gridrow="6" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C4497873A2E8FA5C" box="[268,649,1045,1069]" gridcol="0" gridrow="6" pageId="2" pageNumber="153">AE</th>
<td id="DF7AFC37FF850023C6617873A31CFA5C" box="[804,893,1045,1069]" gridcol="1" gridrow="6" pageId="2" pageNumber="153">108</td>
<td id="DF7AFC37FF850023C6887873A444FA5C" box="[973,1061,1045,1069]" gridcol="2" gridrow="6" pageId="2" pageNumber="153">112</td>
<td id="DF7AFC37FF850023C1307873A4AFFA5C" box="[1141,1230,1045,1069]" gridcol="3" gridrow="6" pageId="2" pageNumber="153">75</td>
<td id="DF7AFC37FF850023C0587873A517FA5C" box="[1309,1398,1045,1069]" gridcol="4" gridrow="6" pageId="2" pageNumber="153">100</td>
</tr>
<tr id="9CAB954BFF850023C449785BA517FA24" box="[268,1398,1085,1109]" gridrow="7" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C449785BA2E8FA24" box="[268,649,1085,1109]" gridcol="0" gridrow="7" pageId="2" pageNumber="153">PE</th>
<td id="DF7AFC37FF850023C661785BA31CFA24" box="[804,893,1085,1109]" gridcol="1" gridrow="7" pageId="2" pageNumber="153">93</td>
<td id="DF7AFC37FF850023C688785BA444FA24" box="[973,1061,1085,1109]" gridcol="2" gridrow="7" pageId="2" pageNumber="153">28</td>
<td id="DF7AFC37FF850023C130785BA4AFFA24" box="[1141,1230,1085,1109]" gridcol="3" gridrow="7" pageId="2" pageNumber="153">35</td>
<td id="DF7AFC37FF850023C058785BA517FA24" box="[1309,1398,1085,1109]" gridcol="4" gridrow="7" pageId="2" pageNumber="153">18</td>
</tr>
<tr id="9CAB954BFF850023C4497803A517FA0F" box="[268,1398,1125,1150]" gridrow="8" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C4497803A2E8FA0F" box="[268,649,1125,1150]" gridcol="0" gridrow="8" pageId="2" pageNumber="153">GP</th>
<td id="DF7AFC37FF850023C6617803A31CFA0F" box="[804,893,1125,1150]" gridcol="1" gridrow="8" pageId="2" pageNumber="153">75</td>
<td id="DF7AFC37FF850023C6887803A444FA0F" box="[973,1061,1125,1150]" gridcol="2" gridrow="8" pageId="2" pageNumber="153">55</td>
<td id="DF7AFC37FF850023C1307803A4AFFA0F" box="[1141,1230,1125,1150]" gridcol="3" gridrow="8" pageId="2" pageNumber="153">-</td>
<td id="DF7AFC37FF850023C0587803A517FA0F" box="[1309,1398,1125,1150]" gridcol="4" gridrow="8" pageId="2" pageNumber="153">-</td>
</tr>
<tr id="9CAB954BFF850023C44978E8A517FAD7" box="[268,1398,1166,1190]" gridrow="9" pageId="2" pageNumber="153">
<th id="DF7AFC37FF850023C44978E8A2E8FAD7" box="[268,649,1166,1190]" gridcol="0" gridrow="9" pageId="2" pageNumber="153">AP</th>
<td id="DF7AFC37FF850023C66178E8A31CFAD7" box="[804,893,1166,1190]" gridcol="1" gridrow="9" pageId="2" pageNumber="153">-</td>
<td id="DF7AFC37FF850023C68878E8A444FAD7" box="[973,1061,1166,1190]" gridcol="2" gridrow="9" pageId="2" pageNumber="153">-</td>
<td id="DF7AFC37FF850023C13078E8A4AFFAD7" box="[1141,1230,1166,1190]" gridcol="3" gridrow="9" pageId="2" pageNumber="153">15</td>
<td id="DF7AFC37FF850023C05878E8A517FAD7" box="[1309,1398,1166,1190]" gridcol="4" gridrow="9" pageId="2" pageNumber="153">50</td>
</tr>
</table>
</paragraph>
<paragraph id="22249709FF85FFDEC582789FA15CF842" blockId="2.[151,1437,1273,2020]" pageId="2" pageNumber="153">
<emphasis id="10EF4B1BFF85FFDEC582789FA2EBFB65" bold="true" box="[199,650,1273,1300]" pageId="2" pageNumber="153">
Morphology and notes.
<emphasis id="10EF4B1BFF85FFDEC4A6789FA2EBFB65" bold="true" box="[483,650,1273,1300]" italics="true" pageId="2" pageNumber="153">Deutonymphs.</emphasis>
</emphasis>
Dorsal and ventral plates covered by foveae. Ds-1 on AD at 0.5 of its total length. OC spindle shaped. Pair of ds-2 and ds-3 on integument between AD and
<collectionCode id="448A0FCCFF85FFDEC1227978A4ECFB49" box="[1127,1165,1310,1336]" country="Netherlands" name="Dutch Plant Protection Service, Culture Collection of Plant Pathogenic Bacteria" pageId="2" pageNumber="153">PD</collectionCode>
, pairs of ds-4 at 0.3 and ds-5 at 0.9 of
<collectionCode id="448A0FCCFF85FFDEC46B7924A133FB2D" box="[302,338,1346,1372]" country="Netherlands" name="Dutch Plant Protection Service, Culture Collection of Plant Pathogenic Bacteria" pageId="2" pageNumber="153">PD</collectionCode>
length, respectively.
<collectionCode id="448A0FCCFF85FFDEC7787924A200FB2D" box="[573,609,1346,1372]" country="Netherlands" name="Dutch Plant Protection Service, Culture Collection of Plant Pathogenic Bacteria" pageId="2" pageNumber="153">PD</collectionCode>
longer than AD (AD:
<collectionCode id="448A0FCCFF85FFDEC6127924A31AFB2D" box="[855,891,1346,1372]" country="Netherlands" name="Dutch Plant Protection Service, Culture Collection of Plant Pathogenic Bacteria" pageId="2" pageNumber="153">PD</collectionCode>
= 0.8) (
<figureCitation id="BAA08B8CFF85FFDEC6977927A457FB2A" box="[978,1078,1345,1371]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Figs. 2A</figureCitation>
;
<figureCitation id="BAA08B8CFF85FFDEC1057927A407FB2A" box="[1088,1126,1345,1371]" captionStart="FIGURE 6" captionStartId="9.[151,250,1512,1536]" captionTargetBox="[264,1322,181,1487]" captionTargetId="figure-16@9.[264,1323,181,1488]" captionTargetPageId="9" captionText="FIGURE 6. Acaromantis monnioti Morselli, 1970 (AC) (FDHAL-24/107) &amp; Simognathus adriaticus Viets, 1940 (D, E) (FDHAL-24/110,111): Light-microscope images of habitus dorsal deutonymph (A) habitus dorsal larva (B) and ventral view of quiescent pupal stage larva (C) habitus dorsal deutonymph (D) habitus dorsal protonymph (E) Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14734737" httpUri="https://zenodo.org/record/14734737/files/figure.png" pageId="2" pageNumber="153">6A</figureCitation>
). AE with two pairs of setae and a pair of epimeral vesicles near insertion of leg
<collectionCode id="448A0FCCFF85FFDEC79A7900A294FBF1" box="[735,757,1382,1408]" pageId="2" pageNumber="153">II</collectionCode>
. PE with one dorsal and three ventral setae.
<collectionCode id="448A0FCCFF85FFDEC1AD7903A56AFB0E" box="[1256,1291,1381,1407]" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="2" pageNumber="153">GP</collectionCode>
large almost quadrangular with a pair of genital acetabula and two pairs of setae (
<figureCitation id="BAA08B8CFF85FFDEC6C279EFA3BFFBD2" box="[903,990,1417,1443]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Fig. 2B</figureCitation>
). Gnathosoma globular, as long as wide, 68 long (
<figureCitation id="BAA08B8CFF85FFDEC5BF79CBA130FBB6" box="[250,337,1453,1479]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Fig. 2B</figureCitation>
). Tibia I long, its base narrow, cylindrical, then rapidly expanding, ending with wide, smooth spine (
<figureCitation id="BAA08B8CFF85FFDEC5DA79B7A099FB9A" box="[159,248,1489,1515]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Fig. 2C</figureCitation>
). Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses; solenidia, famulus and pas included) (
<figureCitation id="BAA08B8CFF85FFDEC4787993A1DFF87E" box="[317,446,1525,1551]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Figs. 2CF</figureCitation>
); Leg I: 1, 1, 1, 4(1), 5, 6; Leg
<collectionCode id="448A0FCCFF85FFDEC65C7990A34EF861" box="[793,815,1526,1552]" pageId="2" pageNumber="153">II</collectionCode>
: 1, 1, 2, 4, 5(2), 6; Leg
<collectionCode id="448A0FCCFF85FFDEC1737990A437F861" box="[1078,1110,1526,1552]" pageId="2" pageNumber="153">III</collectionCode>
: 1, 1, 2, 3, 5(2), 6; Leg
<collectionCode id="448A0FCCFF85FFDEC0187990A51CF861" box="[1373,1405,1526,1552]" pageId="2" pageNumber="153">IV</collectionCode>
: 1, 1, 1, 3, 5(2), 6.
</paragraph>
<paragraph id="22249709FF85FFDEC5827A58A4A2F896" blockId="2.[151,1437,1273,2020]" pageId="2" pageNumber="153">
<emphasis id="10EF4B1BFF85FFDEC5827A58A17EF829" bold="true" box="[199,287,1598,1624]" italics="true" pageId="2" pageNumber="153">Larvae.</emphasis>
Dorsal plates smaller than deutonymph. Ds-1 on AD, ds-2 to ds-4 on integument. Ds-5 on
<collectionCode id="448A0FCCFF85FFDEC0757A58A535F829" box="[1328,1364,1598,1624]" country="Netherlands" name="Dutch Plant Protection Service, Culture Collection of Plant Pathogenic Bacteria" pageId="2" pageNumber="153">PD</collectionCode>
(
<figureCitation id="BAA08B8CFF85FFDEC0217A5BA0DCF80A" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Figs. 2G</figureCitation>
;
<figureCitation id="BAA08B8CFF85FFDEC58F7A07A08FF80A" box="[202,238,1633,1659]" captionStart="FIGURE 6" captionStartId="9.[151,250,1512,1536]" captionTargetBox="[264,1322,181,1487]" captionTargetId="figure-16@9.[264,1323,181,1488]" captionTargetPageId="9" captionText="FIGURE 6. Acaromantis monnioti Morselli, 1970 (AC) (FDHAL-24/107) &amp; Simognathus adriaticus Viets, 1940 (D, E) (FDHAL-24/110,111): Light-microscope images of habitus dorsal deutonymph (A) habitus dorsal larva (B) and ventral view of quiescent pupal stage larva (C) habitus dorsal deutonymph (D) habitus dorsal protonymph (E) Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14734737" httpUri="https://zenodo.org/record/14734737/files/figure.png" pageId="2" pageNumber="153">6B</figureCitation>
). AE with two pairs of setae and a pair of epimeral vesicles. PE with only one ventral seta and no dorsal setae. Genital plate absent (
<figureCitation id="BAA08B8CFF85FFDEC49D7AE3A257F8EE" box="[472,566,1669,1695]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Fig. 2H</figureCitation>
). Gnathosoma globular, 65 long and 47 wide (
<figureCitation id="BAA08B8CFF85FFDEC1127AE3A4D4F8EE" box="[1111,1205,1669,1695]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Fig. 2H</figureCitation>
). Leg chaetotaxy as follows (bipectinated setae between parentheses) (
<figureCitation id="BAA08B8CFF85FFDEC78C7ACFA356F8B5" box="[713,823,1705,1732]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Fig. 2IK</figureCitation>
); Leg I: 1, 2, 5, 5(1), 6; Leg
<collectionCode id="448A0FCCFF85FFDEC13D7ACCA4EFF8B5" box="[1144,1166,1706,1732]" pageId="2" pageNumber="153">II</collectionCode>
: 1, 3, 4, 5(1), 6; Leg
<collectionCode id="448A0FCCFF85FFDEC03D7ACCA5F9F8B5" box="[1400,1432,1706,1732]" pageId="2" pageNumber="153">III</collectionCode>
: 1, 3, 3, 5(2), 6. There is also one more larva specimen which is quiescent pupal stage (
<figureCitation id="BAA08B8CFF85FFDEC1187AABA4D7F896" box="[1117,1206,1741,1767]" captionStart="FIGURE 6" captionStartId="9.[151,250,1512,1536]" captionTargetBox="[264,1322,181,1487]" captionTargetId="figure-16@9.[264,1323,181,1488]" captionTargetPageId="9" captionText="FIGURE 6. Acaromantis monnioti Morselli, 1970 (AC) (FDHAL-24/107) &amp; Simognathus adriaticus Viets, 1940 (D, E) (FDHAL-24/110,111): Light-microscope images of habitus dorsal deutonymph (A) habitus dorsal larva (B) and ventral view of quiescent pupal stage larva (C) habitus dorsal deutonymph (D) habitus dorsal protonymph (E) Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14734737" httpUri="https://zenodo.org/record/14734737/files/figure.png" pageId="2" pageNumber="153">Fig. 6C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6A81C482FF85FFD8C5827A97A088FFF8" lastPageId="4" lastPageNumber="155" pageId="2" pageNumber="153" type="discussion">
<paragraph id="22249709FF85FFD8C5827A97A088FFF8" blockId="2.[151,1437,1273,2020]" lastBlockId="4.[151,1437,151,393]" lastPageId="4" lastPageNumber="155" pageId="2" pageNumber="153">
<emphasis id="10EF4B1BFF85FFDEC5827A97A15AF97A" bold="true" box="[199,315,1777,1803]" pageId="2" pageNumber="153">Remarks.</emphasis>
This species was described by
<bibRefCitation id="460AEAF8FF85FFDEC7E57A97A330F97A" author="Morselli, I." box="[672,849,1777,1803]" pageId="2" pageNumber="153" pagination="109 - 119" refId="ref8592" refString="Morselli, I. (1970) Ricerche sugli Alacaridi delle coste livornesi. II. - Acaromantis monnioti, una nuova specie raccoltasui fondi sabbiosi delle Secche della 206 Meloria (Livorno). Pubblicazioni della Stazione Zoologica di Napoli, 38, 109 - 119." type="journal article" year="1970">Morselli (1970)</bibRefCitation>
from
<collectingCountry id="5A8CD799FF85FFDEC6DD7A97A3AAF97A" box="[920,971,1777,1803]" name="Italy" pageId="2" pageNumber="153">Italy</collectingCountry>
(Livorno) among sand habitats (
<quantity id="E5633AECFF85FFDEC07A7A97A50DF97D" box="[1343,1388,1777,1804]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="153" unit="m" value="4.0">4 m</quantity>
and
<quantity id="E5633AECFF85FFDEC5D27B73A0B1F941" box="[151,208,1813,1840]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="153" unit="m" value="12.0">12 m</quantity>
depths) based on adults, nymphs and larva. Although both adults and juveniles were described in detail, only the adults were illustrated and photographed. The present author was reported adult specimens belonging to this species from sand habitat (
<quantity id="E5633AECFF85FFDEC48C7B3BA194F909" box="[457,501,1885,1912]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="153" unit="m" value="3.0">3 m</quantity>
depth) from Antalya (in
<bibRefCitation id="460AEAF8FF85FFDEC6567B3BA39CF906" author="Stamouli, C. &amp; Akel, E. &amp; Azzurro, E. &amp; Bakiu, R. &amp; Bas, A. A. &amp; Bitar, G. &amp; Boyaci, Y. O. &amp; Cakalli, M. &amp; Corsini-Foka, M. &amp; Crocetta, F. &amp; Dragicevic, B. &amp; Dulcic, J. &amp; Durucan, F. &amp; El Zrelli, R. &amp; Erguden, D. &amp; Filiz, H. &amp; Giardina, F. &amp; Giovos, I. &amp; Gonulal, O. &amp; Hemida, F. &amp; Kassar, A. &amp; Kondylatos, G. &amp; Macali, A. &amp; Mancini, E. &amp; Ovalis, P. &amp; Paladini De Mendoza, F. &amp; Pavicic, M. &amp; Rabaoui, L. &amp; Rizkalla, S. I. &amp; Tiralongo, F. &amp; Turan, C. &amp; Vrdoljak, D. &amp; Yapici, S. &amp; Zenetos, A" box="[787,1021,1885,1912]" pageId="2" pageNumber="153" pagination="534 - 556" refId="ref8978" refString="Stamouli, C., Akel, E. H Kh., Azzurro, E., Bakiu, R., Bas, A. A., Bitar, G., Boyaci, Y. O., Cakalli, M., Corsini-Foka, M., Crocetta, F., Dragicevic, B., Dulcic, J., Durucan, F., El Zrelli, R., Erguden, D., Filiz, H., Giardina, F., Giovos, I., Gonulal, O., Hemida, F., Kassar, A., Kondylatos, G., Macali, A., Mancini, E., Ovalis, P., Paladini De Mendoza, F., Pavicic, M., Rabaoui, L., Rizkalla, S. I., Tiralongo, F., Turan, C., Vrdoljak, D., Yapici, S. &amp; Zenetos, A (2017) New Mediterranean Biodiversity Records (December 2017) Mediterranean Marine Science, 18 (3), 534 - 556. https: // doi. org / 10.12681 / mms. 15823" type="book chapter" year="2017">
Stamouli
<emphasis id="10EF4B1BFF85FFDEC6C47B38A3D5F906" box="[897,948,1885,1911]" italics="true" pageId="2" pageNumber="153">et al</emphasis>
. 2017
</bibRefCitation>
) and it constituted the second report of this species. In this study, the author illustrated and photographed deutonymph and larva specimens for the first time (
<figureCitation id="BAA08B8CFF85FFDEC59C7BC3A13CF9B1" box="[217,349,1957,1984]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="2" pageNumber="153">Figs. 2AK</figureCitation>
;
<figureCitation id="BAA08B8CFF85FFDEC4287BC3A1D2F9CE" box="[365,435,1957,1983]" captionStart="FIGURE 6" captionStartId="9.[151,250,1512,1536]" captionTargetBox="[264,1322,181,1487]" captionTargetId="figure-16@9.[264,1323,181,1488]" captionTargetPageId="9" captionText="FIGURE 6. Acaromantis monnioti Morselli, 1970 (AC) (FDHAL-24/107) &amp; Simognathus adriaticus Viets, 1940 (D, E) (FDHAL-24/110,111): Light-microscope images of habitus dorsal deutonymph (A) habitus dorsal larva (B) and ventral view of quiescent pupal stage larva (C) habitus dorsal deutonymph (D) habitus dorsal protonymph (E) Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14734737" httpUri="https://zenodo.org/record/14734737/files/figure.png" pageId="2" pageNumber="153">6A, B</figureCitation>
). Additionally, a larva specimen in the quiescent pupal stage is presented here (
<figureCitation id="BAA08B8CFF85FFDEC0737BC3A5EEF9CE" box="[1334,1423,1957,1983]" captionStart="FIGURE 6" captionStartId="9.[151,250,1512,1536]" captionTargetBox="[264,1322,181,1487]" captionTargetId="figure-16@9.[264,1323,181,1488]" captionTargetPageId="9" captionText="FIGURE 6. Acaromantis monnioti Morselli, 1970 (AC) (FDHAL-24/107) &amp; Simognathus adriaticus Viets, 1940 (D, E) (FDHAL-24/110,111): Light-microscope images of habitus dorsal deutonymph (A) habitus dorsal larva (B) and ventral view of quiescent pupal stage larva (C) habitus dorsal deutonymph (D) habitus dorsal protonymph (E) Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14734737" httpUri="https://zenodo.org/record/14734737/files/figure.png" pageId="2" pageNumber="153">Fig. 6C</figureCitation>
). The genus
<taxonomicName id="E59BEC8AFF85FFDEC45C7BACA366F992" authority="Trouessart &amp; Neumann, 1893" authorityName="Trouessart &amp; Neumann" authorityYear="1893" box="[281,775,1993,2020]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="10EF4B1BFF85FFDEC45C7BACA1C6F995" box="[281,423,1994,2020]" italics="true" pageId="2" pageNumber="153">Acaromantis</emphasis>
<bibRefCitation id="460AEAF8FF85FFDEC4F47BACA366F992" author="Trouessart, E. &amp; Neumann, G." box="[433,775,1993,2020]" pageId="2" pageNumber="153" pagination="207 - 208" refId="ref9194" refString="Trouessart, E. &amp; Neumann, G. (1893) Description d'un nouveau genre et d'une nouvelle espece de la famille des Halacaridae ou Acariens marins. Le Naturaliste, Serie 2, 15, 207 - 208." type="journal article" year="1893">Trouessart &amp; Neumann, 1893</bibRefCitation>
</taxonomicName>
comprises 11 named species of which only two species,
<taxonomicName id="E59BEC8AFF83FFD8C5D27CF1A1ABFEC0" authority="Morselli, 1970" authorityName="Morselli" authorityYear="1970" box="[151,458,151,177]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="monnioti">
<emphasis id="10EF4B1BFF83FFD8C5D27CF1A17DFEC1" box="[151,284,151,177]" italics="true" pageId="4" pageNumber="155">A. monnioti</emphasis>
<bibRefCitation id="460AEAF8FF83FFD8C4617CF1A1ABFEC0" author="Morselli, I." box="[292,458,151,177]" pageId="4" pageNumber="155" pagination="109 - 119" refId="ref8592" refString="Morselli, I. (1970) Ricerche sugli Alacaridi delle coste livornesi. II. - Acaromantis monnioti, una nuova specie raccoltasui fondi sabbiosi delle Secche della 206 Meloria (Livorno). Pubblicazioni della Stazione Zoologica di Napoli, 38, 109 - 119." type="journal article" year="1970">Morselli, 1970</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="E59BEC8AFF83FFD8C7467CF1A3ADFEC0" authority="Trouessart &amp; Neumann, 1893" authorityName="Trouessart &amp; Neumann" authorityYear="1893" box="[515,972,151,177]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="squilla">
<emphasis id="10EF4B1BFF83FFD8C7467CF1A211FEC0" box="[515,624,151,177]" italics="true" pageId="4" pageNumber="155">A. squilla</emphasis>
<bibRefCitation id="460AEAF8FF83FFD8C73C7CF1A3ADFEC0" author="Trouessart, E. &amp; Neumann, G." box="[633,972,151,177]" pageId="4" pageNumber="155" pagination="207 - 208" refId="ref9194" refString="Trouessart, E. &amp; Neumann, G. (1893) Description d'un nouveau genre et d'une nouvelle espece de la famille des Halacaridae ou Acariens marins. Le Naturaliste, Serie 2, 15, 207 - 208." type="journal article" year="1893">Trouessart &amp; Neumann, 1893</bibRefCitation>
</taxonomicName>
, are known from the Mediterranean Sea.
<taxonomicName id="E59BEC8AFF83FFD8C5D27CDDA17BFEA5" authorityName="Morselli" authorityYear="1970" box="[151,282,187,213]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="monnioti">
<emphasis id="10EF4B1BFF83FFD8C5D27CDDA17BFEA5" box="[151,282,187,213]" italics="true" pageId="4" pageNumber="155">A. monnioti</emphasis>
</taxonomicName>
resembles
<taxonomicName id="E59BEC8AFF83FFD8C4D27CDDA263FEA4" authorityName="Trouessart &amp; Neumann" authorityYear="1893" box="[407,514,187,213]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="squilla">
<emphasis id="10EF4B1BFF83FFD8C4D27CDDA263FEA4" box="[407,514,187,213]" italics="true" pageId="4" pageNumber="155">A. squilla</emphasis>
</taxonomicName>
. However, the adults of former species distinguished from the latter by not having ventral lamellae on telofemur-I and tibia-I of
<taxonomicName id="E59BEC8AFF83FFD8C7D07CB9A361FE88" authorityName="Trouessart &amp; Neumann" authorityYear="1893" box="[661,768,223,249]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="squilla">
<emphasis id="10EF4B1BFF83FFD8C7D07CB9A361FE88" box="[661,768,223,249]" italics="true" pageId="4" pageNumber="155">A. squilla</emphasis>
</taxonomicName>
more conical than
<taxonomicName id="E59BEC8AFF83FFD8C69D7CB9A57CFE88" authority="(Morselli, 1970)" baseAuthorityName="Morselli" baseAuthorityYear="1970" box="[984,1309,223,249]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="monnioti">
<emphasis id="10EF4B1BFF83FFD8C69D7CB9A43AFE89" box="[984,1115,223,249]" italics="true" pageId="4" pageNumber="155">A. monnioti</emphasis>
(
<bibRefCitation id="460AEAF8FF83FFD8C12C7CB9A575FE88" author="Morselli, I." box="[1129,1300,223,249]" pageId="4" pageNumber="155" pagination="109 - 119" refId="ref8592" refString="Morselli, I. (1970) Ricerche sugli Alacaridi delle coste livornesi. II. - Acaromantis monnioti, una nuova specie raccoltasui fondi sabbiosi delle Secche della 206 Meloria (Livorno). Pubblicazioni della Stazione Zoologica di Napoli, 38, 109 - 119." type="journal article" year="1970">Morselli, 1970</bibRefCitation>
)
</taxonomicName>
. Regarding to the comparison of juveniles of the two species, OC reduced to small oblong subcuticular platelet in
<taxonomicName id="E59BEC8AFF83FFD8C05C7D65A5FCFF6D" baseAuthorityName="Morselli" baseAuthorityYear="1970" box="[1305,1437,259,285]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="monnioti">
<emphasis id="10EF4B1BFF83FFD8C05C7D65A5FCFF6D" box="[1305,1437,259,285]" italics="true" pageId="4" pageNumber="155">A. monnioti</emphasis>
</taxonomicName>
both deutonmyph and larva while there is no OC of protonymph stage of
<taxonomicName id="E59BEC8AFF83FFD8C6A27D41A435FF30" authorityName="Trouessart &amp; Neumann" authorityYear="1893" box="[999,1108,295,321]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="squilla">
<emphasis id="10EF4B1BFF83FFD8C6A27D41A435FF30" box="[999,1108,295,321]" italics="true" pageId="4" pageNumber="155">A. squilla</emphasis>
</taxonomicName>
and telofemura I to III have ventral lamellae in
<taxonomicName id="E59BEC8AFF83FFD8C4287D2DA1B9FF14" authorityName="Trouessart &amp; Neumann" authorityYear="1893" box="[365,472,331,357]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="squilla">
<emphasis id="10EF4B1BFF83FFD8C4287D2DA1B9FF14" box="[365,472,331,357]" italics="true" pageId="4" pageNumber="155">A. squilla</emphasis>
</taxonomicName>
while telofemura I to III have not ventral lamellae in
<taxonomicName id="E59BEC8AFF83FFD8C1687D2DA4CEFF15" baseAuthorityName="Morselli" baseAuthorityYear="1970" box="[1069,1199,331,357]" class="Arachnida" family="Halacaridae" genus="Acaromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="monnioti">
<emphasis id="10EF4B1BFF83FFD8C1687D2DA4CEFF15" box="[1069,1199,331,357]" italics="true" pageId="4" pageNumber="155">A. monnioti</emphasis>
</taxonomicName>
according to
<bibRefCitation id="460AEAF8FF83FFD8C0027D2DA087FFF8" author="Bartsch, I." pageId="4" pageNumber="155" pagination="51 - 78" refId="ref6769" refString="Bartsch, I. (1973) Halacaridae (Acari) von der Josephinebank und der grossen Meteorbank aus dem ostlichen Nordatlantik. II. Die Halacaridae aus den Bodengreiferproben. Meteor Forschungs-Ergebnisse, Reihe D, 15, 51 - 78." type="journal article" year="1973">Bartsch (1973)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,495 @@
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<paragraph id="22249709FF8DFFD6C5D27EF3A2EDFCDE" blockId="10.[151,652,661,723]" box="[151,652,661,688]" pageId="10" pageNumber="161">
<heading id="796C2065FF8DFFD6C5D27EF3A2EDFCDE" bold="true" box="[151,652,661,688]" fontSize="11" level="1" pageId="10" pageNumber="161" reason="1">
<taxonomicName id="E59BEC8AFF8DFFD6C5D27EF3A2EDFCDE" authority="Trouessart, 1894" authorityName="Trouessart" authorityYear="1894" box="[151,652,661,688]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF8DFFD6C5D27EF3A2EDFCDE" bold="true" box="[151,652,661,688]" pageId="10" pageNumber="161">
<emphasis id="10EF4B1BFF8DFFD6C5D27EF3A1D4FCC1" bold="true" box="[151,437,661,688]" italics="true" pageId="10" pageNumber="161">Simognathus leiomerus</emphasis>
Trouessart, 1894
</emphasis>
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</paragraph>
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<subSubSection id="6A81C482FF8DFFD6C5D27EDFA2EBFCA2" box="[151,650,697,723]" pageId="10" pageNumber="161" type="description">
<paragraph id="22249709FF8DFFD6C5D27EDFA2EBFCA2" blockId="10.[151,652,661,723]" box="[151,650,697,723]" pageId="10" pageNumber="161">
<figureCitation id="BAA08B8CFF8DFFD6C5D27EDFA14EFCA2" box="[151,303,697,723]" captionStart="FIGURE 7" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[191,1396,186,1846]" captionTargetId="figure-16@11.[187,1401,181,1850]" captionTargetPageId="11" captionText="FIGURE 7. Simognathus leiomerus Trouessart, 1894: Line art illustrations of female (A, B, DI) (FDHAL-24/113) and male (C) (FDHAL-24/118): A Idiosoma, dorsal view B Idiosoma, ventral view B' Close up of female genital opening C Idiosoma, ventral view C' Close up of spermatopositor and male genital opening D Gnathosoma, lateral view E Gnathosoma, dorsal view F Leg I, medial view G Leg II, medial view H Leg III, medial view I Leg IV, medial view. Scale bars: AC, FH=100 µm; D, E=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734741" httpUri="https://zenodo.org/record/14734741/files/figure.png" pageId="10" pageNumber="161">Figures 7AI</figureCitation>
;
<figureCitation id="BAA08B8CFF8DFFD6C47E7EDFA119FCA2" box="[315,376,697,723]" captionStart="FIGURE 8" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[193,1396,186,1846]" captionTargetId="figure-27@12.[189,1398,181,1850]" captionTargetPageId="12" captionText="FIGURE 8. Simognathus leiomerus Trouessart, 1894: Line art illustrations of protonymph (AG) (FDHAL-24/122) and deutonymph (H, I) (FDHAL-24/121) A Idiosoma, dorsal view A'Close up of integument and surrounding area of OC A' Close up of integument B Idiosoma, ventral view B' &amp; B&quot;Close up of genital plate and near integument C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view I' Close up of genital plate Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734743" httpUri="https://zenodo.org/record/14734743/files/figure.png" pageId="10" pageNumber="161">8AI</figureCitation>
;
<figureCitation id="BAA08B8CFF8DFFD6C4C67EDFA1ABFCA2" box="[387,458,697,723]" captionStart="FIGURE 9" captionStartId="13.[151,250,1113,1137]" captionTargetBox="[163,1426,193,1081]" captionTargetId="figure-16@13.[151,1435,181,1089]" captionTargetPageId="13" captionText="FIGURE 9. Simognathus leiomerus Trouessart, 1894: Light-microscope images of female (A, B) (FDHAL-24/113) male (C, D) (FDHAL-24/118) deutonmyph (E, F) (FDHAL-24/121) and protonymph (G, H) (FDHAL-24/122) A Idiosoma, dorsal view B Idiosoma, ventral view C Idiosoma, dorsal view D Idiosoma, ventral view E Idiosoma, dorsal view F Idiosoma, ventral view G Idiosoma, dorsal view H Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734745" httpUri="https://zenodo.org/record/14734745/files/figure.png" pageId="10" pageNumber="161">9AH</figureCitation>
;
<figureCitation id="BAA08B8CFF8DFFD6C4937EDFA24DFCA2" box="[470,556,697,723]" captionStart="FIGURE 10" captionStartId="14.[151,250,1908,1932]" captionTargetBox="[200,1399,196,1885]" captionTargetId="figure-27@14.[188,1399,181,1885]" captionTargetPageId="14" captionText="FIGURE 10. Simognathus leiomerus Trouessart, 1894: Light-microscope images of female (AD) (FDHAL-24/113) A Anterior part of idiosoma and gnathosoma, dorsal view B Gnathosoma, lateral view B' Close up of palps, lateral view C Close up of right ocular plate D Leg I, medial view, detail showing tibia and tarsus I D' Close up of tarsus and tibia I, medial view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734747" httpUri="https://zenodo.org/record/14734747/files/figure.png" pageId="10" pageNumber="161">10AD</figureCitation>
;
<tableCitation id="6F19A2B2FF8DFFD6C7727EDFA2EBFCA2" box="[567,650,697,723]" captionStart="TABLE 4" captionStartId="13.[152,244,1293,1317]" captionText="TABLE 4. Measurements for Simognathus leiomerus (in micrometers)." pageId="10" pageNumber="161">Table 4</tableCitation>
</paragraph>
</subSubSection>
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<paragraph id="22249709FF8DFFD6C5D27E98A3E2FD64" blockId="10.[151,899,765,820]" box="[151,899,765,790]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="028BB027FF8DFFD6C5D27E98A3E2FD64" box="[151,899,765,790]" pageId="10" pageNumber="161">
[
<treatmentCitation id="A33AB118FF8DFFD6C5E57E98A210FD64" author="Monniot, F." box="[160,625,765,790]" page="590" pageId="10" pageNumber="161" year="1961">
<taxonomicName id="E59BEC8AFF8DFFD6C5E57E98A210FD64" authority="Monniot, 1961: 585 - 590" authorityName="Monniot" authorityPageNumber="585 - 590" authorityYear="1961" box="[160,625,765,790]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="10EF4B1BFF8DFFD6C5E57E98A10BFD67" box="[160,362,766,790]" italics="true" pageId="10" pageNumber="161">Simognathus andrei</emphasis>
<bibRefCitation id="460AEAF8FF8DFFD6C4357E98A210FD64" author="Monniot, F." box="[368,625,765,790]" pageId="10" pageNumber="161" pagination="585 - 590" refId="ref8567" refString="Monniot, F. (1961) Simognathus andrei, nouvelle espece d'halacarien recoltee en Mediterranee. Acarologia, 3, 585 - 590." type="journal article" year="1961">Monniot, 1961: 585590</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<figureCitation id="BAA08B8CFF8DFFD6C73F7E98A290FD67" box="[634,753,766,790]" captionStart="FIGURE 1" captionStartId="1.[151,250,1962,1986]" captionTargetBox="[157,1436,955,1930]" captionTargetId="figure-369@1.[151,1436,955,1936]" captionTargetPageId="1" captionText="FIGURE 1. Map showing sampling localities for halacarid mite species inAntalya and Balikesir, Türkiye.Upper right: overview map of Türkiye indicates location of detailed map (Map-Source: Ocean Data View)." figureDoi="http://doi.org/10.5281/zenodo.14734725" httpUri="https://zenodo.org/record/14734725/files/figure.png" pageId="10" pageNumber="161">Figs. 1AC</figureCitation>
,
<figureCitation id="BAA08B8CFF8DFFD6C7B97E98A35CFD67" box="[764,829,766,790]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[193,1394,181,1850]" captionTargetId="figure-16@3.[193,1394,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Acaromantis monnioti Morselli, 1970: Line art illustrations of deutonymph (AF) (FDHAL-24/107) and larva (GK) (FDHAL-24/107) A Idiosoma, dorsal view B Idiosoma, ventral view C Leg I, medial view C' Close up of tibia and tarsus I, medial view D Leg II, ventromedial view E Leg III, medial view F Leg IV, medial view G Idiosoma, dorsal view H Idiosoma, ventral view I Leg I, medial view I' Close up of tibia and tarsus I, medial view J Leg II, ventral view K Leg III, ventral view. Scale bars: A, B=100 µm; CK=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734727" httpUri="https://zenodo.org/record/14734727/files/figure.png" pageId="10" pageNumber="161">2AD</figureCitation>
,
<figureCitation id="BAA08B8CFF8DFFD6C6037E98A3E2FD64" box="[838,899,766,790]" captionStart="FIGURE 3" captionStartId="5.[151,250,1874,1898]" captionTargetBox="[198,1390,181,1850]" captionTargetId="figure-16@5.[198,1390,181,1850]" captionTargetPageId="5" captionText="FIGURE 3. Acaromantis nurhayatae sp. nov.: Line art illustrations of female (AG) (FDHAL-24/106) A Idiosoma, dorsal view A' Close up of integument and surrounding area of OC A&quot; Close up of OC B Idiosoma, ventral view C Gnathosoma, ventrolateral view D Leg I, lateral view D' Close up of tarsus I, medial view E Leg II, medial view E' Close up of tarsus II, medial view E&quot; Tarsus II, lateral view F Leg III, medial view F' Close up of tarsus III, medial view G Leg IV, medial view. Scale bars: A, B, D, F, I, K =100 µm; C, E, G, H, J =50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734729" httpUri="https://zenodo.org/record/14734729/files/figure.png" pageId="10" pageNumber="161">3A, B</figureCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="22249709FF8DFFD6C5D27F7AA290FD45" blockId="10.[151,899,765,820]" box="[151,753,796,820]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="028BB027FF8DFFD6C5D27F7AA290FD45" box="[151,753,796,820]" pageId="10" pageNumber="161">
<treatmentCitation id="A33AB118FF8DFFD6C5D27F7AA220FD45" author="Bartsch, I." box="[151,577,796,820]" page="43" pageId="10" pageNumber="161" year="1980">
<taxonomicName id="E59BEC8AFF8DFFD6C5D27F7AA220FD45" authority="Bartsch, 1980: 43" authorityName="Bartsch" authorityPageNumber="43" authorityYear="1980" box="[151,577,796,820]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF8DFFD6C5D27F7AA1E5FD45" box="[151,388,796,820]" italics="true" pageId="10" pageNumber="161">Simognathus leiomerus</emphasis>
<bibRefCitation id="460AEAF8FF8DFFD6C4CE7F7AA220FD45" author="Bartsch, I." box="[395,577,796,820]" pageId="10" pageNumber="161" pagination="34 - 45" refId="ref6812" refString="Bartsch, I. (1980) Halacaridae (Acari) aus der Bucht von Morlaix (Bretagne). Acarologia, 21, 34 - 45." type="journal article" year="1980">Bartsch, 1980: 43</bibRefCitation>
</taxonomicName>
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, 44, Figs. 2331]
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</paragraph>
</subSubSection>
<subSubSection id="6A81C482FF8DFFD6C5D27F04A2E9FA7D" pageId="10" pageNumber="161" type="materials_examined">
<paragraph id="22249709FF8DFFD6C5D27F04A2E9FA7D" blockId="10.[151,1437,866,2009]" pageId="10" pageNumber="161">
<emphasis id="10EF4B1BFF8DFFD6C5D27F04A1E7FD0D" bold="true" box="[151,390,866,892]" pageId="10" pageNumber="161">Material examined.</emphasis>
<materialsCitation id="92F39D54FF8DFFD6C4D77F04A2E5FA7D" collectingDate="2021-06-16" elevation="6" latitude="36.865498" location="All" longLatPrecision="1" longitude="30.724024" pageId="10" pageNumber="161" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Antalya">
<location id="2744C1D2FF8DFFD6C4D77F04A1D7FD0D" LSID="urn:lsid:plazi:treatment:AA32261FFF8DFFD3C5D27EF3A49FFF6C:2744C1D2FF8DFFD6C4D77F04A1D7FD0D" box="[402,438,866,892]" latitude="36.865498" longLatPrecision="1" longitude="30.724024" name="All" pageId="10" pageNumber="161" stateProvince="Antalya">All</location>
individuals slide mounted, collected at St. 4 (ErenkuŞ,
<collectingRegion id="E05F59EBFF8DFFD6C1167F04A4D1FD0D" box="[1107,1200,866,892]" country="Turkey" name="Antalya" pageId="10" pageNumber="161">Antalya</collectingRegion>
), fine sand,
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<elevation id="A9B6703AFF8DFFD6C00C7F04A5F6FD0C" box="[1353,1431,866,893]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="10" pageNumber="161" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">56 m</elevation>
</quantity>
,
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,
<geoCoordinate id="47AFF1CEFF8DFFD6C4657FE1A1FBFDD0" box="[288,410,903,929]" degrees="30.724024" direction="east" orientation="longitude" pageId="10" pageNumber="161" precision="1" value="30.724024">30.724024</geoCoordinate>
,
<date id="5625B1C9FF8DFFD6C4EC7FE0A228FDD0" box="[425,585,902,929]" pageId="10" pageNumber="161" value="2021-06-16">
<collectingDate id="46614821FF8DFFD6C4EC7FE0A228FDD0" box="[425,585,902,929]" pageId="10" pageNumber="161" value="2021-06-16">16 June 2021</collectingDate>
</date>
. Female (FDHAL-24/113), dissected, Female and deutonymph (FDHAL-24/114), dissected,
<specimenCount id="349D5C80FF8DFFD6C4317FCCA277FDB5" box="[372,534,938,964]" count="3" pageId="10" pageNumber="161" type="female">Three females</specimenCount>
(FDHAL-24/115, FDHAL-24/116, FDHAL-24/117), undissected,
<specimenCount id="349D5C80FF8DFFD6C0557FCCA5FDFDB5" box="[1296,1436,938,964]" count="3" pageId="10" pageNumber="161" type="male">Three males</specimenCount>
(FDHAL-24/118, FDHAL-24/119, FDHAL-24/120), undissected, Deutonymph (FDHAL-24/121), undissected, Protonymph (FDHAL-24/122), undissected
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6A81C482FF8DFFD6C5D27870A3FEF9A9" pageId="10" pageNumber="161" type="description">
<paragraph id="22249709FF8DFFD6C5D27870A2B6FA40" blockId="10.[151,1437,866,2009]" box="[151,727,1046,1073]" pageId="10" pageNumber="161">
<emphasis id="10EF4B1BFF8DFFD6C5D27870A124FA41" bold="true" box="[151,325,1046,1072]" pageId="10" pageNumber="161">Redescription.</emphasis>
Measurements presented in
<tableCitation id="6F19A2B2FF8DFFD6C7C17870A2B6FA40" box="[644,727,1046,1073]" captionStart="TABLE 4" captionStartId="13.[152,244,1293,1317]" captionText="TABLE 4. Measurements for Simognathus leiomerus (in micrometers)." pageId="10" pageNumber="161">Table 4</tableCitation>
</paragraph>
<paragraph id="22249709FF8DFFD6C582785DA4E3F859" blockId="10.[151,1437,866,2009]" pageId="10" pageNumber="161">
<emphasis id="10EF4B1BFF8DFFD6C582785DA221FA24" bold="true" box="[199,576,1082,1109]" pageId="10" pageNumber="161">
Morphology and notes.
<emphasis id="10EF4B1BFF8DFFD6C49A785CA221FA24" box="[479,576,1082,1109]" italics="true" pageId="10" pageNumber="161">Females</emphasis>
</emphasis>
. Dorsal and ventral plates well developed and covered by foveae (
<figureCitation id="BAA08B8CFF8DFFD6C05A785CA5F6FA24" box="[1311,1431,1082,1109]" captionStart="FIGURE 7" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[191,1396,186,1846]" captionTargetId="figure-16@11.[187,1401,181,1850]" captionTargetPageId="11" captionText="FIGURE 7. Simognathus leiomerus Trouessart, 1894: Line art illustrations of female (A, B, DI) (FDHAL-24/113) and male (C) (FDHAL-24/118): A Idiosoma, dorsal view B Idiosoma, ventral view B' Close up of female genital opening C Idiosoma, ventral view C' Close up of spermatopositor and male genital opening D Gnathosoma, lateral view E Gnathosoma, dorsal view F Leg I, medial view G Leg II, medial view H Leg III, medial view I Leg IV, medial view. Scale bars: AC, FH=100 µm; D, E=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734741" httpUri="https://zenodo.org/record/14734741/files/figure.png" pageId="10" pageNumber="161">Fig. 7A, B</figureCitation>
;
<figureCitation id="BAA08B8CFF8DFFD6C5D27839A0BAFA08" box="[151,219,1119,1145]" captionStart="FIGURE 9" captionStartId="13.[151,250,1113,1137]" captionTargetBox="[163,1426,193,1081]" captionTargetId="figure-16@13.[151,1435,181,1089]" captionTargetPageId="13" captionText="FIGURE 9. Simognathus leiomerus Trouessart, 1894: Light-microscope images of female (A, B) (FDHAL-24/113) male (C, D) (FDHAL-24/118) deutonmyph (E, F) (FDHAL-24/121) and protonymph (G, H) (FDHAL-24/122) A Idiosoma, dorsal view B Idiosoma, ventral view C Idiosoma, dorsal view D Idiosoma, ventral view E Idiosoma, dorsal view F Idiosoma, ventral view G Idiosoma, dorsal view H Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734745" httpUri="https://zenodo.org/record/14734745/files/figure.png" pageId="10" pageNumber="161">9A, B</figureCitation>
). Integument brown coloured (
<figureCitation id="BAA08B8CFF8DFFD6C7757838A2C9FA08" box="[560,680,1118,1145]" captionStart="FIGURE 9" captionStartId="13.[151,250,1113,1137]" captionTargetBox="[163,1426,193,1081]" captionTargetId="figure-16@13.[151,1435,181,1089]" captionTargetPageId="13" captionText="FIGURE 9. Simognathus leiomerus Trouessart, 1894: Light-microscope images of female (A, B) (FDHAL-24/113) male (C, D) (FDHAL-24/118) deutonmyph (E, F) (FDHAL-24/121) and protonymph (G, H) (FDHAL-24/122) A Idiosoma, dorsal view B Idiosoma, ventral view C Idiosoma, dorsal view D Idiosoma, ventral view E Idiosoma, dorsal view F Idiosoma, ventral view G Idiosoma, dorsal view H Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734745" httpUri="https://zenodo.org/record/14734745/files/figure.png" pageId="10" pageNumber="161">Fig. 9AH</figureCitation>
) deeper integumental layers with numerous, very delicate canaliculi. Dorsum has six pairs of setae, ds-1 and ds-3 on AD. Ds-2 on anterior end of OC, reduced to a small oblong subcuticular platelet (
<figureCitation id="BAA08B8CFF8DFFD6C5BD78C0A13FFAB0" box="[248,350,1190,1217]" captionStart="FIGURE 10" captionStartId="14.[151,250,1908,1932]" captionTargetBox="[200,1399,196,1885]" captionTargetId="figure-27@14.[188,1399,181,1885]" captionTargetPageId="14" captionText="FIGURE 10. Simognathus leiomerus Trouessart, 1894: Light-microscope images of female (AD) (FDHAL-24/113) A Anterior part of idiosoma and gnathosoma, dorsal view B Gnathosoma, lateral view B' Close up of palps, lateral view C Close up of right ocular plate D Leg I, medial view, detail showing tibia and tarsus I D' Close up of tarsus and tibia I, medial view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734747" httpUri="https://zenodo.org/record/14734747/files/figure.png" pageId="10" pageNumber="161">Fig. 10C</figureCitation>
). Pairs of ds-4 to ds-6 on PD. Marginal and median AE foveated and with three pairs of setae and a pair of epimeral vesicles which are large and 37 long. PE with a dorsal and three ventral setae. Anterior part of GA truncate. Interval from GO to anterior margin of GA equalling 0.6 times the GOs length. GA with 4 pairs of pgs but no sgs (
<figureCitation id="BAA08B8CFF8DFFD6C5AB7974A153FB5C" box="[238,306,1298,1325]" captionStart="FIGURE 7" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[191,1396,186,1846]" captionTargetId="figure-16@11.[187,1401,181,1850]" captionTargetPageId="11" captionText="FIGURE 7. Simognathus leiomerus Trouessart, 1894: Line art illustrations of female (A, B, DI) (FDHAL-24/113) and male (C) (FDHAL-24/118): A Idiosoma, dorsal view B Idiosoma, ventral view B' Close up of female genital opening C Idiosoma, ventral view C' Close up of spermatopositor and male genital opening D Gnathosoma, lateral view E Gnathosoma, dorsal view F Leg I, medial view G Leg II, medial view H Leg III, medial view I Leg IV, medial view. Scale bars: AC, FH=100 µm; D, E=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734741" httpUri="https://zenodo.org/record/14734741/files/figure.png" pageId="10" pageNumber="161">Fig. 7</figureCitation>
B'). Gnathosomal base almost globular, with three segmented palps (
<figureCitation id="BAA08B8CFF8DFFD6C1747974A4D4FB5C" box="[1073,1205,1298,1325]" captionStart="FIGURE 7" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[191,1396,186,1846]" captionTargetId="figure-16@11.[187,1401,181,1850]" captionTargetPageId="11" captionText="FIGURE 7. Simognathus leiomerus Trouessart, 1894: Line art illustrations of female (A, B, DI) (FDHAL-24/113) and male (C) (FDHAL-24/118): A Idiosoma, dorsal view B Idiosoma, ventral view B' Close up of female genital opening C Idiosoma, ventral view C' Close up of spermatopositor and male genital opening D Gnathosoma, lateral view E Gnathosoma, dorsal view F Leg I, medial view G Leg II, medial view H Leg III, medial view I Leg IV, medial view. Scale bars: AC, FH=100 µm; D, E=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734741" httpUri="https://zenodo.org/record/14734741/files/figure.png" pageId="10" pageNumber="161">Figs. 7D, E</figureCitation>
;
<figureCitation id="BAA08B8CFF8DFFD6C1FC7975A48AFB5C" box="[1209,1259,1299,1325]" captionStart="FIGURE 10" captionStartId="14.[151,250,1908,1932]" captionTargetBox="[200,1399,196,1885]" captionTargetId="figure-27@14.[188,1399,181,1885]" captionTargetPageId="14" captionText="FIGURE 10. Simognathus leiomerus Trouessart, 1894: Light-microscope images of female (AD) (FDHAL-24/113) A Anterior part of idiosoma and gnathosoma, dorsal view B Gnathosoma, lateral view B' Close up of palps, lateral view C Close up of right ocular plate D Leg I, medial view, detail showing tibia and tarsus I D' Close up of tarsus and tibia I, medial view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734747" httpUri="https://zenodo.org/record/14734747/files/figure.png" pageId="10" pageNumber="161">10B</figureCitation>
). Second palpal segment with a ventral seta inserted in ventral knob. Tectum triangular. Leg chaetotaxy, from trochanters to tarsus (bipectinated setae between parentheses; solenidia and famulus excluded, pas included) (
<figureCitation id="BAA08B8CFF8DFFD6C132793CA48EFB04" box="[1143,1263,1370,1397]" captionStart="FIGURE 7" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[191,1396,186,1846]" captionTargetId="figure-16@11.[187,1401,181,1850]" captionTargetPageId="11" captionText="FIGURE 7. Simognathus leiomerus Trouessart, 1894: Line art illustrations of female (A, B, DI) (FDHAL-24/113) and male (C) (FDHAL-24/118): A Idiosoma, dorsal view B Idiosoma, ventral view B' Close up of female genital opening C Idiosoma, ventral view C' Close up of spermatopositor and male genital opening D Gnathosoma, lateral view E Gnathosoma, dorsal view F Leg I, medial view G Leg II, medial view H Leg III, medial view I Leg IV, medial view. Scale bars: AC, FH=100 µm; D, E=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734741" httpUri="https://zenodo.org/record/14734741/files/figure.png" pageId="10" pageNumber="161">Figs. 7FI</figureCitation>
); Leg I: 1, 2, 1, 4, 5, 6; Leg II: 1, 2, 2, 4, 5(2), 6; Leg III, 1, 1, 3, 3, 5(2), 6; Leg IV, 1, 1, 2, 3, 5(2), 6. Tarsus I with 3 dorsal setae, dorsolateral solenidion, and a strong spiniform ventral seta and a pair of pas apically. Tarsus I paired claws and pair of long pas. Ventral seta on tibia I tapering; ventromedial seta smooth. Tarsus II with one ventral seta and pair of pas. Paired claws of tarsi II to IV each with accessory process; tarsi II and III with pecten but tarsi IV has no pecten. Lateral claws on Tarsi I and IV smooth, on tarsi IIIII with combs of well-developed tines.
</paragraph>
<paragraph id="22249709FF8DFFD6C5827A54A4F9F800" blockId="10.[151,1437,866,2009]" pageId="10" pageNumber="161">
<emphasis id="10EF4B1BFF8DFFD6C5827A54A175F83C" bold="true" box="[199,276,1586,1613]" italics="true" pageId="10" pageNumber="161">Males.</emphasis>
Male is similar to female except for the genital plate (
<figureCitation id="BAA08B8CFF8DFFD6C61C7A54A3CEF83C" box="[857,943,1586,1613]" captionStart="FIGURE 7" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[191,1396,186,1846]" captionTargetId="figure-16@11.[187,1401,181,1850]" captionTargetPageId="11" captionText="FIGURE 7. Simognathus leiomerus Trouessart, 1894: Line art illustrations of female (A, B, DI) (FDHAL-24/113) and male (C) (FDHAL-24/118): A Idiosoma, dorsal view B Idiosoma, ventral view B' Close up of female genital opening C Idiosoma, ventral view C' Close up of spermatopositor and male genital opening D Gnathosoma, lateral view E Gnathosoma, dorsal view F Leg I, medial view G Leg II, medial view H Leg III, medial view I Leg IV, medial view. Scale bars: AC, FH=100 µm; D, E=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734741" httpUri="https://zenodo.org/record/14734741/files/figure.png" pageId="10" pageNumber="161">Fig. 7C</figureCitation>
). GO surrounded by 18 pgs. Spermatopositor 62 long, extending slightly beyond the ring of pgs. Three pairs of sgs are present (
<figureCitation id="BAA08B8CFF8DFFD6C1697A30A4E0F800" box="[1068,1153,1622,1649]" captionStart="FIGURE 7" captionStartId="11.[151,250,1874,1898]" captionTargetBox="[191,1396,186,1846]" captionTargetId="figure-16@11.[187,1401,181,1850]" captionTargetPageId="11" captionText="FIGURE 7. Simognathus leiomerus Trouessart, 1894: Line art illustrations of female (A, B, DI) (FDHAL-24/113) and male (C) (FDHAL-24/118): A Idiosoma, dorsal view B Idiosoma, ventral view B' Close up of female genital opening C Idiosoma, ventral view C' Close up of spermatopositor and male genital opening D Gnathosoma, lateral view E Gnathosoma, dorsal view F Leg I, medial view G Leg II, medial view H Leg III, medial view I Leg IV, medial view. Scale bars: AC, FH=100 µm; D, E=50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734741" httpUri="https://zenodo.org/record/14734741/files/figure.png" pageId="10" pageNumber="161">Fig. 7C</figureCitation>
<emphasis id="10EF4B1BFF8DFFD6C1C47A3EA4E9F801" bold="true" box="[1153,1160,1624,1648]" pageId="10" pageNumber="161">'</emphasis>
).
</paragraph>
<paragraph id="22249709FF8DFFD6C5827A1CA468F971" blockId="10.[151,1437,866,2009]" pageId="10" pageNumber="161">
<emphasis id="10EF4B1BFF8DFFD6C5827A1CA10FF8E4" bold="true" box="[199,366,1658,1685]" italics="true" pageId="10" pageNumber="161">Deutonymphs.</emphasis>
Dorsal and ventral plates covered by foveate but the plates less developed than adults (
<figureCitation id="BAA08B8CFF8DFFD6C0217A1CA0B5F8C8" captionStart="FIGURE 8" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[193,1396,186,1846]" captionTargetId="figure-27@12.[189,1398,181,1850]" captionTargetPageId="12" captionText="FIGURE 8. Simognathus leiomerus Trouessart, 1894: Line art illustrations of protonymph (AG) (FDHAL-24/122) and deutonymph (H, I) (FDHAL-24/121) A Idiosoma, dorsal view A'Close up of integument and surrounding area of OC A' Close up of integument B Idiosoma, ventral view B' &amp; B&quot;Close up of genital plate and near integument C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view I' Close up of genital plate Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734743" httpUri="https://zenodo.org/record/14734743/files/figure.png" pageId="10" pageNumber="161">Figs. 8H, I</figureCitation>
;
<figureCitation id="BAA08B8CFF8DFFD6C5A57AF9A141F8C8" box="[224,288,1695,1721]" captionStart="FIGURE 9" captionStartId="13.[151,250,1113,1137]" captionTargetBox="[163,1426,193,1081]" captionTargetId="figure-16@13.[151,1435,181,1089]" captionTargetPageId="13" captionText="FIGURE 9. Simognathus leiomerus Trouessart, 1894: Light-microscope images of female (A, B) (FDHAL-24/113) male (C, D) (FDHAL-24/118) deutonmyph (E, F) (FDHAL-24/121) and protonymph (G, H) (FDHAL-24/122) A Idiosoma, dorsal view B Idiosoma, ventral view C Idiosoma, dorsal view D Idiosoma, ventral view E Idiosoma, dorsal view F Idiosoma, ventral view G Idiosoma, dorsal view H Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734745" httpUri="https://zenodo.org/record/14734745/files/figure.png" pageId="10" pageNumber="161">9E, F</figureCitation>
). Pair of ds-1 on AD. OC reduced to small oblong subcuticular platelet. Pairs of ds-2, ds-3 and ds-
<quantity id="E5633AECFF8DFFD6C0347AF8A5FDF8C9" box="[1393,1436,1694,1720]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="161" unit="in" value="4.0">4 in</quantity>
membraneous integument. Ds-5 and ds-6 on PD. AE with two pairs of setae. PE with one dorsal and two ventral setae. GP with a single pair of pgs (
<figureCitation id="BAA08B8CFF8DFFD6C7617A80A206F970" box="[548,615,1766,1793]" captionStart="FIGURE 8" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[193,1396,186,1846]" captionTargetId="figure-27@12.[189,1398,181,1850]" captionTargetPageId="12" captionText="FIGURE 8. Simognathus leiomerus Trouessart, 1894: Line art illustrations of protonymph (AG) (FDHAL-24/122) and deutonymph (H, I) (FDHAL-24/121) A Idiosoma, dorsal view A'Close up of integument and surrounding area of OC A' Close up of integument B Idiosoma, ventral view B' &amp; B&quot;Close up of genital plate and near integument C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view I' Close up of genital plate Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734743" httpUri="https://zenodo.org/record/14734743/files/figure.png" pageId="10" pageNumber="161">Fig. 8</figureCitation>
I). Leg chaetotaxy similar to female.
</paragraph>
<paragraph id="22249709FF8DFFD6C5827B6CA3FEF9A9" blockId="10.[151,1437,866,2009]" pageId="10" pageNumber="161">
<emphasis id="10EF4B1BFF8DFFD6C5827B6CA13CF954" bold="true" box="[199,349,1802,1829]" italics="true" pageId="10" pageNumber="161">Protonymph.</emphasis>
Dorsal and ventral plates covered by foveate but the plates less developed than adults (
<figureCitation id="BAA08B8CFF8DFFD6C0217B6CA0BCF938" captionStart="FIGURE 8" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[193,1396,186,1846]" captionTargetId="figure-27@12.[189,1398,181,1850]" captionTargetPageId="12" captionText="FIGURE 8. Simognathus leiomerus Trouessart, 1894: Line art illustrations of protonymph (AG) (FDHAL-24/122) and deutonymph (H, I) (FDHAL-24/121) A Idiosoma, dorsal view A'Close up of integument and surrounding area of OC A' Close up of integument B Idiosoma, ventral view B' &amp; B&quot;Close up of genital plate and near integument C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view I' Close up of genital plate Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734743" httpUri="https://zenodo.org/record/14734743/files/figure.png" pageId="10" pageNumber="161">Figs. 8AC</figureCitation>
;
<figureCitation id="BAA08B8CFF8DFFD6C5AD7B49A151F938" box="[232,304,1839,1865]" captionStart="FIGURE 9" captionStartId="13.[151,250,1113,1137]" captionTargetBox="[163,1426,193,1081]" captionTargetId="figure-16@13.[151,1435,181,1089]" captionTargetPageId="13" captionText="FIGURE 9. Simognathus leiomerus Trouessart, 1894: Light-microscope images of female (A, B) (FDHAL-24/113) male (C, D) (FDHAL-24/118) deutonmyph (E, F) (FDHAL-24/121) and protonymph (G, H) (FDHAL-24/122) A Idiosoma, dorsal view B Idiosoma, ventral view C Idiosoma, dorsal view D Idiosoma, ventral view E Idiosoma, dorsal view F Idiosoma, ventral view G Idiosoma, dorsal view H Idiosoma, ventral view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734745" httpUri="https://zenodo.org/record/14734745/files/figure.png" pageId="10" pageNumber="161">9G, H</figureCitation>
). Pair of ds-1 on AD. OC reduced to small oblong subcuticular platelet. Pairs of ds-2, ds-3 and ds-
<quantity id="E5633AECFF8DFFD6C0377B48A5FDF939" box="[1394,1436,1838,1864]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="161" unit="in" value="4.0">4 in</quantity>
membraneous integument (
<figureCitation id="BAA08B8CFF8DFFD6C48D7B34A27FF91C" box="[456,542,1874,1901]" captionStart="FIGURE 8" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[193,1396,186,1846]" captionTargetId="figure-27@12.[189,1398,181,1850]" captionTargetPageId="12" captionText="FIGURE 8. Simognathus leiomerus Trouessart, 1894: Line art illustrations of protonymph (AG) (FDHAL-24/122) and deutonymph (H, I) (FDHAL-24/121) A Idiosoma, dorsal view A'Close up of integument and surrounding area of OC A' Close up of integument B Idiosoma, ventral view B' &amp; B&quot;Close up of genital plate and near integument C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view I' Close up of genital plate Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734743" httpUri="https://zenodo.org/record/14734743/files/figure.png" pageId="10" pageNumber="161">Fig. 8A</figureCitation>
<emphasis id="10EF4B1BFF8DFFD6C75B7B32A244F91D" bold="true" box="[542,549,1876,1900]" pageId="10" pageNumber="161">'</emphasis>
). Ds-5 and ds-6 on PD. AE with two pairs of setae. PE with one dorsal and two ventral setae. GP much smaller than in female. No setae on genital plate (
<figureCitation id="BAA08B8CFF8DFFD6C6807B10A478F9E0" box="[965,1049,1910,1937]" captionStart="FIGURE 8" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[193,1396,186,1846]" captionTargetId="figure-27@12.[189,1398,181,1850]" captionTargetPageId="12" captionText="FIGURE 8. Simognathus leiomerus Trouessart, 1894: Line art illustrations of protonymph (AG) (FDHAL-24/122) and deutonymph (H, I) (FDHAL-24/121) A Idiosoma, dorsal view A'Close up of integument and surrounding area of OC A' Close up of integument B Idiosoma, ventral view B' &amp; B&quot;Close up of genital plate and near integument C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view I' Close up of genital plate Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734743" httpUri="https://zenodo.org/record/14734743/files/figure.png" pageId="10" pageNumber="161">Fig. 8B</figureCitation>
<emphasis id="10EF4B1BFF8DFFD6C15C7B1EA441F9E1" bold="true" box="[1049,1056,1912,1936]" pageId="10" pageNumber="161">'</emphasis>
). Leg chaetotaxy, from trochanter to tarsus (solenidia and famulus excluded, pas included) (
<figureCitation id="BAA08B8CFF8DFFD6C6667BFCA3CDF9C4" box="[803,940,1946,1973]" captionStart="FIGURE 8" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[193,1396,186,1846]" captionTargetId="figure-27@12.[189,1398,181,1850]" captionTargetPageId="12" captionText="FIGURE 8. Simognathus leiomerus Trouessart, 1894: Line art illustrations of protonymph (AG) (FDHAL-24/122) and deutonymph (H, I) (FDHAL-24/121) A Idiosoma, dorsal view A'Close up of integument and surrounding area of OC A' Close up of integument B Idiosoma, ventral view B' &amp; B&quot;Close up of genital plate and near integument C Gnathosoma, lateral view D Leg I, medial view E Leg II, medial view F Leg III, medial view G Leg IV, medial view H Idiosoma, dorsal view I Idiosoma, ventral view I' Close up of genital plate Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14734743" httpUri="https://zenodo.org/record/14734743/files/figure.png" pageId="10" pageNumber="161">Figs. 8DG</figureCitation>
); Leg I: 1, 1, 1, 4, 4, 4; Leg II: 1, 1, 2, 3, 4, 45; Leg III, 1, 1, 2, 3, 4, 5; Leg IV, 0, 2, 3, 4, 4. No pecten on claws.
</paragraph>
</subSubSection>
<caption id="76E4C781FF8CFFD7C5D27B34A09FF98A" ID-DOI="http://doi.org/10.5281/zenodo.14734741" ID-Zenodo-Dep="14734741" httpUri="https://zenodo.org/record/14734741/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,250,1874,1898]" targetBox="[191,1396,186,1846]" targetPageId="11" targetType="figure">
<paragraph id="22249709FF8CFFD7C5D27B34A09FF98A" blockId="11.[151,1437,1874,2043]" pageId="11" pageNumber="162">
<emphasis id="10EF4B1BFF8CFFD7C5D27B34A175F91A" bold="true" box="[151,276,1874,1899]" pageId="11" pageNumber="162">FIGURE 7.</emphasis>
<taxonomicName id="E59BEC8AFF8CFFD7C45E7B35A2DCF91A" authority="Trouessart, 1894" authorityName="Trouessart" authorityYear="1894" box="[283,701,1875,1899]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF8CFFD7C45E7B35A268F91A" box="[283,521,1875,1899]" italics="true" pageId="11" pageNumber="162">Simognathus leiomerus</emphasis>
Trouessart, 1894
</taxonomicName>
: Line art illustrations of female
<emphasis id="10EF4B1BFF8CFFD7C1437B34A41FF91B" bold="true" box="[1030,1150,1874,1899]" pageId="11" pageNumber="162">(A, B, DI)</emphasis>
(FDHAL-24/113) and male
<emphasis id="10EF4B1BFF8CFFD7C5D27B10A0DBF9FF" bold="true" box="[151,186,1910,1934]" pageId="11" pageNumber="162">(C)</emphasis>
(FDHAL-24/118):
<emphasis id="10EF4B1BFF8CFFD7C4C67B10A1F5F9FF" bold="true" box="[387,404,1910,1934]" pageId="11" pageNumber="162">A</emphasis>
Idiosoma, dorsal view
<emphasis id="10EF4B1BFF8CFFD7C7C07B11A2F7F9FE" bold="true" box="[645,662,1911,1935]" pageId="11" pageNumber="162">B</emphasis>
Idiosoma, ventral view
<emphasis id="10EF4B1BFF8CFFD7C6CA7B10A3C6F9FF" bold="true" box="[911,935,1910,1934]" pageId="11" pageNumber="162">B'</emphasis>
Close up of female genital opening
<emphasis id="10EF4B1BFF8CFFD7C05B7B10A551F9FF" bold="true" box="[1310,1328,1910,1934]" pageId="11" pageNumber="162">C</emphasis>
Idiosoma, ventral view
<emphasis id="10EF4B1BFF8CFFD7C45E7BFCA155F9C3" bold="true" box="[283,308,1946,1970]" pageId="11" pageNumber="162">C'</emphasis>
Close up of spermatopositor and male genital opening
<emphasis id="10EF4B1BFF8CFFD7C6277BFDA315F9C2" bold="true" box="[866,884,1947,1971]" pageId="11" pageNumber="162">D</emphasis>
Gnathosoma, lateral view
<emphasis id="10EF4B1BFF8CFFD7C1C17BFDA4F4F9C2" bold="true" box="[1156,1173,1947,1971]" pageId="11" pageNumber="162">E</emphasis>
Gnathosoma, dorsal view
<emphasis id="10EF4B1BFF8CFFD7C5D27BD9A0C7F9A6" bold="true" box="[151,166,1983,2007]" pageId="11" pageNumber="162">F</emphasis>
Leg I, medial view
<emphasis id="10EF4B1BFF8CFFD7C4317BD8A1E9F9A7" bold="true" box="[372,392,1982,2006]" pageId="11" pageNumber="162">G</emphasis>
Leg II, medial view
<emphasis id="10EF4B1BFF8CFFD7C71B7BD9A213F9A6" bold="true" box="[606,626,1983,2007]" pageId="11" pageNumber="162">H</emphasis>
Leg III, medial view
<emphasis id="10EF4B1BFF8CFFD7C6157BD9A33BF9A6" bold="true" box="[848,858,1983,2007]" pageId="11" pageNumber="162">I</emphasis>
Leg IV, medial view. Scale bars: AC, FH=100 µm; D, E=50 µm.
</paragraph>
</caption>
<caption id="76E4C781FF8BFFD0C5D27B34A28FF98A" ID-DOI="http://doi.org/10.5281/zenodo.14734743" ID-Zenodo-Dep="14734743" httpUri="https://zenodo.org/record/14734743/files/figure.png" pageId="12" pageNumber="163" startId="12.[151,250,1874,1898]" targetBox="[193,1396,186,1846]" targetPageId="12" targetType="figure">
<paragraph id="22249709FF8BFFD0C5D27B34A28FF98A" blockId="12.[151,1437,1874,2043]" pageId="12" pageNumber="163">
<emphasis id="10EF4B1BFF8BFFD0C5D27B34A179F91A" bold="true" box="[151,280,1874,1899]" pageId="12" pageNumber="163">FIGURE 8.</emphasis>
<taxonomicName id="E59BEC8AFF8BFFD0C4667B35A2B0F91A" authority="Trouessart, 1894" authorityName="Trouessart" authorityYear="1894" box="[291,721,1875,1899]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF8BFFD0C4667B35A274F91A" box="[291,533,1875,1899]" italics="true" pageId="12" pageNumber="163">Simognathus leiomerus</emphasis>
Trouessart, 1894
</taxonomicName>
: Line art illustrations of protonymph
<emphasis id="10EF4B1BFF8BFFD0C12F7B34A4CCF91B" bold="true" box="[1130,1197,1874,1898]" pageId="12" pageNumber="163">(AG)</emphasis>
(FDHAL-24/122) and deutonymph
<emphasis id="10EF4B1BFF8BFFD0C4597B10A139F9FF" bold="true" box="[284,344,1910,1934]" pageId="12" pageNumber="163">(H, I)</emphasis>
(FDHAL-24/121)
<emphasis id="10EF4B1BFF8BFFD0C75D7B10A248F9FF" bold="true" box="[536,553,1910,1934]" pageId="12" pageNumber="163">A</emphasis>
Idiosoma, dorsal view
<emphasis id="10EF4B1BFF8BFFD0C6507B10A34FF9FF" bold="true" box="[789,814,1910,1934]" pageId="12" pageNumber="163">A'</emphasis>
Close up of integument and surrounding area of OC
<emphasis id="10EF4B1BFF8BFFD0C0067B10A53DF9FF" bold="true" box="[1347,1372,1910,1934]" pageId="12" pageNumber="163">A'</emphasis>
Close up of integument
<emphasis id="10EF4B1BFF8BFFD0C40E7BFDA13DF9C2" bold="true" box="[331,348,1947,1971]" pageId="12" pageNumber="163">B</emphasis>
Idiosoma, ventral view
<emphasis id="10EF4B1BFF8BFFD0C7177BFCA2CAF9C3" bold="true" box="[594,683,1946,1971]" pageId="12" pageNumber="163">B' &amp; B&quot;</emphasis>
Close up of genital plate and near integument
<emphasis id="10EF4B1BFF8BFFD0C1C57BFCA4F3F9C3" bold="true" box="[1152,1170,1946,1970]" pageId="12" pageNumber="163">C</emphasis>
Gnathosoma, lateral view
<emphasis id="10EF4B1BFF8BFFD0C5D27BD9A0C8F9A6" bold="true" box="[151,169,1983,2007]" pageId="12" pageNumber="163">D</emphasis>
Leg I, medial view
<emphasis id="10EF4B1BFF8BFFD0C4377BD9A1E2F9A6" bold="true" box="[370,387,1983,2007]" pageId="12" pageNumber="163">E</emphasis>
Leg II, medial view
<emphasis id="10EF4B1BFF8BFFD0C7107BD9A205F9A6" bold="true" box="[597,612,1983,2007]" pageId="12" pageNumber="163">F</emphasis>
Leg III, medial view
<emphasis id="10EF4B1BFF8BFFD0C67B7BD8A333F9A7" bold="true" box="[830,850,1982,2006]" pageId="12" pageNumber="163">G</emphasis>
Leg IV, medial view
<emphasis id="10EF4B1BFF8BFFD0C16F7BD9A45FF9A6" bold="true" box="[1066,1086,1983,2007]" pageId="12" pageNumber="163">H</emphasis>
Idiosoma, dorsal view
<emphasis id="10EF4B1BFF8BFFD0C06D7BD9A553F9A6" bold="true" box="[1320,1330,1983,2007]" pageId="12" pageNumber="163">I</emphasis>
Idiosoma, ventral view
<emphasis id="10EF4B1BFF8BFFD0C45E7B84A14DF98B" bold="true" box="[283,300,2018,2042]" pageId="12" pageNumber="163">I'</emphasis>
Close up of genital plate Scale bars: 50 µm.
</paragraph>
</caption>
<caption id="76E4C781FF8AFFD1C5D2783FA305FAAF" ID-DOI="http://doi.org/10.5281/zenodo.14734745" ID-Zenodo-Dep="14734745" httpUri="https://zenodo.org/record/14734745/files/figure.png" pageId="13" pageNumber="164" startId="13.[151,250,1113,1137]" targetBox="[163,1426,193,1081]" targetPageId="13" targetType="figure">
<paragraph id="22249709FF8AFFD1C5D2783FA305FAAF" blockId="13.[151,1437,1113,1246]" pageId="13" pageNumber="164">
<emphasis id="10EF4B1BFF8AFFD1C5D2783FA175FA00" bold="true" box="[151,276,1113,1137]" pageId="13" pageNumber="164">FIGURE 9.</emphasis>
<taxonomicName id="E59BEC8AFF8AFFD1C459783FA2DEFA03" authority="Trouessart, 1894" authorityName="Trouessart" authorityYear="1894" box="[284,703,1113,1138]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF8AFFD1C459783FA26BFA00" box="[284,522,1113,1137]" italics="true" pageId="13" pageNumber="164">Simognathus leiomerus</emphasis>
Trouessart, 1894
</taxonomicName>
: Light-microscope images of female
<emphasis id="10EF4B1BFF8AFFD1C17B783FA4E1FA00" bold="true" box="[1086,1152,1113,1138]" pageId="13" pageNumber="164">(A, B)</emphasis>
(FDHAL-24/113) male
<emphasis id="10EF4B1BFF8AFFD1C03E783FA0D0FAE4" bold="true" pageId="13" pageNumber="164">(C, D)</emphasis>
(FDHAL-24/118) deutonmyph
<emphasis id="10EF4B1BFF8AFFD1C4B3781BA255FAE4" bold="true" box="[502,564,1149,1173]" pageId="13" pageNumber="164">(E, F)</emphasis>
(FDHAL-24/121) and protonymph
<emphasis id="10EF4B1BFF8AFFD1C6E5781BA384FAE4" bold="true" box="[928,997,1149,1173]" pageId="13" pageNumber="164">(G, H)</emphasis>
(FDHAL-24/122)
<emphasis id="10EF4B1BFF8AFFD1C1E0781BA4D7FAE4" bold="true" box="[1189,1206,1149,1173]" pageId="13" pageNumber="164">A</emphasis>
Idiosoma, dorsal view
<emphasis id="10EF4B1BFF8AFFD1C5D278C4A0C9FACB" bold="true" box="[151,168,1186,1210]" pageId="13" pageNumber="164">B</emphasis>
Idiosoma, ventral view
<emphasis id="10EF4B1BFF8AFFD1C4D878C7A1CEFAC8" bold="true" box="[413,431,1185,1209]" pageId="13" pageNumber="164">C</emphasis>
Idiosoma, dorsal view
<emphasis id="10EF4B1BFF8AFFD1C7DE78C4A2CCFACB" bold="true" box="[667,685,1186,1210]" pageId="13" pageNumber="164">D</emphasis>
Idiosoma, ventral view
<emphasis id="10EF4B1BFF8AFFD1C6E778C4A3D2FACB" bold="true" box="[930,947,1186,1210]" pageId="13" pageNumber="164">E</emphasis>
Idiosoma, dorsal view
<emphasis id="10EF4B1BFF8AFFD1C1DA78C4A4CFFACB" bold="true" box="[1183,1198,1186,1210]" pageId="13" pageNumber="164">F</emphasis>
Idiosoma, ventral view
<emphasis id="10EF4B1BFF8AFFD1C5D278A3A0CAFAAC" bold="true" box="[151,171,1221,1245]" pageId="13" pageNumber="164">G</emphasis>
Idiosoma, dorsal view
<emphasis id="10EF4B1BFF8AFFD1C4DD78A0A1CDFAAF" bold="true" box="[408,428,1222,1246]" pageId="13" pageNumber="164">H</emphasis>
Idiosoma, ventral view. Scale bars: 50 µm.
</paragraph>
</caption>
<caption id="76E4C781FF8AFFD1C5DD796BA3F9FB54" box="[152,920,1293,1317]" pageId="13" pageNumber="164" startId="13.[152,244,1293,1317]" targetType="table">
<paragraph id="22249709FF8AFFD1C5DD796BA3F9FB54" blockId="13.[152,920,1293,1317]" box="[152,920,1293,1317]" pageId="13" pageNumber="164">
<emphasis id="10EF4B1BFF8AFFD1C5DD796BA16EFB54" bold="true" box="[152,271,1293,1317]" pageId="13" pageNumber="164">TABLE 4.</emphasis>
Measurements for
<taxonomicName id="E59BEC8AFF8AFFD1C49B796BA2B6FB54" authorityName="Trouessart" authorityYear="1894" box="[478,727,1293,1317]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF8AFFD1C49B796BA2B6FB54" box="[478,727,1293,1317]" italics="true" pageId="13" pageNumber="164">Simognathus leiomerus</emphasis>
</taxonomicName>
(in micrometers).
</paragraph>
</caption>
<paragraph id="22249709FF8AFFD1C5ED795EA516F979" pageId="13" pageNumber="164">
<table id="509B65A9FF8A0023C5ED7953A5EBF979" box="[168,1418,1333,1800]" gridcols="9" gridrows="12" pageId="13" pageNumber="164">
<tr id="9CAB954BFF8A0023C5ED7953A5EBFB21" box="[168,1418,1333,1360]" gridrow="0" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED7953A26DFB21" box="[168,524,1333,1360]" gridcol="0" gridrow="0" pageId="13" pageNumber="164">
<taxonomicName id="E59BEC8AFF8AFFD1C5ED795EA269FB3E" authority="Trouessart" authorityName="Trouessart" authorityYear="1894" box="[168,520,1336,1360]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF8AFFD1C5ED795EA1F4FB21" box="[168,405,1336,1360]" italics="true" pageId="13" pageNumber="164">Simognathus leiomerus</emphasis>
Trouessart
</taxonomicName>
,
</th>
<th id="DF7AFC37FF8A0023C76B7953A29FFB21" box="[558,766,1333,1360]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="13" pageNumber="164">
<emphasis id="10EF4B1BFF8AFFD1C7237953A2DEFB3C" bold="true" box="[614,703,1333,1357]" pageId="13" pageNumber="164">Females</emphasis>
</th>
<th id="DF7AFC37FF8A0023C65E7953A391FB21" box="[795,1008,1333,1360]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="13" pageNumber="164">
<emphasis id="10EF4B1BFF8AFFD1C6217953A3C4FB3C" bold="true" box="[868,933,1333,1357]" pageId="13" pageNumber="164">Males</emphasis>
</th>
<th id="DF7AFC37FF8A0023C1567953A4DCFB21" box="[1043,1213,1333,1360]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="13" pageNumber="164">
<emphasis id="10EF4B1BFF8AFFD1C1677953A4CEFB3C" bold="true" box="[1058,1199,1333,1357]" pageId="13" pageNumber="164">Deutonymph</emphasis>
</th>
<th id="DF7AFC37FF8A0023C1A47953A5EBFB21" box="[1249,1418,1333,1360]" colspan="2" colspanRight="1" gridcol="7" gridrow="0" pageId="13" pageNumber="164">
<emphasis id="10EF4B1BFF8AFFD1C1B67953A51AFB3C" bold="true" box="[1267,1403,1333,1357]" pageId="13" pageNumber="164">Protonymph</emphasis>
</th>
</tr>
<tr id="9CAB954BFF8A0023C5ED793AA5EBFB07" box="[168,1418,1372,1398]" gridrow="1" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED793AA26DFB07" box="[168,524,1372,1398]" gridcol="0" gridrow="1" pageId="13" pageNumber="164">1894</th>
<td id="DF7AFC37FF8A0023C76B793AA2E6FB07" box="[558,647,1372,1398]" gridcol="1" gridrow="1" pageId="13" pageNumber="164">Length</td>
<td id="DF7AFC37FF8A0023C7E0793AA29FFB07" box="[677,766,1372,1398]" gridcol="2" gridrow="1" pageId="13" pageNumber="164">Width</td>
<td id="DF7AFC37FF8A0023C65E793AA315FB07" box="[795,884,1372,1398]" gridcol="3" gridrow="1" pageId="13" pageNumber="164">Length</td>
<td id="DF7AFC37FF8A0023C6D2793AA391FB07" box="[919,1008,1372,1398]" gridcol="4" gridrow="1" pageId="13" pageNumber="164">Width</td>
<td id="DF7AFC37FF8A0023C156793AA43AFB07" box="[1043,1115,1372,1398]" gridcol="5" gridrow="1" pageId="13" pageNumber="164">Length</td>
<td id="DF7AFC37FF8A0023C13A793AA4DCFB07" box="[1151,1213,1372,1398]" gridcol="6" gridrow="1" pageId="13" pageNumber="164">Width</td>
<td id="DF7AFC37FF8A0023C1A4793AA548FB07" box="[1249,1321,1372,1398]" gridcol="7" gridrow="1" pageId="13" pageNumber="164">Length</td>
<td id="DF7AFC37FF8A0023C009793AA5EBFB07" box="[1356,1418,1372,1398]" gridcol="8" gridrow="1" pageId="13" pageNumber="164">Width</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED79E0A5EBFBEF" box="[168,1418,1414,1438]" gridrow="2" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED79E0A26DFBEF" box="[168,524,1414,1438]" gridcol="0" gridrow="2" pageId="13" pageNumber="164">Idiosoma</th>
<td id="DF7AFC37FF8A0023C76B79E0A2E6FBEF" box="[558,647,1414,1438]" gridcol="1" gridrow="2" pageId="13" pageNumber="164">400420</td>
<td id="DF7AFC37FF8A0023C7E079E0A29FFBEF" box="[677,766,1414,1438]" gridcol="2" gridrow="2" pageId="13" pageNumber="164">200230</td>
<td id="DF7AFC37FF8A0023C65E79E0A315FBEF" box="[795,884,1414,1438]" gridcol="3" gridrow="2" pageId="13" pageNumber="164">400410</td>
<td id="DF7AFC37FF8A0023C6D279E0A391FBEF" box="[919,1008,1414,1438]" gridcol="4" gridrow="2" pageId="13" pageNumber="164">210275</td>
<td id="DF7AFC37FF8A0023C15679E0A43AFBEF" box="[1043,1115,1414,1438]" gridcol="5" gridrow="2" pageId="13" pageNumber="164">426</td>
<td id="DF7AFC37FF8A0023C13A79E0A4DCFBEF" box="[1151,1213,1414,1438]" gridcol="6" gridrow="2" pageId="13" pageNumber="164">250</td>
<td id="DF7AFC37FF8A0023C1A479E0A548FBEF" box="[1249,1321,1414,1438]" gridcol="7" gridrow="2" pageId="13" pageNumber="164">325</td>
<td id="DF7AFC37FF8A0023C00979E0A5EBFBEF" box="[1356,1418,1414,1438]" gridcol="8" gridrow="2" pageId="13" pageNumber="164">212</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED79C8A5EBFBB7" box="[168,1418,1454,1478]" gridrow="3" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED79C8A26DFBB7" box="[168,524,1454,1478]" gridcol="0" gridrow="3" pageId="13" pageNumber="164">Gnathosoma</th>
<td id="DF7AFC37FF8A0023C76B79C8A2E6FBB7" box="[558,647,1454,1478]" gridcol="1" gridrow="3" pageId="13" pageNumber="164">125</td>
<td id="DF7AFC37FF8A0023C7E079C8A29FFBB7" box="[677,766,1454,1478]" gridcol="2" gridrow="3" pageId="13" pageNumber="164">135</td>
<td id="DF7AFC37FF8A0023C65E79C8A315FBB7" box="[795,884,1454,1478]" gridcol="3" gridrow="3" pageId="13" pageNumber="164">137</td>
<td id="DF7AFC37FF8A0023C6D279C8A391FBB7" box="[919,1008,1454,1478]" gridcol="4" gridrow="3" pageId="13" pageNumber="164">137</td>
<td id="DF7AFC37FF8A0023C15679C8A43AFBB7" box="[1043,1115,1454,1478]" gridcol="5" gridrow="3" pageId="13" pageNumber="164">125</td>
<td id="DF7AFC37FF8A0023C13A79C8A4DCFBB7" box="[1151,1213,1454,1478]" gridcol="6" gridrow="3" pageId="13" pageNumber="164">123</td>
<td id="DF7AFC37FF8A0023C1A479C8A548FBB7" box="[1249,1321,1454,1478]" gridcol="7" gridrow="3" pageId="13" pageNumber="164">100</td>
<td id="DF7AFC37FF8A0023C00979C8A5EBFBB7" box="[1356,1418,1454,1478]" gridcol="8" gridrow="3" pageId="13" pageNumber="164">115</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED79B0A5EBFB9E" box="[168,1418,1494,1519]" gridrow="4" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED79B0A26DFB9E" box="[168,524,1494,1519]" gridcol="0" gridrow="4" pageId="13" pageNumber="164">AD</th>
<td id="DF7AFC37FF8A0023C76B79B0A2E6FB9E" box="[558,647,1494,1519]" gridcol="1" gridrow="4" pageId="13" pageNumber="164">150</td>
<td id="DF7AFC37FF8A0023C7E079B0A29FFB9E" box="[677,766,1494,1519]" gridcol="2" gridrow="4" pageId="13" pageNumber="164">125</td>
<td id="DF7AFC37FF8A0023C65E79B0A315FB9E" box="[795,884,1494,1519]" gridcol="3" gridrow="4" pageId="13" pageNumber="164">150157</td>
<td id="DF7AFC37FF8A0023C6D279B0A391FB9E" box="[919,1008,1494,1519]" gridcol="4" gridrow="4" pageId="13" pageNumber="164">137150</td>
<td id="DF7AFC37FF8A0023C15679B0A43AFB9E" box="[1043,1115,1494,1519]" gridcol="5" gridrow="4" pageId="13" pageNumber="164">139</td>
<td id="DF7AFC37FF8A0023C13A79B0A4DCFB9E" box="[1151,1213,1494,1519]" gridcol="6" gridrow="4" pageId="13" pageNumber="164">139</td>
<td id="DF7AFC37FF8A0023C1A479B0A548FB9E" box="[1249,1321,1494,1519]" gridcol="7" gridrow="4" pageId="13" pageNumber="164">118</td>
<td id="DF7AFC37FF8A0023C00979B0A5EBFB9E" box="[1356,1418,1494,1519]" gridcol="8" gridrow="4" pageId="13" pageNumber="164">110</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED7998A5EBF866" box="[168,1418,1534,1559]" gridrow="5" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED7998A26DF866" box="[168,524,1534,1559]" gridcol="0" gridrow="5" pageId="13" pageNumber="164">OC</th>
<td id="DF7AFC37FF8A0023C76B7998A2E6F866" box="[558,647,1534,1559]" gridcol="1" gridrow="5" pageId="13" pageNumber="164">61</td>
<td id="DF7AFC37FF8A0023C7E07998A29FF866" box="[677,766,1534,1559]" gridcol="2" gridrow="5" pageId="13" pageNumber="164">6</td>
<td id="DF7AFC37FF8A0023C65E7998A315F866" box="[795,884,1534,1559]" gridcol="3" gridrow="5" pageId="13" pageNumber="164">5070</td>
<td id="DF7AFC37FF8A0023C6D27998A391F866" box="[919,1008,1534,1559]" gridcol="4" gridrow="5" pageId="13" pageNumber="164">7</td>
<td id="DF7AFC37FF8A0023C1567998A43AF866" box="[1043,1115,1534,1559]" gridcol="5" gridrow="5" pageId="13" pageNumber="164">55</td>
<td id="DF7AFC37FF8A0023C13A7998A4DCF866" box="[1151,1213,1534,1559]" gridcol="6" gridrow="5" pageId="13" pageNumber="164">5</td>
<td id="DF7AFC37FF8A0023C1A47998A548F866" box="[1249,1321,1534,1559]" gridcol="7" gridrow="5" pageId="13" pageNumber="164">42</td>
<td id="DF7AFC37FF8A0023C0097998A5EBF866" box="[1356,1418,1534,1559]" gridcol="8" gridrow="5" pageId="13" pageNumber="164">4</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED7A41A5EBF84E" box="[168,1418,1575,1599]" gridrow="6" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED7A41A26DF84E" box="[168,524,1575,1599]" gridcol="0" gridrow="6" pageId="13" pageNumber="164">PD</th>
<td id="DF7AFC37FF8A0023C76B7A41A2E6F84E" box="[558,647,1575,1599]" gridcol="1" gridrow="6" pageId="13" pageNumber="164">200</td>
<td id="DF7AFC37FF8A0023C7E07A41A29FF84E" box="[677,766,1575,1599]" gridcol="2" gridrow="6" pageId="13" pageNumber="164">150</td>
<td id="DF7AFC37FF8A0023C65E7A41A315F84E" box="[795,884,1575,1599]" gridcol="3" gridrow="6" pageId="13" pageNumber="164">200</td>
<td id="DF7AFC37FF8A0023C6D27A41A391F84E" box="[919,1008,1575,1599]" gridcol="4" gridrow="6" pageId="13" pageNumber="164">167175</td>
<td id="DF7AFC37FF8A0023C1567A41A43AF84E" box="[1043,1115,1575,1599]" gridcol="5" gridrow="6" pageId="13" pageNumber="164">150</td>
<td id="DF7AFC37FF8A0023C13A7A41A4DCF84E" box="[1151,1213,1575,1599]" gridcol="6" gridrow="6" pageId="13" pageNumber="164">148</td>
<td id="DF7AFC37FF8A0023C1A47A41A548F84E" box="[1249,1321,1575,1599]" gridcol="7" gridrow="6" pageId="13" pageNumber="164">105</td>
<td id="DF7AFC37FF8A0023C0097A41A5EBF84E" box="[1356,1418,1575,1599]" gridcol="8" gridrow="6" pageId="13" pageNumber="164">100</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED7A29A5EBF816" box="[168,1418,1615,1639]" gridrow="7" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED7A29A26DF816" box="[168,524,1615,1639]" gridcol="0" gridrow="7" pageId="13" pageNumber="164">AE</th>
<td id="DF7AFC37FF8A0023C76B7A29A2E6F816" box="[558,647,1615,1639]" gridcol="1" gridrow="7" pageId="13" pageNumber="164">150</td>
<td id="DF7AFC37FF8A0023C7E07A29A29FF816" box="[677,766,1615,1639]" gridcol="2" gridrow="7" pageId="13" pageNumber="164">162</td>
<td id="DF7AFC37FF8A0023C65E7A29A315F816" box="[795,884,1615,1639]" gridcol="3" gridrow="7" pageId="13" pageNumber="164">150</td>
<td id="DF7AFC37FF8A0023C6D27A29A391F816" box="[919,1008,1615,1639]" gridcol="4" gridrow="7" pageId="13" pageNumber="164">185</td>
<td id="DF7AFC37FF8A0023C1567A29A43AF816" box="[1043,1115,1615,1639]" gridcol="5" gridrow="7" pageId="13" pageNumber="164">157</td>
<td id="DF7AFC37FF8A0023C13A7A29A4DCF816" box="[1151,1213,1615,1639]" gridcol="6" gridrow="7" pageId="13" pageNumber="164">200</td>
<td id="DF7AFC37FF8A0023C1A47A29A548F816" box="[1249,1321,1615,1639]" gridcol="7" gridrow="7" pageId="13" pageNumber="164">130</td>
<td id="DF7AFC37FF8A0023C0097A29A5EBF816" box="[1356,1418,1615,1639]" gridcol="8" gridrow="7" pageId="13" pageNumber="164">190</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED7A11A5EBF8FE" box="[168,1418,1655,1679]" gridrow="8" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED7A11A26DF8FE" box="[168,524,1655,1679]" gridcol="0" gridrow="8" pageId="13" pageNumber="164">PE</th>
<td id="DF7AFC37FF8A0023C76B7A11A2E6F8FE" box="[558,647,1655,1679]" gridcol="1" gridrow="8" pageId="13" pageNumber="164">155</td>
<td id="DF7AFC37FF8A0023C7E07A11A29FF8FE" box="[677,766,1655,1679]" gridcol="2" gridrow="8" pageId="13" pageNumber="164">53</td>
<td id="DF7AFC37FF8A0023C65E7A11A315F8FE" box="[795,884,1655,1679]" gridcol="3" gridrow="8" pageId="13" pageNumber="164">157</td>
<td id="DF7AFC37FF8A0023C6D27A11A391F8FE" box="[919,1008,1655,1679]" gridcol="4" gridrow="8" pageId="13" pageNumber="164">50</td>
<td id="DF7AFC37FF8A0023C1567A11A43AF8FE" box="[1043,1115,1655,1679]" gridcol="5" gridrow="8" pageId="13" pageNumber="164">163</td>
<td id="DF7AFC37FF8A0023C13A7A11A4DCF8FE" box="[1151,1213,1655,1679]" gridcol="6" gridrow="8" pageId="13" pageNumber="164">50</td>
<td id="DF7AFC37FF8A0023C1A47A11A548F8FE" box="[1249,1321,1655,1679]" gridcol="7" gridrow="8" pageId="13" pageNumber="164">138</td>
<td id="DF7AFC37FF8A0023C0097A11A5EBF8FE" box="[1356,1418,1655,1679]" gridcol="8" gridrow="8" pageId="13" pageNumber="164">38</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED7AF9A5EBF8C9" box="[168,1418,1695,1720]" gridrow="9" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED7AF9A26DF8C9" box="[168,524,1695,1720]" gridcol="0" gridrow="9" pageId="13" pageNumber="164">GA</th>
<td id="DF7AFC37FF8A0023C76B7AF9A2E6F8C9" box="[558,647,1695,1720]" gridcol="1" gridrow="9" pageId="13" pageNumber="164">175</td>
<td id="DF7AFC37FF8A0023C7E07AF9A29FF8C9" box="[677,766,1695,1720]" gridcol="2" gridrow="9" pageId="13" pageNumber="164">137</td>
<td id="DF7AFC37FF8A0023C65E7AF9A315F8C9" box="[795,884,1695,1720]" gridcol="3" gridrow="9" pageId="13" pageNumber="164">167</td>
<td id="DF7AFC37FF8A0023C6D27AF9A391F8C9" box="[919,1008,1695,1720]" gridcol="4" gridrow="9" pageId="13" pageNumber="164">135</td>
<td id="DF7AFC37FF8A0023C1567AF9A43AF8C9" box="[1043,1115,1695,1720]" gridcol="5" gridrow="9" pageId="13" pageNumber="164">75</td>
<td id="DF7AFC37FF8A0023C13A7AF9A4DCF8C9" box="[1151,1213,1695,1720]" gridcol="6" gridrow="9" pageId="13" pageNumber="164">73</td>
<td id="DF7AFC37FF8A0023C1A47AF9A548F8C9" box="[1249,1321,1695,1720]" gridcol="7" gridrow="9" pageId="13" pageNumber="164">54</td>
<td id="DF7AFC37FF8A0023C0097AF9A5EBF8C9" box="[1356,1418,1695,1720]" gridcol="8" gridrow="9" pageId="13" pageNumber="164">48</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED7AAEA5EBF891" box="[168,1418,1736,1760]" gridrow="10" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED7AAEA26DF891" box="[168,524,1736,1760]" gridcol="0" gridrow="10" pageId="13" pageNumber="164">GO</th>
<td id="DF7AFC37FF8A0023C76B7AAEA2E6F891" box="[558,647,1736,1760]" gridcol="1" gridrow="10" pageId="13" pageNumber="164">44</td>
<td id="DF7AFC37FF8A0023C7E07AAEA29FF891" box="[677,766,1736,1760]" gridcol="2" gridrow="10" pageId="13" pageNumber="164">22</td>
<td id="DF7AFC37FF8A0023C65E7AAEA315F891" box="[795,884,1736,1760]" gridcol="3" gridrow="10" pageId="13" pageNumber="164">20</td>
<td id="DF7AFC37FF8A0023C6D27AAEA391F891" box="[919,1008,1736,1760]" gridcol="4" gridrow="10" pageId="13" pageNumber="164">10</td>
<td id="DF7AFC37FF8A0023C1567AAEA43AF891" box="[1043,1115,1736,1760]" gridcol="5" gridrow="10" pageId="13" pageNumber="164">-</td>
<td id="DF7AFC37FF8A0023C13A7AAEA4DCF891" box="[1151,1213,1736,1760]" gridcol="6" gridrow="10" pageId="13" pageNumber="164">-</td>
<td id="DF7AFC37FF8A0023C1A47AAEA548F891" box="[1249,1321,1736,1760]" gridcol="7" gridrow="10" pageId="13" pageNumber="164">-</td>
<td id="DF7AFC37FF8A0023C0097AAEA5EBF891" box="[1356,1418,1736,1760]" gridcol="8" gridrow="10" pageId="13" pageNumber="164">-</td>
</tr>
<tr id="9CAB954BFF8A0023C5ED7A96A5EBF979" box="[168,1418,1776,1800]" gridrow="11" pageId="13" pageNumber="164">
<th id="DF7AFC37FF8A0023C5ED7A96A26DF979" box="[168,524,1776,1800]" gridcol="0" gridrow="11" pageId="13" pageNumber="164">AP</th>
<td id="DF7AFC37FF8A0023C76B7A96A2E6F979" box="[558,647,1776,1800]" gridcol="1" gridrow="11" pageId="13" pageNumber="164">-</td>
<td id="DF7AFC37FF8A0023C7E07A96A29FF979" box="[677,766,1776,1800]" gridcol="2" gridrow="11" pageId="13" pageNumber="164">-</td>
<td id="DF7AFC37FF8A0023C65E7A96A315F979" box="[795,884,1776,1800]" gridcol="3" gridrow="11" pageId="13" pageNumber="164">-</td>
<td id="DF7AFC37FF8A0023C6D27A96A391F979" box="[919,1008,1776,1800]" gridcol="4" gridrow="11" pageId="13" pageNumber="164">-</td>
<td id="DF7AFC37FF8A0023C1567A96A43AF979" box="[1043,1115,1776,1800]" gridcol="5" gridrow="11" pageId="13" pageNumber="164">27</td>
<td id="DF7AFC37FF8A0023C13A7A96A4DCF979" box="[1151,1213,1776,1800]" gridcol="6" gridrow="11" pageId="13" pageNumber="164">138</td>
<td id="DF7AFC37FF8A0023C1A47A96A548F979" box="[1249,1321,1776,1800]" gridcol="7" gridrow="11" pageId="13" pageNumber="164">32</td>
<td id="DF7AFC37FF8A0023C0097A96A5EBF979" box="[1356,1418,1776,1800]" gridcol="8" gridrow="11" pageId="13" pageNumber="164">87</td>
</tr>
</table>
</paragraph>
<subSubSection id="6A81C482FF8AFFD3C5827B3AA49FFF6C" lastPageId="15" lastPageNumber="166" pageId="13" pageNumber="164" type="discussion">
<paragraph id="22249709FF8AFFD3C5827B3AA49FFF6C" blockId="13.[151,1436,1883,2018]" lastBlockId="15.[151,1436,151,285]" lastPageId="15" lastPageNumber="166" pageId="13" pageNumber="164">
<emphasis id="10EF4B1BFF8AFFD1C5827B3AA155F907" bold="true" box="[199,308,1884,1910]" pageId="13" pageNumber="164">Remarks</emphasis>
. This species was described by Trouessart, 1894 from Granville (French Atlantic). Later, the species was reported from Roscoff (French Atlantic) from
<quantity id="E5633AECFF8AFFD1C78F7BE6A295F9EB" box="[714,756,1920,1946]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="164" unit="m" value="7.0">7 m</quantity>
depths by
<bibRefCitation id="460AEAF8FF8AFFD1C6297B19A474F9EB" author="Bartsch, I." box="[876,1045,1919,1946]" pageId="13" pageNumber="164" pagination="34 - 45" refId="ref6812" refString="Bartsch, I. (1980) Halacaridae (Acari) aus der Bucht von Morlaix (Bretagne). Acarologia, 21, 34 - 45." type="journal article" year="1980">Bartsch (1980)</bibRefCitation>
. Afterwards, the species was found among amphioxus sand habitat (
<quantity id="E5633AECFF8AFFD1C74B7BC5A25CF9CF" box="[526,573,1955,1982]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="164" unit="m" value="5.0">5 m</quantity>
depth) from French Mediterranean by
<bibRefCitation id="460AEAF8FF8AFFD1C1467BC5A4DAF9CC" author="Monniot, F." box="[1027,1211,1955,1981]" pageId="13" pageNumber="164" pagination="585 - 590" refId="ref8567" refString="Monniot, F. (1961) Simognathus andrei, nouvelle espece d'halacarien recoltee en Mediterranee. Acarologia, 3, 585 - 590." type="journal article" year="1961">Monniot (1961)</bibRefCitation>
. With regard to the external morphological characters, our specimens correspond to specimens from Bay of Morlaix (
<collectingCountry id="5A8CD799FF8AFFD1C1A07BAEA552F993" box="[1253,1331,1992,2018]" name="France" pageId="13" pageNumber="164">France</collectingCountry>
Atlantic) (
<bibRefCitation id="460AEAF8FF88FFD3C5DA7CF1A123FEC0" author="Monniot, F." box="[159,322,151,177]" pageId="15" pageNumber="166" pagination="585 - 590" refId="ref8567" refString="Monniot, F. (1961) Simognathus andrei, nouvelle espece d'halacarien recoltee en Mediterranee. Acarologia, 3, 585 - 590." type="journal article" year="1961">Monniot 1961</bibRefCitation>
;
<bibRefCitation id="460AEAF8FF88FFD3C4087CF1A184FEC0" author="Bartsch, I." box="[333,485,151,177]" pageId="15" pageNumber="166" pagination="34 - 45" refId="ref6812" refString="Bartsch, I. (1980) Halacaridae (Acari) aus der Bucht von Morlaix (Bretagne). Acarologia, 21, 34 - 45." type="journal article" year="1980">Bartsch 1980</bibRefCitation>
). This is the first report of the species from
<collectingCountry id="5A8CD799FF88FFD3C6837CF1A443FEC0" box="[966,1058,151,177]" name="Turkey" pageId="15" pageNumber="166">Türkiye</collectingCountry>
. The species belongs to
<taxonomicName id="E59BEC8AFF88FFD3C06B7CF1A5FDFEC0" authorityName="Trouessart" authorityYear="1894" box="[1326,1436,151,177]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF88FFD3C06B7CF1A5FDFEC0" box="[1326,1436,151,177]" italics="true" pageId="15" pageNumber="166">leiomerus</emphasis>
</taxonomicName>
group. This species group may be recognised by lacking corneae, OC reduced, completely or to an elongate sclerite (
<bibRefCitation id="460AEAF8FF88FFD3C5DA7CB9A159FE88" author="Bartsch, I." box="[159,312,223,249]" pageId="15" pageNumber="166" pagination="135 - 152" refId="ref6910" refString="Bartsch, I. (1994) The genus Simognathus (Acari: Halacaridae), description of six new species from southern Australia and a tabular key to all species. Acarologia, 35, 135 - 152." type="journal article" year="1994">Bartsch 1994</bibRefCitation>
) and additionally
<taxonomicName id="E59BEC8AFF88FFD3C74F7CB9A2F7FE88" authorityName="Trouessart" authorityYear="1894" box="[522,662,223,249]" class="Arachnida" family="Halacaridae" genus="Simognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="leiomerus">
<emphasis id="10EF4B1BFF88FFD3C74F7CB9A2F7FE88" box="[522,662,223,249]" italics="true" pageId="15" pageNumber="166">S. leiomerus</emphasis>
</taxonomicName>
may recognised by: (1) body uniformly coloured, (2) marginal and median AE foveate, (3) AE separated from GA both female and male, (4) no outlying setae in males.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="2FBEA8018B91FAF24C8CD28F21DDEEA6" ID-DOI="10.11646/zootaxa.5570.2.1" ID-ISSN="1175-5326" ID-ZooBank="7177B69B-6132-485D-87C7-4EA807692757" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737737014959" checkinUser="plazi" docAuthor="Rojas-Arias, Laura, Griswold, Terry, Hinojosa-Díaz, Ismael, Morrone, Juan J. &amp; Barajas, Ricardo Ayala" docDate="2025" docId="D838E015FF81B07BFF5DC57897517C42" docLanguage="en" docName="zootaxa.5570.2.1.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heriades mariana Rojas-Arias, Griswold &amp; Ayala. 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="241" masterDocId="2401986DFFA4B053FFCAC76393587A50" masterDocTitle="Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species" masterLastPageNumber="259" masterPageNumber="201" pageNumber="238" updateTime="1737737247067" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="B0B80475D952FFB6F79234B269CC7AAE">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<mods:namePart id="9FCD0DD13DF861F9C58FEDDA481DC8AF">Rojas-Arias, Laura</mods:namePart>
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<mods:affiliation id="48FB9A4A95EBE1730EACD187DC75E826">Posgrado en Ciencias Biológicas, Universidad Nacional Autónoma de México (Posgrado en Ciencias Biológicas, UNAM. Unidad de</mods:affiliation>
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<mods:namePart id="74C9F9BC4D2AB1532C38B7A8DCB6C2CD">Griswold, Terry</mods:namePart>
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<mods:namePart id="FAED27E1253CF3471036276EED8B1B06">Hinojosa-Díaz, Ismael</mods:namePart>
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<mods:namePart id="06D947C0BAA9714BAD323A1C7D9FE064">Morrone, Juan J.</mods:namePart>
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<mods:namePart id="060FBF9BBCFFF14570DBC4DF8170E8D8">Barajas, Ricardo Ayala</mods:namePart>
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<subSubSection id="188B0288FF81B076FF5DC57890337866" box="[151,875,539,566]" pageId="37" pageNumber="238" type="nomenclature">
<paragraph id="502E5103FF81B076FF5DC57890337866" blockId="37.[151,875,539,602]" box="[151,875,539,566]" pageId="37" pageNumber="238">
<heading id="0B66E66FFF81B076FF5DC57890337866" bold="true" box="[151,875,539,566]" fontSize="11" level="1" pageId="37" pageNumber="238" reason="1">
<emphasis id="62E58D11FF81B076FF5DC57890337866" bold="true" box="[151,875,539,566]" pageId="37" pageNumber="238">
<taxonomicName id="97912A80FF81B076FF5DC578905E7865" authority="Rojas-Arias, Griswold &amp; Ayala." authorityName="Rojas-Arias, Griswold &amp; Ayala." authorityYear="2025" box="[151,774,539,566]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="mariana" status="sp. nov.">
<emphasis id="62E58D11FF81B076FF5DC578922B7865" bold="true" box="[151,371,539,566]" italics="true" pageId="37" pageNumber="238">Heriades mariana</emphasis>
Rojas-Arias, Griswold &amp; Ayala.
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<taxonomicNameLabel id="79D6306AFF81B076FCC7C57F90337866" box="[781,875,540,566]" pageId="37" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="188B0288FF81B076FF5DC55C9203780A" box="[151,347,575,602]" pageId="37" pageNumber="238" type="description">
<paragraph id="502E5103FF81B076FF5DC55C9203780A" blockId="37.[151,875,539,602]" box="[151,347,575,602]" pageId="37" pageNumber="238">
(
<figureCitation id="C8AA4D86FF81B076FF55C55C9258780A" box="[159,256,575,602]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="37" pageNumber="238">Figs. 2L</figureCitation>
,
<figureCitation id="C8AA4D86FF81B076FEC6C523920B780A" box="[268,339,576,602]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStartId-0="38.[151,250,1068,1092]" captionStartId-1="39.[151,250,1943,1967]" captionTargetBox-0="[159,1428,157,1033]" captionTargetBox-1="[172,1397,192,1905]" captionTargetId-0="figure-22@38.[151,1436,149,1042]" captionTargetId-1="figure-18@39.[172,1416,179,1917]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionText-0="FIGURE 22. Known distribution records for Heriades mariana sp. nov." captionText-1="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="37" pageNumber="238">2223</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF81B076FF5DC5E490B57E5A" pageId="37" pageNumber="238" type="diagnosis">
<paragraph id="502E5103FF81B076FF5DC5E490B57E5A" blockId="37.[151,1437,647,2041]" pageId="37" pageNumber="238">
<emphasis id="62E58D11FF81B076FF5DC5E4924A78F1" bold="true" box="[151,274,647,673]" pageId="37" pageNumber="238">Diagnosis.</emphasis>
Interantennal distance short (1.21.6 times antennocular distance); T3 with small punctures (
<figureCitation id="C8AA4D86FF81B076FAF9C5E496D678F2" box="[1331,1422,647,674]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="37" pageNumber="238">Fig. 3K</figureCitation>
); tergal fasciae with rows of thin, white hairs (
<figureCitation id="C8AA4D86FF81B076FD5CC5C891B77896" box="[662,751,683,710]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="37" pageNumber="238">Fig. 3L</figureCitation>
).
<emphasis id="62E58D11FF81B076FCCEC5C890667895" box="[772,830,683,709]" italics="true" pageId="37" pageNumber="238">Male</emphasis>
: head notably wider than long; clypeus, supraclypeal area and frons with golden hairs (
<figureCitation id="C8AA4D86FF81B076FDCDC5AC913778BA" box="[519,623,719,746]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="37" pageNumber="238">Fig. 23A</figureCitation>
); posterior margin of S1 truncate with subtriangular, medial emargination (
<figureCitation id="C8AA4D86FF81B076FF55C59093B9795D" box="[159,225,755,781]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="37" pageNumber="238">Fig. 2</figureCitation>
Lb), in profile, with posterior margin projected and with soft ventral angle less than one third of distance from apex to base (
<figureCitation id="C8AA4D86FF81B076FEE4C47492377962" box="[302,367,791,818]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="37" pageNumber="238">Fig. 2</figureCitation>
La); posterior margin of S5 bilobated, with several lines of capitate hairs on top directed upward, and several lines of simple, shorter hairs on subapical margin (
<figureCitation id="C8AA4D86FF81B076FCA2C45890977906" box="[872,975,827,854]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="37" pageNumber="238">Fig. 23E</figureCitation>
); S6 with two subapical acute spikes, in profile with quadrate tip (
<figureCitation id="C8AA4D86FF81B076FE70C43C9178792A" box="[442,544,863,890]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="37" pageNumber="238">Fig. 23F</figureCitation>
).
<emphasis id="62E58D11FF81B076FDFEC43C91D07929" box="[564,648,863,889]" italics="true" pageId="37" pageNumber="238">Female</emphasis>
: mandible on outer surface with diagonal tubercle from base of outer ridge to ventral margin of mandible (
<figureCitation id="C8AA4D86FF81B076FDF9C4E091CF79CE" box="[563,663,899,926]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="37" pageNumber="238">Fig. 23C</figureCitation>
) (similar to
<taxonomicName id="97912A80FF81B076FCEAC4E790F779CD" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[800,943,899,925]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="vulcanica" status="sp. nov.">
<emphasis id="62E58D11FF81B076FCEAC4E790F779CD" box="[800,943,899,925]" italics="true" pageId="37" pageNumber="238">H. vulcanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF81B076FC7CC4E7975479CE" box="[950,1036,900,926]" pageId="37" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
but different from this on body size and color or hairs on posterior T); apical margin of clypeus without emargination, with four to five small, rounded tubercles; clypeus, supraclypeal area and scutum with medium-sized punctures (0.2 times OD) (
<figureCitation id="C8AA4D86FF81B076FB2EC4A8966279B6" box="[1252,1338,971,998]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="37" pageNumber="238">Fig. 3E</figureCitation>
) and T3 with small and sparse punctures (0.1 times OD), with dull surface (
<figureCitation id="C8AA4D86FF81B076FC4CC48C90B87E5A" box="[902,992,1007,1034]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="37" pageNumber="238">Fig. 3K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF81B07BFF0DC370903C783D" lastPageId="40" lastPageNumber="241" pageId="37" pageNumber="238" type="description">
<paragraph id="502E5103FF81B076FF0DC370964B7E7D" blockId="37.[151,1437,647,2041]" box="[199,1299,1043,1070]" pageId="37" pageNumber="238">
<emphasis id="62E58D11FF81B076FF0DC37092FC7E7E" bold="true" box="[199,420,1043,1070]" pageId="37" pageNumber="238">
Description.
<emphasis id="62E58D11FF81B076FEAAC37092C37E7E" bold="true" box="[352,411,1043,1070]" italics="true" pageId="37" pageNumber="238">Male</emphasis>
:
</emphasis>
Body length,
<quantity id="9769FCE6FF81B076FD89C37091C17E7E" box="[579,665,1043,1070]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="37" pageNumber="238" unit="mm" value="5.2">5.2 mm</quantity>
(4.25.3, n=6); forewing length,
<quantity id="9769FCE6FF81B076FBC5C377973D7E7E" box="[1039,1125,1044,1070]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="37" pageNumber="238" unit="mm" value="3.2">3.2 mm</quantity>
(3.23.5, n=6).
</paragraph>
<paragraph id="502E5103FF81B076FF0DC354973D7C52" blockId="37.[151,1437,647,2041]" pageId="37" pageNumber="238">
<emphasis id="62E58D11FF81B076FF0DC354925C7E01" box="[199,260,1079,1105]" italics="true" pageId="37" pageNumber="238">Head</emphasis>
. Length
<quantity id="9769FCE6FF81B076FEA4C35B929F7E02" box="[366,455,1080,1106]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="238" unit="mm" value="1.4">1.4 mm</quantity>
(1.41.5, n=6), width
<quantity id="9769FCE6FF81B076FD05C35490707E02" box="[719,808,1079,1106]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="37" pageNumber="238" unit="mm" value="1.6">1.6 mm</quantity>
(1.61.7, n=6); eyes about 3 times longer than broad, 2.4 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF81B076FCA2C33F90E77E26" box="[872,959,1116,1142]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="37" pageNumber="238" unit="mm" value="0.7">0.7 mm</quantity>
(0.70.8, n=6); upper interocular distance
<quantity id="9769FCE6FF81B076FF5DC3E393B67ECA" box="[151,238,1152,1178]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="238" unit="mm" value="0.9">0.9 mm</quantity>
(0.91, n=6); interantennal distance 1.6 times antennocular distance; lateral ocelli large, 4.7 times DOP in interocellar area; interocellar distance 1.6 times OD, ocellocular distance 1.9 times OD, OPR 1.7 times OD; labrum with basal, medial, wide elevation with one cusp; clypeus 1.5 times broader than long, lower third convex in profile, apical margin with shallow emargination; scape length 0.8 times alveolocellar distance, 2.4 times longer than wide; pedicel as long as broad, about 1.1 times longer than F1 and about 0.6 times longer than F2; flagellum black; F1 broader than long; remaining flagellomeres 1.6 longer than F1; F11 2.6 times longer than broad.
<emphasis id="62E58D11FF81B076FB08C25796197F1D" box="[1218,1345,1332,1357]" italics="true" pageId="37" pageNumber="238">Mesosoma.</emphasis>
Scutum 2.9 times longer than scutellum, 2.7 times longer than tegula, and 4 longer than axilla; intertegular distance
<quantity id="9769FCE6FF81B076FA8DC23B96C57F22" box="[1351,1437,1368,1394]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="37" pageNumber="238" unit="mm" value="1.1">1.1 mm</quantity>
(1.11.2, n=6); metanotum with transverse ridge, in posterior view straight; propodeum with large pits (
<figureCitation id="C8AA4D86FF81B076FAF4C21896D67FC6" box="[1342,1422,1403,1430]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="37" pageNumber="238">Fig. 3I</figureCitation>
),
<quantity id="9769FCE6FF81B076FF5DC2C393A37FEA" box="[151,251,1440,1466]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="238" unit="mm" value="0.08">0.08 mm</quantity>
long; legs black.
<emphasis id="62E58D11FF81B076FE09C2C391677FE9" box="[451,575,1440,1465]" italics="true" pageId="37" pageNumber="238">Metasoma.</emphasis>
T1
<quantity id="9769FCE6FF81B076FDA1C2C3919A7FEA" box="[619,706,1440,1466]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="238" unit="mm" value="1.4">1.4 mm</quantity>
broad (1.21.4, n=6). T3
<quantity id="9769FCE6FF81B076FC2FC2FC97637FEA" box="[997,1083,1439,1466]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="37" pageNumber="238" unit="mm" value="1.6">1.6 mm</quantity>
broad (1.41.6, n=6); posterior margin of S1 truncate with subtriangular, medial emargination (
<figureCitation id="C8AA4D86FF81B076FC90C2A090C37F8D" box="[858,923,1475,1501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="37" pageNumber="238">Fig. 2</figureCitation>
Lb) in profile, with posterior margin projected and with soft ventral angle less than one third of distance from apex to base (
<figureCitation id="C8AA4D86FF81B076FC3FC284976F7C52" box="[1013,1079,1511,1538]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="37" pageNumber="238">Fig. 2</figureCitation>
La).
</paragraph>
<paragraph id="502E5103FF81B076FF0DC168924D7DA9" blockId="37.[151,1437,647,2041]" pageId="37" pageNumber="238">
<emphasis id="62E58D11FF81B076FF0DC16892097C75" box="[199,337,1547,1573]" italics="true" pageId="37" pageNumber="238">Sculpturing.</emphasis>
Clypeus with small (at base 0.2 times OD), contiguous, not coarse punctures; supraclypeal area with punctures as on base of clypeus; frons with bigger and coarser punctures than those on supraclypeal area; frons in front of ocelli punctation sparser than frons; vertex with bigger, slightly sparser punctures than on supraclypeal area, with micropunctures between punctures; hypostomal area with punctation as on clypeus; pronotum anterior surface with punctures bigger, slightly sparser coarser than on clypeus; disc of scutum with punctures (0.2 times OD) as on vertex, with micropunctures between punctures; rest of mesosoma as on scutum except: scutellum with punctures slightly bigger and sparser; mesepisternum with slightly bigger punctures, with micropunctures between punctures; metepisternum with notably smaller punctures, less dense and coarse; T1 with smaller (0.2 times OD), sparser punctures than on scutum; T2 with punctures smaller than on T1; T3 with smaller, less coarse punctures than on T1, semicircular shape, separated by third DOP; T4 with coarser punctures than on T3; T5 with smaller and slightly coarser punctures than on T4; dorsal surface of T6 with bigger, sparser, irregularly shaped punctures as on T5, on posterior surface with sparser punctation, DBP one DOP; sides of S1 with punctation bigger and denser than on T1, on disc process smaller; sides of S2 with punctures sparser than on T1, disc of S2 microreticulate without punctation.
</paragraph>
<caption id="04EE018BFF82B075FF5DC34F902D7E14" box="[151,885,1068,1092]" pageId="38" pageNumber="239" startId="38.[151,250,1068,1092]" targetBox="[159,1428,157,1033]" targetPageId="38" targetType="figure">
<paragraph id="502E5103FF82B075FF5DC34F902D7E14" blockId="38.[151,885,1068,1092]" box="[151,885,1068,1092]" pageId="38" pageNumber="239">
<emphasis id="62E58D11FF82B075FF5DC34F92787E14" bold="true" box="[151,288,1068,1092]" pageId="38" pageNumber="239">FIGURE 22.</emphasis>
Known distribution records for
<taxonomicName id="97912A80FF82B075FDADC34F90787E14" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[615,800,1068,1092]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="mariana" status="sp. nov.">
<emphasis id="62E58D11FF82B075FDADC34F90787E14" box="[615,800,1068,1092]" italics="true" pageId="38" pageNumber="239">Heriades mariana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF82B075FCECC34E902D7E14" box="[806,885,1069,1092]" pageId="38" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="502E5103FF82B075FF0DC310967B7F80" blockId="38.[151,1437,1138,2029]" pageId="38" pageNumber="239">
<emphasis id="62E58D11FF82B075FF0DC31092687EDC" box="[199,304,1139,1164]" italics="true" pageId="38" pageNumber="239">Vestiture.</emphasis>
Sides of clypeus with plumose, golden equally length hairs, directed ventrally, absent on middle vertical line; supraclypeal area with few plumose, thin, golden, short hairs beneath alveoli; frons with plumose, thin, long, golden hairs directed dorsally (
<figureCitation id="C8AA4D86FF82B075FDA4C3D991807E84" box="[622,728,1210,1236]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="38" pageNumber="239">Fig. 23A</figureCitation>
); hypostomal area with short hairs not joining hairs in clypeus, in frontal view not surpassing mandible; dorsolateral angles of pronotum with pubescence not forming continuous, linear band with hairs on anterior margin of scutum, and without tuft of thick, short hairs; disc of scutum with simple, thin, short, scattered, white hairs on its entire surface; mesepisternum with anterior and posterior edge with plumose, thin, long, white hairs absent medially; disc of metepisternum without characteristic tuft of plumose, thick, short, white hairs; tergal fasciae with rows of thin, long, white hairs, T1 to T3 with simple, short, thin, white hairs in punctures; disc of T4 to T6 with simple, long, thick, whitish hairs in punctures increasing in length towards posterior terga (
<figureCitation id="C8AA4D86FF82B075FF14C2D5921D7F80" box="[222,325,1462,1488]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="38" pageNumber="239">Fig. 23B</figureCitation>
); posterior surface of subapical posterior in S1 with several plumose, short golden hairs.
</paragraph>
<paragraph id="502E5103FF82B075FF0DC2B992CC7C30" blockId="38.[151,1437,1138,2029]" pageId="38" pageNumber="239">
<emphasis id="62E58D11FF82B075FF0DC2B992117FA4" box="[199,329,1498,1524]" italics="true" pageId="38" pageNumber="239">
<taxonomicName id="97912A80FF82B075FF0DC2B9921D7FA4" box="[199,325,1498,1524]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="38" pageNumber="239" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
S6 with an obtuse-angled apex, with two subapical acute spikes, in profile with quadrate tip (
<figureCitation id="C8AA4D86FF82B075FAA5C2B993907C48" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="38" pageNumber="239">Fig. 23E</figureCitation>
); capsule in dorsal view
<quantity id="9769FCE6FF82B075FE1DC29D91747C49" box="[471,556,1534,1561]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="239" unit="mm" value="0.9">0.9 mm</quantity>
length, and
<quantity id="9769FCE6FF82B075FD79C29D90507C49" box="[691,776,1534,1561]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="239" unit="mm" value="0.5">0.5 mm</quantity>
width, 2 times longer than wide, gonobase
<quantity id="9769FCE6FF82B075FB2DC29D96127C49" box="[1255,1354,1534,1561]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="239" unit="mm" value="0.09">0.09 mm</quantity>
length, 5 times wider than long, and gonocoxite
<quantity id="9769FCE6FF82B075FD95C14191EC7C6D" box="[607,692,1570,1597]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="38" pageNumber="239" unit="mm" value="0.3">0.3 mm</quantity>
length, gonostylus
<quantity id="9769FCE6FF82B075FC44C14190BC7C6D" box="[910,996,1570,1597]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="38" pageNumber="239" unit="mm" value="0.8">0.8 mm</quantity>
length with acute apex, penis valve
<quantity id="9769FCE6FF82B075FAB3C141939B7C31" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="38" pageNumber="239" unit="mm" value="0.6">0.6 mm</quantity>
length (
<figureCitation id="C8AA4D86FF82B075FED5C12592DF7C30" box="[287,391,1606,1632]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="38" pageNumber="239">Fig. 23G</figureCitation>
).
</paragraph>
<paragraph id="502E5103FF82B075FF0DC10991EF7DBC" blockId="38.[151,1437,1138,2029]" pageId="38" pageNumber="239">
<emphasis id="62E58D11FF82B075FF0DC109927F7CD4" bold="true" box="[199,295,1642,1669]" pageId="38" pageNumber="239">
<emphasis id="62E58D11FF82B075FF0DC10992467CD5" bold="true" box="[199,286,1642,1669]" italics="true" pageId="38" pageNumber="239">Female</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FF82B075FE0BC109914F7CD5" box="[449,535,1642,1669]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="38" pageNumber="239" unit="mm" value="6.8">6.8 mm</quantity>
(4.56.8, n=3); forewing length
<quantity id="9769FCE6FF82B075FC43C10990877CD5" box="[905,991,1642,1669]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="38" pageNumber="239" unit="mm" value="3.6">3.6 mm</quantity>
(3.43.6, n=3). Similar to male except:
<emphasis id="62E58D11FF82B075FF5DC1ED938C7CF8" box="[151,212,1678,1704]" italics="true" pageId="38" pageNumber="239">Head</emphasis>
. Length
<quantity id="9769FCE6FF82B075FEF6C1ED92CC7CF9" box="[316,404,1678,1705]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="38" pageNumber="239" unit="mm" value="1.7">1.7 mm</quantity>
(1.61.7, n=3), width
<quantity id="9769FCE6FF82B075FD52C1ED91A87CF9" box="[664,752,1678,1705]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="38" pageNumber="239" unit="mm" value="1.8">1.8 mm</quantity>
(1.61.8, n=3); eyes about 3.9 times longer than broad, 1.7 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF82B075FCF7C1D190CC7C9D" box="[829,916,1714,1741]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="239" unit="mm" value="0.9">0.9 mm</quantity>
(0.80.9, n=3); upper interocular distance
<quantity id="9769FCE6FF82B075FAB3C1D1939B7CA1" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="38" pageNumber="239" unit="mm" value="1.2">1.2 mm</quantity>
(1.11.2, n=3); interantennal distance 1.2 times antennocular distance; interocellar distance 1.7 times OD, ocellocular distance 1.4 times OD, OPR 1.8 times OD; labrum with one rounded, basal, medial tubercle; mandible on outer surface with outer and condylar ridge parallel with tubercle diagonal from base of ouer ridge, reaching ventral margin of mandible; dorsal margin of mandible with carina; clypeus without emargination and apical margin with five rounded, small, lower tubercles (
<figureCitation id="C8AA4D86FF82B075FDF5C00591FE7DD0" box="[575,678,1894,1920]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="38" pageNumber="239">Fig. 23C</figureCitation>
); scape 2.9 times longer than broad; pedicel 1.4 longer than broad, about 1.6 times longer than F1 and 2.2 longer than F2, remaining flagellomeres 1.3 longer than F1.
<emphasis id="62E58D11FF82B075FAD7C0E896CD7DF4" box="[1309,1429,1931,1956]" italics="true" pageId="38" pageNumber="239">Mesosoma</emphasis>
. Scutum 2.8 times longer than scutellum; intertegular distance
<quantity id="9769FCE6FF82B075FC87C0CD90FB7D99" box="[845,931,1966,1993]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="38" pageNumber="239" unit="mm" value="1.3">1.3 mm</quantity>
(1.21.3, n=3).
<emphasis id="62E58D11FF82B075FB92C0CC97957D98" box="[1112,1229,1967,1992]" italics="true" pageId="38" pageNumber="239">Metasoma</emphasis>
. T1
<quantity id="9769FCE6FF82B075FACAC0CD960E7D99" box="[1280,1366,1966,1993]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="38" pageNumber="239" unit="mm" value="1.4">1.4 mm</quantity>
broad (1.31.4, n=3); T3
<quantity id="9769FCE6FF82B075FEA6C0B1929A7DBD" box="[364,450,2002,2029]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="38" pageNumber="239" unit="mm" value="1.7">1.7 mm</quantity>
broad (1.51.7, n=3).
</paragraph>
<caption id="04EE018BFF83B074FF5DC0F4907A7D83" pageId="39" pageNumber="240" startId="39.[151,250,1943,1967]" targetBox="[172,1397,192,1905]" targetPageId="39" targetType="figure">
<paragraph id="502E5103FF83B074FF5DC0F4907A7D83" blockId="39.[151,1436,1943,2003]" pageId="39" pageNumber="240">
<emphasis id="62E58D11FF83B074FF5DC0F492477DFF" bold="true" box="[151,287,1943,1967]" pageId="39" pageNumber="240">FIGURE 23.</emphasis>
<taxonomicName id="97912A80FF83B074FEEFC0F492857DFF" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[293,477,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="mariana" status="sp. nov.">
<emphasis id="62E58D11FF83B074FEEFC0F492857DFF" box="[293,477,1943,1967]" italics="true" pageId="39" pageNumber="240">Heriades mariana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF83B074FE29C0FB91697DFF" box="[483,561,1944,1967]" pageId="39" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
Male:
<emphasis id="62E58D11FF83B074FDBDC0F491C87DFF" bold="true" box="[631,656,1943,1967]" pageId="39" pageNumber="240">A.</emphasis>
frontal view
<emphasis id="62E58D11FF83B074FCDFC0F490747DFF" bold="true" box="[789,812,1943,1967]" pageId="39" pageNumber="240">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FF83B074FBC3C0F497797DFF" bold="true" box="[1033,1057,1943,1967]" pageId="39" pageNumber="240">C.</emphasis>
frontal view
<emphasis id="62E58D11FF83B074FB6DC0F497987DFF" bold="true" box="[1191,1216,1943,1967]" pageId="39" pageNumber="240">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FF83B074FA4FC0F496C47DFF" bold="true" box="[1413,1436,1943,1967]" pageId="39" pageNumber="240">E.</emphasis>
S5.
<emphasis id="62E58D11FF83B074FF75C0D8938A7D83" bold="true" box="[191,210,1979,2003]" pageId="39" pageNumber="240">F.</emphasis>
S6.
<emphasis id="62E58D11FF83B074FECAC0D892427D83" bold="true" box="[256,282,1979,2003]" pageId="39" pageNumber="240">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FF83B074FDB5C0D891C17D83" bold="true" box="[639,665,1979,2003]" pageId="39" pageNumber="240">H.</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="502E5103FF8CB07BFF0DC7F4964C7872" blockId="40.[151,1437,151,1555]" pageId="40" pageNumber="241">
<emphasis id="62E58D11FF8CB07BFF0DC7F492097AE1" box="[199,337,151,177]" italics="true" pageId="40" pageNumber="241">Sculpturing.</emphasis>
Clypeus with medium-sized (0.2 times OD), contiguous, coarse punctures becoming smaller on lower two thirds, with few micropunctures and space between punctures resembling lines; supraclypeal area with punctures slightly bigger and sparser than base of clypeus: frons with slightly bigger, coarser, and contiguous punctures than supraclypeal area; frons in front of ocelli with slightly bigger punctures on sides, than those on frons; vertex bigger and sparser than on clypeus; gena evenly punctate as on frons in front of ocelli; scutum with slightly smaller and denser punctures than on vertex; rest of mesosoma as on scutum except: scutellum with bigger and sparser punctures; mesepisternum with bigger and slightly sparser punctures, with micropuntures between punctures; metasomal terga bright, microreticulate between punctures; T1 and T2 with smaller punctures (0.2 times OD) than on scutum, sparser (0.33 times DOP), with rounded shape; T3 with punctures large, 0.7 times DOP of scutum, irregularly shaped, and sparser (1.5 times DOP); dorsal surface of T6 with bigger, coarser and denser punctures than on T5; sterna with punctation smaller and sparser than on scutum, microreticulate on posterior margin.
</paragraph>
<paragraph id="502E5103FF8CB07BFF0DC54D903C783D" blockId="40.[151,1437,151,1555]" pageId="40" pageNumber="241">
<emphasis id="62E58D11FF8CB07BFF0DC54D92687817" box="[199,304,558,583]" italics="true" pageId="40" pageNumber="241">Vestiture.</emphasis>
Sides of clypeus with few plumose, short, whitish hairs; frons with few plumose, thin, whitish hairs (
<figureCitation id="C8AA4D86FF8CB07BFF55C530925E783D" box="[159,262,595,621]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="40" pageNumber="241">Fig. 23C</figureCitation>
); gena with plumose, whitish hairs evenly distributed.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8CB07BFF0DC51B97DF7E02" pageId="40" pageNumber="241" type="materials_examined">
<paragraph id="502E5103FF8CB07BFF0DC51B90A679C7" blockId="40.[151,1437,151,1555]" pageId="40" pageNumber="241">
<emphasis id="62E58D11FF8CB07BFF0DC51B922278C2" bold="true" box="[199,378,632,658]" pageId="40" pageNumber="241">Type material:</emphasis>
<materialsCitation id="E0F95B5EFF8CB07BFE49C51B907F78E7" collectingDate="1991-02-12" collectionCode="BBSL" collectorName="F. D. Parker" country="Costa Rica" elevation="19" latitude="10.391666" location="Canas" longLatPrecision="21" longitude="-85.149445" pageId="40" pageNumber="241" specimenCode="BBSL 178527" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="62E58D11FF8CB07BFE49C51B970E78C2" bold="true" box="[387,1110,632,658]" pageId="40" pageNumber="241">
<typeStatus id="8F2AEFA1FF8CB07BFE49C51B92A878C2" box="[387,496,632,658]" pageId="40" pageNumber="241" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="0037F978FF8CB07BFD8EC51B91B478C2" box="[580,748,632,658]" collectionCode="BBSL" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="40" pageNumber="241" type="Museum">BBSL 178527</specimenCode>
).
<collectingCountry id="28861193FF8CB07BFCC9C51B90ED78C2" box="[771,949,632,658]" name="Costa Rica" pageId="40" pageNumber="241">COSTA RICA</collectingCountry>
:
<collectingRegion id="92559FE1FF8CB07BFC0EC51B970A78C2" box="[964,1106,632,658]" country="Costa Rica" name="Guanacaste" pageId="40" pageNumber="241">Guanacaste</collectingRegion>
,
</emphasis>
EJN
<quantity id="9769FCE6FF8CB07BFB50C51B978178C2" box="[1178,1241,632,658]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="40" pageNumber="241" unit="km" value="14.0">14km</quantity>
S
<location id="554E07D8FF8CB07BFB36C51B961B78C2" LSID="urn:lsid:plazi:treatment:D838E015FF81B07BFF5DC57897517C42:554E07D8FF8CB07BFB36C51B961B78C2" box="[1276,1347,632,658]" country="Costa Rica" latitude="10.391666" longLatPrecision="21" longitude="-85.149445" name="Canas" pageId="40" pageNumber="241" stateProvince="Guanacaste">Cañas</location>
, [
<quantity id="9769FCE6FF8CB07BFA92C51B96CF78C3" box="[1368,1431,632,659]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="40" pageNumber="241" unit="m" value="19.0">
<elevation id="DBBCB630FF8CB07BFA92C51B96CF78C3" box="[1368,1431,632,659]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="40" pageNumber="241" unit="m" value="19.0">19 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8CB07BFF5DC5FE927878E7" box="[151,288,669,695]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="40" pageNumber="241" precision="15" seconds="30" value="10.391666">10°2330”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8CB07BFEEEC5FE92E078E8" box="[292,440,669,696]" degrees="85" direction="west" minutes="08" orientation="longitude" pageId="40" pageNumber="241" precision="15" seconds="58" value="-85.149445">85°0858”W</geoCoordinate>
],
<date id="242F77C3FF8CB07BFE0DC5FE911878E7" box="[455,576,669,695]" pageId="40" pageNumber="241" value="1991-02-12">
<collectingDate id="346B8E2BFF8CB07BFE0DC5FE911878E7" box="[455,576,669,695]" pageId="40" pageNumber="241" value="1991-02-12">12/II/1991</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFD82C5FD919078E7" box="[584,712,669,696]" pageId="40" pageNumber="241">F.D. Parker</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFD19C5FE904778E7" box="[723,799,669,695]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="40" pageNumber="241" type="Museum">BBSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8CB07BFCE5C5FE97FE788D" collectingDate="1991-03-16" collectionCode="BBSL" collectorName="F. D. Parker" country="Costa Rica" elevation="19" latitude="10.391666" location="Canas" longLatPrecision="21" longitude="-85.149445" pageId="40" pageNumber="241" specimenCode="BBSL178545" specimenCount="1" stateProvince="Guanacaste" typeStatus="allotype">
<emphasis id="62E58D11FF8CB07BFCE5C5FE96C478E7" bold="true" box="[815,1436,669,695]" pageId="40" pageNumber="241">
<typeStatus id="8F2AEFA1FF8CB07BFCE5C5FE90CC78E7" box="[815,916,669,695]" pageId="40" pageNumber="241" type="allotype">Allotype</typeStatus>
(
<specimenCode id="0037F978FF8CB07BFC55C5FE971B78E7" box="[927,1091,669,695]" collectionCode="BBSL" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="40" pageNumber="241" type="Museum">BBSL178545</specimenCode>
):
<collectingCountry id="28861193FF8CB07BFB9EC5FE965978E7" box="[1108,1281,669,695]" name="Costa Rica" pageId="40" pageNumber="241">COSTA RICA</collectingCountry>
:
<collectingRegion id="92559FE1FF8CB07BFAC3C5FE96CF78E7" box="[1289,1431,669,695]" country="Costa Rica" name="Guanacaste" pageId="40" pageNumber="241">Guanacaste</collectingRegion>
,
</emphasis>
EJN
<quantity id="9769FCE6FF8CB07BFF18C5A0924A788D" box="[210,274,707,733]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="40" pageNumber="241" unit="km" value="14.0">14km</quantity>
S
<location id="554E07D8FF8CB07BFEFCC5A09225788D" LSID="urn:lsid:plazi:treatment:D838E015FF81B07BFF5DC57897517C42:554E07D8FF8CB07BFEFCC5A09225788D" box="[310,381,707,733]" country="Costa Rica" latitude="10.391666" longLatPrecision="21" longitude="-85.149445" name="Canas" pageId="40" pageNumber="241" stateProvince="Guanacaste">Cañas</location>
, [
<quantity id="9769FCE6FF8CB07BFE59C5A0928C788D" box="[403,468,707,733]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="40" pageNumber="241" unit="m" value="19.0">
<elevation id="DBBCB630FF8CB07BFE59C5A0928C788D" box="[403,468,707,733]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="40" pageNumber="241" unit="m" value="19.0">19 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8CB07BFE29C5A09134788D" box="[483,620,707,733]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="40" pageNumber="241" precision="15" seconds="30" value="10.391666">10°2330”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8CB07BFDBDC5A19052788D" box="[631,778,706,733]" degrees="85" direction="west" minutes="08" orientation="longitude" pageId="40" pageNumber="241" precision="15" seconds="58" value="-85.149445">85°0858”W</geoCoordinate>
],
<date id="242F77C3FF8CB07BFCEBC5A090FB788D" box="[801,931,707,733]" pageId="40" pageNumber="241" value="1991-03-16">
<collectingDate id="346B8E2BFF8CB07BFCEBC5A090FB788D" box="[801,931,707,733]" pageId="40" pageNumber="241" value="1991-03-16">16/III/1991</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFC78C5A09767788D" box="[946,1087,707,733]" pageId="40" pageNumber="241">F. D. Parker</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFB98C5A097C6788D" box="[1106,1182,707,733]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="40" pageNumber="241" type="Museum">BBSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8CB07BFB7EC5A096CE7952" collectingDate="1991-03-16" collectionCode="BBSL" collectorName="F. D. Parker" country="Costa Rica" elevation="19" latitude="10.391666" location="14 km S Canas" longLatPrecision="21" longitude="-85.149445" pageId="40" pageNumber="241" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="62E58D11FF8CB07BFB7EC5A092DA7952" bold="true" pageId="40" pageNumber="241">
<typeStatus id="8F2AEFA1FF8CB07BFB7EC5A09668788D" box="[1204,1328,707,733]" pageId="40" pageNumber="241" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="28861193FF8CB07BFAF5C5A193BB7952" name="Costa Rica" pageId="40" pageNumber="241">COSTA RICA</collectingCountry>
:
<collectingRegion id="92559FE1FF8CB07BFF25C58B92257952" box="[239,381,744,770]" country="Costa Rica" name="Guanacaste" pageId="40" pageNumber="241">Guanacaste</collectingRegion>
,
</emphasis>
EJN
<location id="554E07D8FF8CB07BFE0BC58B913D7952" LSID="urn:lsid:plazi:treatment:D838E015FF81B07BFF5DC57897517C42:554E07D8FF8CB07BFE0BC58B913D7952" box="[449,613,744,770]" country="Costa Rica" latitude="10.391666" longLatPrecision="21" longitude="-85.149445" name="14 km S Canas" pageId="40" pageNumber="241" stateProvince="Guanacaste">
<quantity id="9769FCE6FF8CB07BFE0BC58B91587952" box="[449,512,744,770]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="40" pageNumber="241" unit="km" value="14.0">14km</quantity>
S Cañas
</location>
, [
<quantity id="9769FCE6FF8CB07BFDB2C58B91ED7953" box="[632,693,744,771]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="40" pageNumber="241" unit="m" value="19.0">
<elevation id="DBBCB630FF8CB07BFDB2C58B91ED7953" box="[632,693,744,771]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="40" pageNumber="241" unit="m" value="19.0">19 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8CB07BFD0BC58B90127952" box="[705,842,744,770]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="40" pageNumber="241" precision="15" seconds="30" value="10.391666">10°2330”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8CB07BFC9BC58B90BD7953" box="[849,997,744,771]" degrees="85" direction="west" minutes="08" orientation="longitude" pageId="40" pageNumber="241" precision="15" seconds="58" value="-85.149445">85°0858”W</geoCoordinate>
],
<date id="242F77C3FF8CB07BFC32C58B97237952" box="[1016,1147,744,770]" pageId="40" pageNumber="241" value="1991-03-16">
<collectingDate id="346B8E2BFF8CB07BFC32C58B97237952" box="[1016,1147,744,770]" pageId="40" pageNumber="241" value="1991-03-16">16/III/1991</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFB4CC58A96547952" box="[1158,1292,744,771]" pageId="40" pageNumber="241">F. D. Parker</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFAD1C58B963A7952" box="[1307,1378,744,770]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="40" pageNumber="241" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8CB07BFAA3C58B96D27952" box="[1385,1418,744,770]" count="1" pageId="40" pageNumber="241" type="female">1♀</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF8CB07BFF5DC46E90957977" box="[151,973,781,808]" collectingDate="1989-01-21" collectingDateMax="1989-01-22" collectingDateMin="1989-01-21" collectionCode="BBSL" collectorName="Parker" country="Costa Rica" elevation="103" latitude="10.433056" longLatPrecision="21" longitude="-85.08417" pageId="40" pageNumber="241" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<quantity id="9769FCE6FF8CB07BFF5DC46E93B97978" box="[151,225,781,808]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="40" pageNumber="241" unit="m" value="103.0">
<elevation id="DBBCB630FF8CB07BFF5DC46E93B97978" box="[151,225,781,808]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="40" pageNumber="241" unit="m" value="103.0">103 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8CB07BFF27C46E922E7978" box="[237,374,781,808]" degrees="10" direction="north" minutes="25" orientation="latitude" pageId="40" pageNumber="241" precision="15" seconds="59" value="10.433056">10°2559”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8CB07BFEB4C46E91557978" box="[382,525,781,808]" degrees="85" direction="west" minutes="05" orientation="longitude" pageId="40" pageNumber="241" precision="15" seconds="03" value="-85.08417">85°0503”W</geoCoordinate>
,
<date id="242F77C3FF8CB07BFDD3C46E91F67977" box="[537,686,781,807]" pageId="40" pageNumber="241" value="1989-01-21" valueMax="1989-01-22" valueMin="1989-01-21">
<collectingDate id="346B8E2BFF8CB07BFDD3C46E91F67977" box="[537,686,781,807]" pageId="40" pageNumber="241" value="1989-01-21" valueMax="1989-01-22" valueMin="1989-01-21">21-22/I/1989</collectingDate>
</date>
, F. D.
<collectorName id="FD6434D5FF8CB07BFD32C46E90197977" box="[760,833,781,807]" pageId="40" pageNumber="241">Parker</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFC9BC46E90C07977" box="[849,920,781,807]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="40" pageNumber="241" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8CB07BFC55C46E909B7977" box="[927,963,781,807]" count="1" pageId="40" pageNumber="241" type="male">1♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF8CB07BFC10C46E96CF7977" box="[986,1431,781,808]" collectingDate="1989-02-16" collectingDateMax="1989-02-20" collectingDateMin="1989-02-16" collectionCode="BBSL" collectorName="Parker" country="Costa Rica" pageId="40" pageNumber="241" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<date id="242F77C3FF8CB07BFC10C46E97207977" box="[986,1144,781,807]" pageId="40" pageNumber="241" value="1989-02-16" valueMax="1989-02-20" valueMin="1989-02-16">
<collectingDate id="346B8E2BFF8CB07BFC10C46E97207977" box="[986,1144,781,807]" pageId="40" pageNumber="241" value="1989-02-16" valueMax="1989-02-20" valueMin="1989-02-16">16-20/II/1989</collectingDate>
</date>
, F. D.
<collectorName id="FD6434D5FF8CB07BFB08C46E96537977" box="[1218,1291,781,807]" pageId="40" pageNumber="241">Parker</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFAD1C46E963A7977" box="[1307,1378,781,807]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="40" pageNumber="241" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8CB07BFAA3C46E96D57977" box="[1385,1421,781,807]" count="1" pageId="40" pageNumber="241" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8CB07BFF5DC45092467922" collectingDate="1987-02-17" collectingDateMax="1987-02-27" collectingDateMin="1987-02-17" collectionCode="BBSL" collectorName="W. L. Rubink-Mak. Tr." country="Costa Rica" elevation="19" latitude="10.391666" location="Canas" longLatPrecision="21" longitude="-85.149445" municipality="La Taboga For. Res." pageId="40" pageNumber="241" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<collectingMunicipality id="B04ACB79FF8CB07BFF5DC45092DD791D" box="[151,389,819,845]" pageId="40" pageNumber="241">La Taboga For. Res.</collectingMunicipality>
9kmSW
<location id="554E07D8FF8CB07BFE3CC4509165791D" LSID="urn:lsid:plazi:treatment:D838E015FF81B07BFF5DC57897517C42:554E07D8FF8CB07BFE3CC4509165791D" box="[502,573,819,845]" country="Costa Rica" latitude="10.391666" longLatPrecision="21" longitude="-85.149445" municipality="La Taboga For. Res." name="Canas" pageId="40" pageNumber="241" stateProvince="Guanacaste">Canas</location>
, [
<quantity id="9769FCE6FF8CB07BFD99C45091CC791D" box="[595,660,819,845]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="40" pageNumber="241" unit="m" value="19.0">
<elevation id="DBBCB630FF8CB07BFD99C45091CC791D" box="[595,660,819,845]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="40" pageNumber="241" unit="m" value="19.0">19 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8CB07BFD6EC4509075791D" box="[676,813,819,845]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="40" pageNumber="241" precision="15" seconds="30" value="10.391666">10°2330”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8CB07BFCF2C4519094791D" box="[824,972,818,845]" degrees="85" direction="west" minutes="08" orientation="longitude" pageId="40" pageNumber="241" precision="15" seconds="58" value="-85.149445">85°0858”W</geoCoordinate>
],
<date id="242F77C3FF8CB07BFC29C45097D9791D" box="[995,1153,819,845]" pageId="40" pageNumber="241" value="1987-02-17" valueMax="1987-02-27" valueMin="1987-02-17">
<collectingDate id="346B8E2BFF8CB07BFC29C45097D9791D" box="[995,1153,819,845]" pageId="40" pageNumber="241" value="1987-02-17" valueMax="1987-02-27" valueMin="1987-02-17">17-27/II/1987</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFB5AC45096C5791D" box="[1168,1437,819,845]" pageId="40" pageNumber="241">W. L. Rubink-Mak. Tr.</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFF55C43B93BE7922" box="[159,230,856,882]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="40" pageNumber="241" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8CB07BFF3AC43B924A7922" box="[240,274,856,882]" count="1" pageId="40" pageNumber="241" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8CB07BFEE4C43B90A279C7" collectingDate="1972-02-14" collectionCode="USDA" collectorName="E. R. Heithaus" country="Costa Rica" county="Hacienda" elevation="50" latitude="10.3425" location="El" longLatPrecision="21" longitude="-85.299446" municipality="Canas" pageId="40" pageNumber="241" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Guanacaste" typeStatus="paratype">
<collectingCounty id="B94F298FFF8CB07BFEE4C43B92C07922" box="[302,408,856,882]" pageId="40" pageNumber="241">Hacienda</collectingCounty>
COMELCO
<quantity id="9769FCE6FF8CB07BFDFDC43B91DC7922" box="[567,644,856,882]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="40" pageNumber="241" unit="km" value="24.0">24 km</quantity>
. NW
<collectingMunicipality id="B04ACB79FF8CB07BFD06C43B904B7922" box="[716,787,856,882]" pageId="40" pageNumber="241">Canas</collectingMunicipality>
, Inter-Am Hwy,
<location id="554E07D8FF8CB07BFC2EC43B90A57922" LSID="urn:lsid:plazi:treatment:D838E015FF81B07BFF5DC57897517C42:554E07D8FF8CB07BFC2EC43B90A57922" box="[996,1021,856,882]" country="Costa Rica" county="Hacienda" latitude="10.3425" longLatPrecision="21" longitude="-85.299446" municipality="Canas" name="El" pageId="40" pageNumber="241" stateProvince="Guanacaste">El</location>
<quantity id="9769FCE6FF8CB07BFBC2C43B971F7923" box="[1032,1095,856,883]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="40" pageNumber="241" unit="m" value="50.0">
<elevation id="DBBCB630FF8CB07BFBC2C43B971F7923" box="[1032,1095,856,883]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="40" pageNumber="241" unit="m" value="50.0">50 m</elevation>
</quantity>
, 1130.
<quantity id="9769FCE6FF8CB07BFB55C43B97A27923" box="[1183,1274,856,883]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="40" pageNumber="241" unit="m" value="1200.0">
<elevation id="DBBCB630FF8CB07BFB55C43B97A27923" box="[1183,1274,856,883]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="40" pageNumber="241" unit="m" value="1200.0">1200 m</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FF8CB07BFADBC43B96C47922" box="[1297,1436,856,882]" degrees="10" direction="north" minutes="20" orientation="latitude" pageId="40" pageNumber="241" precision="15" seconds="33" value="10.3425">10°2033”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8CB07BFF5DC41E927379C8" box="[151,299,893,920]" degrees="85" direction="west" minutes="17" orientation="longitude" pageId="40" pageNumber="241" precision="15" seconds="58" value="-85.299446">85°1758”W</geoCoordinate>
], 12, 15/I, 3, 7,
<date id="242F77C3FF8CB07BFE11C41E910C79C7" box="[475,596,893,919]" pageId="40" pageNumber="241" value="1972-02-14">
<collectingDate id="346B8E2BFF8CB07BFE11C41E910C79C7" box="[475,596,893,919]" pageId="40" pageNumber="241" value="1972-02-14">14/II/1972</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFD95C41D905B79C7" box="[607,771,893,920]" pageId="40" pageNumber="241">E. R. Heithaus</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFCDBC41E900679C7" box="[785,862,893,919]" country="USA" name="United States Department of Agriculture" pageId="40" pageNumber="241">USDA</collectionCode>
ARS
<specimenCount id="46979A8AFF8CB07BFC6AC41E909B79C7" box="[928,963,893,919]" count="3" pageId="40" pageNumber="241" type="female">3♀</specimenCount>
<specimenCount id="46979A8AFF8CB07BFC00C41E90A979C7" box="[970,1009,893,919]" count="4" pageId="40" pageNumber="241" type="male">4♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="502E5103FF8CB07BFF0DC4C197DF7E02" blockId="40.[151,1437,151,1555]" pageId="40" pageNumber="241">
<emphasis id="62E58D11FF8CB07BFF0DC4C1916D79EC" bold="true" box="[199,565,930,956]" pageId="40" pageNumber="241">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFF8CB07BFDF1C4C1976079B2" collectingDate="1986-02-15" collectionCode="KUNHM-ENT" collectorName="W. T. Wcislo" country="Costa Rica" elevation="78" latitude="9.777778" location="Eastern Boundary of Reserva Biologica Carara" longLatPrecision="21" longitude="-84.598335" pageId="40" pageNumber="241" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Puntarenas">
<emphasis id="62E58D11FF8CB07BFDF1C4C190DC79ED" bold="true" box="[571,900,930,957]" pageId="40" pageNumber="241">
<collectingCountry id="28861193FF8CB07BFDF1C4C191B279EC" box="[571,746,930,956]" name="Costa Rica" pageId="40" pageNumber="241">COSTA RICA</collectingCountry>
:
<collectingRegion id="92559FE1FF8CB07BFD3FC4C0902779ED" box="[757,895,931,957]" country="Costa Rica" name="Puntarenas" pageId="40" pageNumber="241">Puntarenas</collectingRegion>
,
</emphasis>
<location id="554E07D8FF8CB07BFC41C4C096C179ED" LSID="urn:lsid:plazi:treatment:D838E015FF81B07BFF5DC57897517C42:554E07D8FF8CB07BFC41C4C096C179ED" box="[907,1433,931,957]" country="Costa Rica" latitude="9.777778" longLatPrecision="21" longitude="-84.598335" name="Eastern Boundary of Reserva Biologica Carara" pageId="40" pageNumber="241" stateProvince="Puntarenas">Eastern Boundary of Reserva Biológica Carara</location>
, [
<quantity id="9769FCE6FF8CB07BFF54C4AB938379B3" box="[158,219,968,995]" metricMagnitude="1" metricUnit="m" metricValue="7.8" pageId="40" pageNumber="241" unit="m" value="78.0">
<elevation id="DBBCB630FF8CB07BFF54C4AB938379B3" box="[158,219,968,995]" metricMagnitude="1" metricUnit="m" metricValue="7.8" pageId="40" pageNumber="241" unit="m" value="78.0">78 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8CB07BFF2CC4AB923979B2" box="[230,353,968,994]" degrees="9" direction="north" minutes="46" orientation="latitude" pageId="40" pageNumber="241" precision="15" seconds="40" value="9.777778">9°4640”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8CB07BFEADC4AB92A479B3" box="[359,508,968,995]" degrees="84" direction="west" minutes="35" orientation="longitude" pageId="40" pageNumber="241" precision="15" seconds="54" value="-84.598335">84°3554”W</geoCoordinate>
],
<date id="242F77C3FF8CB07BFDC7C4AB91DF79B3" box="[525,647,968,995]" pageId="40" pageNumber="241" value="1986-02-15">
<collectingDate id="346B8E2BFF8CB07BFDC7C4AB91DF79B3" box="[525,647,968,995]" pageId="40" pageNumber="241" value="1986-02-15">15/II/1986</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFD5AC4AA907879B2" box="[656,800,968,995]" pageId="40" pageNumber="241">W. T. Wcislo</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFCE4C4AB908179B2" box="[814,985,968,994]" pageId="40" pageNumber="241">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFF8CB07BFC15C4AB975A79B2" box="[991,1026,968,994]" count="1" pageId="40" pageNumber="241" type="female">1♀</specimenCount>
<specimenCount id="46979A8AFF8CB07BFBC3C4AB977779B2" box="[1033,1071,968,994]" count="4" pageId="40" pageNumber="241" type="male">4♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8CB07BFB88C4AB91B47E57" collectingDate="1989-04-22" collectionCode="BBSL" collectorName="F. D. Parker" country="Costa Rica" elevation="1236" latitude="9.918056" location="San Jose" longLatPrecision="21" longitude="-84.15389" pageId="40" pageNumber="241" specimenCount="1" specimenCount-male="1" stateProvince="San Jose">
<emphasis id="62E58D11FF8CB07BFB88C4AB97E879B2" bold="true" box="[1090,1200,967,994]" pageId="40" pageNumber="241">
<collectingRegion id="92559FE1FF8CB07BFB88C4AB97F379B2" box="[1090,1195,967,994]" country="Costa Rica" name="San Jose" pageId="40" pageNumber="241">San José</collectingRegion>
,
</emphasis>
[
<quantity id="9769FCE6FF8CB07BFB77C4AB964E79B3" box="[1213,1302,968,995]" metricMagnitude="3" metricUnit="m" metricValue="1.236" pageId="40" pageNumber="241" unit="m" value="1236.0">
<elevation id="DBBCB630FF8CB07BFB77C4AB964E79B3" box="[1213,1302,968,995]" metricMagnitude="3" metricUnit="m" metricValue="1.236" pageId="40" pageNumber="241" unit="m" value="1236.0">1236 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8CB07BFAEBC4AB96C479B3" box="[1313,1436,968,995]" degrees="9" direction="north" minutes="55" orientation="latitude" pageId="40" pageNumber="241" precision="15" seconds="05" value="9.918056">9°5505”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8CB07BFF5DC48E92737E57" box="[151,299,1005,1031]" degrees="84" direction="west" minutes="09" orientation="longitude" pageId="40" pageNumber="241" precision="15" seconds="14" value="-84.15389">84°0914”W</geoCoordinate>
],
<date id="242F77C3FF8CB07BFE8AC48E929C7E57" box="[320,452,1005,1031]" pageId="40" pageNumber="241" value="1989-04-22">
<collectingDate id="346B8E2BFF8CB07BFE8AC48E929C7E57" box="[320,452,1005,1031]" pageId="40" pageNumber="241" value="1989-04-22">22/IV/1989</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFE1BC48D91047E57" box="[465,604,1005,1032]" pageId="40" pageNumber="241">F. D. Parker</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFDA7C48E91EC7E57" box="[621,692,1005,1031]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="40" pageNumber="241" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8CB07BFD77C48E91BA7E57" box="[701,738,1005,1031]" count="1" pageId="40" pageNumber="241" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8CB07BFD30C48E92AD7E7D" collectingDate="1986-05-19" collectionCode="KUNHM-ENT" collectorName="W. T. Wcislo" country="Costa Rica" elevation="1105" latitude="9.935278" location="Escazu" longLatPrecision="21" longitude="-84.113335" pageId="40" pageNumber="241" specimenCount="1" specimenCount-male="1" stateProvince="San Jose">
<location id="554E07D8FF8CB07BFD30C48E90147E57" LSID="urn:lsid:plazi:treatment:D838E015FF81B07BFF5DC57897517C42:554E07D8FF8CB07BFD30C48E90147E57" box="[762,844,1005,1031]" country="Costa Rica" latitude="9.935278" longLatPrecision="21" longitude="-84.113335" name="Escazu" pageId="40" pageNumber="241" stateProvince="San Jose">Escazú</location>
, [
<quantity id="9769FCE6FF8CB07BFCAAC48E90E37E58" box="[864,955,1005,1032]" metricMagnitude="3" metricUnit="m" metricValue="1.105" pageId="40" pageNumber="241" unit="m" value="1105.0">
<elevation id="DBBCB630FF8CB07BFCAAC48E90E37E58" box="[864,955,1005,1032]" metricMagnitude="3" metricUnit="m" metricValue="1.105" pageId="40" pageNumber="241" unit="m" value="1105.0">1105 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8CB07BFC03C48E971C7E58" box="[969,1092,1005,1032]" degrees="9" direction="north" minutes="56" orientation="latitude" pageId="40" pageNumber="241" precision="15" seconds="07" value="9.935278">9°5607”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8CB07BFB87C48E97B97E57" box="[1101,1249,1005,1031]" degrees="84" direction="west" minutes="06" orientation="longitude" pageId="40" pageNumber="241" precision="15" seconds="48" value="-84.113335">84°0648”W</geoCoordinate>
],
<date id="242F77C3FF8CB07BFB3CC48E96287E57" box="[1270,1392,1005,1031]" pageId="40" pageNumber="241" value="1986-05-19">
<collectingDate id="346B8E2BFF8CB07BFB3CC48E96287E57" box="[1270,1392,1005,1031]" pageId="40" pageNumber="241" value="1986-05-19">19/V/1986</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFAB7C48D925C7E7D" pageId="40" pageNumber="241">W. T. Wcislo</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFED9C37092E67E7D" box="[275,446,1043,1069]" pageId="40" pageNumber="241">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFF8CB07BFE0CC37092B37E7D" box="[454,491,1043,1069]" count="1" pageId="40" pageNumber="241" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8CB07BFDC8C371926C7E02" collectingDate="2005-04-01" collectionCode="ECOAB" collectorName="J. Esponda" country="Mexico" elevation="315" location="Mapastepec" pageId="40" pageNumber="241" specimenCount="1" specimenCount-female="1" stateProvince="Chiapas">
<emphasis id="62E58D11FF8CB07BFDC8C37191AA7E7C" bold="true" box="[514,754,1042,1068]" pageId="40" pageNumber="241">
<collectingCountry id="28861193FF8CB07BFDC8C37191257E7C" box="[514,637,1042,1068]" name="Mexico" pageId="40" pageNumber="241">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF8CB07BFD43C37191B57E7C" box="[649,749,1042,1068]" country="Mexico" name="Chiapas" pageId="40" pageNumber="241">Chiapas</collectingRegion>
,
</emphasis>
<location id="554E07D8FF8CB07BFD30C37090DB7E7D" LSID="urn:lsid:plazi:treatment:D838E015FF81B07BFF5DC57897517C42:554E07D8FF8CB07BFD30C37090DB7E7D" box="[762,899,1043,1069]" country="Mexico" name="Mapastepec" pageId="40" pageNumber="241" stateProvince="Chiapas">Mapastepec</location>
,
<quantity id="9769FCE6FF8CB07BFC45C37190817E7D" box="[911,985,1042,1069]" metricMagnitude="2" metricUnit="m" metricValue="3.15" pageId="40" pageNumber="241" unit="m" value="315.0">
<elevation id="DBBCB630FF8CB07BFC45C37190817E7D" box="[911,985,1042,1069]" metricMagnitude="2" metricUnit="m" metricValue="3.15" pageId="40" pageNumber="241" unit="m" value="315.0">315 m</elevation>
</quantity>
, 15.5219 -92.80,
<date id="242F77C3FF8CB07BFB6AC371964D7E7D" box="[1184,1301,1042,1069]" pageId="40" pageNumber="241" value="2005-04-01">
<collectingDate id="346B8E2BFF8CB07BFB6AC371964D7E7D" box="[1184,1301,1042,1069]" pageId="40" pageNumber="241" value="2005-04-01">1/IV/2005</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFAE8C37096C47E7D" box="[1314,1436,1043,1069]" pageId="40" pageNumber="241">J. Esponda</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFF55C35B92587E02" box="[159,256,1080,1106]" pageId="40" pageNumber="241">ECOAB</collectionCode>
<specimenCount id="46979A8AFF8CB07BFECCC35B92707E02" box="[262,296,1080,1106]" count="1" pageId="40" pageNumber="241" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8CB07BFE8AC35B97DA7E02" box="[320,1154,1080,1107]" collectingDate="2014-12-14" collectionCode="ECOAB" collectorName="J. Merida" country="Mexico" elevation="990" location="Tapachula" pageId="40" pageNumber="241" specimenCount="1" specimenCount-female="1" stateProvince="Chiapas">
<location id="554E07D8FF8CB07BFE8AC35B92EE7E02" LSID="urn:lsid:plazi:treatment:D838E015FF81B07BFF5DC57897517C42:554E07D8FF8CB07BFE8AC35B92EE7E02" box="[320,438,1080,1106]" country="Mexico" name="Tapachula" pageId="40" pageNumber="241" stateProvince="Chiapas">Tapachula</location>
,
<quantity id="9769FCE6FF8CB07BFE0BC35B91527E03" box="[449,522,1080,1107]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="40" pageNumber="241" unit="m" value="990.0">
<elevation id="DBBCB630FF8CB07BFE0BC35B91527E03" box="[449,522,1080,1107]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="40" pageNumber="241" unit="m" value="990.0">990 m</elevation>
</quantity>
, 15.13574 -92.21,
<date id="242F77C3FF8CB07BFD16C35B90317E02" box="[732,873,1080,1106]" pageId="40" pageNumber="241" value="2014-12-14">
<collectingDate id="346B8E2BFF8CB07BFD16C35B90317E02" box="[732,873,1080,1106]" pageId="40" pageNumber="241" value="2014-12-14">14/XII/2014</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8CB07BFCBEC35A90867E02" box="[884,990,1080,1107]" pageId="40" pageNumber="241">J. Mérida</collectorName>
(
<collectionCode id="3680C9C6FF8CB07BFC27C35B97167E02" box="[1005,1102,1080,1106]" pageId="40" pageNumber="241">ECOAB</collectionCode>
<specimenCount id="46979A8AFF8CB07BFB9EC35B972E7E02" box="[1108,1142,1080,1106]" count="1" pageId="40" pageNumber="241" type="female">1♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8CB07BFF0DC33E91997EB8" pageId="40" pageNumber="241" type="distribution">
<paragraph id="502E5103FF8CB07BFF0DC33E91997EB8" blockId="40.[151,1437,151,1555]" pageId="40" pageNumber="241">
<emphasis id="62E58D11FF8CB07BFF0DC33E92397E27" bold="true" box="[199,353,1117,1143]" pageId="40" pageNumber="241">Distribution.</emphasis>
Present from
<collectingRegion id="92559FE1FF8CB07BFDC0C33E913F7E27" box="[522,615,1117,1143]" country="Mexico" name="Chiapas" pageId="40" pageNumber="241">Chiapas</collectingRegion>
,
<collectingCountry id="28861193FF8CB07BFDBCC33E91947E27" box="[630,716,1117,1143]" name="Mexico" pageId="40" pageNumber="241">Mexico</collectingCountry>
to
<collectingCountry id="28861193FF8CB07BFD3CC33E902F7E27" box="[758,887,1117,1143]" name="Costa Rica" pageId="40" pageNumber="241">Costa Rica</collectingCountry>
, mostly in the Pacific Lowlands biogeographic province (
<bibRefCitation id="34002CF2FF8CB07BFEC2C3E092A87ECD" author="Morrone, J. J. &amp; Escalante, T. &amp; Rodriguez-Tapia, G. &amp; Carmona, A. &amp; Arana, M. &amp; Mercado-Gomez, J. D." box="[264,496,1155,1181]" pageId="40" pageNumber="241" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. &amp; Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="62E58D11FF8CB07BFEB9C3E092FD7ECD" box="[371,421,1155,1181]" italics="true" pageId="40" pageNumber="241">et al</emphasis>
. 2022
</bibRefCitation>
). Most of the known specimens are from
<collectingCountry id="28861193FF8CB07BFC0EC3E0971A7ECD" box="[964,1090,1155,1181]" name="Costa Rica" pageId="40" pageNumber="241">Costa Rica</collectingCountry>
, but
<specimenCount id="46979A8AFF8CB07BFBB3C3E097AE7ECD" box="[1145,1270,1155,1181]" count="2" pageId="40" pageNumber="241" type="female">two female</specimenCount>
specimens are from
<collectingRegion id="92559FE1FF8CB07BFF1EC3CB92697E92" box="[212,305,1192,1218]" country="Mexico" name="Chiapas" pageId="40" pageNumber="241">Chiapas</collectingRegion>
, in southern
<collectingCountry id="28861193FF8CB07BFE74C3CB914C7E92" box="[446,532,1192,1218]" name="Mexico" pageId="40" pageNumber="241">Mexico</collectingCountry>
(
<figureCitation id="C8AA4D86FF8CB07BFDE8C3CB912E7E92" box="[546,630,1192,1218]" captionStart="FIGURE 22" captionStartId="38.[151,250,1068,1092]" captionTargetBox="[159,1428,157,1033]" captionTargetId="figure-22@38.[151,1436,149,1042]" captionTargetPageId="38" captionText="FIGURE 22. Known distribution records for Heriades mariana sp. nov." pageId="40" pageNumber="241">Fig. 22</figureCitation>
). This species is endemic from Central America, associate to the tropical and cloud forest. Altitudinal range:
<quantity id="9769FCE6FF8CB07BFDEFC3AE91E57EB8" box="[549,701,1229,1256]" metricMagnitude="2" metricUnit="m" metricValue="6.275" metricValueMax="12.36" metricValueMin="0.19" pageId="40" pageNumber="241" unit="m" value="627.5" valueMax="1236.0" valueMin="19.0">19 to 1236 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8CB07BFF0DC39092BC7F07" pageId="40" pageNumber="241" type="etymology">
<paragraph id="502E5103FF8CB07BFF0DC39092BC7F07" blockId="40.[151,1437,151,1555]" pageId="40" pageNumber="241">
<emphasis id="62E58D11FF8CB07BFF0DC39092167F5D" bold="true" box="[199,334,1267,1293]" pageId="40" pageNumber="241">Etymology.</emphasis>
This species is named after melittologist Dra.
<taxonomicName id="97912A80FF8CB07BFC87C39097047F5D" authority="Acuna Cordero" authorityName="Acuna Cordero" box="[845,1116,1267,1293]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="mariana">Mariana Acuña Cordero</taxonomicName>
, who had contributed greatly on the knowledge of native bees from
<collectingCountry id="28861193FF8CB07BFDF2C27B91EC7F62" box="[568,692,1304,1330]" name="Costa Rica" pageId="40" pageNumber="241">Costa Rica</collectingCountry>
. She works in an integral way on ecology, natural history, taxonomy and environmental education.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8CB07BFF0DC20197517C42" pageId="40" pageNumber="241" type="discussion">
<paragraph id="502E5103FF8CB07BFF0DC20197517C42" blockId="40.[151,1437,151,1555]" pageId="40" pageNumber="241">
<emphasis id="62E58D11FF8CB07BFF0DC20192107F2C" bold="true" box="[199,328,1378,1404]" pageId="40" pageNumber="241">Comments</emphasis>
. There is variation on base of labrum, in the size of medial tubercle. This species is similar to
<taxonomicName id="97912A80FF8CB07BFA4BC20093B37FF2" authorityName="Michener" authorityYear="1943" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="62E58D11FF8CB07BFA4BC20093B37FF2" italics="true" pageId="40" pageNumber="241">H. currani</emphasis>
</taxonomicName>
however the male differs by having two subapical spikes on S6 and females with apical margin of clypeus with more than two tubercles all of the same size. On females the tubercle on outer surface of mandible varies from straight to slightly curved. The spikes of S6 on males are similar to
<taxonomicName id="97912A80FF8CB07BFC67C2B097747FBD" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[941,1068,1491,1517]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="amanda" status="sp. nov.">
<emphasis id="62E58D11FF8CB07BFC67C2B097747FBD" box="[941,1068,1491,1517]" italics="true" pageId="40" pageNumber="241">H. amanda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF8CB07BFBFCC2B097D77FBD" box="[1078,1167,1491,1517]" pageId="40" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
but differ in being one structure and not a few hairs together, and in profile having a quadrangular tip.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,610 @@
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<mods:title id="38633B372C6BA65CD989727BAB28D6C9">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<subSubSection id="188B0288FF85B072FF5DC25990677F04" box="[151,831,1338,1365]" pageId="33" pageNumber="234" type="nomenclature">
<paragraph id="502E5103FF85B072FF5DC25990677F04" blockId="33.[151,831,1338,1400]" box="[151,831,1338,1365]" pageId="33" pageNumber="234">
<heading id="0B66E66FFF85B072FF5DC25990677F04" bold="true" box="[151,831,1338,1365]" fontSize="11" level="1" pageId="33" pageNumber="234" reason="1">
<emphasis id="62E58D11FF85B072FF5DC25990677F04" bold="true" box="[151,831,1338,1365]" pageId="33" pageNumber="234">
<taxonomicName id="97912A80FF85B072FF5DC25991827F04" authority="Rojas-Arias, Griswold &amp; Ayala." authorityName="Rojas-Arias, Griswold &amp; Ayala." authorityYear="2025" box="[151,730,1338,1365]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
<emphasis id="62E58D11FF85B072FF5DC259921F7F05" bold="true" box="[151,327,1338,1365]" italics="true" pageId="33" pageNumber="234">Heriades luisa</emphasis>
Rojas-Arias, Griswold &amp; Ayala.
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF85B072FD2BC25990677F04" box="[737,831,1338,1364]" pageId="33" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="188B0288FF85B072FF5DC23D92067F28" box="[151,350,1374,1400]" pageId="33" pageNumber="234" type="description">
<paragraph id="502E5103FF85B072FF5DC23D92067F28" blockId="33.[151,831,1338,1400]" box="[151,350,1374,1400]" pageId="33" pageNumber="234">
(
<figureCitation id="C8AA4D86FF85B072FF55C23D925B7F28" box="[159,259,1374,1400]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="33" pageNumber="234">Figs. 2N</figureCitation>
,
<figureCitation id="C8AA4D86FF85B072FEC5C23D920F7F28" box="[271,343,1374,1400]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStartId-0="34.[151,250,1098,1122]" captionStartId-1="35.[151,250,1943,1967]" captionTargetBox-0="[159,1428,187,1064]" captionTargetBox-1="[161,1406,179,1888]" captionTargetId-0="figure-22@34.[151,1436,179,1072]" captionTargetId-1="figure-18@35.[161,1426,179,1917]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="FIGURE 20. Known distribution records for Heriades luisa sp. nov." captionText-1="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="33" pageNumber="234">2021</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF85B072FF5DC2C597267CB0" pageId="33" pageNumber="234" type="diagnosis">
<paragraph id="502E5103FF85B072FF5DC2C597267CB0" blockId="33.[151,1437,1446,2013]" pageId="33" pageNumber="234">
<emphasis id="62E58D11FF85B072FF5DC2C5924A7F90" bold="true" box="[151,274,1446,1472]" pageId="33" pageNumber="234">Diagnosis.</emphasis>
Clypeus in profile concave and strongly projected on apical margin; clypeus, supraclypeal area and scutum with small punctures (~0.2 times OD); propodeum with large pits (
<figureCitation id="C8AA4D86FF85B072FC6AC2A990A87FB4" box="[928,1008,1482,1508]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="33" pageNumber="234">Fig. 3I</figureCitation>
).
<emphasis id="62E58D11FF85B072FBC2C2A9971A7FB4" box="[1032,1090,1482,1508]" italics="true" pageId="33" pageNumber="234">Male</emphasis>
: with well-defined paraocular pubescence; frons with erected hairs, but clypeus with plumose, thin, short, white hairs, almost transparent in frontal view (
<figureCitation id="C8AA4D86FF85B072FF15C17192117C7C" box="[223,329,1554,1580]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="33" pageNumber="234">Fig. 21A</figureCitation>
); posterior margin of S1 produced to median rounded angle, without emargination (
<figureCitation id="C8AA4D86FF85B072FACDC17196137C7C" box="[1287,1355,1554,1580]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="33" pageNumber="234">Fig. 2</figureCitation>
Nb), in profile, with soft ventral angle less than one third of distance from apex to base (
<figureCitation id="C8AA4D86FF85B072FBE9C155973E7C00" box="[1059,1126,1590,1616]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="33" pageNumber="234">Fig. 2</figureCitation>
Na); posterior margin of S5 with two quadrate lobes, two to three lines of capitate hairs on top of it, and two small tufts of a few simple, thick, short hairs on top of two small, inner, smooth elevations (
<figureCitation id="C8AA4D86FF85B072FCE3C11D90C87CC8" box="[809,912,1662,1688]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="33" pageNumber="234">Fig. 21E</figureCitation>
); S6 with one subapical tuft of a few capitate, thick, short erected (
<figureCitation id="C8AA4D86FF85B072FEB2C1C192837CEC" box="[376,475,1698,1724]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="33" pageNumber="234">Fig. 21F</figureCitation>
).
<emphasis id="62E58D11FF85B072FE27C1C191197CEC" box="[493,577,1698,1724]" italics="true" pageId="33" pageNumber="234">Female</emphasis>
: outer surface of mandible without tubercle; apical margin of clypeus without emargination and five well defined tubercles; apical tooth longer than the rest (
<figureCitation id="C8AA4D86FF85B072FBC1C1A5972A7CB0" box="[1035,1138,1734,1760]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="33" pageNumber="234">Fig. 21C</figureCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="502E5103FF85B072FF0DC18996207D54" blockId="33.[151,1437,1446,2013]" box="[199,1400,1770,1797]" pageId="33" pageNumber="234">
<emphasis id="62E58D11FF85B072FF0DC18992FC7D54" bold="true" box="[199,420,1770,1797]" pageId="33" pageNumber="234">
Description.
<emphasis id="62E58D11FF85B072FEAAC18992C37D55" bold="true" box="[352,411,1770,1797]" italics="true" pageId="33" pageNumber="234">Male</emphasis>
:
</emphasis>
Body length,
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(
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6.7
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7.7 mm
</quantity>
, n=3); forewing length,
<quantity id="9769FCE6FF85B072FB8BC18997CF7D55" box="[1089,1175,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="33" pageNumber="234" unit="mm" value="4.2">4.2 mm</quantity>
(
<quantity id="9769FCE6FF85B072FB6CC18996687D55" box="[1190,1328,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="33" pageNumber="234" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">
4.1
<emphasis id="62E58D11FF85B072FB00C18997807D54" bold="true" box="[1226,1240,1770,1796]" pageId="33" pageNumber="234"></emphasis>
4.2 mm
</quantity>
, n=3).
</paragraph>
<paragraph id="502E5103FF85B071FF0DC06D90227FF7" blockId="33.[151,1437,1446,2013]" lastBlockId="34.[151,1437,1168,1987]" lastPageId="34" lastPageNumber="235" pageId="33" pageNumber="234">
<emphasis id="62E58D11FF85B072FF0DC06D925C7D78" box="[199,260,1806,1832]" italics="true" pageId="33" pageNumber="234">Head</emphasis>
. Length
<quantity id="9769FCE6FF85B072FEA3C06D92E77D79" box="[361,447,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="234" unit="mm" value="1.5">1.5 mm</quantity>
(
<quantity id="9769FCE6FF85B072FE04C06D91017D79" box="[462,601,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="33" pageNumber="234" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.51.8 mm</quantity>
, n=3), width
<quantity id="9769FCE6FF85B072FD3AC06D901E7D79" box="[752,838,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="33" pageNumber="234" unit="mm" value="1.8">1.8 mm</quantity>
(
<quantity id="9769FCE6FF85B072FC9FC06D90B87D79" box="[853,992,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="33" pageNumber="234" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.81.9 mm</quantity>
, n=3); eyes about 3.3 times longer than broad, 1.7 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF85B072FC69C05190A07D1D" box="[931,1016,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="mm" value="0.9">0.9 mm</quantity>
(
<quantity id="9769FCE6FF85B072FBCFC05197D77D1D" box="[1029,1167,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="33" pageNumber="234" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9 mm</quantity>
, n=3); upper interocular distance
<quantity id="9769FCE6FF85B072FF30C03592097D21" box="[250,337,1878,1905]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="33" pageNumber="234" unit="mm" value="1.1">1.1 mm</quantity>
(
<quantity id="9769FCE6FF85B072FEABC035928E7D21" box="[353,470,1878,1905]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="33" pageNumber="234" unit="mm" value="1.05" valueMax="1.1" valueMin="1.0">11.1 mm</quantity>
, n=3); interantennal distance 1.5 times antennocular distance; lateral ocelli large (
<figureCitation id="C8AA4D86FF85B072FAA5C03593E27DC4" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="33" pageNumber="234">Fig. 3C</figureCitation>
), 3.2 times DOP in interocellar area; interocellar distance 1.8 times OD, ocellocular distance 1.8 times OD, OPR 1.8 times OD; labrum with basal, medial, wide elevation with one cusp; clypeus 1.7 times broader than long, in profile strongly convex; apical margin without emargination; scape length 0.9 times alveolocellar distance, 3.2 times longer than wide; pedicel longer than broad, about 1.2 times longer than F1 and about 1.1 times longer than F2; flagellum black, F1 longer than broad, remaining flagellomeres 1.36 longer than F1; F11 2.0 times longer than broad.
<emphasis id="62E58D11FF86B071FF23C3BA92307EA2" box="[233,360,1241,1266]" italics="true" pageId="34" pageNumber="235">Mesosoma.</emphasis>
Scutum 2.31 times longer than scutellum, 3.4 times longer than tegula, and 4.2 longer than axilla; intertegular distance
<quantity id="9769FCE6FF86B071FE07C39E917A7F47" box="[461,546,1277,1303]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="34" pageNumber="235" unit="mm" value="1.3">1.3 mm</quantity>
(
<quantity id="9769FCE6FF86B071FDFAC39E91ED7F47" box="[560,693,1277,1303]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="34" pageNumber="235" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3-1.4 mm</quantity>
, n=3); metanotum with transverse ridge, in posterior view straight; propodeum with large pits (I),
<quantity id="9769FCE6FF86B071FE3AC242910C7F6B" box="[496,596,1313,1339]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="34" pageNumber="235" unit="mm" value="0.08">0.08 mm</quantity>
long; legs black.
<emphasis id="62E58D11FF86B071FCD1C24290CF7F6A" box="[795,919,1313,1338]" italics="true" pageId="34" pageNumber="235">Metasoma.</emphasis>
T1
<quantity id="9769FCE6FF86B071FC09C24397427F6B" box="[963,1050,1312,1339]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="235" unit="mm" value="1.5">1.5 mm</quantity>
broad (
<quantity id="9769FCE6FF86B071FBBAC243965B7F6B" box="[1136,1283,1312,1339]" metricMagnitude="-3" metricUnit="m" metricValue="1.615" metricValueMax="1.7" metricValueMin="1.53" pageId="34" pageNumber="235" unit="mm" value="1.615" valueMax="1.7" valueMin="1.53">1.53-1.7 mm</quantity>
, n=3); T3
<quantity id="9769FCE6FF86B071FAB3C242939B7F0F" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="34" pageNumber="235" unit="mm" value="1.7">1.7 mm</quantity>
broad (
<quantity id="9769FCE6FF86B071FED0C22692F87F0F" box="[282,416,1349,1375]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="34" pageNumber="235" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7-1.9 mm</quantity>
, n=3); T6 with posterior margin not projected; posterior margin of S1 produced to median rounded angle, without emargination (
<figureCitation id="C8AA4D86FF86B071FD80C20B91D67FD2" box="[586,654,1384,1410]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="34" pageNumber="235">Fig. 2</figureCitation>
Nb), in profile, with posterior margin projected and with soft ventral angle less than one third of distance from apex to base (
<figureCitation id="C8AA4D86FF86B071FCCDC2EF90117FF7" box="[775,841,1420,1447]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="34" pageNumber="235">Fig. 2</figureCitation>
Na).
</paragraph>
<caption id="04EE018BFF86B071FF5DC32990087E32" box="[151,848,1098,1122]" pageId="34" pageNumber="235" startId="34.[151,250,1098,1122]" targetBox="[159,1428,187,1064]" targetPageId="34" targetType="figure">
<paragraph id="502E5103FF86B071FF5DC32990087E32" blockId="34.[151,848,1098,1122]" box="[151,848,1098,1122]" pageId="34" pageNumber="235">
<emphasis id="62E58D11FF86B071FF5DC32992787E32" bold="true" box="[151,288,1098,1122]" pageId="34" pageNumber="235">FIGURE 20.</emphasis>
Known distribution records for
<taxonomicName id="97912A80FF86B071FDADC32991A37E32" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[615,763,1098,1122]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
<emphasis id="62E58D11FF86B071FDADC32991A37E32" box="[615,763,1098,1122]" italics="true" pageId="34" pageNumber="235">Heriades luisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF86B071FCC8C32890087E32" box="[770,848,1099,1122]" pageId="34" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="502E5103FF86B071FF0DC2D390EC7D92" blockId="34.[151,1437,1168,1987]" pageId="34" pageNumber="235">
<emphasis id="62E58D11FF86B071FF0DC2D392097F9A" box="[199,337,1456,1482]" italics="true" pageId="34" pageNumber="235">Sculpturing.</emphasis>
Clypeus with small (at base 0.2 times OD), contiguous, not coarse punctures, smaller towards disc and apical margin; supraclypeal area with smaller, denser punctures than those on base of clypeus; frons with bigger and coarser than those on supraclypeal area, and in frons in front of ocelli even bigger than frons; vertex with bigger, sparser and coarser punctures than on supraclypeal area, without micropunctures between punctures; hypostomal area with punctation slightly smaller and sparser than on clypeus; pronotum anterior surface with punctures sparser than on clypeus, denser on dorsolateral angles; disc of scutum with slightly denser punctures (0.2 times OD) than on vertex, without micropunctures between punctures; rest of mesosoma as on scutum except: scutellum with punctures bigger and sparser; mesepisternum with slightly bigger and slightly sparser punctures, irregularly shaped with micropunctures between punctures; metepisternum with notably smaller punctures, less dense and coarse; T1 with smaller (0.2 times OD), slightly sparser punctures; T2 with punctures smaller and sparser than on T1; T3 with smaller, less coarse punctures than on T1, semicircular shape, separated by half DOP; T4 with slightly bigger, denser and coarser punctures than on T3; T5 with smaller punctures than on T4; dorsal surface of T6 with smaller, denser, irregularly shaped punctures than on T5, on posterior surface with bigger and sparser punctation, DBP one third of DOP; sides of S1 with punctation bigger and sparser than on T1, on disc smaller and irregularly shaped; sides of S2 with punctures as on T1, disc of S2 microreticulate without punctation.
</paragraph>
<caption id="04EE018BFF87B070FF5DC0F491A37D83" pageId="35" pageNumber="236" startId="35.[151,250,1943,1967]" targetBox="[161,1406,179,1888]" targetPageId="35" targetType="figure">
<paragraph id="502E5103FF87B070FF5DC0F491A37D83" blockId="35.[151,1436,1943,2003]" pageId="35" pageNumber="236">
<emphasis id="62E58D11FF87B070FF5DC0F492477DFF" bold="true" box="[151,287,1943,1967]" pageId="35" pageNumber="236">FIGURE 21.</emphasis>
<taxonomicName id="97912A80FF87B070FEEEC0F492E07DFF" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[292,440,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
<emphasis id="62E58D11FF87B070FEEEC0F492E07DFF" box="[292,440,1943,1967]" italics="true" pageId="35" pageNumber="236">Heriades luisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF87B070FE74C0FB91547DFF" box="[446,524,1944,1967]" pageId="35" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
Male:
<emphasis id="62E58D11FF87B070FD98C0F491327DFF" bold="true" box="[594,618,1943,1967]" pageId="35" pageNumber="236">A.</emphasis>
frontal view
<emphasis id="62E58D11FF87B070FD3AC0F4905F7DFF" bold="true" box="[752,775,1943,1967]" pageId="35" pageNumber="236">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FF87B070FC29C0F490A37DFF" bold="true" box="[995,1019,1943,1967]" pageId="35" pageNumber="236">C.</emphasis>
frontal view
<emphasis id="62E58D11FF87B070FB4BC0F497C17DFF" bold="true" box="[1153,1177,1943,1967]" pageId="35" pageNumber="236">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FF87B070FA94C0F4962E7DFF" bold="true" box="[1374,1398,1943,1967]" pageId="35" pageNumber="236">E.</emphasis>
S5.
<emphasis id="62E58D11FF87B070FF5DC0D893F27D83" bold="true" box="[151,170,1979,2003]" pageId="35" pageNumber="236">F.</emphasis>
S6.
<emphasis id="62E58D11FF87B070FF12C0D893AB7D83" bold="true" box="[216,243,1979,2003]" pageId="35" pageNumber="236">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FF87B070FD9DC0D891297D83" bold="true" box="[599,625,1979,2003]" pageId="35" pageNumber="236">H.</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="502E5103FF80B077FF0DC7F4961A7B81" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
<emphasis id="62E58D11FF80B077FF0DC7F492687AE0" box="[199,304,151,176]" italics="true" pageId="36" pageNumber="237">Vestiture.</emphasis>
Sides of clypeus with plumose, white hairs, of similar length, directed ventrally, scattered on middle vertical line; supraclypeal area with few plumose, thin, golden, short hairs; frons with plumose, thin, long, white hairs directed dorsally (
<figureCitation id="C8AA4D86FF80B077FE6FC7BC91577AA9" box="[421,527,223,249]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="36" pageNumber="237">Fig. 21A</figureCitation>
); pubescence in dorsolateral angles of pronotum not forming continuous, linear band with pubescence on anterior margin of scutum, and without tuft of thick, short hairs; disc of scutum with simple, thin, short, scattered, white hairs on its entire surface; mesepisternum with plumose, thin, short, white hairs, increasing in length towards dorsal surface; disc of metepisternum without characteristic tuft of plumose, thick, short, white hairs; tergal fasciae with rows of thin, long, white hairs (
<figureCitation id="C8AA4D86FF80B077FC7BC60C97517BD9" box="[945,1033,367,393]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="36" pageNumber="237">Fig. 3L</figureCitation>
); T1 to T3 with simple, short, thin, white hairs in punctures; disc of T4 to T6 with simple, long, thick, whitish hairs in punctures increasing in length towards posterior terga (
<figureCitation id="C8AA4D86FF80B077FE62C6D491497B81" box="[424,529,439,465]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="36" pageNumber="237">Fig. 21D</figureCitation>
); posterior surface of S1 with few plumose, scattered, long whitish hairs.
</paragraph>
<paragraph id="502E5103FF80B077FF0DC6B89294789D" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
<emphasis id="62E58D11FF80B077FF0DC6B892117BA5" box="[199,329,475,501]" italics="true" pageId="36" pageNumber="237">
<taxonomicName id="97912A80FF80B077FF0DC6B8921D7BA5" box="[199,325,475,501]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="36" pageNumber="237" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
Posterior margin of S5 sub-quadrate, strongly bilobated, with two quadrate lobes on top, three lines of capitate hairs on top of lobes, and two tufts of a few simple, thick, short hairs on top of two small, inner, smooth elevations (at base of lobes), with transverse, subapical line (
<figureCitation id="C8AA4D86FF80B077FC9CC54090E6786D" box="[854,958,547,573]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="36" pageNumber="237">Fig. 21E</figureCitation>
); S6 with pointy, acute-angled apex, with very short and thin lamella, with one subapical tuft of a few capitate thick, short hairs erected (
<figureCitation id="C8AA4D86FF80B077FB04C524966B7831" box="[1230,1331,583,609]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="36" pageNumber="237">Fig. 21F</figureCitation>
); capsule in dorsal view
<quantity id="9769FCE6FF80B077FE8AC50892FD78D5" box="[320,421,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="36" pageNumber="237" unit="mm" value="1.03">1.03 mm</quantity>
length, and
<quantity id="9769FCE6FF80B077FDF8C50991CF78D5" box="[562,663,618,645]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="36" pageNumber="237" unit="mm" value="0.52">0.52 mm</quantity>
width, 1.98 times longer than wide, gonobase
<quantity id="9769FCE6FF80B077FB60C508965778D5" box="[1194,1295,619,645]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="36" pageNumber="237" unit="mm" value="0.08">0.08 mm</quantity>
length, 5.75 times wider than long, and gonocoxite
<quantity id="9769FCE6FF80B077FD9CC5ED91E478F9" box="[598,700,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="36" pageNumber="237" unit="mm" value="0.45">0.45 mm</quantity>
length, gonostylus
<quantity id="9769FCE6FF80B077FC56C5ED975A78F9" box="[924,1026,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="36" pageNumber="237" unit="mm" value="0.95">0.95 mm</quantity>
length with acute apex, penis valve
<quantity id="9769FCE6FF80B077FF5DC5D093A3789D" box="[151,251,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="36" pageNumber="237" unit="mm" value="0.71">0.71 mm</quantity>
length (
<figureCitation id="C8AA4D86FF80B077FE9DC5D09298789D" box="[343,448,691,717]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="36" pageNumber="237">Fig. 21G</figureCitation>
).
</paragraph>
<paragraph id="502E5103FF80B077FF0DC5B592557E65" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
<emphasis id="62E58D11FF80B077FF0DC5B5927F78A1" bold="true" box="[199,295,726,753]" pageId="36" pageNumber="237">
<emphasis id="62E58D11FF80B077FF0DC5B5924678A1" bold="true" box="[199,286,726,753]" italics="true" pageId="36" pageNumber="237">Female</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FF80B077FE75C5B4914D78A1" box="[447,533,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="36" pageNumber="237" unit="mm" value="8.3">8.3 mm</quantity>
(
<quantity id="9769FCE6FF80B077FDEEC5B591F678A1" box="[548,686,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="8.05" metricValueMax="8.6" metricValueMin="7.5" pageId="36" pageNumber="237" unit="mm" value="8.05" valueMax="8.6" valueMin="7.5">
7.5
<emphasis id="62E58D11FF80B077FD82C5B4910E78A1" bold="true" box="[584,598,727,753]" pageId="36" pageNumber="237"></emphasis>
8.6 mm
</quantity>
, n=5); forewing length
<quantity id="9769FCE6FF80B077FC7CC5B4975478A1" box="[950,1036,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="36" pageNumber="237" unit="mm" value="4.6">4.6 mm</quantity>
(
<quantity id="9769FCE6FF80B077FBD0C5B497FD78A1" box="[1050,1189,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="36" pageNumber="237" unit="mm" value="4.85" valueMax="5.1" valueMin="4.6">
4.6
<emphasis id="62E58D11FF80B077FBF5C5B4971578A1" bold="true" box="[1087,1101,727,753]" pageId="36" pageNumber="237"></emphasis>
5.1 mm
</quantity>
, n=5); similar to male except:
<emphasis id="62E58D11FF80B077FF24C59892737945" box="[238,299,763,789]" italics="true" pageId="36" pageNumber="237">Head</emphasis>
. Length
<quantity id="9769FCE6FF80B077FE44C59892BB7945" box="[398,483,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="36" pageNumber="237" unit="mm" value="1.9">1.9 mm</quantity>
(
<quantity id="9769FCE6FF80B077FE38C59891237945" box="[498,635,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="36" pageNumber="237" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">
1.9
<emphasis id="62E58D11FF80B077FDDCC598917C7945" bold="true" box="[534,548,763,789]" pageId="36" pageNumber="237"></emphasis>
2.2 mm
</quantity>
, n=5), width
<quantity id="9769FCE6FF80B077FCC5C598903C7945" box="[783,868,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="36" pageNumber="237" unit="mm" value="1.9">1.9 mm</quantity>
(
<quantity id="9769FCE6FF80B077FCB8C59890BF7945" box="[882,999,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="36" pageNumber="237" unit="mm" value="1.95" valueMax="2.0" valueMin="1.9">
1.9
<emphasis id="62E58D11FF80B077FC5CC59890FC7945" bold="true" box="[918,932,763,789]" pageId="36" pageNumber="237"></emphasis>
2 mm
</quantity>
, n=5); eyes about 4.1 times longer than broad, 1.2 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF80B077FC77C47C975A7969" box="[957,1026,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="237" unit="mm" value="1.0">1 mm</quantity>
; upper interocular distance
<quantity id="9769FCE6FF80B077FAF7C47C96C07969" box="[1341,1432,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="36" pageNumber="237" unit="mm" value="1.1">1.1 mm</quantity>
; interantennal distance 1.6 times antennocular distance; interocellar distance 1.8 times OD, ocellocular distance 1.8 times OD, OPR 1.9 times OD; labrum with one rounded, basal, median tubercle; dorsal margin of mandible with carina; clypeus without emargination and apical margin with five rounded, small tubercles (
<figureCitation id="C8AA4D86FF80B077FB2BC4E8961179F5" box="[1249,1353,907,933]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="36" pageNumber="237">Fig. 21C</figureCitation>
); scape 3.2 times longer than broad; pedicel 1.3 longer than broad, about 2 times longer than F1 and 1.6 longer than F2; remaining flagellomeres 1.8 longer than F1.
<emphasis id="62E58D11FF80B077FD0AC4B0906079BC" box="[704,824,979,1004]" italics="true" pageId="36" pageNumber="237">Mesosoma</emphasis>
. Scutum 2.9 times longer than scutellum; intertegular distance
<quantity id="9769FCE6FF80B077FF31C495920A7E41" box="[251,338,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="237" unit="mm" value="1.5">1.5 mm</quantity>
(
<quantity id="9769FCE6FF80B077FEA8C49492B57E41" box="[354,493,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="36" pageNumber="237" unit="mm" value="1.4" valueMax="1.6" valueMin="1.2">
1.2
<emphasis id="62E58D11FF80B077FE4CC49492CC7E41" bold="true" box="[390,404,1015,1041]" pageId="36" pageNumber="237"></emphasis>
1.6 mm
</quantity>
, n=5).
<emphasis id="62E58D11FF80B077FDF7C49491EA7E40" box="[573,690,1015,1040]" italics="true" pageId="36" pageNumber="237">Metasoma</emphasis>
. T1
<quantity id="9769FCE6FF80B077FD22C49490677E41" box="[744,831,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="36" pageNumber="237" unit="mm" value="1.7">1.7 mm</quantity>
broad (
<quantity id="9769FCE6FF80B077FC5CC49497797E41" box="[918,1057,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="36" pageNumber="237" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">
1.7
<emphasis id="62E58D11FF80B077FC70C49490907E41" bold="true" box="[954,968,1015,1041]" pageId="36" pageNumber="237"></emphasis>
1.8 mm
</quantity>
, n=5). T3
<quantity id="9769FCE6FF80B077FB53C49497A87E41" box="[1177,1264,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="36" pageNumber="237" unit="mm" value="1.9">1.9 mm</quantity>
broad (
<quantity id="9769FCE6FF80B077FA8DC494939D7E65" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="36" pageNumber="237" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">
1.9
<emphasis id="62E58D11FF80B077FAA1C49496217E41" bold="true" box="[1387,1401,1015,1041]" pageId="36" pageNumber="237"></emphasis>
2.1 mm
</quantity>
, n=5).
</paragraph>
<paragraph id="502E5103FF80B077FF0DC35C90637FB5" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
<emphasis id="62E58D11FF80B077FF0DC35C92097E09" box="[199,337,1087,1113]" italics="true" pageId="36" pageNumber="237">Sculpturing.</emphasis>
Clypeus with small (0.3 times OD), contiguous, coarse punctures becoming smaller on lower two thirds, irregularly shaped, without micropunctures between punctures; supraclypeal area with punctures slightly smaller than base of clypeus; frons with bigger, coarser, and contiguous punctures than on supraclypeal area; on frons in front of ocelli slightly biggerpunctures on sides than on frons; vertex with bigger and sparser than on clypeus, with irregular surface between punctures; juxtantennal carina absent but strongly angulated; gena evenly punctate as on frons in front of ocelli; scutum with smaller and denser punctures than on vertex, with micropunctures between punctures; rest of mesosoma as on scutum except: scutellum with bigger and slightly sparser punctures; mesepisternum with bigger and slightly sparser punctures, with micropuntures between punctures; terga bright, microreticulate between punctures; T1 and T2 with smaller (0.1 times OD) punctures than on scutum, sparser (0.3 times DOP), with rounded shape; T3 with punctures 0.7 times DOP of scutum, rounded, and sparser (2.2 times DOP); dorsal surface of T6 with bigger, coarser and denser punctures than on T5; sterna with punctation smaller and sparser than on scutum, microreticulate on posterior margin.
</paragraph>
<paragraph id="502E5103FF80B077FF0DC28C91CF7C25" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
<emphasis id="62E58D11FF80B077FF0DC28C92687C58" box="[199,304,1519,1544]" italics="true" pageId="36" pageNumber="237">Vestiture.</emphasis>
Clypeus with few simple, short, white hairs in punctures; frons with few plumose, thin, white hairs (
<figureCitation id="C8AA4D86FF80B077FF55C17092507C7D" box="[159,264,1555,1581]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="36" pageNumber="237">Fig. 21C</figureCitation>
); gena with plumose, white hairs increasing in length towards hypostomal area; T4 to T6 with simple, thin, golden hairs in punctures, increasing in length towards posterior terga; subapical margin of T6 with linear tuft of plumose, short, yellowish hairs (
<figureCitation id="C8AA4D86FF80B077FDE8C13891D37C25" box="[546,651,1627,1653]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="36" pageNumber="237">Fig. 21D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF80B077FF0DC11C92617D8D" pageId="36" pageNumber="237" type="materials_examined">
<paragraph id="502E5103FF80B077FF0DC11C977D7D1D" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
<emphasis id="62E58D11FF80B077FF0DC11C92217CC8" bold="true" box="[199,377,1662,1689]" pageId="36" pageNumber="237">Type material:</emphasis>
<materialsCitation id="E0F95B5EFF80B077FE4BC11C92AB7CED" collectingDate="2009-01-24" collectionCode="ECOAB" collectorName="P. Sagot" country="Mexico" elevation="5" location="Arriaga" pageId="36" pageNumber="237" specimenCode="ECOAB 46959" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="62E58D11FF80B077FE4BC11C97587CC8" bold="true" box="[385,1024,1662,1689]" pageId="36" pageNumber="237">
<typeStatus id="8F2AEFA1FF80B077FE4BC11C92B67CC9" box="[385,494,1663,1689]" pageId="36" pageNumber="237" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="0037F978FF80B077FD8AC11D91A17CC8" box="[576,761,1662,1688]" collectionCode="ECOAB" pageId="36" pageNumber="237">ECOAB 46959</specimenCode>
):
<collectingCountry id="28861193FF80B077FCDAC11D90D37CC8" box="[784,907,1662,1688]" name="Mexico" pageId="36" pageNumber="237">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF80B077FC52C11D90A47CC8" box="[920,1020,1662,1688]" country="Mexico" name="Chiapas" pageId="36" pageNumber="237">Chiapas</collectingRegion>
,
</emphasis>
<location id="554E07D8FF80B077FBC3C11C97397CC9" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FBC3C11C97397CC9" box="[1033,1121,1663,1689]" country="Mexico" name="Arriaga" pageId="36" pageNumber="237" stateProvince="Chiapas">Arriaga</location>
, 465,
<quantity id="9769FCE6FF80B077FB6AC11D97967CC9" box="[1184,1230,1662,1689]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">
<elevation id="DBBCB630FF80B077FB6AC11D97967CC9" box="[1184,1230,1662,1689]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">5 m</elevation>
</quantity>
, 16.34207 -93.95,
<date id="242F77C3FF80B077FF5DC1C0925F7CED" box="[151,263,1699,1725]" pageId="36" pageNumber="237" value="2009-01-24">
<collectingDate id="346B8E2BFF80B077FF5DC1C0925F7CED" box="[151,263,1699,1725]" pageId="36" pageNumber="237" value="2009-01-24">24/I/2009</collectingDate>
</date>
,
<collectorName id="FD6434D5FF80B077FEDCC1C0922C7CED" box="[278,372,1699,1725]" pageId="36" pageNumber="237">P. Sagot</collectorName>
(
<collectionCode id="3680C9C6FF80B077FE4CC1C092B37CED" box="[390,491,1699,1725]" pageId="36" pageNumber="237">ECOAB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF80B077FE35C1C191367CB1" collectingDate="2009-11-18" collectionCode="ECOAB" country="Mexico" elevation="5" location="Arriaga" pageId="36" pageNumber="237" specimenCode="ECOAB 46961" specimenCount="1" stateProvince="Chiapas" typeStatus="allotype">
<emphasis id="62E58D11FF80B077FE35C1C197657CEC" bold="true" box="[511,1085,1698,1724]" pageId="36" pageNumber="237">
<typeStatus id="8F2AEFA1FF80B077FE35C1C1913C7CEC" box="[511,612,1698,1724]" pageId="36" pageNumber="237" type="allotype">Allotype</typeStatus>
(
<specimenCode id="0037F978FF80B077FDBDC1C1906A7CEC" box="[631,818,1698,1724]" collectionCode="ECOAB" pageId="36" pageNumber="237">ECOAB 46961</specimenCode>
):
<collectingCountry id="28861193FF80B077FC80C1C1909D7CEC" box="[842,965,1698,1724]" name="Mexico" pageId="36" pageNumber="237">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF80B077FC1EC1C197607CEC" box="[980,1080,1698,1724]" country="Mexico" name="Chiapas" pageId="36" pageNumber="237">Chiapas</collectingRegion>
,
</emphasis>
<location id="554E07D8FF80B077FB8DC1C097C77CED" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FB8DC1C097C77CED" box="[1095,1183,1699,1725]" country="Mexico" name="Arriaga" pageId="36" pageNumber="237" stateProvince="Chiapas">Arriaga</location>
, 465,
<quantity id="9769FCE6FF80B077FB2BC1C196487CED" box="[1249,1296,1698,1725]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">
<elevation id="DBBCB630FF80B077FB2BC1C196487CED" box="[1249,1296,1698,1725]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">5 m</elevation>
</quantity>
, 16.34207 - 93.95,
<date id="242F77C3FF80B077FF22C1A492337CB1" box="[232,363,1735,1761]" pageId="36" pageNumber="237" value="2009-11-18">
<collectingDate id="346B8E2BFF80B077FF22C1A492337CB1" box="[232,363,1735,1761]" pageId="36" pageNumber="237" value="2009-11-18">18/XI/2009</collectingDate>
</date>
, C.
<taxonomicName id="97912A80FF80B077FE55C1A491367CB1" authority="(ECOAB)" baseAuthorityName="ECOAB" box="[415,622,1735,1761]" class="Magnoliopsida" family="Fabaceae" genus="Balboa" kingdom="Plantae" order="Fabales" pageId="36" pageNumber="237" phylum="Tracheophyta" rank="genus">Balboa (ECOAB)</taxonomicName>
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF80B077FDB6C1A497E47D55" collectingDate="2009-11-18" collectionCode="ECOAB" country="Mexico" elevation="5" location="Arriaga" pageId="36" pageNumber="237" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="62E58D11FF80B077FDB6C1A490A07CB0" bold="true" box="[636,1016,1734,1761]" pageId="36" pageNumber="237">
<typeStatus id="8F2AEFA1FF80B077FDB6C1A491AE7CB1" box="[636,758,1735,1761]" pageId="36" pageNumber="237" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="28861193FF80B077FCCFC1A590D87CB0" box="[773,896,1734,1760]" name="Mexico" pageId="36" pageNumber="237">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF80B077FC45C1A590AB7CB0" box="[911,1011,1734,1760]" country="Mexico" name="Chiapas" pageId="36" pageNumber="237">Chiapas</collectingRegion>
,
</emphasis>
<location id="554E07D8FF80B077FBCBC1A497017CB1" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FBCBC1A497017CB1" box="[1025,1113,1735,1761]" country="Mexico" name="Arriaga" pageId="36" pageNumber="237" stateProvince="Chiapas">Arriaga</location>
, 465,
<quantity id="9769FCE6FF80B077FB50C1A597927CB1" box="[1178,1226,1734,1761]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">
<elevation id="DBBCB630FF80B077FB50C1A597927CB1" box="[1178,1226,1734,1761]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">5 m</elevation>
</quantity>
, 16.34207 -93.95, 24, 24/I, 14/III, 17,
<date id="242F77C3FF80B077FEBCC18892A27D55" box="[374,506,1771,1797]" pageId="36" pageNumber="237" value="2009-11-18">
<collectingDate id="346B8E2BFF80B077FEBCC18892A27D55" box="[374,506,1771,1797]" pageId="36" pageNumber="237" value="2009-11-18">18/XI/2009</collectingDate>
</date>
, C.
<taxonomicName id="97912A80FF80B077FDEDC18890B47D55" authority=", J. Merida, M. Guzman, P. Sagot" authorityName="J. Merida, M. Guzman, P. Sagot" box="[551,1004,1771,1797]" class="Magnoliopsida" family="Fabaceae" genus="Balboa" kingdom="Plantae" order="Fabales" pageId="36" pageNumber="237" phylum="Tracheophyta" rank="genus">Balboa, J. Mérida, M. Guzmán, P. Sagot</taxonomicName>
(
<collectionCode id="3680C9C6FF80B077FC31C18897047D55" box="[1019,1116,1771,1797]" pageId="36" pageNumber="237">ECOAB</collectionCode>
<specimenCount id="46979A8AFF80B077FBA9C18897DE7D55" box="[1123,1158,1771,1797]" count="6" pageId="36" pageNumber="237" type="female">6♀</specimenCount>
<specimenCount id="46979A8AFF80B077FB44C18997EA7D55" box="[1166,1202,1770,1797]" count="5" pageId="36" pageNumber="237" type="male">5♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF80B077FB03C189912B7D79" collectingDate="2009-01-24" collectionCode="ECOAB" collectorName="Sagot" country="Mexico" elevation="5" pageId="36" pageNumber="237" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
465,
<quantity id="9769FCE6FF80B077FB31C189967F7D55" box="[1275,1319,1770,1797]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">
<elevation id="DBBCB630FF80B077FB31C189967F7D55" box="[1275,1319,1770,1797]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">5 m</elevation>
</quantity>
, 16.34207 -93.95,
<date id="242F77C3FF80B077FF3AC06C92077D79" box="[240,351,1807,1833]" pageId="36" pageNumber="237" value="2009-01-24">
<collectingDate id="346B8E2BFF80B077FF3AC06C92077D79" box="[240,351,1807,1833]" pageId="36" pageNumber="237" value="2009-01-24">24/I/2009</collectingDate>
</date>
, P.
<collectorName id="FD6434D5FF80B077FE40C06C92927D79" box="[394,458,1807,1833]" pageId="36" pageNumber="237">Sagot</collectorName>
(
<collectionCode id="3680C9C6FF80B077FE11C06C91647D79" box="[475,572,1807,1833]" pageId="36" pageNumber="237">ECOAB</collectionCode>
<specimenCount id="46979A8AFF80B077FD8FC06C91317D79" box="[581,617,1807,1833]" count="1" pageId="36" pageNumber="237" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF80B077FD48C06C96CF7D79" box="[642,1431,1807,1833]" collectingDate="2016-03-24" collectionCode="ECOAB" collectorName="P. Sagot" country="Mexico" elevation="412" location="Bochil" pageId="36" pageNumber="237" specimenCount="4" specimenCount-female="4" stateProvince="Chiapas" typeStatus="paratype">
<location id="554E07D8FF80B077FD48C06C91977D79" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FD48C06C91977D79" box="[642,719,1807,1833]" country="Mexico" name="Bochil" pageId="36" pageNumber="237" stateProvince="Chiapas">Bochil</location>
,
<quantity id="9769FCE6FF80B077FD17C06C90707D79" box="[733,808,1807,1833]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="36" pageNumber="237" unit="m" value="412.0">
<elevation id="DBBCB630FF80B077FD17C06C90707D79" box="[733,808,1807,1833]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="36" pageNumber="237" unit="m" value="412.0">412 m</elevation>
</quantity>
, 16.97647 -93.06,
<date id="242F77C3FF80B077FBCBC06C97DC7D79" box="[1025,1156,1807,1833]" pageId="36" pageNumber="237" value="2016-03-24">
<collectingDate id="346B8E2BFF80B077FBCBC06C97DC7D79" box="[1025,1156,1807,1833]" pageId="36" pageNumber="237" value="2016-03-24">24/III/2016</collectingDate>
</date>
,
<collectorName id="FD6434D5FF80B077FB5BC06C97B67D79" box="[1169,1262,1807,1833]" pageId="36" pageNumber="237">P. Sagot</collectorName>
(
<collectionCode id="3680C9C6FF80B077FB35C06C96387D79" box="[1279,1376,1807,1833]" pageId="36" pageNumber="237">ECOAB</collectionCode>
<specimenCount id="46979A8AFF80B077FAA3C06C96D67D79" box="[1385,1422,1807,1833]" count="4" pageId="36" pageNumber="237" type="female">4♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF80B077FF5DC05097797D1D" box="[151,1057,1842,1869]" collectingDate="2021-01-25" collectionCode="ECOAB" collectorName="Sagot" country="Mexico" elevation="553" location="Chiapa de Corzo" pageId="36" pageNumber="237" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Chiapas" typeStatus="paratype">
<location id="554E07D8FF80B077FF5DC050920E7D1D" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FF5DC050920E7D1D" box="[151,342,1843,1869]" country="Mexico" name="Chiapa de Corzo" pageId="36" pageNumber="237" stateProvince="Chiapas">Chiapa de Corzo</location>
,
<quantity id="9769FCE6FF80B077FEABC05192F27D1D" box="[353,426,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" pageId="36" pageNumber="237" unit="m" value="553.0">
<elevation id="DBBCB630FF80B077FEABC05192F27D1D" box="[353,426,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" pageId="36" pageNumber="237" unit="m" value="553.0">553 m</elevation>
</quantity>
, 16.46994 -92.99,
<date id="242F77C3FF80B077FDB6C05191B47D1D" box="[636,748,1842,1869]" pageId="36" pageNumber="237" value="2021-01-25">
<collectingDate id="346B8E2BFF80B077FDB6C05191B47D1D" box="[636,748,1842,1869]" pageId="36" pageNumber="237" value="2021-01-25">25/I/2021</collectingDate>
</date>
, P.
<collectorName id="FD6434D5FF80B077FCD8C050900A7D1D" box="[786,850,1843,1869]" pageId="36" pageNumber="237">Sagot</collectorName>
(
<collectionCode id="3680C9C6FF80B077FCAAC05090997D1D" box="[864,961,1843,1869]" pageId="36" pageNumber="237">ECOAB</collectionCode>
<specimenCount id="46979A8AFF80B077FC02C05090B37D1D" box="[968,1003,1843,1869]" count="1" pageId="36" pageNumber="237" type="female">1♀</specimenCount>
<specimenCount id="46979A8AFF80B077FC38C050974F7D1D" box="[1010,1047,1843,1869]" count="3" pageId="36" pageNumber="237" type="male">3♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="502E5103FF80B077FF0DC03592617D8D" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
<emphasis id="62E58D11FF80B077FF0DC035916C7D20" bold="true" box="[199,564,1878,1904]" pageId="36" pageNumber="237">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFF80B077FDF3C03591867DC5" collectingDate="1987-11-02" collectionCode="BBSL" collectorName="T. Griswold" country="Mexico" elevation="250" latitude="17.989445" location="Los Amates Playa Azul" longLatPrecision="21" longitude="-102.365" pageId="36" pageNumber="237" specimenCount="1" specimenCount-female="1" stateProvince="Michoacan de Ocampo">
<emphasis id="62E58D11FF80B077FDF3C03590117D21" bold="true" box="[569,841,1878,1905]" pageId="36" pageNumber="237">
<collectingCountry id="28861193FF80B077FDF3C03591EC7D20" box="[569,692,1878,1904]" name="Mexico" pageId="36" pageNumber="237">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF80B077FD74C035901C7D21" box="[702,836,1878,1905]" country="Mexico" name="Michoacan de Ocampo" pageId="36" pageNumber="237">Michoacán</collectingRegion>
,
</emphasis>
<location id="554E07D8FF80B077FC85C03497097D21" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FC85C03497097D21" box="[847,1105,1879,1905]" country="Mexico" latitude="17.989445" longLatPrecision="21" longitude="-102.365" name="Los Amates Playa Azul" pageId="36" pageNumber="237" stateProvince="Michoacan de Ocampo">Los Amates Playa Azul</location>
26kmN, [
<quantity id="9769FCE6FF80B077FB74C03596517D21" box="[1214,1289,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="237" unit="m" value="250.0">
<elevation id="DBBCB630FF80B077FB74C03596517D21" box="[1214,1289,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="237" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF80B077FAD9C03596C47D21" box="[1299,1436,1878,1905]" degrees="17" direction="north" minutes="59" orientation="latitude" pageId="36" pageNumber="237" precision="15" seconds="22" value="17.989445">17°5922”N</geoCoordinate>
<geoCoordinate id="35A537C4FF80B077FF5DC01992617DC5" box="[151,313,1914,1941]" degrees="102" direction="west" minutes="21" orientation="longitude" pageId="36" pageNumber="237" precision="15" seconds="54" value="-102.365">102°2154”W</geoCoordinate>
],
<date id="242F77C3FF80B077FE81C01892997DC5" box="[331,449,1915,1941]" pageId="36" pageNumber="237" value="1987-11-02">
<collectingDate id="346B8E2BFF80B077FE81C01892997DC5" box="[331,449,1915,1941]" pageId="36" pageNumber="237" value="1987-11-02">2/XI/1987</collectingDate>
</date>
,
<collectorName id="FD6434D5FF80B077FE01C018910A7DC5" box="[459,594,1915,1941]" pageId="36" pageNumber="237">T. Griswold</collectorName>
, (
<collectionCode id="3680C9C6FF80B077FDAEC01891F37DC5" box="[612,683,1915,1941]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="36" pageNumber="237" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF80B077FD7BC018918A7DC5" box="[689,722,1915,1941]" count="1" pageId="36" pageNumber="237" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF80B077FD20C019911B7DE9" collectingDate="1993-04-07" collectionCode="BBSL" collectorName="La Ventosa &amp; A. Rodriguez" country="Mexico" elevation="29" latitude="16.551666" location="La Ventosa" longLatPrecision="21" longitude="-94.94583" pageId="36" pageNumber="237" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
<emphasis id="62E58D11FF80B077FD20C01990137DC4" bold="true" box="[746,843,1914,1940]" pageId="36" pageNumber="237">
<collectingRegion id="92559FE1FF80B077FD20C019901E7DC4" box="[746,838,1914,1940]" country="Mexico" name="Oaxaca" pageId="36" pageNumber="237">Oaxaca</collectingRegion>
,
</emphasis>
<collectorName id="FD6434D5FF80B077FC9BC01890957DC5" box="[849,973,1915,1941]" pageId="36" pageNumber="237">
<location id="554E07D8FF80B077FC9BC01890957DC5" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FC9BC01890957DC5" box="[849,973,1915,1941]" country="Mexico" latitude="16.551666" longLatPrecision="21" longitude="-94.94583" name="La Ventosa" pageId="36" pageNumber="237" stateProvince="Oaxaca">La Ventosa</location>
</collectorName>
10kmN,
<quantity id="9769FCE6FF80B077FBFEC01897367DC5" box="[1076,1134,1915,1941]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="36" pageNumber="237" unit="m" value="29.0">
<elevation id="DBBCB630FF80B077FBFEC01897367DC5" box="[1076,1134,1915,1941]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="36" pageNumber="237" unit="m" value="29.0">29 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF80B077FBB3C018965A7DC5" box="[1145,1282,1915,1941]" degrees="16" direction="north" minutes="33" orientation="latitude" pageId="36" pageNumber="237" precision="15" seconds="06" value="16.551666">16°3306”N</geoCoordinate>
<geoCoordinate id="35A537C4FF80B077FAC2C01996C07DC5" box="[1288,1432,1914,1941]" degrees="94" direction="west" minutes="56" orientation="longitude" pageId="36" pageNumber="237" precision="15" seconds="45" value="-94.94583">94°5645”W</geoCoordinate>
,
<date id="242F77C3FF80B077FF5DC0FC92557DE9" box="[151,269,1951,1977]" pageId="36" pageNumber="237" value="1993-04-07">
<collectingDate id="346B8E2BFF80B077FF5DC0FC92557DE9" box="[151,269,1951,1977]" pageId="36" pageNumber="237" value="1993-04-07">7/IV/1993</collectingDate>
</date>
,
<collectorName id="FD6434D5FF80B077FED0C0FC92EC7DE9" box="[282,436,1951,1977]" pageId="36" pageNumber="237">A. Rodriguez</collectorName>
(
<collectionCode id="3680C9C6FF80B077FE0FC0FC91547DE9" box="[453,524,1951,1977]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="36" pageNumber="237" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF80B077FDDFC0FC916E7DE9" box="[533,566,1951,1977]" count="1" pageId="36" pageNumber="237" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF80B077FD98C0FC926C7D8D" collectingDate="2018-11-19" collectionCode="ECOAB" collectorName="N. Arnold" country="Mexico" elevation="1154" location="Santa Maria Zoquitlan" pageId="36" pageNumber="237" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
<location id="554E07D8FF80B077FD98C0FC900F7DE9" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FD98C0FC900F7DE9" box="[594,855,1951,1977]" country="Mexico" name="Santa Maria Zoquitlan" pageId="36" pageNumber="237" stateProvince="Oaxaca">Santa Maria Zoquitlan</location>
,
<quantity id="9769FCE6FF80B077FCAEC0FD90E57DE9" box="[868,957,1950,1977]" metricMagnitude="3" metricUnit="m" metricValue="1.154" pageId="36" pageNumber="237" unit="m" value="1154.0">
<elevation id="DBBCB630FF80B077FCAEC0FD90E57DE9" box="[868,957,1950,1977]" metricMagnitude="3" metricUnit="m" metricValue="1.154" pageId="36" pageNumber="237" unit="m" value="1154.0">1154 m</elevation>
</quantity>
, 16.51338 -96.34,
<date id="242F77C3FF80B077FB52C0FC96447DE9" box="[1176,1308,1951,1977]" pageId="36" pageNumber="237" value="2018-11-19">
<collectingDate id="346B8E2BFF80B077FB52C0FC96447DE9" box="[1176,1308,1951,1977]" pageId="36" pageNumber="237" value="2018-11-19">19/XI/2018</collectingDate>
</date>
,
<collectorName id="FD6434D5FF80B077FAE3C0FC96C47DE9" box="[1321,1436,1951,1977]" pageId="36" pageNumber="237">N. Arnold</collectorName>
(
<collectionCode id="3680C9C6FF80B077FF55C0A092587D8D" box="[159,256,1987,2013]" pageId="36" pageNumber="237">ECOAB</collectionCode>
<specimenCount id="46979A8AFF80B077FECCC0A092707D8D" box="[262,296,1987,2013]" count="1" pageId="36" pageNumber="237" type="female">1♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF81B076FF0DC7F5966A7AA8" pageId="37" pageNumber="238" type="distribution">
<paragraph id="502E5103FF81B076FF0DC7F5966A7AA8" blockId="37.[151,1437,150,460]" pageId="37" pageNumber="238">
<emphasis id="62E58D11FF81B076FF0DC7F592397AE0" bold="true" box="[199,353,150,176]" pageId="37" pageNumber="238">Distribution.</emphasis>
This species is present along the Pacific slope of southern
<collectingCountry id="28861193FF81B076FBD9C7F497337AE1" box="[1043,1131,151,177]" name="Mexico" pageId="37" pageNumber="238">Mexico</collectingCountry>
, in areas with tropical dry forest, and is present in the states of
<collectingRegion id="92559FE1FF81B076FD8FC7D8919C7A85" box="[581,708,187,213]" country="Mexico" name="Michoacan de Ocampo" pageId="37" pageNumber="238">Michoacán</collectingRegion>
,
<collectingRegion id="92559FE1FF81B076FD18C7D8907E7A85" box="[722,806,187,213]" country="Mexico" name="Oaxaca" pageId="37" pageNumber="238">Oaxaca</collectingRegion>
and
<collectingRegion id="92559FE1FF81B076FCA8C7D890E77A85" box="[866,959,187,213]" country="Mexico" name="Chiapas" pageId="37" pageNumber="238">Chiapas</collectingRegion>
, being endemic to
<collectingCountry id="28861193FF81B076FB57C7D897AD7A85" box="[1181,1269,187,213]" name="Mexico" pageId="37" pageNumber="238">Mexico</collectingCountry>
, in the Pacific Lowlands biogeographic province (
<bibRefCitation id="34002CF2FF81B076FDEEC7BD90507AA8" author="Morrone, J. J. &amp; Escalante, T. &amp; Rodriguez-Tapia, G. &amp; Carmona, A. &amp; Arana, M. &amp; Mercado-Gomez, J. D." box="[548,776,222,248]" pageId="37" pageNumber="238" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. &amp; Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="62E58D11FF81B076FD45C7BC91987AA8" box="[655,704,222,248]" italics="true" pageId="37" pageNumber="238">et al</emphasis>
. 2022
</bibRefCitation>
) (
<figureCitation id="C8AA4D86FF81B076FCD5C7BD902B7AA8" box="[799,883,222,248]" captionStart="FIGURE 20" captionStartId="34.[151,250,1098,1122]" captionTargetBox="[159,1428,187,1064]" captionTargetId="figure-22@34.[151,1436,179,1072]" captionTargetPageId="34" captionText="FIGURE 20. Known distribution records for Heriades luisa sp. nov." pageId="37" pageNumber="238">Fig. 20</figureCitation>
). Altitudinal range: sea level to
<quantity id="9769FCE6FF81B076FB1DC7BD96767AA8" box="[1239,1326,222,249]" metricMagnitude="3" metricUnit="m" metricValue="1.154" pageId="37" pageNumber="238" unit="m" value="1154.0">1154 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF81B076FF0DC66296277B6E" pageId="37" pageNumber="238" type="etymology">
<paragraph id="502E5103FF81B076FF0DC66296277B6E" blockId="37.[151,1437,150,460]" pageId="37" pageNumber="238">
<emphasis id="62E58D11FF81B076FF0DC66292167B4B" bold="true" box="[199,334,257,283]" pageId="37" pageNumber="238">Etymology.</emphasis>
This species is dedicated to Dr. Luisa Ruz, a Chilean taxonomist and melitologist who has contributed greatly to the knowledge of native bees in South America, especially bees of the family
<taxonomicName id="97912A80FF81B076FB3FC64796237B6E" authorityName="Latreille" authorityYear="1802" box="[1269,1403,292,318]" class="Insecta" family="Andrenidae" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="family">Andrenidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF81B076FF0DC624918E7B9B" pageId="37" pageNumber="238" type="discussion">
<paragraph id="502E5103FF81B076FF0DC624918E7B9B" blockId="37.[151,1437,150,460]" pageId="37" pageNumber="238">
<emphasis id="62E58D11FF81B076FF0DC62492177B31" bold="true" box="[199,335,327,353]" pageId="37" pageNumber="238">Comments.</emphasis>
Females show variation in tubercles on the apical margin of clypeus in frontal view only, from well-defined rounded tubercles to crenulated margin. Although body size is large for the genus, some small females can be found in
<collectingRegion id="92559FE1FF81B076FE85C6ED92967BF8" box="[335,462,398,424]" country="Mexico" name="Michoacan de Ocampo" pageId="37" pageNumber="238">Michoacán</collectingRegion>
, Los Amates, and
<collectingRegion id="92559FE1FF81B076FD6AC6ED91A57BF8" box="[672,765,398,424]" country="Mexico" name="Chiapas" pageId="37" pageNumber="238">Chiapas</collectingRegion>
, Arriaga,
<collectingCountry id="28861193FF81B076FCA6C6ED909D7BF8" box="[876,965,398,424]" name="Mexico" pageId="37" pageNumber="238">Mexico</collectingCountry>
. Males seem to vary on S6 with posterior tuft in a V-line or a rounded shape of capitate hairs.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,448 @@
<document id="414984BB888607A81B3B230B0F015ABF" ID-DOI="10.11646/zootaxa.5570.2.1" ID-ISSN="1175-5326" ID-ZooBank="7177B69B-6132-485D-87C7-4EA807692757" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737737014959" checkinUser="plazi" docAuthor="Rojas-Arias, Laura, Griswold, Terry, Hinojosa-Díaz, Ismael, Morrone, Juan J. &amp; Barajas, Ricardo Ayala" docDate="2025" docId="D838E015FF8CB07AFF5DC10591A37DA3" docLanguage="en" docName="zootaxa.5570.2.1.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heriades micheneri Timberlake 1947" docType="treatment" docVersion="1" lastPageNumber="242" masterDocId="2401986DFFA4B053FFCAC76393587A50" masterDocTitle="Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species" masterLastPageNumber="259" masterPageNumber="201" pageNumber="241" updateTime="1737737247067" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="AFD45A94CFF3A514FC632B07B1A8D935">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<mods:namePart id="727741BBFE6A0D179585293FFB54B556">Rojas-Arias, Laura</mods:namePart>
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<mods:affiliation id="3D58A85A21B089709D63B5AE4865E918">Posgrado en Ciencias Biológicas, Universidad Nacional Autónoma de México (Posgrado en Ciencias Biológicas, UNAM. Unidad de</mods:affiliation>
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<mods:namePart id="434125D3EEEE63AB755D69D1C9E290E2">Griswold, Terry</mods:namePart>
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<mods:namePart id="3982458EA7366DEBC0533A77E5E24A30">Hinojosa-Díaz, Ismael</mods:namePart>
<mods:nameIdentifier id="CBEB2BBFA5E6D83F1E36D8DFF8902A32" type="ORCID">0000-0002-5493-850X</mods:nameIdentifier>
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<mods:namePart id="EBE132D699690CB9AC2E86B0F97B2584">Morrone, Juan J.</mods:namePart>
<mods:affiliation id="420CF2727294EE355AB2FE134A7E82C4">Museo de Zoología “ Alfonso L. Herrera ”, Departamento de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional Autónoma</mods:affiliation>
</mods:name>
<mods:name id="FB7221BAB269D05584771A1B2C51F1D1" type="personal">
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</mods:role>
<mods:namePart id="914DC644660B270DD564A579964A9F76">Barajas, Ricardo Ayala</mods:namePart>
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<mods:title id="A0317702E465672C8F4A85FB792180B1">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FD40E6DA529E38FF7092A3B64D7679A5">
<mods:date id="8FF72825103CDEADF1971AB7CBD48B81">2025</mods:date>
<mods:detail id="52646D94DF374E59E6647C33D4ECF4B8" type="pubDate">
<mods:number id="4205D0A160D8C770FEF8FB0F84A2A755">2025-01-17</mods:number>
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<mods:detail id="252D0E32596D9321BE4190DC1DBB6B97" type="volume">
<mods:number id="07687D0B9FCE5ABEF00D9D76669C2E8E">5570</mods:number>
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<mods:number id="AC3AB48CB0DF1C490EB919176FE2429C">2</mods:number>
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<mods:start id="73949FAF4D3AC984768BAA28E48548EB">201</mods:start>
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<mods:url id="F96BEFB5B7B1543727E31F38788DFB3E">https://doi.org/10.11646/zootaxa.5570.2.1</mods:url>
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<mods:identifier id="4F1F64470557008ECBE750057E829EAC" type="DOI">10.11646/zootaxa.5570.2.1</mods:identifier>
<mods:identifier id="16E2BE9841A088410C8267F24E1132A4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="D838E015FF8CB07AFF5DC10591A37DA3" LSID="urn:lsid:plazi:treatment:D838E015FF8CB07AFF5DC10591A37DA3" httpUri="http://treatment.plazi.org/id/D838E015FF8CB07AFF5DC10591A37DA3" lastPageId="41" lastPageNumber="242" pageId="40" pageNumber="241">
<subSubSection id="188B0288FF8CB07BFF5DC10591317CD0" box="[151,617,1638,1665]" pageId="40" pageNumber="241" type="nomenclature">
<paragraph id="502E5103FF8CB07BFF5DC10591317CD0" blockId="40.[151,617,1638,1701]" box="[151,617,1638,1665]" pageId="40" pageNumber="241">
<heading id="0B66E66FFF8CB07BFF5DC10591317CD0" bold="true" box="[151,617,1638,1665]" fontSize="11" level="1" pageId="40" pageNumber="241" reason="1">
<taxonomicName id="97912A80FF8CB07BFF5DC10591317CD0" ID-CoL="3KYS3" authority="Timberlake, 1947" authorityName="Timberlake" authorityYear="1947" box="[151,617,1638,1665]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="micheneri">
<emphasis id="62E58D11FF8CB07BFF5DC10591317CD0" bold="true" box="[151,617,1638,1665]" pageId="40" pageNumber="241">
<emphasis id="62E58D11FF8CB07BFF5DC10592DE7CD1" bold="true" box="[151,390,1638,1665]" italics="true" pageId="40" pageNumber="241">Heriades micheneri</emphasis>
Timberlake, 1947
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8CB07BFF5DC1E892927D13" pageId="40" pageNumber="241" type="description">
<paragraph id="502E5103FF8CB07BFF5DC1E892077CF5" blockId="40.[151,617,1638,1701]" box="[151,351,1674,1701]" pageId="40" pageNumber="241">
(
<figureCitation id="C8AA4D86FF8CB07BFF55C1E8925B7CF5" box="[159,259,1675,1701]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="40" pageNumber="241">Figs. 2A</figureCitation>
,
<figureCitation id="C8AA4D86FF8CB07BFEC5C1E9920F7CF5" box="[271,343,1674,1701]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStartId-0="41.[151,250,1098,1122]" captionStartId-1="42.[151,250,1943,1967]" captionTargetBox-0="[160,1428,187,1063]" captionTargetBox-1="[160,1414,187,1905]" captionTargetId-0="figure-18@41.[151,1436,179,1072]" captionTargetId-1="figure-22@42.[160,1427,179,1917]" captionTargetPageId-0="41" captionTargetPageId-1="42" captionText-0="FIGURE 24. Known distribution records for Heriades micheneri." captionText-1="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="40" pageNumber="241">2425</figureCitation>
)
</paragraph>
<paragraph id="502E5103FF8CB07BFF5DC1AC91DA7D56" blockId="40.[151,1436,1743,1859]" pageId="40" pageNumber="241">
<taxonomicName id="97912A80FF8CB07BFF5DC1AC91F17CB7" authority="Timberlake, 1947" authorityName="Timberlake" authorityYear="1947" box="[151,681,1743,1767]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="micheneri" subGenus="Neotrypetes">
<emphasis id="62E58D11FF8CB07BFF5DC1AC92B67CB7" box="[151,494,1743,1767]" italics="true" pageId="40" pageNumber="241">Heriades (Neotrypetes) micheneri</emphasis>
Timberlake, 1947
</taxonomicName>
. Two new species of bees from
<collectingRegion id="92559FE1FF8CB07BFC28C1AC976C7CB7" box="[994,1076,1743,1767]" country="United States of America" name="Arizona" pageId="40" pageNumber="241">Arizona</collectingRegion>
(
<taxonomicName id="97912A80FF8CB07BFB8AC1AC96697CB7" authority=", Apoidea" authorityName="Apoidea" box="[1088,1329,1743,1767]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Hymenoptera" pageId="40" pageNumber="241" phylum="Tracheophyta" rank="order">Hymenoptera, Apoidea</taxonomicName>
).
<emphasis id="62E58D11FF8CB07BFA8AC1AC92C47D56" italics="true" pageId="40" pageNumber="241">The Pan- Pacific Entomologist</emphasis>
. Vol. XXIII. No. 1:26.
</paragraph>
<paragraph id="502E5103FF8CB07BFF0DC06E90B57D74" blockId="40.[151,1436,1743,1859]" box="[199,1005,1804,1829]" pageId="40" pageNumber="241">
Type locality: Patagonia,
<collectingRegion id="92559FE1FF8CB07BFE02C06F91447D74" box="[456,540,1804,1828]" country="United States of America" name="Arizona" pageId="40" pageNumber="241">Arizona</collectingRegion>
, on damp sand,
<date id="242F77C3FF8CB07BFD75C06E90027D75" box="[703,858,1805,1829]" pageId="40" pageNumber="241" value="1940-08-08">August 8, 1940</date>
(Timberlake).
</paragraph>
<paragraph id="502E5103FF8CB07BFF0DC04892927D13" blockId="40.[151,1436,1743,1859]" box="[199,458,1835,1859]" pageId="40" pageNumber="241">
<typeStatus id="8F2AEFA1FF8CB07BFF0DC048927D7D13" box="[199,293,1835,1859]" pageId="40" pageNumber="241" type="holotype">Holotype</typeStatus>
female: UCRC.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8CB07AFF5DC01797F27D4A" lastPageId="41" lastPageNumber="242" pageId="40" pageNumber="241" type="diagnosis">
<paragraph id="502E5103FF8CB07AFF5DC01796067FE2" blockId="40.[151,1436,1908,2009]" lastBlockId="41.[151,1437,1171,2035]" lastPageId="41" lastPageNumber="242" pageId="40" pageNumber="241">
<emphasis id="62E58D11FF8CB07BFF5DC017924A7DDE" bold="true" box="[151,274,1908,1934]" pageId="40" pageNumber="241">Diagnosis.</emphasis>
Small-sized bees (males
<quantity id="9769FCE6FF8CB07BFDFDC017919A7DDF" box="[567,706,1908,1935]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="5.8" metricValueMin="5.5" pageId="40" pageNumber="241" unit="mm" value="5.65" valueMax="5.8" valueMin="5.5">5.55.8 mm</quantity>
, n=5; females
<quantity id="9769FCE6FF8CB07BFCA6C01790AF7DDF" box="[876,1015,1908,1935]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.1" metricValueMin="5.6" pageId="40" pageNumber="241" unit="mm" value="5.85" valueMax="6.1" valueMin="5.6">5.66.1 mm</quantity>
, n=3), ocelli medium-size (
<figureCitation id="C8AA4D86FF8CB07BFAFEC01796D67DDE" box="[1332,1422,1908,1934]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="40" pageNumber="241">Fig. 3B</figureCitation>
); punctation large along the whole body (
<figureCitation id="C8AA4D86FF8CB07BFD81C0FA91C77DE3" box="[587,671,1945,1971]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="40" pageNumber="241">Fig. 3F</figureCitation>
); dorsolateral angles of pronotum with tuft of plumose, short, whitish to yellowish hairs; tergal fasciae with row of thick, whitish hairs (
<figureCitation id="C8AA4D86FF8CB07BFC56C0DC97737D89" box="[924,1067,1982,2009]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="40" pageNumber="241">Fig. 25C, D</figureCitation>
); body with plumose hairs in well-defined tufts; frons, disc of scutum, mesepisternum and terga with bare areas.
<emphasis id="62E58D11FF8DB07AFBFDC3F097297EFD" box="[1079,1137,1171,1197]" italics="true" pageId="41" pageNumber="242">Male</emphasis>
: outer surface of mandible without tubercles; clypeus covered by plumose, long, white hairs directed medially (
<figureCitation id="C8AA4D86FF8DB07AFBABC3DB97947E83" box="[1121,1228,1208,1235]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="41" pageNumber="242">Fig. 25A</figureCitation>
); hypostomal area with plumose, long, white hairs, length surpassing mandible; S1 with strong ventral process, in profile when sides of process projected forming a right angle (
<figureCitation id="C8AA4D86FF8DB07AFD4EC260919F7F4D" box="[644,711,1283,1309]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="41" pageNumber="242">Fig. 2</figureCitation>
Aa), in posterior view with top semi circled (
<figureCitation id="C8AA4D86FF8DB07AFB0DC26096537F4D" box="[1223,1291,1283,1309]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="41" pageNumber="242">Fig. 2</figureCitation>
Ac); S5 with two quadrangular lobes with acute strong angles on sides, and a few lines of simple, short thick hairs, increasing length outwards, with one transverse line of capitate hairs increasing in length outwards (
<figureCitation id="C8AA4D86FF8DB07AFB5EC22D97A37F38" box="[1172,1275,1357,1384]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="41" pageNumber="242">Fig. 25E</figureCitation>
); S6 with two tufts of simple, short, thin hairs, directed downward, all hairs the same length (
<figureCitation id="C8AA4D86FF8DB07AFBDAC210972D7FDE" box="[1040,1141,1395,1422]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="41" pageNumber="242">Fig. 25F</figureCitation>
).
<emphasis id="62E58D11FF8DB07AFB43C21097857FDD" box="[1161,1245,1395,1421]" italics="true" pageId="41" pageNumber="242">Female</emphasis>
: similar to male, but without plumose hairs on clypeus (
<figureCitation id="C8AA4D86FF8DB07AFD80C2FB91E87FE3" box="[586,688,1432,1459]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="41" pageNumber="242">Fig. 25C</figureCitation>
), this species is similar to
<taxonomicName id="97912A80FF8DB07AFC1DC2FA97397FE2" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[983,1121,1432,1458]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="danuncia" status="sp. nov">
<emphasis id="62E58D11FF8DB07AFC1DC2FA97397FE2" box="[983,1121,1432,1458]" italics="true" pageId="41" pageNumber="242">H. danuncia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF8DB07AFBA2C2FA97E17FE3" box="[1128,1209,1433,1459]" pageId="41" pageNumber="242" rank="species">sp. nov</taxonomicNameLabel>
(
<figureCitation id="C8AA4D86FF8DB07AFB03C2FB96097FE2" box="[1225,1361,1432,1458]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="41" pageNumber="242">Fig. 23B, D</figureCitation>
).
</paragraph>
<caption id="04EE018BFF8DB07AFF5DC329906D7E32" box="[151,821,1098,1122]" pageId="41" pageNumber="242" startId="41.[151,250,1098,1122]" targetBox="[160,1428,187,1063]" targetPageId="41" targetType="figure">
<paragraph id="502E5103FF8DB07AFF5DC329906D7E32" blockId="41.[151,821,1098,1122]" box="[151,821,1098,1122]" pageId="41" pageNumber="242">
<emphasis id="62E58D11FF8DB07AFF5DC32992787E32" bold="true" box="[151,288,1098,1122]" pageId="41" pageNumber="242">FIGURE 24.</emphasis>
Known distribution records for
<taxonomicName id="97912A80FF8DB07AFDADC32990777E32" box="[615,815,1098,1122]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="micheneri">
<emphasis id="62E58D11FF8DB07AFDADC32990777E32" box="[615,815,1098,1122]" italics="true" pageId="41" pageNumber="242">Heriades micheneri</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="502E5103FF8DB07AFF0DC2DF97F27D4A" blockId="41.[151,1437,1171,2035]" pageId="41" pageNumber="242">
<emphasis id="62E58D11FF8DB07AFF0DC2DF91637F86" bold="true" box="[199,571,1468,1494]" pageId="41" pageNumber="242">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFF8DB07AFD8EC2DF90CF7FAA" collectingDate="1968-08-10" collectionCode="BBSL" collectorName="T. A. Sears &amp; R. C. Gardner &amp; C. S. Glaser" country="Mexico" elevation="1426" latitude="27.522223" location="Santo Nino" longLatPrecision="1" longitude="-104.83722" pageId="41" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="Chihuahua">
<emphasis id="62E58D11FF8DB07AFD8EC2DF90027F86" bold="true" box="[580,858,1468,1494]" pageId="41" pageNumber="242">
<collectingCountry id="28861193FF8DB07AFD8EC2DF91E77F86" box="[580,703,1468,1494]" name="Mexico" pageId="41" pageNumber="242">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF8DB07AFD07C2DF900D7F86" box="[717,853,1468,1494]" country="Mexico" name="Chihuahua" pageId="41" pageNumber="242">Chihuahua</collectingRegion>
,
</emphasis>
<location id="554E07D8FF8DB07AFCAEC2DF90B07F86" LSID="urn:lsid:plazi:treatment:D838E015FF8CB07AFF5DC10591A37DA3:554E07D8FF8DB07AFCAEC2DF90B07F86" box="[868,1000,1468,1494]" country="Mexico" latitude="27.522223" longLatPrecision="1" longitude="-104.83722" name="Santo Nino" pageId="41" pageNumber="242" stateProvince="Chihuahua">Santo Niño</location>
,
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<elevation id="DBBCB630FF8DB07AFC3FC2DF97167F87" box="[1013,1102,1468,1495]" metricMagnitude="3" metricUnit="m" metricValue="1.426" pageId="41" pageNumber="242" unit="m" value="1426.0">1426 m</elevation>
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,
<date id="242F77C3FF8DB07AFF5DC28392767FAA" box="[151,302,1504,1530]" pageId="41" pageNumber="242" value="1968-08-10">
<collectingDate id="346B8E2BFF8DB07AFF5DC28392767FAA" box="[151,302,1504,1530]" pageId="41" pageNumber="242" value="1968-08-10">10/VIII/1968</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8DB07AFEF1C28292E47FAA" box="[315,444,1504,1531]" pageId="41" pageNumber="242">T. A. Sears</collectorName>
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<collectorName id="FD6434D5FF8DB07AFE00C28291367FAA" box="[458,622,1504,1531]" pageId="41" pageNumber="242">R. C. Gardner</collectorName>
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<collectorName id="FD6434D5FF8DB07AFDB1C283905F7FAA" box="[635,775,1504,1530]" pageId="41" pageNumber="242">C. S. Glaser</collectorName>
(
<collectionCode id="3680C9C6FF8DB07AFCD2C28390077FAA" box="[792,863,1504,1530]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="41" pageNumber="242" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8DB07AFCA2C28390D57FAA" box="[872,909,1504,1530]" count="1" pageId="41" pageNumber="242" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8DB07AFC6FC28397797C4E" collectingDate="1968-07-30" collectionCode="BBSL" collectorName="T. A. Sears &amp; R. C. Gardner &amp; C. S. Glaser" country="Mexico" elevation="442" latitude="26.728611" location="Choix." longLatPrecision="4" longitude="-108.2975" pageId="41" pageNumber="242" specimenCount="1" specimenCount-female="1" stateProvince="Sinaloa">
<emphasis id="62E58D11FF8DB07AFC6FC283975C7FAA" bold="true" box="[933,1028,1504,1530]" pageId="41" pageNumber="242">
<collectingRegion id="92559FE1FF8DB07AFC6FC28390A77FAA" box="[933,1023,1504,1530]" country="Mexico" name="Sinaloa" pageId="41" pageNumber="242">Sinaloa</collectingRegion>
,
</emphasis>
<location id="554E07D8FF8DB07AFBC4C28397027FAA" LSID="urn:lsid:plazi:treatment:D838E015FF8CB07AFF5DC10591A37DA3:554E07D8FF8DB07AFBC4C28397027FAA" box="[1038,1114,1504,1530]" country="Mexico" latitude="26.728611" longLatPrecision="4" longitude="-108.2975" name="Choix." pageId="41" pageNumber="242" stateProvince="Sinaloa">Choix.</location>
1miN,
<quantity id="9769FCE6FF8DB07AFB79C28397A67FAB" box="[1203,1278,1504,1531]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="41" pageNumber="242" unit="m" value="442.0">
<elevation id="DBBCB630FF8DB07AFB79C28397A67FAB" box="[1203,1278,1504,1531]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="41" pageNumber="242" unit="m" value="442.0">442 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8DB07AFAC6C28396C47FAA" box="[1292,1436,1504,1530]" degrees="26.72861111" direction="north" orientation="latitude" pageId="41" pageNumber="242" precision="1" value="26.728611">26.72861111</geoCoordinate>
<geoCoordinate id="35A537C4FF8DB07AFF5DC16792547C4F" box="[151,268,1540,1567]" degrees="108.2975" direction="west" orientation="longitude" pageId="41" pageNumber="242" precision="5" value="-108.2975">-108.2975</geoCoordinate>
,
<date id="242F77C3FF8DB07AFED1C16792F07C4E" box="[283,424,1540,1566]" pageId="41" pageNumber="242" value="1968-07-30">
<collectingDate id="346B8E2BFF8DB07AFED1C16792F07C4E" box="[283,424,1540,1566]" pageId="41" pageNumber="242" value="1968-07-30">30/VII/1968</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8DB07AFE7DC16691647C4E" box="[439,572,1540,1567]" pageId="41" pageNumber="242">T. A. Sears</collectorName>
,
<collectorName id="FD6434D5FF8DB07AFD81C16691AA7C4E" box="[587,754,1540,1567]" pageId="41" pageNumber="242">R. C. Gardner</collectorName>
,
<collectorName id="FD6434D5FF8DB07AFCCBC16790C87C4E" box="[769,912,1540,1566]" pageId="41" pageNumber="242">C. S. Glaser</collectorName>
(
<collectionCode id="3680C9C6FF8DB07AFC68C16790B17C4E" box="[930,1001,1540,1566]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="41" pageNumber="242" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8DB07AFC3EC167974D7C4E" box="[1012,1045,1540,1566]" count="1" pageId="41" pageNumber="242" type="female">1♀</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF8DB07AFBF8C16797477C12" collectingDate="1968-08-31" collectionCode="BBSL" collectorName="T. A. Sears &amp; R. C. Gardner &amp; Glaser" country="Mexico" elevation="442" latitude="26.728611" longLatPrecision="4" longitude="-108.2975" pageId="41" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="Sinaloa">
4miNW,
<quantity id="9769FCE6FF8DB07AFB56C16797B07C4F" box="[1180,1256,1540,1567]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="41" pageNumber="242" unit="m" value="442.0">
<elevation id="DBBCB630FF8DB07AFB56C16797B07C4F" box="[1180,1256,1540,1567]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="41" pageNumber="242" unit="m" value="442.0">442 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8DB07AFB32C16796D07C4E" box="[1272,1416,1540,1566]" degrees="26.72861111" direction="north" orientation="latitude" pageId="41" pageNumber="242" precision="1" value="26.728611">26.72861111</geoCoordinate>
<geoCoordinate id="35A537C4FF8DB07AFA59C166925B7C13" degrees="108.2975" direction="west" orientation="longitude" pageId="41" pageNumber="242" precision="5" value="-108.2975">- 108.2975</geoCoordinate>
,
<date id="242F77C3FF8DB07AFED8C14B92F07C12" box="[274,424,1576,1602]" pageId="41" pageNumber="242" value="1968-08-31">
<collectingDate id="346B8E2BFF8DB07AFED8C14B92F07C12" box="[274,424,1576,1602]" pageId="41" pageNumber="242" value="1968-08-31">31/VIII/1968</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8DB07AFE7DC14A91637C12" box="[439,571,1576,1603]" pageId="41" pageNumber="242">T. A. Sears</collectorName>
,
<collectorName id="FD6434D5FF8DB07AFD80C14A91A97C12" box="[586,753,1576,1603]" pageId="41" pageNumber="242">R. C. Gardner</collectorName>
, C. S.
<collectorName id="FD6434D5FF8DB07AFC8EC14B90D57C12" box="[836,909,1576,1602]" pageId="41" pageNumber="242">Glaser</collectorName>
(
<collectionCode id="3680C9C6FF8DB07AFC6AC14B90BF7C12" box="[928,999,1576,1602]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="41" pageNumber="242" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8DB07AFC3BC14B974D7C12" box="[1009,1045,1576,1602]" count="1" pageId="41" pageNumber="242" type="male">1♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF8DB07AFBE5C14B97317C36" collectingDate="1968-09-05" collectionCode="BBSL" collectorName="T. A. Sears &amp; R. C. Gardner &amp; Glaser" country="Mexico" elevation="465" latitude="26.728056" longLatPrecision="1" longitude="-108.29611" pageId="41" pageNumber="242" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sinaloa">
5.5miNW,
<quantity id="9769FCE6FF8DB07AFB67C14B97A27C13" box="[1197,1274,1576,1603]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="41" pageNumber="242" unit="m" value="465.0">
<elevation id="DBBCB630FF8DB07AFB67C14B97A27C13" box="[1197,1274,1576,1603]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="41" pageNumber="242" unit="m" value="465.0">465 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8DB07AFAC3C14B96C47C13" box="[1289,1436,1576,1603]" degrees="26.72805556" direction="north" orientation="latitude" pageId="41" pageNumber="242" precision="1" value="26.728056">26.72805556</geoCoordinate>
<geoCoordinate id="35A537C4FF8DB07AFF5DC12F926B7C36" box="[151,307,1612,1638]" degrees="108.2961111" direction="west" orientation="longitude" pageId="41" pageNumber="242" precision="1" value="-108.29611">-108.2961111</geoCoordinate>
,
<date id="242F77C3FF8DB07AFE89C12F92E17C37" box="[323,441,1612,1639]" pageId="41" pageNumber="242" value="1968-09-05">
<collectingDate id="346B8E2BFF8DB07AFE89C12F92E17C37" box="[323,441,1612,1639]" pageId="41" pageNumber="242" value="1968-09-05">5/IX/1968</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8DB07AFE02C12E91167C36" box="[456,590,1612,1639]" pageId="41" pageNumber="242">T. A. Sears</collectorName>
,
<collectorName id="FD6434D5FF8DB07AFD94C12E905F7C36" box="[606,775,1612,1639]" pageId="41" pageNumber="242">R. C. Gardner</collectorName>
, C. S.
<collectorName id="FD6434D5FF8DB07AFC97C12F90FE7C36" box="[861,934,1612,1638]" pageId="41" pageNumber="242">Glaser</collectorName>
(
<collectionCode id="3680C9C6FF8DB07AFC70C12F97597C36" box="[954,1025,1612,1638]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="41" pageNumber="242" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8DB07AFBC6C12F97777C36" box="[1036,1071,1612,1638]" count="1" pageId="41" pageNumber="242" type="female">1♀</specimenCount>
<specimenCount id="46979A8AFF8DB07AFBF1C12F97077C36" box="[1083,1119,1612,1638]" count="1" pageId="41" pageNumber="242" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8DB07AFBB0C12F90D77CDA" collectingDate="1991-09-03" collectorName="L. M. Godinez" country="Mexico" elevation="440" latitude="27.025555" location="Alamos" longLatPrecision="1" longitude="-108.94139" pageId="41" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="Sonora">
<emphasis id="62E58D11FF8DB07AFBB0C12F978E7C36" bold="true" box="[1146,1238,1612,1638]" pageId="41" pageNumber="242">
<collectingRegion id="92559FE1FF8DB07AFBB0C12F97897C36" box="[1146,1233,1612,1638]" country="Mexico" name="Sonora" pageId="41" pageNumber="242">Sonora</collectingRegion>
,
</emphasis>
<location id="554E07D8FF8DB07AFB2AC12F96627C36" LSID="urn:lsid:plazi:treatment:D838E015FF8CB07AFF5DC10591A37DA3:554E07D8FF8DB07AFB2AC12F96627C36" box="[1248,1338,1612,1638]" country="Mexico" latitude="27.025555" longLatPrecision="1" longitude="-108.94139" name="Alamos" pageId="41" pageNumber="242" stateProvince="Sonora">Alamos</location>
,
<quantity id="9769FCE6FF8DB07AFA80C12F96CF7C37" box="[1354,1431,1612,1639]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="41" pageNumber="242" unit="m" value="440.0">
<elevation id="DBBCB630FF8DB07AFA80C12F96CF7C37" box="[1354,1431,1612,1639]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="41" pageNumber="242" unit="m" value="440.0">440 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8DB07AFF5DC11392727CDB" box="[151,298,1648,1675]" degrees="27.02555556" direction="north" orientation="latitude" pageId="41" pageNumber="242" precision="1" value="27.025555">27.02555556</geoCoordinate>
<geoCoordinate id="35A537C4FF8DB07AFEF8C11392897CDA" box="[306,465,1648,1674]" degrees="108.9413889" direction="west" orientation="longitude" pageId="41" pageNumber="242" precision="1" value="-108.94139">-108.9413889</geoCoordinate>
,
<date id="242F77C3FF8DB07AFE17C113910B7CDA" box="[477,595,1648,1674]" pageId="41" pageNumber="242" value="1991-09-03">
<collectingDate id="346B8E2BFF8DB07AFE17C113910B7CDA" box="[477,595,1648,1674]" pageId="41" pageNumber="242" value="1991-09-03">3/IX/1991</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8DB07AFD94C112905C7CDA" box="[606,772,1648,1675]" pageId="41" pageNumber="242">L. M. Godinez</collectorName>
(EBCh
<specimenCount id="46979A8AFF8DB07AFCABC11390DD7CDA" box="[865,901,1648,1674]" count="1" pageId="41" pageNumber="242" type="male">1♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF8DB07AFC56C11391747CFE" collectingDate="1975-09-01" collectionCode="BBSL" collectorName="Villegas" country="Mexico" elevation="387" latitude="27.025555" longLatPrecision="1" longitude="-108.94139" pageId="41" pageNumber="242" specimenCount="1" specimenCount-female="1" stateProvince="Sonora">
13KmW,
<quantity id="9769FCE6FF8DB07AFBC2C113970A7CDB" box="[1032,1106,1648,1675]" metricMagnitude="2" metricUnit="m" metricValue="3.87" pageId="41" pageNumber="242" unit="m" value="387.0">
<elevation id="DBBCB630FF8DB07AFBC2C113970A7CDB" box="[1032,1106,1648,1675]" metricMagnitude="2" metricUnit="m" metricValue="3.87" pageId="41" pageNumber="242" unit="m" value="387.0">387 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8DB07AFB94C11397A97CDB" box="[1118,1265,1648,1675]" degrees="27.02555556" direction="north" orientation="latitude" pageId="41" pageNumber="242" precision="1" value="27.025555">27.02555556</geoCoordinate>
<geoCoordinate id="35A537C4FF8DB07AFB33C11396C07CDA" box="[1273,1432,1648,1674]" degrees="108.9413889" direction="west" orientation="longitude" pageId="41" pageNumber="242" precision="1" value="-108.94139">-108.9413889</geoCoordinate>
,
<date id="242F77C3FF8DB07AFF5DC1F792437CFF" box="[151,283,1684,1711]" pageId="41" pageNumber="242" value="1975-09-01">
<collectingDate id="346B8E2BFF8DB07AFF5DC1F792437CFF" box="[151,283,1684,1711]" pageId="41" pageNumber="242" value="1975-09-01">01/IX/1975</collectingDate>
</date>
, B.
<collectorName id="FD6434D5FF8DB07AFE8CC1F792FA7CFE" box="[326,418,1684,1710]" pageId="41" pageNumber="242">Villegas</collectorName>
(
<collectionCode id="3680C9C6FF8DB07AFE7AC1F792AF7CFE" box="[432,503,1684,1710]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="41" pageNumber="242" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8DB07AFE34C1F791787CFE" box="[510,544,1684,1710]" count="1" pageId="41" pageNumber="242" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8DB07AFDF2C1F792B97C82" collectingDate="1977-09-27" collectionCode="BBSL" collectorName="J. Chemsak &amp; Michelbachers" country="Mexico" elevation="469" latitude="29.689722" location="Carbo" longLatPrecision="1" longitude="-110.956665" pageId="41" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="Sonora">
<location id="554E07D8FF8DB07AFDF2C1F791277CFE" LSID="urn:lsid:plazi:treatment:D838E015FF8CB07AFF5DC10591A37DA3:554E07D8FF8DB07AFDF2C1F791277CFE" box="[568,639,1684,1710]" country="Mexico" latitude="29.689722" longLatPrecision="1" longitude="-110.956665" name="Carbo" pageId="41" pageNumber="242" stateProvince="Sonora">Carbo</location>
, 15miS, [
<quantity id="9769FCE6FF8DB07AFD22C1F7906C7CFF" box="[744,820,1684,1711]" metricMagnitude="2" metricUnit="m" metricValue="4.6899999999999995" pageId="41" pageNumber="242" unit="m" value="469.0">
<elevation id="DBBCB630FF8DB07AFD22C1F7906C7CFF" box="[744,820,1684,1711]" metricMagnitude="2" metricUnit="m" metricValue="4.6899999999999995" pageId="41" pageNumber="242" unit="m" value="469.0">469 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8DB07AFC8AC1F7908B7CFE" box="[832,979,1684,1710]" degrees="29.68972222" direction="north" orientation="latitude" pageId="41" pageNumber="242" precision="1" value="29.689722">29.68972222</geoCoordinate>
<geoCoordinate id="35A537C4FF8DB07AFC10C1F797237CFF" box="[986,1147,1684,1711]" degrees="110.9566667" direction="west" orientation="longitude" pageId="41" pageNumber="242" precision="1" value="-110.956665">-110.9566667</geoCoordinate>
],
<date id="242F77C3FF8DB07AFB46C1F796487CFE" box="[1164,1296,1684,1710]" pageId="41" pageNumber="242" value="1977-09-27">
<collectingDate id="346B8E2BFF8DB07AFB46C1F796487CFE" box="[1164,1296,1684,1710]" pageId="41" pageNumber="242" value="1977-09-27">27/IX/1977</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8DB07AFAD1C1F696C47CFE" box="[1307,1436,1684,1711]" pageId="41" pageNumber="242">J. Chemsak</collectorName>
&amp;
<collectorName id="FD6434D5FF8DB07AFF79C1DB92007C82" box="[179,344,1720,1746]" pageId="41" pageNumber="242">Michelbachers</collectorName>
(
<collectionCode id="3680C9C6FF8DB07AFEAFC1DB92F47C82" box="[357,428,1720,1746]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="41" pageNumber="242" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8DB07AFE78C1DB928F7C82" box="[434,471,1720,1746]" count="1" pageId="41" pageNumber="242" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8DB07AFE26C1DB92537CA6" collectingDate="1970-09-30" collectionCode="BBSL" collectorName="RM Bohart" country="Mexico" elevation="25" latitude="27.974445" location="San Carlos" longLatPrecision="1" longitude="-111.03778" pageId="41" pageNumber="242" specimenCount="58" specimenCount-female="15" specimenCount-male="43" stateProvince="Sonora">
<location id="554E07D8FF8DB07AFE26C1DB91307C82" LSID="urn:lsid:plazi:treatment:D838E015FF8CB07AFF5DC10591A37DA3:554E07D8FF8DB07AFE26C1DB91307C82" box="[492,616,1720,1746]" country="Mexico" latitude="27.974445" longLatPrecision="1" longitude="-111.03778" name="San Carlos" pageId="41" pageNumber="242" stateProvince="Sonora">San Carlos</location>
, [
<quantity id="9769FCE6FF8DB07AFDB3C1DB91ED7C83" box="[633,693,1720,1747]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="41" pageNumber="242" unit="m" value="25.0">
<elevation id="DBBCB630FF8DB07AFDB3C1DB91ED7C83" box="[633,693,1720,1747]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="41" pageNumber="242" unit="m" value="25.0">25 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8DB07AFD0AC1DB900B7C82" box="[704,851,1720,1746]" degrees="27.97444444" direction="north" orientation="latitude" pageId="41" pageNumber="242" precision="1" value="27.974445">27.97444444</geoCoordinate>
<geoCoordinate id="35A537C4FF8DB07AFC93C1DB90A17C82" box="[857,1017,1720,1746]" degrees="111.0377778" direction="west" orientation="longitude" pageId="41" pageNumber="242" precision="1" value="-111.03778">-111.0377778</geoCoordinate>
],
<date id="242F77C3FF8DB07AFBC0C1DB97D67C82" box="[1034,1166,1720,1746]" pageId="41" pageNumber="242" value="1970-09-30">
<collectingDate id="346B8E2BFF8DB07AFBC0C1DB97D67C82" box="[1034,1166,1720,1746]" pageId="41" pageNumber="242" value="1970-09-30">30/IX/1970</collectingDate>
</date>
, GE,
<collectorName id="FD6434D5FF8DB07AFB00C1DA96107C82" box="[1226,1352,1720,1747]" pageId="41" pageNumber="242">RM Bohart</collectorName>
(
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<specimenCount id="46979A8AFF8DB07AFF5DC1BF93907CA7" box="[151,200,1756,1783]" count="15" pageId="41" pageNumber="242" type="female">15♀</specimenCount>
<specimenCount id="46979A8AFF8DB07AFF04C1BF925A7CA6" box="[206,258,1756,1782]" count="43" pageId="41" pageNumber="242" type="male">43♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF8DB07AFEDCC1BF97897CA6" box="[278,1233,1756,1783]" collectingDate="1983-08-15" collectionCode="BBSL" collectorName="Fisher" country="Mexico" elevation="25" latitude="27.974445" longLatPrecision="1" longitude="-111.03778" pageId="41" pageNumber="242" specimenCount="2" specimenCount-male="2" stateProvince="Sonora">
6kmNNW,
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<elevation id="DBBCB630FF8DB07AFE5FC1BF92977CA7" box="[405,463,1756,1783]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="41" pageNumber="242" unit="m" value="25.0">25 m</elevation>
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,
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, 11,
<date id="242F77C3FF8DB07AFC88C1BF90817CA7" box="[834,985,1756,1783]" pageId="41" pageNumber="242" value="1983-08-15">
<collectingDate id="346B8E2BFF8DB07AFC88C1BF90817CA7" box="[834,985,1756,1783]" pageId="41" pageNumber="242" value="1983-08-15">15/VIII/1983</collectingDate>
</date>
, E.
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(
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)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8DB07AFB11C1BF97FE7D4A" collectingDate="1963-07-14" collectionCode="BBSL" collectorName="R. L. Westcott" country="Mexico" latitude="27.946112" location="Guacamayas" longLatPrecision="1" longitude="-111.06167" municipality="Bahia San Carlos" pageId="41" pageNumber="242" specimenCount="2" specimenCount-male="2" stateProvince="Sonora">
<collectingMunicipality id="B04ACB79FF8DB07AFB11C1BF96C47CA6" box="[1243,1436,1756,1782]" pageId="41" pageNumber="242">Bahía San Carlos</collectingMunicipality>
nr.
<location id="554E07D8FF8DB07AFF71C06392157D4A" LSID="urn:lsid:plazi:treatment:D838E015FF8CB07AFF5DC10591A37DA3:554E07D8FF8DB07AFF71C06392157D4A" box="[187,333,1792,1818]" country="Mexico" latitude="27.946112" longLatPrecision="1" longitude="-111.06167" municipality="Bahia San Carlos" name="Guacamayas" pageId="41" pageNumber="242" stateProvince="Sonora">Guacamayas</location>
, [0 m,
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],
<date id="242F77C3FF8DB07AFD28C06390377D4A" box="[738,879,1792,1818]" pageId="41" pageNumber="242" value="1963-07-14">
<collectingDate id="346B8E2BFF8DB07AFD28C06390377D4A" box="[738,879,1792,1818]" pageId="41" pageNumber="242" value="1963-07-14">14/VII/1963</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8DB07AFCB0C06297427D4B" box="[890,1050,1793,1819]" pageId="41" pageNumber="242">R. L. Westcott</collectorName>
(
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<specimenCount id="46979A8AFF8DB07AFBBDC06397C57D4A" box="[1143,1181,1792,1818]" count="2" pageId="41" pageNumber="242" type="male">2♂</specimenCount>
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.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8DB07AFF0DC04796357DD6" pageId="41" pageNumber="242" type="distribution">
<paragraph id="502E5103FF8DB07AFF0DC04796357DD6" blockId="41.[151,1437,1171,2035]" pageId="41" pageNumber="242">
<emphasis id="62E58D11FF8DB07AFF0DC04792067D6E" bold="true" box="[199,350,1828,1854]" pageId="41" pageNumber="242">Distribution.</emphasis>
This species is present in the states of
<collectingRegion id="92559FE1FF8DB07AFD34C047900E7D6E" box="[766,854,1828,1854]" country="United States of America" name="Arizona" pageId="41" pageNumber="242">Arizona</collectingRegion>
and
<collectingRegion id="92559FE1FF8DB07AFC40C04697407D6E" box="[906,1048,1828,1855]" country="United States of America" name="New Mexico" pageId="41" pageNumber="242">New Mexico</collectingRegion>
in the
<collectingCountry id="28861193FF8DB07AFBAAC04797AD7D6E" box="[1120,1269,1828,1854]" name="United States of America" pageId="41" pageNumber="242">United States</collectingCountry>
, and in
<collectingCountry id="28861193FF8DB07AFA82C04796C47D6E" box="[1352,1436,1828,1854]" name="Mexico" pageId="41" pageNumber="242">Mexico</collectingCountry>
in the states of
<collectingRegion id="92559FE1FF8DB07AFEF3C02B92EB7D32" box="[313,435,1864,1890]" country="Mexico" name="Chihuahua" pageId="41" pageNumber="242">Chihuahua</collectingRegion>
,
<collectingRegion id="92559FE1FF8DB07AFE76C02B91527D32" box="[444,522,1864,1890]" country="Mexico" name="Sonora" pageId="41" pageNumber="242">Sonora</collectingRegion>
and northern
<collectingRegion id="92559FE1FF8DB07AFD6AC02B91AD7D32" box="[672,757,1864,1890]" country="Mexico" name="Sinaloa" pageId="41" pageNumber="242">Sinaloa</collectingRegion>
, mostly in xeric areas of the Sonoran, Sierra Madre Occidental and Chihuahuan biogeographic provinces (
<bibRefCitation id="34002CF2FF8DB07AFDA4C00F90087DD6" author="Morrone, J. J. &amp; Escalante, T. &amp; Rodriguez-Tapia, G. &amp; Carmona, A. &amp; Arana, M. &amp; Mercado-Gomez, J. D." box="[622,848,1900,1926]" pageId="41" pageNumber="242" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. &amp; Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="62E58D11FF8DB07AFD1CC00E90567DD6" box="[726,782,1900,1926]" italics="true" pageId="41" pageNumber="242">et al.</emphasis>
2022
</bibRefCitation>
). Altitudinal range: sea level to
<quantity id="9769FCE6FF8DB07AFB66C00F97A77DD7" box="[1196,1279,1900,1927]" metricMagnitude="3" metricUnit="m" metricValue="1.426" pageId="41" pageNumber="242" unit="m" value="1426.0">1426 m</quantity>
(
<figureCitation id="C8AA4D86FF8DB07AFAC4C00F96387DD6" box="[1294,1376,1900,1926]" captionStart="FIGURE 24" captionStartId="41.[151,250,1098,1122]" captionTargetBox="[160,1428,187,1063]" captionTargetId="figure-18@41.[151,1436,179,1072]" captionTargetPageId="41" captionText="FIGURE 24. Known distribution records for Heriades micheneri." pageId="41" pageNumber="242">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8DB07AFF0DC0F391A37DA3" pageId="41" pageNumber="242" type="discussion">
<paragraph id="502E5103FF8DB07AFF0DC0F391A37DA3" blockId="41.[151,1437,1171,2035]" pageId="41" pageNumber="242">
<emphasis id="62E58D11FF8DB07AFF0DC0F392107DFA" bold="true" box="[199,328,1936,1962]" pageId="41" pageNumber="242">Comments</emphasis>
. The size of ocelli varies but seems to be always bigger than the surrounding punctures on the vertex. The color on tufts of dorsolateral angles of pronotum and others varies from a yellowish white to orange but more frequently they are orange on
<taxonomicName id="97912A80FF8DB07AFDE2C0BA91C67DA2" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[552,670,2008,2034]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="amanda" status="sp. nov.">
<emphasis id="62E58D11FF8DB07AFDE2C0BA91C67DA2" box="[552,670,2008,2034]" italics="true" pageId="41" pageNumber="242">H amanda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF8DB07AFD6FC0BA91A37DA3" box="[677,763,2009,2035]" pageId="41" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,779 @@
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<mods:title id="981645D1703A0F3CFB6394576AF66437">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<taxonomicName id="97912A80FF8FB078FF5DC3F591807EE0" authority="Rojas-Arias, Griswold &amp; Ayala." authorityName="Rojas-Arias, Griswold &amp; Ayala." authorityYear="2025" box="[151,728,1174,1201]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="244" phylum="Arthropoda" rank="species" species="nora" status="sp. nov.">
<emphasis id="62E58D11FF8FB078FF5DC3F5921D7EE1" bold="true" box="[151,325,1174,1201]" italics="true" pageId="43" pageNumber="244">Heriades nora</emphasis>
Rojas-Arias, Griswold &amp; Ayala.
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<subSubSection id="188B0288FF8FB078FF5DC3D992277E84" box="[151,383,1210,1236]" pageId="43" pageNumber="244" type="description">
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(
<figureCitation id="C8AA4D86FF8FB078FF55C3D993B57E84" box="[159,237,1210,1236]" captionStart="FIGURE 1" captionStartId="2.[151,250,1938,1962]" captionTargetBox="[151,1436,918,1912]" captionTargetId="figure-284@2.[151,1436,918,1912]" captionTargetPageId="2" captionText="FIGURE 1. S1 of male of H. nora, on ventral view as an example, showing (a) profile with posterior margin of sterna and whole metasoma to the right, (b) posterior margin, and (c) ventral process of S1 posterior view with dorsal surface up." pageId="43" pageNumber="244">Figs. 1</figureCitation>
,
<figureCitation id="C8AA4D86FF8FB078FF31C3D9927C7E84" box="[251,292,1210,1236]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="43" pageNumber="244">2M</figureCitation>
,
<figureCitation id="C8AA4D86FF8FB078FEFAC3D9922F7E84" box="[304,375,1210,1236]" captionStart-0="FIGURE 26" captionStart-1="FIGURE 27" captionStartId-0="43.[151,250,1068,1092]" captionStartId-1="45.[151,250,1943,1967]" captionTargetBox-0="[159,1428,157,1034]" captionTargetBox-1="[159,1419,192,1905]" captionTargetId-0="figure-18@43.[151,1436,149,1042]" captionTargetId-1="figure-18@45.[159,1428,179,1917]" captionTargetPageId-0="43" captionTargetPageId-1="45" captionText-0="FIGURE 26. Known distribution records for Heriades nora sp. nov." captionText-1="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="43" pageNumber="244">2627</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8FB078FF5DC26196147C30" pageId="43" pageNumber="244" type="diagnosis">
<paragraph id="502E5103FF8FB078FF5DC26196147C30" blockId="43.[151,1437,1282,2029]" pageId="43" pageNumber="244">
<emphasis id="62E58D11FF8FB078FF5DC261924A7F4C" bold="true" box="[151,274,1282,1308]" pageId="43" pageNumber="244">Diagnosis.</emphasis>
Base of clypeus with small punctures (0.1 times OD); tergal fasciae with row of thick, white hairs (
<figureCitation id="C8AA4D86FF8FB078FAA5C26193EB7F10" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="43" pageNumber="244">Fig. 3J</figureCitation>
).
<emphasis id="62E58D11FF8FB078FF00C245925C7F10" box="[202,260,1318,1344]" italics="true" pageId="43" pageNumber="244">Male</emphasis>
: clypeus with plumose, thick, dense, short, white to goldish hairs directed medially, joining long hairs on hypostomal area (
<figureCitation id="C8AA4D86FF8FB078FE49C22992B47F34" box="[387,492,1354,1380]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="43" pageNumber="244">Fig. 27A</figureCitation>
); posterior margin of S1 truncate without emargination (
<figureCitation id="C8AA4D86FF8FB078FBAFC22997F07F34" box="[1125,1192,1354,1380]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="43" pageNumber="244">Fig. 2</figureCitation>
Mb); posterior half of S2 and posterior third of S3 with plumose, thin, long, dense, white hairs; posterior margin of S5 wavy, with two small tufts of capitate hairs on lateral margins directed radially outward (
<figureCitation id="C8AA4D86FF8FB078FC1CC2F197657FFC" box="[982,1085,1426,1452]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="43" pageNumber="244">Fig. 27E</figureCitation>
); S6 with one subapical tuft of several hooked, thin hairs increasing in length outwards (
<figureCitation id="C8AA4D86FF8FB078FCECC2D590D47F80" box="[806,908,1462,1488]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="43" pageNumber="244">Fig. 27F</figureCitation>
).
<emphasis id="62E58D11FF8FB078FC6BC2D590AD7F80" box="[929,1013,1462,1488]" italics="true" pageId="43" pageNumber="244">Female</emphasis>
: with outer ridge on outer surface of mandible with low elevation on its most basal point; clypeus and supraclypeal area with small punctures; clypeus with emargination and a more than five small, very rounded tubercles, almost crenulated margin (
<figureCitation id="C8AA4D86FF8FB078FB1EC29D96637C48" box="[1236,1339,1534,1560]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="43" pageNumber="244">Fig. 27C</figureCitation>
); similar to
<taxonomicName id="97912A80FF8FB078FF7EC14092547C6C" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[180,268,1570,1596]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="244" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
<emphasis id="62E58D11FF8FB078FF7EC14092547C6C" box="[180,268,1570,1596]" italics="true" pageId="43" pageNumber="244">H. luisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF8FB078FED9C14092327C6D" box="[275,362,1571,1597]" pageId="43" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
but differing from this by having clypeus in profile not projected on apical margin, scutum with large punctures (0.55 times OD), propodeum with large pits (
<figureCitation id="C8AA4D86FF8FB078FC8EC12590C87C30" box="[836,912,1606,1632]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="43" pageNumber="244">Fig. 3I</figureCitation>
) and T3 with large punctures (
<figureCitation id="C8AA4D86FF8FB078FB22C12596677C30" box="[1256,1343,1606,1632]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="43" pageNumber="244">Fig. 3L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8FB07DFF0DC10997177AA9" lastPageId="46" lastPageNumber="247" pageId="43" pageNumber="244" type="description">
<paragraph id="502E5103FF8FB078FF0DC109964B7CD4" blockId="43.[151,1437,1282,2029]" box="[199,1299,1642,1669]" pageId="43" pageNumber="244">
<emphasis id="62E58D11FF8FB078FF0DC10992FC7CD4" bold="true" box="[199,420,1642,1669]" pageId="43" pageNumber="244">
Description.
<emphasis id="62E58D11FF8FB078FEAAC10992C37CD5" bold="true" box="[352,411,1642,1669]" italics="true" pageId="43" pageNumber="244">Male</emphasis>
:
</emphasis>
Body length,
<quantity id="9769FCE6FF8FB078FD89C10991C17CD5" box="[579,665,1642,1669]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="43" pageNumber="244" unit="mm" value="6.3">6.3 mm</quantity>
(5.56.5, n=6); forewing length,
<quantity id="9769FCE6FF8FB078FBC5C109973D7CD5" box="[1039,1125,1642,1669]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="43" pageNumber="244" unit="mm" value="3.7">3.7 mm</quantity>
(3.64.2, n=6).
</paragraph>
<paragraph id="502E5103FF8FB07FFF0DC1ED97617AA8" blockId="43.[151,1437,1282,2029]" lastBlockId="44.[151,1437,150,2036]" lastPageId="44" lastPageNumber="245" pageId="43" pageNumber="244">
<emphasis id="62E58D11FF8FB078FF0DC1ED925C7CF8" box="[199,260,1678,1704]" italics="true" pageId="43" pageNumber="244">Head</emphasis>
. Length
<quantity id="9769FCE6FF8FB078FEA2C1ED92E77CF9" box="[360,447,1678,1705]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="43" pageNumber="244" unit="mm" value="1.4">1.4 mm</quantity>
(1.41.7, n=6), width
<quantity id="9769FCE6FF8FB078FD71C1ED90497CF9" box="[699,785,1678,1705]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="43" pageNumber="244" unit="mm" value="1.6">1.6 mm</quantity>
(1.561.85, n=6); eyes about 3.4 times longer than broad, 2.1 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF8FB078FCA2C1D190E77C9D" box="[872,959,1714,1741]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="43" pageNumber="244" unit="mm" value="0.8">0.8 mm</quantity>
(0.80.9, n=6); upper interocular distance
<quantity id="9769FCE6FF8FB078FF5DC1B593817CA1" box="[151,217,1750,1777]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="244" unit="mm" value="1.0">1 mm</quantity>
(11.1, n=6); interantennal distance 2.1 times antennocular distance; lateral ocelli large (
<figureCitation id="C8AA4D86FF8FB078FB0DC1B596477CA0" box="[1223,1311,1750,1776]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="43" pageNumber="244">Fig. 3C</figureCitation>
), 3.3 times DOP in interocellar area; interocellar distance 1.6 times OD; ocellocular distance 1.5 times OD, OPR 1.6 times OD; labrum without basal elevation; clypeus 1.2 times broader than long, lower half convex in profile, apical margin without emargination; scape length 0.7 times alveolocellar distance, 2.3 times longer than wide; pedicel as long as broad, about 1.3 times longer than F1 and about 0.6 times longer than F2; flagellum black, F1 broader than long, remaining flagellomeres 2 times longer than F1; F11 1.83 times longer than broad.
<emphasis id="62E58D11FF8FB078FB85C0E897967DF4" box="[1103,1230,1931,1956]" italics="true" pageId="43" pageNumber="244">Mesosoma.</emphasis>
Scutum 2.6 times longer than scutellum, 3.1 times longer than tegula, and 3.7 longer than axilla; intertegular distance
<quantity id="9769FCE6FF8FB078FAC9C0CD96027D99" box="[1283,1370,1966,1993]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="43" pageNumber="244" unit="mm" value="1.2">1.2 mm</quantity>
(1.1 1.4, n=6); metanotum with transverse ridge, in posterior view straight; propodeum with large pits (
<figureCitation id="C8AA4D86FF8FB078FB13C0B1967D7DBC" box="[1241,1317,2002,2028]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="43" pageNumber="244">Fig. 3I</figureCitation>
),
<quantity id="9769FCE6FF8FB078FAF0C0B196C57DBD" box="[1338,1437,2002,2029]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="43" pageNumber="244" unit="mm" value="0.09">0.09 mm</quantity>
long; legs black.
<emphasis id="62E58D11FF88B07FFE93C7F4928D7AE0" box="[345,469,151,176]" italics="true" pageId="44" pageNumber="245">Metasoma.</emphasis>
T1
<quantity id="9769FCE6FF88B07FFDC9C7F491027AE1" box="[515,602,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="44" pageNumber="245" unit="mm" value="1.4">1.4 mm</quantity>
broad (1.31.5, n=6); T3
<quantity id="9769FCE6FF88B07FFC4BC7F490807AE1" box="[897,984,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="44" pageNumber="245" unit="mm" value="1.6">1.6 mm</quantity>
broad (1.51.8: n=6); T6 with posterior margin not projected; S1 with posterior margin truncate without emargination (
<figureCitation id="C8AA4D86FF88B07FFBC4C7D997087A84" box="[1038,1104,186,212]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="44" pageNumber="245">Fig. 2</figureCitation>
Mb), in profile, with elongate margin of process that stand out above morphological posterior margin (
<figureCitation id="C8AA4D86FF88B07FFC0BC7BD975B7AA8" box="[961,1027,222,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="44" pageNumber="245">Fig. 2</figureCitation>
Ma).
</paragraph>
<paragraph id="502E5103FF88B07FFF0DC66197BE78BC" blockId="44.[151,1437,150,2036]" pageId="44" pageNumber="245">
<emphasis id="62E58D11FF88B07FFF0DC66192177B4C" box="[199,335,258,284]" italics="true" pageId="44" pageNumber="245">Sculpturing.</emphasis>
Clypeus with small (at base 0.1 times OD), contiguous, coarse punctures; supraclypeal area with punctures as on base of clypeus; frons with bigger and coarser punctures than those on supraclypeal area; frons in front of ocelli sparser and slightly smaller than frons; vertex with bigger, sparser punctures than on supraclypeal area, without micropunctures between punctures; hypostomal area with punctation as on clypeus; pronotum anterior surface with punctures bigger coarser than on clypeus; disc of scutum with bigger (0.3 times OD), sparser punctures than on vertex, with micropunctures between punctures; rest of mesosoma as on scutum except: scutellum with punctures slightly bigger and sparser punctures; mesepisternum with slightly bigger punctures, irregularly shaped towards ventral surface, with micropunctures between punctures; metepisternum with notably smaller punctures, less dense and coarse; T1 with smaller (0.2 times OD), denser punctures than on scutum; T2 with punctures smaller than on T1; T3 with bigger, coarser, and denser punctures than on T1, slightly rounded shape, separated by less than third DOP; T4 with slightly smaller punctures than on T3; T5 with smaller and slightly coarser punctures than on T4, DBP resembling lines; dorsal surface of T6 with bigger, equally sparser punctures as on T5, on posterior surface with bigger and slightly sparser punctation, DBP one DOP; sides of S1 with punctation bigger and sparser than on T1, and on disc smaller and denser; sides of S2 with punctures sparser than on T1, disc of S2 microreticulate without punctation.
</paragraph>
<paragraph id="502E5103FF88B07FFF0DC594972F7E02" blockId="44.[151,1437,150,2036]" pageId="44" pageNumber="245">
<emphasis id="62E58D11FF88B07FFF0DC59492687940" box="[199,304,759,784]" italics="true" pageId="44" pageNumber="245">Vestiture.</emphasis>
Clypeus with hairs, scattered on middle vertical line, directed medially; supraclypeal area with plumose, thin, white to gold, short hairs directed medially and dorsally; frons with plumose, thin, long, white to golden hairs directed dorsally; hypostomal area with hairs joining hairs in clypeus (
<figureCitation id="C8AA4D86FF88B07FFB80C45D97EC7908" box="[1098,1204,830,856]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="44" pageNumber="245">Fig. 27A</figureCitation>
); dorsolateral angles of pronotum with pubescence not forming continuous, linear band with hairs on anterior margin of scutum, and without tuft of thick, short hairs; disc of scutum with simple, thin, short, scattered, white hairs on its entire surface; mesepisternum with plumose, thin, long, white hairs absent on dorsal third; disc of metepisternum without tuft of plumose, thick, short, white hairs; tergal fasciae with row of thick, short, white hairs; T1 to T3 with simple, short, thin, white hairs in punctures; disc of T4 to T6 with simple, long, thick, whitish hairs in punctures increasing in length towards posterior terga (
<figureCitation id="C8AA4D86FF88B07FFE37C377913D7E7E" box="[509,613,1044,1070]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="44" pageNumber="245">Fig. 27B</figureCitation>
); posterior surface of subapical margin in S1 with tuft of plumose short golden hairs; S2 and posterior margin of S3 with plumose, long, thin, dense, white hairs.
</paragraph>
<paragraph id="502E5103FF88B07FFF0DC33896387F55" blockId="44.[151,1437,150,2036]" pageId="44" pageNumber="245">
<emphasis id="62E58D11FF88B07FFF0DC33892117E25" box="[199,329,1115,1141]" italics="true" pageId="44" pageNumber="245">
<taxonomicName id="97912A80FF88B07FFF0DC338921D7E25" box="[199,325,1115,1141]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="44" pageNumber="245" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
Posterior margin of S5 wavy, with two tufts of capitate hairs on lateral margins directed outward, with transverse, waved, thin, dark, subapical line (
<figureCitation id="C8AA4D86FF88B07FFD08C31C907E7EC9" box="[706,806,1151,1177]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="44" pageNumber="245">Fig. 27E</figureCitation>
); S6 with rounded, right-angled apex, with one subapical tuft of several hooked, thin hairs increasing in length radially outwards (
<figureCitation id="C8AA4D86FF88B07FFC1CC3C097647EED" box="[982,1084,1187,1213]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="44" pageNumber="245">Fig. 27F</figureCitation>
); capsule in dorsal view
<quantity id="9769FCE6FF88B07FFA90C3C096C57EED" box="[1370,1437,1187,1213]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="44" pageNumber="245" unit="mm" value="1.0">1 mm</quantity>
length, and
<quantity id="9769FCE6FF88B07FFEEAC3A492207EB1" box="[288,376,1223,1249]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="44" pageNumber="245" unit="mm" value="0.4">0.4 mm</quantity>
width, 2.3 times longer than wide, gonobase
<quantity id="9769FCE6FF88B07FFC40C3A490A97EB1" box="[906,1009,1223,1249]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="44" pageNumber="245" unit="mm" value="0.08">0.08 mm</quantity>
length, 5 times wider than long, and gonocoxite
<quantity id="9769FCE6FF88B07FFED1C38892297F55" box="[283,369,1259,1285]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="44" pageNumber="245" unit="mm" value="0.3">0.3 mm</quantity>
length, gonostylus
<quantity id="9769FCE6FF88B07FFD86C38891FA7F55" box="[588,674,1259,1285]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="245" unit="mm" value="0.9">0.9 mm</quantity>
length with acute apex, penis valve
<quantity id="9769FCE6FF88B07FFBF3C38997D77F55" box="[1081,1167,1258,1285]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="44" pageNumber="245" unit="mm" value="0.6">0.6 mm</quantity>
length (
<figureCitation id="C8AA4D86FF88B07FFB20C389960B7F55" box="[1258,1363,1258,1285]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="44" pageNumber="245">Fig. 27G</figureCitation>
).
</paragraph>
<paragraph id="502E5103FF88B07FFF0DC26D97E97C3A" blockId="44.[151,1437,150,2036]" pageId="44" pageNumber="245">
<emphasis id="62E58D11FF88B07FFF0DC26D927F7F78" bold="true" box="[199,295,1294,1321]" pageId="44" pageNumber="245">
<emphasis id="62E58D11FF88B07FFF0DC26D92467F79" bold="true" box="[199,286,1294,1321]" italics="true" pageId="44" pageNumber="245">Female</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FF88B07FFE0FC26D91507F79" box="[453,520,1294,1321]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="44" pageNumber="245" unit="mm" value="6.0">6 mm</quantity>
(5.66.7, n=6); forewing length
<quantity id="9769FCE6FF88B07FFC4AC26D90807F79" box="[896,984,1294,1321]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="44" pageNumber="245" unit="mm" value="3.8">3.8 mm</quantity>
(3.74.4, n=6). Similar to male except:
<emphasis id="62E58D11FF88B07FFF5DC251938C7F1C" box="[151,212,1330,1356]" italics="true" pageId="44" pageNumber="245">Head</emphasis>
. Length
<quantity id="9769FCE6FF88B07FFEFFC25192D17F1C" box="[309,393,1330,1356]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="44" pageNumber="245" unit="mm" value="1.6">1.6 mm</quantity>
(1.41.8, n=6), width
<quantity id="9769FCE6FF88B07FFDB4C251918B7F1C" box="[638,723,1330,1356]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="44" pageNumber="245" unit="mm" value="1.7">1.7 mm</quantity>
(1.61.8, n=6); eyes about 3.6 times longer than broad, 1.4 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF88B07FFCCBC23590017F20" box="[769,857,1366,1392]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="245" unit="mm" value="0.9">0.9 mm</quantity>
(0.80.9, n=6); upper interocular distance
<quantity id="9769FCE6FF88B07FFA8FC23596C57F20" box="[1349,1437,1366,1392]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="44" pageNumber="245" unit="mm" value="1.1">1.1 mm</quantity>
(1 1.2, n=6); interantennal distance 1.8 times antennocular distance; interocellar distance 2 times OD, ocellocular distance 1.9 times OD, OPR 1.4 times OD; labrum with one rounded, basal, medial tubercle; mandible on outer surface with outer and condylar ridge parallel without tubercle; dorsal margin of mandible with carina; clypeus with shallow emargination on medial half of length and two rounded, small, lower tubercles, crenulated on sides; scape 3.2 times longer than broad; pedicel 1.4 longer than broad, about 2.2 times longer than F1 and F2, remaining flagellomeres 1.7 longer than F1.
<emphasis id="62E58D11FF88B07FFDC5C14E91DF7C16" box="[527,647,1581,1606]" italics="true" pageId="44" pageNumber="245">Mesosoma</emphasis>
. Scutum 2.5 times longer than scutellum; intertegular distance
<quantity id="9769FCE6FF88B07FFA8DC14F96C57C17" box="[1351,1437,1580,1607]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="44" pageNumber="245" unit="mm" value="1.4">1.4 mm</quantity>
(1.11.4, n=6).
<emphasis id="62E58D11FF88B07FFE8CC13292E37C3A" box="[326,443,1617,1642]" italics="true" pageId="44" pageNumber="245">Metasoma</emphasis>
. T1
<quantity id="9769FCE6FF88B07FFE24C133911C7C3A" box="[494,580,1616,1643]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="44" pageNumber="245" unit="mm" value="1.5">1.5 mm</quantity>
broad (1.21.5, n=6); T3
<quantity id="9769FCE6FF88B07FFCACC13390E47C3A" box="[870,956,1616,1642]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="44" pageNumber="245" unit="mm" value="1.7">1.7 mm</quantity>
broad (1.51.8, n=6).
</paragraph>
<paragraph id="502E5103FF88B07FFF0DC110964C7DA3" blockId="44.[151,1437,150,2036]" pageId="44" pageNumber="245">
<emphasis id="62E58D11FF88B07FFF0DC11092097CDD" box="[199,337,1651,1677]" italics="true" pageId="44" pageNumber="245">Sculpturing.</emphasis>
Mandibles dull with micropunctures, carinae smooth and shiny; clypeus with medium-sized (0.2 times OD), contiguous, coarse punctures becoming smaller on lower two thirds, with few micropunctures between punctures; supraclypeal area with punctures slightly smaller and sparser than base of clypeus; frons with bigger, coarser and contiguous punctures than those on supralypeal area; frons in front of ocelli with slightly smaller punctures than those on frons; vertex bigger, sparser and coarser than on clypeus; gena evenly punctate as on frons in front of ocelli; scutum with slightly bigger and denser punctures than on vertex; rest of mesosoma as on scutum except: scutellum with bigger and slightly sparser punctures; mesepisternum with bigger punctures; metasomal terga bright with micropunctures between punctures; T1 and T2 with smaller (0.2 times OD) punctures than on scutum, sparser (1 time DOP), with slightly semicircular shape; T3 with punctures 0.3 times DOP as on scutum, irregularly shaped, and sparser (0.3 times DOP); dorsal surface of T6 with smaller, denser, and more coarsely punctures than on T5; sterna with punctation smaller and sparser than on scutum, microreticulate on posterior margin.
</paragraph>
<caption id="04EE018BFF89B07EFF5DC0F491A37D83" pageId="45" pageNumber="246" startId="45.[151,250,1943,1967]" targetBox="[159,1419,192,1905]" targetPageId="45" targetType="figure">
<paragraph id="502E5103FF89B07EFF5DC0F491A37D83" blockId="45.[151,1436,1943,2003]" pageId="45" pageNumber="246">
<emphasis id="62E58D11FF89B07EFF5DC0F492477DFF" bold="true" box="[151,287,1943,1967]" pageId="45" pageNumber="246">FIGURE 27.</emphasis>
<taxonomicName id="97912A80FF89B07EFEEFC0F492EF7DFF" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[293,439,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="246" phylum="Arthropoda" rank="species" species="nora" status="sp. nov.">
<emphasis id="62E58D11FF89B07EFEEFC0F492EF7DFF" box="[293,439,1943,1967]" italics="true" pageId="45" pageNumber="246">Heriades nora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF89B07EFE77C0FB91537DFF" box="[445,523,1944,1967]" pageId="45" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
Male:
<emphasis id="62E58D11FF89B07EFD9BC0F491317DFF" bold="true" box="[593,617,1943,1967]" pageId="45" pageNumber="246">A.</emphasis>
frontal view
<emphasis id="62E58D11FF89B07EFD25C0F4905E7DFF" bold="true" box="[751,774,1943,1967]" pageId="45" pageNumber="246">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FF89B07EFC28C0F490A37DFF" bold="true" box="[994,1019,1943,1967]" pageId="45" pageNumber="246">C.</emphasis>
frontal view
<emphasis id="62E58D11FF89B07EFB4AC0F497C17DFF" bold="true" box="[1152,1177,1943,1967]" pageId="45" pageNumber="246">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FF89B07EFA94C0F4962E7DFF" bold="true" box="[1374,1398,1943,1967]" pageId="45" pageNumber="246">E.</emphasis>
S5.
<emphasis id="62E58D11FF89B07EFF5DC0D893F27D83" bold="true" box="[151,170,1979,2003]" pageId="45" pageNumber="246">F.</emphasis>
S6.
<emphasis id="62E58D11FF89B07EFF12C0D893AB7D83" bold="true" box="[216,243,1979,2003]" pageId="45" pageNumber="246">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FF89B07EFD9DC0D891297D83" bold="true" box="[599,625,1979,2003]" pageId="45" pageNumber="246">H.</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="502E5103FF8AB07DFF0DC7F497177AA9" blockId="46.[151,1437,151,1329]" pageId="46" pageNumber="247">
<emphasis id="62E58D11FF8AB07DFF0DC7F492687AE0" box="[199,304,151,176]" italics="true" pageId="46" pageNumber="247">Vestiture.</emphasis>
Clypeus with few plumose, short, white hairs on sides; frons with few plumose, thin, white hairs; gena with plumose, white hairs evenly distributed (
<figureCitation id="C8AA4D86FF8AB07DFD1CC7D890667A85" box="[726,830,187,213]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="46" pageNumber="247">Fig. 27C</figureCitation>
); T4 to T6 with simple, thin, white hairs in punctures; subapical margin of T6 with linear tuft of plumose, short, yellowish hairs (
<figureCitation id="C8AA4D86FF8AB07DFC10C7BC971B7AA9" box="[986,1091,223,249]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="46" pageNumber="247">Fig. 27D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8AB07DFF0DC660914C7999" pageId="46" pageNumber="247" type="materials_examined">
<paragraph id="502E5103FF8AB07DFF0DC66090C07831" blockId="46.[151,1437,151,1329]" pageId="46" pageNumber="247">
<emphasis id="62E58D11FF8AB07DFF0DC66092237B4C" bold="true" box="[199,379,258,285]" pageId="46" pageNumber="247">Type material:</emphasis>
<materialsCitation id="E0F95B5EFF8AB07DFE4CC66097C87B11" collectingDate="2013-10-11" collectionCode="CZUG" collectorName="de Quila &amp; E. Razo &amp; A. Murias &amp; C. Uribe" country="Mexico" county="Male" elevation="1723" latitude="20.261917" location="Camino" longLatPrecision="14" longitude="-104.06417" municipality="Teocolotlan" pageId="46" pageNumber="247" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="62E58D11FF8AB07DFE4CC66090917B4C" bold="true" box="[390,969,258,285]" pageId="46" pageNumber="247">
<typeStatus id="8F2AEFA1FF8AB07DFE4CC66092AB7B4D" box="[390,499,259,285]" pageId="46" pageNumber="247" type="holotype">Holotype</typeStatus>
<collectingCounty id="B94F298FFF8AB07DFE34C66091637B4D" box="[510,571,259,285]" pageId="46" pageNumber="247">Male</collectingCounty>
(MEG0393):
<collectingCountry id="28861193FF8AB07DFD2DC661903A7B4C" box="[743,866,258,284]" name="Mexico" pageId="46" pageNumber="247">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF8AB07DFCBBC661909C7B4C" box="[881,964,258,284]" country="Mexico" name="Jalisco" pageId="46" pageNumber="247">Jalisco</collectingRegion>
,
</emphasis>
<collectingMunicipality id="B04ACB79FF8AB07DFC1EC66097037B4D" box="[980,1115,259,285]" pageId="46" pageNumber="247">Teocolotlán</collectingMunicipality>
,
<location id="554E07D8FF8AB07DFBA3C660979A7B4D" LSID="urn:lsid:plazi:treatment:D838E015FF8FB07DFF5DC3F597A57F61:554E07D8FF8AB07DFBA3C660979A7B4D" box="[1129,1218,259,285]" country="Mexico" county="Male" latitude="20.261917" longLatPrecision="14" longitude="-104.06417" municipality="Teocolotlan" name="Camino" pageId="46" pageNumber="247" stateProvince="Jalisco">Camino</location>
a
<location id="554E07D8FF8AB07DFB2EC66096C07B4D" LSID="urn:lsid:plazi:treatment:D838E015FF8FB07DFF5DC3F597A57F61:554E07D8FF8AB07DFB2EC66096C07B4D" box="[1252,1432,259,285]" country="Mexico" county="Male" latitude="20.261917" longLatPrecision="14" longitude="-104.06417" municipality="Teocolotlan" name="Sierra de Quila" pageId="46" pageNumber="247" stateProvince="Jalisco">
Sierra
<collectorName id="FD6434D5FF8AB07DFAF8C66096C07B4D" box="[1330,1432,259,285]" pageId="46" pageNumber="247">de Quila</collectorName>
</location>
,
<quantity id="9769FCE6FF8AB07DFF5DC64493B67B11" box="[151,238,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.723" pageId="46" pageNumber="247" unit="m" value="1723.0">
<elevation id="DBBCB630FF8AB07DFF5DC64493B67B11" box="[151,238,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.723" pageId="46" pageNumber="247" unit="m" value="1723.0">1723 m</elevation>
</quantity>
,
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<geoCoordinate id="35A537C4FF8AB07DFE55C64591647B11" box="[415,572,294,321]" degrees="104" direction="west" minutes="03" orientation="longitude" pageId="46" pageNumber="247" precision="15" seconds="51" value="-104.06417">104°0351”W</geoCoordinate>
,
<date id="242F77C3FF8AB07DFD82C64491997B11" box="[584,705,295,321]" pageId="46" pageNumber="247" value="2013-10-11">
<collectingDate id="346B8E2BFF8AB07DFD82C64491997B11" box="[584,705,295,321]" pageId="46" pageNumber="247" value="2013-10-11">11/X/2013</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8AB07DFD06C644907F7B11" box="[716,807,295,321]" pageId="46" pageNumber="247">E. Razo</collectorName>
,
<collectorName id="FD6434D5FF8AB07DFCFBC64490FB7B11" box="[817,931,295,321]" pageId="46" pageNumber="247">A. Murias</collectorName>
&amp;
<collectorName id="FD6434D5FF8AB07DFC0DC644977F7B11" box="[967,1063,295,321]" pageId="46" pageNumber="247">C. Uribe</collectorName>
(
<collectionCode id="3680C9C6FF8AB07DFBFCC64497D17B11" box="[1078,1161,295,321]" country="Mexico" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="46" pageNumber="247">CZUG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8AB07DFB53C645917F7BD9" collectingDate="2013-11-22" collectionCode="CZUG" collectorName="la Cienaga &amp; C. Uribe &amp; E. Razo &amp; A. Urias" country="Mexico" elevation="1481" latitude="20.253805" location="Camino" longLatPrecision="14" longitude="-104.06084" municipality="Tecolotlan" pageId="46" pageNumber="247" specimenCount="1" stateProvince="Jalisco" typeStatus="allotype">
<emphasis id="62E58D11FF8AB07DFB53C645922E7B34" bold="true" pageId="46" pageNumber="247">
<typeStatus id="8F2AEFA1FF8AB07DFB53C64597A67B10" box="[1177,1278,294,320]" pageId="46" pageNumber="247" type="allotype">Allotype</typeStatus>
(MEG0394):
<collectingCountry id="28861193FF8AB07DFF5DC629924A7B34" box="[151,274,330,356]" name="Mexico" pageId="46" pageNumber="247">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF8AB07DFED5C629922A7B34" box="[287,370,330,356]" country="Mexico" name="Jalisco" pageId="46" pageNumber="247">Jalisco</collectingRegion>
,
</emphasis>
<collectingMunicipality id="B04ACB79FF8AB07DFEB5C62892A07B35" box="[383,504,331,357]" pageId="46" pageNumber="247">Tecolotlán</collectingMunicipality>
,
<location id="554E07D8FF8AB07DFDC9C62891047B35" LSID="urn:lsid:plazi:treatment:D838E015FF8FB07DFF5DC3F597A57F61:554E07D8FF8AB07DFDC9C62891047B35" box="[515,604,331,357]" country="Mexico" latitude="20.253805" longLatPrecision="14" longitude="-104.06084" municipality="Tecolotlan" name="Camino" pageId="46" pageNumber="247" stateProvince="Jalisco">Camino</location>
a
<collectorName id="FD6434D5FF8AB07DFDB2C62891A97B35" box="[632,753,331,357]" pageId="46" pageNumber="247">la Ciénaga</collectorName>
BTC,
<quantity id="9769FCE6FF8AB07DFCF4C62890CE7B35" box="[830,918,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.4809999999999999" pageId="46" pageNumber="247" unit="m" value="1481.0">
<elevation id="DBBCB630FF8AB07DFCF4C62890CE7B35" box="[830,918,331,357]" metricMagnitude="3" metricUnit="m" metricValue="1.4809999999999999" pageId="46" pageNumber="247" unit="m" value="1481.0">1481 m</elevation>
</quantity>
,
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<geoCoordinate id="35A537C4FF8AB07DFB83C62897BE7B35" box="[1097,1254,331,357]" degrees="104" direction="west" minutes="03" orientation="longitude" pageId="46" pageNumber="247" precision="15" seconds="39" value="-104.06084">104°0339”W</geoCoordinate>
,
<date id="242F77C3FF8AB07DFB38C628962E7B35" box="[1266,1398,331,357]" pageId="46" pageNumber="247" value="2013-11-22">
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</date>
,
<collectorName id="FD6434D5FF8AB07DFA49C62893827BD9" pageId="46" pageNumber="247">C. Uribe</collectorName>
,
<collectorName id="FD6434D5FF8AB07DFF2FC60C92667BD9" box="[229,318,367,393]" pageId="46" pageNumber="247">E. Razo</collectorName>
&amp;
<collectorName id="FD6434D5FF8AB07DFE95C60C92E67BD9" box="[351,446,367,393]" pageId="46" pageNumber="247">A. Urias</collectorName>
(
<collectionCode id="3680C9C6FF8AB07DFE06C60C91477BD9" box="[460,543,367,393]" country="Mexico" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="46" pageNumber="247">CZUG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8AB07DFDFBC60C97257BFD" collectingDate="1985-10-01" collectingDateMax="1985-10-08" collectingDateMin="1985-10-01" collectionCode="BBSL" collectorName="R. Ayala &amp; T. L. Griswold" country="Mexico" elevation="91" latitude="19.498333" location="Chamela." longLatPrecision="21" longitude="-105.04444" pageId="46" pageNumber="247" specimenCount="99" specimenCount-female="4" specimenCount-male="4" stateProvince="Jalisco" typeStatus="paratype">
<emphasis id="62E58D11FF8AB07DFDFBC60C90CF7BD8" bold="true" box="[561,919,366,393]" pageId="46" pageNumber="247">
<typeStatus id="8F2AEFA1FF8AB07DFDFBC60C91F57BD9" box="[561,685,367,393]" pageId="46" pageNumber="247" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="28861193FF8AB07DFD73C60D906C7BD8" box="[697,820,366,392]" name="Mexico" pageId="46" pageNumber="247">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF8AB07DFCF5C60D90CA7BD8" box="[831,914,366,392]" country="Mexico" name="Jalisco" pageId="46" pageNumber="247">Jalisco</collectingRegion>
,
</emphasis>
<location id="554E07D8FF8AB07DFC54C60C97517BD9" LSID="urn:lsid:plazi:treatment:D838E015FF8FB07DFF5DC3F597A57F61:554E07D8FF8AB07DFC54C60C97517BD9" box="[926,1033,367,393]" country="Mexico" latitude="19.498333" longLatPrecision="21" longitude="-105.04444" name="Chamela." pageId="46" pageNumber="247" stateProvince="Jalisco">Chamela.</location>
[
<specimenCount id="46979A8AFF8AB07DFBDCC60C97007BD9" box="[1046,1112,367,393]" count="91" pageId="46" pageNumber="247" type="generic">
<quantity id="9769FCE6FF8AB07DFBDCC60C970C7BD9" box="[1046,1108,367,393]" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="46" pageNumber="247" unit="m" value="91.0">
<elevation id="DBBCB630FF8AB07DFBDCC60C970C7BD9" box="[1046,1108,367,393]" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="46" pageNumber="247" unit="m" value="91.0">91 m</elevation>
</quantity>
.
</specimenCount>
<geoCoordinate id="35A537C4FF8AB07DFB95C60D97B07BD9" box="[1119,1256,366,393]" degrees="19" direction="north" minutes="29" orientation="latitude" pageId="46" pageNumber="247" precision="15" seconds="54" value="19.498333">19°2954”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8AB07DFB25C60D96C97BD9" box="[1263,1425,366,393]" degrees="105" direction="west" minutes="02" orientation="longitude" pageId="46" pageNumber="247" precision="15" seconds="40" value="-105.04444">105°0240”W</geoCoordinate>
], 19/VIII,
<collectingDate id="346B8E2BFF8AB07DFF36C6F191517BFD" box="[252,521,402,429]" isEnumeration="true" pageId="46" pageNumber="247">
<date id="242F77C3FF8AB07DFF36C6F1922F7BFD" box="[252,375,402,429]" pageId="46" pageNumber="247" value="1985-10-22">22/X/1985</date>
,
<date id="242F77C3FF8AB07DFE4FC6F191517BFD" box="[389,521,402,429]" pageId="46" pageNumber="247" value="1985-11-25">25/XI/1985</date>
</collectingDate>
,
<collectorName id="FD6434D5FF8AB07DFDD2C6F091257BFD" box="[536,637,403,429]" pageId="46" pageNumber="247">R. Ayala</collectorName>
,
<date id="242F77C3FF8AB07DFD46C6F190487BFD" box="[652,784,402,429]" pageId="46" pageNumber="247" value="1985-10-01" valueMax="1985-10-08" valueMin="1985-10-01">
<collectingDate id="346B8E2BFF8AB07DFD46C6F190487BFD" box="[652,784,402,429]" pageId="46" pageNumber="247" value="1985-10-01" valueMax="1985-10-08" valueMin="1985-10-01">1-8/X/1985</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8AB07DFCD7C6F090907BFD" box="[797,968,403,429]" pageId="46" pageNumber="247">T. L. Griswold</collectorName>
(
<collectionCode id="3680C9C6FF8AB07DFC10C6F097797BFD" box="[986,1057,403,429]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="46" pageNumber="247" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8AB07DFBE1C6F097147BFD" box="[1067,1100,403,429]" count="4" pageId="46" pageNumber="247" type="female">4♀</specimenCount>
<specimenCount id="46979A8AFF8AB07DFB86C6F0972C7BFD" box="[1100,1140,403,429]" count="4" pageId="46" pageNumber="247" type="male">4♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8AB07DFB46C6F0939E7BA5" collectingDate="1985-11-13" collectingDateMax="1989-07-20" collectingDateMin="1985-11-13" collectionCode="BBSL" collectorName="R. Ayala &amp; R. A. Usela &amp; Griswold" country="Mexico" elevation="91" latitude="19.498333" location="Est. Biol" longLatPrecision="21" longitude="-105.04444" municipality="Chamela" pageId="46" pageNumber="247" specimenCount="3" specimenCount-female="3" stateProvince="Jalisco" typeStatus="paratype">
<collectingMunicipality id="B04ACB79FF8AB07DFB46C6F097AA7BFD" box="[1164,1266,403,429]" pageId="46" pageNumber="247">Chamela</collectingMunicipality>
,
<location id="554E07D8FF8AB07DFACAC6F096307BFD" LSID="urn:lsid:plazi:treatment:D838E015FF8FB07DFF5DC3F597A57F61:554E07D8FF8AB07DFACAC6F096307BFD" box="[1280,1384,403,429]" country="Mexico" latitude="19.498333" longLatPrecision="21" longitude="-105.04444" municipality="Chamela" name="Est. Biol" pageId="46" pageNumber="247" stateProvince="Jalisco">Est. Biol</location>
, [
<quantity id="9769FCE6FF8AB07DFAB4C6F093F77B81" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="46" pageNumber="247" unit="m" value="91.0">
<elevation id="DBBCB630FF8AB07DFAB4C6F093F77B81" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="46" pageNumber="247" unit="m" value="91.0">91 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8AB07DFF76C6D5921E7B81" box="[188,326,438,465]" degrees="19" direction="north" minutes="29" orientation="latitude" pageId="46" pageNumber="247" precision="15" seconds="54" value="19.498333">19°2954”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8AB07DFE84C6D592A97B81" box="[334,497,438,465]" degrees="105" direction="west" minutes="02" orientation="longitude" pageId="46" pageNumber="247" precision="15" seconds="40" value="-105.04444">105°0240”W</geoCoordinate>
],
<date id="242F77C3FF8AB07DFDCEC6D591DF7B81" box="[516,647,438,465]" pageId="46" pageNumber="247" value="1985-11-18">
<collectingDate id="346B8E2BFF8AB07DFDCEC6D591DF7B81" box="[516,647,438,465]" pageId="46" pageNumber="247" value="1985-11-18">18/XI/1985</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8AB07DFD5EC6D491A07B81" box="[660,760,439,465]" pageId="46" pageNumber="247">R. Ayala</collectorName>
,
<date id="242F77C3FF8AB07DFCCFC6D590DE7B81" box="[773,902,438,465]" pageId="46" pageNumber="247" value="1985-11-13">
<collectingDate id="346B8E2BFF8AB07DFCCFC6D590DE7B81" box="[773,902,438,465]" pageId="46" pageNumber="247" value="1985-11-13">13/11/1985</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8AB07DFC58C6D497427B81" box="[914,1050,439,465]" pageId="46" pageNumber="247">R. A. Usela</collectorName>
,
<date id="242F77C3FF8AB07DFBECC6D497EB7B81" box="[1062,1203,439,465]" pageId="46" pageNumber="247" value="1989-07-20">
<collectingDate id="346B8E2BFF8AB07DFBECC6D497EB7B81" box="[1062,1203,439,465]" pageId="46" pageNumber="247" value="1989-07-20">20/VII/1989</collectingDate>
</date>
, T.
<collectorName id="FD6434D5FF8AB07DFB17C6D4961D7B81" box="[1245,1349,439,465]" pageId="46" pageNumber="247">Griswold</collectorName>
(
<collectionCode id="3680C9C6FF8AB07DFA9CC6D496C57B81" box="[1366,1437,439,465]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="46" pageNumber="247" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF8AB07DFF5DC6B893E37BA5" box="[151,187,475,501]" count="3" pageId="46" pageNumber="247" type="female">3♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8AB07DFF1EC6B8974F7BA5" box="[212,1047,474,501]" collectingDate="2019-06-29" collectionCode="ECOAB" collectorName="J. Merida" country="Mexico" elevation="92" location="La Huerta" pageId="46" pageNumber="247" specimenCount="6" specimenCount-male="6" stateProvince="Jalisco" typeStatus="paratype">
<location id="554E07D8FF8AB07DFF1EC6B892127BA5" LSID="urn:lsid:plazi:treatment:D838E015FF8FB07DFF5DC3F597A57F61:554E07D8FF8AB07DFF1EC6B892127BA5" box="[212,330,475,501]" country="Mexico" name="La Huerta" pageId="46" pageNumber="247" stateProvince="Jalisco">La Huerta</location>
,
<quantity id="9769FCE6FF8AB07DFE92C6B892CD7BA5" box="[344,405,475,501]" metricMagnitude="1" metricUnit="m" metricValue="9.2" pageId="46" pageNumber="247" unit="m" value="92.0">
<elevation id="DBBCB630FF8AB07DFE92C6B892CD7BA5" box="[344,405,475,501]" metricMagnitude="1" metricUnit="m" metricValue="9.2" pageId="46" pageNumber="247" unit="m" value="92.0">92 m</elevation>
</quantity>
, 19.4983 -105.04,
<date id="242F77C3FF8AB07DFDA5C6B891AB7BA5" box="[623,755,475,501]" pageId="46" pageNumber="247" value="2019-06-29">
<collectingDate id="346B8E2BFF8AB07DFDA5C6B891AB7BA5" box="[623,755,475,501]" pageId="46" pageNumber="247" value="2019-06-29">29/VI/2019</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8AB07DFCCBC6B890357BA5" box="[769,877,475,501]" pageId="46" pageNumber="247">J. Mérida</collectorName>
(
<collectionCode id="3680C9C6FF8AB07DFCB4C6B890877BA5" box="[894,991,475,501]" pageId="46" pageNumber="247">ECOAB</collectionCode>
<specimenCount id="46979A8AFF8AB07DFC22C6B897567BA5" box="[1000,1038,475,501]" count="6" pageId="46" pageNumber="247" type="male">6♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF8AB07DFBEEC6B897C37849" collectingDate="2014-05-23" collectionCode="CZUG" collectorName="la Cienaga &amp; E. Razo &amp; Munoz" country="Mexico" elevation="1704" latitude="20.260834" location="Camino" longLatPrecision="21" longitude="-104.06722" municipality="Tecolotlan" pageId="46" pageNumber="247" specimenCount="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<collectingMunicipality id="B04ACB79FF8AB07DFBEEC6B897C57BA5" box="[1060,1181,475,501]" pageId="46" pageNumber="247">Tecolotlán</collectingMunicipality>
,
<location id="554E07D8FF8AB07DFB61C6B8965C7BA5" LSID="urn:lsid:plazi:treatment:D838E015FF8FB07DFF5DC3F597A57F61:554E07D8FF8AB07DFB61C6B8965C7BA5" box="[1195,1284,475,501]" country="Mexico" latitude="20.260834" longLatPrecision="21" longitude="-104.06722" municipality="Tecolotlan" name="Camino" pageId="46" pageNumber="247" stateProvince="Jalisco">Camino</location>
a
<collectorName id="FD6434D5FF8AB07DFAE9C6B896C57BA5" box="[1315,1437,475,501]" pageId="46" pageNumber="247">la Ciénaga</collectorName>
BTC,
<quantity id="9769FCE6FF8AB07DFF14C69C92607849" box="[222,312,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.704" pageId="46" pageNumber="247" unit="m" value="1704.0">
<elevation id="DBBCB630FF8AB07DFF14C69C92607849" box="[222,312,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.704" pageId="46" pageNumber="247" unit="m" value="1704.0">1704 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8AB07DFE8DC69D92887849" box="[327,464,510,537]" degrees="20" direction="north" minutes="15" orientation="latitude" pageId="46" pageNumber="247" precision="15" seconds="39" value="20.260834">20°1539”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8AB07DFE13C69C912F7849" box="[473,631,511,537]" degrees="104" direction="west" minutes="04" orientation="longitude" pageId="46" pageNumber="247" precision="15" seconds="02" value="-104.06722">104°0402”W</geoCoordinate>
,
<date id="242F77C3FF8AB07DFD4FC69C90587849" box="[645,768,511,537]" pageId="46" pageNumber="247" value="2014-05-23">
<collectingDate id="346B8E2BFF8AB07DFD4FC69C90587849" box="[645,768,511,537]" pageId="46" pageNumber="247" value="2014-05-23">23/V/2014</collectingDate>
</date>
,
<collectorName id="FD6434D5FF8AB07DFCC7C69C90317849" box="[781,873,511,537]" pageId="46" pageNumber="247">E. Razo</collectorName>
&amp; A.
<collectorName id="FD6434D5FF8AB07DFC7FC69C975C7849" box="[949,1028,511,537]" pageId="46" pageNumber="247">Muñoz</collectorName>
(
<collectionCode id="3680C9C6FF8AB07DFBDCC69C973C7849" box="[1046,1124,511,537]" country="Mexico" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="46" pageNumber="247">CZUG</collectionCode>
<specimenCount id="46979A8AFF8AB07DFBA4C69C97CA7849" box="[1134,1170,511,537]" count="1" pageId="46" pageNumber="247" type="male">1♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF8AB07DFB61C69C90ED786D" collectingDate="2013-11-23" collectionCode="CZUG" collectorName="Uribe &amp; Razo &amp; Urias" country="Mexico" elevation="1723" latitude="20.261667" longLatPrecision="21" longitude="-104.06417" pageId="46" pageNumber="247" specimenCount="3" specimenCount-male="3" stateProvince="Jalisco" typeStatus="paratype">
<quantity id="9769FCE6FF8AB07DFB61C69C965D7849" box="[1195,1285,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.723" pageId="46" pageNumber="247" unit="m" value="1723.0">
<elevation id="DBBCB630FF8AB07DFB61C69C965D7849" box="[1195,1285,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.723" pageId="46" pageNumber="247" unit="m" value="1723.0">1723 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8AB07DFAD9C69D96C47849" box="[1299,1436,510,537]" degrees="20" direction="north" minutes="15" orientation="latitude" pageId="46" pageNumber="247" precision="15" seconds="42" value="20.261667">20°1542”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8AB07DFF5DC541926C786D" box="[151,308,546,573]" degrees="104" direction="west" minutes="03" orientation="longitude" pageId="46" pageNumber="247" precision="15" seconds="51" value="-104.06417">104°0351”W</geoCoordinate>
,
<date id="242F77C3FF8AB07DFE8BC540929D786D" box="[321,453,547,573]" pageId="46" pageNumber="247" value="2013-11-23">
<collectingDate id="346B8E2BFF8AB07DFE8BC540929D786D" box="[321,453,547,573]" pageId="46" pageNumber="247" value="2013-11-23">23/XI/2013</collectingDate>
</date>
, C.
<collectorName id="FD6434D5FF8AB07DFE39C540916E786D" box="[499,566,547,573]" pageId="46" pageNumber="247">Uribe</collectorName>
, E.
<collectorName id="FD6434D5FF8AB07DFDA9C54091C5786D" box="[611,669,547,573]" pageId="46" pageNumber="247">Razo</collectorName>
&amp; A.
<collectorName id="FD6434D5FF8AB07DFD2EC5409079786D" box="[740,801,547,573]" pageId="46" pageNumber="247">Urias</collectorName>
(
<collectionCode id="3680C9C6FF8AB07DFCFBC5409027786D" box="[817,895,547,573]" country="Mexico" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="46" pageNumber="247">CZUG</collectionCode>
<specimenCount id="46979A8AFF8AB07DFC4DC54090F4786D" box="[903,940,547,573]" count="3" pageId="46" pageNumber="247" type="male">3♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF8AB07DFC09C54090CC7831" collectingDate="25 / X / 2012, 22 / XI / 2013" collectionCode="CZUG" collectorName="Uribe &amp; Razo &amp; Urias." country="Mexico" elevation="1481" latitude="20.253805" longLatPrecision="14" longitude="-104.06084" pageId="46" pageNumber="247" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<quantity id="9769FCE6FF8AB07DFC09C5409743786D" box="[963,1051,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.4809999999999999" pageId="46" pageNumber="247" unit="m" value="1481.0">
<elevation id="DBBCB630FF8AB07DFC09C5409743786D" box="[963,1051,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.4809999999999999" pageId="46" pageNumber="247" unit="m" value="1481.0">1481 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF8AB07DFBE2C541979E786D" box="[1064,1222,546,573]" degrees="20" direction="north" minutes="15" orientation="latitude" pageId="46" pageNumber="247" precision="1" seconds="13.7" value="20.253805">20°1513.7”N</geoCoordinate>
<geoCoordinate id="35A537C4FF8AB07DFB04C5409634786D" box="[1230,1388,547,573]" degrees="104" direction="west" minutes="03" orientation="longitude" pageId="46" pageNumber="247" precision="15" seconds="39" value="-104.06084">104°0339”W</geoCoordinate>
,
<collectingDate id="346B8E2BFF8AB07DFAB2C54192257831" isEnumeration="true" pageId="46" pageNumber="247">
<date id="242F77C3FF8AB07DFAB2C54193B67831" pageId="46" pageNumber="247" value="2012-10-25">25/ X/2012</date>
,
<date id="242F77C3FF8AB07DFF33C52492257831" box="[249,381,583,609]" pageId="46" pageNumber="247" value="2013-11-22">22/XI/2013</date>
</collectingDate>
, C.
<collectorName id="FD6434D5FF8AB07DFE63C52492B47831" box="[425,492,583,609]" pageId="46" pageNumber="247">Uribe</collectorName>
, E.
<collectorName id="FD6434D5FF8AB07DFDDCC52491087831" box="[534,592,583,609]" pageId="46" pageNumber="247">Razo</collectorName>
&amp; A.
<collectorName id="FD6434D5FF8AB07DFD5EC52491807831" box="[660,728,583,609]" pageId="46" pageNumber="247">Urias.</collectorName>
(
<collectionCode id="3680C9C6FF8AB07DFD2CC524906C7831" box="[742,820,583,609]" country="Mexico" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="46" pageNumber="247">CZUG</collectionCode>
<specimenCount id="46979A8AFF8AB07DFCF1C52490067831" box="[827,862,583,609]" count="1" pageId="46" pageNumber="247" type="female">1♀</specimenCount>
<specimenCount id="46979A8AFF8AB07DFCAFC52490D17831" box="[869,905,583,609]" count="1" pageId="46" pageNumber="247" type="male">1♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="502E5103FF8AB07DFF0DC509914C7999" blockId="46.[151,1437,151,1329]" pageId="46" pageNumber="247">
<emphasis id="62E58D11FF8AB07DFF0DC509916178D4" bold="true" box="[199,569,618,644]" pageId="46" pageNumber="247">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFF8AB07DFD8BC509902E78F9" collectingDate="1968-08-10" collectionCode="BBSL" collectorName="Sears &amp; Gardner &amp; Glaser" country="Mexico" elevation="2013" latitude="28.511389" location="Santo Nino" longLatPrecision="20" longitude="-106.71583" pageId="46" pageNumber="247" specimenCount="1" specimenCount-male="1" stateProvince="Chihuahua">
<emphasis id="62E58D11FF8AB07DFD8BC509900E78D4" bold="true" box="[577,854,618,644]" pageId="46" pageNumber="247">
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:
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, C. S.
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<location id="554E07D8FF8AB07DFC35C5EC97CE78F9" LSID="urn:lsid:plazi:treatment:D838E015FF8FB07DFF5DC3F597A57F61:554E07D8FF8AB07DFC35C5EC97CE78F9" box="[1023,1174,655,681]" country="Mexico" latitude="17.518888" longLatPrecision="1" longitude="-99.545555" name="Chilpancingo" pageId="46" pageNumber="247" stateProvince="Guerrero">Chilpancingo</location>
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S,
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E,
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<materialsCitation id="E0F95B5EFF8AB07DFB79C5D197F478A1" collectingDate="1987-11-01" collectionCode="KUNHM-ENT" collectorName="Arteaga El Descansadero" country="Mexico" elevation="300" latitude="18.356112" location="Michoacan" longLatPrecision="21" longitude="-102.29806" pageId="46" pageNumber="247" specimenCount="1" specimenCount-male="1" stateProvince="Michoacan de Ocampo">
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80kmN
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],
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<materialsCitation id="E0F95B5EFF8AB07DFB7FC5B497457945" collectingDate="1996-06-06" collectionCode="KUNHM-ENT" collectorName="R. Brooks" country="Mexico" elevation="990" latitude="18.464445" location="Ajuchitlan" longLatPrecision="21" longitude="-98.99166" pageId="46" pageNumber="247" specimenCount="7" specimenCount-male="7" stateProvince="Morelos">
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<location id="554E07D8FF8AB07DFAD5C5B496C078A1" LSID="urn:lsid:plazi:treatment:D838E015FF8FB07DFF5DC3F597A57F61:554E07D8FF8AB07DFAD5C5B496C078A1" box="[1311,1432,727,753]" country="Mexico" latitude="18.464445" longLatPrecision="21" longitude="-98.99166" name="Ajuchitlan" pageId="46" pageNumber="247" stateProvince="Morelos">Ajuchitlán</location>
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<collectorName id="FD6434D5FF8AB07DFBE1C59893BA7969" pageId="46" pageNumber="247">Sierra de Huatla C.E.A.M.I.S.H. Huatla</collectorName>
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N 4.0km W, [
<quantity id="9769FCE6FF8AB07DFE12C47D916C7969" box="[472,564,798,825]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="46" pageNumber="247" unit="m" value="1050.0">
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, R.
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(
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<collectingMunicipality id="B04ACB79FF8AB07DFE26C40490FF79D1" box="[492,935,871,897]" pageId="46" pageNumber="247">Huimilpan Parque Nacional Cimatario</collectingMunicipality>
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.
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<collectingRegion id="92559FE1FF8AB07DFD98C4E8919279F5" box="[594,714,907,933]" country="Mexico" name="Zacatecas" pageId="46" pageNumber="247">Zacatecas</collectingRegion>
,
</emphasis>
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,
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<elevation id="DBBCB630FF8AB07DFC67C4E8975E79F5" box="[941,1030,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.483" pageId="46" pageNumber="247" unit="m" value="1483.0">1483 m</elevation>
</quantity>
,
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,
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<collectingDate id="346B8E2BFF8AB07DFAB2C4E893AF7999" pageId="46" pageNumber="247" value="2012-09-21">21/ IX/2012</collectingDate>
</date>
, A.
<collectorName id="FD6434D5FF8AB07DFEE9C4CC92247999" box="[291,380,943,969]" pageId="46" pageNumber="247">Salcedo</collectorName>
(
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<specimenCount id="46979A8AFF8AB07DFE2BC4CC915D7999" box="[481,517,943,969]" count="1" pageId="46" pageNumber="247" type="male">1♂</specimenCount>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8AB07DFF0DC4B197797E65" pageId="46" pageNumber="247" type="distribution">
<paragraph id="502E5103FF8AB07DFF0DC4B197797E65" blockId="46.[151,1437,151,1329]" pageId="46" pageNumber="247">
<emphasis id="62E58D11FF8AB07DFF0DC4B1923979BC" bold="true" box="[199,353,978,1004]" pageId="46" pageNumber="247">Distribution.</emphasis>
This species is present mostly in the center of
<collectingCountry id="28861193FF8AB07DFCA2C4B0909879BD" box="[872,960,979,1005]" name="Mexico" pageId="46" pageNumber="247">Mexico</collectingCountry>
, in areas with tropical dry forest and in the mountains in ecotone with pine-oak forest. There are records from the Sierra Madre Occidental, from
<collectingRegion id="92559FE1FF8AB07DFACDC49496D87E41" box="[1287,1408,1015,1041]" country="Mexico" name="Chihuahua" pageId="46" pageNumber="247">Chihuahua</collectingRegion>
to
<collectingRegion id="92559FE1FF8AB07DFF5DC37893A77E65" box="[151,255,1051,1077]" country="Mexico" name="Guerrero" pageId="46" pageNumber="247">Guerrero</collectingRegion>
, and it is endemic to
<collectingCountry id="28861193FF8AB07DFE27C378911B7E65" box="[493,579,1051,1077]" name="Mexico" pageId="46" pageNumber="247">Mexico</collectingCountry>
(
<figureCitation id="C8AA4D86FF8AB07DFD9BC37891FE7E65" box="[593,678,1051,1077]" captionStart="FIGURE 26" captionStartId="43.[151,250,1068,1092]" captionTargetBox="[159,1428,157,1034]" captionTargetId="figure-18@43.[151,1436,149,1042]" captionTargetPageId="43" captionText="FIGURE 26. Known distribution records for Heriades nora sp. nov." pageId="46" pageNumber="247">Fig. 26</figureCitation>
). Altitudinal range:
<quantity id="9769FCE6FF8AB07DFC4FC37897457E65" box="[901,1053,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.1605" metricValueMax="2.23" metricValueMin="0.091" pageId="46" pageNumber="247" unit="m" value="1160.5" valueMax="2230.0" valueMin="91.0">91 to 2230 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8AB07DFF0DC35C96167EF1" pageId="46" pageNumber="247" type="etymology">
<paragraph id="502E5103FF8AB07DFF0DC35C96167EF1" blockId="46.[151,1437,151,1329]" pageId="46" pageNumber="247">
<emphasis id="62E58D11FF8AB07DFF0DC35C92137E09" bold="true" box="[199,331,1087,1113]" pageId="46" pageNumber="247">Etymology.</emphasis>
This species is dedicated to the memory of Nora Patricia López, an environment biologist who worked in the protection of red Guacamayas (
<taxonomicName id="97912A80FF8AB07DFDE4C30091FE7E2C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[558,678,1123,1149]" class="Aves" family="Psittacidae" genus="Ara" kingdom="Animalia" order="Psittaciformes" pageId="46" pageNumber="247" phylum="Chordata" rank="species" species="macao">
<emphasis id="62E58D11FF8AB07DFDE4C30091FE7E2C" box="[558,678,1123,1149]" italics="true" pageId="46" pageNumber="247">Ara macao</emphasis>
</taxonomicName>
), a species with high extinction risk in
<collectingRegion id="92559FE1FF8AB07DFB86C30097FF7E2D" box="[1100,1191,1123,1149]" country="Mexico" name="Chiapas" pageId="46" pageNumber="247">Chiapas</collectingRegion>
,
<collectingCountry id="28861193FF8AB07DFB7BC30096507E2D" box="[1201,1288,1123,1149]" name="Mexico" pageId="46" pageNumber="247">Mexico</collectingCountry>
. She is one of the many environmentalists murdered for their activism on behalf of biodiversity protection in Latin America.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF8AB07DFF0DC3C997A57F61" pageId="46" pageNumber="247" type="discussion">
<paragraph id="502E5103FF8AB07DFF0DC3C997A57F61" blockId="46.[151,1437,151,1329]" pageId="46" pageNumber="247">
<emphasis id="62E58D11FF8AB07DFF0DC3C992177E94" bold="true" box="[199,335,1194,1220]" pageId="46" pageNumber="247">Comments.</emphasis>
This species is abundant in collections. Females present variation in the size of tubercles on apical margin of clypeus. Females punctures on T3 varies slightly on size, but always large, also varies on shape from rounded to the more usual horseshoe-shaped. Males present variation in hairs of clypeus, on some long enough to cover low margin, and on others shorter, showing a few smaller hairs in the middle of apical margin.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,513 @@
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<mods:title id="9979051A32AA59E6C71EA96B21BF9D1F">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<paragraph id="502E5103FF90B067FF5DC3AD90227EB9" blockId="52.[151,890,1230,1293]" box="[151,890,1230,1257]" pageId="52" pageNumber="253">
<heading id="0B66E66FFF90B067FF5DC3AD90227EB9" bold="true" box="[151,890,1230,1257]" fontSize="11" level="1" pageId="52" pageNumber="253" reason="1">
<emphasis id="62E58D11FF90B067FF5DC3AD90227EB9" bold="true" box="[151,890,1230,1257]" pageId="52" pageNumber="253">
<taxonomicName id="97912A80FF90B067FF5DC3AD904D7EB8" authority="Rojas-Arias, Griswold &amp; Ayala." authorityName="Rojas-Arias, Griswold &amp; Ayala." authorityYear="2025" box="[151,789,1230,1257]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="vulcanica" status="sp. nov.">
<emphasis id="62E58D11FF90B067FF5DC3AD92D97EB9" bold="true" box="[151,385,1230,1257]" italics="true" pageId="52" pageNumber="253">Heriades vulcanica</emphasis>
Rojas-Arias, Griswold &amp; Ayala.
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<subSubSection id="188B0288FF90B067FF5DC39092067F5D" box="[151,350,1267,1293]" pageId="52" pageNumber="253" type="description">
<paragraph id="502E5103FF90B067FF5DC39092067F5D" blockId="52.[151,890,1230,1293]" box="[151,350,1267,1293]" pageId="52" pageNumber="253">
(
<figureCitation id="C8AA4D86FF90B067FF55C390925B7F5D" box="[159,259,1267,1293]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="52" pageNumber="253">Figs. 2K</figureCitation>
,
<figureCitation id="C8AA4D86FF90B067FEC5C390920E7F5D" box="[271,342,1267,1293]" captionStart-0="FIGURE 32" captionStart-1="FIGURE 33" captionStartId-0="54.[151,250,1108,1132]" captionStartId-1="55.[151,250,1943,1967]" captionTargetBox-0="[159,1428,187,1083]" captionTargetBox-1="[155,1419,189,1905]" captionTargetId-0="figure-22@54.[151,1436,179,1083]" captionTargetId-1="figure-18@55.[155,1432,179,1917]" captionTargetPageId-0="54" captionTargetPageId-1="55" captionText-0="FIGURE 32. Known distribution records for Heriades vulcanica sp. nov." captionText-1="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="52" pageNumber="253">3233</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF90B067FF5DC25997807CC9" pageId="52" pageNumber="253" type="diagnosis">
<paragraph id="502E5103FF90B067FF5DC25997807CC9" blockId="52.[151,1437,1338,2013]" pageId="52" pageNumber="253">
<emphasis id="62E58D11FF90B067FF5DC259924A7F04" bold="true" box="[151,274,1338,1364]" pageId="52" pageNumber="253">Diagnosis.</emphasis>
Large-sized bees (males 7.27.7, n=3; female
<quantity id="9769FCE6FF90B067FCD4C25890217F05" box="[798,889,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="52" pageNumber="253" unit="mm" value="6.8">6.8 mm</quantity>
); head and mesosoma with plumose, long, think, scattered, white, spread hairs; posterior terga with hairs yellow to orangish; tergal fasciae with rows of thin hairs; T3 with small punctures (0.1 times OD) (
<figureCitation id="C8AA4D86FF90B067FDA1C2E091AD7FCD" box="[619,757,1411,1437]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="52" pageNumber="253">Fig. 33B, D</figureCitation>
).
<emphasis id="62E58D11FF90B067FCC0C2E0901C7FCD" box="[778,836,1411,1437]" italics="true" pageId="52" pageNumber="253">Male</emphasis>
: head slightly subtriangular, face with plumose, thin, scattered, white to yellowish hairs, antenna black with thick scape (
<figureCitation id="C8AA4D86FF90B067FCB4C2C490BE7F91" box="[894,998,1447,1473]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="52" pageNumber="253">Fig. 33A</figureCitation>
); S1 produced to median rounded angle (
<figureCitation id="C8AA4D86FF90B067FF55C2A893B97FB5" box="[159,225,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="52" pageNumber="253">Fig. 2</figureCitation>
Kb), in profile, with soft ventral angle one third of distance from apex to base (
<figureCitation id="C8AA4D86FF90B067FB86C2A897D67FB5" box="[1100,1166,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="52" pageNumber="253">Fig. 2</figureCitation>
Ka); posterior margin of S5 with two subtriangular lobes, and two rounded tufts of capitate hairs (
<figureCitation id="C8AA4D86FF90B067FC05C28C976D7C59" box="[975,1077,1519,1545]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="52" pageNumber="253">Fig. 33E</figureCitation>
); S6 with two subapical tufts of several, short, thin hairs directed downward and medially (
<figureCitation id="C8AA4D86FF90B067FCE6C17090C87C7D" box="[812,912,1555,1581]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="52" pageNumber="253">Fig. 33F</figureCitation>
), similar to that on
<taxonomicName id="97912A80FF90B067FBA1C17097AC7C7D" authorityName="Cockerell" authorityYear="1897" box="[1131,1268,1555,1581]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="62E58D11FF90B067FBA1C17097AC7C7D" box="[1131,1268,1555,1581]" italics="true" pageId="52" pageNumber="253">H. crucifera</emphasis>
</taxonomicName>
,
<taxonomicName id="97912A80FF90B067FAC8C170962B7C7D" authorityName="Crawford" authorityYear="1913" box="[1282,1395,1555,1581]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="leavitti">
<emphasis id="62E58D11FF90B067FAC8C170962B7C7D" box="[1282,1395,1555,1581]" italics="true" pageId="52" pageNumber="253">H. leavitti</emphasis>
</taxonomicName>
,
<taxonomicName id="97912A80FF90B067FA4BC17093BA7C00" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="62E58D11FF90B067FA4BC17093BA7C00" italics="true" pageId="52" pageNumber="253">H. texana</emphasis>
</taxonomicName>
and
<taxonomicName id="97912A80FF90B067FED0C15492CC7C00" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[282,404,1591,1616]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="cristina" status="sp. nov.">
<emphasis id="62E58D11FF90B067FED0C15492CC7C00" box="[282,404,1591,1616]" italics="true" pageId="52" pageNumber="253">H. cristina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF90B067FE51C15492AB7C01" box="[411,499,1591,1617]" pageId="52" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
but different in the other characters mentioned.
<emphasis id="62E58D11FF90B067FBDCC15497327C01" box="[1046,1130,1591,1617]" italics="true" pageId="52" pageNumber="253">Female</emphasis>
: outer surface of mandible with very strong, angulated tubercle forming a subtriangular shape on ventral margin of mandible (
<figureCitation id="C8AA4D86FF90B067FB38C138960F7C25" box="[1266,1367,1627,1653]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="52" pageNumber="253">Fig. 33C</figureCitation>
) (
<figureCitation id="C8AA4D86FF90B067FAA5C13893917CC9" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="52" pageNumber="253">Fig. 33C</figureCitation>
), similar to
<taxonomicName id="97912A80FF90B067FE85C11C92E47CC8" box="[335,444,1663,1688]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="62E58D11FF90B067FE85C11C92E47CC8" box="[335,444,1663,1688]" italics="true" pageId="52" pageNumber="253">H. texana</emphasis>
</taxonomicName>
but different in color of hairs on T3 and malar area without projection.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF90B065FF0DC1C1976D7EAD" lastPageId="54" lastPageNumber="255" pageId="52" pageNumber="253" type="description">
<paragraph id="502E5103FF90B067FF0DC1C1964C7CED" blockId="52.[151,1437,1338,2013]" box="[199,1300,1698,1725]" pageId="52" pageNumber="253">
<emphasis id="62E58D11FF90B067FF0DC1C192FD7CED" bold="true" box="[199,421,1698,1725]" pageId="52" pageNumber="253">Description. Male:</emphasis>
Body length,
<quantity id="9769FCE6FF90B067FD8EC1C091C27CED" box="[580,666,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="52" pageNumber="253" unit="mm" value="7.2">7.2 mm</quantity>
(7.27.7, n=3); forewing length,
<quantity id="9769FCE6FF90B067FBDAC1C1973E7CED" box="[1040,1126,1698,1725]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="52" pageNumber="253" unit="mm" value="5.6">5.6 mm</quantity>
(5.15.6, n=3).
</paragraph>
<paragraph id="502E5103FF90B066FF0DC1A4904A7B35" blockId="52.[151,1437,1338,2013]" lastBlockId="53.[151,1437,150,2013]" lastPageId="53" lastPageNumber="254" pageId="52" pageNumber="253">
<emphasis id="62E58D11FF90B067FF0DC1A4925C7CB1" box="[199,260,1735,1761]" italics="true" pageId="52" pageNumber="253">Head</emphasis>
. Length
<quantity id="9769FCE6FF90B067FEADC1A492E47CB1" box="[359,444,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="52" pageNumber="253" unit="mm" value="1.7">1.7 mm</quantity>
(1.21.7, n=3), width
<quantity id="9769FCE6FF90B067FD7FC1A490537CB1" box="[693,779,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="52" pageNumber="253" unit="mm" value="1.9">1.9 mm</quantity>
(1.92.1, n=3); eyes about 3.3 times longer than broad, 1.4 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF90B067FCE7C18890DA7D55" box="[813,898,1771,1797]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="52" pageNumber="253" unit="mm" value="0.9">0.9 mm</quantity>
(0.91, n=3); upper interocular distance
<quantity id="9769FCE6FF90B067FA8DC18896C57D55" box="[1351,1437,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="52" pageNumber="253" unit="mm" value="1.2">1.2 mm</quantity>
(1.21.3, n=3); interantennal distance 1.9 times antennocular distance; lateral ocelli large (
<figureCitation id="C8AA4D86FF90B067FB4CC06C97847D79" box="[1158,1244,1807,1833]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="52" pageNumber="253">Fig. 3C</figureCitation>
), 3 times DOP in interocellar area; interocellar distance 1.9 times OD, ocellocular distance 1.9 times OD; OPR 2.1 times OD; labrum with basal, medial, elevation; clypeus 1.8 times broader than long, lower third convex in profile, apical margin without emargination; juxtantennal carina absent but strongly angulated; scape black, length 0.8 times alveolocellar distance, 2.7 times longer than wide; pedicel as long as broad, black, as long as F1 and about 0.4 times longer than F2; flagellum black; F1 broader than long, remaining flagellomeres 2.3 longer than first; F11 dark brown 2.4 times longer than broad (
<figureCitation id="C8AA4D86FF91B066FEA0C7F4928A7AE1" box="[362,466,151,177]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="53" pageNumber="254">Fig. 33A</figureCitation>
).
<emphasis id="62E58D11FF91B066FE2FC7F4913C7AE0" box="[485,612,151,176]" italics="true" pageId="53" pageNumber="254">Mesosoma.</emphasis>
Scutum 2.5 times longer than scutellum, 2.8 times longer than tegula, and 3.4 longer than axilla; intertegular distance
<quantity id="9769FCE6FF91B066FD4DC7D891867A85" box="[647,734,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="53" pageNumber="254" unit="mm" value="1.4">1.4 mm</quantity>
(1.41.6, n=3); metanotum with transverse ridge, in posterior view straight. Propodeum with large pits,
<quantity id="9769FCE6FF91B066FDA1C7BC91987AA9" box="[619,704,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="254" unit="mm" value="0.1">0.1 mm</quantity>
long; legs black.
<emphasis id="62E58D11FF91B066FC4FC7BC97597AA8" box="[901,1025,223,248]" italics="true" pageId="53" pageNumber="254">Metasoma.</emphasis>
T1
<quantity id="9769FCE6FF91B066FBE7C7BC97DB7AA9" box="[1069,1155,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="53" pageNumber="254" unit="mm" value="1.7">1.7 mm</quantity>
broad (1.52, n=3); T3
<quantity id="9769FCE6FF91B066FA44C7BC939B7B4D" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="53" pageNumber="254" unit="mm" value="2.0">2 mm</quantity>
broad (1.82, n=3); T6 with posterior margin projected (
<figureCitation id="C8AA4D86FF91B066FC8AC66090FF7B4D" box="[832,935,259,285]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="53" pageNumber="254">Fig. 33B</figureCitation>
); posterior margin of S1 produced to median rounded angle, without emargination (
<figureCitation id="C8AA4D86FF91B066FD84C64491CB7B11" box="[590,659,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="53" pageNumber="254">Fig. 2</figureCitation>
Kb), in profile, with posterior margin not projected and soft ventral angle one third of distance from apex to base (
<figureCitation id="C8AA4D86FF91B066FD54C62891B97B35" box="[670,737,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="53" pageNumber="254">Fig. 2</figureCitation>
Ka).
</paragraph>
<paragraph id="502E5103FF91B066FF0DC60C97BF7969" blockId="53.[151,1437,150,2013]" pageId="53" pageNumber="254">
<emphasis id="62E58D11FF91B066FF0DC60C92087BD9" box="[199,336,367,393]" italics="true" pageId="53" pageNumber="254">Sculpturing.</emphasis>
Clypeus with small (at base 0.2 times OD), contiguous, coarse punctures; supraclypeal area with punctures bigger, more rounded and slightly sparser than on base of clypeus; frons with smaller, coarser and denser punctures than those on supraclypeal area; vertex with bigger, slightly sparser punctures than on supraclypeal area, with micropunctures between punctures; hypostomal area with punctation as on clypeus; pronotum anterior surface with punctures bigger and sparser than on clypeus, denser on pronotal lobes; disc of scutum with slightly smaller (0.3 times OD), punctures than on vertex, with micropunctures between punctures; rest of mesosoma as on scutum except: scutellum with punctures sparser; mesepisternum with equal DOP as on scutum but sparser on its entire surface; metepisternum with notably smaller punctures, less dense and coarse; T1 and T2 with smaller (0.2 times OD), slightly denser punctures than on scutum; T2 with punctures smaller and sparser than on T1; T3 with smaller punctures than on T1, rounded shape, separated by less than half DOP; T4 with bigger, denser and coarser punctures than on T3; T5 with smaller and denser punctures than on T4; dorsal surface of T6 with punctures as on T5, on posterior surface with notably smaller and slightly sparser punctation, DBP one half DOP; sides of S1 with punctation as on T1, and in process smaller; sides of S2 with punctures as on T1, disc of S2 microreticulate without punctation.
</paragraph>
<paragraph id="502E5103FF91B066FF0DC42092E47E65" blockId="53.[151,1437,150,2013]" pageId="53" pageNumber="254">
<emphasis id="62E58D11FF91B066FF0DC4209268790C" box="[199,304,835,860]" italics="true" pageId="53" pageNumber="254">Vestiture.</emphasis>
Supraclypeal area with plumose, thin, white, long hairs directed medially; frons with plumose, thin, orangish, long hairs directed dorsally (
<figureCitation id="C8AA4D86FF91B066FDF5C40491FE79D1" box="[575,678,871,897]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="53" pageNumber="254">Fig. 33A</figureCitation>
); dorsolateral angles with pubescence not forming continuous, linear tuft with hairs on anterior margin of scutum, and without tuft of thick, short hairs; disc of scutum and mesepisternum with plumose, thin, long, white, scattered hairs, evenly distributed; disc of metepisternum without tuft of thick, short, white hairs; tergal fasciae with rows of thin, long, orange hairs; T1 to T3 with simple, short, thin, orangish hairs in punctures; disc of T4 to T6 with simple, long, thick, orange hairs in punctures increasing in length towards posterior terga (
<figureCitation id="C8AA4D86FF91B066FE82C37892F77E65" box="[328,431,1051,1077]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="53" pageNumber="254">Fig. 33B</figureCitation>
).
</paragraph>
<paragraph id="502E5103FF91B066FF0DC35C91B27E95" blockId="53.[151,1437,150,2013]" pageId="53" pageNumber="254">
<emphasis id="62E58D11FF91B066FF0DC35C92117E09" box="[199,329,1087,1113]" italics="true" pageId="53" pageNumber="254">
<taxonomicName id="97912A80FF91B066FF0DC35C921D7E09" box="[199,325,1087,1113]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="53" pageNumber="254" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
S6 with right-angled apex, with two subapical tufts of several, short, thin hairs directed downward and medially (
<figureCitation id="C8AA4D86FF91B066FEF7C30092FC7E2D" box="[317,420,1123,1149]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="53" pageNumber="254">Fig. 33F</figureCitation>
); capsule in dorsal view
<quantity id="9769FCE6FF91B066FD0FC30090457E2D" box="[709,797,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="53" pageNumber="254" unit="mm" value="1.3">1.3 mm</quantity>
length, and
<quantity id="9769FCE6FF91B066FC65C30097507E2D" box="[943,1032,1123,1149]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="53" pageNumber="254" unit="mm" value="0.6">0.6 mm</quantity>
width, 2.3 times longer than wide, gonobase
<quantity id="9769FCE6FF91B066FECDC3E492327EF1" box="[263,362,1159,1185]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="53" pageNumber="254" unit="mm" value="0.09">0.09 mm</quantity>
length,5.8 times wider than long, and gonocoxite
<quantity id="9769FCE6FF91B066FC5CC3E590B37EF1" box="[918,1003,1158,1185]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="53" pageNumber="254" unit="mm" value="0.5">0.5 mm</quantity>
length, gonostylus
<quantity id="9769FCE6FF91B066FB09C3E496417EF1" box="[1219,1305,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="53" pageNumber="254" unit="mm" value="1.2">1.2 mm</quantity>
length with rounded apex, penis valve
<quantity id="9769FCE6FF91B066FE09C3C891417E95" box="[451,537,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="53" pageNumber="254" unit="mm" value="0.8">0.8 mm</quantity>
length (
<figureCitation id="C8AA4D86FF91B066FDBEC3C891867E95" box="[628,734,1195,1221]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="53" pageNumber="254">Fig. 33G</figureCitation>
).
</paragraph>
<paragraph id="502E5103FF91B066FF0DC3AD91697C7D" blockId="53.[151,1437,150,2013]" pageId="53" pageNumber="254">
<emphasis id="62E58D11FF91B066FF0DC3AD927F7EB9" bold="true" box="[199,295,1230,1257]" pageId="53" pageNumber="254">
<emphasis id="62E58D11FF91B066FF0DC3AD92467EB9" bold="true" box="[199,286,1230,1257]" italics="true" pageId="53" pageNumber="254">Female</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FF91B066FE0BC3AC91437EB9" box="[449,539,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="53" pageNumber="254" unit="mm" value="6.8">6.8 mm</quantity>
; forewing length
<quantity id="9769FCE6FF91B066FD28C3AC90637EB9" box="[738,827,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="53" pageNumber="254" unit="mm" value="4.9">4.9 mm</quantity>
. Similar to male except:
<emphasis id="62E58D11FF91B066FB9FC3AC97CA7EB9" box="[1109,1170,1231,1257]" italics="true" pageId="53" pageNumber="254">Head</emphasis>
. Length
<quantity id="9769FCE6FF91B066FB3DC3AC96087EB9" box="[1271,1360,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="53" pageNumber="254" unit="mm" value="1.9">1.9 mm</quantity>
, width
<quantity id="9769FCE6FF91B066FF5DC39093AA7F5D" box="[151,242,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="53" pageNumber="254" unit="mm" value="2.1">2.1 mm</quantity>
; eyes about 3.5 times longer than broad, 1.1 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF91B066FF5DC27493AB7F61" box="[151,243,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="53" pageNumber="254" unit="mm" value="1.2">1.2 mm</quantity>
; upper interocular distance
<quantity id="9769FCE6FF91B066FDFEC27491C87F61" box="[564,656,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="53" pageNumber="254" unit="mm" value="1.3">1.3 mm</quantity>
; interantennal distance 1.1 times antennocular distance; interocellar distance 1.6 times OD, ocellocular distance 1.9 times OD, OPR 2.5 times OD; labrum with rounded, basal, medial tubercle; mandible on outer surface with outer and condylar ridge parallel, with strong, angulated tubercle forming a subtriangular shape on ventral margin of mandible; dorsal margin of mandible with carina; clypeus with medial small subtriangular emargination (
<figureCitation id="C8AA4D86FF91B066FDD1C2C491DA7F91" box="[539,642,1447,1473]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="53" pageNumber="254">Fig. 33C</figureCitation>
), in profile elevated on lower fifth; scape 3.1 times longer than broad; pedicel 1.3 longer than broad, about 1.6 times longer than F1 and 1.3 longer than F2, remaining flagellomeres 1.6 times longer than F1.
<emphasis id="62E58D11FF91B066FE43C28C91597C58" box="[393,513,1519,1544]" italics="true" pageId="53" pageNumber="254">Mesosoma</emphasis>
. Scutum 2.5 times longer than scutellum; intertegular distance
<quantity id="9769FCE6FF91B066FB75C28D964F7C59" box="[1215,1303,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="254" unit="mm" value="1.5">1.5 mm</quantity>
.
<emphasis id="62E58D11FF91B066FAEBC28C96CE7C58" box="[1313,1430,1519,1544]" italics="true" pageId="53" pageNumber="254">Metasoma</emphasis>
. T1
<quantity id="9769FCE6FF91B066FF77C171924B7C7D" box="[189,275,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="254" unit="mm" value="1.5">1.5 mm</quantity>
broad. T3
<quantity id="9769FCE6FF91B066FE47C17092BB7C7D" box="[397,483,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="53" pageNumber="254" unit="mm" value="1.8">1.8 mm</quantity>
broad.
</paragraph>
<paragraph id="502E5103FF91B066FF0DC15491B17D8D" blockId="53.[151,1437,150,2013]" pageId="53" pageNumber="254">
<emphasis id="62E58D11FF91B066FF0DC15492097C01" box="[199,337,1591,1617]" italics="true" pageId="53" pageNumber="254">Sculpturing.</emphasis>
Mandibles with carinae and tubercle smooth and shiny; clypeus with medium-sized (0.2 times OD), contiguous, coarse, punctures, shiny and smooth between punctures; supraclypeal area with punctures smaller than base of clypeus; frons with bigger, coarser, and denser punctation, with micropunctures between punctures than on supraclypeal area: frons in front of ocelli with slightly smaller punctures than those on frons; vertex slightly bigger, sparser, and coarser than on clypeus, with micropunctures between punctures; gena evenly punctate, bigger and denser than on frons in front of ocelli; scutum with slightly bigger and denser punctures than on vertex; rest of mesosoma as on scutum except: scutellum with sparser punctures; mesepisternum with slightly sparser punctures; metasomal terga bright; T1 with smaller (0.7 times OD), slightly denser punctures than on scutum, with rounded shape; T2 with smaller, sparser than on T1, rounded shaped punctures; T3 with punctures smaller, sparser (0.3 times DOP), notably less coarse, than on scutum, semicircular shaped, and sparser (1 time DOP); dorsal surface of T6 with smaller, denser, and more coarsely punctures than on T5; sterna with punctation smaller and sparser than on scutum, with integument microreticulate on posterior margin.
</paragraph>
<caption id="04EE018BFF92B065FF5DC33790DA7E3C" box="[151,898,1108,1133]" pageId="54" pageNumber="255" startId="54.[151,250,1108,1132]" targetBox="[159,1428,187,1083]" targetPageId="54" targetType="figure">
<paragraph id="502E5103FF92B065FF5DC33790DA7E3C" blockId="54.[151,898,1108,1133]" box="[151,898,1108,1133]" pageId="54" pageNumber="255">
<emphasis id="62E58D11FF92B065FF5DC33792787E3C" bold="true" box="[151,288,1108,1132]" pageId="54" pageNumber="255">FIGURE 32.</emphasis>
Known distribution records for
<taxonomicName id="97912A80FF92B065FDADC33690747E3D" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[615,812,1109,1133]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="255" phylum="Arthropoda" rank="species" species="vulcanica" status="sp. nov.">
<emphasis id="62E58D11FF92B065FDADC33690747E3D" box="[615,812,1109,1133]" italics="true" pageId="54" pageNumber="255">Heriades vulcanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF92B065FCF9C33690DA7E3C" box="[819,898,1109,1132]" pageId="54" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="502E5103FF92B065FF0DC3FF976D7EAD" blockId="54.[151,1437,1179,1961]" pageId="54" pageNumber="255">
<emphasis id="62E58D11FF92B065FF0DC3FF92687EE5" box="[199,304,1180,1205]" italics="true" pageId="54" pageNumber="255">Vestiture.</emphasis>
Clypeus, frons, paraocular area and gena with plumose, long, scattered, white hairs (
<figureCitation id="C8AA4D86FF92B065FB38C3F896027EE5" box="[1266,1370,1179,1205]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="54" pageNumber="255">Fig. 33C</figureCitation>
); disc of T1 to T6 with simple, scattered, thin, orangish hairs in punctures, increasing in length towards posterior terga; subapical margin of T6 with linear tuft of plumose, short, whitish hairs (
<figureCitation id="C8AA4D86FF92B065FC0AC38097707EAD" box="[960,1064,1251,1277]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="54" pageNumber="255">Fig. 33D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF92B065FF0DC26492687C35" pageId="54" pageNumber="255" type="materials_examined">
<paragraph id="502E5103FF92B065FF0DC26492FF7FA9" blockId="54.[151,1437,1179,1961]" pageId="54" pageNumber="255">
<emphasis id="62E58D11FF92B065FF0DC264922F7F71" bold="true" box="[199,375,1287,1313]" pageId="54" pageNumber="255">Type material:</emphasis>
<materialsCitation id="E0F95B5EFF92B065FEB7C264977E7F15" collectingDate="1952-08-01" collectionCode="BBSL" collectorName="No. LR &amp; E. E. Gilbert &amp; C. D. MacNeil" country="Mexico" elevation="2361" latitude="16.761667" location="Casas" longLatPrecision="21" longitude="-92.633606" pageId="54" pageNumber="255" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="62E58D11FF92B065FEB7C26490887F71" bold="true" box="[381,976,1286,1313]" pageId="54" pageNumber="255">
<typeStatus id="8F2AEFA1FF92B065FEB7C26492B27F71" box="[381,490,1287,1313]" pageId="54" pageNumber="255" type="holotype">Holotype</typeStatus>
male (
<collectorName id="FD6434D5FF92B065FDF3C26591CA7F70" box="[569,658,1286,1312]" pageId="54" pageNumber="255">No. LR</collectorName>
0007):
<collectingCountry id="28861193FF92B065FD2BC26490047F71" box="[737,860,1287,1313]" name="Mexico" pageId="54" pageNumber="255">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF92B065FCADC26490937F71" box="[871,971,1287,1313]" country="Mexico" name="Chiapas" pageId="54" pageNumber="255">Chiapas</collectingRegion>
,
</emphasis>
San Cristobal de las
<location id="554E07D8FF92B065FB71C26497A47F71" LSID="urn:lsid:plazi:treatment:D838E015FF90B065FF5DC3AD90EA7DF9:554E07D8FF92B065FB71C26497A47F71" box="[1211,1276,1287,1313]" country="Mexico" latitude="16.761667" longLatPrecision="21" longitude="-92.633606" name="Casas" pageId="54" pageNumber="255" stateProvince="Chiapas">Casas</location>
39miE, [
<quantity id="9769FCE6FF92B065FAA8C26493F77F15" metricMagnitude="3" metricUnit="m" metricValue="2.3609999999999998" pageId="54" pageNumber="255" unit="m" value="2361.0">
<elevation id="DBBCB630FF92B065FAA8C26493F77F15" metricMagnitude="3" metricUnit="m" metricValue="2.3609999999999998" pageId="54" pageNumber="255" unit="m" value="2361.0">2361 m</elevation>
</quantity>
,
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<geoCoordinate id="35A537C4FF92B065FE81C24892867F15" box="[331,478,1323,1349]" degrees="92" direction="west" minutes="38" orientation="longitude" pageId="54" pageNumber="255" precision="15" seconds="01" value="-92.633606">92°3801”W</geoCoordinate>
],
<date id="242F77C3FF92B065FE3BC24891227F16" box="[497,634,1323,1350]" pageId="54" pageNumber="255" value="1952-08-01">
<collectingDate id="346B8E2BFF92B065FE3BC24891227F16" box="[497,634,1323,1350]" pageId="54" pageNumber="255" value="1952-08-01">1/VIII/1952</collectingDate>
</date>
,
<collectorName id="FD6434D5FF92B065FD4FC248904C7F15" box="[645,788,1323,1349]" pageId="54" pageNumber="255">E. E. Gilbert</collectorName>
,
<collectorName id="FD6434D5FF92B065FCD5C248909C7F15" box="[799,964,1323,1349]" pageId="54" pageNumber="255">C. D. MacNeil</collectorName>
(
<collectionCode id="3680C9C6FF92B065FC18C24897467F15" box="[978,1054,1323,1349]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="54" pageNumber="255" type="Museum">BBSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF92B065FBE5C24896C17F39" collectingDate="1960-04-11" collectionCode="USNM" collectorName="J. F. G. Clarke" country="Mexico" elevation="6" latitude="19.809444" location="Allen Pt Ascension Bay" longLatPrecision="21" longitude="-87.4775" pageId="54" pageNumber="255" specimenCount="1" stateProvince="Quintana Roo" typeStatus="allotype">
<emphasis id="62E58D11FF92B065FBE5C24892127F39" bold="true" pageId="54" pageNumber="255">
<typeStatus id="8F2AEFA1FF92B065FBE5C24897CC7F15" box="[1071,1172,1323,1349]" pageId="54" pageNumber="255" type="allotype">Allotype</typeStatus>
(LR0008):
<collectingCountry id="28861193FF92B065FAD7C24896C07F15" box="[1309,1432,1323,1349]" name="Mexico" pageId="54" pageNumber="255">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF92B065FF5DC22C921D7F39" box="[151,325,1359,1385]" country="Mexico" name="Quintana Roo" pageId="54" pageNumber="255">Quintana Roo</collectingRegion>
,
</emphasis>
<location id="554E07D8FF92B065FE9FC22C91337F39" LSID="urn:lsid:plazi:treatment:D838E015FF90B065FF5DC3AD90EA7DF9:554E07D8FF92B065FE9FC22C91337F39" box="[341,619,1359,1385]" country="Mexico" latitude="19.809444" longLatPrecision="21" longitude="-87.4775" name="Allen Pt Ascension Bay" pageId="54" pageNumber="255" stateProvince="Quintana Roo">Allen Pt Ascension Bay</location>
,
<quantity id="9769FCE6FF92B065FDB3C22C91F27F39" box="[633,682,1359,1385]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="54" pageNumber="255" unit="m" value="6.0">
<elevation id="DBBCB630FF92B065FDB3C22C91F27F39" box="[633,682,1359,1385]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="54" pageNumber="255" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF92B065FD73C22C901A7F39" box="[697,834,1359,1385]" degrees="19" direction="north" minutes="48" orientation="latitude" pageId="54" pageNumber="255" precision="15" seconds="34" value="19.809444">19°4834”N</geoCoordinate>
<geoCoordinate id="35A537C4FF92B065FC86C22C90847F39" box="[844,988,1359,1385]" degrees="87" direction="west" minutes="28" orientation="longitude" pageId="54" pageNumber="255" precision="15" seconds="39" value="-87.4775">87°2839”W</geoCoordinate>
,
<date id="242F77C3FF92B065FC21C22C97367F39" box="[1003,1134,1359,1385]" pageId="54" pageNumber="255" value="1960-04-11">
<collectingDate id="346B8E2BFF92B065FC21C22C97367F39" box="[1003,1134,1359,1385]" pageId="54" pageNumber="255" value="1960-04-11">11/IV/1960</collectingDate>
</date>
,
<collectorName id="FD6434D5FF92B065FBB6C22C96707F39" box="[1148,1320,1359,1385]" pageId="54" pageNumber="255">J. F. G. Clarke</collectorName>
(
<collectionCode id="3680C9C6FF92B065FAF0C22C96C97F39" box="[1338,1425,1359,1385]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="54" pageNumber="255" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF92B065FF5DC210939E7FE1" collectingDate="2012-07-02" collectionCode="ECOAB" collectorName="P. Sagot" country="Mexico" elevation="2087" location="Amatenango del Valle" pageId="54" pageNumber="255" specimenCount="2" specimenCount-male="2" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="62E58D11FF92B065FF5DC21091497FDD" bold="true" box="[151,529,1395,1421]" pageId="54" pageNumber="255">
<typeStatus id="8F2AEFA1FF92B065FF5DC210924B7FDD" box="[151,275,1395,1421]" pageId="54" pageNumber="255" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="28861193FF92B065FEEBC21092C47FDD" box="[289,412,1395,1421]" name="Mexico" pageId="54" pageNumber="255">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF92B065FE62C21091547FDD" box="[424,524,1395,1421]" country="Mexico" name="Chiapas" pageId="54" pageNumber="255">Chiapas</collectingRegion>
,
</emphasis>
<location id="554E07D8FF92B065FDD0C210904F7FDD" LSID="urn:lsid:plazi:treatment:D838E015FF90B065FF5DC3AD90EA7DF9:554E07D8FF92B065FDD0C210904F7FDD" box="[538,791,1395,1421]" country="Mexico" name="Amatenango del Valle" pageId="54" pageNumber="255" stateProvince="Chiapas">Amatenango del Valle</location>
,
<quantity id="9769FCE6FF92B065FCEEC21090247FDD" box="[804,892,1395,1421]" metricMagnitude="3" metricUnit="m" metricValue="2.0869999999999997" pageId="54" pageNumber="255" unit="m" value="2087.0">
<elevation id="DBBCB630FF92B065FCEEC21090247FDD" box="[804,892,1395,1421]" metricMagnitude="3" metricUnit="m" metricValue="2.0869999999999997" pageId="54" pageNumber="255" unit="m" value="2087.0">2087 m</elevation>
</quantity>
, 16.5067 -92.38,
<date id="242F77C3FF92B065FB8EC210979B7FDD" box="[1092,1219,1395,1421]" pageId="54" pageNumber="255" value="2012-07-02">
<collectingDate id="346B8E2BFF92B065FB8EC210979B7FDD" box="[1092,1219,1395,1421]" pageId="54" pageNumber="255" value="2012-07-02">2/VII/2012</collectingDate>
</date>
,
<collectorName id="FD6434D5FF92B065FB1AC21096747FDD" box="[1232,1324,1395,1421]" pageId="54" pageNumber="255">P. Sagot</collectorName>
(
<collectionCode id="3680C9C6FF92B065FAF1C21096C47FDD" box="[1339,1436,1395,1421]" pageId="54" pageNumber="255">ECOAB</collectionCode>
<specimenCount id="46979A8AFF92B065FF5DC2F493E57FE1" box="[151,189,1431,1457]" count="2" pageId="54" pageNumber="255" type="male">2♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF92B065FF1FC2F492DC7F85" collectingDate="1952-08-01" collectionCode="BBSL" collectorName="E. E. Gilbert &amp; C. D. MacNeil" country="Saint Kitts and Nevis" elevation="2361" latitude="16.761667" location="Casas" longLatPrecision="21" longitude="-92.633606" pageId="54" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
<collectingCountry id="28861193FF92B065FF1FC2F492287FE1" box="[213,368,1431,1457]" name="Saint Kitts and Nevis" pageId="54" pageNumber="255">San Cristobal</collectingCountry>
de las
<location id="554E07D8FF92B065FE02C2F491517FE1" LSID="urn:lsid:plazi:treatment:D838E015FF90B065FF5DC3AD90EA7DF9:554E07D8FF92B065FE02C2F491517FE1" box="[456,521,1431,1457]" country="Saint Kitts and Nevis" latitude="16.761667" longLatPrecision="21" longitude="-92.633606" name="Casas" pageId="54" pageNumber="255" stateProvince="Chiapas">Casas</location>
39miE, [
<quantity id="9769FCE6FF92B065FDBDC2F4918C7FE1" box="[631,724,1431,1457]" metricMagnitude="3" metricUnit="m" metricValue="2.3609999999999998" pageId="54" pageNumber="255" unit="m" value="2361.0">
<elevation id="DBBCB630FF92B065FDBDC2F4918C7FE1" box="[631,724,1431,1457]" metricMagnitude="3" metricUnit="m" metricValue="2.3609999999999998" pageId="54" pageNumber="255" unit="m" value="2361.0">2361 m</elevation>
</quantity>
,
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<geoCoordinate id="35A537C4FF92B065FCBCC2F497527FE1" box="[886,1034,1431,1457]" degrees="92" direction="west" minutes="38" orientation="longitude" pageId="54" pageNumber="255" precision="15" seconds="01" value="-92.633606">92°3801”W</geoCoordinate>
],
<date id="242F77C3FF92B065FBEAC2F497F17FE2" box="[1056,1193,1431,1458]" pageId="54" pageNumber="255" value="1952-08-01">
<collectingDate id="346B8E2BFF92B065FBEAC2F497F17FE2" box="[1056,1193,1431,1458]" pageId="54" pageNumber="255" value="1952-08-01">1/VIII/1952</collectingDate>
</date>
,
<collectorName id="FD6434D5FF92B065FB7DC2F496167FE1" box="[1207,1358,1431,1457]" pageId="54" pageNumber="255">E. E. Gilbert</collectorName>
,
<collectorName id="FD6434D5FF92B065FA97C2F493A17F85" pageId="54" pageNumber="255">C. D. MacNeil</collectorName>
(
<collectionCode id="3680C9C6FF92B065FEC2C2D892177F85" box="[264,335,1467,1493]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="54" pageNumber="255" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF92B065FE9FC2D892227F85" box="[341,378,1467,1493]" count="1" pageId="54" pageNumber="255" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF92B065FE45C2D892FB7FA9" collectingDate="1960-04-11" collectionCode="USNM" collectorName="J. F. G. Clarke" country="Mexico" elevation="6" latitude="19.809444" location="Allen Pt Ascension Bay" longLatPrecision="21" longitude="-87.4775" pageId="54" pageNumber="255" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Quintana Roo" typeStatus="paratype">
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<collectingRegion id="92559FE1FF92B065FE45C2D891627F85" box="[399,570,1467,1493]" inferredBySuffix="true" pageId="54" pageNumber="255">Quintana Roo</collectingRegion>
,
</emphasis>
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,
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,
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,
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</date>
,
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(
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<specimenCount id="46979A8AFF92B065FEBEC2BC92C07FA9" box="[372,408,1503,1529]" count="1" pageId="54" pageNumber="255" type="male">1♂</specimenCount>
)
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.
</paragraph>
<paragraph id="502E5103FF92B065FF0DC16092687C35" blockId="54.[151,1437,1179,1961]" pageId="54" pageNumber="255">
<emphasis id="62E58D11FF92B065FF0DC160916F7C4D" bold="true" box="[199,567,1539,1565]" pageId="54" pageNumber="255">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFF92B065FDF4C160914C7C11" collectingDate="1964-07-22" collectionCode="AMNH" collectorName="J. C. &amp; D. Pallister" country="Mexico" elevation="3" latitude="21.28111" location="Progreso" longLatPrecision="21" longitude="-89.665276" pageId="54" pageNumber="255" specimenCount="4" specimenCount-male="4" stateProvince="Yucatan">
<emphasis id="62E58D11FF92B065FDF4C16090757C4D" bold="true" box="[574,813,1538,1565]" pageId="54" pageNumber="255">
<collectingCountry id="28861193FF92B065FDF4C16091E17C4D" box="[574,697,1539,1565]" name="Mexico" pageId="54" pageNumber="255">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF92B065FD0EC16190707C4D" box="[708,808,1538,1565]" country="Mexico" name="Yucatan" pageId="54" pageNumber="255">Yucatán</collectingRegion>
,
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<location id="554E07D8FF92B065FCFEC16090C27C4D" LSID="urn:lsid:plazi:treatment:D838E015FF90B065FF5DC3AD90EA7DF9:554E07D8FF92B065FCFEC16090C27C4D" box="[820,922,1539,1565]" country="Mexico" latitude="21.28111" longLatPrecision="21" longitude="-89.665276" name="Progreso" pageId="54" pageNumber="255" stateProvince="Yucatan">Progreso</location>
,
<quantity id="9769FCE6FF92B065FC6CC160908B7C4D" box="[934,979,1539,1565]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="54" pageNumber="255" unit="m" value="3.0">
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</quantity>
,
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<geoCoordinate id="35A537C4FF92B065FBBAC16097A77C4E" box="[1136,1279,1539,1566]" degrees="89" direction="west" minutes="39" orientation="longitude" pageId="54" pageNumber="255" precision="15" seconds="55" value="-89.665276">89°3955”W</geoCoordinate>
,
<date id="242F77C3FF92B065FAC1C16096C17C4D" box="[1291,1433,1539,1565]" pageId="54" pageNumber="255" value="1964-07-22">
<collectingDate id="346B8E2BFF92B065FAC1C16096C17C4D" box="[1291,1433,1539,1565]" pageId="54" pageNumber="255" value="1964-07-22">22/VII/1964</collectingDate>
</date>
,
<collectorName id="FD6434D5FF92B065FF5DC144922C7C11" box="[151,372,1575,1601]" pageId="54" pageNumber="255">J. C. &amp; D. Pallister</collectorName>
(
<collectionCode id="3680C9C6FF92B065FE4EC14492837C11" box="[388,475,1575,1601]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="54" pageNumber="255">AMNH</collectionCode>
<specimenCount id="46979A8AFF92B065FE2FC14491537C11" box="[485,523,1575,1601]" count="4" pageId="54" pageNumber="255" type="male">4♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF92B065FDEBC14492747C35" collectingDate="1964-07-25" collectionCode="AMNH" collectorName="J. C. &amp; D. Pallister" country="Mexico" elevation="5" latitude="21.166666" location="Puerto Sisal" longLatPrecision="21" longitude="-90.024445" pageId="54" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Yucatan">
<location id="554E07D8FF92B065FDEBC14491F47C11" LSID="urn:lsid:plazi:treatment:D838E015FF90B065FF5DC3AD90EA7DF9:554E07D8FF92B065FDEBC14491F47C11" box="[545,684,1575,1601]" country="Mexico" latitude="21.166666" longLatPrecision="21" longitude="-90.024445" name="Puerto Sisal" pageId="54" pageNumber="255" stateProvince="Yucatan">Puerto Sisal</location>
,
<quantity id="9769FCE6FF92B065FD73C14491B07C11" box="[697,744,1575,1602]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="54" pageNumber="255" unit="m" value="5.0">
<elevation id="DBBCB630FF92B065FD73C14491B07C11" box="[697,744,1575,1602]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="54" pageNumber="255" unit="m" value="5.0">5 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF92B065FD3CC14490277C11" box="[758,895,1575,1601]" degrees="21" direction="north" minutes="10" orientation="latitude" pageId="54" pageNumber="255" precision="15" seconds="00" value="21.166666">21°1000”N</geoCoordinate>
<geoCoordinate id="35A537C4FF92B065FC42C14497407C11" box="[904,1048,1575,1601]" degrees="90" direction="west" minutes="01" orientation="longitude" pageId="54" pageNumber="255" precision="15" seconds="28" value="-90.024445">90°0128”W</geoCoordinate>
,
<date id="242F77C3FF92B065FBEFC14497EA7C12" box="[1061,1202,1575,1602]" pageId="54" pageNumber="255" value="1964-07-25">
<collectingDate id="346B8E2BFF92B065FBEFC14497EA7C12" box="[1061,1202,1575,1602]" pageId="54" pageNumber="255" value="1964-07-25">25/VII/1964</collectingDate>
</date>
,
<collectorName id="FD6434D5FF92B065FB0AC14496C47C11" box="[1216,1436,1575,1601]" pageId="54" pageNumber="255">J. C. &amp; D. Pallister</collectorName>
(
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<specimenCount id="46979A8AFF92B065FF37C12892797C35" box="[253,289,1611,1637]" count="1" pageId="54" pageNumber="255" type="male">1♂</specimenCount>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF92B065FF0DC10C96357CFD" pageId="54" pageNumber="255" type="distribution">
<paragraph id="502E5103FF92B065FF0DC10C96357CFD" blockId="54.[151,1437,1179,1961]" pageId="54" pageNumber="255">
<emphasis id="62E58D11FF92B065FF0DC10C92047CD9" bold="true" box="[199,348,1647,1673]" pageId="54" pageNumber="255">Distribution.</emphasis>
This species is present in the
<collectingRegion id="92559FE1FF92B065FD5DC10C91B67CD9" box="[663,750,1647,1673]" country="Mexico" name="Yucatan" pageId="54" pageNumber="255">Yucatán</collectingRegion>
Peninsula and
<collectingRegion id="92559FE1FF92B065FC5AC10C90B27CD9" box="[912,1002,1647,1673]" country="Mexico" name="Chiapas" pageId="54" pageNumber="255">Chiapas</collectingRegion>
, being endemic to
<collectingCountry id="28861193FF92B065FB78C10C96507CD9" box="[1202,1288,1647,1673]" name="Mexico" pageId="54" pageNumber="255">Mexico</collectingCountry>
, mostly in the
<collectingRegion id="92559FE1FF92B065FF5DC1F093B67CFD" box="[151,238,1683,1709]" country="Mexico" name="Yucatan" pageId="54" pageNumber="255">Yucatán</collectingRegion>
Peninsula biogeographic province (
<bibRefCitation id="34002CF2FF92B065FDBFC1F0900A7CFD" author="Morrone, J. J. &amp; Escalante, T. &amp; Rodriguez-Tapia, G. &amp; Carmona, A. &amp; Arana, M. &amp; Mercado-Gomez, J. D." box="[629,850,1683,1709]" pageId="54" pageNumber="255" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. &amp; Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="62E58D11FF92B065FD17C1F790547CFD" box="[733,780,1683,1709]" italics="true" pageId="54" pageNumber="255">et al</emphasis>
. 2022
</bibRefCitation>
) (
<figureCitation id="C8AA4D86FF92B065FCA2C1F090E27CFD" box="[872,954,1683,1709]" captionStart="FIGURE 32" captionStartId="54.[151,250,1108,1132]" captionTargetBox="[159,1428,187,1083]" captionTargetId="figure-22@54.[151,1436,179,1083]" captionTargetPageId="54" captionText="FIGURE 32. Known distribution records for Heriades vulcanica sp. nov." pageId="54" pageNumber="255">Fig. 32</figureCitation>
). Altitudinal range: sea level to
<quantity id="9769FCE6FF92B065FAD9C1F096307CFD" box="[1299,1384,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="2.3609999999999998" pageId="54" pageNumber="255" unit="m" value="2361.0">2361 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF92B065FF0DC1D492717D6D" pageId="54" pageNumber="255" type="etymology">
<paragraph id="502E5103FF92B065FF0DC1D492717D6D" blockId="54.[151,1437,1179,1961]" pageId="54" pageNumber="255">
<emphasis id="62E58D11FF92B065FF0DC1D492167C81" bold="true" box="[199,334,1719,1745]" pageId="54" pageNumber="255">Etymology.</emphasis>
This species is dedicated to María Rojas Arias, and “La Vulcanizadora”, a film production company, founded by her and Andrés Jurado. She is a visual artist and filmmaker from
<collectingCountry id="28861193FF92B065FBBEC1B897BD7CA5" box="[1140,1253,1755,1781]" name="Colombia" pageId="54" pageNumber="255">Colombia</collectingCountry>
, she works with official and unofficial archival materials working with events that constituted national speeches, war policies, and colonization.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF92B065FF0DC02490EA7DF9" pageId="54" pageNumber="255" type="discussion">
<paragraph id="502E5103FF92B065FF0DC02490EA7DF9" blockId="54.[151,1437,1179,1961]" pageId="54" pageNumber="255">
<emphasis id="62E58D11FF92B065FF0DC02492177D31" bold="true" box="[199,335,1863,1889]" pageId="54" pageNumber="255">Comments.</emphasis>
On scutum DBP of males vary, from 0.2 to less. It was not possible to add more morphological information of female as there was just
<specimenCount id="46979A8AFF92B065FD97C00891A37DD5" box="[605,763,1899,1925]" count="1" pageId="54" pageNumber="255" type="generic">one specimen</specimenCount>
. This species is one of the few with large body size for the subgenus, and with orange hairs on posterior margin of posterior terga.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,719 @@
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<treatment id="D838E015FF95B067FF5DC3FA97327E2D" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D" httpUri="http://treatment.plazi.org/id/D838E015FF95B067FF5DC3FA97327E2D" lastPageId="52" lastPageNumber="253" pageId="49" pageNumber="250">
<subSubSection id="188B0288FF95B062FF5DC3FA90297EE3" box="[151,881,1177,1204]" pageId="49" pageNumber="250" type="nomenclature">
<paragraph id="502E5103FF95B062FF5DC3FA90297EE3" blockId="49.[151,881,1177,1239]" box="[151,881,1177,1204]" pageId="49" pageNumber="250">
<heading id="0B66E66FFF95B062FF5DC3FA90297EE3" bold="true" box="[151,881,1177,1204]" fontSize="11" level="1" pageId="49" pageNumber="250" reason="1">
<emphasis id="62E58D11FF95B062FF5DC3FA90297EE3" bold="true" box="[151,881,1177,1204]" pageId="49" pageNumber="250">
<taxonomicName id="97912A80FF95B062FF5DC3FA90547EE3" authority="Rojas-Arias, Griswold &amp; Ayala." authorityName="Rojas-Arias, Griswold &amp; Ayala." authorityYear="2025" box="[151,780,1177,1204]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="250" phylum="Arthropoda" rank="species" species="viridiana" status="sp. nov.">
<emphasis id="62E58D11FF95B062FF5DC3FA92217EE4" bold="true" box="[151,377,1177,1204]" italics="true" pageId="49" pageNumber="250">Heriades viridiana</emphasis>
Rojas-Arias, Griswold &amp; Ayala.
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF95B062FCDEC3FA90297EE3" box="[788,881,1177,1203]" pageId="49" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF95B062FF5DC3DE92067E87" box="[151,350,1213,1239]" pageId="49" pageNumber="250" type="description">
<paragraph id="502E5103FF95B062FF5DC3DE92067E87" blockId="49.[151,881,1177,1239]" box="[151,350,1213,1239]" pageId="49" pageNumber="250">
(
<figureCitation id="C8AA4D86FF95B062FF55C3DE925B7E87" box="[159,259,1213,1239]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="49" pageNumber="250">Figs. 2G</figureCitation>
,
<figureCitation id="C8AA4D86FF95B062FEC5C3DE920F7E87" box="[271,343,1213,1239]" captionStart-0="FIGURE 30" captionStart-1="FIGURE 31" captionStartId-0="49.[151,250,1071,1095]" captionStartId-1="51.[151,250,1943,1967]" captionTargetBox-0="[159,1428,157,1045]" captionTargetBox-1="[182,1392,192,1905]" captionTargetId-0="figure-18@49.[151,1436,149,1045]" captionTargetId-1="figure-18@51.[182,1405,179,1917]" captionTargetPageId-0="49" captionTargetPageId-1="51" captionText-0="FIGURE 30. Known distribution records for Heriades viridiana sp. nov." captionText-1="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="49" pageNumber="250">3031</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF95B062FF5DC266971F7C34" pageId="49" pageNumber="250" type="diagnosis">
<paragraph id="502E5103FF95B062FF5DC266971F7C34" blockId="49.[151,1437,1285,2029]" pageId="49" pageNumber="250">
<emphasis id="62E58D11FF95B062FF5DC266924A7F4F" bold="true" box="[151,274,1285,1311]" pageId="49" pageNumber="250">Diagnosis.</emphasis>
Tergal fasciae made of one row of thick, white hairs (
<figureCitation id="C8AA4D86FF95B062FCA2C26690EF7F4F" box="[872,951,1285,1311]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="49" pageNumber="250">Fig. 3J</figureCitation>
).
<emphasis id="62E58D11FF95B062FC06C266975E7F4F" box="[972,1030,1285,1311]" italics="true" pageId="49" pageNumber="250">Male</emphasis>
: at least ventral surface of F2 to F11 bright orange, scape, pedicel and F1 black; clypeus with plumose, orangish, dense hairs, with similar length evenly distributed, joining hairs on hypostomal area (
<figureCitation id="C8AA4D86FF95B062FD61C22E904E7F37" box="[683,790,1357,1383]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="49" pageNumber="250">Fig. 31A</figureCitation>
); S1 with strong process, in profile with sides projected forming an acute angle, in posterior view of process with one acute angle (
<figureCitation id="C8AA4D86FF95B062FC3FC21297617FDB" box="[1013,1081,1393,1419]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="49" pageNumber="250">Fig. 2</figureCitation>
Ga); S5 with two subtriangular lobes, and a line of capitate hairs increasing in length toward sides (
<figureCitation id="C8AA4D86FF95B062FC4EC2F690B07FFF" box="[900,1000,1429,1455]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="49" pageNumber="250">Fig. 31E</figureCitation>
); S6 with two subapical tufts of several hooked, long hairs, all hairs with similar length and directed medially and on posterior margin a few simple, sort hairs medially (
<figureCitation id="C8AA4D86FF95B062FE82C2BE92F47FA7" box="[328,428,1501,1527]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="49" pageNumber="250">Fig. 31F</figureCitation>
).
<emphasis id="62E58D11FF95B062FE0BC2BE914D7FA7" box="[449,533,1501,1527]" italics="true" pageId="49" pageNumber="250">Female</emphasis>
: mandible with outer surface without tubercle; clypeus, apical margin with five or more tubercles; T3 with punctures medium-sized (0.33 times DOP as on scutum), and sparse (0.5 times DOP), similar to
<taxonomicName id="97912A80FF95B062FEC2C14592DF7C6F" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[264,391,1573,1599]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="250" phylum="Arthropoda" rank="species" species="briceida" status="sp. nov.">
<emphasis id="62E58D11FF95B062FEC2C14592DF7C6F" box="[264,391,1573,1599]" italics="true" pageId="49" pageNumber="250">H. briceida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF95B062FE44C14592BC7C10" box="[398,484,1574,1600]" pageId="49" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
and seemly differing only in not having clypeus with stronger emargination and with tubercles in the middle of apical margin, and mesepisternum with sparser punctures.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF95B067FF0DC10E91387A85" lastPageId="52" lastPageNumber="253" pageId="49" pageNumber="250" type="description">
<paragraph id="502E5103FF95B062FF0DC10E97A67CD7" blockId="49.[151,1437,1285,2029]" box="[199,1278,1645,1672]" pageId="49" pageNumber="250">
<emphasis id="62E58D11FF95B062FF0DC10E92FC7CD7" bold="true" box="[199,420,1645,1672]" pageId="49" pageNumber="250">
Description.
<emphasis id="62E58D11FF95B062FEAAC10E92C37CD8" bold="true" box="[352,411,1645,1672]" italics="true" pageId="49" pageNumber="250">Male</emphasis>
:
</emphasis>
Body length,
<quantity id="9769FCE6FF95B062FD89C10E91C17CD8" box="[579,665,1645,1672]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="49" pageNumber="250" unit="mm" value="6.6">6.6 mm</quantity>
(5.76.6, n=6); forewing length,
<quantity id="9769FCE6FF95B062FBC5C10E973D7CD8" box="[1039,1125,1645,1672]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="49" pageNumber="250" unit="mm" value="3.3">3.3 mm</quantity>
(3.34, n=6).
</paragraph>
<paragraph id="502E5103FF95B061FF0DC1F292317B33" blockId="49.[151,1437,1285,2029]" lastBlockId="50.[151,1437,151,2035]" lastPageId="50" lastPageNumber="251" pageId="49" pageNumber="250">
<emphasis id="62E58D11FF95B062FF0DC1F2925C7CFB" box="[199,260,1681,1707]" italics="true" pageId="49" pageNumber="250">Head</emphasis>
. Length
<quantity id="9769FCE6FF95B062FEA3C1F292987CFC" box="[361,448,1681,1708]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="49" pageNumber="250" unit="mm" value="1.5">1.5 mm</quantity>
(1.41.6, n=6), width
<quantity id="9769FCE6FF95B062FD74C1F2904C7CFC" box="[702,788,1681,1708]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="49" pageNumber="250" unit="mm" value="1.8">1.8 mm</quantity>
(1.61.9, n=6); eyes about 3 times longer than broad, 2.1 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF95B062FC83C1D690F97C9F" box="[841,929,1717,1743]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="49" pageNumber="250" unit="mm" value="0.9">0.9 mm</quantity>
(0.80.9, n=6); upper interocular distance
<quantity id="9769FCE6FF95B062FA44C1D6939B7CA3" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="49" pageNumber="250" unit="mm" value="1.0">1 mm</quantity>
(0.91.1, n=6); interantennal distance 2 times antennocular distance; lateral ocelli large (
<figureCitation id="C8AA4D86FF95B062FB0BC1BB96407CA3" box="[1217,1304,1752,1779]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="49" pageNumber="250">Fig. 3C</figureCitation>
) (3.9 times DOP in interocellar area); interocellar distance 2 times OD; labrum with wide, basal, medial tubercle posterior margin subquadrangular; dorsal margin of mandible with carina; clypeus with shallow emargination on all length and crenulated of apical margin, 2 times broader than long, lower half convex in profile, apical margin without emargination (
<figureCitation id="C8AA4D86FF95B062FEF4C00492F37DD2" box="[318,427,1895,1922]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="49" pageNumber="250">Fig. 31A</figureCitation>
); ocellocular distance 1.4 times OD, OPR 2 times OD; scape black, length 0.7 times alveolocellar distance, 2.1 times longer than wide; pedicel black, about 2 times longer than F1 and about 0.9 times longer than F2; F1 dark brown, broader than long, remaining flagellomeres yellow and slightly dark brown on dorsal surface, 2.3 longer than first; F11 1.4 times longer than broad.
<emphasis id="62E58D11FF95B062FC99C0B0908B7DBC" box="[851,979,2003,2028]" italics="true" pageId="49" pageNumber="250">Mesosoma.</emphasis>
Scutum 2.7 times longer than scutellum, 3.1 times longer than tegula, and 3.4 longer than axilla; intertegular distance
<quantity id="9769FCE6FF96B061FBC1C7F4973B7AE1" box="[1035,1123,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="50" pageNumber="251" unit="mm" value="1.3">1.3 mm</quantity>
(1.11.3, n=6); metanotum with transverse ridge, in posterior view with two small medial subtriangular elevations; propodeum with small to midsized pits (
<figureCitation id="C8AA4D86FF96B061FEF7C7BD92C07AA8" box="[317,408,222,248]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="50" pageNumber="251">Fig. 3H</figureCitation>
),
<quantity id="9769FCE6FF96B061FE67C7BD914A7AA9" box="[429,530,222,249]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="50" pageNumber="251" unit="mm" value="0.06">0.06 mm</quantity>
long; legs dark amber.
<emphasis id="62E58D11FF96B061FCD7C7BC90C27AA8" box="[797,922,223,248]" italics="true" pageId="50" pageNumber="251">Metasoma.</emphasis>
T1
<quantity id="9769FCE6FF96B061FC02C7BD97477AA9" box="[968,1055,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="50" pageNumber="251" unit="mm" value="1.5">1.5 mm</quantity>
broad (1.31.6, n=6); T3
<quantity id="9769FCE6FF96B061FA8CC7BD96C57AA9" box="[1350,1437,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="50" pageNumber="251" unit="mm" value="1.7">1.7 mm</quantity>
broad (1.51.8, n=6); posterior margin of S1 truncate with shallow, broad emargination and subapical smooth carina (
<figureCitation id="C8AA4D86FF96B061FF55C64693B97B6F" box="[159,225,293,319]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="50" pageNumber="251">Fig. 2</figureCitation>
Gb), in profile, with posterior margin not projected and strong process apex on one half of distance from apex to base (
<figureCitation id="C8AA4D86FF96B061FF3FC62A92607B33" box="[245,312,329,355]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="50" pageNumber="251">Fig. 2</figureCitation>
Ga).
</paragraph>
<paragraph id="502E5103FF96B061FF0DC60E922A7964" blockId="50.[151,1437,151,2035]" pageId="50" pageNumber="251">
<emphasis id="62E58D11FF96B061FF0DC60E92097BD7" box="[199,337,365,391]" italics="true" pageId="50" pageNumber="251">Sculpturing.</emphasis>
Clypeus with small (at base 0.7 times OD), contiguous, coarse punctures, equally sized; supraclypeal area with punctures smaller than base of clypeus; frons with bigger, coarser and denser punctures than those on supraclipeal area; frons in front of ocelli with sparser punctation than on frons; vertex with bigger, sparser punctures than on supraclypeal area, with micropunctures between punctures; hypostomal area with punctation as on clypeus, bigger and sparser on gena; pronotum anterior surface with punctures sparser and less coarse than on clypeus, denser and slightly coarser on pronotal lobe; disc of scutum with bigger (0.4 times OD), almost contiguous punctures than on vertex; mesepisternum with equal DBP and DOP punctures as on scutum; metepisternum with smaller punctures, less dense and coarse; T1 and T2 with smaller (0.2 times OD) punctures than on scutum, equally sparser; T3 with bigger, less sparse, coarse punctures than on T1, slightly rounded, separated by less than third DOP; T4 and T5 with denser and coarser punctation than on T3; dorsal surface of T6 with punctation as on T5, on posterior surface with notably smaller and sparser punctation, DBP half DOP; sides of S1 with punctation bigger and parser than T1, and in process notably smaller and denser; sides of S2 with punctures smaller than on T1, disc of S2 microreticulate without punctation.
</paragraph>
<paragraph id="502E5103FF96B061FF0DC45D96337E5A" blockId="50.[151,1437,151,2035]" pageId="50" pageNumber="251">
<emphasis id="62E58D11FF96B061FF0DC45D92777907" box="[199,303,830,855]" italics="true" pageId="50" pageNumber="251">Vestiture.</emphasis>
Clypeus with plumose, orangish, equally length, dense hairs, evenly distributed; supraclypeal area and frons with plumose, thin, white to orangish hairs directed dorsally, equally length; hypostomal area with plumose, long, white hairs, joining hairs in clypeus, in frontal view surpassing mandible (
<figureCitation id="C8AA4D86FF96B061FBE3C4E697CA79CF" box="[1065,1170,901,927]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="50" pageNumber="251">Fig. 31A</figureCitation>
); dorsolateral angles of pronotum with plumose, thin, scattered, white hairs; disc of metepisternum without tuft of thick, short, white hairs; tergal fasciae with row of thick, short, orangish hairs, present from T1 to T4; disc of T4 to T6 with simple, long, thick, white hairs in punctures (
<figureCitation id="C8AA4D86FF96B061FE7AC493914D7E5A" box="[432,533,1008,1034]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="50" pageNumber="251">Fig. 31B</figureCitation>
); S1 with posterior surface of process with plumose, thin, dense, white hairs.
</paragraph>
<paragraph id="502E5103FF96B061FF0DC37797D37EB1" blockId="50.[151,1437,151,2035]" pageId="50" pageNumber="251">
<emphasis id="62E58D11FF96B061FF0DC37792117E7E" box="[199,329,1044,1070]" italics="true" pageId="50" pageNumber="251">
<taxonomicName id="97912A80FF96B061FF0DC377921D7E7E" box="[199,325,1044,1070]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="50" pageNumber="251" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
Posterior margin of S5 with two subtriangular lobes directed backwards, and a line of capitate hairs directed and increasing in length outward, with transverse, thin, dark, subapical, bilobed line (
<figureCitation id="C8AA4D86FF96B061FB0DC35B96767E02" box="[1223,1326,1080,1106]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="50" pageNumber="251">Fig. 31E</figureCitation>
); S6 with rounded, obtuse-angled posterior margin, without blackish subapical margin, with two subapical tufts of several hooked, long, thin hairs, with similar length directed medially (
<figureCitation id="C8AA4D86FF96B061FCAAC31C909C7EC9" box="[864,964,1151,1177]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="50" pageNumber="251">Fig. 31F</figureCitation>
); capsule in dorsal view
<quantity id="9769FCE6FF96B061FB16C31C96467ECA" box="[1244,1310,1151,1178]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="50" pageNumber="251" unit="mm" value="1.0">1 mm</quantity>
length,
<quantity id="9769FCE6FF96B061FAB3C31C939B7EED" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="50" pageNumber="251" unit="mm" value="0.4">0.4 mm</quantity>
width, 2.5 times longer than wide, gonobase
<quantity id="9769FCE6FF96B061FD7DC3C090427EED" box="[695,794,1187,1213]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="50" pageNumber="251" unit="mm" value="0.06">0.06 mm</quantity>
length, 6.5 times wider than long, and gonocoxite
<quantity id="9769FCE6FF96B061FA82C3C096C57EED" box="[1352,1437,1187,1213]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="50" pageNumber="251" unit="mm" value="0.4">0.4 mm</quantity>
length, gonostylus
<quantity id="9769FCE6FF96B061FEA6C3A492F57EB1" box="[364,429,1223,1249]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="50" pageNumber="251" unit="mm" value="1.0">1 mm</quantity>
length with rounded apex, penis valve
<quantity id="9769FCE6FF96B061FCAEC3A490E27EB1" box="[868,954,1223,1249]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="50" pageNumber="251" unit="mm" value="0.7">0.7 mm</quantity>
length (
<figureCitation id="C8AA4D86FF96B061FBDCC3A497277EB1" box="[1046,1151,1223,1249]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="50" pageNumber="251">Fig. 31G</figureCitation>
).
</paragraph>
<paragraph id="502E5103FF96B061FF0DC38996297C72" blockId="50.[151,1437,151,2035]" pageId="50" pageNumber="251">
<emphasis id="62E58D11FF96B061FF0DC38992707F54" bold="true" box="[199,296,1258,1284]" pageId="50" pageNumber="251">Female:</emphasis>
Body length
<quantity id="9769FCE6FF96B061FE0BC389914F7F55" box="[449,535,1258,1285]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="50" pageNumber="251" unit="mm" value="6.3">6.3 mm</quantity>
(5.86.3, n=2); forewing length
<quantity id="9769FCE6FF96B061FC43C38990877F55" box="[905,991,1258,1285]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="50" pageNumber="251" unit="mm" value="4.3">4.3 mm</quantity>
(3.54.3, n=2). Similar to male except:
<emphasis id="62E58D11FF96B061FF5DC26D938C7F78" box="[151,212,1294,1320]" italics="true" pageId="50" pageNumber="251">Head</emphasis>
. Length
<quantity id="9769FCE6FF96B061FEF6C26D92CC7F79" box="[316,404,1294,1321]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="50" pageNumber="251" unit="mm" value="1.8">1.8 mm</quantity>
(1.61.8, n=2), width
<quantity id="9769FCE6FF96B061FD52C26D91A87F79" box="[664,752,1294,1321]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="50" pageNumber="251" unit="mm" value="1.9">1.9 mm</quantity>
(1.61.9, n=2); eyes about 3.7 times longer than broad, 1.4 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FF96B061FC9DC25190E97F1C" box="[855,945,1330,1356]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="50" pageNumber="251" unit="mm" value="0.9">0.9 mm</quantity>
(0.80.9, n=2); upper interocular distance
<quantity id="9769FCE6FF96B061FF5DC23593B77F20" box="[151,239,1366,1392]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="50" pageNumber="251" unit="mm" value="1.1">1.1 mm</quantity>
(11.1, n=2); interantennal distance 1.6 times antennocular distance; interocellar distance 1.9 times OD, ocellocular distance 1.5 times OD, OPR 1.9 times OD; labrum without rounded, basal, medial tubercle; dorsal margin of mandible with carina; clypeus with shallow emargination on all length and crenulated on apical margin (
<figureCitation id="C8AA4D86FF96B061FF55C2A292507F8B" box="[159,264,1473,1499]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="50" pageNumber="251">Fig. 31C</figureCitation>
); scape 3 times longer than broad; pedicel 1.2 longer than broad, about 2 times longer than F1 and F2, remaining flagellomeres 2 times longer than first.
<emphasis id="62E58D11FF96B061FD03C28690197FAE" box="[713,833,1509,1534]" italics="true" pageId="50" pageNumber="251">Mesosoma</emphasis>
. Scutum 3.1 times longer than scutellum; intertegular distance
<quantity id="9769FCE6FF96B061FF30C16B92087C73" box="[250,336,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="50" pageNumber="251" unit="mm" value="1.3">1.3 mm</quantity>
(1.21.3, n=2).
<emphasis id="62E58D11FF96B061FDCFC16A91D97C72" box="[517,641,1545,1570]" italics="true" pageId="50" pageNumber="251">Metasoma.</emphasis>
T1
<quantity id="9769FCE6FF96B061FD64C16B905C7C73" box="[686,772,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="50" pageNumber="251" unit="mm" value="1.7">1.7 mm</quantity>
broad (1.41.7, n=2); T3
<quantity id="9769FCE6FF96B061FBECC16B97247C73" box="[1062,1148,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="50" pageNumber="251" unit="mm" value="1.9">1.9 mm</quantity>
broad (1.61.9, n=2).
</paragraph>
<paragraph id="502E5103FF96B061FF0DC14F92007DA3" blockId="50.[151,1437,151,2035]" pageId="50" pageNumber="251">
<emphasis id="62E58D11FF96B061FF0DC14F92097C16" box="[199,337,1580,1606]" italics="true" pageId="50" pageNumber="251">Sculpturing.</emphasis>
Clypeus with punctures medium-sized (0.2 times OD), dense (0.2 DBP) and coarse, between punctures shiny and smooth; supraclypeal area with punctures as on upper margin of clypeus; frons with bigger, coarser, denser and slightly sparser punctation than on supraclypeal area, with micropunctures between punctures; frons in front of ocelli slightly smaller punctures than those on frons; vertex with punctures bigger, coarser and slightly sparser than on clypeus, with micropunctures between punctures; gena evenly punctate, with bigger, less coarse punctures and with more DBP than on frons in front of ocelli; scutum with bigger and denser punctures than on vertex; rest of mesosoma as in scutum except: scutellum with sparser punctures; mesepisternum with slightly smaller punctures on ventral half; metasomal terga smooth and shiny, microreticulate between punctures; T1 with smaller (0.2 times OD), slightly denser punctures than on scutum, with slightly semicircular shape; T2 with smaller, sparser, semicircular shaped punctures than on T1; T3 with punctures smaller (0.2 times DOP), less coarse, than on scutum, horseshoe-shaped, and sparser (0.3 times DOP); dorsal surface of T6 with smaller, denser, and more coarsely punctures than on T5; sterna with punctation smaller and sparser than on scutum, with microreticulate on posterior margin.
</paragraph>
<caption id="04EE018BFF97B060FF5DC0F490187D83" pageId="51" pageNumber="252" startId="51.[151,250,1943,1967]" targetBox="[182,1392,192,1905]" targetPageId="51" targetType="figure">
<paragraph id="502E5103FF97B060FF5DC0F490187D83" blockId="51.[151,1437,1943,2003]" pageId="51" pageNumber="252">
<emphasis id="62E58D11FF97B060FF5DC0F492787DFF" bold="true" box="[151,288,1943,1967]" pageId="51" pageNumber="252">FIGURE 31.</emphasis>
<taxonomicName id="97912A80FF97B060FEEDC0F492B07DFF" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[295,488,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="252" phylum="Arthropoda" rank="species" species="viridiana" status="sp. nov.">
<emphasis id="62E58D11FF97B060FEEDC0F492B07DFF" box="[295,488,1943,1967]" italics="true" pageId="51" pageNumber="252">Heriades viridiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF97B060FE24C0FB91657DFF" box="[494,573,1944,1967]" pageId="51" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
Male:
<emphasis id="62E58D11FF97B060FD4FC0F491C67DFF" bold="true" box="[645,670,1943,1967]" pageId="51" pageNumber="252">A.</emphasis>
frontal view
<emphasis id="62E58D11FF97B060FCEDC0F490667DFF" bold="true" box="[807,830,1943,1967]" pageId="51" pageNumber="252">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FF97B060FBD5C0F4976F7DFF" bold="true" box="[1055,1079,1943,1967]" pageId="51" pageNumber="252">C.</emphasis>
frontal view
<emphasis id="62E58D11FF97B060FB0AC0F497817DFF" bold="true" box="[1216,1241,1943,1967]" pageId="51" pageNumber="252">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FF97B060FF5DC0D893F67D83" bold="true" box="[151,174,1979,2003]" pageId="51" pageNumber="252">E.</emphasis>
S5.
<emphasis id="62E58D11FF97B060FF16C0D893B77D83" bold="true" box="[220,239,1979,2003]" pageId="51" pageNumber="252">F.</emphasis>
S6.
<emphasis id="62E58D11FF97B060FED7C0D8926F7D83" bold="true" box="[285,311,1979,2003]" pageId="51" pageNumber="252">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FF97B060FD56C0D891EE7D83" bold="true" box="[668,694,1979,2003]" pageId="51" pageNumber="252">H.</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="502E5103FF90B067FF0DC7F491387A85" blockId="52.[151,1437,151,1149]" pageId="52" pageNumber="253">
<emphasis id="62E58D11FF90B067FF0DC7F492687AE0" box="[199,304,151,176]" italics="true" pageId="52" pageNumber="253">Vestiture.</emphasis>
Clypeus and frons with few short, white hairs in punctures (
<figureCitation id="C8AA4D86FF90B067FC15C7F4971F7AE1" box="[991,1095,151,177]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="52" pageNumber="253">Fig. 31C</figureCitation>
); subapical margin of T6 with linear tuft of plumose, short, white hairs.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF90B067FF0DC7BC970E7969" pageId="52" pageNumber="253" type="materials_examined">
<paragraph id="502E5103FF90B067FF0DC7BC96CF7BFD" blockId="52.[151,1437,151,1149]" pageId="52" pageNumber="253">
<emphasis id="62E58D11FF90B067FF0DC7BC92267AA8" bold="true" box="[199,382,222,249]" pageId="52" pageNumber="253">Type material:</emphasis>
<materialsCitation id="E0F95B5EFF90B067FE46C7BC91D07B4D" collectingDate="1988-10-08" collectorName="No. LR &amp; La Laja &amp; R. Ayala" country="Mexico" elevation="441" latitude="18.93111" location="La Laja" longLatPrecision="21" longitude="-101.86694" pageId="52" pageNumber="253" specimenCount="1" stateProvince="Michoacan de Ocampo" typeStatus="holotype">
<emphasis id="62E58D11FF90B067FE46C7BC977F7AA9" bold="true" box="[396,1063,222,249]" pageId="52" pageNumber="253">
<typeStatus id="8F2AEFA1FF90B067FE46C7BC92A17AA9" box="[396,505,223,249]" pageId="52" pageNumber="253" type="holotype">Holotype</typeStatus>
male (
<collectorName id="FD6434D5FF90B067FD9CC7BD91E07AA8" box="[598,696,222,248]" pageId="52" pageNumber="253">No. LR</collectorName>
0005):
<collectingCountry id="28861193FF90B067FCC4C7BD90D17AA8" box="[782,905,222,248]" name="Mexico" pageId="52" pageNumber="253">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF90B067FC56C7BD977A7AA9" box="[924,1058,222,249]" country="Mexico" name="Michoacan de Ocampo" pageId="52" pageNumber="253">Michoacán</collectingRegion>
,
</emphasis>
<collectorName id="FD6434D5FF90B067FBFFC7BC97CD7AA9" box="[1077,1173,223,249]" pageId="52" pageNumber="253">
<location id="554E07D8FF90B067FBFFC7BC97CD7AA9" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FBFFC7BC97CD7AA9" box="[1077,1173,223,249]" country="Mexico" latitude="18.93111" longLatPrecision="21" longitude="-101.86694" name="La Laja" pageId="52" pageNumber="253" stateProvince="Michoacan de Ocampo">La Laja</location>
</collectorName>
, [
<quantity id="9769FCE6FF90B067FB64C7BC96587AA9" box="[1198,1280,223,249]" metricMagnitude="2" metricUnit="m" metricValue="4.41" pageId="52" pageNumber="253" unit="m" value="441.0">
<elevation id="DBBCB630FF90B067FB64C7BC96587AA9" box="[1198,1280,223,249]" metricMagnitude="2" metricUnit="m" metricValue="4.41" pageId="52" pageNumber="253" unit="m" value="441.0">441 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF90B067FAD9C7BD96C47AA9" box="[1299,1436,222,249]" degrees="18" direction="north" minutes="55" orientation="latitude" pageId="52" pageNumber="253" precision="15" seconds="52" value="18.93111">18°5552”N</geoCoordinate>
<geoCoordinate id="35A537C4FF90B067FF5DC66192617B4D" box="[151,313,258,285]" degrees="101" direction="west" minutes="52" orientation="longitude" pageId="52" pageNumber="253" precision="15" seconds="01" value="-101.86694">101°5201”W</geoCoordinate>
],
<date id="242F77C3FF90B067FE87C66092E27B4D" box="[333,442,259,285]" pageId="52" pageNumber="253" value="1988-10-08">
<collectingDate id="346B8E2BFF90B067FE87C66092E27B4D" box="[333,442,259,285]" pageId="52" pageNumber="253" value="1988-10-08">8/X/1988</collectingDate>
</date>
,
<collectorName id="FD6434D5FF90B067FE0FC660917D7B4D" box="[453,549,259,285]" pageId="52" pageNumber="253">R. Ayala</collectorName>
(EBCh)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF90B067FD58C661972E7B11" collectingDate="1998-09-04" collectionCode="BBSL" collectorName="Izucar de Matamoros SE" country="Mexico" latitude="18.456" location="Izucar de Matamoros" longLatPrecision="13" longitude="-98.373665" pageId="52" pageNumber="253" specimenCode="BBSL335211" specimenCount="1" stateProvince="Morelos" typeStatus="allotype">
<emphasis id="62E58D11FF90B067FD58C66197F57B4D" bold="true" box="[658,1197,258,285]" pageId="52" pageNumber="253">
<typeStatus id="8F2AEFA1FF90B067FD58C66191AF7B4C" box="[658,759,258,284]" pageId="52" pageNumber="253" type="allotype">Allotype</typeStatus>
(
<specimenCode id="0037F978FF90B067FCCDC66190F07B4C" box="[775,936,258,284]" collectionCode="BBSL" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="52" pageNumber="253" type="Museum">BBSL335211</specimenCode>
):
<collectingCountry id="28861193FF90B067FC77C66197607B4C" box="[957,1080,258,284]" name="Mexico" pageId="52" pageNumber="253">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF90B067FB8EC66097FF7B4D" box="[1092,1191,259,285]" country="Mexico" name="Morelos" pageId="52" pageNumber="253">Morelos</collectingRegion>
,
</emphasis>
<collectingRegion id="92559FE1FF90B067FB7FC660965C7B4D" box="[1205,1284,259,285]" country="Mexico" name="Puebla" pageId="52" pageNumber="253">Puebla</collectingRegion>
, Tehuitzingo,
<collectorName id="FD6434D5FF90B067FF5DC64492957B11" box="[151,461,295,321]" pageId="52" pageNumber="253">
<location id="554E07D8FF90B067FF5DC64492C17B11" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FF5DC64492C17B11" box="[151,409,295,321]" country="Mexico" latitude="18.456" longLatPrecision="13" longitude="-98.373665" name="Izucar de Matamoros" pageId="52" pageNumber="253" stateProvince="Morelos">Izucar de Matamoros</location>
SE
</collectorName>
,
<geoCoordinate id="35A537C4FF90B067FE28C644913E7B11" box="[482,614,295,321]" degrees="18" direction="north" minutes="27.36" orientation="latitude" pageId="52" pageNumber="253" precision="9" value="18.456">18°27.36N</geoCoordinate>
<geoCoordinate id="35A537C4FF90B067FDBDC64490597B11" box="[631,769,295,321]" degrees="98" direction="west" minutes="22.42" orientation="longitude" pageId="52" pageNumber="253" precision="9" value="-98.373665">98°22.42W</geoCoordinate>
,
<date id="242F77C3FF90B067FCDCC64490D47B11" box="[790,908,295,321]" pageId="52" pageNumber="253" value="1998-09-04">
<collectingDate id="346B8E2BFF90B067FCDCC64490D47B11" box="[790,908,295,321]" pageId="52" pageNumber="253" value="1998-09-04">4/IX/1998</collectingDate>
</date>
, Griswold (
<collectionCode id="3680C9C6FF90B067FBE8C64497367B11" box="[1058,1134,295,321]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="52" pageNumber="253" type="Museum">BBSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF90B067FB41C64497867B35" collectingDate="1988-10-08" collectorName="La Laja &amp; R. Ayala" country="Mexico" elevation="441" latitude="18.93111" location="La Laja" longLatPrecision="21" longitude="-101.86694" pageId="52" pageNumber="253" specimenCount="25" specimenCount-female="1" specimenCount-male="24" stateProvince="Michoacan de Ocampo" typeStatus="paratype">
<emphasis id="62E58D11FF90B067FB41C644927A7B35" bold="true" pageId="52" pageNumber="253">
<typeStatus id="8F2AEFA1FF90B067FB41C644965F7B11" box="[1163,1287,295,321]" pageId="52" pageNumber="253" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="28861193FF90B067FAD7C64596C07B10" box="[1309,1432,294,320]" name="Mexico" pageId="52" pageNumber="253">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF90B067FF5DC62992457B35" box="[151,285,330,357]" country="Mexico" name="Michoacan de Ocampo" pageId="52" pageNumber="253">Michoacán</collectingRegion>
,
</emphasis>
<collectorName id="FD6434D5FF90B067FEE0C62892DD7B35" box="[298,389,331,357]" pageId="52" pageNumber="253">
<location id="554E07D8FF90B067FEE0C62892DD7B35" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FEE0C62892DD7B35" box="[298,389,331,357]" country="Mexico" latitude="18.93111" longLatPrecision="21" longitude="-101.86694" name="La Laja" pageId="52" pageNumber="253" stateProvince="Michoacan de Ocampo">La Laja</location>
</collectorName>
, [
<quantity id="9769FCE6FF90B067FE5DC62892BC7B35" box="[407,484,331,357]" metricMagnitude="2" metricUnit="m" metricValue="4.41" pageId="52" pageNumber="253" unit="m" value="441.0">
<elevation id="DBBCB630FF90B067FE5DC62892BC7B35" box="[407,484,331,357]" metricMagnitude="2" metricUnit="m" metricValue="4.41" pageId="52" pageNumber="253" unit="m" value="441.0">441 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF90B067FE3AC62991227B35" box="[496,634,330,357]" degrees="18" direction="north" minutes="55" orientation="latitude" pageId="52" pageNumber="253" precision="15" seconds="52" value="18.93111">18°5552”N</geoCoordinate>
<geoCoordinate id="35A537C4FF90B067FD4BC629907B7B35" box="[641,803,330,357]" degrees="101" direction="west" minutes="52" orientation="longitude" pageId="52" pageNumber="253" precision="15" seconds="01" value="-101.86694">101°5201”W</geoCoordinate>
],
<date id="242F77C3FF90B067FCFDC62890FC7B35" box="[823,932,331,357]" pageId="52" pageNumber="253" value="1988-10-08">
<collectingDate id="346B8E2BFF90B067FCFDC62890FC7B35" box="[823,932,331,357]" pageId="52" pageNumber="253" value="1988-10-08">8/X/1988</collectingDate>
</date>
,
<collectorName id="FD6434D5FF90B067FC7AC62897487B35" box="[944,1040,331,357]" pageId="52" pageNumber="253">R. Ayala</collectorName>
(EBCh,
<specimenCount id="46979A8AFF90B067FBBFC62897C07B35" box="[1141,1176,331,357]" count="1" pageId="52" pageNumber="253" type="female">1♀</specimenCount>
<specimenCount id="46979A8AFF90B067FB6AC628978D7B35" box="[1184,1237,331,357]" count="24" pageId="52" pageNumber="253" type="male">24♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF90B067FB23C62892D67BFD" collectingDate="1996-09-07" collectionCode="KUNHM-ENT" collectorName="Sierra de Huautla, C. E. A. M. I. S. H &amp; R. Brooks" country="Mexico" elevation="1050" latitude="18.45" location="Sierra de Huautla" longLatPrecision="21" longitude="-99.03333" pageId="52" pageNumber="253" specimenCount="10" specimenCount-male="10" stateProvince="Morelos" typeStatus="paratype">
<emphasis id="62E58D11FF90B067FB23C62896097B35" bold="true" box="[1257,1361,331,357]" pageId="52" pageNumber="253">
<collectingRegion id="92559FE1FF90B067FB23C62896147B35" box="[1257,1356,331,357]" country="Mexico" name="Morelos" pageId="52" pageNumber="253">Morelos</collectingRegion>
,
</emphasis>
<collectorName id="FD6434D5FF90B067FA93C62892917BD9" pageId="52" pageNumber="253">
<location id="554E07D8FF90B067FA93C628924C7BD9" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FA93C628924C7BD9" country="Mexico" latitude="18.45" longLatPrecision="21" longitude="-99.03333" name="Sierra de Huautla" pageId="52" pageNumber="253" stateProvince="Morelos">Sierra de Huautla</location>
, C.E.A.M.I.S.H
</collectorName>
. Huatla
<quantity id="9769FCE6FF90B067FDEEC60D91337BD9" box="[548,619,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="52" pageNumber="253" unit="km" value="2.5">2.5km</quantity>
N 4.0km W, [
<quantity id="9769FCE6FF90B067FCCEC60D90057BD9" box="[772,861,366,393]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="52" pageNumber="253" unit="m" value="1050.0">
<elevation id="DBBCB630FF90B067FCCEC60D90057BD9" box="[772,861,366,393]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="52" pageNumber="253" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF90B067FCA3C60C90BB7BD9" box="[873,995,367,393]" degrees="18" direction="north" minutes="27" orientation="latitude" pageId="52" pageNumber="253" precision="15" seconds="0" value="18.45">18°270”N</geoCoordinate>
<geoCoordinate id="35A537C4FF90B067FC20C60C973A7BD9" box="[1002,1122,367,393]" degrees="99" direction="west" minutes="2" orientation="longitude" pageId="52" pageNumber="253" precision="15" seconds="0" value="-99.03333">99°20”W</geoCoordinate>
], 1, 5,
<date id="242F77C3FF90B067FB61C60C96787BD9" box="[1195,1312,367,393]" pageId="52" pageNumber="253" value="1996-09-07">
<collectingDate id="346B8E2BFF90B067FB61C60C96787BD9" box="[1195,1312,367,393]" pageId="52" pageNumber="253" value="1996-09-07">7/IX/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FF90B067FAE1C60C96C47BD9" box="[1323,1436,367,393]" pageId="52" pageNumber="253">R. Brooks</collectorName>
(
<collectionCode id="3680C9C6FF90B067FF54C6F092117BFD" box="[158,329,403,429]" pageId="52" pageNumber="253">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFF90B067FE9BC6F092DD7BFD" box="[337,389,403,429]" count="10" pageId="52" pageNumber="253" type="male">10♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF90B067FE5DC6F096C97BFD" box="[407,1425,402,429]" collectingDate="1996-09-06" collectionCode="KUNHM-ENT" collectorName="R. Brooks" country="Mexico" elevation="990" latitude="18.464445" location="Ajuchitlan" longLatPrecision="21" longitude="-98.99166" pageId="52" pageNumber="253" specimenCount="1" specimenCount-female="1" stateProvince="Morelos" typeStatus="paratype">
<location id="554E07D8FF90B067FE5DC6F091487BFD" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FE5DC6F091487BFD" box="[407,528,403,429]" country="Mexico" latitude="18.464445" longLatPrecision="21" longitude="-98.99166" name="Ajuchitlan" pageId="52" pageNumber="253" stateProvince="Morelos">Ajuchitlán</location>
, [
<quantity id="9769FCE6FF90B067FDE8C6F091367BFD" box="[546,622,403,429]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="52" pageNumber="253" unit="m" value="990.0">
<elevation id="DBBCB630FF90B067FDE8C6F091367BFD" box="[546,622,403,429]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="52" pageNumber="253" unit="m" value="990.0">990 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF90B067FDB0C6F1905B7BFD" box="[634,771,402,429]" degrees="18" direction="north" minutes="27" orientation="latitude" pageId="52" pageNumber="253" precision="15" seconds="52" value="18.464445">18°2752”N</geoCoordinate>
<geoCoordinate id="35A537C4FF90B067FCC0C6F190C67BFD" box="[778,926,402,429]" degrees="98" direction="west" minutes="59" orientation="longitude" pageId="52" pageNumber="253" precision="15" seconds="30" value="-98.99166">98°5930”W</geoCoordinate>
],
<date id="242F77C3FF90B067FC7BC6F0977E7BFD" box="[945,1062,403,429]" pageId="52" pageNumber="253" value="1996-09-06">
<collectingDate id="346B8E2BFF90B067FC7BC6F0977E7BFD" box="[945,1062,403,429]" pageId="52" pageNumber="253" value="1996-09-06">6/IX/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FF90B067FBF8C6F097FB7BFD" box="[1074,1187,403,429]" pageId="52" pageNumber="253">R. Brooks</collectorName>
(
<collectionCode id="3680C9C6FF90B067FB78C6F096057BFD" box="[1202,1373,403,429]" pageId="52" pageNumber="253">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFF90B067FAAEC6F096DD7BFD" box="[1380,1413,403,429]" count="1" pageId="52" pageNumber="253" type="female">1♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="502E5103FF90B067FF0DC6D5970E7969" blockId="52.[151,1437,151,1149]" pageId="52" pageNumber="253">
<emphasis id="62E58D11FF90B067FF0DC6D591617B80" bold="true" box="[199,569,438,464]" pageId="52" pageNumber="253">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFF90B067FD8BC6D492DC7BA5" collectingDate="1924-08-10" collectionCode="KUNHM-ENT" collectorName="Pollug" country="United States of America" location="Baboquivari Mts." pageId="52" pageNumber="253" specimenCount="1" specimenCount-male="1" stateProvince="Arizona">
<emphasis id="62E58D11FF90B067FD8BC6D490CD7B80" bold="true" box="[577,917,438,465]" pageId="52" pageNumber="253">
<collectingCountry id="28861193FF90B067FD8BC6D4907A7B80" box="[577,802,438,465]" name="United States of America" pageId="52" pageNumber="253">UNITED STATES</collectingCountry>
:
<collectingRegion id="92559FE1FF90B067FCE7C6D590D77B80" box="[813,911,438,464]" country="United States of America" name="Arizona" pageId="52" pageNumber="253">Arizona</collectingRegion>
,
</emphasis>
<location id="554E07D8FF90B067FC57C6D4973B7B81" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FC57C6D4973B7B81" box="[925,1123,439,465]" country="United States of America" name="Baboquivari Mts." pageId="52" pageNumber="253" stateProvince="Arizona">Baboquivari Mts.</location>
<date id="242F77C3FF90B067FBA1C6D496597B81" box="[1131,1281,439,465]" pageId="52" pageNumber="253" value="1924-08-10">
<collectingDate id="346B8E2BFF90B067FBA1C6D496597B81" box="[1131,1281,439,465]" pageId="52" pageNumber="253" value="1924-08-10">10/VIII/1924</collectingDate>
</date>
, O. C.
<collectorName id="FD6434D5FF90B067FA99C6D496C47B81" box="[1363,1436,439,465]" pageId="52" pageNumber="253">Pollug</collectorName>
(
<collectionCode id="3680C9C6FF90B067FF54C6B892117BA5" box="[158,329,475,501]" pageId="52" pageNumber="253">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFF90B067FE9FC6B892227BA5" box="[341,378,475,501]" count="1" pageId="52" pageNumber="253" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF90B067FE5EC6B9910F7849" collectingDate="1946-06-03" collectionCode="BBSL, KUNHM-ENT" collectorName="J. &amp; D. Pallister" country="Mexico" elevation="684" latitude="17.944443" location="Mexcala" longLatPrecision="21" longitude="-99.601944" pageId="52" pageNumber="253" specimenCount="2" specimenCount-male="2" stateProvince="Guerrero">
<emphasis id="62E58D11FF90B067FE5EC6B991CE7BA4" bold="true" box="[404,662,474,500]" pageId="52" pageNumber="253">
<collectingCountry id="28861193FF90B067FE5EC6B991577BA4" box="[404,527,474,500]" name="Mexico" pageId="52" pageNumber="253">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF90B067FDD5C6B991C97BA4" box="[543,657,474,500]" country="Mexico" name="Guerrero" pageId="52" pageNumber="253">Guerrero</collectingRegion>
,
</emphasis>
<location id="554E07D8FF90B067FD68C6B8905D7BA5" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FD68C6B8905D7BA5" box="[674,773,475,501]" country="Mexico" latitude="17.944443" longLatPrecision="21" longitude="-99.601944" name="Mexcala" pageId="52" pageNumber="253" stateProvince="Guerrero">Mexcala</location>
, [
<quantity id="9769FCE6FF90B067FCD6C6B890347BA5" box="[796,876,475,501]" metricMagnitude="2" metricUnit="m" metricValue="6.84" pageId="52" pageNumber="253" unit="m" value="684.0">
<elevation id="DBBCB630FF90B067FCD6C6B890347BA5" box="[796,876,475,501]" metricMagnitude="2" metricUnit="m" metricValue="6.84" pageId="52" pageNumber="253" unit="m" value="684.0">684 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF90B067FCB6C6B9975E7BA5" box="[892,1030,474,501]" degrees="17" direction="north" minutes="56" orientation="latitude" pageId="52" pageNumber="253" precision="15" seconds="40" value="17.944443">17°5640”N</geoCoordinate>
<geoCoordinate id="35A537C4FF90B067FBDBC6B897FE7BA5" box="[1041,1190,475,501]" degrees="99" direction="west" minutes="36" orientation="longitude" pageId="52" pageNumber="253" precision="15" seconds="07" value="-99.601944">99°3607”W</geoCoordinate>
],
<date id="242F77C3FF90B067FB76C6B8966A7BA5" box="[1212,1330,475,501]" pageId="52" pageNumber="253" value="1946-06-03">
<collectingDate id="346B8E2BFF90B067FB76C6B8966A7BA5" box="[1212,1330,475,501]" pageId="52" pageNumber="253" value="1946-06-03">3/VI/1946</collectingDate>
</date>
,
<collectorName id="FD6434D5FF90B067FA88C6B893AB7849" pageId="52" pageNumber="253">J. &amp; D. Pallister</collectorName>
(
<collectionCode id="3680C9C6FF90B067FECAC69C921F7849" box="[256,327,511,537]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="52" pageNumber="253" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFF90B067FE86C69C92287849" box="[332,368,511,537]" count="1" pageId="52" pageNumber="253" type="male">1♂</specimenCount>
,
<collectionCode id="3680C9C6FF90B067FEB1C69C917C7849" box="[379,548,511,537]" pageId="52" pageNumber="253">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFF90B067FDE3C69C91157849" box="[553,589,511,537]" count="1" pageId="52" pageNumber="253" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF90B067FDABC69C928F786D" collectionCode="KUNHM-ENT" collectorName="U. Kans. Mex. Exped." country="Mexico" elevation="777" latitude="17.518888" location="Chilpancingo" longLatPrecision="21" longitude="-99.545555" pageId="52" pageNumber="253" specimenCount="5" specimenCount-male="5" stateProvince="Guerrero">
<location id="554E07D8FF90B067FDABC69C91A37849" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FDABC69C91A37849" box="[609,763,511,537]" country="Mexico" latitude="17.518888" longLatPrecision="21" longitude="-99.545555" name="Chilpancingo" pageId="52" pageNumber="253" stateProvince="Guerrero">Chilpancingo</location>
,
<quantity id="9769FCE6FF90B067FCCEC69C901D7849" box="[772,837,511,537]" metricMagnitude="4" metricUnit="m" metricValue="2.7358848" pageId="52" pageNumber="253" unit="mi" value="17.0">17mi.</quantity>
N,
<quantity id="9769FCE6FF90B067FCAFC69D90EE7849" box="[869,950,510,537]" metricMagnitude="2" metricUnit="m" metricValue="7.7724" pageId="52" pageNumber="253" unit="ft" value="2550.0">
<elevation id="DBBCB630FF90B067FCAFC69D90EE7849" box="[869,950,510,537]" metricMagnitude="2" metricUnit="m" metricValue="7.7724" pageId="52" pageNumber="253" unit="ft" value="2550.0">2550 ft</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FF90B067FC0DC69C970A7849" box="[967,1106,511,537]" degrees="17" direction="north" minutes="31" orientation="latitude" pageId="52" pageNumber="253" precision="15" seconds="08" value="17.518888">17°3108”N</geoCoordinate>
<geoCoordinate id="35A537C4FF90B067FB9DC69C97B47849" box="[1111,1260,511,537]" degrees="99" direction="west" minutes="32" orientation="longitude" pageId="52" pageNumber="253" precision="15" seconds="44" value="-99.545555">99°3244”W</geoCoordinate>
],
<collectorName id="FD6434D5FF90B067FB36C69C93BE786D" pageId="52" pageNumber="253">U. Kans. Mex. Exped.</collectorName>
(
<collectionCode id="3680C9C6FF90B067FF3FC54092F8786D" box="[245,416,547,573]" pageId="52" pageNumber="253">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFF90B067FE62C5419296786D" box="[424,462,546,573]" count="5" pageId="52" pageNumber="253" type="male">5♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF90B067FE2EC54191C47831" collectingDate="2014-04-12" collectionCode="UAQE" collectorName="la Cienaga &amp; E. Razo &amp; A. Munoz" country="Mexico" elevation="1704" latitude="20.260834" location="Camino" longLatPrecision="21" longitude="-104.06722" municipality="Tecolotlan" pageId="52" pageNumber="253" specimenCount="1" specimenCount-male="1" stateProvince="Jalisco">
<emphasis id="62E58D11FF90B067FE2EC5419164786C" bold="true" box="[484,572,546,572]" pageId="52" pageNumber="253">
<collectingRegion id="92559FE1FF90B067FE2EC541916F786C" box="[484,567,546,572]" country="Mexico" name="Jalisco" pageId="52" pageNumber="253">Jalisco</collectingRegion>
,
</emphasis>
<collectingMunicipality id="B04ACB79FF90B067FD8EC54091E5786D" box="[580,701,547,573]" pageId="52" pageNumber="253">Tecolotlán</collectingMunicipality>
,
<location id="554E07D8FF90B067FD02C5409079786D" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FD02C5409079786D" box="[712,801,547,573]" country="Mexico" latitude="20.260834" longLatPrecision="21" longitude="-104.06722" municipality="Tecolotlan" name="Camino" pageId="52" pageNumber="253" stateProvince="Jalisco">Camino</location>
a
<collectorName id="FD6434D5FF90B067FCF4C54090EE786D" box="[830,950,547,573]" pageId="52" pageNumber="253">la Ciénaga</collectorName>
BTC,
<quantity id="9769FCE6FF90B067FBCEC5409704786D" box="[1028,1116,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.704" pageId="52" pageNumber="253" unit="m" value="1704.0">
<elevation id="DBBCB630FF90B067FBCEC5409704786D" box="[1028,1116,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.704" pageId="52" pageNumber="253" unit="m" value="1704.0">1704 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF90B067FBA3C54197AA786D" box="[1129,1266,546,573]" degrees="20" direction="north" minutes="15" orientation="latitude" pageId="52" pageNumber="253" precision="15" seconds="39" value="20.260834">20°1539”N</geoCoordinate>
<geoCoordinate id="35A537C4FF90B067FB30C54096C0786D" box="[1274,1432,547,573]" degrees="104" direction="west" minutes="04" orientation="longitude" pageId="52" pageNumber="253" precision="15" seconds="02" value="-104.06722">104°0402”W</geoCoordinate>
,
<date id="242F77C3FF90B067FF5DC52492437831" box="[151,283,583,609]" pageId="52" pageNumber="253" value="2014-04-12">
<collectingDate id="346B8E2BFF90B067FF5DC52492437831" box="[151,283,583,609]" pageId="52" pageNumber="253" value="2014-04-12">12/IV/2014</collectingDate>
</date>
,
<collectorName id="FD6434D5FF90B067FEEFC52492247831" box="[293,380,583,609]" pageId="52" pageNumber="253">E. Razo</collectorName>
&amp;
<collectorName id="FD6434D5FF90B067FE51C52491537831" box="[411,523,583,609]" pageId="52" pageNumber="253">A. Muñoz</collectorName>
(
<collectionCode id="3680C9C6FF90B067FDD2C52491307831" box="[536,616,583,609]" pageId="52" pageNumber="253">UAQE</collectionCode>
<specimenCount id="46979A8AFF90B067FDA7C52491CA7831" box="[621,658,583,609]" count="1" pageId="52" pageNumber="253" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF90B067FD6CC525907C78D5" collectingDate="1961-06-26" collectionCode="KUNHM-ENT" collectorName="U. Kans. Mex. Exped." country="Mexico" latitude="16.385555" location="Tehuantepec" longLatPrecision="21" longitude="-95.34361" pageId="52" pageNumber="253" specimenCount="15" specimenCount-female="15" stateProvince="Oaxaca">
<emphasis id="62E58D11FF90B067FD6CC525905F7830" bold="true" box="[678,775,582,608]" pageId="52" pageNumber="253">
<collectingRegion id="92559FE1FF90B067FD6CC525905A7830" box="[678,770,582,608]" country="Mexico" name="Oaxaca" pageId="52" pageNumber="253">Oaxaca</collectingRegion>
,
</emphasis>
<location id="554E07D8FF90B067FCC6C52490C47831" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FCC6C52490C47831" box="[780,924,583,609]" country="Mexico" latitude="16.385555" longLatPrecision="21" longitude="-95.34361" name="Tehuantepec" pageId="52" pageNumber="253" stateProvince="Oaxaca">Tehuantepec</location>
,
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NW, 700, [
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, 3min.n.w., [
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, [
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NE, [
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],
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,
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,
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(EBCh
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)
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<materialsCitation id="E0F95B5EFF90B067FBC8C5B490997945" collectingDate="1970-09-18" collectionCode="BBSL" collectorName="R. M. Bohart Colr" country="Mexico" elevation="1739" latitude="21.578056" location="Jalpa" longLatPrecision="21" longitude="-102.968056" pageId="52" pageNumber="253" specimenCount="2" specimenCount-male="2" stateProvince="Zacatecas">
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<collectingRegion id="92559FE1FF90B067FBC8C5B4972278A1" box="[1026,1146,727,753]" country="Mexico" name="Zacatecas" pageId="52" pageNumber="253">Zacatecas</collectingRegion>
,
</emphasis>
<location id="554E07D8FF90B067FB41C5B4979F78A1" LSID="urn:lsid:plazi:treatment:D838E015FF95B067FF5DC3FA97327E2D:554E07D8FF90B067FB41C5B4979F78A1" box="[1163,1223,727,753]" country="Mexico" latitude="21.578056" longLatPrecision="21" longitude="-102.968056" name="Jalpa" pageId="52" pageNumber="253" stateProvince="Zacatecas">Jalpa</location>
, 10miS, [
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,
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],
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(
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)
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,
<materialsCitation id="E0F95B5EFF90B067FC06C59896CF7945" box="[972,1431,763,789]" collectingDate="1970-09-17" collectionCode="BBSL" collectorName="Bohart" country="Mexico" pageId="52" pageNumber="253" specimenCount="1" specimenCount-male="1" stateProvince="Zacatecas">
<date id="242F77C3FF90B067FC06C59897087945" box="[972,1104,763,789]" pageId="52" pageNumber="253" value="1970-09-17">
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</date>
, GE, RM
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(
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)
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,
<collectorName id="FD6434D5FF90B067FEC0C47C923D7969" box="[266,357,799,825]" pageId="52" pageNumber="253">La Joya</collectorName>
,
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,
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</paragraph>
</subSubSection>
<subSubSection id="188B0288FF90B067FF0DC421962F79F5" pageId="52" pageNumber="253" type="distribution">
<paragraph id="502E5103FF90B067FF0DC421962F79F5" blockId="52.[151,1437,151,1149]" pageId="52" pageNumber="253">
<emphasis id="62E58D11FF90B067FF0DC4219239790C" bold="true" box="[199,353,834,860]" pageId="52" pageNumber="253">Distribution.</emphasis>
This species is distributed along the mountain range of the Sierra Madre Occidental in Zacatecas, the Sierra Madre del Sur in Oaxaca, and central Mexico in Morelos (
<figureCitation id="C8AA4D86FF90B067FC09C404974379D1" box="[963,1051,871,897]" captionStart="FIGURE 30" captionStartId="49.[151,250,1071,1095]" captionTargetBox="[159,1428,157,1045]" captionTargetId="figure-18@49.[151,1436,149,1045]" captionTargetPageId="49" captionText="FIGURE 30. Known distribution records for Heriades viridiana sp. nov." pageId="52" pageNumber="253">Fig. 30</figureCitation>
). A male from the
<collectingCountry id="28861193FF90B067FB36C40496C079D1" box="[1276,1432,871,897]" name="United States of America" pageId="52" pageNumber="253">United States</collectingCountry>
,
<collectingRegion id="92559FE1FF90B067FF5DC4E893AC79F5" box="[151,244,907,933]" country="United States of America" name="Arizona" pageId="52" pageNumber="253">Arizona</collectingRegion>
, extends the distribution of the species out of the Neotropical region. Altitudinal range:
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.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF90B067FF0DC4CC927B7E41" pageId="52" pageNumber="253" type="etymology">
<paragraph id="502E5103FF90B067FF0DC4CC927B7E41" blockId="52.[151,1437,151,1149]" pageId="52" pageNumber="253">
<emphasis id="62E58D11FF90B067FF0DC4CC92167999" bold="true" box="[199,334,943,969]" pageId="52" pageNumber="253">Etymology.</emphasis>
This species is dedicated to M. Sc. Viridiana Vega Badillo, taxonomist of
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and academic technician in the entomological collection IEXA of INECOL in Xalapa,
<collectingRegion id="92559FE1FF90B067FC20C4B0970979BD" box="[1002,1105,979,1005]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="52" pageNumber="253">Veracruz</collectingRegion>
, for her contributions to the entomology.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF90B067FF0DC37997327E2D" pageId="52" pageNumber="253" type="discussion">
<paragraph id="502E5103FF90B067FF0DC37997327E2D" blockId="52.[151,1437,151,1149]" pageId="52" pageNumber="253">
<emphasis id="62E58D11FF90B067FF0DC37992177E64" bold="true" box="[199,335,1050,1076]" pageId="52" pageNumber="253">Comments.</emphasis>
Color of hairs in clypeus and paraocular area of males vary, from whitish to orange, males of this species are similar to
<taxonomicName id="97912A80FF90B067FE4DC35C915D7E09" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[391,517,1087,1113]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="briceida" status="sp. nov.">
<emphasis id="62E58D11FF90B067FE4DC35C915D7E09" box="[391,517,1087,1113]" italics="true" pageId="52" pageNumber="253">H. briceida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF90B067FDC6C35C913A7E09" box="[524,610,1087,1113]" pageId="52" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
but can be recognized by having a pointy top of process on S
<quantity id="9769FCE6FF90B067FAC1C35C966B7E09" box="[1291,1331,1087,1113]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="52" pageNumber="253" unit="in" value="1.0">1 in</quantity>
posterior view, and whiter hairs on clypeus. About females see comments on
<taxonomicName id="97912A80FF90B067FC47C30097547E2D" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[909,1036,1123,1149]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="253" phylum="Arthropoda" rank="species" species="briceida" status="sp. nov.">
<emphasis id="62E58D11FF90B067FC47C30097547E2D" box="[909,1036,1123,1149]" italics="true" pageId="52" pageNumber="253">H. briceida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF90B067FBDEC30097327E2D" box="[1044,1130,1123,1149]" pageId="52" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,634 @@
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<mods:title id="5E0798298BDDC085289F8E91A28331C5">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<paragraph id="502E5103FFA8B05FFF5DC2E1903E7FCD" blockId="12.[151,870,1410,1473]" box="[151,870,1410,1437]" pageId="12" pageNumber="213">
<heading id="0B66E66FFFA8B05FFF5DC2E1903E7FCD" bold="true" box="[151,870,1410,1437]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<emphasis id="62E58D11FFA8B05FFF5DC2E1903E7FCD" bold="true" box="[151,870,1410,1437]" pageId="12" pageNumber="213">
<taxonomicName id="97912A80FFA8B05FFF5DC2E190597FCC" authority="Rojas-Arias, Griswold &amp; Ayala." authorityName="Rojas-Arias, Griswold &amp; Ayala." authorityYear="2025" box="[151,769,1410,1437]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="briceida" status="sp. nov.">
<emphasis id="62E58D11FFA8B05FFF5DC2E192367FCD" bold="true" box="[151,366,1410,1437]" italics="true" pageId="12" pageNumber="213">Heriades briceida</emphasis>
Rojas-Arias, Griswold &amp; Ayala.
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<subSubSection id="188B0288FFA8B05FFF5DC2C4921A7F91" box="[151,322,1447,1473]" pageId="12" pageNumber="213" type="description">
<paragraph id="502E5103FFA8B05FFF5DC2C4921A7F91" blockId="12.[151,870,1410,1473]" box="[151,322,1447,1473]" pageId="12" pageNumber="213">
(
<figureCitation id="C8AA4D86FFA8B05FFF55C2C4925B7F91" box="[159,259,1447,1473]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="12" pageNumber="213">Figs. 2D</figureCitation>
,
<figureCitation id="C8AA4D86FFA8B05FFEC5C2C492637F91" box="[271,315,1447,1473]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="13.[151,250,1098,1122]" captionStartId-1="15.[151,250,1943,1967]" captionTargetBox-0="[159,1428,187,1064]" captionTargetBox-1="[163,1410,183,1910]" captionTargetId-0="figure-18@13.[151,1436,179,1072]" captionTargetId-1="figure-18@15.[163,1425,179,1917]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionText-0="FIGURE 6. Known distribution records for Heriades briceida sp. nov." captionText-1="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="12" pageNumber="213">67</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFA8B05FFF5DC28D92547DC5" pageId="12" pageNumber="213" type="diagnosis">
<paragraph id="502E5103FFA8B05FFF5DC28D92547DC5" blockId="12.[151,1437,1518,1977]" pageId="12" pageNumber="213">
<emphasis id="62E58D11FFA8B05FFF5DC28D924A7C58" bold="true" box="[151,274,1518,1544]" pageId="12" pageNumber="213">Diagnosis.</emphasis>
Ocelli medium-sized (
<figureCitation id="C8AA4D86FFA8B05FFDD3C28C912B7C59" box="[537,627,1519,1545]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="12" pageNumber="213">Fig. 3B</figureCitation>
); tergal fasciae with row of thick, white hairs (
<figureCitation id="C8AA4D86FFA8B05FFB5EC28C97BC7C59" box="[1172,1252,1519,1545]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="12" pageNumber="213">Fig. 3I</figureCitation>
).
<emphasis id="62E58D11FFA8B05FFB36C28C966E7C59" box="[1276,1334,1519,1545]" italics="true" pageId="12" pageNumber="213">Male</emphasis>
: clypeus with plumose, dense, thin, white hairs directed medially, all with similar length (
<figureCitation id="C8AA4D86FFA8B05FFBDDC17097287C7D" box="[1047,1136,1555,1581]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="12" pageNumber="213">Fig. 7A</figureCitation>
); hairs on hypostomal area and fore tarsal segments larger than on clypeus; posterior margin of S1 truncate (
<figureCitation id="C8AA4D86FFA8B05FFBEDC15497327C01" box="[1063,1130,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="12" pageNumber="213">Fig. 2</figureCitation>
Db); disc of S1 with strong process, in profile, with sides projected forming an acute angle (
<figureCitation id="C8AA4D86FFA8B05FFCAEC13890FE7C25" box="[868,934,1627,1653]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="12" pageNumber="213">Fig. 2</figureCitation>
Da), and in posterior view with emargination on top, two rounded corners, and white hairs on posterior surface (
<figureCitation id="C8AA4D86FFA8B05FFC6EC11C90B37CC9" box="[932,1003,1663,1689]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="12" pageNumber="213">Fig. 2</figureCitation>
Dc); posterior margin of S5 with two subtriangular lobes, and one row of capitate hairs directed laterally (
<figureCitation id="C8AA4D86FFA8B05FFC5AC1C090BF7CED" box="[912,999,1699,1725]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="12" pageNumber="213">Fig. 7E</figureCitation>
); posterior margin of S6 with two lines of a few capitate, long, thick, hooked hairs, directed medially, and a few simple, long hairs medially (
<figureCitation id="C8AA4D86FFA8B05FFAFDC1A496D77CB1" box="[1335,1423,1735,1761]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="12" pageNumber="213">Fig. 7F</figureCitation>
).
<emphasis id="62E58D11FFA8B05FFF5DC18893B37D55" box="[151,235,1771,1797]" italics="true" pageId="12" pageNumber="213">Female</emphasis>
: outer surface of mandible without tubercle; apical margin of clypeus with five or more tubercles, or several not well-defined tubercles; frons with plumose, white hairs (
<figureCitation id="C8AA4D86FFA8B05FFCF6C06C90CC7D79" box="[828,916,1807,1833]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="12" pageNumber="213">Fig. 7C</figureCitation>
); T3 with medium-sized (0.3 times DOP as on scutum), and sparse punctures (0.5 times DOP (
<figureCitation id="C8AA4D86FFA8B05FFD7BC05090517D1D" box="[689,777,1843,1869]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="12" pageNumber="213">Fig. 7D</figureCitation>
) similar to
<taxonomicName id="97912A80FFA8B05FFC41C050974D7D1D" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[907,1045,1843,1869]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="viridiana">
<emphasis id="62E58D11FFA8B05FFC41C050974D7D1D" box="[907,1045,1843,1869]" italics="true" pageId="12" pageNumber="213">H. viridiana</emphasis>
</taxonomicName>
sp nov. but differing in not having clypeus with softer emargination, without tubercles in middle of apical margin, and mesepisternum with denser punctures.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFA8B05DFF0DC0FD96407FCD" lastPageId="14" lastPageNumber="215" pageId="12" pageNumber="213" type="description">
<paragraph id="502E5103FFA8B05FFF0DC0FD97A87DE9" blockId="12.[151,1437,1518,1977]" box="[199,1264,1950,1977]" pageId="12" pageNumber="213">
<emphasis id="62E58D11FFA8B05FFF0DC0FD92FC7DE9" bold="true" box="[199,420,1950,1977]" pageId="12" pageNumber="213">
Description.
<emphasis id="62E58D11FFA8B05FFEAAC0FD92C37DE9" bold="true" box="[352,411,1950,1977]" italics="true" pageId="12" pageNumber="213">Male</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FFA8B05FFDF6C0FD91CA7DE9" box="[572,658,1950,1977]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="12" pageNumber="213" unit="mm" value="5.9">5.9 mm</quantity>
(5.97.3, n=4); forewing length
<quantity id="9769FCE6FFA8B05FFBCBC0FC970F7DE9" box="[1025,1111,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="213" unit="mm" value="3.4">3.4 mm</quantity>
(3.44, n=4).
</paragraph>
<caption id="04EE018BFFA9B05EFF5DC329903F7E32" box="[151,871,1098,1122]" pageId="13" pageNumber="214" startId="13.[151,250,1098,1122]" targetBox="[159,1428,187,1064]" targetPageId="13" targetType="figure">
<paragraph id="502E5103FFA9B05EFF5DC329903F7E32" blockId="13.[151,871,1098,1122]" box="[151,871,1098,1122]" pageId="13" pageNumber="214">
<emphasis id="62E58D11FFA9B05EFF5DC329924B7E32" bold="true" box="[151,275,1098,1122]" pageId="13" pageNumber="214">FIGURE 6.</emphasis>
Known distribution records for
<taxonomicName id="97912A80FFA9B05EFD90C32990497E32" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[602,785,1098,1122]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="briceida" status="sp. nov.">
<emphasis id="62E58D11FFA9B05EFD90C32990497E32" box="[602,785,1098,1122]" italics="true" pageId="13" pageNumber="214">Heriades briceida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA9B05EFCD2C328903F7E32" box="[792,871,1099,1122]" pageId="13" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="502E5103FFA9B05EFF0DC3F3938F7CF2" blockId="13.[151,1437,1168,2023]" pageId="13" pageNumber="214">
<emphasis id="62E58D11FFA9B05EFF0DC3F3925C7EFA" box="[199,260,1168,1194]" italics="true" pageId="13" pageNumber="214">Head</emphasis>
. Length
<quantity id="9769FCE6FFA9B05EFEA6C3F3929B7EFB" box="[364,451,1168,1195]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="214" unit="mm" value="1.6">1.6 mm</quantity>
(1.61.8, n=4), width
<quantity id="9769FCE6FFA9B05EFD0CC3F390467EFB" box="[710,798,1168,1195]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="214" unit="mm" value="1.7">1.7 mm</quantity>
(1.71.9, n=4); eyes about 3.6 times longer than broad, 1.8 times wider than gena in profile, lower interocular distance
<quantity id="9769FCE6FFA9B05EFCA2C3D790E77E9F" box="[872,959,1204,1231]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="214" unit="mm" value="0.9">0.9 mm</quantity>
(0.80.9, n=4), upper interocular distance
<quantity id="9769FCE6FFA9B05EFF5DC3BB93817EA3" box="[151,217,1240,1267]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="214" unit="mm" value="1.0">1 mm</quantity>
(11.1, n=4); interantennal distance 1.9 times antennocular distance; lateral ocelli medium-sized, 2.2 times DOP on interocellar area, median ocellus slightly smaller; interocellar distance 2.2 times OD, ocellocular distance 1.9 times OD; OPR 2.2 times OD; labrum with basal, medial, low elevation; clypeus 1.7 times broader than long, apical margin without emargination; juxtantennal carina absent but strongly angulated; scape black, length 0.7 times alveolocellar distance, 2.3 times longer than broad; pedicel black about 1.4 times longer than F1 and about 0.6 time longer than F2; flagellum black; F1 slightly broader than long; F2 1.3 times longer F1; F11 2.4 times longer than broad.
<emphasis id="62E58D11FFA9B05EFEA0C2D292B17F9A" box="[362,489,1457,1482]" italics="true" pageId="13" pageNumber="214">Mesosoma.</emphasis>
Scutum 2.6 times longer than scutellum, 3.3 times longer than tegula, and 4.8 longer than axilla; intertegular distance
<quantity id="9769FCE6FFA9B05EFDC0C2B791397FBF" box="[522,609,1492,1519]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="214" unit="mm" value="1.3">1.3 mm</quantity>
(1.21.3, n=4); metanotum with transverse ridge, in posterior view ridge straight; propodeum with middle pits,
<quantity id="9769FCE6FFA9B05EFD83C29B91F67C43" box="[585,686,1528,1555]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="214" unit="mm" value="0.08">0.08 mm</quantity>
long (
<figureCitation id="C8AA4D86FFA9B05EFD3DC29B900A7C42" box="[759,850,1528,1554]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="13" pageNumber="214">Fig. 3H</figureCitation>
); legs black.
<emphasis id="62E58D11FFA9B05EFC22C29A973C7C42" box="[1000,1124,1529,1554]" italics="true" pageId="13" pageNumber="214">Metasoma.</emphasis>
T1
<quantity id="9769FCE6FFA9B05EFB58C29B97B17C43" box="[1170,1257,1528,1555]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="214" unit="mm" value="1.5">1.5 mm</quantity>
broad (1.51.8, n=4). T3
<quantity id="9769FCE6FFA9B05EFECDC17F92397C67" box="[263,353,1564,1591]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="214" unit="mm" value="1.7">1.7 mm</quantity>
broad (1.71.9, n=4); posterior margin of T6 not projected; posterior margin of S1 truncate, subquadrangular, with large, shallow emargination (
<figureCitation id="C8AA4D86FFA9B05EFD2CC12390727C0A" box="[742,810,1600,1626]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="13" pageNumber="214">Fig. 2</figureCitation>
Db) sided by two rounded, small cusps, in profile with strong ventral process, in more than one half of distance from apex to base, posterior view with two cusps (
<figureCitation id="C8AA4D86FFA9B05EFAA5C10793FE7CF2" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="13" pageNumber="214">Fig. 2</figureCitation>
Dc).
</paragraph>
<paragraph id="502E5103FFA9B05DFF0DC1CF97EF7AA9" blockId="13.[151,1437,1168,2023]" lastBlockId="14.[151,1437,150,2013]" lastPageId="14" lastPageNumber="215" pageId="13" pageNumber="214">
<emphasis id="62E58D11FFA9B05EFF0DC1CF92097C96" box="[199,337,1708,1734]" italics="true" pageId="13" pageNumber="214">Sculpturing.</emphasis>
Clypeus with small (at base 0.4 times OD), contiguous, coarse punctures, getting smaller towards sides and apical margin; frons with punctures bigger, coarser and denser, and on frons in front of ocelli slightly smaller than on frons; vertex bigger than on supraclypeal area, with micropunctures between punctures; hypostomal area with punctures smaller than on clypeus; pronotum, anterior surface and dorsolateral angles with punctures bigger, less coarse than on clypeus; disc of scutum with punctures slightly smaller (0.5 times OD), sparser than on vertex (
<figureCitation id="C8AA4D86FFA9B05EFF55C00393A27D2A" box="[159,250,1888,1914]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="13" pageNumber="214">Fig. 7B</figureCitation>
); mesepisternum with punctures as on scutum with micropunctures between punctures; metepisternum with smaller punctures, as coarse and defined as on mesepisternum; T1 with (0.4 times OD) punctures as on scutum, becoming smaller towards posterior margin; T2 as on T1 but smaller; T3 with bigger, slightly sparser and equally coarse punctures than on T1, slightly rounded, separated by less than one third DOP, with micropunctures between punctures; T4 and T5 with bigger, denser and coarser than on T3; S1 with punctures smaller, sparser than on T3, in posterior margin with notably smaller, less dense punctation; sides of S2 with similar punctation than on T1; sides of S2 with bigger punctation than on S1, becoming smaller towards disc and posterior margin.
</paragraph>
<paragraph id="502E5103FFAAB05DFF0DC66092217B81" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="215">
<emphasis id="62E58D11FFAAB05DFF0DC66092687B4C" box="[199,304,259,284]" italics="true" pageId="14" pageNumber="215">Vestiture.</emphasis>
Clypeus with hairs directed medially; frons with plumose, thin, white hairs with similar length, directed dorsally (
<figureCitation id="C8AA4D86FFAAB05DFEA2C644929C7B11" box="[360,452,295,321]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="14" pageNumber="215">Fig. 7A</figureCitation>
); pronotal lobe on posterior area of carina with thin, white hairs; scutum and disc of mesepisternum with simple, short, golden on its entire surface; anterior and posterior margin of mesepisternum with few plumose, thin, short, scattered, white hairs; disc of metepisternum without appressed tuft thick, short hairs; tergal fasciae of thick hairs, present from T1 to T3; T4 to T6 with simple, thick, sparse, short, white hairs in punctures (
<figureCitation id="C8AA4D86FFAAB05DFED9C6D492347B81" box="[275,364,439,465]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="14" pageNumber="215">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="502E5103FFAAB05DFF0DC6B8918478F9" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="215">
<emphasis id="62E58D11FFAAB05DFF0DC6B892117BA5" box="[199,329,475,501]" italics="true" pageId="14" pageNumber="215">
<taxonomicName id="97912A80FFAAB05DFF0DC6B8921D7BA5" box="[199,325,475,501]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="14" pageNumber="215" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
S5 slightly bilobated with two subtriangular lobes on posterior margin, and a line of capitate hairs directed to sides, with transverse line of preapical hairs, thin and amber (
<figureCitation id="C8AA4D86FFAAB05DFC22C69C97197849" box="[1000,1089,511,537]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="14" pageNumber="215">Fig. 7E</figureCitation>
); S6 with acute apex, without blackened preapical margin, with two submarginal lines with few capitate, long, thick, hairs, with same length; genital capsule in dorsal view,
<quantity id="9769FCE6FFAAB05DFE3EC52491137831" box="[500,587,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="215" unit="mm" value="1.2">1.2 mm</quantity>
length and
<quantity id="9769FCE6FFAAB05DFD1BC52590707831" box="[721,808,582,609]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="215" unit="mm" value="0.5">0.5 mm</quantity>
width (2.5 times longer than wide) (
<figureCitation id="C8AA4D86FFAAB05DFB0DC52496457831" box="[1223,1309,583,609]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="14" pageNumber="215">Fig. 7F</figureCitation>
); gonobase
<quantity id="9769FCE6FFAAB05DFF5DC50893A778D5" box="[151,255,619,645]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="215" unit="mm" value="0.08">0.08 mm</quantity>
length, of 5.6 times wider than long, and gonocoxite
<quantity id="9769FCE6FFAAB05DFCBDC508908978D5" box="[887,977,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="215" unit="mm" value="0.4">0.4 mm</quantity>
length; gonostylus
<quantity id="9769FCE6FFAAB05DFB7DC508964878D5" box="[1207,1296,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="215" unit="mm" value="1.1">1.1 mm</quantity>
length with rounded apex; penis valve
<quantity id="9769FCE6FFAAB05DFE09C5EC914178F9" box="[451,537,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="215" unit="mm" value="0.7">0.7 mm</quantity>
length (
<figureCitation id="C8AA4D86FFAAB05DFDBFC5EC919778F9" box="[629,719,655,681]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="14" pageNumber="215">Fig. 7G</figureCitation>
).
</paragraph>
<paragraph id="502E5103FFAAB05DFF0DC5D1907D79BD" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="215">
<emphasis id="62E58D11FFAAB05DFF0DC5D1927F789D" bold="true" box="[199,295,690,717]" pageId="14" pageNumber="215">
<emphasis id="62E58D11FFAAB05DFF0DC5D19246789D" bold="true" box="[199,286,690,717]" italics="true" pageId="14" pageNumber="215">Female</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FFAAB05DFE0BC5D0914F789D" box="[449,535,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="14" pageNumber="215" unit="mm" value="6.1">6.1 mm</quantity>
(5.66.1, n=3); forewing length
<quantity id="9769FCE6FFAAB05DFC43C5D09087789D" box="[905,991,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="215" unit="mm" value="3.7">3.7 mm</quantity>
(3.33.8, n=3). Similar to male except:
<emphasis id="62E58D11FFAAB05DFF5DC5B4938C78A1" box="[151,212,727,753]" italics="true" pageId="14" pageNumber="215">Head</emphasis>
. Length
<quantity id="9769FCE6FFAAB05DFEFFC5B492D178A1" box="[309,393,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="215" unit="mm" value="1.6">1.6 mm</quantity>
(1.51.6, n=3), width
<quantity id="9769FCE6FFAAB05DFDB4C5B4918B78A1" box="[638,723,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="215" unit="mm" value="1.7">1.7 mm</quantity>
(1.61.8, n=3); eyes about 3.5 times longer than broad, 1.7 times wider than gena in profile, lower interocular distance
<quantity id="9769FCE6FFAAB05DFCCBC59890017945" box="[769,857,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="215" unit="mm" value="0.9">0.9 mm</quantity>
(0.80.9, n=3), upper interocular distance
<quantity id="9769FCE6FFAAB05DFA8FC59896C57945" box="[1349,1437,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="215" unit="mm" value="1.1">1.1 mm</quantity>
(11.1, n=3); interantennal distance 1.5 times antennocular distance; interocellar distance 2.5 times OD, ocellocular distance 2 times OD, OPR 2.2 times OD; dorsal margin of mandible with carina; clypeus with shallow emargination on all length and crenulated of apical margin; scape 3.4 times longer than broad; pedicel 2.9 times longer than broad, about 2 times longer than F1 and 2.4 longer than F2, remaining flagellomeres 1.8 times longer than first.
<emphasis id="62E58D11FFAAB05DFAD7C4E896CD79F4" box="[1309,1429,907,932]" italics="true" pageId="14" pageNumber="215">Mesosoma</emphasis>
. Scutum 2.8 times longer than scutellum; intertegular distance
<quantity id="9769FCE6FFAAB05DFC9BC4CC90FF7999" box="[849,935,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="215" unit="mm" value="1.4">1.4 mm</quantity>
(1.31.4, n=3).
<emphasis id="62E58D11FFAAB05DFB97C4CC978A7998" box="[1117,1234,943,968]" italics="true" pageId="14" pageNumber="215">Metasoma</emphasis>
T1
<quantity id="9769FCE6FFAAB05DFB35C4CD960D7999" box="[1279,1365,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="215" unit="mm" value="1.5">1.5 mm</quantity>
broad (1.21.5, n=3). T3
<quantity id="9769FCE6FFAAB05DFEA1C4B0929979BD" box="[363,449,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="215" unit="mm" value="1.8">1.8 mm</quantity>
broad (1.61.8, n=3) (
<figureCitation id="C8AA4D86FFAAB05DFD74C4B0904079BD" box="[702,792,979,1005]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="14" pageNumber="215">Fig. 7D</figureCitation>
).
</paragraph>
<paragraph id="502E5103FFAAB05DFF0DC49496747F05" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="215">
<emphasis id="62E58D11FFAAB05DFF0DC49492107E41" box="[199,328,1015,1041]" italics="true" pageId="14" pageNumber="215">Sculpturing</emphasis>
. Clypeus with medium-sized punctures (0.4 times OD), dense (0.3 DBP) and coarse punctation, with micropunctures between punctures; supraclypeal area with punctures smaller than base of clypeus, bigger, coarser, and equal DBP on frons, slightly smaller and sparser, with micropuntures between punctures on frons in front of ocelli; vertex sparser and slightly bigger than on clypeus; gena evenly punctate, slightly bigger and more DBP than on frons in front of ocelli; scutum with bigger and denser punctures than on vertex; rest of mesosoma as on scutum except: scutellum with bigger and sparser punctures; mesepisternum with punctures slightly angulated on ventral half; terga bright, microreticulate between punctures; T1 with punctures smaller (0.3 times OD), sparser (0.3 times DOP) than on scutum, with rounded shape; T2 with punctures slightly bigger than on T3; T3 with punctures 0.3 times DOP as on scutum, semicircular shaped, and sparser (0.5 times DOP); dorsal surface of T6 with smaller, denser, and more coarsely punctures than on T5; sterna with punctation smaller and sparser than on scutum (
<figureCitation id="C8AA4D86FFAAB05DFB0CC25896477F05" box="[1222,1311,1339,1365]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="14" pageNumber="215">Fig. 7D</figureCitation>
).
</paragraph>
<paragraph id="502E5103FFAAB05DFF0DC23C96407FCD" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="215">
<emphasis id="62E58D11FFAAB05DFF0DC23C92687F28" box="[199,304,1375,1400]" italics="true" pageId="14" pageNumber="215">Vestiture.</emphasis>
Clypeus with few plumose, short, white hairs on sides; frons without plumose hairs; gena with plumose, white hairs evenly distributed; disc of T4 to T6 without plumose hairs in punctures (
<figureCitation id="C8AA4D86FFAAB05DFB78C2E096527FCD" box="[1202,1290,1411,1437]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="14" pageNumber="215">Fig. 7C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFAAB05DFF0DC2C490E27D21" pageId="14" pageNumber="215" type="materials_examined">
<paragraph id="502E5103FFAAB05DFF0DC2C496707CC9" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="215">
<emphasis id="62E58D11FFAAB05DFF0DC2C492217F90" bold="true" box="[199,377,1446,1473]" pageId="14" pageNumber="215">Type material:</emphasis>
<materialsCitation id="E0F95B5EFFAAB05DFE48C2C491BB7FB5" collectingDate="1989-07-10" collectingDateMax="1989-07-14" collectingDateMin="1989-07-10" collectionCode="BBSL" collectorName="No. LR" country="Mexico" elevation="71" latitude="19.49639" location="Jalisco" longLatPrecision="21" longitude="-105.04611" pageId="14" pageNumber="215" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="62E58D11FFAAB05DFE48C2C490977F90" bold="true" box="[386,975,1446,1473]" pageId="14" pageNumber="215">
<typeStatus id="8F2AEFA1FFAAB05DFE48C2C492B77F91" box="[386,495,1447,1473]" pageId="14" pageNumber="215" type="holotype">Holotype</typeStatus>
male (
<collectorName id="FD6434D5FFAAB05DFD88C2C591C67F90" box="[578,670,1446,1472]" pageId="14" pageNumber="215">No. LR</collectorName>
0003):
<collectingCountry id="28861193FFAAB05DFD25C2C590327F90" box="[751,874,1446,1472]" name="Mexico" pageId="14" pageNumber="215">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFAAB05DFCB2C2C590937F90" box="[888,971,1446,1472]" country="Mexico" name="Jalisco" pageId="14" pageNumber="215">Jalisco</collectingRegion>
,
</emphasis>
Chamela, Est. Biol. [
<quantity id="9769FCE6FFAAB05DFB0DC2C4965D7F91" box="[1223,1285,1447,1473]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="215" unit="m" value="71.0">
<elevation id="DBBCB630FFAAB05DFB0DC2C4965D7F91" box="[1223,1285,1447,1473]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="215" unit="m" value="71.0">71 m</elevation>
</quantity>
,
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],
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</date>
, Griswold (
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)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFAAB05DFD3BC2A990157C59" collectingDate="1986-07-15" collectionCode="BBSL" collectorName="R. Ayala" country="Mexico" elevation="71" latitude="19.49639" location="Chamela" longLatPrecision="21" longitude="-105.04611" pageId="14" pageNumber="215" specimenCount="1" stateProvince="Jalisco" typeStatus="allotype">
<emphasis id="62E58D11FFAAB05DFD3BC2A997947FB4" bold="true" box="[753,1228,1482,1508]" pageId="14" pageNumber="215">
<typeStatus id="8F2AEFA1FFAAB05DFD3BC2A9900E7FB4" box="[753,854,1482,1508]" pageId="14" pageNumber="215" type="allotype">Allotype</typeStatus>
(LR0004):
<collectingCountry id="28861193FFAAB05DFC23C2A9973C7FB4" box="[1001,1124,1482,1508]" name="Mexico" pageId="14" pageNumber="215">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFAAB05DFBBEC2A9979F7FB4" box="[1140,1223,1482,1508]" country="Mexico" name="Jalisco" pageId="14" pageNumber="215">Jalisco</collectingRegion>
,
</emphasis>
<location id="554E07D8FFAAB05DFB12C2A896667FB5" LSID="urn:lsid:plazi:treatment:D838E015FFA8B043FF5DC2E1973D7BD8:554E07D8FFAAB05DFB12C2A896667FB5" box="[1240,1342,1483,1509]" country="Mexico" latitude="19.49639" longLatPrecision="21" longitude="-105.04611" name="Chamela" pageId="14" pageNumber="215" stateProvince="Jalisco">Chamela</location>
, [
<quantity id="9769FCE6FFAAB05DFA9CC2A896CF7FB5" box="[1366,1431,1483,1509]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="215" unit="m" value="71.0">
<elevation id="DBBCB630FFAAB05DFA9CC2A896CF7FB5" box="[1366,1431,1483,1509]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="215" unit="m" value="71.0">71 m</elevation>
</quantity>
,
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],
<date id="242F77C3FFAAB05DFE2FC28D912A7C59" box="[485,626,1518,1545]" pageId="14" pageNumber="215" value="1986-07-15">
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</date>
,
<collectorName id="FD6434D5FFAAB05DFD48C28C91BD7C59" box="[642,741,1519,1545]" pageId="14" pageNumber="215">R. Ayala</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFD33C28C901D7C59" box="[761,837,1519,1545]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="14" pageNumber="215" type="Museum">BBSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFAAB05DFC96C28C91957C01" collectingDate="3 / VII / 1984, 11 / VII / 1985, 17 / X / 1985, 4 - 15 / VII / 1986" collectingDateMax="8 / VIII / 1985, 26 - 30 / IX / 1985" collectingDateMin="3 / VII / 1984, 11 / VII / 1985, 17 / X / 1985, 4 - 15 / VII / 1986" collectionCode="BBSL" collectorName="R. Ayala &amp; F. D. Parker &amp; Griswold" country="Mexico" elevation="71" latitude="19.49639" location="Chamela" longLatPrecision="21" longitude="-105.04611" pageId="14" pageNumber="215" specimenCount="13" specimenCount-female="5" specimenCount-male="8" stateProvince="Jalisco" typeStatus="paratype">
<emphasis id="62E58D11FFAAB05DFC96C28C97957C58" bold="true" box="[860,1229,1518,1545]" pageId="14" pageNumber="215">
<typeStatus id="8F2AEFA1FFAAB05DFC96C28C90807C59" box="[860,984,1519,1545]" pageId="14" pageNumber="215" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="28861193FFAAB05DFC22C28D973B7C58" box="[1000,1123,1518,1544]" name="Mexico" pageId="14" pageNumber="215">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFAAB05DFBB9C28D97907C58" box="[1139,1224,1518,1544]" country="Mexico" name="Jalisco" pageId="14" pageNumber="215">Jalisco</collectingRegion>
:
</emphasis>
<location id="554E07D8FFAAB05DFB13C28C96677C59" LSID="urn:lsid:plazi:treatment:D838E015FFA8B043FF5DC2E1973D7BD8:554E07D8FFAAB05DFB13C28C96677C59" box="[1241,1343,1519,1545]" country="Mexico" latitude="19.49639" longLatPrecision="21" longitude="-105.04611" name="Chamela" pageId="14" pageNumber="215" stateProvince="Jalisco">Chamela</location>
, [
<quantity id="9769FCE6FFAAB05DFA9CC28C96CF7C59" box="[1366,1431,1519,1545]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="215" unit="m" value="71.0">
<elevation id="DBBCB630FFAAB05DFA9CC28C96CF7C59" box="[1366,1431,1519,1545]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="215" unit="m" value="71.0">71 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFAAB05DFF5DC17092787C7D" box="[151,288,1555,1581]" degrees="19" direction="north" minutes="29" orientation="latitude" pageId="14" pageNumber="215" precision="15" seconds="47" value="19.49639">19°2947”N</geoCoordinate>
<geoCoordinate id="35A537C4FFAAB05DFEE1C17192957C7D" box="[299,461,1554,1581]" degrees="105" direction="west" minutes="02" orientation="longitude" pageId="14" pageNumber="215" precision="15" seconds="46" value="-105.04611">105°0246”W</geoCoordinate>
],
<collectingDate id="346B8E2BFFAAB05DFE29C17097617C7D" box="[483,1081,1554,1581]" isEnumeration="true" pageId="14" pageNumber="215">
<date id="242F77C3FFAAB05DFE29C170913A7C7D" box="[483,610,1555,1581]" pageId="14" pageNumber="215" value="1984-07-03">3/VII/1984</date>
,
<date id="242F77C3FFAAB05DFDBBC17191A47C7D" box="[625,764,1554,1581]" pageId="14" pageNumber="215" value="1985-07-11">11/VII/1985</date>
,
<date id="242F77C3FFAAB05DFCC1C17190DE7C7D" box="[779,902,1554,1581]" pageId="14" pageNumber="215" value="1985-10-17">17/X/1985</date>
,
<date id="242F77C3FFAAB05DFC5FC17197617C7D" box="[917,1081,1554,1581]" pageId="14" pageNumber="215" value="1986-07-04" valueMax="1986-07-15" valueMin="1986-07-04">4-15/VII/1986</date>
</collectingDate>
,
<collectorName id="FD6434D5FFAAB05DFB82C17097F57C7D" box="[1096,1197,1555,1581]" pageId="14" pageNumber="215">R. Ayala</collectorName>
,
<collectingDate id="346B8E2BFFAAB05DFB76C17193AF7C01" isEnumeration="true" pageId="14" pageNumber="215">
<date id="242F77C3FFAAB05DFB76C171961D7C7D" box="[1212,1349,1554,1581]" pageId="14" pageNumber="215" value="1985-08-08">8/VIII/1985</date>
,
<date id="242F77C3FFAAB05DFA99C17093AF7C01" pageId="14" pageNumber="215" value="1985-09-26" valueMax="1985-09-30" valueMin="1985-09-26">26-30/ IX/1985</date>
</collectingDate>
,
<collectorName id="FD6434D5FFAAB05DFEC9C15492D57C01" box="[259,397,1591,1617]" pageId="14" pageNumber="215">F. D. Parker</collectorName>
, T.
<collectorName id="FD6434D5FFAAB05DFE7FC15491457C01" box="[437,541,1591,1617]" pageId="14" pageNumber="215">Griswold</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFDE7C154912C7C01" box="[557,628,1591,1617]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="14" pageNumber="215" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFAAB05DFDB1C15591C47C01" box="[635,668,1590,1617]" count="5" pageId="14" pageNumber="215" type="female">5♀</specimenCount>
<specimenCount id="46979A8AFFAAB05DFD56C155919B7C01" box="[668,707,1590,1617]" count="8" pageId="14" pageNumber="215" type="male">8♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFAAB05DFD13C15492A77C25" collectingDate="1989-07-10" collectingDateMax="1989-07-14" collectingDateMin="1989-07-10" collectionCode="BBSL" collectorName="Griswold" country="Mexico" elevation="71" latitude="19.49639" location="Est. Biol." longLatPrecision="21" longitude="-105.04611" municipality="Chamela" pageId="14" pageNumber="215" specimenCount="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<collectingMunicipality id="B04ACB79FFAAB05DFD13C15490677C01" box="[729,831,1591,1617]" pageId="14" pageNumber="215">Chamela</collectingMunicipality>
,
<location id="554E07D8FFAAB05DFC81C15490ED7C01" LSID="urn:lsid:plazi:treatment:D838E015FFA8B043FF5DC2E1973D7BD8:554E07D8FFAAB05DFC81C15490ED7C01" box="[843,949,1591,1617]" country="Mexico" latitude="19.49639" longLatPrecision="21" longitude="-105.04611" municipality="Chamela" name="Est. Biol." pageId="14" pageNumber="215" stateProvince="Jalisco">Est. Biol.</location>
[
<quantity id="9769FCE6FFAAB05DFC09C15497597C01" box="[963,1025,1591,1617]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="215" unit="m" value="71.0">
<elevation id="DBBCB630FFAAB05DFC09C15497597C01" box="[963,1025,1591,1617]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="215" unit="m" value="71.0">71 m</elevation>
</quantity>
,
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],
<date id="242F77C3FFAAB05DFA99C15492597C25" pageId="14" pageNumber="215" value="1989-07-10" valueMax="1989-07-14" valueMin="1989-07-10">
<collectingDate id="346B8E2BFFAAB05DFA99C15492597C25" pageId="14" pageNumber="215" value="1989-07-10" valueMax="1989-07-14" valueMin="1989-07-10">10-14/ VII/1989</collectingDate>
</date>
,
<collectorName id="FD6434D5FFAAB05DFEC6C138922C7C25" box="[268,372,1627,1653]" pageId="14" pageNumber="215">Griswold</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFE49C13892927C25" box="[387,458,1627,1653]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="14" pageNumber="215" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFAAB05DFE1BC13892AE7C25" box="[465,502,1627,1653]" count="1" pageId="14" pageNumber="215" type="male">1♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFFAAB05DFDC6C138965D7C25" box="[524,1285,1627,1653]" collectingDate="1989-07-20" collectionCode="KUNHM-ENT" collectorName="Brooks &amp; Michener &amp; A. Roig" country="Mexico" pageId="14" pageNumber="215" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
<date id="242F77C3FFAAB05DFDC6C13891C17C25" box="[524,665,1627,1653]" pageId="14" pageNumber="215" value="1989-07-20">
<collectingDate id="346B8E2BFFAAB05DFDC6C13891C17C25" box="[524,665,1627,1653]" pageId="14" pageNumber="215" value="1989-07-20">20/VII/1989</collectingDate>
</date>
, R.
<collectorName id="FD6434D5FFAAB05DFD0CC13890427C25" box="[710,794,1627,1653]" pageId="14" pageNumber="215">Brooks</collectorName>
, C.
<collectorName id="FD6434D5FFAAB05DFC8CC13890EB7C25" box="[838,947,1627,1653]" pageId="14" pageNumber="215">Michener</collectorName>
,
<collectorName id="FD6434D5FFAAB05DFC77C138974D7C25" box="[957,1045,1627,1653]" pageId="14" pageNumber="215">A. Roig</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFBEFC13897887C25" box="[1061,1232,1627,1653]" pageId="14" pageNumber="215">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFFAAB05DFB1DC13897A07C25" box="[1239,1272,1627,1653]" count="1" pageId="14" pageNumber="215" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFAAB05DFAD8C138967B7CC9" collectingDate="1985-10-04" collectionCode="BBSL" collectorName="La Huerta &amp; T. Griswold" country="Mexico" elevation="26" latitude="19.279167" location="Careyes" longLatPrecision="21" longitude="-104.790276" municipality="Cuiztmala Playa" pageId="14" pageNumber="215" specimenCount="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
[
<collectorName id="FD6434D5FFAAB05DFAD2C13896CA7C25" box="[1304,1426,1627,1653]" pageId="14" pageNumber="215">La Huerta</collectorName>
],
<collectingMunicipality id="B04ACB79FFAAB05DFF5DC11C92177CC9" box="[151,335,1663,1689]" pageId="14" pageNumber="215">Cuiztmala Playa</collectingMunicipality>
8kmS
<location id="554E07D8FFAAB05DFE55C11C92A47CC9" LSID="urn:lsid:plazi:treatment:D838E015FFA8B043FF5DC2E1973D7BD8:554E07D8FFAAB05DFE55C11C92A47CC9" box="[415,508,1663,1689]" country="Mexico" latitude="19.279167" longLatPrecision="21" longitude="-104.790276" municipality="Cuiztmala Playa" name="Careyes" pageId="14" pageNumber="215" stateProvince="Jalisco">Careyes</location>
, [
<quantity id="9769FCE6FFAAB05DFDC4C11C91137CC9" box="[526,587,1663,1689]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="215" unit="m" value="26.0">
<elevation id="DBBCB630FFAAB05DFDC4C11C91137CC9" box="[526,587,1663,1689]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="215" unit="m" value="26.0">26 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFAAB05DFD9DC11D91B87CC9" box="[599,736,1662,1689]" degrees="19" direction="north" minutes="16" orientation="latitude" pageId="14" pageNumber="215" precision="15" seconds="45" value="19.279167">19°1645”N</geoCoordinate>
<geoCoordinate id="35A537C4FFAAB05DFD2DC11D90D27CC9" box="[743,906,1662,1689]" degrees="104" direction="west" minutes="47" orientation="longitude" pageId="14" pageNumber="215" precision="15" seconds="25" value="-104.790276">104°4725”W</geoCoordinate>
],
<date id="242F77C3FFAAB05DFC56C11D97517CC9" box="[924,1033,1662,1689]" pageId="14" pageNumber="215" value="1985-10-04">
<collectingDate id="346B8E2BFFAAB05DFC56C11D97517CC9" box="[924,1033,1662,1689]" pageId="14" pageNumber="215" value="1985-10-04">4/X/1985</collectingDate>
</date>
,
<collectorName id="FD6434D5FFAAB05DFBD9C11C97C07CC9" box="[1043,1176,1663,1689]" pageId="14" pageNumber="215">T. Griswold</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFB6DC11C97B67CC9" box="[1191,1262,1663,1689]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="14" pageNumber="215" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFAAB05DFB3FC11C96417CC9" box="[1269,1305,1663,1689]" count="1" pageId="14" pageNumber="215" type="male">1♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="502E5103FFAAB05DFF0DC1C190E27D21" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="215">
<emphasis id="62E58D11FFAAB05DFF0DC1C191607CEC" bold="true" box="[199,568,1698,1724]" pageId="14" pageNumber="215">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFFAAB05DFD8AC1C190527CB1" collectingDate="1962-08-16" collectionCode="KUNHM-ENT" collectorName="Ordway &amp; Naumann" country="Mexico" latitude="18.889166" location="Yautepec" longLatPrecision="21" longitude="-99.06278" pageId="14" pageNumber="215" specimenCount="1" specimenCount-female="1" stateProvince="Morelos">
<emphasis id="62E58D11FFAAB05DFD8AC1C190777CED" bold="true" box="[576,815,1698,1725]" pageId="14" pageNumber="215">
<collectingCountry id="28861193FFAAB05DFD8AC1C191E37CEC" box="[576,699,1698,1724]" name="Mexico" pageId="14" pageNumber="215">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFAAB05DFD0DC1C090727CED" box="[711,810,1699,1725]" country="Mexico" name="Morelos" pageId="14" pageNumber="215">Morelos</collectingRegion>
,
</emphasis>
<location id="554E07D8FFAAB05DFCFCC1C090C57CED" LSID="urn:lsid:plazi:treatment:D838E015FFA8B043FF5DC2E1973D7BD8:554E07D8FFAAB05DFCFCC1C090C57CED" box="[822,925,1699,1725]" country="Mexico" latitude="18.889166" longLatPrecision="21" longitude="-99.06278" name="Yautepec" pageId="14" pageNumber="215" stateProvince="Morelos">Yautepec</location>
<quantity id="9769FCE6FFAAB05DFC6FC1C090B57CED" box="[933,1005,1699,1725]" metricMagnitude="4" metricUnit="m" metricValue="1.17482112" pageId="14" pageNumber="215" unit="mi" value="7.3">7.3mi.</quantity>
S, 3000, [
<geoCoordinate id="35A537C4FFAAB05DFBA3C1C197AD7CED" box="[1129,1269,1698,1725]" degrees="18" direction="north" minutes="53" orientation="latitude" pageId="14" pageNumber="215" precision="15" seconds="21" value="18.889166">18°5321”N</geoCoordinate>
<geoCoordinate id="35A537C4FFAAB05DFB36C1C096C97CED" box="[1276,1425,1699,1725]" degrees="99" direction="west" minutes="03" orientation="longitude" pageId="14" pageNumber="215" precision="15" seconds="46" value="-99.06278">99°0346”W</geoCoordinate>
],
<date id="242F77C3FFAAB05DFF5DC1A492757CB1" box="[151,301,1735,1761]" pageId="14" pageNumber="215" value="1962-08-16">
<collectingDate id="346B8E2BFFAAB05DFF5DC1A492757CB1" box="[151,301,1735,1761]" pageId="14" pageNumber="215" value="1962-08-16">16/VIII/1962</collectingDate>
</date>
,
<collectorName id="FD6434D5FFAAB05DFEFDC1A492C97CB1" box="[311,401,1735,1761]" pageId="14" pageNumber="215">Ordway</collectorName>
&amp;
<collectorName id="FD6434D5FFAAB05DFE78C1A491477CB1" box="[434,543,1735,1761]" pageId="14" pageNumber="215">Naumann</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFDE1C1A4918E7CB1" box="[555,726,1735,1761]" pageId="14" pageNumber="215">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFFAAB05DFD16C1A491A57CB1" box="[732,765,1735,1761]" count="1" pageId="14" pageNumber="215" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFAAB05DFCDEC1A590FD7D55" collectingDate="1963-08-14" collectionCode="BBSL" collectorName="Parker &amp; Stange" country="Mexico" elevation="213" latitude="16.868055" location="23 mi South Matias Romero" longLatPrecision="21" longitude="-95.04417" pageId="14" pageNumber="215" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
<emphasis id="62E58D11FFAAB05DFCDEC1A5902D7CB0" bold="true" box="[788,885,1734,1760]" pageId="14" pageNumber="215">
<collectingRegion id="92559FE1FFAAB05DFCDEC1A590287CB0" box="[788,880,1734,1760]" country="Mexico" name="Oaxaca" pageId="14" pageNumber="215">Oaxaca</collectingRegion>
,
</emphasis>
<location id="554E07D8FFAAB05DFCB1C1A497F77CB1" LSID="urn:lsid:plazi:treatment:D838E015FFA8B043FF5DC2E1973D7BD8:554E07D8FFAAB05DFCB1C1A497F77CB1" box="[891,1199,1735,1761]" country="Mexico" latitude="16.868055" longLatPrecision="21" longitude="-95.04417" name="23 mi South Matias Romero" pageId="14" pageNumber="215" stateProvince="Oaxaca">
<quantity id="9769FCE6FFAAB05DFCB1C1A490ED7CB1" box="[891,949,1735,1761]" metricMagnitude="4" metricUnit="m" metricValue="3.7014912" pageId="14" pageNumber="215" unit="mi" value="23.0">23mi</quantity>
South Matias Romero
</location>
, [
<quantity id="9769FCE6FFAAB05DFB75C1A496517CB1" box="[1215,1289,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="14" pageNumber="215" unit="m" value="213.0">
<elevation id="DBBCB630FFAAB05DFB75C1A496517CB1" box="[1215,1289,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="14" pageNumber="215" unit="m" value="213.0">213 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFAAB05DFAD9C1A596C47CB1" box="[1299,1436,1734,1761]" degrees="16" direction="north" minutes="52" orientation="latitude" pageId="14" pageNumber="215" precision="15" seconds="05" value="16.868055">16°5205”N</geoCoordinate>
<geoCoordinate id="35A537C4FFAAB05DFF5DC18992737D55" box="[151,299,1770,1797]" degrees="95" direction="west" minutes="02" orientation="longitude" pageId="14" pageNumber="215" precision="15" seconds="39" value="-95.04417">95°0239”W</geoCoordinate>
],
<date id="242F77C3FFAAB05DFE8BC18892807D55" box="[321,472,1771,1797]" pageId="14" pageNumber="215" value="1963-08-14">
<collectingDate id="346B8E2BFFAAB05DFE8BC18892807D55" box="[321,472,1771,1797]" pageId="14" pageNumber="215" value="1963-08-14">14/VIII/1963</collectingDate>
</date>
, F. D.
<collectorName id="FD6434D5FFAAB05DFDE3C188912C7D55" box="[553,628,1771,1797]" pageId="14" pageNumber="215">Parker</collectorName>
, L. A.
<collectorName id="FD6434D5FFAAB05DFD02C188904C7D55" box="[712,788,1771,1797]" pageId="14" pageNumber="215">Stange</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFCECC18890357D55" box="[806,877,1771,1797]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="14" pageNumber="215" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFAAB05DFCBDC18890C37D55" box="[887,923,1771,1797]" count="1" pageId="14" pageNumber="215" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFAAB05DFC7EC18890C57D79" collectingDate="1963-08-06" collectionCode="BBSL" collectorName="F. D. Parker &amp; L. A. Stange" country="Mexico" elevation="680" latitude="16.56125" location="El Camaron" longLatPrecision="1" longitude="-96.027466" pageId="14" pageNumber="215" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
3miWest
<location id="554E07D8FFAAB05DFBEBC18897F67D55" LSID="urn:lsid:plazi:treatment:D838E015FFA8B043FF5DC2E1973D7BD8:554E07D8FFAAB05DFBEBC18897F67D55" box="[1057,1198,1771,1797]" country="Mexico" latitude="16.56125" longLatPrecision="1" longitude="-96.027466" name="El Camaron" pageId="14" pageNumber="215" stateProvince="Oaxaca">El Camarón</location>
, [
<quantity id="9769FCE6FFAAB05DFB08C18896497D55" box="[1218,1297,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="215" unit="m" value="680.0">
<elevation id="DBBCB630FFAAB05DFB08C18896497D55" box="[1218,1297,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="14" pageNumber="215" unit="m" value="680.0">680 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFAAB05DFAEAC18996D17D55" box="[1312,1417,1770,1797]" degrees="16.56125" direction="north" orientation="latitude" pageId="14" pageNumber="215" precision="1" value="16.56125">16.56125</geoCoordinate>
<geoCoordinate id="35A537C4FFAAB05DFA59C188924C7D79" degrees="96.027467" direction="west" orientation="longitude" pageId="14" pageNumber="215" precision="1" value="-96.027466">- 96.027467</geoCoordinate>
],
<date id="242F77C3FFAAB05DFEE6C06C92EC7D79" box="[300,436,1807,1833]" pageId="14" pageNumber="215" value="1963-08-06">
<collectingDate id="346B8E2BFFAAB05DFEE6C06C92EC7D79" box="[300,436,1807,1833]" pageId="14" pageNumber="215" value="1963-08-06">6/VIII/1963</collectingDate>
</date>
,
<collectorName id="FD6434D5FFAAB05DFE0CC06C91037D79" box="[454,603,1807,1833]" pageId="14" pageNumber="215">F. D. Parker</collectorName>
,
<collectorName id="FD6434D5FFAAB05DFDA7C06C905D7D79" box="[621,773,1807,1833]" pageId="14" pageNumber="215">L. A. Stange</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFCD1C06C903A7D79" box="[795,866,1807,1833]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="14" pageNumber="215" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFAAB05DFCA5C06C90CB7D79" box="[879,915,1807,1833]" count="1" pageId="14" pageNumber="215" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFAAB05DFC65C06C90FF7D1D" collectingDate="1952-07-21" collectionCode="BERK" collectorName="E. E. Gilbert &amp; C. D. MacNeil" country="Mexico" elevation="47" latitude="16.322779" location="Tehuantepec" longLatPrecision="21" longitude="-95.24249" pageId="14" pageNumber="215" specimenCount="2" specimenCount-male="2" stateProvince="Oaxaca">
<location id="554E07D8FFAAB05DFC65C06C97677D79" LSID="urn:lsid:plazi:treatment:D838E015FFA8B043FF5DC2E1973D7BD8:554E07D8FFAAB05DFC65C06C97677D79" box="[943,1087,1807,1833]" country="Mexico" latitude="16.322779" longLatPrecision="21" longitude="-95.24249" name="Tehuantepec" pageId="14" pageNumber="215" stateProvince="Oaxaca">Tehuantepec</location>
, 44miW, [
<quantity id="9769FCE6FFAAB05DFB77C06C96597D79" box="[1213,1281,1807,1833]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="14" pageNumber="215" unit="m" value="47.0">
<elevation id="DBBCB630FFAAB05DFB77C06C96597D79" box="[1213,1281,1807,1833]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="14" pageNumber="215" unit="m" value="47.0">47 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFAAB05DFAD9C06C96C47D79" box="[1299,1436,1807,1833]" degrees="16" direction="north" minutes="19" orientation="latitude" pageId="14" pageNumber="215" precision="15" seconds="22" value="16.322779">16°1922”N</geoCoordinate>
<geoCoordinate id="35A537C4FFAAB05DFF5DC05192737D1D" box="[151,299,1842,1869]" degrees="95" direction="west" minutes="14" orientation="longitude" pageId="14" pageNumber="215" precision="15" seconds="33" value="-95.24249">95°1433”W</geoCoordinate>
],
<date id="242F77C3FFAAB05DFEF4C05192937D1D" box="[318,459,1842,1869]" pageId="14" pageNumber="215" value="1952-07-21">
<collectingDate id="346B8E2BFFAAB05DFEF4C05192937D1D" box="[318,459,1842,1869]" pageId="14" pageNumber="215" value="1952-07-21">21/VII/1952</collectingDate>
</date>
,
<collectorName id="FD6434D5FFAAB05DFE1CC050913E7D1D" box="[470,614,1843,1869]" pageId="14" pageNumber="215">E. E. Gilbert</collectorName>
,
<collectorName id="FD6434D5FFAAB05DFDBBC050904E7D1D" box="[625,790,1843,1869]" pageId="14" pageNumber="215">C. D. MacNeil</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFCEEC05090287D1D" box="[804,880,1843,1869]" pageId="14" pageNumber="215">BERK</collectionCode>
<specimenCount id="46979A8AFFAAB05DFCB2C05090C67D1D" box="[888,926,1843,1869]" count="2" pageId="14" pageNumber="215" type="male">2♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFAAB05DFC7BC05090ED7D21" collectingDate="1961-06-26" collectionCode="KUNHM-ENT" collectorName="U. Kans. Mex. Exped." country="Mexico" elevation="533" latitude="16.368889" location="Tehuantepec" longLatPrecision="21" longitude="-95.331665" pageId="14" pageNumber="215" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
<location id="554E07D8FFAAB05DFC7BC05097197D1D" LSID="urn:lsid:plazi:treatment:D838E015FFA8B043FF5DC2E1973D7BD8:554E07D8FFAAB05DFC7BC05097197D1D" box="[945,1089,1843,1869]" country="Mexico" latitude="16.368889" longLatPrecision="21" longitude="-95.331665" name="Tehuantepec" pageId="14" pageNumber="215" stateProvince="Oaxaca">Tehuantepec</location>
, 28miW,
<quantity id="9769FCE6FFAAB05DFB66C05197A77D1D" box="[1196,1279,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="5.334" pageId="14" pageNumber="215" unit="ft" value="1750.0">
<elevation id="DBBCB630FFAAB05DFB66C05197A77D1D" box="[1196,1279,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="5.334" pageId="14" pageNumber="215" unit="ft" value="1750.0">1750 ft</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FFAAB05DFADBC05096C47D1D" box="[1297,1436,1843,1869]" degrees="16" direction="north" minutes="22" orientation="latitude" pageId="14" pageNumber="215" precision="15" seconds="08" value="16.368889">16°2208”N</geoCoordinate>
<geoCoordinate id="35A537C4FFAAB05DFF5DC03592737D21" box="[151,299,1878,1905]" degrees="95" direction="west" minutes="19" orientation="longitude" pageId="14" pageNumber="215" precision="15" seconds="54" value="-95.331665">95°1954”W</geoCoordinate>
],
<date id="242F77C3FFAAB05DFEF4C03492997D21" box="[318,449,1879,1905]" pageId="14" pageNumber="215" value="1961-06-26">
<collectingDate id="346B8E2BFFAAB05DFEF4C03492997D21" box="[318,449,1879,1905]" pageId="14" pageNumber="215" value="1961-06-26">26/VI/1961</collectingDate>
</date>
,
<collectorName id="FD6434D5FFAAB05DFE07C034919F7D21" box="[461,711,1879,1905]" pageId="14" pageNumber="215">U. Kans. Mex. Exped.</collectorName>
(
<collectionCode id="3680C9C6FFAAB05DFD1FC03490D87D21" box="[725,896,1879,1905]" pageId="14" pageNumber="215">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFFAAB05DFC4DC03490F07D21" box="[903,936,1879,1905]" count="1" pageId="14" pageNumber="215" type="female">1♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFAAB05DFF0DC019926B7D8D" pageId="14" pageNumber="215" type="distribution">
<paragraph id="502E5103FFAAB05DFF0DC019926B7D8D" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="215">
<emphasis id="62E58D11FFAAB05DFF0DC01992397DC4" bold="true" box="[199,353,1914,1940]" pageId="14" pageNumber="215">Distribution.</emphasis>
This species is distributed along the Pacific slope of
<collectingCountry id="28861193FFAAB05DFC07C018977D7DC5" box="[973,1061,1915,1941]" name="Mexico" pageId="14" pageNumber="215">Mexico</collectingCountry>
, with records in central
<collectingCountry id="28861193FFAAB05DFA8AC01896C07DC5" box="[1344,1432,1915,1941]" name="Mexico" pageId="14" pageNumber="215">Mexico</collectingCountry>
, being endemic to it, in the states of
<collectingRegion id="92559FE1FFAAB05DFDE5C0FC91D47DE9" box="[559,652,1951,1977]" country="Mexico" name="Morelos" pageId="14" pageNumber="215">Morelos</collectingRegion>
and
<collectingRegion id="92559FE1FFAAB05DFD0FC0FC90447DE9" box="[709,796,1951,1977]" country="Mexico" name="Oaxaca" pageId="14" pageNumber="215">Oaxaca</collectingRegion>
, present in tropical dry forest (
<figureCitation id="C8AA4D86FFAAB05DFBB0C0FC97987DE9" box="[1146,1216,1951,1977]" captionStart="FIGURE 6" captionStartId="13.[151,250,1098,1122]" captionTargetBox="[159,1428,187,1064]" captionTargetId="figure-18@13.[151,1436,179,1072]" captionTargetPageId="13" captionText="FIGURE 6. Known distribution records for Heriades briceida sp. nov." pageId="14" pageNumber="215">Fig. 6</figureCitation>
). Altitudinal range:
<quantity id="9769FCE6FFAAB05DFF5DC0A092777D8D" box="[151,303,1986,2013]" metricMagnitude="2" metricUnit="m" metricValue="6.385" metricValueMax="12.51" metricValueMin="0.26" pageId="14" pageNumber="215" unit="m" value="638.5" valueMax="1251.0" valueMin="26.0">26 to 1251 m</quantity>
.
</paragraph>
</subSubSection>
<caption id="04EE018BFFABB05CFF5DC0F4907A7D83" pageId="15" pageNumber="216" startId="15.[151,250,1943,1967]" targetBox="[163,1410,183,1910]" targetPageId="15" targetType="figure">
<paragraph id="502E5103FFABB05CFF5DC0F4907A7D83" blockId="15.[151,1436,1943,2003]" pageId="15" pageNumber="216">
<emphasis id="62E58D11FFABB05CFF5DC0F4924B7DFF" bold="true" box="[151,275,1943,1967]" pageId="15" pageNumber="216">FIGURE 7.</emphasis>
<taxonomicName id="97912A80FFABB05CFED3C0F492897DFF" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[281,465,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="briceida" status="sp. nov.">
<emphasis id="62E58D11FFABB05CFED3C0F492897DFF" box="[281,465,1943,1967]" italics="true" pageId="15" pageNumber="216">Heriades briceida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFABB05CFE1DC0FB917E7DFF" box="[471,550,1944,1967]" pageId="15" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
Male:
<emphasis id="62E58D11FFABB05CFDA7C0F491DE7DFF" bold="true" box="[621,646,1943,1967]" pageId="15" pageNumber="216">A.</emphasis>
frontal view
<emphasis id="62E58D11FFABB05CFCC4C0F4907D7DFF" bold="true" box="[782,805,1943,1967]" pageId="15" pageNumber="216">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FFABB05CFBCEC0F497447DFF" bold="true" box="[1028,1052,1943,1967]" pageId="15" pageNumber="216">C.</emphasis>
frontal view
<emphasis id="62E58D11FFABB05CFB6EC0F497E57DFF" bold="true" box="[1188,1213,1943,1967]" pageId="15" pageNumber="216">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FFABB05CFA4FC0F496C47DFF" bold="true" box="[1413,1436,1943,1967]" pageId="15" pageNumber="216">E.</emphasis>
S5.
<emphasis id="62E58D11FFABB05CFF75C0D8938A7D83" bold="true" box="[191,210,1979,2003]" pageId="15" pageNumber="216">F.</emphasis>
S6.
<emphasis id="62E58D11FFABB05CFECAC0D892427D83" bold="true" box="[256,282,1979,2003]" pageId="15" pageNumber="216">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FFABB05CFDB5C0D891C17D83" bold="true" box="[639,665,1979,2003]" pageId="15" pageNumber="216">H.</emphasis>
ventral view.
</paragraph>
</caption>
<subSubSection id="188B0288FFB4B043FF0DC7F5975A7A84" pageId="16" pageNumber="217" type="etymology">
<paragraph id="502E5103FFB4B043FF0DC7F5975A7A84" blockId="16.[151,1437,150,393]" pageId="16" pageNumber="217">
<emphasis id="62E58D11FFB4B043FF0DC7F592167AE0" bold="true" box="[199,334,150,176]" pageId="16" pageNumber="217">Etymology.</emphasis>
This species is dedicated to Briceida Cuevas Cob, bilingual Mayan poet and founder of the Asociación de Escritores en Lenguas Indígenas de
<collectingRegion id="92559FE1FFB4B043FC6FC7D990A67A84" box="[933,1022,186,212]" country="Mexico" name="Mexico" pageId="16" pageNumber="217">Mexico</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFB4B043FF0DC7BD973D7BD8" pageId="16" pageNumber="217" type="discussion">
<paragraph id="502E5103FFB4B043FF0DC7BD973D7BD8" blockId="16.[151,1437,150,393]" pageId="16" pageNumber="217">
<emphasis id="62E58D11FFB4B043FF0DC7BD92177AA8" bold="true" box="[199,335,222,248]" pageId="16" pageNumber="217">Comments.</emphasis>
Males of this species are similar to
<taxonomicName id="97912A80FFB4B043FD26C7BC902F7AA8" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[748,887,222,248]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="viridiana" status="sp. nov.">
<emphasis id="62E58D11FFB4B043FD26C7BC902F7AA8" box="[748,887,222,248]" italics="true" pageId="16" pageNumber="217">H. viridiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFB4B043FCB5C7BC90817AA9" box="[895,985,223,249]" pageId="16" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
, but can be recognized by having black flagellum, process of S1, in posterior view, with rounded corners, similar to two cusps on top, and in tufts and pubescence of apical margin of S5 and S6. We used location data to establish the female of this species.Although we highlight some differences with females of
<taxonomicName id="97912A80FFB4B043FDB4C628905F7B34" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[638,775,330,356]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="viridiana" status="sp. nov.">
<emphasis id="62E58D11FFB4B043FDB4C628905F7B34" box="[638,775,330,356]" italics="true" pageId="16" pageNumber="217">H. viridiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFB4B043FCC4C628903F7B35" box="[782,871,331,357]" pageId="16" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
, further analysis needs to be done due to the small number of specimens, and the similarities among females of other Neotropical species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,767 @@
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<mods:title id="32597C1B3E099D8A879632BD1D67E8F0">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<mods:namePart id="65B21D885949ED7ED627450AD60AFD62">Rojas-Arias, Laura</mods:namePart>
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<mods:affiliation id="C9C5164A521D9A37A4341684820E6CC2">Posgrado en Ciencias Biológicas, Universidad Nacional Autónoma de México (Posgrado en Ciencias Biológicas, UNAM. Unidad de</mods:affiliation>
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<subSubSection id="188B0288FFACB05BFF5DC5309007783E" box="[151,863,595,622]" pageId="8" pageNumber="209" type="nomenclature">
<paragraph id="502E5103FFACB05BFF5DC5309007783E" blockId="8.[151,863,595,659]" box="[151,863,595,622]" pageId="8" pageNumber="209">
<heading id="0B66E66FFFACB05BFF5DC5309007783E" bold="true" box="[151,863,595,622]" fontSize="11" level="1" pageId="8" pageNumber="209" reason="1">
<emphasis id="62E58D11FFACB05BFF5DC5309007783E" bold="true" box="[151,863,595,622]" pageId="8" pageNumber="209">
<taxonomicName id="97912A80FFACB05BFF5DC53091A2783E" authority="Rojas-Arias, Griswold &amp; Ayala" authorityName="Rojas-Arias, Griswold &amp; Ayala" authorityYear="2025" box="[151,762,595,622]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="amanda" status="sp. nov.">
<emphasis id="62E58D11FFACB05BFF5DC5309236783E" bold="true" box="[151,366,595,622]" italics="true" pageId="8" pageNumber="209">Heriades amanda</emphasis>
Rojas-Arias, Griswold &amp; Ayala
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFACB05BFCCBC5379007783E" box="[769,863,596,622]" pageId="8" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
</subSubSection>
<subSubSection id="188B0288FFACB05BFF5DC51B921978C3" box="[151,321,632,659]" pageId="8" pageNumber="209" type="description">
<paragraph id="502E5103FFACB05BFF5DC51B921978C3" blockId="8.[151,863,595,659]" box="[151,321,632,659]" pageId="8" pageNumber="209">
(
<figureCitation id="C8AA4D86FFACB05BFF55C51B925A78C2" box="[159,258,632,658]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="8" pageNumber="209">Figs. 2B</figureCitation>
,
<figureCitation id="C8AA4D86FFACB05BFEC7C51B926178C3" box="[269,313,632,659]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="9.[151,250,1083,1107]" captionStartId-1="11.[151,250,1944,1968]" captionTargetBox-0="[160,1428,172,1049]" captionTargetBox-1="[163,1409,179,1905]" captionTargetId-0="figure-18@9.[151,1436,164,1057]" captionTargetId-1="figure-18@11.[163,1423,179,1918]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="FIGURE 4. Known distribution records for Heriades amanda sp. nov." captionText-1="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="8" pageNumber="209">45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFACB05BFF5DC5DD976579A2" pageId="8" pageNumber="209" type="diagnosis">
<paragraph id="502E5103FFACB05BFF5DC5DD976579A2" blockId="8.[151,1437,702,2032]" pageId="8" pageNumber="209">
<emphasis id="62E58D11FFACB05BFF5DC5DD924A7888" bold="true" box="[151,274,702,728]" pageId="8" pageNumber="209">Diagnosis.</emphasis>
Disc of metepisternum with tuft of plumose, thick, short, white hairs (
<figureCitation id="C8AA4D86FFACB05BFBF8C5DD97F77888" box="[1074,1199,702,729]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="8" pageNumber="209">Fig. 5B, D</figureCitation>
). Male: clypeus with small, contiguous punctures (at base 3.5 times OD); large interantennal distance (3.4 times antennocular distance); frons, upper and middle part of paraocular area with pubescence of golden hairs; scutum with small punctures (0.2 times OD) (
<figureCitation id="C8AA4D86FFACB05BFED0C44B922B7913" box="[282,371,808,835]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="8" pageNumber="209">Fig. 5A</figureCitation>
); posterior margin of S1 produced to median rounded angle, in profile, with low ventral angle in more than one third of distance from apex to base (
<figureCitation id="C8AA4D86FFACB05BFD1FC42890417935" box="[725,793,843,869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="8" pageNumber="209">Fig. 2</figureCitation>
Ba); S5 with two rectangular lobes and two rounded tufts of capitate hairs directed medially (
<figureCitation id="C8AA4D86FFACB05BFDE8C40D912079D9" box="[546,632,878,905]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="8" pageNumber="209">Fig. 5E</figureCitation>
); S6 with posterior margin produced to right-angled apex, with blackish subapical margin, with two subapical tufts of 3 to 4 thick, long, hairs together resembling an acute spike (
<figureCitation id="C8AA4D86FFACB05BFAF0C4F296D779FC" box="[1338,1423,913,940]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="8" pageNumber="209">Fig. 5F</figureCitation>
). Female: clypeus with 6 tubercles on apical margin and large punctures (0.3 times OD); frons with hooked, golden hairs; T3 with small (half DOP as on scutum), and sparse punctures (2 times DOP).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFACB05FFF0DC49890467B11" lastPageId="12" lastPageNumber="213" pageId="8" pageNumber="209" type="description">
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<emphasis id="62E58D11FFACB05BFF0DC49892FC7E45" bold="true" box="[199,420,1019,1046]" pageId="8" pageNumber="209">
Description.
<emphasis id="62E58D11FFACB05BFEAAC49892C37E46" bold="true" box="[352,411,1019,1046]" italics="true" pageId="8" pageNumber="209">Male</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FFACB05BFDF6C49891CA7E46" box="[572,658,1019,1046]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="8" pageNumber="209" unit="mm" value="5.6">5.6 mm</quantity>
(4.75.9, n=6); forewing length
<quantity id="9769FCE6FFACB05BFBCBC498970F7E46" box="[1025,1111,1019,1046]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="209" unit="mm" value="3.5">3.5 mm</quantity>
(3.13.5, n=6).
</paragraph>
<paragraph id="502E5103FFACB05AFF0DC37D91877F1F" blockId="8.[151,1437,702,2032]" lastBlockId="9.[151,1437,1153,2044]" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209">
<emphasis id="62E58D11FFACB05BFF0DC37D925C7E68" box="[199,260,1054,1080]" italics="true" pageId="8" pageNumber="209">Head</emphasis>
. Rounded, about as long as broad; length
<quantity id="9769FCE6FFACB05BFD06C37D90767E69" box="[716,814,1054,1081]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="8" pageNumber="209" unit="mm" value="1.58">1.58 mm</quantity>
(1.51.6, n=6); width
<quantity id="9769FCE6FFACB05BFBE9C37D972F7E69" box="[1059,1143,1054,1081]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="209" unit="mm" value="1.7">1.7 mm</quantity>
(1.51.7, n=6); eyes about 3.3 times longer than broad, 1.8 times wider than gena in profile, convergent below; lower interocular distance
<quantity id="9769FCE6FFACB05BFAB3C321939B7E2F" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="209" unit="mm" value="0.8">0.8 mm</quantity>
(0.70.8, n=6); upper interocular distance
<quantity id="9769FCE6FFACB05BFD6CC30691B07E2F" box="[678,744,1125,1151]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="209" unit="mm" value="1.0">1 mm</quantity>
(0.91.1, n=6); interantennal distance 3.4 times antennocular distance; lateral ocelli large, 3 times DOP in interocellar area, median ocellus comparable; interocellar distance 2 times OD; ocellocular distance 1.7 times OD; OPR 0.9 times OD; labrum with basal median elevation with two cusps, posterior margin slightly rounded; mandible with two teeth, apical subtriangular and preapical quadrate, both almost same width; proboscis in repose reaching posteriormost edge of hypostomal carina, not surpassing it; hypostomal edge In ventral view carinate, bent laterally on anterior and posterior sections; malar space linear, without posterior projection; clypeus truncate, 1.5 times broader than long, lower half convex in profile, apical margin with concave medial emargination occupying 0.3 of clypeal width; juxtantennal carina absent; antenna black; scape length 0.7 times alveolocellar distance, 2.4 times longer than wide; pedicel as long as broad, about 1.2 times longer than F1 and about as long as F2; F1 as broad as long, slightly smaller than F2, remaining flagellomeres 1.4 longer than first; F11 1.8 times longer than broad.
<emphasis id="62E58D11FFACB05BFD25C2A690367F8E" box="[751,878,1477,1502]" italics="true" pageId="8" pageNumber="209">Mesosoma.</emphasis>
Propleura with strong angle not carinate; pronotal lobe carinate reaching rounded dorsolateral angles of pronotum; mesepisternum bent on dorsal and ventral edges; scutum 3 times longer than scutellum, 3 times longer than tegula, 3.6 longer than axilla, bent anteriorly with vertical and dorsal surface; intertegular distance
<quantity id="9769FCE6FFACB05BFDA6C14D919D7C19" box="[620,709,1582,1609]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="209" unit="mm" value="1.2">1.2 mm</quantity>
(11.2, n=6); axilla subtriangular, not pointy; metanotum with transverse ridge, in posterior view with medial acute elevation; propodeum with small to midsized pits,
<quantity id="9769FCE6FFACB05BFA8EC13196C57C3C" box="[1348,1437,1618,1644]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="209" unit="mm" value="0.1">0.1 mm</quantity>
long; legs dark; fore coxa subtriangular, bulky, convex except on anterior margin; fore trochanter conic, 2 times longer than broad; fore femur swollen, 2.3 times longer than wide; fore tibia slightly conic, 3.1 times longer than broad, with posterior external spike of
<quantity id="9769FCE6FFACB05BFD86C1D891E87C86" box="[588,688,1723,1750]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="209" unit="mm" value="0.04">0.04 mm</quantity>
long; fore basitarsus 2.5 times longer than broad, with two lateral, rounded projections; fore tarsal segment 2 about as long as broad, 3 and 4 becoming shorter; tarsal segment 5 trapezoid, 1.8 times longer than broad on its posterior margin, with two bifurcated claws, arolium conic or slightly ligulate; mid coxa oval, with two vertical, divergent carinae, where femur fits; mid trochanter 1.5 times longer than broad; femur swollen, 2.2 times longer than broad; mid tibia slightly bent, 7 times longer than broad, with posterior spike similar to the one in fore tibia; mid basitarsus 4.2 times longer than broad; rest of mid tarsal segments as in fore leg; hind coxa flat dorsolaterally, subquadrangular medially and rounded on external sides; hind trochanter 1.2 times longer than broad; hind femur 2.2 times longer than broad; hind tibia 3.5 times longer than broad, posterior margin with external spike and inner spur; hind basitarsus 3.1 times longer than broad.
<emphasis id="62E58D11FFACB05BFBD0C0B597C07DBF" box="[1050,1176,2006,2031]" italics="true" pageId="8" pageNumber="209">Metasoma.</emphasis>
Nearly parallel-sized; T1
<quantity id="9769FCE6FFADB05AFF75C3E2924E7ECC" box="[191,278,1153,1180]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="210" unit="mm" value="1.4">1.4 mm</quantity>
broad (1.21.4, n=6); T3
<quantity id="9769FCE6FFADB05AFDF5C3E291CE7ECC" box="[575,662,1153,1180]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="210" unit="mm" value="1.6">1.6 mm</quantity>
broad (1.41.6, n=6); T6 with posterior margin projected, in profile with rounded elevation
<quantity id="9769FCE6FFADB05AFE54C3C692AB7E90" box="[414,499,1189,1216]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="210" unit="mm" value="0.5">0.5 mm</quantity>
from apex to base,
<quantity id="9769FCE6FFADB05AFD01C3C690787E90" box="[715,800,1189,1216]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="210" unit="mm" value="0.3">0.3 mm</quantity>
wide; S1 posterior margin produced to median angle, not emarginate, slightly rounded, with two lateral, rounded projections on each side, in profile, with posterior margin projected and soft ventral angle more than one third of distance from apex to base; S2 truncate, with two wide, soft lobes at posterior margin, with two rounded elevations on each side of disc; S3 and S4 similar to S2 but less sclerotized; terga, S1 and S2 with black integument.
</paragraph>
<caption id="04EE018BFFADB05AFF5DC358903C7E02" box="[151,868,1083,1107]" pageId="9" pageNumber="210" startId="9.[151,250,1083,1107]" targetBox="[160,1428,172,1049]" targetPageId="9" targetType="figure">
<paragraph id="502E5103FFADB05AFF5DC358903C7E02" blockId="9.[151,868,1083,1107]" box="[151,868,1083,1107]" pageId="9" pageNumber="210">
<emphasis id="62E58D11FFADB05AFF5DC358924B7E03" bold="true" box="[151,275,1083,1107]" pageId="9" pageNumber="210">FIGURE 4.</emphasis>
Known distribution records for
<taxonomicName id="97912A80FFADB05AFD90C35890577E03" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[602,783,1083,1107]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="amanda" status="sp. nov.">
<emphasis id="62E58D11FFADB05AFD90C35890577E03" box="[602,783,1083,1107]" italics="true" pageId="9" pageNumber="210">Heriades amanda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFADB05AFCDFC358903C7E02" box="[789,868,1083,1106]" pageId="9" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="502E5103FFADB059FF0DC23A93B37B33" blockId="9.[151,1437,1153,2044]" lastBlockId="10.[151,1437,150,2036]" lastPageId="10" lastPageNumber="211" pageId="9" pageNumber="210">
<emphasis id="62E58D11FFADB05AFF0DC23A92097F23" box="[199,337,1369,1395]" italics="true" pageId="9" pageNumber="210">Sculpturing.</emphasis>
Mandible with outer surfaces bright with micropunctures without larger punctures, carinae smooth and shiny; labrum with dense micropunctures; clypeus with small (at base 0.3 times OD), contiguous, coarse punctures, smaller towards apex and medially, shiny between punctures; supraclypeal area with punctures smaller, denser than base of clypeus, slightly bigger, more coarsely but equally sparser on frons, even bigger on frons in front of ocelli and vertex, frons and vertex with microreticulate integument as rest of face; hypostomal area with fine, minute punctures, DBP half DOP or less, bigger, sparser towards gena; gena evenly punctate as on frons in front of ocelli; anterior surface of propleuron with punctation as on disc of clypeus, lateral surface with dense micropunctures as on labrum; dorsal surface of pronotum as on anterior surface of propleuron, anterior surface with medium-sized and sparser punctation, becoming smaller, denser and coarser on dorsolateral angles; disc of scutum with smaller (0.2 times OD), denser punctures than on vertex, bigger towards external margin; rest of mesosoma as on scutum except: scutellum with punctures bigger, sparser, smaller towards external margin; mesepisternum with punctures sparser, with micropunctures between punctures; metepisternum with smaller punctures, less dense, coarse; lateral surface of propodeum with even smaller punctures; posterior surface of propodeum with smaller, less coarse, sparser punctures, except on wide impunctate vertical midline; tegula with fine, minute, scattered punctures; fore coxa with punctation as on propleuron; fore trochanter and femur with smaller sparser punctation than coxa; fore tibia and tarsal segments with less coarse, sparser punctation than on femur; mid and hind legs similar to fore legs except: mid femur with denser punctation, hind coxa with bigger, sparser punctation, hind tibia with bigger, coarser, sparser punctation; metasomal terga smooth, shiny between punctures, becoming smaller towards posterior margin, with line of small punctures where fasciae attach; T1 and T2 with punctation as on scutum, slightly sparser; T3 with smaller, sparser punctures than on T1, slightly horseshoe-shaped, separated by half DOP; T4 and T5 with bigger, denser, coarser punctation than on T3; dorsal surface of T6 with smaller, equally sparser punctation than on T5, on posterior surface with bigger, notably sparser punctation, DBP half DOP; S1 with punctation as on T1, but slightly bigger; sides of S2 similar to on T1; integument of disc in S2 with integument without punctation; S3 and S4 as disc of S2, but less sclerotized; S5 to S8 with less sclerotized, smooth, shiny, amber integument than on S3 and S4.
</paragraph>
<paragraph id="502E5103FFAEB059FF0DC60D961C792B" blockId="10.[151,1437,150,2036]" pageId="10" pageNumber="211">
<emphasis id="62E58D11FFAEB059FF0DC60D92687BD7" box="[199,304,366,391]" italics="true" pageId="10" pageNumber="211">Vestiture.</emphasis>
Mandible with plumose, white hairs at base and simple, golden hairs at apex; labrum without tuft; clypeus covered with plumose, white, of similar length evenly distributed on its entire surface; paraocular area with plumose, white hairs on its entire surface; frons with plumose, thin, golden hairs directed medially; preoccipital area with rows of plumose, white hairs directed dorsally; hypostomal area with plumose, long, white hairs, in frontal view surpassing mandible; dorsal surface of pronotum with rows of plumose, short, white hairs; dorsolateral angles with plumose, white hairs continuing on anterior margin of scutum forming, a single continuous, transverse, linear band; pronotal lobe on posterior area of carina with plumose, thin, dense, white hairs; scutum with transverse pubescence of plumose, white hairs on sides continuing on posterior margin of scutellum, directed dorsally, increasing in length towards scutellum; scutum and mesepisternum with simple, thin, short, scattered, golden hairs on punctures; lateral surface of mesepisternum with lower two thirds and ventral surface with plumose, thin, white hairs; disc of metepisternum with tuft of plumose, thick, short, white hair; sides of propodeum, and posterior surface of propodeum with hairs as on mesepisternum; anterior surface of T1 with plumose, thin, scattered, white hairs, on posterior area of transverse carina with row of plumose, short, sparse, erected, white hairs; terga with fasciae of row of dense, short, white hairs, increasing in length towards sides, increasing in number of rows on T1; T4 to T6 with plumose, scattered, white hairs in punctures, with row of simple, scattered, short hairs on posterior margin.
</paragraph>
<paragraph id="502E5103FFAEB059FF0DC4E691417EED" blockId="10.[151,1437,150,2036]" pageId="10" pageNumber="211">
<emphasis id="62E58D11FFAEB059FF0DC4E6921179CF" box="[199,329,901,927]" italics="true" pageId="10" pageNumber="211">
<taxonomicName id="97912A80FFAEB059FF0DC4E6921D79CF" box="[199,325,901,927]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="10" pageNumber="211" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
Posterior margin of S5 with two rectangular lobes, two rounded tufts of capitate hairs directed medially, simple short hairs directed outward, with transverse, thin, dark, subapical line (
<figureCitation id="C8AA4D86FFAEB059FBB4C4CA978D7994" box="[1150,1237,937,964]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="10" pageNumber="211">Fig. 5E</figureCitation>
; Posterior margin of S6 produced to right-angled apex, with blackish subapical margin, with two subapical tufts of 3 to 4 long, thick, hairs, together resembling an acute spike (
<figureCitation id="C8AA4D86FFAEB059FDA0C49391E77E5B" box="[618,703,1008,1035]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="10" pageNumber="211">Fig. 5F</figureCitation>
); S7 produced to rounded wide medial angle, with sclerotized line on posterior margin; S8 arrow-shaped, simple hairs spread on posterior third, medial line from base to two thirds of S8 and lateral, thin extensions, joining together at tips, surrounding genital capsule; genital capsule in dorsal view
<quantity id="9769FCE6FFAEB059FF5DC33F93B77E26" box="[151,239,1116,1142]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="211" unit="mm" value="0.8">0.8 mm</quantity>
length,
<quantity id="9769FCE6FFAEB059FE87C33F92FC7E26" box="[333,420,1116,1142]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="211" unit="mm" value="0.3">0.3 mm</quantity>
width, 2.7 times longer than wide, gonobase
<quantity id="9769FCE6FFAEB059FC64C33F974C7E26" box="[942,1044,1116,1142]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="211" unit="mm" value="0.08">0.08 mm</quantity>
length, 3.8 times wider than long, gonocoxite
<quantity id="9769FCE6FFAEB059FED1C31C922A7ECA" box="[283,370,1151,1178]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="211" unit="mm" value="0.3">0.3 mm</quantity>
length, gonostylus
<quantity id="9769FCE6FFAEB059FD85C31C91FE7ECA" box="[591,678,1151,1178]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="211" unit="mm" value="0.7">0.7 mm</quantity>
length with rounded apex, penis valve
<quantity id="9769FCE6FFAEB059FBA9C31C97E27ECA" box="[1123,1210,1151,1178]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="211" unit="mm" value="0.6">0.6 mm</quantity>
length, with lateral, rounded lamellas in posterior half.
</paragraph>
<paragraph id="502E5103FFAEB059FF0DC3A591227CDE" blockId="10.[151,1437,150,2036]" pageId="10" pageNumber="211">
<emphasis id="62E58D11FFAEB059FF0DC3A5927F7EB1" bold="true" box="[199,295,1222,1249]" pageId="10" pageNumber="211">
<emphasis id="62E58D11FFAEB059FF0DC3A592467EB1" bold="true" box="[199,286,1222,1249]" italics="true" pageId="10" pageNumber="211">Female</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FFAEB059FE0FC3A591457EB1" box="[453,541,1222,1249]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="211" unit="mm" value="5.1">5.1 mm</quantity>
(5.16.8, n=6); forewing length 4.0 mm (3.44, n=6). Similar to male except:
<emphasis id="62E58D11FFAEB059FF5DC389938C7F54" box="[151,212,1258,1284]" italics="true" pageId="10" pageNumber="211">Head</emphasis>
. Length
<quantity id="9769FCE6FFAEB059FEF6C38992CC7F55" box="[316,404,1258,1285]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="211" unit="mm" value="1.8">1.8 mm</quantity>
(1.51.8, n=6); width
<quantity id="9769FCE6FFAEB059FD53C38991A97F55" box="[665,753,1258,1285]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="211" unit="mm" value="1.8">1.8 mm</quantity>
(1.61.9, n=6); eyes about 3.7 times longer than broad, 1.9 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FFAEB059FCE7C26D90DA7F79" box="[813,898,1294,1321]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="211" unit="mm" value="0.9">0.9 mm</quantity>
(0.81, n=6); upper interocular distance
<quantity id="9769FCE6FFAEB059FA8DC26D96C57F79" box="[1351,1437,1294,1321]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="211" unit="mm" value="1.1">1.1 mm</quantity>
(11.2, n=6); interantennal distance 1.8 times antennocular distance; interocellar distance 2.2 times OD, ocellocular distance 1.7 times OD; OPR 1-time OD; mandible with strong outer and condylar ridges parallel, without tubercle on outer surface, with toothlike elevation on dorsal margin; three teeth, apical longer than second, both subtriangular and almost same width, third right-angulated; clypeus without emargination and apical margin with six rounded tubercles; scape 3.2 times longer than broad; pedicel 1.3 longer than broad, about 1.6 times longer than F1 and 1.3 longer than F2; F1 broader than long, remaining flagellomeres 1.8 longer than F1.
<emphasis id="62E58D11FFAEB059FBADC28697877FAE" box="[1127,1247,1509,1534]" italics="true" pageId="10" pageNumber="211">Mesosoma</emphasis>
. Scutum 3 times longer than scutellum; intertegular distance
<quantity id="9769FCE6FFAEB059FD4AC16B918D7C73" box="[640,725,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="211" unit="mm" value="1.4">1.4 mm</quantity>
(1.21.4, n=6).
<emphasis id="62E58D11FFAEB059FC42C16A90A57C72" box="[904,1021,1545,1570]" italics="true" pageId="10" pageNumber="211">Metasoma</emphasis>
. T1
<quantity id="9769FCE6FFAEB059FBE5C16B97DC7C73" box="[1071,1156,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="211" unit="mm" value="1.5">1.5 mm</quantity>
broad (1.31.6, n=6); T3
<quantity id="9769FCE6FFAEB059FF5DC14F93B67C17" box="[151,238,1580,1607]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="211" unit="mm" value="1.7">1.7 mm</quantity>
broad (1.51.8, n=6); T6 with posterior margin slightly projected and subapical margin, in profile, almost straight; all sterna truncate, with lighter margin; S1 with shallow, wide emargination along margin and small process in profile near base; S6 with rounded apex.
</paragraph>
<paragraph id="502E5103FFAEB059FF0DC1F4976F7DA3" blockId="10.[151,1437,150,2036]" pageId="10" pageNumber="211">
<emphasis id="62E58D11FFAEB059FF0DC1F492097CE1" box="[199,337,1687,1713]" italics="true" pageId="10" pageNumber="211">Sculpturing.</emphasis>
Mandibles dull with micropunctures without larger puctures, carinae smooth and shiny; labrum with punctation as on mandible; clypeus with large (0.3 times OD), dense, coarse and contiguous punctures, microreticulate between punctures; supraclypeal area with punctures bigger than those on base of clypeus, bigger, less coarse, and sparser on frons, smaller on frons in front of ocelli; vertex as on clypeus; gena evenly punctate as on frons in front of ocelli, with large punctation and less DBP; scutum with punctation as on vertex, smaller towards external margin; rest of mesosoma as on scutum except: scutellum with bigger and sparser punctures; mesepisternum with punctures slightly smaller; metasomal terga dull, microreticulate between punctures; T1 and T2 with punctation as on scutum, slightly sparser. T3 with punctures half DOP as on scutum, horseshoe-shaped, and sparser (2 times DOP) than on T1; dorsal surface of T6 with smaller, denser, and more coarsely punctures than on T5; sterna with punctation as on scutum, with micropunctures on posterior margin.
</paragraph>
<caption id="04EE018BFFAFB058FF5DC0FB907A7D84" pageId="11" pageNumber="212" startId="11.[151,250,1944,1968]" targetBox="[163,1409,179,1905]" targetPageId="11" targetType="figure">
<paragraph id="502E5103FFAFB058FF5DC0FB907A7D84" blockId="11.[151,1436,1944,2004]" pageId="11" pageNumber="212">
<emphasis id="62E58D11FFAFB058FF5DC0FB924B7DE0" bold="true" box="[151,275,1944,1968]" pageId="11" pageNumber="212">FIGURE 5.</emphasis>
<taxonomicName id="97912A80FFAFB058FED0C0FB92977DE0" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[282,463,1944,1968]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="amanda" status="sp. nov.">
<emphasis id="62E58D11FFAFB058FED0C0FB92977DE0" box="[282,463,1944,1968]" italics="true" pageId="11" pageNumber="212">Heriades amanda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFAFB058FE1FC0FB917C7DFF" box="[469,548,1944,1967]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
Male:
<emphasis id="62E58D11FFAFB058FDA1C0FB91DC7DE0" bold="true" box="[619,644,1944,1968]" pageId="11" pageNumber="212">A.</emphasis>
frontal view
<emphasis id="62E58D11FFAFB058FCC6C0FB907B7DE0" bold="true" box="[780,803,1944,1968]" pageId="11" pageNumber="212">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FFAFB058FBC9C0FB97437DE0" bold="true" box="[1027,1051,1944,1968]" pageId="11" pageNumber="212">C.</emphasis>
frontal view
<emphasis id="62E58D11FFAFB058FB6EC0FB97E47DE0" bold="true" box="[1188,1212,1944,1968]" pageId="11" pageNumber="212">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FFAFB058FA4FC0FB96C47DE0" bold="true" box="[1413,1436,1944,1968]" pageId="11" pageNumber="212">E.</emphasis>
S5.
<emphasis id="62E58D11FFAFB058FF75C0DF938A7D84" bold="true" box="[191,210,1980,2004]" pageId="11" pageNumber="212">F.</emphasis>
S6.
<emphasis id="62E58D11FFAFB058FECAC0DF92427D84" bold="true" box="[256,282,1980,2004]" pageId="11" pageNumber="212">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FFAFB058FDB5C0DF91C17D84" bold="true" box="[639,665,1980,2004]" pageId="11" pageNumber="212">H.</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="502E5103FFA8B05FFF0DC7F490467B11" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="213">
<emphasis id="62E58D11FFA8B05FFF0DC7F492687AE0" box="[199,304,151,176]" italics="true" pageId="12" pageNumber="213">Vestiture.</emphasis>
Mandible with few golden hairs at apex and ventral margin; clypeus with simple, golden hairs evenly distributed; frons with hooked, golden hairs; hypostomal area with few simple, golden hairs; gena with plumose, white hairs evenly distributed; T4 to T6 with simple, scattered, thin, white hairs in punctures; subapical margin of T6 with linear tuft of plumose, short, yellowish hairs; scopa with hooked, thin, long hairs and posterior margin of S6 with transverse tuft of plumose, short, yellowish hairs.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFA8B05FFF0DC628901479BD" pageId="12" pageNumber="213" type="materials_examined">
<paragraph id="502E5103FFA8B05FFF0DC62897887849" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="213">
<emphasis id="62E58D11FFA8B05FFF0DC62892207B34" bold="true" box="[199,376,330,357]" pageId="12" pageNumber="213">Type material:</emphasis>
<materialsCitation id="E0F95B5EFFA8B05FFEB5C62892307BD9" collectingDate="1985-10-08" collectionCode="BBSL" collectorName="No. LR &amp; F. D. Parker &amp; T. L. Griswold." country="Mexico" location="Jalisco" pageId="12" pageNumber="213" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="62E58D11FFA8B05FFEB5C628909D7B34" bold="true" box="[383,965,330,357]" pageId="12" pageNumber="213">
<typeStatus id="8F2AEFA1FFA8B05FFEB5C62892B47B35" box="[383,492,331,357]" pageId="12" pageNumber="213" type="holotype">Holotype</typeStatus>
male (
<collectorName id="FD6434D5FFA8B05FFDF6C62991C07B34" box="[572,664,330,356]" pageId="12" pageNumber="213">No. LR</collectorName>
0001):
<collectingCountry id="28861193FFA8B05FFD2DC62990077B34" box="[743,863,330,356]" name="Mexico" pageId="12" pageNumber="213">MEXICO</collectingCountry>
.
<collectingRegion id="92559FE1FFA8B05FFCA1C62990987B34" box="[875,960,330,356]" country="Mexico" name="Jalisco" pageId="12" pageNumber="213">Jalisco</collectingRegion>
:
</emphasis>
Chamela. 1,
<date id="242F77C3FFA8B05FFB91C62997907B35" box="[1115,1224,330,357]" pageId="12" pageNumber="213" value="1985-10-08">
<collectingDate id="346B8E2BFFA8B05FFB91C62997907B35" box="[1115,1224,330,357]" pageId="12" pageNumber="213" value="1985-10-08">8/X/1985</collectingDate>
</date>
.
<collectorName id="FD6434D5FFA8B05FFB19C62896047B35" box="[1235,1372,331,357]" pageId="12" pageNumber="213">F. D. Parker</collectorName>
,
<collectorName id="FD6434D5FFA8B05FFAADC628925F7BD9" pageId="12" pageNumber="213">T. L. Griswold.</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFEDEC60C92387BD9" box="[276,352,367,393]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="12" pageNumber="213" type="Museum">BBSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFEBBC60D914E7BFD" collectingDate="1985-04-15" collectingDateMax="1985-04-24" collectingDateMin="1985-04-15" collectionCode="BBSL" collectorName="F. D. Parker" country="Mexico" elevation="19" latitude="19.524721" location="Chamela" longLatPrecision="21" longitude="-105.05056" pageId="12" pageNumber="213" specimenCount="1" stateProvince="Jalisco" typeStatus="allotype">
<emphasis id="62E58D11FFA8B05FFEBBC60D90607BD8" bold="true" box="[369,824,366,392]" pageId="12" pageNumber="213">
<typeStatus id="8F2AEFA1FFA8B05FFEBBC60D928E7BD8" box="[369,470,366,392]" pageId="12" pageNumber="213" type="allotype">Allotype</typeStatus>
(LR0002):
<collectingCountry id="28861193FFA8B05FFD91C60D918B7BD8" box="[603,723,366,392]" name="Mexico" pageId="12" pageNumber="213">MEXICO</collectingCountry>
.
<collectingRegion id="92559FE1FFA8B05FFD14C60D906B7BD8" box="[734,819,366,392]" country="Mexico" name="Jalisco" pageId="12" pageNumber="213">Jalisco</collectingRegion>
:
</emphasis>
<location id="554E07D8FFA8B05FFCF4C60C90FA7BD9" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFCF4C60C90FA7BD9" box="[830,930,367,393]" country="Mexico" latitude="19.524721" longLatPrecision="21" longitude="-105.05056" name="Chamela" pageId="12" pageNumber="213" stateProvince="Jalisco">Chamela</location>
(Rio), [
<quantity id="9769FCE6FFA8B05FFC3FC60C97747BD9" box="[1013,1068,367,393]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="213" unit="m" value="19.0">
<elevation id="DBBCB630FFA8B05FFC3FC60C97747BD9" box="[1013,1068,367,393]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="213" unit="m" value="19.0">19m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFA8B05FFBFFC60C97E67BD9" box="[1077,1214,367,393]" degrees="19" direction="north" minutes="31" orientation="latitude" pageId="12" pageNumber="213" precision="15" seconds="29" value="19.524721">19°3129”N</geoCoordinate>
<geoCoordinate id="35A537C4FFA8B05FFB0EC60D963E7BD9" box="[1220,1382,366,393]" degrees="105" direction="west" minutes="03" orientation="longitude" pageId="12" pageNumber="213" precision="15" seconds="02" value="-105.05056">105°0302”W</geoCoordinate>
],
<date id="242F77C3FFA8B05FFABDC60D92437BFD" pageId="12" pageNumber="213" value="1985-04-15" valueMax="1985-04-24" valueMin="1985-04-15">
<collectingDate id="346B8E2BFFA8B05FFABDC60D92437BFD" pageId="12" pageNumber="213" value="1985-04-15" valueMax="1985-04-24" valueMin="1985-04-15">15- 24/IV/1985</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFEE2C6F092E97BFD" box="[296,433,403,429]" pageId="12" pageNumber="213">F. D. Parker</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFE08C6F091567BFD" box="[450,526,403,429]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="12" pageNumber="213" type="Museum">BBSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFDE8C6F091807B81" collectingDate="18 / II / 1986, 23 / VI / 1990" collectionCode="BBSL" collectorName="R. Ayala" country="Mexico" elevation="71" latitude="19.49639" location="Chamela" longLatPrecision="21" longitude="-105.04611" pageId="12" pageNumber="213" specimenCount="17" specimenCount-female="11" specimenCount-male="6" stateProvince="Jalisco" typeStatus="paratype">
<emphasis id="62E58D11FFA8B05FFDE8C6F090D67BFC" bold="true" box="[546,910,402,429]" pageId="12" pageNumber="213">
<typeStatus id="8F2AEFA1FFA8B05FFDE8C6F091C67BFD" box="[546,670,403,429]" pageId="12" pageNumber="213" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="28861193FFA8B05FFD66C6F1907F7BFC" box="[684,807,402,428]" name="Mexico" pageId="12" pageNumber="213">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFA8B05FFCFEC6F190D17BFC" box="[820,905,402,428]" country="Mexico" name="Jalisco" pageId="12" pageNumber="213">Jalisco</collectingRegion>
:
</emphasis>
<location id="554E07D8FFA8B05FFC5DC6F090A57BFD" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFC5DC6F090A57BFD" box="[919,1021,403,429]" country="Mexico" latitude="19.49639" longLatPrecision="21" longitude="-105.04611" name="Chamela" pageId="12" pageNumber="213" stateProvince="Jalisco">Chamela</location>
, [
<quantity id="9769FCE6FFA8B05FFBDBC6F097177BFD" box="[1041,1103,403,429]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="12" pageNumber="213" unit="m" value="71.0">
<elevation id="DBBCB630FFA8B05FFBDBC6F097177BFD" box="[1041,1103,403,429]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="12" pageNumber="213" unit="m" value="71.0">71 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFA8B05FFB97C6F097BE7BFD" box="[1117,1254,403,429]" degrees="19" direction="north" minutes="29" orientation="latitude" pageId="12" pageNumber="213" precision="15" seconds="47" value="19.49639">19°2947”N</geoCoordinate>
<geoCoordinate id="35A537C4FFA8B05FFB24C6F196C97BFD" box="[1262,1425,402,429]" degrees="105" direction="west" minutes="02" orientation="longitude" pageId="12" pageNumber="213" precision="15" seconds="46" value="-105.04611">105°0246”W</geoCoordinate>
],
<collectingDate id="346B8E2BFFA8B05FFF5DC6D492FB7B81" box="[151,419,439,465]" isEnumeration="true" pageId="12" pageNumber="213">
<date id="242F77C3FFA8B05FFF5DC6D492487B81" box="[151,272,439,465]" pageId="12" pageNumber="213" value="1986-02-18">18/II/1986</date>
,
<date id="242F77C3FFA8B05FFED5C6D492FB7B81" box="[287,419,439,465]" pageId="12" pageNumber="213" value="1990-06-23">23/VI/1990</date>
</collectingDate>
,
<collectorName id="FD6434D5FFA8B05FFE78C6D4914D7B81" box="[434,533,439,465]" pageId="12" pageNumber="213">R. Ayala</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFDE2C6D491377B81" box="[552,623,439,465]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="12" pageNumber="213" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFA8B05FFDB0C6D491FF7B81" box="[634,679,439,465]" count="11" pageId="12" pageNumber="213" type="female">11♀</specimenCount>
<specimenCount id="46979A8AFFA8B05FFD6DC6D491967B81" box="[679,718,439,465]" count="6" pageId="12" pageNumber="213" type="male">6♂</specimenCount>
)
</materialsCitation>
, 15,
<date id="242F77C3FFA8B05FFCDCC6D490C07B81" box="[790,920,439,465]" pageId="12" pageNumber="213" value="1986-03-19">19/III/1986</date>
, R. Ayala (EBCh
<specimenCount id="46979A8AFFA8B05FFBA4C6D497C87B81" box="[1134,1168,439,465]" count="3" pageId="12" pageNumber="213" type="female">3♀</specimenCount>
<specimenCount id="46979A8AFFA8B05FFB45C6D497EC7B81" box="[1167,1204,439,465]" count="1" pageId="12" pageNumber="213" type="male">1♂</specimenCount>
),
<materialsCitation id="E0F95B5EFFA8B05FFB05C6D5907C7BA5" collectingDate="1985-10-01" collectingDateMax="1985-10-08" collectingDateMin="1985-10-01" collectionCode="BBSL" collectorName="F. D. Parker &amp; T. L. Griswold &amp; Rozen" country="Mexico" pageId="12" pageNumber="213" specimenCount="4" specimenCount-male="4" stateProvince="Jalisco" typeStatus="paratype">
<date id="242F77C3FFA8B05FFB05C6D5960B7B81" box="[1231,1363,438,465]" pageId="12" pageNumber="213" value="1985-10-01" valueMax="1985-10-08" valueMin="1985-10-01">
<collectingDate id="346B8E2BFFA8B05FFB05C6D5960B7B81" box="[1231,1363,438,465]" pageId="12" pageNumber="213" value="1985-10-01" valueMax="1985-10-08" valueMin="1985-10-01">1-8/X/1985</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFAA8C6D493BA7BA5" pageId="12" pageNumber="213">F. D. Parker</collectorName>
,
<collectorName id="FD6434D5FFA8B05FFF27C6B892CB7BA5" box="[237,403,475,501]" pageId="12" pageNumber="213">T. L. Griswold</collectorName>
,
<date id="242F77C3FFA8B05FFE55C6B991537BA5" box="[415,523,474,501]" pageId="12" pageNumber="213" value="1985-10-06">
<collectingDate id="346B8E2BFFA8B05FFE55C6B991537BA5" box="[415,523,474,501]" pageId="12" pageNumber="213" value="1985-10-06">6/X/1985</collectingDate>
</date>
, J. G.
<collectorName id="FD6434D5FFA8B05FFD9BC6B891C17BA5" box="[593,665,475,501]" pageId="12" pageNumber="213">Rozen</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFD6DC6B891B67BA5" box="[679,750,475,501]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="12" pageNumber="213" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFA8B05FFD3FC6B890437BA5" box="[757,795,475,501]" count="4" pageId="12" pageNumber="213" type="male">4♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFCE5C6B8911E7849" collectingDate="1985-04-15" collectingDateMax="1985-04-24" collectingDateMin="1985-04-15" collectionCode="BBSL" collectorName="F. D. Parker" country="Mexico" elevation="19" latitude="19.524721" location="Rio" longLatPrecision="21" longitude="-105.05056" municipality="Chamela" pageId="12" pageNumber="213" specimenCount="13" specimenCount-female="13" stateProvince="Jalisco" typeStatus="paratype">
<collectingMunicipality id="B04ACB79FFA8B05FFCE5C6B890CB7BA5" box="[815,915,475,501]" pageId="12" pageNumber="213">Chamela</collectingMunicipality>
(
<location id="554E07D8FFA8B05FFC68C6B890977BA5" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFC68C6B890977BA5" box="[930,975,475,501]" country="Mexico" latitude="19.524721" longLatPrecision="21" longitude="-105.05056" municipality="Chamela" name="Rio" pageId="12" pageNumber="213" stateProvince="Jalisco">Rio</location>
), [
<quantity id="9769FCE6FFA8B05FFC20C6B8977F7BA5" box="[1002,1063,475,501]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="213" unit="m" value="19.0">
<elevation id="DBBCB630FFA8B05FFC20C6B8977F7BA5" box="[1002,1063,475,501]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="213" unit="m" value="19.0">19 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFA8B05FFBF8C6B897E37BA5" box="[1074,1211,475,501]" degrees="19" direction="north" minutes="31" orientation="latitude" pageId="12" pageNumber="213" precision="15" seconds="29" value="19.524721">19°3129”N</geoCoordinate>
<geoCoordinate id="35A537C4FFA8B05FFB08C6B9963D7BA5" box="[1218,1381,474,501]" degrees="105" direction="west" minutes="03" orientation="longitude" pageId="12" pageNumber="213" precision="15" seconds="02" value="-105.05056">105°0302”W</geoCoordinate>
],
<date id="242F77C3FFA8B05FFABDC6B992437849" pageId="12" pageNumber="213" value="1985-04-15" valueMax="1985-04-24" valueMin="1985-04-15">
<collectingDate id="346B8E2BFFA8B05FFABDC6B992437849" pageId="12" pageNumber="213" value="1985-04-15" valueMax="1985-04-24" valueMin="1985-04-15">15- 24/IV/1985</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFEECC69C92F57849" box="[294,429,511,537]" pageId="12" pageNumber="213">F. D. Parker</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFE76C69C915B7849" box="[444,515,511,537]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="12" pageNumber="213" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFA8B05FFDC3C69C91637849" box="[521,571,511,537]" count="13" pageId="12" pageNumber="213" type="female">13♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFD98C69C97927849" box="[594,1226,510,537]" collectingDate="1986-04-15" collectingDateMax="1986-04-24" collectingDateMin="1986-04-15" collectionCode="BBSL" collectorName="F. D. Parker" country="Mexico" location="Rio" municipality="Chamela" pageId="12" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
<collectingMunicipality id="B04ACB79FFA8B05FFD98C69C91EE7849" box="[594,694,511,537]" pageId="12" pageNumber="213">Chamela</collectingMunicipality>
(
<location id="554E07D8FFA8B05FFD0EC69C91A97849" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFD0EC69C91A97849" box="[708,753,511,537]" country="Mexico" municipality="Chamela" name="Rio" pageId="12" pageNumber="213" stateProvince="Jalisco">Rio</location>
),
<date id="242F77C3FFA8B05FFCCFC69D90F77849" box="[773,943,510,537]" pageId="12" pageNumber="213" value="1986-04-15" valueMax="1986-04-24" valueMin="1986-04-15">
<collectingDate id="346B8E2BFFA8B05FFCCFC69D90F77849" box="[773,943,510,537]" pageId="12" pageNumber="213" value="1986-04-15" valueMax="1986-04-24" valueMin="1986-04-15">15-24/IV/1986</collectingDate>
</date>
.
<collectorName id="FD6434D5FFA8B05FFC70C69C97187849" box="[954,1088,511,537]" pageId="12" pageNumber="213">F. D. Parker</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFB85C69C97CE7849" box="[1103,1174,511,537]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="12" pageNumber="213" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFA8B05FFB57C69C97E67849" box="[1181,1214,511,537]" count="1" pageId="12" pageNumber="213" type="female">1♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="502E5103FFA8B05FFF0DC541901479BD" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="213">
<emphasis id="62E58D11FFA8B05FFF0DC5419163786C" bold="true" box="[199,571,546,572]" pageId="12" pageNumber="213">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFFA8B05FFD8EC54091547831" collectingDate="2005-02-23" collectionCode="ECOAB" collectorName="M. Guzman" country="Mexico" elevation="596" location="Acacoyagua" pageId="12" pageNumber="213" specimenCode="ECOAB 1" specimenCount="57" specimenCount-female="38" specimenCount-male="19" stateProvince="Chiapas">
<specimenCount id="46979A8AFFA8B05FFD8EC540912D786D" box="[580,629,547,573]" count="38" pageId="12" pageNumber="213" type="female">38♀</specimenCount>
<specimenCount id="46979A8AFFA8B05FFDB4C54091EB786D" box="[638,691,547,573]" count="19" pageId="12" pageNumber="213" type="male">19♂</specimenCount>
.
<emphasis id="62E58D11FFA8B05FFD0AC54190EA786C" bold="true" box="[704,946,546,572]" pageId="12" pageNumber="213">
<collectingCountry id="28861193FFA8B05FFD0AC5419060786C" box="[704,824,546,572]" name="Mexico" pageId="12" pageNumber="213">MEXICO</collectingCountry>
.
<collectingRegion id="92559FE1FFA8B05FFC8CC54190F4786C" box="[838,940,546,572]" country="Mexico" name="Chiapas" pageId="12" pageNumber="213">Chiapas</collectingRegion>
:
</emphasis>
<location id="554E07D8FFA8B05FFC73C540971E786D" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFC73C540971E786D" box="[953,1094,547,573]" country="Mexico" name="Acacoyagua" pageId="12" pageNumber="213" stateProvince="Chiapas">Acacoyagua</location>
,
<quantity id="9769FCE6FFA8B05FFB9EC54197C7786D" box="[1108,1183,546,573]" metricMagnitude="2" metricUnit="m" metricValue="5.96" pageId="12" pageNumber="213" unit="m" value="596.0">
<elevation id="DBBCB630FFA8B05FFB9EC54197C7786D" box="[1108,1183,546,573]" metricMagnitude="2" metricUnit="m" metricValue="5.96" pageId="12" pageNumber="213" unit="m" value="596.0">596 m</elevation>
</quantity>
, 15.44473 -92.70,
<date id="242F77C3FFA8B05FFAB2C54093B57831" pageId="12" pageNumber="213" value="2005-02-23">
<collectingDate id="346B8E2BFFA8B05FFAB2C54093B57831" pageId="12" pageNumber="213" value="2005-02-23">23/ II/2005</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFF3DC52492257831" box="[247,381,583,609]" pageId="12" pageNumber="213">M. Guzmán</collectorName>
(
<specimenCode id="0037F978FFA8B05FFE41C52491587831" box="[395,512,583,609]" collectionCode="ECOAB" pageId="12" pageNumber="213">ECOAB 1</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFDD2C52592A378D5" collectingDate="2019-12-29" collectionCode="ECOAB" collectorName="P. Sagot &amp; E. Caballero &amp; Gomez" country="Mexico" elevation="318" location="Azoyu" pageId="12" pageNumber="213" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guerrero">
<emphasis id="62E58D11FFA8B05FFDD2C52591CA7830" bold="true" box="[536,658,582,608]" pageId="12" pageNumber="213">
<collectingRegion id="92559FE1FFA8B05FFDD2C52591D57830" box="[536,653,582,608]" country="Mexico" name="Guerrero" pageId="12" pageNumber="213">Guerrero</collectingRegion>
:
</emphasis>
<location id="554E07D8FFA8B05FFD5DC52491BC7831" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFD5DC52491BC7831" box="[663,740,583,609]" country="Mexico" name="Azoyu" pageId="12" pageNumber="213" stateProvince="Guerrero">Azoyu</location>
,
<quantity id="9769FCE6FFA8B05FFD25C52490607831" box="[751,824,583,609]" metricMagnitude="2" metricUnit="m" metricValue="3.18" pageId="12" pageNumber="213" unit="m" value="318.0">
<elevation id="DBBCB630FFA8B05FFD25C52490607831" box="[751,824,583,609]" metricMagnitude="2" metricUnit="m" metricValue="3.18" pageId="12" pageNumber="213" unit="m" value="318.0">318 m</elevation>
</quantity>
, 16.77561 -98.60,
<date id="242F77C3FFA8B05FFBC3C52497CF7831" box="[1033,1175,583,609]" pageId="12" pageNumber="213" value="2019-12-29">
<collectingDate id="346B8E2BFFA8B05FFBC3C52497CF7831" box="[1033,1175,583,609]" pageId="12" pageNumber="213" value="2019-12-29">29/XII/2019</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFB68C52497A77831" box="[1186,1279,583,609]" pageId="12" pageNumber="213">P. Sagot</collectorName>
,
<collectorName id="FD6434D5FFA8B05FFAC3C52496C07831" box="[1289,1432,583,609]" pageId="12" pageNumber="213">E. Caballero</collectorName>
, E.
<collectorName id="FD6434D5FFA8B05FFF7DC508925078D5" box="[183,264,619,645]" pageId="12" pageNumber="213">Gómez</collectorName>
L. (
<collectionCode id="3680C9C6FFA8B05FFEF2C50892C178D5" box="[312,409,619,645]" pageId="12" pageNumber="213">ECOAB</collectionCode>
<specimenCount id="46979A8AFFA8B05FFE6BC508929C78D5" box="[417,452,619,645]" count="1" pageId="12" pageNumber="213" type="female">1♀</specimenCount>
<specimenCount id="46979A8AFFA8B05FFE07C50892A978D5" box="[461,497,619,645]" count="1" pageId="12" pageNumber="213" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFDC2C50896CF78D5" box="[520,1431,619,645]" collectingDate="2019-12-26" collectionCode="ECOAB" collectorName="E. Caballero" country="Mexico" elevation="1090" location="Malinaltepec" pageId="12" pageNumber="213" specimenCount="1" specimenCount-male="1" stateProvince="Guerrero">
<location id="554E07D8FFA8B05FFDC2C50891C578D5" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFDC2C50891C578D5" box="[520,669,619,645]" country="Mexico" name="Malinaltepec" pageId="12" pageNumber="213" stateProvince="Guerrero">Malinaltepec</location>
,
<quantity id="9769FCE6FFA8B05FFD63C508905A78D5" box="[681,770,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="12" pageNumber="213" unit="m" value="1090.0">
<elevation id="DBBCB630FFA8B05FFD63C508905A78D5" box="[681,770,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="12" pageNumber="213" unit="m" value="1090.0">1090 m</elevation>
</quantity>
, 17.0244 -98.78,
<date id="242F77C3FFA8B05FFC03C508970F78D5" box="[969,1111,619,645]" pageId="12" pageNumber="213" value="2019-12-26">
<collectingDate id="346B8E2BFFA8B05FFC03C508970F78D5" box="[969,1111,619,645]" pageId="12" pageNumber="213" value="2019-12-26">26/XII/2019</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFBA9C50897A878D5" box="[1123,1264,619,645]" pageId="12" pageNumber="213">E. Caballero</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFACAC508963978D5" box="[1280,1377,619,645]" pageId="12" pageNumber="213">ECOAB</collectionCode>
<specimenCount id="46979A8AFFA8B05FFAA3C50896D578D5" box="[1385,1421,619,645]" count="1" pageId="12" pageNumber="213" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFF5DC5ED92D8789D" collectingDate="1987-10-29" collectionCode="KUNHM-ENT" collectorName="L. Godinez" country="Mexico" elevation="1390" latitude="19.179722" location="Las Canas" longLatPrecision="21" longitude="-101.753334" pageId="12" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Michoacan de Ocampo">
<emphasis id="62E58D11FFA8B05FFF5DC5ED927D78F9" bold="true" box="[151,293,654,681]" pageId="12" pageNumber="213">
<collectingRegion id="92559FE1FFA8B05FFF5DC5ED927878F9" box="[151,288,654,681]" country="Mexico" name="Michoacan de Ocampo" pageId="12" pageNumber="213">Michoacán</collectingRegion>
:
</emphasis>
<location id="554E07D8FFA8B05FFEFAC5EC92FF78F9" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFEFAC5EC92FF78F9" box="[304,423,655,681]" country="Mexico" latitude="19.179722" longLatPrecision="21" longitude="-101.753334" name="Las Canas" pageId="12" pageNumber="213" stateProvince="Michoacan de Ocampo">Las Canas</location>
<locationDeviation id="419C4821FFA8B05FFE79C5EC918D78F9" box="[435,725,655,681]" location="Ario de Rosales" pageId="12" pageNumber="213">
<quantity id="9769FCE6FFA8B05FFE79C5EC92AB78F9" box="[435,499,655,681]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="213" unit="km" value="21.0">21km</quantity>
S Ario de Rosales
</locationDeviation>
,
<quantity id="9769FCE6FFA8B05FFD2EC5EC906778F9" box="[740,831,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.39" pageId="12" pageNumber="213" unit="m" value="1390.0">
<elevation id="DBBCB630FFA8B05FFD2EC5EC906778F9" box="[740,831,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.39" pageId="12" pageNumber="213" unit="m" value="1390.0">1390 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFA8B05FFC85C5EC908078F9" box="[847,984,655,681]" degrees="19" direction="north" minutes="10" orientation="latitude" pageId="12" pageNumber="213" precision="15" seconds="47" value="19.179722">19°1047”N</geoCoordinate>
<geoCoordinate id="35A537C4FFA8B05FFC29C5ED97D978F9" box="[995,1153,654,681]" degrees="101" direction="west" minutes="45" orientation="longitude" pageId="12" pageNumber="213" precision="15" seconds="12" value="-101.753334">101°4512”W</geoCoordinate>
,
<date id="242F77C3FFA8B05FFB5AC5EC965378F9" box="[1168,1291,655,681]" pageId="12" pageNumber="213" value="1987-10-29">
<collectingDate id="346B8E2BFFA8B05FFB5AC5EC965378F9" box="[1168,1291,655,681]" pageId="12" pageNumber="213" value="1987-10-29">29/X/1987</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFAD0C5EC96C478F9" box="[1306,1436,655,681]" pageId="12" pageNumber="213">L. Godinez</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFF54C5D09211789D" box="[158,329,691,717]" pageId="12" pageNumber="213">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFFA8B05FFE98C5D0922C789D" box="[338,372,691,717]" count="1" pageId="12" pageNumber="213" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFE44C5D196CF789D" box="[398,1431,690,717]" collectingDate="2010-01-08" collectionCode="ECOAB" collectorName="P. Sagot" country="Mexico" elevation="115" location="Santa Maria Huatulco" pageId="12" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
<emphasis id="62E58D11FFA8B05FFE44C5D192AA789C" bold="true" box="[398,498,690,716]" pageId="12" pageNumber="213">
<collectingRegion id="92559FE1FFA8B05FFE44C5D192B5789C" box="[398,493,690,716]" country="Mexico" name="Oaxaca" pageId="12" pageNumber="213">Oaxaca</collectingRegion>
:
</emphasis>
<location id="554E07D8FFA8B05FFE31C5D091AF789D" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFE31C5D091AF789D" box="[507,759,691,717]" country="Mexico" name="Santa Maria Huatulco" pageId="12" pageNumber="213" stateProvince="Oaxaca">Santa María Huatulco</location>
,
<quantity id="9769FCE6FFA8B05FFCCEC5D19015789D" box="[772,845,690,717]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="213" unit="m" value="115.0">
<elevation id="DBBCB630FFA8B05FFCCEC5D19015789D" box="[772,845,690,717]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="213" unit="m" value="115.0">115 m</elevation>
</quantity>
, 15.75778 -96.26,
<date id="242F77C3FFA8B05FFBEFC5D097DE789D" box="[1061,1158,691,717]" pageId="12" pageNumber="213" value="2010-01-08">
<collectingDate id="346B8E2BFFA8B05FFBEFC5D097DE789D" box="[1061,1158,691,717]" pageId="12" pageNumber="213" value="2010-01-08">8/I/2010</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFB59C5D097B7789D" box="[1171,1263,691,717]" pageId="12" pageNumber="213">P. Sagot</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFB35C5D09638789D" box="[1279,1376,691,717]" pageId="12" pageNumber="213">ECOAB</collectionCode>
<specimenCount id="46979A8AFFA8B05FFAA3C5D096D2789D" box="[1385,1418,691,717]" count="1" pageId="12" pageNumber="213" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFF5DC5B4970078A1" box="[151,1112,726,753]" collectingDate="2015-11-10" collectionCode="ECOAB" collectorName="N. Arnold" country="Mexico" elevation="312" location="San Juan Colorado" pageId="12" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
<location id="554E07D8FFA8B05FFF5DC5B4922C78A1" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFF5DC5B4922C78A1" box="[151,372,727,753]" country="Mexico" name="San Juan Colorado" pageId="12" pageNumber="213" stateProvince="Oaxaca">San Juan Colorado</location>
,
<quantity id="9769FCE6FFA8B05FFE48C5B4929678A1" box="[386,462,727,753]" metricMagnitude="2" metricUnit="m" metricValue="3.12" pageId="12" pageNumber="213" unit="m" value="312.0">
<elevation id="DBBCB630FFA8B05FFE48C5B4929678A1" box="[386,462,727,753]" metricMagnitude="2" metricUnit="m" metricValue="3.12" pageId="12" pageNumber="213" unit="m" value="312.0">312 m</elevation>
</quantity>
, 16.48506 -97.90,
<date id="242F77C3FFA8B05FFD63C5B5907478A1" box="[681,812,726,753]" pageId="12" pageNumber="213" value="2015-11-10">
<collectingDate id="346B8E2BFFA8B05FFD63C5B5907478A1" box="[681,812,726,753]" pageId="12" pageNumber="213" value="2015-11-10">10/XI/2015</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFCF1C5B490F678A1" box="[827,942,727,753]" pageId="12" pageNumber="213">N. Arnold</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFC0AC5B4977978A1" box="[960,1057,727,753]" pageId="12" pageNumber="213">ECOAB</collectionCode>
<specimenCount id="46979A8AFFA8B05FFBE0C5B4971478A1" box="[1066,1100,727,753]" count="1" pageId="12" pageNumber="213" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFBA2C5B491A27945" collectingDate="2016-01-09" collectionCode="ECOAB" collectorName="P. Sagot" country="Mexico" elevation="30" location="Santa Maria Tonameca" pageId="12" pageNumber="213" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
<location id="554E07D8FFA8B05FFBA2C5B4962A78A1" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFBA2C5B4962A78A1" box="[1128,1394,727,753]" country="Mexico" name="Santa Maria Tonameca" pageId="12" pageNumber="213" stateProvince="Oaxaca">Santa María Tonameca</location>
,
<quantity id="9769FCE6FFA8B05FFA4AC5B493F77945" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="213" unit="m" value="30.0">
<elevation id="DBBCB630FFA8B05FFA4AC5B493F77945" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="213" unit="m" value="30.0">30 m</elevation>
</quantity>
, 15.67066 -96.54,
<date id="242F77C3FFA8B05FFE4DC59892B07945" box="[391,488,763,789]" pageId="12" pageNumber="213" value="2016-01-09">
<collectingDate id="346B8E2BFFA8B05FFE4DC59892B07945" box="[391,488,763,789]" pageId="12" pageNumber="213" value="2016-01-09">9/I/2016</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFE3FC598910A7945" box="[501,594,763,789]" pageId="12" pageNumber="213">P. Sagot</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFDA8C598919B7945" box="[610,707,763,789]" pageId="12" pageNumber="213">ECOAB</collectionCode>
<specimenCount id="46979A8AFFA8B05FFD01C59891A87945" box="[715,752,763,789]" count="1" pageId="12" pageNumber="213" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFCCDC59891507969" collectingDate="2018-12-11" collectionCode="ECOAB" collectorName="P. Sagot" country="Mexico" elevation="863" location="Santo Domingo Teojomulco" pageId="12" pageNumber="213" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
<location id="554E07D8FFA8B05FFCCDC59897127945" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFCCDC59897127945" box="[775,1098,763,789]" country="Mexico" name="Santo Domingo Teojomulco" pageId="12" pageNumber="213" stateProvince="Oaxaca">Santo Domingo Teojomulco</location>
,
<quantity id="9769FCE6FFA8B05FFB9DC59897FA7945" box="[1111,1186,763,789]" metricMagnitude="2" metricUnit="m" metricValue="8.629999999999999" pageId="12" pageNumber="213" unit="m" value="863.0">
<elevation id="DBBCB630FFA8B05FFB9DC59897FA7945" box="[1111,1186,763,789]" metricMagnitude="2" metricUnit="m" metricValue="8.629999999999999" pageId="12" pageNumber="213" unit="m" value="863.0">863 m</elevation>
</quantity>
, 16.56159 -97.33,
<date id="242F77C3FFA8B05FFAB3C59892597969" pageId="12" pageNumber="213" value="2018-12-11">
<collectingDate id="346B8E2BFFA8B05FFAB3C59892597969" pageId="12" pageNumber="213" value="2018-12-11">11/ XII/2018</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFEC1C47C923D7969" box="[267,357,799,825]" pageId="12" pageNumber="213">P. Sagot</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFEB8C47C928B7969" box="[370,467,799,825]" pageId="12" pageNumber="213">ECOAB</collectionCode>
<specimenCount id="46979A8AFFA8B05FFE13C47C92A57969" box="[473,509,799,825]" count="1" pageId="12" pageNumber="213" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFDD9C47C926C790D" collectingDate="2018-12-18" collectionCode="ECOAB" collectorName="O. Caballero &amp; P. Sagot" country="Mexico" elevation="246" location="San Pedro Tututepec" pageId="12" pageNumber="213" specimenCount="2" specimenCount-male="2" stateProvince="Oaxaca">
<location id="554E07D8FFA8B05FFDD9C47C91A47969" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFDD9C47C91A47969" box="[531,764,799,825]" country="Mexico" name="San Pedro Tututepec" pageId="12" pageNumber="213" stateProvince="Oaxaca">San Pedro Tututepec</location>
,
<quantity id="9769FCE6FFA8B05FFCCCC47C90167969" box="[774,846,799,825]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="12" pageNumber="213" unit="m" value="246.0">
<elevation id="DBBCB630FFA8B05FFCCCC47C90167969" box="[774,846,799,825]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="12" pageNumber="213" unit="m" value="246.0">246 m</elevation>
</quantity>
, 15.9671 -97.19,
<date id="242F77C3FFA8B05FFBDAC47C97C57969" box="[1040,1181,799,825]" pageId="12" pageNumber="213" value="2018-12-18">
<collectingDate id="346B8E2BFFA8B05FFBDAC47C97C57969" box="[1040,1181,799,825]" pageId="12" pageNumber="213" value="2018-12-18">18/XII/2018</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFB6DC47C96617969" box="[1191,1337,799,825]" pageId="12" pageNumber="213">O. Caballero</collectorName>
,
<collectorName id="FD6434D5FFA8B05FFA89C47C96C47969" box="[1347,1436,799,825]" pageId="12" pageNumber="213">P. Sagot</collectorName>
(
<collectionCode id="3680C9C6FFA8B05FFF55C4209258790D" box="[159,256,835,861]" pageId="12" pageNumber="213">ECOAB</collectionCode>
<specimenCount id="46979A8AFFA8B05FFECFC4209273790D" box="[261,299,835,861]" count="2" pageId="12" pageNumber="213" type="male">2♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFEF5C420967E790D" box="[319,1318,834,861]" collectingDate="1989-02-06" collectorName="G. Rodriguez" country="Mexico" elevation="85" latitude="15.868611" location="Puerto Escondido" longLatPrecision="21" longitude="-97.07083" pageId="12" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
<location id="554E07D8FFA8B05FFEF5C420915F790D" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFEF5C420915F790D" box="[319,519,835,861]" country="Mexico" latitude="15.868611" longLatPrecision="21" longitude="-97.07083" name="Puerto Escondido" pageId="12" pageNumber="213" stateProvince="Oaxaca">Puerto Escondido</location>
, [
<quantity id="9769FCE6FFA8B05FFDD2C421910C790D" box="[536,596,834,861]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="85.0">
<elevation id="DBBCB630FFA8B05FFDD2C421910C790D" box="[536,596,834,861]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="213" unit="m" value="85.0">85 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFA8B05FFD95C42191B0790D" box="[607,744,834,861]" degrees="15" direction="north" minutes="52" orientation="latitude" pageId="12" pageNumber="213" precision="15" seconds="07" value="15.868611">15°5207”N</geoCoordinate>
<geoCoordinate id="35A537C4FFA8B05FFD24C42190D9790D" box="[750,897,834,861]" degrees="97" direction="west" minutes="04" orientation="longitude" pageId="12" pageNumber="213" precision="15" seconds="15" value="-97.07083">97°0415”W</geoCoordinate>
],
<date id="242F77C3FFA8B05FFC59C42090A7790D" box="[915,1023,835,861]" pageId="12" pageNumber="213" value="1989-02-06">
<collectingDate id="346B8E2BFFA8B05FFC59C42090A7790D" box="[915,1023,835,861]" pageId="12" pageNumber="213" value="1989-02-06">6/II/1989</collectingDate>
</date>
,
<collectorName id="FD6434D5FFA8B05FFBC2C42097C6790D" box="[1032,1182,835,861]" pageId="12" pageNumber="213">G. Rodriguez</collectorName>
(EBCh
<specimenCount id="46979A8AFFA8B05FFB32C4209641790D" box="[1272,1305,835,861]" count="1" pageId="12" pageNumber="213" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFA8B05FFAFBC420970479D1" collectingDate="1989-03-05" collectorName="R. Ayala" country="Mexico" elevation="442" latitude="16.691666" location="Km" longLatPrecision="21" longitude="-98.02945" municipality="Putla" pageId="12" pageNumber="213" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
Pinotepa-
<collectingMunicipality id="B04ACB79FFA8B05FFF5DC404938B79D1" box="[151,211,871,897]" pageId="12" pageNumber="213">Putla</collectingMunicipality>
,
<location id="554E07D8FFA8B05FFF28C404925479D1" LSID="urn:lsid:plazi:treatment:D838E015FFACB05FFF5DC53090A77F61:554E07D8FFA8B05FFF28C404925479D1" box="[226,268,871,897]" country="Mexico" latitude="16.691666" longLatPrecision="21" longitude="-98.02945" municipality="Putla" name="Km" pageId="12" pageNumber="213" stateProvince="Oaxaca">Km</location>
112, [
<quantity id="9769FCE6FFA8B05FFE9CC40492C479D1" box="[342,412,871,897]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="12" pageNumber="213" unit="m" value="442.0">
<elevation id="DBBCB630FFA8B05FFE9CC40492C479D1" box="[342,412,871,897]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="12" pageNumber="213" unit="m" value="442.0">442m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFA8B05FFE60C404916B79D1" box="[426,563,871,897]" degrees="16" direction="north" minutes="41" orientation="latitude" pageId="12" pageNumber="213" precision="15" seconds="30" value="16.691666">16°4130”N</geoCoordinate>
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:
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, [
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~
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N, [
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&amp;
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(
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<specimenCount id="46979A8AFFA8B05FFCD3C4B0906279BD" box="[793,826,979,1005]" count="1" pageId="12" pageNumber="213" type="female">1♀</specimenCount>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFA8B05FFF0DC49590897E09" pageId="12" pageNumber="213" type="distribution">
<paragraph id="502E5103FFA8B05FFF0DC49590897E09" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="213">
<emphasis id="62E58D11FFA8B05FFF0DC49592397E40" bold="true" box="[199,353,1014,1040]" pageId="12" pageNumber="213">Distribution.</emphasis>
This species is distributed along the west coast in
<collectingRegion id="92559FE1FFA8B05FFC5EC49490B27E41" box="[916,1002,1015,1041]" country="Mexico" name="Mexico" pageId="12" pageNumber="213">Mexico</collectingRegion>
on the Pacific slope, between northern
<collectingRegion id="92559FE1FFA8B05FFF5DC37893B37E65" box="[151,235,1051,1077]" country="Mexico" name="Sinaloa" pageId="12" pageNumber="213">Sinaloa</collectingRegion>
and
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, in areas with a tropical dry forest, mostly corresponding to the Pacific Lowlands biogeographic province (
<bibRefCitation id="34002CF2FFA8B05FFEC2C35C92B37E09" author="Morrone, J. J. &amp; Escalante, T. &amp; Rodriguez-Tapia, G. &amp; Carmona, A. &amp; Arana, M. &amp; Mercado-Gomez, J. D." box="[264,491,1087,1113]" pageId="12" pageNumber="213" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. &amp; Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="62E58D11FFA8B05FFEB8C35C92FC7E09" box="[370,420,1087,1113]" italics="true" pageId="12" pageNumber="213">et al</emphasis>
. 2022
</bibRefCitation>
) (
<figureCitation id="C8AA4D86FFA8B05FFDC8C35C911F7E09" box="[514,583,1087,1113]" captionStart="FIGURE 4" captionStartId="9.[151,250,1083,1107]" captionTargetBox="[160,1428,172,1049]" captionTargetId="figure-18@9.[151,1436,164,1057]" captionTargetPageId="9" captionText="FIGURE 4. Known distribution records for Heriades amanda sp. nov." pageId="12" pageNumber="213">Fig. 4</figureCitation>
). Altitudinal range:
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</paragraph>
</subSubSection>
<subSubSection id="188B0288FFA8B05FFF0DC30096C17E95" pageId="12" pageNumber="213" type="etymology">
<paragraph id="502E5103FFA8B05FFF0DC30096C17E95" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="213">
<emphasis id="62E58D11FFA8B05FFF0DC30092167E2D" bold="true" box="[199,334,1123,1149]" pageId="12" pageNumber="213">Etymology.</emphasis>
This species is dedicated to the Mexican curator and art historian Amanda de la Garza, General Director of Visual Arts and the University Museum of Contemporary Art from UNAM (MUAC). Her professional practice has focused on contemporary art and archival exhibitions, contributing greatly to Mexicos cultural scene.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFA8B05FFF0DC3AD90A77F61" pageId="12" pageNumber="213" type="discussion">
<paragraph id="502E5103FFA8B05FFF0DC3AD90A77F61" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="213">
<emphasis id="62E58D11FFA8B05FFF0DC3AD92177EB8" bold="true" box="[199,335,1230,1256]" pageId="12" pageNumber="213">Comments.</emphasis>
There is variation among the females on the DBP on scutum, from one third to one fourth, as well as on the DOP on T3 varying from larger than on T1 to smaller than on T3, but as coarse as T3. Among males there is a small variation on the interantennal distance and on DBP and DOP on T3.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="FB5DD16A065ACC01319E076A367642AD" type="ZooBank">7177B69B-6132-485D-87C7-4EA807692757</mods:identifier>
</mods:mods>
<treatment id="D838E015FFB7B045FF5DC3C792137D79" LSID="urn:lsid:plazi:treatment:D838E015FFB7B045FF5DC3C792137D79" httpUri="http://treatment.plazi.org/id/D838E015FFB7B045FF5DC3C792137D79" lastPageId="22" lastPageNumber="223" pageId="19" pageNumber="220">
<subSubSection id="188B0288FFB7B040FF5DC3C790077EEE" box="[151,863,1188,1215]" pageId="19" pageNumber="220" type="nomenclature">
<paragraph id="502E5103FFB7B040FF5DC3C790077EEE" blockId="19.[151,863,1188,1250]" box="[151,863,1188,1215]" pageId="19" pageNumber="220">
<heading id="0B66E66FFFB7B040FF5DC3C790077EEE" bold="true" box="[151,863,1188,1215]" fontSize="11" level="1" pageId="19" pageNumber="220" reason="1">
<emphasis id="62E58D11FFB7B040FF5DC3C790077EEE" bold="true" box="[151,863,1188,1215]" pageId="19" pageNumber="220">
<taxonomicName id="97912A80FFB7B040FF5DC3C791A27EEE" authority="Rojas-Arias, Griswold &amp; Ayala." authorityName="Rojas-Arias, Griswold &amp; Ayala." authorityYear="2025" box="[151,762,1188,1215]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="cristina" status="sp. nov.">
<emphasis id="62E58D11FFB7B040FF5DC3C7923F7EEE" bold="true" box="[151,359,1188,1215]" italics="true" pageId="19" pageNumber="220">Heriades cristina</emphasis>
Rojas-Arias, Griswold &amp; Ayala.
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFB7B040FCC8C3C790077EEE" box="[770,863,1188,1214]" pageId="19" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFB7B040FF5DC3AB920A7EB2" box="[151,338,1224,1250]" pageId="19" pageNumber="220" type="description">
<paragraph id="502E5103FFB7B040FF5DC3AB920A7EB2" blockId="19.[151,863,1188,1250]" box="[151,338,1224,1250]" pageId="19" pageNumber="220">
(
<figureCitation id="C8AA4D86FFB7B040FF55C3AB93A07EB2" box="[159,248,1224,1250]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="19" pageNumber="220">Figs. 2I</figureCitation>
,
<figureCitation id="C8AA4D86FFB7B040FECEC3AB92127EB2" box="[260,330,1224,1250]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="20.[151,250,1068,1092]" captionStartId-1="21.[151,250,1943,1967]" captionTargetBox-0="[159,1428,157,1033]" captionTargetBox-1="[179,1395,190,1905]" captionTargetId-0="figure-22@20.[151,1436,149,1042]" captionTargetId-1="figure-18@21.[179,1408,179,1917]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURE 10. Known distribution records for Heriades cristina sp. nov." captionText-1="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="19" pageNumber="220">1011</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFB7B040FF5DC27392B07C1A" pageId="19" pageNumber="220" type="diagnosis">
<paragraph id="502E5103FFB7B040FF5DC27392B07C1A" blockId="19.[151,1437,1296,2043]" pageId="19" pageNumber="220">
<emphasis id="62E58D11FFB7B040FF5DC273924A7F7A" bold="true" box="[151,274,1296,1322]" pageId="19" pageNumber="220">Diagnosis.</emphasis>
Tergal fasciae with rows of thin, white hairs (
<figureCitation id="C8AA4D86FFB7B040FCD1C273902B7F7A" box="[795,883,1296,1322]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="19" pageNumber="220">Fig. 3L</figureCitation>
).
<emphasis id="62E58D11FFB7B040FC4DC27390997F7A" box="[903,961,1296,1322]" italics="true" pageId="19" pageNumber="220">Male</emphasis>
: antenna orange to clear amber, including scape, pedicel and F1 (
<figureCitation id="C8AA4D86FFB7B040FE6CC25791497F1E" box="[422,529,1332,1358]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="19" pageNumber="220">Fig. 11A</figureCitation>
); legs yellowish to clear amber; scutum with large punctation (0.3 times OD) (
<figureCitation id="C8AA4D86FFB7B040FF55C23B92507F22" box="[159,264,1368,1394]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="19" pageNumber="220">Fig. 11B</figureCitation>
); S5 with two rounded tufts of capitate hairs directed medially, and simple short hairs towards sides, with transverse, lobed, thin, light, subapical line (
<figureCitation id="C8AA4D86FFB7B040FD1EC21F90627FC6" box="[724,826,1404,1430]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="19" pageNumber="220">Fig. 11E</figureCitation>
); posterior margin of S6 produced ton obtuse-angled apex, with two subapical tufts of several, short, thin hairs, directed downward (
<figureCitation id="C8AA4D86FFB7B040FBD4C2C397D87FEA" box="[1054,1152,1440,1466]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="19" pageNumber="220">Fig. 11F</figureCitation>
), similar to
<taxonomicName id="97912A80FFB7B040FAC1C2C296CE7FEA" authorityName="Cockerell" authorityYear="1897" box="[1291,1430,1440,1466]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="62E58D11FFB7B040FAC1C2C296CE7FEA" box="[1291,1430,1440,1466]" italics="true" pageId="19" pageNumber="220">H. crucifera</emphasis>
</taxonomicName>
,
<taxonomicName id="97912A80FFB7B040FF5DC2A6925D7F8E" box="[151,261,1477,1502]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="62E58D11FFB7B040FF5DC2A6925D7F8E" box="[151,261,1477,1502]" italics="true" pageId="19" pageNumber="220">H. texana</emphasis>
</taxonomicName>
, and
<taxonomicName id="97912A80FFB7B040FE89C2A692EC7F8E" authorityName="Crawford" authorityYear="1913" box="[323,436,1476,1502]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="leavitti">
<emphasis id="62E58D11FFB7B040FE89C2A692EC7F8E" box="[323,436,1476,1502]" italics="true" pageId="19" pageNumber="220">H. leavitti</emphasis>
</taxonomicName>
but different in the rest of characters here mentioned.
<emphasis id="62E58D11FFB7B040FBD3C2A797357F8E" box="[1049,1133,1476,1502]" italics="true" pageId="19" pageNumber="220">Female</emphasis>
: outer surface of mandible without tubercle, clypeus with two tubercles and a soft depression between them, similar to
<taxonomicName id="97912A80FFB7B040FB5EC28A965C7C52" authorityName="Crawford" authorityYear="1913" box="[1172,1284,1512,1538]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="leavitti">
<emphasis id="62E58D11FFB7B040FB5EC28A965C7C52" box="[1172,1284,1512,1538]" italics="true" pageId="19" pageNumber="220">H. leavitti</emphasis>
</taxonomicName>
, but differing from this by having apical margin of clypeus with stronger emargination and larger punctures on T3 (0.7 times DOP) with bright integument.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFB7B045FF04C13791C07E09" lastPageId="22" lastPageNumber="223" pageId="19" pageNumber="220" type="description">
<paragraph id="502E5103FFB7B040FF04C13796427C3F" blockId="19.[151,1437,1296,2043]" box="[206,1306,1620,1647]" pageId="19" pageNumber="220">
<emphasis id="62E58D11FFB7B040FF04C13792F37C3E" bold="true" box="[206,427,1620,1647]" pageId="19" pageNumber="220">
Description.
<emphasis id="62E58D11FFB7B040FEADC13792FA7C3F" bold="true" box="[359,418,1620,1647]" italics="true" pageId="19" pageNumber="220">Male</emphasis>
:
</emphasis>
Body length,
<quantity id="9769FCE6FFB7B040FD80C13791F87C3F" box="[586,672,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="19" pageNumber="220" unit="mm" value="5.9">5.9 mm</quantity>
(5.35.9, n=5); forewing length,
<quantity id="9769FCE6FFB7B040FBDCC13797347C3F" box="[1046,1132,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="19" pageNumber="220" unit="mm" value="3.7">3.7 mm</quantity>
(3.43.7, n=5).
</paragraph>
<paragraph id="502E5103FFB7B047FF0DC11B975D7E84" blockId="19.[151,1437,1296,2043]" lastBlockId="20.[151,1437,1138,2029]" lastPageId="20" lastPageNumber="221" pageId="19" pageNumber="220">
<emphasis id="62E58D11FFB7B040FF0DC11B925C7CC2" box="[199,260,1656,1682]" italics="true" pageId="19" pageNumber="220">Head</emphasis>
. Length
<quantity id="9769FCE6FFB7B040FEADC11B92E47CC3" box="[359,444,1656,1683]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="220" unit="mm" value="1.5">1.5 mm</quantity>
(1.51.6, n=5), width
<quantity id="9769FCE6FFB7B040FD7FC11B90537CC3" box="[693,779,1656,1683]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="19" pageNumber="220" unit="mm" value="1.6">1.6 mm</quantity>
(1.51.7, n=5); eyes about 3.3 times longer than broad, 1.9 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FFB7B040FCE7C1FF90DA7CE7" box="[813,898,1692,1719]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="220" unit="mm" value="0.8">0.8 mm</quantity>
(0.70.8, n=5); upper interocular distance
<quantity id="9769FCE6FFB7B040FA96C1FF96C57CE7" box="[1372,1437,1692,1719]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="220" unit="mm" value="1.0">1 mm</quantity>
(0.91, n=5); interantennal distance 2.9 times antennocular distance; lateral ocelli large (
<figureCitation id="C8AA4D86FFB7B040FB4CC1A397867C8A" box="[1158,1246,1728,1754]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="19" pageNumber="220">Fig. 3C</figureCitation>
), 2.3 times DOP in interocellar area; interocellar distance 1.6 times OD, ocellocular distance 1 time OD, OPR 1 time OD; labrum without elevation; clypeus 1.6 times broader than long, in profile lower half convex and apical margin without emargination; juxtantennal carina absent but strongly angulated; antenna yellow (
<figureCitation id="C8AA4D86FFB7B040FBE2C04F97D77D16" box="[1064,1167,1836,1862]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="19" pageNumber="220">Fig. 11A</figureCitation>
); scape length 0.8 times alveolocellar distance, 2.3 times longer than wide; pedicel as long as broad, about 1.5 times longer than F1 and about 0.6 times longer than F2; F1 longer than broad, remaining flagellomeres 2.8 longer than first; F11 dark brown 2.6 times longer than broad.
<emphasis id="62E58D11FFB7B040FE1EC0FA910B7DE2" box="[468,595,1945,1970]" italics="true" pageId="19" pageNumber="220">Mesosoma.</emphasis>
Scutum 1.9 times longer than scutellum, 3 times longer than tegula, and 3.4 longer than axilla; intertegular distance
<quantity id="9769FCE6FFB7B040FD9FC0DF91F37D87" box="[597,683,1980,2007]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="220" unit="mm" value="1.1">1.1 mm</quantity>
(1.11.3, n=5); metanotum with transverse ridge, in posterior view straight; propodeum with large pits (
<figureCitation id="C8AA4D86FFB7B040FDF2C08391DF7DAA" box="[568,647,2016,2042]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="19" pageNumber="220">Fig. 3I</figureCitation>
),
<quantity id="9769FCE6FFB7B040FD57C08391AB7DAB" box="[669,755,2016,2043]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="220" unit="mm" value="0.1">0.1 mm</quantity>
long; legs yellow to amber.
<emphasis id="62E58D11FFB7B040FBF2C08297EF7DAA" box="[1080,1207,2017,2042]" italics="true" pageId="19" pageNumber="220">Metasoma.</emphasis>
T1
<quantity id="9769FCE6FFB7B040FB2EC08396637DAB" box="[1252,1339,2016,2043]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="19" pageNumber="220" unit="mm" value="1.4">1.4 mm</quantity>
broad (1.31.4, n=5); T3
<quantity id="9769FCE6FFB0B047FEB8C31192927EDD" box="[370,458,1138,1165]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="20" pageNumber="221" unit="mm" value="1.6">1.6 mm</quantity>
broad (1.51.6, n=5); T6 with posterior margin not projected; posterior margin of S1 produced to median rounded angle, without emargination (
<figureCitation id="C8AA4D86FFB0B047FCEDC3F590327EE0" box="[807,874,1174,1200]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="20" pageNumber="221">Fig. 2</figureCitation>
Ib), in profile, with posterior margin projected and low ventral angle more than one third of distance from apex to base (
<figureCitation id="C8AA4D86FFB0B047FC56C3D990877E84" box="[924,991,1210,1236]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="20" pageNumber="221">Fig. 2</figureCitation>
Ia).
</paragraph>
<caption id="04EE018BFFB0B047FF5DC34F90357E14" box="[151,877,1068,1092]" pageId="20" pageNumber="221" startId="20.[151,250,1068,1092]" targetBox="[159,1428,157,1033]" targetPageId="20" targetType="figure">
<paragraph id="502E5103FFB0B047FF5DC34F90357E14" blockId="20.[151,877,1068,1092]" box="[151,877,1068,1092]" pageId="20" pageNumber="221">
<emphasis id="62E58D11FFB0B047FF5DC34F92787E14" bold="true" box="[151,288,1068,1092]" pageId="20" pageNumber="221">FIGURE 10.</emphasis>
Known distribution records for
<taxonomicName id="97912A80FFB0B047FDADC34F904F7E14" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[615,791,1068,1092]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="cristina" status="sp. nov.">
<emphasis id="62E58D11FFB0B047FDADC34F904F7E14" box="[615,791,1068,1092]" italics="true" pageId="20" pageNumber="221">Heriades cristina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFB0B047FCD4C34E90357E14" box="[798,877,1069,1092]" pageId="20" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="502E5103FFB0B047FF0DC3BD922A7CA0" blockId="20.[151,1437,1138,2029]" pageId="20" pageNumber="221">
<emphasis id="62E58D11FFB0B047FF0DC3BD92097EA8" box="[199,337,1246,1272]" italics="true" pageId="20" pageNumber="221">Sculpturing.</emphasis>
Clypeus with small (at base 0.1 times OD), almost contiguous, coarse punctures; supraclypeal area with punctures slightly bigger, more rounded than base of clypeus, on frons slightly bigger, coarser and sparser punctures; vertex with bigger, sparser punctures than on supraclypeal area, microreticulate between punctures; hypostomal area with punctation smaller, sparser and less coarse than on clypeus; anterior surface of pronotum with punctures bigger and sparser than on clypeus, in pronotal lobes denser and coarser; disc of scutum with bigger punctures (0.3 times OD) than on vertex, with micropunctures between punctures; rest of mesosoma as on scutum except: scutellum with punctures sparser, mesepisternum with 1-time DOP than punctures on scutum, but sparser towards ventral surface; metepisternum with notably smaller punctures, less dense and coarse; T1 and T2 with smaller (0.2 times OD), denser punctures than on scutum, without micropunctures between punctures; T2 with punctures smaller and sparser than on T1; T3 with smaller and sparser punctures than on T1, slightly rounded to semicircular shaped, separated by more than half DOP; T4 and T5 with bigger, denser and coarser punctures than on T3; dorsal surface of T6 with smaller, irregularly shaped punctures as on T5, on posterior surface with notably smaller and slightly sparser punctation, DBP one half DOP; sides of S1 with punctation bigger and parser than T1, and smaller on medial process of S1; sides of S2 with bigger punctures than on T1 and disc of S2 microreticulate without punctation.
</paragraph>
<paragraph id="502E5103FFB0B047FF0DC19897BC7DBC" blockId="20.[151,1437,1138,2029]" pageId="20" pageNumber="221">
<emphasis id="62E58D11FFB0B047FF0DC19892687D44" box="[199,304,1787,1812]" italics="true" pageId="20" pageNumber="221">Vestiture.</emphasis>
Supraclypeal area with plumose, thin, white, short hairs directed medially; frons with plumose, thin, long, white hairs directed dorsally (
<figureCitation id="C8AA4D86FFB0B047FDF8C07D91C47D68" box="[562,668,1822,1848]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="20" pageNumber="221">Fig. 11A</figureCitation>
); dorsolateral angles of pronotum with thin, short, scattered, white hairs, not forming continuous, single, linear tuft with hairs on anterior margin of scutum; disc of scutum with simple, thin, short, scattered, white hairs on its entire surface; mesepisternum with plumose, thin, white, scattered hairs, denser on anterior and posterior margins; disc of metepisternum without tuft of thick, short, white hairs; tergal fasciae with row of thick, short, white hairs, decreasing in thickness towards posterior terga; disc of T1 to T6 with simple, long, thin, white hairs in punctures increasing in length towards posterior terga (
<figureCitation id="C8AA4D86FFB0B047FBBBC0B1978F7DBC" box="[1137,1239,2002,2028]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="20" pageNumber="221">Fig. 11B</figureCitation>
).
</paragraph>
<caption id="04EE018BFFB1B046FF5DC0F4907A7D83" pageId="21" pageNumber="222" startId="21.[151,250,1943,1967]" targetBox="[179,1395,190,1905]" targetPageId="21" targetType="figure">
<paragraph id="502E5103FFB1B046FF5DC0F4907A7D83" blockId="21.[151,1436,1943,2003]" pageId="21" pageNumber="222">
<emphasis id="62E58D11FFB1B046FF5DC0F492467DFF" bold="true" box="[151,286,1943,1967]" pageId="21" pageNumber="222">FIGURE 11.</emphasis>
<taxonomicName id="97912A80FFB1B046FEEEC0F4928C7DFF" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[292,468,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="cristina" status="sp. nov.">
<emphasis id="62E58D11FFB1B046FEEEC0F4928C7DFF" box="[292,468,1943,1967]" italics="true" pageId="21" pageNumber="222">Heriades cristina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFB1B046FE11C0FB91727DFF" box="[475,554,1944,1967]" pageId="21" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
Male:
<emphasis id="62E58D11FFB1B046FDBBC0F491D27DFF" bold="true" box="[625,650,1943,1967]" pageId="21" pageNumber="222">A.</emphasis>
frontal view
<emphasis id="62E58D11FFB1B046FCDBC0F490707DFF" bold="true" box="[785,808,1943,1967]" pageId="21" pageNumber="222">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FFB1B046FBCCC0F497467DFF" bold="true" box="[1030,1054,1943,1967]" pageId="21" pageNumber="222">C.</emphasis>
frontal view
<emphasis id="62E58D11FFB1B046FB6FC0F497E67DFF" bold="true" box="[1189,1214,1943,1967]" pageId="21" pageNumber="222">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FFB1B046FA4FC0F496C47DFF" bold="true" box="[1413,1436,1943,1967]" pageId="21" pageNumber="222">E.</emphasis>
S5.
<emphasis id="62E58D11FFB1B046FF75C0D8938A7D83" bold="true" box="[191,210,1979,2003]" pageId="21" pageNumber="222">F.</emphasis>
S6.
<emphasis id="62E58D11FFB1B046FECAC0D892427D83" bold="true" box="[256,282,1979,2003]" pageId="21" pageNumber="222">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FFB1B046FDB5C0D891C17D83" bold="true" box="[639,665,1979,2003]" pageId="21" pageNumber="222">H.</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="502E5103FFB2B045FF0DC7F497A47B4D" blockId="22.[151,1437,151,1833]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC7F492117AE1" box="[199,329,151,177]" italics="true" pageId="22" pageNumber="223">
<taxonomicName id="97912A80FFB2B045FF0DC7F4921D7AE1" box="[199,325,151,177]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="22" pageNumber="223" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
S6 with an obtuse-angled apex, with two subapical tufts of several, short, thin hairs, directed down, with blackish subapical margin on sides, not connected medially (
<figureCitation id="C8AA4D86FFB2B045FCB0C7D890857A85" box="[890,989,187,213]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="22" pageNumber="223">Fig. 11F</figureCitation>
); capsule in dorsal view
<quantity id="9769FCE6FFB2B045FB38C7D896117A85" box="[1266,1353,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="223" unit="mm" value="0.9">0.9 mm</quantity>
length, and
<quantity id="9769FCE6FFB2B045FF0DC7BD92457AA9" box="[199,285,222,249]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="223" unit="mm" value="0.5">0.5 mm</quantity>
width, 2.1 times longer than wide, gonobase
<quantity id="9769FCE6FFB2B045FCD0C7BC90277AA9" box="[794,895,223,249]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="22" pageNumber="223" unit="mm" value="0.07">0.07 mm</quantity>
length, 6 times wider than long, and gonocoxite
<quantity id="9769FCE6FFB2B045FF5DC66093B57B4D" box="[151,237,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="223" unit="mm" value="0.4">0.4 mm</quantity>
length, gonostylus
<quantity id="9769FCE6FFB2B045FE02C66091467B4D" box="[456,542,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="223" unit="mm" value="0.9">0.9 mm</quantity>
length with rounded apex, penis valve
<quantity id="9769FCE6FFB2B045FC1CC66097747B4D" box="[982,1068,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="223" unit="mm" value="0.6">0.6 mm</quantity>
length (
<figureCitation id="C8AA4D86FFB2B045FB42C66097B67B4D" box="[1160,1262,259,285]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="22" pageNumber="223">Fig. 11G</figureCitation>
).
</paragraph>
<paragraph id="502E5103FFB2B045FF0DC64592397831" blockId="22.[151,1437,151,1833]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC645927F7B11" bold="true" box="[199,295,294,321]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC64592467B11" bold="true" box="[199,286,294,321]" italics="true" pageId="22" pageNumber="223">Female</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FFB2B045FE0FC64491787B11" box="[453,544,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="22" pageNumber="223" unit="mm" value="6.8">6.8 mm</quantity>
; forewing length
<quantity id="9769FCE6FFB2B045FD20C64490687B11" box="[746,816,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="223" unit="mm" value="4.0">4 mm</quantity>
. Similar to male except:
<emphasis id="62E58D11FFB2B045FB9AC64497D57B11" box="[1104,1165,295,321]" italics="true" pageId="22" pageNumber="223">Head</emphasis>
. Length
<quantity id="9769FCE6FFB2B045FB3FC64596177B11" box="[1269,1359,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="223" unit="mm" value="1.5">1.5 mm</quantity>
, width
<quantity id="9769FCE6FFB2B045FF5DC62893AA7B35" box="[151,242,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="223" unit="mm" value="1.8">1.8 mm</quantity>
; eyes about 4.2 times longer than broad, 1.8 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FFB2B045FF5DC60C93AB7BD9" box="[151,243,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="223" unit="mm" value="0.9">0.9 mm</quantity>
; upper interocular distance
<quantity id="9769FCE6FFB2B045FDFEC60C91C87BD9" box="[564,656,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="223" unit="mm" value="1.1">1.1 mm</quantity>
; interantennal distance 1.7 times antennocular distance; interocellar distance 1.6 times OD, ocellocular distance 1.3 times OD, OPR 1.4 times OD; labrum with rounded, basal, medial tubercle; dorsal margin of mandible with carina; clypeus with medial emargination and depression, sided by two small tubercles and crenulated on sides of apical margin; scape 3.1 times longer than broad; pedicel 1.1 longer than broad, about 1.1 times longer than F1 and 2.2 longer than F2, remaining flagellomeres 2 times longer than F1.
<emphasis id="62E58D11FFB2B045FF5DC5409257786C" box="[151,271,547,572]" italics="true" pageId="22" pageNumber="223">Mesosoma</emphasis>
. Scutum 2.9 times longer than scutellum; intertegular distance
<quantity id="9769FCE6FFB2B045FC29C5409766786D" box="[995,1086,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="223" unit="mm" value="1.4">1.4 mm</quantity>
.
<emphasis id="62E58D11FFB2B045FB80C54097E7786C" box="[1098,1215,547,572]" italics="true" pageId="22" pageNumber="223">Metasoma</emphasis>
. T1
<quantity id="9769FCE6FFB2B045FB3FC5419615786D" box="[1269,1357,546,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="223" unit="mm" value="1.5">1.5 mm</quantity>
broad. T3
<quantity id="9769FCE6FFB2B045FF77C524924B7831" box="[189,275,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="223" unit="mm" value="1.8">1.8 mm</quantity>
broad.
</paragraph>
<paragraph id="502E5103FFB2B045FF0DC508924B79BD" blockId="22.[151,1437,151,1833]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC508920978D5" box="[199,337,619,645]" italics="true" pageId="22" pageNumber="223">Sculpturing.</emphasis>
Clypeus with medium-sized (0.2 times OD), contiguous, coarse, punctures, shiny and smooth between punctures; supraclypeal area with punctures as on base of clypeus; frons with bigger, coarser and denser with micropunctures between punctures than on supraclypeal area; vertex bigger and coarser punctures than on clypeus, with micropunctures between punctures; gena evenly punctate, bigger and denser than on frons in front of ocelli; scutum with bigger punctures than on vertex; rest of mesosoma as on scutum except: scutellum with slightly sparser punctures; mesepisternum with slightly sparser and irregularly shaped punctures; metasomal terga bright, microreticulate between punctures; T1 with smaller (0.11 times OD), slightly denser punctures than on scutum, with rounded shape; T2 with smaller, sparser than on T1, semicircular shaped punctures; T3 with punctures bigger (0.7 times DOP), notably less coarse, than on scutum, semicircular shaped, and sparser (1 time DOP); dorsal surface of T6 with smaller, denser, and more coarsely punctures than on T5; sterna with punctation smaller and sparser than on scutum.
</paragraph>
<paragraph id="502E5103FFB2B045FF0DC49491C07E09" blockId="22.[151,1437,151,1833]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC49492687E40" box="[199,304,1015,1040]" italics="true" pageId="22" pageNumber="223">Vestiture.</emphasis>
Sides of clypeus with plumose, white, hairs; disc of clypeus and frons with simple, short, white hairs in punctures (
<figureCitation id="C8AA4D86FFB2B045FEE5C37892CB7E65" box="[303,403,1051,1077]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="22" pageNumber="223">Fig. 11C</figureCitation>
); disc of T1 to T6 with hairs increasing in length towards posterior terga; posterior margin of T6 with transverse tuft of short, whitish hairs.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFB2B045FF0DC300904D7F91" pageId="22" pageNumber="223" type="materials_examined">
<paragraph id="502E5103FFB2B045FF0DC30092947F05" blockId="22.[151,1437,151,1833]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC300922F7E2C" bold="true" box="[199,375,1122,1149]" pageId="22" pageNumber="223">Type material:</emphasis>
<materialsCitation id="E0F95B5EFFB2B045FEB7C30091517EF1" collectingDate="1996-03-26" collectionCode="KUNHM-ENT" collectorName="R. Tut" country="Mexico" elevation="3" latitude="20.859167" location="Celestun" longLatPrecision="21" longitude="-90.39722" pageId="22" pageNumber="223" specimenCode="SEMC1387972" specimenCount="1" typeStatus="holotype">
<emphasis id="62E58D11FFB2B045FEB7C30090A37E2D" bold="true" box="[381,1019,1122,1149]" pageId="22" pageNumber="223">
<typeStatus id="8F2AEFA1FFB2B045FEB7C30092B27E2D" box="[381,490,1123,1149]" pageId="22" pageNumber="223" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="0037F978FFB2B045FDF2C30191AA7E2C" box="[568,754,1122,1148]" collectionCode="SEMC" country="USA" name="University of Kansas - Biodiversity Institute" pageId="22" pageNumber="223" type="Museum">SEMC1387972</specimenCode>
):
<collectingCountry id="28861193FFB2B045FCCCC30190D97E2C" box="[774,897,1122,1148]" name="Mexico" pageId="22" pageNumber="223">MEXICO</collectingCountry>
:
<location id="554E07D8FFB2B045FC46C30190AE7E2D" LSID="urn:lsid:plazi:treatment:D838E015FFB7B045FF5DC3C792137D79:554E07D8FFB2B045FC46C30190AE7E2D" box="[908,1014,1122,1149]" country="Mexico" latitude="20.859167" longLatPrecision="21" longitude="-90.39722" name="Celestun" pageId="22" pageNumber="223">Celestún</location>
,
</emphasis>
[
<quantity id="9769FCE6FFB2B045FBC0C300976F7E2D" box="[1034,1079,1123,1149]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="223" unit="m" value="3.0">
<elevation id="DBBCB630FFB2B045FBC0C300976F7E2D" box="[1034,1079,1123,1149]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="223" unit="m" value="3.0">3 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFB2B045FB89C30197947E2D" box="[1091,1228,1122,1149]" degrees="20" direction="north" minutes="51" orientation="latitude" pageId="22" pageNumber="223" precision="15" seconds="33" value="20.859167">20°5133”N</geoCoordinate>
<geoCoordinate id="35A537C4FFB2B045FB18C301963F7E2D" box="[1234,1383,1122,1149]" degrees="90" direction="west" minutes="23" orientation="longitude" pageId="22" pageNumber="223" precision="15" seconds="50" value="-90.39722">90°2350”W</geoCoordinate>
],
<date id="242F77C3FFB2B045FAB2C30093AE7EF1" pageId="22" pageNumber="223" value="1996-03-26">
<collectingDate id="346B8E2BFFB2B045FAB2C30093AE7EF1" pageId="22" pageNumber="223" value="1996-03-26">26/ III/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FFB2B045FF35C3E4921B7EF1" box="[255,323,1159,1185]" pageId="22" pageNumber="223">R. Tut</collectorName>
(
<collectionCode id="3680C9C6FFB2B045FE9AC3E491597EF1" box="[336,513,1159,1185]" pageId="22" pageNumber="223">KUNHM-ENT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFB2B045FDDBC3E591AB7E95" collectingDate="1996-04-25" collectionCode="KUNHM-ENT" collectorName="K. Tibbo" country="Mexico" elevation="24" latitude="20.709167" location="Yokdzonot" longLatPrecision="21" longitude="-88.73139" pageId="22" pageNumber="223" specimenCode="SEMC1387971" specimenCount="1" stateProvince="Yucatan" typeStatus="allotype">
<emphasis id="62E58D11FFB2B045FDDBC3E597647EF1" bold="true" box="[529,1084,1158,1185]" pageId="22" pageNumber="223">
<typeStatus id="8F2AEFA1FFB2B045FDDBC3E5912E7EF0" box="[529,630,1158,1184]" pageId="22" pageNumber="223" type="allotype">Allotype</typeStatus>
(
<specimenCode id="0037F978FFB2B045FD49C3E590657EF0" box="[643,829,1158,1184]" collectionCode="SEMC" country="USA" name="University of Kansas - Biodiversity Institute" pageId="22" pageNumber="223" type="Museum">SEMC1387971</specimenCode>
):
<collectingCountry id="28861193FFB2B045FC85C3E590927EF0" box="[847,970,1158,1184]" name="Mexico" pageId="22" pageNumber="223">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFB2B045FC18C3E5976E7EF1" box="[978,1078,1158,1185]" country="Mexico" name="Yucatan" pageId="22" pageNumber="223">Yucatán</collectingRegion>
,
</emphasis>
<location id="554E07D8FFB2B045FB8BC3E497E67EF1" LSID="urn:lsid:plazi:treatment:D838E015FFB7B045FF5DC3C792137D79:554E07D8FFB2B045FB8BC3E497E67EF1" box="[1089,1214,1159,1185]" country="Mexico" latitude="20.709167" longLatPrecision="21" longitude="-88.73139" name="Yokdzonot" pageId="22" pageNumber="223" stateProvince="Yucatan">Yokdzonot</location>
, [
<quantity id="9769FCE6FFB2B045FB04C3E496517EF1" box="[1230,1289,1159,1185]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="223" unit="m" value="24.0">
<elevation id="DBBCB630FFB2B045FB04C3E496517EF1" box="[1230,1289,1159,1185]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="223" unit="m" value="24.0">24 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFB2B045FAD9C3E496C47EF1" box="[1299,1436,1159,1185]" degrees="20" direction="north" minutes="42" orientation="latitude" pageId="22" pageNumber="223" precision="15" seconds="33" value="20.709167">20°4233”N</geoCoordinate>
<geoCoordinate id="35A537C4FFB2B045FF5DC3C992737E95" box="[151,299,1194,1221]" degrees="88" direction="west" minutes="43" orientation="longitude" pageId="22" pageNumber="223" precision="15" seconds="53" value="-88.73139">88°4353”W</geoCoordinate>
],
<date id="242F77C3FFB2B045FEF7C3C992987E95" box="[317,448,1194,1221]" pageId="22" pageNumber="223" value="1996-04-25">
<collectingDate id="346B8E2BFFB2B045FEF7C3C992987E95" box="[317,448,1194,1221]" pageId="22" pageNumber="223" value="1996-04-25">25/IV/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FFB2B045FE01C3C891757E95" box="[459,557,1195,1221]" pageId="22" pageNumber="223">K. Tibbo</collectorName>
(
<collectionCode id="3680C9C6FFB2B045FDF0C3C891B37E95" box="[570,747,1195,1221]" pageId="22" pageNumber="223">KUNHM-ENT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFB2B045FD36C3C890367F5D" collectingDate="1996-07-10" collectionCode="CER-UADY" collectorName="A. Canto &amp; C. Navarro &amp; F. Leon &amp; H. Contreras &amp; H. Navarro &amp; L. Novelo &amp; R. Rodriguez &amp; R. Tut &amp; S. Norena" country="Mexico" elevation="3" latitude="20.859304" location="Celestun" longLatPrecision="1" longitude="-90.3972" pageId="22" pageNumber="223" specimenCount="33" specimenCount-female="23" specimenCount-male="10" stateProvince="Yucatan" typeStatus="paratype">
<emphasis id="62E58D11FFB2B045FD36C3C897297E95" bold="true" box="[764,1137,1194,1221]" pageId="22" pageNumber="223">
<typeStatus id="8F2AEFA1FFB2B045FD36C3C890207E95" box="[764,888,1195,1221]" pageId="22" pageNumber="223" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="28861193FFB2B045FC49C3C990A67E94" box="[899,1022,1194,1220]" name="Mexico" pageId="22" pageNumber="223">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFB2B045FBCDC3C997337E95" box="[1031,1131,1194,1221]" country="Mexico" name="Yucatan" pageId="22" pageNumber="223">Yucatán</collectingRegion>
,
</emphasis>
<location id="554E07D8FFB2B045FBBDC3C897847E95" LSID="urn:lsid:plazi:treatment:D838E015FFB7B045FF5DC3C792137D79:554E07D8FFB2B045FBBDC3C897847E95" box="[1143,1244,1195,1221]" country="Mexico" latitude="20.859304" longLatPrecision="1" longitude="-90.3972" name="Celestun" pageId="22" pageNumber="223" stateProvince="Yucatan">Celestún</location>
, [
<quantity id="9769FCE6FFB2B045FB24C3C896437E95" box="[1262,1307,1195,1221]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="223" unit="m" value="3.0">
<elevation id="DBBCB630FFB2B045FB24C3C896437E95" box="[1262,1307,1195,1221]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="223" unit="m" value="3.0">3 m</elevation>
</quantity>
,
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<geoCoordinate id="35A537C4FFB2B045FF5DC3AD92457EB9" box="[151,285,1230,1257]" degrees="90.397205" direction="west" orientation="longitude" pageId="22" pageNumber="223" precision="1" value="-90.3972">-90.397205</geoCoordinate>
], 26/03, 16/IV, 07/V, 10, 18/VI,
<date id="242F77C3FFB2B045FDB3C3AC905D7EB9" box="[633,773,1231,1257]" pageId="22" pageNumber="223" value="1996-07-10">
<collectingDate id="346B8E2BFFB2B045FDB3C3AC905D7EB9" box="[633,773,1231,1257]" pageId="22" pageNumber="223" value="1996-07-10">10/VII/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FFB2B045FCC4C3AC902C7EB9" box="[782,884,1231,1257]" pageId="22" pageNumber="223">A. Canto</collectorName>
,
<collectorName id="FD6434D5FFB2B045FCB7C3AC90A37EB9" box="[893,1019,1231,1257]" pageId="22" pageNumber="223">C. Navarro</collectorName>
,
<collectorName id="FD6434D5FFB2B045FBCEC3AC97017EB9" box="[1028,1113,1231,1257]" pageId="22" pageNumber="223">F. León</collectorName>
,
<collectorName id="FD6434D5FFB2B045FBA9C3AC97AB7EB9" box="[1123,1267,1231,1257]" pageId="22" pageNumber="223">H. Contreras</collectorName>
,
<collectorName id="FD6434D5FFB2B045FB36C3AC96237EB9" box="[1276,1403,1231,1257]" pageId="22" pageNumber="223">H. Navarro</collectorName>
,
<collectorName id="FD6434D5FFB2B045FA4EC3AC93B47F5D" pageId="22" pageNumber="223">L. Novelo</collectorName>
,
<collectorName id="FD6434D5FFB2B045FF3CC39092D57F5D" box="[246,397,1267,1293]" pageId="22" pageNumber="223">R. Rodriguez</collectorName>
,
<collectorName id="FD6434D5FFB2B045FE5DC39092867F5D" box="[407,478,1267,1293]" pageId="22" pageNumber="223">R. Tut</collectorName>
,
<collectorName id="FD6434D5FFB2B045FE22C390910E7F5D" box="[488,598,1267,1293]" pageId="22" pageNumber="223">S. Noreña</collectorName>
(
<collectionCode id="3680C9C6FFB2B045FDA9C39091AD7F5D" box="[611,757,1267,1293]" pageId="22" pageNumber="223">CER-UADY</collectionCode>
<specimenCount id="46979A8AFFB2B045FD31C39090747F5D" box="[763,812,1267,1293]" count="23" pageId="22" pageNumber="223" type="female">23♀</specimenCount>
<specimenCount id="46979A8AFFB2B045FCFBC390903E7F5D" box="[817,870,1267,1293]" count="10" pageId="22" pageNumber="223" type="male">10♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFB2B045FCB2C39091897F61" collectingDate="1995-05-24" collectionCode="CER-UADY" collectorName="El Cuyo &amp; H. Moo &amp; R. Tut" country="Mexico" elevation="6" latitude="21.515833" location="El Cuyo" longLatPrecision="1" longitude="-87.67833" pageId="22" pageNumber="223" specimenCount="9" specimenCount-female="7" specimenCount-male="2" stateProvince="Yucatan" typeStatus="paratype">
[Mérida],
<collectorName id="FD6434D5FFB2B045FC22C390971E7F5D" box="[1000,1094,1267,1293]" pageId="22" pageNumber="223">
<location id="554E07D8FFB2B045FC22C390971E7F5D" LSID="urn:lsid:plazi:treatment:D838E015FFB7B045FF5DC3C792137D79:554E07D8FFB2B045FC22C390971E7F5D" box="[1000,1094,1267,1293]" country="Mexico" latitude="21.515833" longLatPrecision="1" longitude="-87.67833" name="El Cuyo" pageId="22" pageNumber="223" stateProvince="Yucatan">El Cuyo</location>
</collectorName>
, [
<quantity id="9769FCE6FFB2B045FB9DC39097DD7F5D" box="[1111,1157,1267,1293]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="223" unit="m" value="6.0">
<elevation id="DBBCB630FFB2B045FB9DC39097DD7F5D" box="[1111,1157,1267,1293]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="223" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFB2B045FB45C391965E7F5D" box="[1167,1286,1266,1293]" degrees="21.515833" direction="north" orientation="latitude" pageId="22" pageNumber="223" precision="1" value="21.515833">21.515833</geoCoordinate>
<geoCoordinate id="35A537C4FFB2B045FAC6C39096C97F5D" box="[1292,1425,1267,1293]" degrees="87.678333" direction="west" orientation="longitude" pageId="22" pageNumber="223" precision="1" value="-87.67833">-87.678333</geoCoordinate>
],
<date id="242F77C3FFB2B045FF5DC27592497F61" box="[151,273,1302,1329]" pageId="22" pageNumber="223" value="1995-05-24">
<collectingDate id="346B8E2BFFB2B045FF5DC27592497F61" box="[151,273,1302,1329]" pageId="22" pageNumber="223" value="1995-05-24">24/V/1995</collectingDate>
</date>
,
<collectorName id="FD6434D5FFB2B045FED4C27492227F61" box="[286,378,1303,1329]" pageId="22" pageNumber="223">H. Moo</collectorName>
,
<collectorName id="FD6434D5FFB2B045FE4CC27492957F61" box="[390,461,1303,1329]" pageId="22" pageNumber="223">R. Tut</collectorName>
(
<collectionCode id="3680C9C6FFB2B045FE16C27491367F61" box="[476,622,1303,1329]" pageId="22" pageNumber="223">CER-UADY</collectionCode>
<specimenCount id="46979A8AFFB2B045FDBCC27491C27F61" box="[630,666,1303,1329]" count="7" pageId="22" pageNumber="223" type="female">7♀</specimenCount>
<specimenCount id="46979A8AFFB2B045FD68C274919F7F61" box="[674,711,1303,1329]" count="2" pageId="22" pageNumber="223" type="male">2♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFB2B045FD16C274929F7F05" collectingDate="1996-04-25" collectionCode="KUNHM-ENT" collectorName="K. Tibbo" country="Mexico" elevation="24" latitude="20.709167" location="Yokdzonot" longLatPrecision="21" longitude="-88.73139" pageId="22" pageNumber="223" specimenCount="1" specimenCount-female="1" stateProvince="Yucatan" typeStatus="paratype">
<location id="554E07D8FFB2B045FD16C27490017F61" LSID="urn:lsid:plazi:treatment:D838E015FFB7B045FF5DC3C792137D79:554E07D8FFB2B045FD16C27490017F61" box="[732,857,1303,1329]" country="Mexico" latitude="20.709167" longLatPrecision="21" longitude="-88.73139" name="Yokdzonot" pageId="22" pageNumber="223" stateProvince="Yucatan">Yokdzonot</location>
, [
<quantity id="9769FCE6FFB2B045FCA6C27490F27F61" box="[876,938,1303,1329]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="223" unit="m" value="24.0">
<elevation id="DBBCB630FFB2B045FCA6C27490F27F61" box="[876,938,1303,1329]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="223" unit="m" value="24.0">24 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFB2B045FC7DC27497187F61" box="[951,1088,1303,1329]" degrees="20" direction="north" minutes="42" orientation="latitude" pageId="22" pageNumber="223" precision="15" seconds="33" value="20.709167">20°4233”N</geoCoordinate>
<geoCoordinate id="35A537C4FFB2B045FB83C27597847F61" box="[1097,1244,1302,1329]" degrees="88" direction="west" minutes="43" orientation="longitude" pageId="22" pageNumber="223" precision="15" seconds="53" value="-88.73139">88°4353”W</geoCoordinate>
],
<date id="242F77C3FFB2B045FB3BC275962C7F61" box="[1265,1396,1302,1329]" pageId="22" pageNumber="223" value="1996-04-25">
<collectingDate id="346B8E2BFFB2B045FB3BC275962C7F61" box="[1265,1396,1302,1329]" pageId="22" pageNumber="223" value="1996-04-25">25/IV/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FFB2B045FA4BC27493817F05" pageId="22" pageNumber="223">K. Tibbo</collectorName>
(
<collectionCode id="3680C9C6FFB2B045FF2DC25892CA7F05" box="[231,402,1339,1365]" pageId="22" pageNumber="223">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFFB2B045FE53C25892E37F05" box="[409,443,1339,1365]" count="1" pageId="22" pageNumber="223" type="female">1♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="502E5103FFB2B045FF0DC23D904D7F91" blockId="22.[151,1437,151,1833]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC23D916A7F28" bold="true" box="[199,562,1374,1400]" pageId="22" pageNumber="223">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFFB2B045FDFDC23D90547FCD" collectingDate="1952-08-03" collectionCode="BERK" collectorName="E. E. Gilbert &amp; C. D. MacNeil" country="Mexico" elevation="1694" latitude="16.27389" location="Comitan" longLatPrecision="21" longitude="-92.13139" pageId="22" pageNumber="223" specimenCount="2" specimenCount-female="2" stateProvince="Chiapas">
<emphasis id="62E58D11FFB2B045FDFDC23D907C7F28" bold="true" box="[567,804,1374,1400]" pageId="22" pageNumber="223">
<collectingCountry id="28861193FFB2B045FDFDC23D91EA7F28" box="[567,690,1374,1400]" name="Mexico" pageId="22" pageNumber="223">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFB2B045FD71C23D90477F28" box="[699,799,1374,1400]" country="Mexico" name="Chiapas" pageId="22" pageNumber="223">Chiapas</collectingRegion>
,
</emphasis>
<location id="554E07D8FFB2B045FCE3C23C90D47F29" LSID="urn:lsid:plazi:treatment:D838E015FFB7B045FF5DC3C792137D79:554E07D8FFB2B045FCE3C23C90D47F29" box="[809,908,1375,1401]" country="Mexico" latitude="16.27389" longLatPrecision="21" longitude="-92.13139" name="Comitan" pageId="22" pageNumber="223" stateProvince="Chiapas">Comitán</location>
, 16miNW, [
<quantity id="9769FCE6FFB2B045FBC7C23C973D7F29" box="[1037,1125,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.694" pageId="22" pageNumber="223" unit="m" value="1694.0">
<elevation id="DBBCB630FFB2B045FBC7C23C973D7F29" box="[1037,1125,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.694" pageId="22" pageNumber="223" unit="m" value="1694.0">1694 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFB2B045FBA4C23C97AF7F29" box="[1134,1271,1375,1401]" degrees="16" direction="north" minutes="16" orientation="latitude" pageId="22" pageNumber="223" precision="15" seconds="26" value="16.27389">16°1626”N</geoCoordinate>
<geoCoordinate id="35A537C4FFB2B045FB36C23D96C97F29" box="[1276,1425,1374,1401]" degrees="92" direction="west" minutes="07" orientation="longitude" pageId="22" pageNumber="223" precision="15" seconds="53" value="-92.13139">92°0753”W</geoCoordinate>
],
<date id="242F77C3FFB2B045FF5DC2E192787FCD" box="[151,288,1410,1437]" pageId="22" pageNumber="223" value="1952-08-03">
<collectingDate id="346B8E2BFFB2B045FF5DC2E192787FCD" box="[151,288,1410,1437]" pageId="22" pageNumber="223" value="1952-08-03">3/VIII/1952</collectingDate>
</date>
,
<collectorName id="FD6434D5FFB2B045FEE7C2E092997FCD" box="[301,449,1411,1437]" pageId="22" pageNumber="223">E. E. Gilbert</collectorName>
,
<collectorName id="FD6434D5FFB2B045FE04C2E0912F7FCD" box="[462,631,1411,1437]" pageId="22" pageNumber="223">C. D. MacNeil</collectorName>
(
<collectionCode id="3680C9C6FFB2B045FD42C2E0918C7FCD" box="[648,724,1411,1437]" pageId="22" pageNumber="223">BERK</collectionCode>
<specimenCount id="46979A8AFFB2B045FD17C2E0905A7FCD" box="[733,770,1411,1437]" count="2" pageId="22" pageNumber="223" type="female">2♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFB2B045FCD0C2E0904A7F91" collectingDate="2015-06-23" collectingDateMax="2018-03-14" collectingDateMin="2015-06-23" collectionCode="ECOAB" collectorName="N. Arnold &amp; A. Tapia" country="Mexico" elevation="17" location="Comalcaco" pageId="22" pageNumber="223" specimenCount="3" specimenCount-male="3" stateProvince="Tabasco">
<emphasis id="62E58D11FFB2B045FCD0C2E090D97FCD" bold="true" box="[794,897,1411,1437]" pageId="22" pageNumber="223">
<collectingRegion id="92559FE1FFB2B045FCD0C2E090257FCD" box="[794,893,1411,1437]" country="Mexico" name="Tabasco" pageId="22" pageNumber="223">Tabasco</collectingRegion>
,
</emphasis>
<location id="554E07D8FFB2B045FC41C2E097547FCD" LSID="urn:lsid:plazi:treatment:D838E015FFB7B045FF5DC3C792137D79:554E07D8FFB2B045FC41C2E097547FCD" box="[907,1036,1411,1437]" country="Mexico" name="Comalcaco" pageId="22" pageNumber="223" stateProvince="Tabasco">Comalcaco</location>
,
<quantity id="9769FCE6FFB2B045FBD3C2E0970E7FCD" box="[1049,1110,1411,1437]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="22" pageNumber="223" unit="m" value="17.0">
<elevation id="DBBCB630FFB2B045FBD3C2E0970E7FCD" box="[1049,1110,1411,1437]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="22" pageNumber="223" unit="m" value="17.0">17 m</elevation>
</quantity>
, 18.30629 -93.26,
<quantity id="9769FCE6FFB2B045FAE4C2E096337FCD" box="[1326,1387,1411,1437]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="223" unit="m" value="13.0">
<elevation id="DBBCB630FFB2B045FAE4C2E096337FCD" box="[1326,1387,1411,1437]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="223" unit="m" value="13.0">13 m</elevation>
</quantity>
,
<date id="242F77C3FFB2B045FAB2C2E093AF7F91" pageId="22" pageNumber="223" value="2015-06-23">
<collectingDate id="346B8E2BFFB2B045FAB2C2E093AF7F91" pageId="22" pageNumber="223" value="2015-06-23">23/ VI/2015</collectingDate>
</date>
,
<collectorName id="FD6434D5FFB2B045FEC9C2C4922D7F91" box="[259,373,1447,1473]" pageId="22" pageNumber="223">N. Arnold</collectorName>
,
<date id="242F77C3FFB2B045FE4BC2C4915B7F91" box="[385,515,1447,1473]" pageId="22" pageNumber="223" value="2018-03-14">
<collectingDate id="346B8E2BFFB2B045FE4BC2C4915B7F91" box="[385,515,1447,1473]" pageId="22" pageNumber="223" value="2018-03-14">14/III/2018</collectingDate>
</date>
,
<collectorName id="FD6434D5FFB2B045FDC6C2C491347F91" box="[524,620,1447,1473]" pageId="22" pageNumber="223">A. Tapia</collectorName>
(
<collectionCode id="3680C9C6FFB2B045FDB0C2C491837F91" box="[634,731,1447,1473]" pageId="22" pageNumber="223">ECOAB</collectionCode>
<specimenCount id="46979A8AFFB2B045FD28C2C490507F91" box="[738,776,1447,1473]" count="3" pageId="22" pageNumber="223" type="male">3♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFB2B045FF0DC2A992207C7D" pageId="22" pageNumber="223" type="distribution">
<paragraph id="502E5103FFB2B045FF0DC2A992207C7D" blockId="22.[151,1437,151,1833]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC2A992397FB4" bold="true" box="[199,353,1482,1508]" pageId="22" pageNumber="223">Distribution.</emphasis>
This species is present in the
<collectingRegion id="92559FE1FFB2B045FD6CC2A890587FB5" box="[678,768,1483,1509]" country="Mexico" name="Yucatan" pageId="22" pageNumber="223">Yucatán</collectingRegion>
Peninsula,
<collectingRegion id="92559FE1FFB2B045FCB5C2A890857FB5" box="[895,989,1483,1509]" country="Mexico" name="Tabasco" pageId="22" pageNumber="223">Tabasco</collectingRegion>
, and
<collectingRegion id="92559FE1FFB2B045FBDFC2A8972A7FB5" box="[1045,1138,1483,1509]" country="Mexico" name="Chiapas" pageId="22" pageNumber="223">Chiapas</collectingRegion>
, being endemic to
<collectingCountry id="28861193FFB2B045FA8AC2A896C07FB5" box="[1344,1432,1483,1509]" name="Mexico" pageId="22" pageNumber="223">Mexico</collectingCountry>
, in the
<collectingRegion id="92559FE1FFB2B045FF10C28C926C7C59" box="[218,308,1519,1545]" country="Mexico" name="Yucatan" pageId="22" pageNumber="223">Yucatán</collectingRegion>
Peninsula and Veracruzan biogeographic provinces (
<bibRefCitation id="34002CF2FFB2B045FCB4C28C97067C59" author="Morrone, J. J. &amp; Escalante, T. &amp; Rodriguez-Tapia, G. &amp; Carmona, A. &amp; Arana, M. &amp; Mercado-Gomez, J. D." box="[894,1118,1519,1545]" pageId="22" pageNumber="223" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. &amp; Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="62E58D11FFB2B045FC2DC28C974F7C59" box="[999,1047,1519,1545]" italics="true" pageId="22" pageNumber="223">et al</emphasis>
. 2022
</bibRefCitation>
) (
<figureCitation id="C8AA4D86FFB2B045FBB9C28C979E7C59" box="[1139,1222,1519,1545]" captionStart="FIGURE 10" captionStartId="20.[151,250,1068,1092]" captionTargetBox="[159,1428,157,1033]" captionTargetId="figure-22@20.[151,1436,149,1042]" captionTargetPageId="20" captionText="FIGURE 10. Known distribution records for Heriades cristina sp. nov." pageId="22" pageNumber="223">Fig. 10</figureCitation>
). Altitudinal range: sea level to
<quantity id="9769FCE6FFB2B045FED6C170922C7C7D" box="[284,372,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="1.694" pageId="22" pageNumber="223" unit="m" value="1694.0">1694 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFB2B045FF0DC15491CF7CC9" pageId="22" pageNumber="223" type="etymology">
<paragraph id="502E5103FFB2B045FF0DC15491CF7CC9" blockId="22.[151,1437,151,1833]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC15492167C01" bold="true" box="[199,334,1591,1617]" pageId="22" pageNumber="223">Etymology.</emphasis>
This species is dedicated to M. Sc. Cristina Mayorga, who contributed greatly to the taxonomy of Hemiptera of
<collectingRegion id="92559FE1FFB2B045FE90C13892E87C25" box="[346,432,1627,1653]" country="Mexico" name="Mexico" pageId="22" pageNumber="223">Mexico</collectingRegion>
and serves as a technician of the National Collection of Insects CNIN of the National Autonomous University of
<collectingRegion id="92559FE1FFB2B045FE00C11C91787CC9" box="[458,544,1663,1689]" country="Mexico" name="Mexico" pageId="22" pageNumber="223">Mexico</collectingRegion>
(UNAM).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFB2B045FF0DC1C192137D79" pageId="22" pageNumber="223" type="discussion">
<paragraph id="502E5103FFB2B045FF0DC1C192137D79" blockId="22.[151,1437,151,1833]" pageId="22" pageNumber="223">
<emphasis id="62E58D11FFB2B045FF0DC1C192177CEC" bold="true" box="[199,335,1698,1724]" pageId="22" pageNumber="223">Comments.</emphasis>
The DBP on supraclypeal area of males vary from 0.25 to 0.17. The length and number of hairs on scape in males also vary from short and scarce to long and dense. This species can be easily recognized by large punctures on scutum. Males by yellow antennae and dense number of hairs on hypostomal area, clypeus and fore tarsal segments.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,610 @@
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<mods:title id="962451B31C047EA224E5E1CCC70EF995">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<treatment id="D838E015FFBBB072FF5DC1CA96137EB8" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8" httpUri="http://treatment.plazi.org/id/D838E015FFBBB072FF5DC1CA96137EB8" lastPageId="33" lastPageNumber="234" pageId="31" pageNumber="232">
<subSubSection id="188B0288FFBBB04CFF5DC1CA91727C93" box="[151,554,1705,1732]" pageId="31" pageNumber="232" type="nomenclature">
<paragraph id="502E5103FFBBB04CFF5DC1CA91727C93" blockId="31.[151,554,1705,1767]" box="[151,554,1705,1732]" pageId="31" pageNumber="232">
<heading id="0B66E66FFFBBB04CFF5DC1CA91727C93" bold="true" box="[151,554,1705,1732]" fontSize="11" level="1" pageId="31" pageNumber="232" reason="1">
<taxonomicName id="97912A80FFBBB04CFF5DC1CA91727C93" ID-CoL="3KYRS" authority="Crawford, 1913" authorityName="Crawford" authorityYear="1913" box="[151,554,1705,1732]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="leavitti">
<emphasis id="62E58D11FFBBB04CFF5DC1CA91727C93" bold="true" box="[151,554,1705,1732]" pageId="31" pageNumber="232">
<emphasis id="62E58D11FFBBB04CFF5DC1CA92057C94" bold="true" box="[151,349,1705,1732]" italics="true" pageId="31" pageNumber="232">Heriades leavitti</emphasis>
Crawford, 1913
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBBB04CFF5DC1AE92327DD6" pageId="31" pageNumber="232" type="description">
<paragraph id="502E5103FFBBB04CFF5DC1AE920F7CB7" blockId="31.[151,554,1705,1767]" box="[151,343,1741,1767]" pageId="31" pageNumber="232">
(
<figureCitation id="C8AA4D86FFBBB04CFF55C1AE93A47CB7" box="[159,252,1741,1767]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="31" pageNumber="232">Figs. 2F</figureCitation>
,
<figureCitation id="C8AA4D86FFBBB04CFEC2C1AE92177CB7" box="[264,335,1741,1767]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStartId-0="31.[151,250,1098,1122]" captionStartId-1="32.[151,250,1943,1967]" captionTargetBox-0="[159,1428,187,1063]" captionTargetBox-1="[163,1412,184,1905]" captionTargetId-0="figure-18@31.[151,1436,179,1072]" captionTargetId-1="figure-22@32.[163,1425,179,1917]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="FIGURE 18. Known distribution records for Heriades leavitti." captionText-1="FIGURE 19. Heriades leavitti. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="31" pageNumber="232">1819</figureCitation>
)
</paragraph>
<paragraph id="502E5103FFBBB04CFF5DC07192327DD6" blockId="31.[151,1436,1810,1987]" pageId="31" pageNumber="232">
<taxonomicName id="97912A80FFBBB04CFF5DC07192B77D7A" authority="Crawford, 1913" authorityName="Crawford" authorityYear="1913" box="[151,495,1810,1834]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="leavitti">
<emphasis id="62E58D11FFBBB04CFF5DC071921A7D7A" box="[151,322,1810,1834]" italics="true" pageId="31" pageNumber="232">Heriades leavitti</emphasis>
<materialsCitation id="E0F95B5EFFBBB04CFE80C07192B77D7A" box="[330,495,1810,1834]" location="The Canadian Entomologist" municipality="Some" pageId="31" pageNumber="232" specimenCount="2" stateProvince="New Brunswick">Crawford, 1913</materialsCitation>
</taxonomicName>
.
<collectingMunicipality id="B04ACB79FFBBB04CFE30C071916C7D7A" box="[506,564,1810,1834]" pageId="31" pageNumber="232">Some</collectingMunicipality>
bees from the
<collectingRegion id="92559FE1FFBBB04CFD1AC07190207D7A" box="[720,888,1810,1834]" country="Canada" name="New Brunswick" pageId="31" pageNumber="232">New Brunswick</collectingRegion>
, with descriptions of a new species of
<taxonomicName id="97912A80FFBBB04CFAC1C07196307D7A" authorityName="Spinola" authorityYear="1808" box="[1291,1384,1810,1834]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis id="62E58D11FFBBB04CFAC1C07196307D7A" box="[1291,1384,1810,1834]" italics="true" pageId="31" pageNumber="232">Heriades</emphasis>
</taxonomicName>
.
<emphasis id="62E58D11FFBBB04CFABCC07192E07D18" italics="true" pageId="31" pageNumber="232">
<location id="554E07D8FFBBB04CFABCC07192E07D18" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FFBBB04CFABCC07192E07D18" municipality="Some" name="The Canadian Entomologist" pageId="31" pageNumber="232" stateProvince="New Brunswick">The Canadian Entomologist</location>
</emphasis>
:270.
<typeStatus id="8F2AEFA1FFBBB04CFF0DC02C93A17D37" box="[199,249,1871,1895]" pageId="31" pageNumber="232">Type</typeStatus>
locality: Nerepis,
<collectingRegion id="92559FE1FFBBB04CFE7FC02C91047D37" box="[437,604,1871,1895]" country="Canada" name="New Brunswick" pageId="31" pageNumber="232">New Brunswick</collectingRegion>
.
<typeStatus id="8F2AEFA1FFBBB04CFF0DC00D93A17DD6" box="[199,249,1902,1926]" pageId="31" pageNumber="232">Type</typeStatus>
in USNM.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBBB04CFF5DC0EF910F7D93" pageId="31" pageNumber="232" type="reference_group">
<paragraph id="502E5103FFBBB04CFF5DC0EF911F7DF4" blockId="31.[151,1436,1810,1987]" box="[151,583,1932,1956]" pageId="31" pageNumber="232">
<taxonomicName id="97912A80FFBBB04CFF5DC0EF911B7DF4" ID-CoL="8SG9J" authority="Graenicher, 1928: 281" authorityName="Graenicher" authorityPageNumber="281" authorityYear="1928" box="[151,579,1932,1956]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="crawfordi">
<emphasis id="62E58D11FFBBB04CFF5DC0EF92057DF4" box="[151,349,1932,1956]" italics="true" pageId="31" pageNumber="232">Heriades crawfordi</emphasis>
Graenicher, 1928:281
</taxonomicName>
.
</paragraph>
<paragraph id="502E5103FFBBB04CFF5DC0CF910F7D93" blockId="31.[151,1436,1810,1987]" box="[151,599,1963,1987]" pageId="31" pageNumber="232">
<taxonomicName id="97912A80FFBBB04CFF5DC0CF910B7D93" authority="Robertson, 1929: 131" authorityName="Robertson" authorityPageNumber="131" authorityYear="1929" box="[151,595,1963,1987]" class="Insecta" family="Megachilidae" genus="Neotrypetes" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="truncates">
<emphasis id="62E58D11FFBBB04CFF5DC0CF922D7D93" box="[151,373,1964,1987]" italics="true" pageId="31" pageNumber="232">Neotrypetes truncates</emphasis>
Robertson, 1929:131
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="04EE018BFF84B073FF5DC0F491B97D83" pageId="32" pageNumber="233" startId="32.[151,250,1943,1967]" targetBox="[163,1412,184,1905]" targetPageId="32" targetType="figure">
<paragraph id="502E5103FF84B073FF5DC0F491B97D83" blockId="32.[151,1436,1943,2003]" pageId="32" pageNumber="233">
<emphasis id="62E58D11FF84B073FF5DC0F492787DFF" bold="true" box="[151,288,1943,1967]" pageId="32" pageNumber="233">FIGURE 19.</emphasis>
<emphasis id="62E58D11FF84B073FEEDC0F492807DFF" box="[295,472,1943,1967]" italics="true" pageId="32" pageNumber="233">
<taxonomicName id="97912A80FF84B073FEEDC0F4928D7DFF" authorityName="Crawford" authorityYear="1913" box="[295,469,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="leavitti">Heriades leavitti</taxonomicName>
.
</emphasis>
Male:
<emphasis id="62E58D11FF84B073FDEBC0F491617DFF" bold="true" box="[545,569,1943,1967]" pageId="32" pageNumber="233">A.</emphasis>
frontal view
<emphasis id="62E58D11FF84B073FD09C0F491827DFF" bold="true" box="[707,730,1943,1967]" pageId="32" pageNumber="233">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FF84B073FC76C0F4908C7DFF" bold="true" box="[956,980,1943,1967]" pageId="32" pageNumber="233">C.</emphasis>
frontal view
<emphasis id="62E58D11FF84B073FB95C0F4972F7DFF" bold="true" box="[1119,1143,1943,1967]" pageId="32" pageNumber="233">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FF84B073FA88C0F496027DFF" bold="true" box="[1346,1370,1943,1967]" pageId="32" pageNumber="233">E.</emphasis>
S5.
<emphasis id="62E58D11FF84B073FA43C0F496C47DFF" bold="true" box="[1417,1436,1943,1967]" pageId="32" pageNumber="233">F.</emphasis>
S6.
<emphasis id="62E58D11FF84B073FF75C0D893817D83" bold="true" box="[191,217,1979,2003]" pageId="32" pageNumber="233">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FF84B073FDF7C0D891007D83" bold="true" box="[573,600,1979,2003]" pageId="32" pageNumber="233">H.</emphasis>
ventral view.
</paragraph>
</caption>
<subSubSection id="188B0288FF85B072FF5DC7F590AB7998" pageId="33" pageNumber="234" type="diagnosis">
<paragraph id="502E5103FF85B072FF5DC7F596D57B80" blockId="33.[151,1437,150,1257]" pageId="33" pageNumber="234">
<emphasis id="62E58D11FF85B072FF5DC7F5924A7AE0" bold="true" box="[151,274,150,176]" pageId="33" pageNumber="234">Diagnosis.</emphasis>
T3 with punctation medium-sized (0.7 times DOP as on scutum), and sparse punctures (1 time DOP) (
<figureCitation id="C8AA4D86FF85B072FF55C7D993A87A84" box="[159,240,186,212]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="33" pageNumber="234">Fig. 3J</figureCitation>
); tergal fasciae with rows of thin, white hairs (
<figureCitation id="C8AA4D86FF85B072FD35C7D9900F7A84" box="[767,855,186,212]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="33" pageNumber="234">Fig. 3L</figureCitation>
); wings usually dusky.
<emphasis id="62E58D11FF85B072FB95C7D997C17A84" box="[1119,1177,186,212]" italics="true" pageId="33" pageNumber="234">Male</emphasis>
: large ocelli (
<figureCitation id="C8AA4D86FF85B072FAFFC7D996D67A84" box="[1333,1422,186,212]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="33" pageNumber="234">Fig. 3C</figureCitation>
); posterior side of malar area rounded, not produced to angle; posterior margin of S5 with two rectangular lobes, with two rounded tufts of capitate hairs directed inward, and simple short hairs directed outward, with transverse subapical line (
<figureCitation id="C8AA4D86FF85B072FE80C64592EB7B10" box="[330,435,294,320]" captionStart="FIGURE 19" captionStartId="32.[151,250,1943,1967]" captionTargetBox="[163,1412,184,1905]" captionTargetId="figure-22@32.[163,1425,179,1917]" captionTargetPageId="32" captionText="FIGURE 19. Heriades leavitti. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="33" pageNumber="234">Fig. 19E</figureCitation>
); S6 with posterior margin produced to right-angled apex, without blackish subapical margin, with two tufts of simple, short, thin hairs, directed downward (
<figureCitation id="C8AA4D86FF85B072FC7AC629974B7B34" box="[944,1043,330,356]" captionStart="FIGURE 19" captionStartId="32.[151,250,1943,1967]" captionTargetBox="[163,1412,184,1905]" captionTargetId="figure-22@32.[163,1425,179,1917]" captionTargetPageId="32" captionText="FIGURE 19. Heriades leavitti. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="33" pageNumber="234">Fig. 19F</figureCitation>
).
<emphasis id="62E58D11FF85B072FBEDC62997237B34" box="[1063,1147,330,356]" italics="true" pageId="33" pageNumber="234">Female</emphasis>
: apical margin of clypeus with two tubercles sided by others smaller and irregular shaped (
<figureCitation id="C8AA4D86FF85B072FCBBC60D90807BD8" box="[881,984,366,392]" captionStart="FIGURE 19" captionStartId="32.[151,250,1943,1967]" captionTargetBox="[163,1412,184,1905]" captionTargetId="figure-22@32.[163,1425,179,1917]" captionTargetPageId="32" captionText="FIGURE 19. Heriades leavitti. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="33" pageNumber="234">Fig. 19C</figureCitation>
), similar to
<taxonomicName id="97912A80FF85B072FBAAC60C97B47BD8" baseAuthorityName="Cresson" baseAuthorityYear="1872" box="[1120,1260,366,392]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="variolosa">
<emphasis id="62E58D11FF85B072FBAAC60C97B47BD8" box="[1120,1260,366,392]" italics="true" pageId="33" pageNumber="234">H. variolosa</emphasis>
</taxonomicName>
and
<taxonomicName id="97912A80FF85B072FAEEC60C96C57BD8" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1316,1437,367,392]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="cristina" status="sp. nov.">
<emphasis id="62E58D11FF85B072FAEEC60C96C57BD8" box="[1316,1437,367,392]" italics="true" pageId="33" pageNumber="234">H. cristina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFF85B072FF5DC6F093A87BFD" box="[151,240,403,429]" pageId="33" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
, but different from first by having usually darker wings and by second for apical margin of clypeus straight, without emargination but with depression between tubercles and T3 with smaller punctation and dull integument.
</paragraph>
<paragraph id="502E5103FF85B072FF0DC6B990AB7998" blockId="33.[151,1437,150,1257]" pageId="33" pageNumber="234">
<emphasis id="62E58D11FF85B072FF0DC6B991617BA4" bold="true" box="[199,569,474,500]" pageId="33" pageNumber="234">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFF85B072FD8BC6B992847848" collectingDate="1989-05-20" collectionCode="KUNHM-ENT" collectorName="Yanega" country="Mexico" elevation="601" latitude="22.5775" location="San Luis Potosi" longLatPrecision="5" longitude="-99.40139" pageId="33" pageNumber="234" specimenCount="1" specimenCount-female="1" stateProvince="San Luis Potosi">
<emphasis id="62E58D11FF85B072FD8BC6B990D37BA4" bold="true" box="[577,907,474,500]" pageId="33" pageNumber="234">
<collectingCountry id="28861193FF85B072FD8BC6B991E47BA4" box="[577,700,474,500]" name="Mexico" pageId="33" pageNumber="234">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FF85B072FD03C6B990DE7BA4" box="[713,902,474,500]" country="Mexico" name="San Luis Potosi" pageId="33" pageNumber="234">San Luis Potosi</collectingRegion>
,
</emphasis>
<quantity id="9769FCE6FF85B072FC5EC6B990867BA5" box="[916,990,474,501]" metricMagnitude="2" metricUnit="m" metricValue="6.01" pageId="33" pageNumber="234" unit="m" value="601.0">
<elevation id="DBBCB630FF85B072FC5EC6B990867BA5" box="[916,990,474,501]" metricMagnitude="2" metricUnit="m" metricValue="6.01" pageId="33" pageNumber="234" unit="m" value="601.0">601 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FC21C6B9971E7BA5" box="[1003,1094,474,501]" degrees="22.5775" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="5" value="22.5775">22.5775</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FB84C6B997B67BA4" box="[1102,1262,474,500]" degrees="99.40138889" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-99.40139">-99.40138889</geoCoordinate>
,
<date id="242F77C3FF85B072FB30C6B9962D7BA4" box="[1274,1397,474,500]" pageId="33" pageNumber="234" value="1989-05-20">
<collectingDate id="346B8E2BFF85B072FB30C6B9962D7BA4" box="[1274,1397,474,500]" pageId="33" pageNumber="234" value="1989-05-20">20/V/1989</collectingDate>
</date>
, D.
<collectorName id="FD6434D5FF85B072FF5DC69C93B27849" box="[151,234,511,537]" pageId="33" pageNumber="234">Yanega</collectorName>
(
<collectionCode id="3680C9C6FF85B072FF30C69D92FD7848" box="[250,421,510,536]" pageId="33" pageNumber="234">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFF85B072FE65C69D92887848" box="[431,464,510,536]" count="1" pageId="33" pageNumber="234" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FE21C69D939E786C" collectionCode="IEXA-INECOL" collectorName="G. Quintos" country="Mexico" elevation="1200" latitude="19.4451" location="Coatepec" longLatPrecision="5" longitude="-96.96448" pageId="33" pageNumber="234" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Veracruz de Ignacio de la Llave">
<emphasis id="62E58D11FF85B072FE21C69D91077848" bold="true" box="[491,607,510,536]" pageId="33" pageNumber="234">
<collectingRegion id="92559FE1FF85B072FE21C69D91027848" box="[491,602,510,536]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="33" pageNumber="234">Veracruz</collectingRegion>
,
</emphasis>
<location id="554E07D8FF85B072FDA3C69D918C7848" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FDA3C69D918C7848" box="[617,724,510,536]" country="Mexico" latitude="19.4451" longLatPrecision="5" longitude="-96.96448" name="Coatepec" pageId="33" pageNumber="234" stateProvince="Veracruz de Ignacio de la Llave">Coatepec</location>
,
<quantity id="9769FCE6FF85B072FD2BC69D90627849" box="[737,826,510,537]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="234" unit="m" value="1200.0">
<elevation id="DBBCB630FF85B072FD2BC69D90627849" box="[737,826,510,537]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="234" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
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<geoCoordinate id="35A537C4FF85B072FC66C69D977A7848" box="[940,1058,510,536]" degrees="96.96448" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-96.96448">-96.96448</geoCoordinate>
,
<collectorName id="FD6434D5FF85B072FBE5C69D97F47848" box="[1071,1196,510,536]" pageId="33" pageNumber="234">G. Quintos</collectorName>
(
<collectionCode id="3680C9C6FF85B072FB76C69D96287848" box="[1212,1392,510,536]" pageId="33" pageNumber="234">IEXA-INECOL</collectionCode>
<specimenCount id="46979A8AFF85B072FAB3C69D96C47848" box="[1401,1436,510,536]" count="1" pageId="33" pageNumber="234" type="female">1♀</specimenCount>
<specimenCount id="46979A8AFF85B072FF5DC54193E5786C" box="[151,189,546,572]" count="2" pageId="33" pageNumber="234" type="male">2♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FF19C5419798786C" box="[211,1216,546,573]" collectingDate="2021-11-25" collectionCode="IEXA-INECOL" collectorName="G. Quintos" country="Mexico" elevation="1110" latitude="19.51586" location="Xalapa" longLatPrecision="1" longitude="-96.87426" pageId="33" pageNumber="234" specimenCount="2" specimenCount-male="2" stateProvince="Veracruz de Ignacio de la Llave">
<location id="554E07D8FF85B072FF19C541927D786C" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FF19C541927D786C" box="[211,293,546,572]" country="Mexico" latitude="19.51586" longLatPrecision="1" longitude="-96.87426" name="Xalapa" pageId="33" pageNumber="234" stateProvince="Veracruz de Ignacio de la Llave">Xalapa</location>
,
<quantity id="9769FCE6FF85B072FEF9C54192D2786D" box="[307,394,546,573]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="33" pageNumber="234" unit="m" value="1110.0">
<elevation id="DBBCB630FF85B072FEF9C54192D2786D" box="[307,394,546,573]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="33" pageNumber="234" unit="m" value="1110.0">1110 m</elevation>
</quantity>
.
<geoCoordinate id="35A537C4FF85B072FE5DC5419158786D" box="[407,512,546,573]" degrees="19.51586" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="19.51586">19.51586</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FDC3C5419126786C" box="[521,638,546,572]" degrees="96.87426" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-96.87426">-96.87426</geoCoordinate>
, 01,
<date id="242F77C3FF85B072FD7DC5419063786D" box="[695,827,546,573]" pageId="33" pageNumber="234" value="2021-11-25">
<collectingDate id="346B8E2BFF85B072FD7DC5419063786D" box="[695,827,546,573]" pageId="33" pageNumber="234" value="2021-11-25">25/XI/2021</collectingDate>
</date>
,
<collectorName id="FD6434D5FF85B072FC82C541909D786C" box="[840,965,546,572]" pageId="33" pageNumber="234">G. Quintos</collectorName>
(
<collectionCode id="3680C9C6FF85B072FC1FC54197D1786C" box="[981,1161,546,572]" pageId="33" pageNumber="234">IEXA-INECOL</collectionCode>
<specimenCount id="46979A8AFF85B072FB58C54197EF786C" box="[1170,1207,546,572]" count="2" pageId="33" pageNumber="234" type="male">2♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF85B072FB04C541906D7830" collectingDate="2021-09-10" collectionCode="IEXA-INECOL" collectorName="Quintos" country="Mexico" elevation="1150" latitude="19.51213" longLatPrecision="1" longitude="-96.94586" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="9769FCE6FF85B072FB04C541967E786D" box="[1230,1318,546,573]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="33" pageNumber="234" unit="m" value="1150.0">
<elevation id="DBBCB630FF85B072FB04C541967E786D" box="[1230,1318,546,573]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="33" pageNumber="234" unit="m" value="1150.0">1150 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FAF9C54196C4786D" box="[1331,1436,546,573]" degrees="19.51213" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="19.51213">19.51213</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FF5DC52592557831" box="[151,269,582,609]" degrees="96.94586" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-96.94586">-96.94586</geoCoordinate>
,
<date id="242F77C3FF85B072FED7C52592F97830" box="[285,417,582,608]" pageId="33" pageNumber="234" value="2021-09-10">
<collectingDate id="346B8E2BFF85B072FED7C52592F97830" box="[285,417,582,608]" pageId="33" pageNumber="234" value="2021-09-10">10/IX/2021</collectingDate>
</date>
, G.
<collectorName id="FD6434D5FF85B072FE10C525916B7830" box="[474,563,582,608]" pageId="33" pageNumber="234">Quintos</collectorName>
(
<collectionCode id="3680C9C6FF85B072FD8DC52591A37830" box="[583,763,582,608]" pageId="33" pageNumber="234">IEXA-INECOL</collectionCode>
<specimenCount id="46979A8AFF85B072FCCDC52590747830" box="[775,812,582,608]" count="1" pageId="33" pageNumber="234" type="male">1♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF85B072FC82C525926A78D4" collectingDate="2021-09-10" collectionCode="IEXA-INECOL" country="Mexico" elevation="1200" latitude="19.51337" longLatPrecision="1" longitude="-96.91743" pageId="33" pageNumber="234" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="9769FCE6FF85B072FC82C52590FC7831" box="[840,932,582,609]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="234" unit="m" value="1200.0">
<elevation id="DBBCB630FF85B072FC82C52590FC7831" box="[840,932,582,609]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="234" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FC7CC52597477831" box="[950,1055,582,609]" degrees="19.51337" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="19.51337">19.51337</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FBE6C52597F97830" box="[1068,1185,582,608]" degrees="96.91743" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-96.91743">-96.91743</geoCoordinate>
,
<date id="242F77C3FF85B072FB78C525966E7830" box="[1202,1334,582,608]" pageId="33" pageNumber="234" value="2021-09-10">
<collectingDate id="346B8E2BFF85B072FB78C525966E7830" box="[1202,1334,582,608]" pageId="33" pageNumber="234" value="2021-09-10">10/IX/2021</collectingDate>
</date>
, (
<collectionCode id="3680C9C6FF85B072FA84C52593A578D4" pageId="33" pageNumber="234">IEXA-INECOL</collectionCode>
<specimenCount id="46979A8AFF85B072FECEC509927D78D4" box="[260,293,618,644]" count="1" pageId="33" pageNumber="234" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FEF4C50A939E78F8" collectingDate="1996-03-26" collectionCode="CER-UADY" collectorName="F. Leon &amp; R. Rodriguez" country="Mexico" elevation="3" latitude="20.859304" location="Celestun" longLatPrecision="1" longitude="-90.3972" pageId="33" pageNumber="234" specimenCount="2" specimenCount-male="2" stateProvince="Yucatan">
<emphasis id="62E58D11FF85B072FEF4C50A92FF78D4" bold="true" box="[318,423,617,644]" pageId="33" pageNumber="234">
<collectingRegion id="92559FE1FF85B072FEF4C50A92FA78D4" box="[318,418,617,644]" country="Mexico" name="Yucatan" pageId="33" pageNumber="234">Yucatán</collectingRegion>
,
</emphasis>
<location id="554E07D8FF85B072FE65C509914C78D4" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FE65C509914C78D4" box="[431,532,618,644]" country="Mexico" latitude="20.859304" longLatPrecision="1" longitude="-90.3972" name="Celestun" pageId="33" pageNumber="234" stateProvince="Yucatan">Celestún</location>
,
<quantity id="9769FCE6FF85B072FDD5C509911478D5" box="[543,588,618,645]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="234" unit="m" value="3.0">
<elevation id="DBBCB630FF85B072FDD5C509911478D5" box="[543,588,618,645]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="234" unit="m" value="3.0">3 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FD9DC509919678D5" box="[599,718,618,645]" degrees="20.859304" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="20.859304">20.859304</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FD1DC509900378D5" box="[727,859,618,645]" degrees="90.397205" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-90.3972">-90.397205</geoCoordinate>
,
<date id="242F77C3FF85B072FCA2C50990B278D4" box="[872,1002,618,644]" pageId="33" pageNumber="234" value="1996-03-26">
<collectingDate id="346B8E2BFF85B072FCA2C50990B278D4" box="[872,1002,618,644]" pageId="33" pageNumber="234" value="1996-03-26">26/03/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FF85B072FC33C508970B78D4" box="[1017,1107,618,645]" pageId="33" pageNumber="234">F. León</collectorName>
,
<collectorName id="FD6434D5FF85B072FBABC50897A278D4" box="[1121,1274,618,645]" pageId="33" pageNumber="234">R. Rodriguez</collectorName>
(
<collectionCode id="3680C9C6FF85B072FAC1C50996C578D4" box="[1291,1437,618,644]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FF5DC5ED93E578F8" box="[151,189,654,680]" count="2" pageId="33" pageNumber="234" type="male">2♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFF85B072FF1FC5EC979578F8" box="[213,1229,654,681]" collectingDate="2006-07-16" collectionCode="CER-UADY" country="Mexico" elevation="6" latitude="21.037079" location="Reserva Estatal El Palmar" longLatPrecision="1" longitude="-90.18214" pageId="33" pageNumber="234" specimenCount="3" specimenCount-male="3" stateProvince="Yucatan">
<location id="554E07D8FF85B072FF1FC5EC915C78F8" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FF1FC5EC915C78F8" box="[213,516,654,681]" country="Mexico" latitude="21.037079" longLatPrecision="1" longitude="-90.18214" name="Reserva Estatal El Palmar" pageId="33" pageNumber="234" stateProvince="Yucatan">Reserva Estatal El Palmar</location>
,
<quantity id="9769FCE6FF85B072FDD9C5ED911B78F9" box="[531,579,654,681]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="234" unit="m" value="6.0">
<elevation id="DBBCB630FF85B072FDD9C5ED911B78F9" box="[531,579,654,681]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="234" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FD98C5ED919178F8" box="[594,713,654,680]" degrees="21.037079" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="21.037079">21.037079</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FD19C5ED900F78F8" box="[723,855,654,680]" degrees="90.182137" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-90.18214">-90.182137</geoCoordinate>
,
<date id="242F77C3FF85B072FCAFC5ED90A878F8" box="[869,1008,654,680]" pageId="33" pageNumber="234" value="2006-07-16">
<collectingDate id="346B8E2BFF85B072FCAFC5ED90A878F8" box="[869,1008,654,680]" pageId="33" pageNumber="234" value="2006-07-16">16/VII/2006</collectingDate>
</date>
(
<collectionCode id="3680C9C6FF85B072FBCBC5ED97CB78F8" box="[1025,1171,654,680]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FB57C5ED979B78F8" box="[1181,1219,654,680]" count="3" pageId="33" pageNumber="234" type="male">3♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FB11C5ED9023789C" collectingDate="1996-03-14" collectionCode="CER-UADY" collectorName="R. Rodriguez" country="Mexico" elevation="30" latitude="20.6412" location="Cuncunul" longLatPrecision="5" longitude="-88.296295" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Yucatan">
<location id="554E07D8FF85B072FB11C5ED961378F8" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FB11C5ED961378F8" box="[1243,1355,654,680]" country="Mexico" latitude="20.6412" longLatPrecision="5" longitude="-88.296295" name="Cuncunul" pageId="33" pageNumber="234" stateProvince="Yucatan">Cuncunul</location>
,
<quantity id="9769FCE6FF85B072FA93C5ED96CF78F9" box="[1369,1431,654,681]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="234" unit="m" value="30.0">
<elevation id="DBBCB630FF85B072FA93C5ED96CF78F9" box="[1369,1431,654,681]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="234" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FF5DC5D193AA789C" box="[151,242,690,716]" degrees="20.6412" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="5" value="20.6412">20.6412</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FF32C5D19224789C" box="[248,380,690,716]" degrees="88.296294" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-88.296295">-88.296294</geoCoordinate>
,
<date id="242F77C3FF85B072FE4CC5D19150789C" box="[390,520,690,716]" pageId="33" pageNumber="234" value="1996-03-14">
<collectingDate id="346B8E2BFF85B072FE4CC5D19150789C" box="[390,520,690,716]" pageId="33" pageNumber="234" value="1996-03-14">14/III/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FF85B072FDD8C5D091FF789C" box="[530,679,690,717]" pageId="33" pageNumber="234">R. Rodriguez</collectorName>
(
<collectionCode id="3680C9C6FF85B072FD7EC5D1901E789C" box="[692,838,690,716]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FC86C5D19029789C" box="[844,881,690,716]" count="1" pageId="33" pageNumber="234" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FC4CC5D0915D78A0" collectingDate="1996-05-13" collectionCode="CER-UADY" collectorName="H. Contreras" country="Mexico" elevation="29" latitude="20.388369" location="Dzan" longLatPrecision="1" longitude="-89.46909" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Yucatan">
<location id="554E07D8FF85B072FC4CC5D0909C789D" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FC4CC5D0909C789D" box="[902,964,691,717]" country="Mexico" latitude="20.388369" longLatPrecision="1" longitude="-89.46909" name="Dzan" pageId="33" pageNumber="234" stateProvince="Yucatan">Dzan</location>
,
<quantity id="9769FCE6FF85B072FC04C5D19750789D" box="[974,1032,690,717]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="33" pageNumber="234" unit="m" value="29.0">
<elevation id="DBBCB630FF85B072FC04C5D19750789D" box="[974,1032,690,717]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="33" pageNumber="234" unit="m" value="29.0">29 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FBD9C5D197D2789C" box="[1043,1162,690,716]" degrees="20.388369" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="20.388369">20.388369</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FB5AC5D1964B789C" box="[1168,1299,690,716]" degrees="89.469093" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-89.46909">-89.469093</geoCoordinate>
,
<date id="242F77C3FF85B072FAD7C5D196C0789C" box="[1309,1432,690,716]" pageId="33" pageNumber="234" value="1996-05-13">
<collectingDate id="346B8E2BFF85B072FAD7C5D196C0789C" box="[1309,1432,690,716]" pageId="33" pageNumber="234" value="1996-05-13">13/V/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FF85B072FF5DC5B4927278A0" box="[151,298,726,753]" pageId="33" pageNumber="234">H. Contreras</collectorName>
(
<collectionCode id="3680C9C6FF85B072FEF1C5B5929578A0" box="[315,461,726,752]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FE1CC5B592A378A0" box="[470,507,726,752]" count="1" pageId="33" pageNumber="234" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FDDEC5B591257944" collectingDate="2006-05-10" collectionCode="CER-UADY" collectorName="V. Gaumeri" country="Mexico" elevation="6" latitude="21.502728" location="Reserva Estatal Dzilam" longLatPrecision="1" longitude="-88.49085" municipality="Dzliam de Bravo" pageId="33" pageNumber="234" specimenCount="9" specimenCount-male="9" stateProvince="Yucatan">
<collectingMunicipality id="B04ACB79FF85B072FDDEC5B5918478A1" box="[532,732,726,753]" pageId="33" pageNumber="234">Dzliam de Bravo</collectingMunicipality>
,
<location id="554E07D8FF85B072FD21C5B490A278A0" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FD21C5B490A278A0" box="[747,1018,726,753]" country="Mexico" latitude="21.502728" longLatPrecision="1" longitude="-88.49085" municipality="Dzliam de Bravo" name="Reserva Estatal Dzilam" pageId="33" pageNumber="234" stateProvince="Yucatan">Reserva Estatal Dzilam</location>
,
<quantity id="9769FCE6FF85B072FBC3C5B5976178A1" box="[1033,1081,726,753]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="234" unit="m" value="6.0">
<elevation id="DBBCB630FF85B072FBC3C5B5976178A1" box="[1033,1081,726,753]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="234" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FB82C5B5979578A1" box="[1096,1229,726,753]" degrees="21.5027272" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="21.502728">21.5027272</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FB1DC5B5960278A1" box="[1239,1370,726,753]" degrees="88.490851" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-88.49085">-88.490851</geoCoordinate>
, 05/I,
<date id="242F77C3FF85B072FF5DC59992497944" box="[151,273,762,788]" pageId="33" pageNumber="234" value="2006-05-10">
<collectingDate id="346B8E2BFF85B072FF5DC59992497944" box="[151,273,762,788]" pageId="33" pageNumber="234" value="2006-05-10">10/V/2006</collectingDate>
</date>
,
<collectorName id="FD6434D5FF85B072FED5C59892FA7944" box="[287,418,762,789]" pageId="33" pageNumber="234">V. Gaumeri</collectorName>
(
<collectionCode id="3680C9C6FF85B072FE79C599911D7944" box="[435,581,762,788]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FD84C599912B7944" box="[590,627,762,788]" count="9" pageId="33" pageNumber="234" type="male">9♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FD40C59992BB7968" collectingDate="1995-05-24" collectingDateMax="1996-01-11" collectingDateMin="1995-05-24" collectionCode="CER-UADY" collectorName="R. Tut &amp; L. Novelo" country="Mexico" elevation="6" latitude="21.515833" location="El Cuyo" longLatPrecision="1" longitude="-87.67833" pageId="33" pageNumber="234" specimenCount="2" specimenCount-male="2" stateProvince="Yucatan">
<location id="554E07D8FF85B072FD40C59991B47944" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FD40C59991B47944" box="[650,748,762,788]" country="Mexico" latitude="21.515833" longLatPrecision="1" longitude="-87.67833" name="El Cuyo" pageId="33" pageNumber="234" stateProvince="Yucatan">El Cuyo</location>
,
<quantity id="9769FCE6FF85B072FD30C59990717945" box="[762,809,762,789]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="234" unit="m" value="6.0">
<elevation id="DBBCB630FF85B072FD30C59990717945" box="[762,809,762,789]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="234" unit="m" value="6.0">6 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FCFDC59990F67945" box="[823,942,762,789]" degrees="21.515833" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="21.515833">21.515833</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FC72C59997637944" box="[952,1083,762,788]" degrees="87.678333" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-87.67833">-87.678333</geoCoordinate>
,
<date id="242F77C3FF85B072FB82C599979A7945" box="[1096,1218,762,789]" pageId="33" pageNumber="234" value="1995-05-24">
<collectingDate id="346B8E2BFF85B072FB82C599979A7945" box="[1096,1218,762,789]" pageId="33" pageNumber="234" value="1995-05-24">24/V/1995</collectingDate>
</date>
,
<collectorName id="FD6434D5FF85B072FB1BC59896447945" box="[1233,1308,763,789]" pageId="33" pageNumber="234">R. Tut</collectorName>
,
<date id="242F77C3FF85B072FAE3C59996C07944" box="[1321,1432,762,788]" pageId="33" pageNumber="234" value="1996-01-11">
<collectingDate id="346B8E2BFF85B072FAE3C59996C07944" box="[1321,1432,762,788]" pageId="33" pageNumber="234" value="1996-01-11">11/I/1996</collectingDate>
</date>
,
<collectorName id="FD6434D5FF85B072FF5DC47C92527968" box="[151,266,798,825]" pageId="33" pageNumber="234">L. Novelo</collectorName>
(
<collectionCode id="3680C9C6FF85B072FED0C47D92F47968" box="[282,428,798,824]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FE7EC47D92827968" box="[436,474,798,824]" count="2" pageId="33" pageNumber="234" type="male">2♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FE3AC47D92B5790C" collectingDate="2006-07-24" collectionCode="CER-UADY" country="Mexico" elevation="11" latitude="20.653353" location="Reserva Estatal Lagunas de Yalahau" longLatPrecision="1" longitude="-89.21638" pageId="33" pageNumber="234" specimenCount="2" specimenCount-male="2" stateProvince="Yucatan">
[Homún],
<location id="554E07D8FF85B072FDACC47C975B7968" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FDACC47C975B7968" box="[614,1027,798,825]" country="Mexico" latitude="20.653353" longLatPrecision="1" longitude="-89.21638" name="Reserva Estatal Lagunas de Yalahau" pageId="33" pageNumber="234" stateProvince="Yucatan">Reserva Estatal Lagunas de Yalahau</location>
,
<quantity id="9769FCE6FF85B072FBDAC47D97147969" box="[1040,1100,798,825]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="33" pageNumber="234" unit="m" value="11.0">
<elevation id="DBBCB630FF85B072FBDAC47D97147969" box="[1040,1100,798,825]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="33" pageNumber="234" unit="m" value="11.0">11 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FB93C47D97887969" box="[1113,1232,798,825]" degrees="20.653353" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="20.653353">20.653353</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FB12C47D96047968" box="[1240,1372,798,824]" degrees="89.216377" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-89.21638">-89.216377</geoCoordinate>
, 14/I,
<date id="242F77C3FF85B072FF5DC421924F790C" box="[151,279,834,860]" pageId="33" pageNumber="234" value="2006-07-24">
<collectingDate id="346B8E2BFF85B072FF5DC421924F790C" box="[151,279,834,860]" pageId="33" pageNumber="234" value="2006-07-24">24/07/2006</collectingDate>
</date>
(
<collectionCode id="3680C9C6FF85B072FEEFC42192EF790C" box="[293,439,834,860]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FE74C42192BC790C" box="[446,484,834,860]" count="2" pageId="33" pageNumber="234" type="male">2♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FE33C42096CF790C" box="[505,1431,834,861]" collectingDate="1995-06-15" collectionCode="CER-UADY" collectorName="S. Norena" country="Mexico" elevation="13" latitude="21.197361" location="La Sierra" longLatPrecision="1" longitude="-87.71918" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Yucatan">
<location id="554E07D8FF85B072FE33C420913B790C" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FE33C420913B790C" box="[505,611,834,861]" country="Mexico" latitude="21.197361" longLatPrecision="1" longitude="-87.71918" name="La Sierra" pageId="33" pageNumber="234" stateProvince="Yucatan">La Sierra</location>
,
<quantity id="9769FCE6FF85B072FDA4C42191F1790D" box="[622,681,834,861]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="234" unit="m" value="13.0">
<elevation id="DBBCB630FF85B072FDA4C42191F1790D" box="[622,681,834,861]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="234" unit="m" value="13.0">13 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FD7FC4219074790C" box="[693,812,834,860]" degrees="21.197361" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="21.197361">21.197361</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FCFEC42190EF790C" box="[820,951,834,860]" degrees="87.719176" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-87.71918">-87.719176</geoCoordinate>
,
<date id="242F77C3FF85B072FC08C421971E790D" box="[962,1094,834,861]" pageId="33" pageNumber="234" value="1995-06-15">
<collectingDate id="346B8E2BFF85B072FC08C421971E790D" box="[962,1094,834,861]" pageId="33" pageNumber="234" value="1995-06-15">15/VI/1995</collectingDate>
</date>
,
<collectorName id="FD6434D5FF85B072FB9BC4219799790D" box="[1105,1217,834,861]" pageId="33" pageNumber="234">S. Noreña</collectorName>
(
<collectionCode id="3680C9C6FF85B072FB1AC421963A790C" box="[1232,1378,834,860]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FAA3C42196D5790C" box="[1385,1421,834,860]" count="1" pageId="33" pageNumber="234" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FF5DC40597A379D0" box="[151,1275,870,897]" collectingDate="2006-11-07" collectionCode="CER-UADY" country="Mexico" elevation="11" latitude="21.094807" location="Reserva Estatal Dzibilchaltun" longLatPrecision="1" longitude="-89.597755" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Yucatan">
[Mérida],
<location id="554E07D8FF85B072FECDC404910B79D0" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FECDC404910B79D0" box="[263,595,870,897]" country="Mexico" latitude="21.094807" longLatPrecision="1" longitude="-89.597755" name="Reserva Estatal Dzibilchaltun" pageId="33" pageNumber="234" stateProvince="Yucatan">Reserva Estatal Dzibilchaltún</location>
,
<quantity id="9769FCE6FF85B072FD97C40591CD79D1" box="[605,661,870,897]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="33" pageNumber="234" unit="m" value="11.0">
<elevation id="DBBCB630FF85B072FD97C40591CD79D1" box="[605,661,870,897]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="33" pageNumber="234" unit="m" value="11.0">11 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FD55C405904E79D0" box="[671,790,870,896]" degrees="21.094807" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="21.094807">21.094807</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FCD1C40590C779D1" box="[795,927,870,897]" degrees="89.597755" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-89.597755">-89.597755</geoCoordinate>
,
<date id="242F77C3FF85B072FC62C405977179D0" box="[936,1065,870,896]" pageId="33" pageNumber="234" value="2006-11-07">
<collectingDate id="346B8E2BFF85B072FC62C405977179D0" box="[936,1065,870,896]" pageId="33" pageNumber="234" value="2006-11-07">07/XI/2006</collectingDate>
</date>
(
<collectionCode id="3680C9C6FF85B072FBFCC405979079D0" box="[1078,1224,870,896]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FB07C40597A979D0" box="[1229,1265,870,896]" count="1" pageId="33" pageNumber="234" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFF85B072FACFC40590CB79F4" collectingDate="1995-05-26" collectionCode="CER-UADY" collectorName="Moo" country="Mexico" elevation="3" latitude="21.596481" location="Rio Lagartos" longLatPrecision="1" longitude="-88.1579" pageId="33" pageNumber="234" specimenCount="1" specimenCount-male="1" stateProvince="Yucatan">
<location id="554E07D8FF85B072FACFC40596C079D1" LSID="urn:lsid:plazi:treatment:D838E015FFBBB072FF5DC1CA96137EB8:554E07D8FF85B072FACFC40596C079D1" box="[1285,1432,870,897]" country="Mexico" latitude="21.596481" longLatPrecision="1" longitude="-88.1579" name="Rio Lagartos" pageId="33" pageNumber="234" stateProvince="Yucatan">Río Lagartos</location>
,
<quantity id="9769FCE6FF85B072FF5DC4E9939D79F5" box="[151,197,906,933]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="234" unit="m" value="3.0">
<elevation id="DBBCB630FF85B072FF5DC4E9939D79F5" box="[151,197,906,933]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="234" unit="m" value="3.0">3 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FF85B072FF1BC4E9921079F5" box="[209,328,906,933]" degrees="21.596482" direction="north" orientation="latitude" pageId="33" pageNumber="234" precision="1" value="21.596481">21.596482</geoCoordinate>
<geoCoordinate id="35A537C4FF85B072FE9AC4E9928B79F5" box="[336,467,906,933]" degrees="88.157898" direction="west" orientation="longitude" pageId="33" pageNumber="234" precision="1" value="-88.1579">-88.157898</geoCoordinate>
,
<date id="242F77C3FF85B072FE15C4E9910179F5" box="[479,601,906,933]" pageId="33" pageNumber="234" value="1995-05-26">
<collectingDate id="346B8E2BFF85B072FE15C4E9910179F5" box="[479,601,906,933]" pageId="33" pageNumber="234" value="1995-05-26">26/V/1995</collectingDate>
</date>
, H.
<collectorName id="FD6434D5FF85B072FD42C4E891E579F5" box="[648,701,907,933]" pageId="33" pageNumber="234">Moo</collectorName>
(
<collectionCode id="3680C9C6FF85B072FD06C4E9900679F4" box="[716,862,906,932]" pageId="33" pageNumber="234">CER-UADY</collectionCode>
<specimenCount id="46979A8AFF85B072FCAFC4E990D279F4" box="[869,906,906,932]" count="1" pageId="33" pageNumber="234" type="male">1♂</specimenCount>
)
</materialsCitation>
,
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<date id="242F77C3FF85B072FC6AC4E9977379F4" box="[928,1067,906,932]" pageId="33" pageNumber="234" value="1996-07-27">
<collectingDate id="346B8E2BFF85B072FC6AC4E9977379F4" box="[928,1067,906,932]" pageId="33" pageNumber="234" value="1996-07-27">27/VII/1996</collectingDate>
</date>
(
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)
</materialsCitation>
.
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, 8
<collectorName id="FD6434D5FF85B072FAA6C4E8938C7998" pageId="33" pageNumber="234">K N Ticul</collectorName>
,
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,
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</date>
,
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(
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<specimenCount id="46979A8AFF85B072FC0AC4CD90BC7998" box="[960,996,942,968]" count="1" pageId="33" pageNumber="234" type="male">1♂</specimenCount>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF85B072FF0DC4B197717E2D" pageId="33" pageNumber="234" type="distribution">
<paragraph id="502E5103FF85B072FF0DC4B197717E2D" blockId="33.[151,1437,150,1257]" pageId="33" pageNumber="234">
<emphasis id="62E58D11FF85B072FF0DC4B1923979BC" bold="true" box="[199,353,978,1004]" pageId="33" pageNumber="234">Distribution.</emphasis>
This species is mainly present in the
<collectingCountry id="28861193FF85B072FCE9C4B190E579BC" box="[803,957,978,1004]" name="United States of America" pageId="33" pageNumber="234">United States</collectingCountry>
and
<collectingCountry id="28861193FF85B072FC31C4B1970B79BC" box="[1019,1107,978,1004]" name="Canada" pageId="33" pageNumber="234">Canada</collectingCountry>
; however, we found records that extend the distribution to
<collectingCountry id="28861193FF85B072FE30C495910A7E40" box="[506,594,1014,1040]" name="Mexico" pageId="33" pageNumber="234">Mexico</collectingCountry>
, along the Gulf of
<collectingCountry id="28861193FF85B072FCFDC49590D57E40" box="[823,909,1014,1040]" name="Mexico" pageId="33" pageNumber="234">Mexico</collectingCountry>
slope and
<collectingRegion id="92559FE1FF85B072FBC5C49597317E40" box="[1039,1129,1014,1040]" country="Mexico" name="Yucatan" pageId="33" pageNumber="234">Yucatán</collectingRegion>
Peninsula. The records of Quintos-Andrade,
<emphasis id="62E58D11FF85B072FEADC37892C07E64" box="[359,408,1050,1076]" italics="true" pageId="33" pageNumber="234">et al</emphasis>
. (2023) of
<taxonomicName id="97912A80FF85B072FDDFC37891F97E64" baseAuthorityName="Cresson" baseAuthorityYear="1872" box="[533,673,1050,1076]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="variolosa">
<emphasis id="62E58D11FF85B072FDDFC37891F97E64" box="[533,673,1050,1076]" italics="true" pageId="33" pageNumber="234">H. variolosa</emphasis>
</taxonomicName>
from
<collectingCountry id="28861193FF85B072FD2EC37990647E64" box="[740,828,1050,1076]" name="Mexico" pageId="33" pageNumber="234">Mexico</collectingCountry>
,
<collectingRegion id="92559FE1FF85B072FC8DC37890F67E65" box="[839,942,1051,1077]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="33" pageNumber="234">Veracruz</collectingRegion>
, Coatepec and Xalapa belongs to
<emphasis id="62E58D11FF85B072FAECC37896C57E64" box="[1318,1437,1050,1076]" italics="true" pageId="33" pageNumber="234">
<taxonomicName id="97912A80FF85B072FAECC37896C07E64" authorityName="Crawford" authorityYear="1913" box="[1318,1432,1050,1076]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="leavitti">H. leavitti</taxonomicName>
.
</emphasis>
These areas correspond to the Sierra Madre Oriental, Veracruzan and
<collectingRegion id="92559FE1FF85B072FC65C35D97517E08" box="[943,1033,1086,1112]" country="Mexico" name="Yucatan" pageId="33" pageNumber="234">Yucatán</collectingRegion>
Peninsula biogeographic provinces (
<bibRefCitation id="34002CF2FF85B072FF55C30192DA7E2C" author="Morrone, J. J. &amp; Escalante, T. &amp; Rodriguez-Tapia, G. &amp; Carmona, A. &amp; Arana, M. &amp; Mercado-Gomez, J. D." box="[159,386,1122,1149]" pageId="33" pageNumber="234" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. &amp; Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="62E58D11FF85B072FEC3C30092637E2C" box="[265,315,1122,1148]" italics="true" pageId="33" pageNumber="234">et al</emphasis>
. 2022
</bibRefCitation>
) (
<figureCitation id="C8AA4D86FF85B072FE53C30192B47E2C" box="[409,492,1122,1148]" captionStart="FIGURE 18" captionStartId="31.[151,250,1098,1122]" captionTargetBox="[159,1428,187,1063]" captionTargetId="figure-18@31.[151,1436,179,1072]" captionTargetPageId="31" captionText="FIGURE 18. Known distribution records for Heriades leavitti." pageId="33" pageNumber="234">Fig. 18</figureCitation>
). The altitudinal range is from sea level to
<quantity id="9769FCE6FF85B072FC07C301977D7E2D" box="[973,1061,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="234" unit="m" value="1200.0">1200 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FF85B072FF0DC3E596137EB8" pageId="33" pageNumber="234" type="discussion">
<paragraph id="502E5103FF85B072FF0DC3E596137EB8" blockId="33.[151,1437,150,1257]" pageId="33" pageNumber="234">
<emphasis id="62E58D11FF85B072FF0DC3E592177EF0" bold="true" box="[199,335,1158,1184]" pageId="33" pageNumber="234">Comments.</emphasis>
This species is similar to
<taxonomicName id="97912A80FF85B072FDBFC3E490597EF0" baseAuthorityName="Cresson" baseAuthorityYear="1872" box="[629,769,1158,1184]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="variolosa">
<emphasis id="62E58D11FF85B072FDBFC3E490597EF0" box="[629,769,1158,1184]" italics="true" pageId="33" pageNumber="234">H. variolosa</emphasis>
</taxonomicName>
; females of
<taxonomicName id="97912A80FF85B072FC47C3E490A67EF0" authorityName="Crawford" authorityYear="1913" box="[909,1022,1158,1184]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="leavitti">
<emphasis id="62E58D11FF85B072FC47C3E490A67EF0" box="[909,1022,1158,1184]" italics="true" pageId="33" pageNumber="234">H. leavitti</emphasis>
</taxonomicName>
can be recognized by having greater DBP on scutum and dusky wings; and males can be easily recognized by having two tufts of simple, thin, short hairs on S6, not a few simple, scattered capitate hairs, and two rounded tufts of capitate hairs on S5, not one line.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,643 @@
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<heading id="0B66E66FFFBDB04AFF5DC2E991767FF4" bold="true" box="[151,558,1418,1445]" fontSize="11" level="1" pageId="25" pageNumber="226" reason="1">
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<emphasis id="62E58D11FFBDB04AFF5DC2E991767FF4" bold="true" box="[151,558,1418,1445]" pageId="25" pageNumber="226">
<emphasis id="62E58D11FFBDB04AFF5DC2E9923F7FF4" bold="true" box="[151,359,1418,1445]" italics="true" pageId="25" pageNumber="226">Heriades currani</emphasis>
Michener, 1943
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(
<figureCitation id="C8AA4D86FFBDB04AFF55C2CD92587F98" box="[159,256,1454,1480]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="25" pageNumber="226">Figs. 2E</figureCitation>
,
<figureCitation id="C8AA4D86FFBDB04AFEC6C2CD920B7F99" box="[268,339,1454,1481]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStartId-0="25.[151,250,1098,1122]" captionStartId-1="26.[151,250,1943,1967]" captionTargetBox-0="[160,1428,187,1064]" captionTargetBox-1="[161,1421,179,1898]" captionTargetId-0="figure-18@25.[151,1436,179,1072]" captionTargetId-1="figure-22@26.[161,1426,179,1917]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURE 14. Known distribution records for Heriades currani." captionText-1="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="25" pageNumber="226">1415</figureCitation>
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<paragraph id="502E5103FFBDB04AFF5DC29092897C37" blockId="25.[151,1436,1523,1669]" pageId="25" pageNumber="226">
<taxonomicName id="97912A80FFBDB04AFF5DC29091C27C5B" authority="Michener, 1943" authorityName="Michener" authorityYear="1943" box="[151,666,1523,1547]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="currani" subGenus="Neotrypetes">
<emphasis id="62E58D11FFBDB04AFF5DC29092BC7C5A" box="[151,484,1523,1547]" italics="true" pageId="25" pageNumber="226">Heriades (Neotrypetes) currani</emphasis>
Michener, 1943
</taxonomicName>
. A new bee of the genus
<taxonomicName id="97912A80FFBDB04AFC0EC29097797C5B" authorityName="Spinola" authorityYear="1808" box="[964,1057,1523,1547]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="62E58D11FFBDB04AFC0EC29097797C5B" box="[964,1057,1523,1547]" italics="true" pageId="25" pageNumber="226">Heriades</emphasis>
</taxonomicName>
from
<collectingCountry id="28861193FFBDB04AFBA6C29097987C5B" box="[1132,1216,1523,1547]" name="Panama" pageId="25" pageNumber="226">Panamá</collectingCountry>
.
<emphasis id="62E58D11FFBDB04AFB05C29092CC7C79" italics="true" pageId="25" pageNumber="226">
<taxonomicName id="97912A80FFBDB04AFB05C29096C47C5B" authority=", Cambridge" authorityName="Cambridge" box="[1231,1436,1523,1547]" class="Insecta" family="Psychidae" genus="Psyche" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="genus">Psyche, Cambridge</taxonomicName>
Entomological Club
</emphasis>
:109. Type locality:
<collectingCountry id="28861193FFBDB04AFE93C15392F47C18" box="[345,428,1584,1608]" name="Panama" pageId="25" pageNumber="226">Panamá</collectingCountry>
,
<collectingCountry id="28861193FFBDB04AFE7CC15391DE7C18" box="[438,646,1584,1608]" name="Panama" pageId="25" pageNumber="226">Republic of Panamá</collectingCountry>
,
<date id="242F77C3FFBDB04AFD5AC15390DF7C18" box="[656,903,1584,1608]" pageId="25" pageNumber="226" value="1915-02" valueMax="1915-03-31" valueMin="1915-02-00">February to March 1915</date>
(C. H. Curran)
<typeStatus id="8F2AEFA1FFBDB04AFF0DC12C927D7C37" box="[199,293,1615,1639]" pageId="25" pageNumber="226" type="holotype">Holotype</typeStatus>
female: AMNH.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBDB04AFF5DC10E904E7CD5" box="[151,790,1645,1669]" pageId="25" pageNumber="226" type="reference_group">
<paragraph id="502E5103FFBDB04AFF5DC10E904E7CD5" blockId="25.[151,1436,1523,1669]" box="[151,790,1645,1669]" pageId="25" pageNumber="226">
<taxonomicName id="97912A80FFBDB04AFF5DC10E91287CD5" ID-CoL="8SNLF" authority="Gonzalez &amp; Griswold, 2012" authorityName="Gonzalez &amp; Griswold" authorityYear="2011" box="[151,624,1645,1669]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="62E58D11FFBDB04AFF5DC10E92107CD5" box="[151,328,1645,1669]" italics="true" pageId="25" pageNumber="226">Heriades tayrona</emphasis>
<bibRefCitation id="34002CF2FFBDB04AFE85C10E91287CD5" author="Gonzalez, V. H. &amp; Griswold, T. &amp; Praz, C. J. &amp; Danforth, B. N." box="[335,624,1645,1669]" pageId="25" pageNumber="226" pagination="261 - 286" refId="ref43188" refString="Gonzalez, V. H., Griswold, T., Praz, C. J. &amp; Danforth, B. N. (2012) Phylogeny of the bee family Megachilidae (Hymenoptera: Apoidea) based on adult morphology. Systematic Entomology, 37 (2), 261 - 286. https: // doi. org / 10.1111 / j. 1365 - 3113.2012.00620. x" type="journal article" year="2012">Gonzalez &amp; Griswold, 2012</bibRefCitation>
</taxonomicName>
.
<emphasis id="62E58D11FFBDB04AFDB3C10E904E7CD5" bold="true" box="[633,790,1645,1669]" pageId="25" pageNumber="226">New synonym.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBDB048FF5DC1D09127789C" lastPageId="27" lastPageNumber="228" pageId="25" pageNumber="226" type="diagnosis">
<paragraph id="502E5103FFBDB04AFF5DC1D091EA7DBE" blockId="25.[151,1437,1715,2030]" pageId="25" pageNumber="226">
<emphasis id="62E58D11FFBDB04AFF5DC1D0924A7C9D" bold="true" box="[151,274,1715,1741]" pageId="25" pageNumber="226">Diagnosis.</emphasis>
Tergal fasciae with rows of thin, white hairs (
<figureCitation id="C8AA4D86FFBDB04AFCE6C1D090DE7C9E" box="[812,902,1715,1742]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="25" pageNumber="226">Fig. 3L</figureCitation>
).
<emphasis id="62E58D11FFBDB04AFC57C1D0908F7C9D" box="[925,983,1715,1741]" italics="true" pageId="25" pageNumber="226">Male</emphasis>
: clypeus with hairs scarce and short in longitudinal middle areas, and on hypostomal area (
<figureCitation id="C8AA4D86FFBDB04AFD11C1B4901B7CA2" box="[731,835,1751,1778]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="25" pageNumber="226">Fig. 15A</figureCitation>
); ocelli large (
<figureCitation id="C8AA4D86FFBDB04AFC29C1B497637CA2" box="[995,1083,1751,1778]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="25" pageNumber="226">Fig. 2C</figureCitation>
); scutum with large punctation; posterior margin of S5 with one line of capitate hairs (
<figureCitation id="C8AA4D86FFBDB04AFD3DC19890047D46" box="[759,860,1787,1814]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="25" pageNumber="226">Fig. 15E</figureCitation>
); S6 with one subapical, transverse line of capitate, erected, long hairs (
<figureCitation id="C8AA4D86FFBDB04AFEB8C07C928D7D6A" box="[370,469,1823,1850]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="25" pageNumber="226">Fig. 15F</figureCitation>
); posterior margin of S1 truncate (
<figureCitation id="C8AA4D86FFBDB04AFC98C07C90CC7D69" box="[850,916,1823,1849]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="25" pageNumber="226">Fig. 2</figureCitation>
Eb) and, in profile, a low ventral angle in more than one third of distance from apex to base (
<figureCitation id="C8AA4D86FFBDB04AFD6BC02091BD7D0E" box="[673,741,1859,1886]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="25" pageNumber="226">Fig. 2</figureCitation>
Ea).
<emphasis id="62E58D11FFBDB04AFCD6C02090287D0D" box="[796,880,1859,1885]" italics="true" pageId="25" pageNumber="226">Female</emphasis>
: outer surface of mandible with strong elevation at base of condylar ridge, slightly rounded in frontal view (
<figureCitation id="C8AA4D86FFBDB04AFCF8C00490CE7DD2" box="[818,918,1895,1922]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="25" pageNumber="226">Fig. 15C</figureCitation>
) and with a smaller one at base of outer ridge fading in the surface, both with a Y-shaped; clypeus apical margin with two well defined tubercles with variation in size, or not well defined (
<figureCitation id="C8AA4D86FFBDB04AFE7EC0CC91427D9A" box="[436,538,1967,1994]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="25" pageNumber="226">Fig. 15C</figureCitation>
); scutum with large punctures (0.3 times OD), in T3 small (0.4 times DOP as on scutum) and sparse (2.5 times DOP) (
<figureCitation id="C8AA4D86FFBDB04AFDF6C0B091FC7DBE" box="[572,676,2003,2030]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="25" pageNumber="226">Fig. 15D</figureCitation>
).
</paragraph>
<caption id="04EE018BFFBEB049FF5DC0F491B97D83" pageId="26" pageNumber="227" startId="26.[151,250,1943,1967]" targetBox="[161,1421,179,1898]" targetPageId="26" targetType="figure">
<paragraph id="502E5103FFBEB049FF5DC0F491B97D83" blockId="26.[151,1436,1943,2003]" pageId="26" pageNumber="227">
<emphasis id="62E58D11FFBEB049FF5DC0F492787DFF" bold="true" box="[151,288,1943,1967]" pageId="26" pageNumber="227">FIGURE 15.</emphasis>
<emphasis id="62E58D11FFBEB049FEEDC0F492857DFF" box="[295,477,1943,1967]" italics="true" pageId="26" pageNumber="227">
<taxonomicName id="97912A80FFBEB049FEEDC0F492817DFF" authorityName="Michener" authorityYear="1943" box="[295,473,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="currani">Heriades currani</taxonomicName>
.
</emphasis>
Male:
<emphasis id="62E58D11FFBEB049FDEFC0F491667DFF" bold="true" box="[549,574,1943,1967]" pageId="26" pageNumber="227">A.</emphasis>
frontal view
<emphasis id="62E58D11FFBEB049FD0DC0F491867DFF" bold="true" box="[711,734,1943,1967]" pageId="26" pageNumber="227">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FFBEB049FC75C0F4908F7DFF" bold="true" box="[959,983,1943,1967]" pageId="26" pageNumber="227">C.</emphasis>
frontal view
<emphasis id="62E58D11FFBEB049FBAAC0F497217DFF" bold="true" box="[1120,1145,1943,1967]" pageId="26" pageNumber="227">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FFBEB049FA89C0F496027DFF" bold="true" box="[1347,1370,1943,1967]" pageId="26" pageNumber="227">E.</emphasis>
S5.
<emphasis id="62E58D11FFBEB049FA43C0F496C47DFF" bold="true" box="[1417,1436,1943,1967]" pageId="26" pageNumber="227">F.</emphasis>
S6.
<emphasis id="62E58D11FFBEB049FF75C0D893817D83" bold="true" box="[191,217,1979,2003]" pageId="26" pageNumber="227">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FFBEB049FDF7C0D891007D83" bold="true" box="[573,600,1979,2003]" pageId="26" pageNumber="227">H.</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="502E5103FFBFB048FF0DC7F59127789C" blockId="27.[151,1437,150,1149]" pageId="27" pageNumber="228">
<emphasis id="62E58D11FFBFB048FF0DC7F591657AE0" bold="true" box="[199,573,150,176]" pageId="27" pageNumber="228">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFFBFB048FD82C7F597177A84" collectingDate="2011-12-17" collectionCode="BBSL" collectorName="V. H. Gonzalez" country="Colombia" elevation="99" latitude="11.264566" location="Bastidas" longLatPrecision="1" longitude="-74.1654" municipality="Santa Marta" pageId="27" pageNumber="228" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena">
<emphasis id="62E58D11FFBFB048FD82C7F590DE7AE0" bold="true" box="[584,902,150,176]" pageId="27" pageNumber="228">
<collectingCountry id="28861193FFBFB048FD82C7F591B37AE0" box="[584,747,150,176]" name="Colombia" pageId="27" pageNumber="228">COLOMBIA</collectingCountry>
:
<collectingRegion id="92559FE1FFBFB048FD30C7F590DA7AE0" box="[762,898,150,176]" country="Colombia" name="Magdalena" pageId="27" pageNumber="228">Magdalena</collectingRegion>
,
</emphasis>
<collectingMunicipality id="B04ACB79FFBFB048FC5BC7F597787AE1" box="[913,1056,150,177]" pageId="27" pageNumber="228">Santa Marta</collectingMunicipality>
, on road from
<location id="554E07D8FFBFB048FB1BC7F596687AE0" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FB1BC7F596687AE0" box="[1233,1328,150,176]" country="Colombia" latitude="11.264566" longLatPrecision="1" longitude="-74.1654" municipality="Santa Marta" name="Bastidas" pageId="27" pageNumber="228" stateProvince="Magdalena">Bastidas</location>
to
<taxonomicName id="97912A80FFBFB048FA91C7F593B77A84" authority="Concha" authorityName="Concha" class="Magnoliopsida" family="Asteraceae" genus="Bahia" kingdom="Plantae" order="Asterales" pageId="27" pageNumber="228" phylum="Tracheophyta" rank="genus">Bahia Concha</taxonomicName>
,
<quantity id="9769FCE6FFBFB048FF31C7D9926F7A85" box="[251,311,186,213]" metricMagnitude="1" metricUnit="m" metricValue="9.9" pageId="27" pageNumber="228" unit="m" value="99.0">
<elevation id="DBBCB630FFBFB048FF31C7D9926F7A85" box="[251,311,186,213]" metricMagnitude="1" metricUnit="m" metricValue="9.9" pageId="27" pageNumber="228" unit="m" value="99.0">99 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFBFB048FE89C7D9928C7A85" box="[323,468,186,213]" degrees="11" direction="north" minutes="15.874" orientation="latitude" pageId="27" pageNumber="228" precision="1" value="11.264566">11°15.874N</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FE16C7D9912B7A84" box="[476,627,186,212]" degrees="74" direction="west" minutes="09.924" orientation="longitude" pageId="27" pageNumber="228" precision="1" value="-74.1654">74°09.924W</geoCoordinate>
,
<date id="242F77C3FFBFB048FD4AC7D990537A84" box="[640,779,186,212]" pageId="27" pageNumber="228" value="2011-12-17">
<collectingDate id="346B8E2BFFBFB048FD4AC7D990537A84" box="[640,779,186,212]" pageId="27" pageNumber="228" value="2011-12-17">17/XII/2011</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBFB048FCDDC7D8909B7A84" box="[791,963,186,213]" pageId="27" pageNumber="228">V. H. Gonzalez</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FC18C7D997417A84" box="[978,1049,186,212]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="27" pageNumber="228" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBFB048FBEAC7D9971D7A84" box="[1056,1093,186,212]" count="1" pageId="27" pageNumber="228" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FB91C7D890DF7AA8" collectingDate="2001-05-04" collectingDateMax="2001-05-23" collectingDateMin="2001-05-04" collectionCode="USDA-ARS" collectorName="Henriquez" country="Colombia" elevation="10" latitude="11.333333" longLatPrecision="1295" longitude="-74.03333" pageId="27" pageNumber="228" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena">
PNN
<taxonomicName id="97912A80FFBFB048FB50C7D896337A84" authority="Neguanje" authorityName="Neguanje" box="[1178,1387,186,213]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="tayrona">Tayrona Neguanje</taxonomicName>
, [
<quantity id="9769FCE6FFBFB048FAB4C7D993F77AA9" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="228" unit="m" value="10.0">
<elevation id="DBBCB630FFBFB048FAB4C7D993F77AA9" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="228" unit="m" value="10.0">10 m</elevation>
</quantity>
]
<geoCoordinate id="35A537C4FFBFB048FF77C7BD92447AA8" box="[189,284,222,248]" degrees="11" direction="north" minutes="20" orientation="latitude" pageId="27" pageNumber="228" precision="925" value="11.333333">11°20N</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FEE8C7BD92237AA8" box="[290,379,222,248]" degrees="74" direction="west" minutes="2" orientation="longitude" pageId="27" pageNumber="228" precision="925" value="-74.03333">74°2W</geoCoordinate>
,
<date id="242F77C3FFBFB048FE4CC7BD91407AA8" box="[390,536,222,248]" pageId="27" pageNumber="228" value="2001-05-04" valueMax="2001-05-23" valueMin="2001-05-04">
<collectingDate id="346B8E2BFFBFB048FE4CC7BD91407AA8" box="[390,536,222,248]" pageId="27" pageNumber="228" value="2001-05-04" valueMax="2001-05-23" valueMin="2001-05-04">4-23/V/2001</collectingDate>
</date>
, R.
<collectorName id="FD6434D5FFBFB048FD88C7BD91EF7AA8" box="[578,695,222,248]" pageId="27" pageNumber="228">Henriquez</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FD0EC7BD900A7AA8" box="[708,850,222,248]" pageId="27" pageNumber="228">USDA-ARS</collectionCode>
<specimenCount id="46979A8AFFBFB048FC93C7BD90227AA8" box="[857,890,222,248]" count="1" pageId="27" pageNumber="228" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FC58C7BD90997B4C" collectingDate="1989-03-05" collectionCode="BBSL" collectorName="S. Canas. &amp; F. D. Parker" country="Costa Rica" elevation="19" latitude="10.391666" location="Guanacaste" longLatPrecision="1" longitude="-85.149445" pageId="27" pageNumber="228" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guanacaste">
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<collectingCountry id="28861193FFBFB048FC58C7BD97197AA8" box="[914,1089,222,248]" name="Costa Rica" pageId="27" pageNumber="228">COSTA RICA</collectingCountry>
:
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,
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], 1,
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(
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)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FC04C66090097B10" collectingDate="1991-03-16" collectionCode="BBSL" collectorName="Parker" country="Costa Rica" elevation="19" latitude="10.391666" location="14 km S Canas" longLatPrecision="1" longitude="-85.149445" pageId="27" pageNumber="228" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste">
EJN
<location id="554E07D8FFBFB048FBCCC66197F47B4C" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FBCCC66197F47B4C" box="[1030,1196,258,284]" country="Costa Rica" latitude="10.391666" longLatPrecision="1" longitude="-85.149445" name="14 km S Canas" pageId="27" pageNumber="228" stateProvince="Guanacaste">
<quantity id="9769FCE6FFBFB048FBCCC661971E7B4C" box="[1030,1094,258,284]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="228" unit="km" value="14.0">14km</quantity>
S Cañas
</location>
, [
<quantity id="9769FCE6FFBFB048FB75C66197A57B4D" box="[1215,1277,258,285]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="27" pageNumber="228" unit="m" value="19.0">
<elevation id="DBBCB630FFBFB048FB75C66197A57B4D" box="[1215,1277,258,285]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="27" pageNumber="228" unit="m" value="19.0">19 m</elevation>
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,
<geoCoordinate id="35A537C4FFBFB048FAC3C66196C47B4C" box="[1289,1436,258,284]" degrees="10.39166667" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="1" value="10.391666">10.39166667</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FF5DC64592627B11" box="[151,314,294,321]" degrees="85.14944444" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="1" value="-85.149445">-85.14944444</geoCoordinate>
], 12/II,
<date id="242F77C3FFBFB048FE53C64591447B10" box="[409,540,294,320]" pageId="27" pageNumber="228" value="1991-03-16">
<collectingDate id="346B8E2BFFBFB048FE53C64591447B10" box="[409,540,294,320]" pageId="27" pageNumber="228" value="1991-03-16">16/III/1991</collectingDate>
</date>
, F. D.
<collectorName id="FD6434D5FFBFB048FDBEC64591E57B10" box="[628,701,294,320]" pageId="27" pageNumber="228">Parker</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FD1BC64590407B10" box="[721,792,294,320]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="27" pageNumber="228" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBFB048FCEEC645901D7B10" box="[804,837,294,320]" count="1" pageId="27" pageNumber="228" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FCA9C64596C17B34" collectingDate="1971-12-04" collectingDateMax="1972-02-07" collectingDateMin="1971-12-04" collectionCode="BBSL" collectorName="NW Canas &amp; E. R. Heithaus" country="Costa Rica" elevation="1165" latitude="10.3425" location="Hacienda" longLatPrecision="5" longitude="-85.299446" pageId="27" pageNumber="228" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Guanacaste">
<location id="554E07D8FFBFB048FCA9C64590957B10" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FCA9C64590957B10" box="[867,973,294,320]" country="Costa Rica" latitude="10.3425" longLatPrecision="5" longitude="-85.299446" name="Hacienda" pageId="27" pageNumber="228" stateProvince="Guanacaste">Hacienda</location>
COMELCO
<quantity id="9769FCE6FFBFB048FBA4C64597E57B10" box="[1134,1213,294,320]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="27" pageNumber="228" unit="km" value="24.0">24 km</quantity>
.
<collectorName id="FD6434D5FFBFB048FB07C64496167B10" box="[1229,1358,294,321]" pageId="27" pageNumber="228">NW Canas</collectorName>
, Inter-Am Hwy.,
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<elevation id="DBBCB630FFBFB048FED5C62992E07B35" box="[287,440,330,357]" metricMagnitude="3" metricUnit="m" metricValue="1.165" metricValueMax="1.2" metricValueMin="1.13" pageId="27" pageNumber="228" unit="m" value="1165.0" valueMax="1200.0" valueMin="1130.0">1130-1200 m</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FFBFB048FE06C62991727B35" box="[460,554,330,357]" degrees="10.3425" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="5" value="10.3425">10.3425</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FDF9C629918E7B35" box="[563,726,330,357]" degrees="85.29944444" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="1" value="-85.299446">-85.29944444</geoCoordinate>
],
<date id="242F77C3FFBFB048FD23C629902E7B34" box="[745,886,330,356]" pageId="27" pageNumber="228" value="1971-12-04">
<collectingDate id="346B8E2BFFBFB048FD23C629902E7B34" box="[745,886,330,356]" pageId="27" pageNumber="228" value="1971-12-04">04/XII/1971</collectingDate>
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, 04,
<date id="242F77C3FFBFB048FC65C62997707B34" box="[943,1064,330,356]" pageId="27" pageNumber="228" value="1972-02-07">
<collectingDate id="346B8E2BFFBFB048FC65C62997707B34" box="[943,1064,330,356]" pageId="27" pageNumber="228" value="1972-02-07">07/II/1972</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBFB048FBFFC62897857B34" box="[1077,1245,330,357]" pageId="27" pageNumber="228">E. R. Heithaus</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FB27C629966C7B34" box="[1261,1332,330,356]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="27" pageNumber="228" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBFB048FAF7C62996387B34" box="[1341,1376,330,356]" count="2" pageId="27" pageNumber="228" type="female">2♀</specimenCount>
<specimenCount id="46979A8AFFBFB048FAA3C62996D67B34" box="[1385,1422,330,356]" count="3" pageId="27" pageNumber="228" type="male">3♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FF5DC60D97FB7BD8" box="[151,1187,366,393]" collectingDate="2001-11-18" collectionCode="KUNHM-ENT" collectorName="R. Brooks" country="Costa Rica" elevation="10" latitude="10.396918" location="Palo Verde Sta" longLatPrecision="1" longitude="-85.32351" pageId="27" pageNumber="228" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste">
<location id="554E07D8FFBFB048FF5DC60D921A7BD8" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FF5DC60D921A7BD8" box="[151,322,366,392]" country="Costa Rica" latitude="10.396918" longLatPrecision="1" longitude="-85.32351" name="Palo Verde Sta" pageId="27" pageNumber="228" stateProvince="Guanacaste">Palo Verde Sta</location>
,
<quantity id="9769FCE6FFBFB048FE9AC60D92D57BD9" box="[336,397,366,393]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="228" unit="m" value="10.0">
<elevation id="DBBCB630FFBFB048FE9AC60D92D57BD9" box="[336,397,366,393]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="228" unit="m" value="10.0">10 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFBFB048FE51C60D914A7BD8" box="[411,530,366,392]" degrees="10.396918" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="1" value="10.396918">10.396918</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FDD6C60D91C67BD9" box="[540,670,366,393]" degrees="85.323511" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="1" value="-85.32351">-85.323511</geoCoordinate>
,
<date id="242F77C3FFBFB048FD61C60D90777BD8" box="[683,815,366,392]" pageId="27" pageNumber="228" value="2001-11-18">
<collectingDate id="346B8E2BFFBFB048FD61C60D90777BD8" box="[683,815,366,392]" pageId="27" pageNumber="228" value="2001-11-18">18/XI/2001</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBFB048FCF6C60C90E87BD8" box="[828,944,366,393]" pageId="27" pageNumber="228">R. Brooks</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FC0BC60D97347BD8" box="[961,1132,366,392]" pageId="27" pageNumber="228">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFFBFB048FBBFC60D97CE7BD8" box="[1141,1174,366,392]" count="1" pageId="27" pageNumber="228" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FB78C60D90267BFC" collectingDate="2001-11-22" collectionCode="KUNHM-ENT" country="Costa Rica" elevation="520" latitude="10.755658" location="Liberia" longLatPrecision="1" longitude="-85.37553" municipality="Km" pageId="27" pageNumber="228" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste">
21.7
<collectingMunicipality id="B04ACB79FFBFB048FB28C60D96547BD8" box="[1250,1292,366,392]" pageId="27" pageNumber="228">Km</collectingMunicipality>
NE
<location id="554E07D8FFBFB048FA8EC60D96C07BD8" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FA8EC60D96C07BD8" box="[1348,1432,366,392]" country="Costa Rica" latitude="10.755658" longLatPrecision="1" longitude="-85.37553" municipality="Km" name="Liberia" pageId="27" pageNumber="228" stateProvince="Guanacaste">Liberia</location>
,
<quantity id="9769FCE6FFBFB048FF5DC6F193BB7BFD" box="[151,227,402,429]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="27" pageNumber="228" unit="m" value="520.0">
<elevation id="DBBCB630FFBFB048FF5DC6F193BB7BFD" box="[151,227,402,429]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="27" pageNumber="228" unit="m" value="520.0">520 m</elevation>
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,
<geoCoordinate id="35A537C4FFBFB048FF3BC6F192307BFD" box="[241,360,402,429]" degrees="10.755658" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="1" value="10.755658">10.755658</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FEBBC6F192AD7BFD" box="[369,501,402,429]" degrees="85.375528" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="1" value="-85.37553">-85.375528</geoCoordinate>
,
<date id="242F77C3FFBFB048FDC8C6F191DF7BFC" box="[514,647,402,428]" pageId="27" pageNumber="228" value="2001-11-22">
<collectingDate id="346B8E2BFFBFB048FDC8C6F191DF7BFC" box="[514,647,402,428]" pageId="27" pageNumber="228" value="2001-11-22">22/XI/2001</collectingDate>
</date>
, (
<collectionCode id="3680C9C6FFBFB048FD51C6F1901E7BFC" box="[667,838,402,428]" pageId="27" pageNumber="228">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFFBFB048FC9AC6F190297BFC" box="[848,881,402,428]" count="1" pageId="27" pageNumber="228" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FC47C6F1918A7B80" collectingDate="1991-11-09" collectionCode="BBSL" collectorName="F. D. Parker" country="Costa Rica" elevation="26" latitude="10.4775" location="Potrero" longLatPrecision="5" longitude="-85.780556" municipality="Sugar Beach" pageId="27" pageNumber="228" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste">
<collectingMunicipality id="B04ACB79FFBFB048FC47C6F197457BFC" box="[909,1053,402,428]" pageId="27" pageNumber="228">Sugar Beach</collectingMunicipality>
3kmN
<location id="554E07D8FFBFB048FBBCC6F097937BFD" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FBBCC6F097937BFD" box="[1142,1227,403,429]" country="Costa Rica" latitude="10.4775" longLatPrecision="5" longitude="-85.780556" municipality="Sugar Beach" name="Potrero" pageId="27" pageNumber="228" stateProvince="Guanacaste">Potrero</location>
, [
<quantity id="9769FCE6FFBFB048FB2AC6F196787BFD" box="[1248,1312,402,429]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="27" pageNumber="228" unit="m" value="26.0">
<elevation id="DBBCB630FFBFB048FB2AC6F196787BFD" box="[1248,1312,402,429]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="27" pageNumber="228" unit="m" value="26.0">26 m</elevation>
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,
<geoCoordinate id="35A537C4FFBFB048FAE4C6F196D17BFD" box="[1326,1417,402,429]" degrees="10.4775" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="5" value="10.4775">10.4775</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FA59C6F092687B81" degrees="85.78055556" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="1" value="-85.780556">- 85.78055556</geoCoordinate>
],
<date id="242F77C3FFBFB048FE8BC6D592EF7B80" box="[321,439,438,464]" pageId="27" pageNumber="228" value="1991-11-09">
<collectingDate id="346B8E2BFFBFB048FE8BC6D592EF7B80" box="[321,439,438,464]" pageId="27" pageNumber="228" value="1991-11-09">9/XI/1991</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBFB048FE08C6D491107B80" box="[450,584,438,465]" pageId="27" pageNumber="228">F. D. Parker</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FD9CC6D591C57B80" box="[598,669,438,464]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="27" pageNumber="228" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBFB048FD6EC6D591907B80" box="[676,712,438,464]" count="1" pageId="27" pageNumber="228" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FD14C6D591157BA4" collectingDate="1976-03-01" collectionCode="BBSL" collectorName="PU Puntarenas &amp; R. M. Bohart" country="Costa Rica" elevation="6" latitude="9.978056" location="Puntarenas" longLatPrecision="1" longitude="-84.821945" pageId="27" pageNumber="228" specimenCount="2" specimenCount-female="2" stateProvince="Puntarenas">
<emphasis id="62E58D11FFBFB048FD14C6D590357B80" bold="true" box="[734,877,438,464]" pageId="27" pageNumber="228">
<collectingRegion id="92559FE1FFBFB048FD14C6D590307B80" box="[734,872,438,464]" country="Costa Rica" name="Puntarenas" pageId="27" pageNumber="228">Puntarenas</collectingRegion>
,
</emphasis>
<collectorName id="FD6434D5FFBFB048FCBEC6D497447B81" box="[884,1052,439,465]" pageId="27" pageNumber="228">PU Puntarenas</collectorName>
,
<quantity id="9769FCE6FFBFB048FBEDC6D5970C7B81" box="[1063,1108,438,465]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="27" pageNumber="228" unit="m" value="6.0">
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,
<geoCoordinate id="35A537C4FFBFB048FB95C6D597AA7B81" box="[1119,1266,438,465]" degrees="9.978055556" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="1" value="9.978056">9.978055556</geoCoordinate>
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,
<date id="242F77C3FFBFB048FF5DC6B992427BA4" box="[151,282,474,500]" pageId="27" pageNumber="228" value="1976-03-01">
<collectingDate id="346B8E2BFFBFB048FF5DC6B992427BA4" box="[151,282,474,500]" pageId="27" pageNumber="228" value="1976-03-01">01/III/1976</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBFB048FEEDC6B892E67BA4" box="[295,446,474,501]" pageId="27" pageNumber="228">R. M. Bohart</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FE05C6B9914E7BA4" box="[463,534,474,500]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="27" pageNumber="228" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBFB048FDD4C6B9911B7BA4" box="[542,579,474,500]" count="2" pageId="27" pageNumber="228" type="female">2♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FD91C6B991B47848" collectingDate="10 / III / 1985, 10 / III / 1985" collectionCode="KUNHM-ENT" collectorName="Km W &amp; W. Wcislo" country="Costa Rica" elevation="1395" latitude="10.0622225" location="Alajuela" longLatPrecision="1" longitude="-84.13167" pageId="27" pageNumber="228" specimenCount="1" specimenCount-female="1" stateProvince="San Jose">
<emphasis id="62E58D11FFBFB048FD91C6B991937BA4" bold="true" box="[603,715,473,500]" pageId="27" pageNumber="228">
<collectingRegion id="92559FE1FFBFB048FD91C6B9919E7BA4" box="[603,710,473,500]" country="Costa Rica" name="San Jose" pageId="27" pageNumber="228">San José</collectingRegion>
,
</emphasis>
15
<collectorName id="FD6434D5FFBFB048FD27C6B990657BA5" box="[749,829,474,501]" pageId="27" pageNumber="228">Km W</collectorName>
,
<location id="554E07D8FFBFB048FC82C6B990F27BA4" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FC82C6B990F27BA4" box="[840,938,474,500]" country="Costa Rica" latitude="10.0622225" longLatPrecision="1" longitude="-84.13167" name="Alajuela" pageId="27" pageNumber="228" stateProvince="San Jose">Alájuela</location>
, [
<quantity id="9769FCE6FFBFB048FC74C6B997417BA5" box="[958,1049,474,501]" metricMagnitude="3" metricUnit="m" metricValue="1.395" pageId="27" pageNumber="228" unit="m" value="1395.0">
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</quantity>
,
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],
<collectingDate id="346B8E2BFFBFB048FAB3C6B992DA7849" isEnumeration="true" pageId="27" pageNumber="228">
<date id="242F77C3FFBFB048FAB3C6B993AD7849" pageId="27" pageNumber="228" value="1985-03-10">10/ III/1985</date>
,
<date id="242F77C3FFBFB048FECAC69D92DA7849" box="[256,386,510,537]" pageId="27" pageNumber="228" value="1985-03-10">10/III/1985</date>
</collectingDate>
,
<collectorName id="FD6434D5FFBFB048FE46C69C92A77848" box="[396,511,510,537]" pageId="27" pageNumber="228">W. Wcislo</collectorName>
(
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<specimenCount id="46979A8AFFBFB048FD74C69D91877848" box="[702,735,510,536]" count="1" pageId="27" pageNumber="228" type="female">1♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FD3DC69D975E786C" collectingDate="2009-03-13" collectionCode="ECOAB" collectorName="J. Merida" country="Mexico" elevation="5" latitude="16.32131" location="Arriaga" longLatPrecision="1" longitude="-93.9832" pageId="27" pageNumber="228" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Chiapas">
<emphasis id="62E58D11FFBFB048FD3DC69D90BD7848" bold="true" box="[759,997,510,536]" pageId="27" pageNumber="228">
<collectingCountry id="28861193FFBFB048FD3DC69D902A7848" box="[759,882,510,536]" name="Mexico" pageId="27" pageNumber="228">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFBFB048FCB6C69D90B87848" box="[892,992,510,536]" country="Mexico" name="Chiapas" pageId="27" pageNumber="228">Chiapas</collectingRegion>
,
</emphasis>
<location id="554E07D8FFBFB048FC20C69D971A7848" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FC20C69D971A7848" box="[1002,1090,510,536]" country="Mexico" latitude="16.32131" longLatPrecision="1" longitude="-93.9832" name="Arriaga" pageId="27" pageNumber="228" stateProvince="Chiapas">Arriaga</location>
, 367,
<quantity id="9769FCE6FFBFB048FBB4C69D97F17849" box="[1150,1193,510,537]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="228" unit="m" value="5.0">
<elevation id="DBBCB630FFBFB048FBB4C69D97F17849" box="[1150,1193,510,537]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="228" unit="m" value="5.0">5 m</elevation>
</quantity>
,
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, 20/I,
<date id="242F77C3FFBFB048FF1BC541920C786C" box="[209,340,546,572]" pageId="27" pageNumber="228" value="2009-03-13">
<collectingDate id="346B8E2BFFBFB048FF1BC541920C786C" box="[209,340,546,572]" pageId="27" pageNumber="228" value="2009-03-13">13/III/2009</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBFB048FE94C5409291786C" box="[350,457,546,573]" pageId="27" pageNumber="228">J. Mérida</collectorName>
, C.
<taxonomicName id="97912A80FFBFB048FE39C5419062786C" authority=", P. Sagot, M. Guzman" authorityName="P. Sagot, M. Guzman" box="[499,826,546,573]" class="Magnoliopsida" family="Fabaceae" genus="Balboa" kingdom="Plantae" order="Fabales" pageId="27" pageNumber="228" phylum="Tracheophyta" rank="genus">Balboa, P. Sagot, M. Guzmán</taxonomicName>
(
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<specimenCount id="46979A8AFFBFB048FC64C5419089786C" box="[942,977,546,572]" count="4" pageId="27" pageNumber="228" type="female">4♀</specimenCount>
<specimenCount id="46979A8AFFBFB048FC1DC54190A5786C" box="[983,1021,546,572]" count="2" pageId="27" pageNumber="228" type="male">2♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FBDAC54192A87830" collectingDate="2009-03-12" collectionCode="ECOAB" country="Mexico" elevation="5" latitude="16.1352" location="Tonala" longLatPrecision="7" longitude="-93.6824" pageId="27" pageNumber="228" specimenCount="4" specimenCount-female="4" stateProvince="Chiapas">
<location id="554E07D8FFBFB048FBDAC5419707786C" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FBDAC5419707786C" box="[1040,1119,546,572]" country="Mexico" latitude="16.1352" longLatPrecision="7" longitude="-93.6824" name="Tonala" pageId="27" pageNumber="228" stateProvince="Chiapas">Tonalá</location>
, 407,
<quantity id="9769FCE6FFBFB048FB50C541979D786D" box="[1178,1221,546,573]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="228" unit="m" value="5.0">
<elevation id="DBBCB630FFBFB048FB50C541979D786D" box="[1178,1221,546,573]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="228" unit="m" value="5.0">5 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFBFB048FB1AC5419673786D" box="[1232,1323,546,573]" degrees="16.1352" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="5" value="16.1352">16.1352</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FAFBC54196C0786C" box="[1329,1432,546,572]" degrees="93.6824" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="5" value="-93.6824">-93.6824</geoCoordinate>
, 09,
<date id="242F77C3FFBFB048FF08C525921D7830" box="[194,325,582,608]" pageId="27" pageNumber="228" value="2009-03-12">
<collectingDate id="346B8E2BFFBFB048FF08C525921D7830" box="[194,325,582,608]" pageId="27" pageNumber="228" value="2009-03-12">12/III/2009</collectingDate>
</date>
, (
<collectionCode id="3680C9C6FFBFB048FE92C52592E17830" box="[344,441,582,608]" pageId="27" pageNumber="228">ECOAB</collectionCode>
<specimenCount id="46979A8AFFBFB048FE0BC52592BE7830" box="[449,486,582,608]" count="4" pageId="27" pageNumber="228" type="female">4♀</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FE37C52592BE78D4" collectingDate="1990-09-01" collectionCode="KUNHM-ENT" collectorName="L. M. Godinez" country="Mexico" elevation="10" latitude="15.866944" location="Puerto Escondido" longLatPrecision="1" longitude="-97.09194" pageId="27" pageNumber="228" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
<emphasis id="62E58D11FFBFB048FE37C52591067830" bold="true" box="[509,606,582,608]" pageId="27" pageNumber="228">
<collectingRegion id="92559FE1FFBFB048FE37C52591017830" box="[509,601,582,608]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="228">Oaxaca</collectingRegion>
,
</emphasis>
<location id="554E07D8FFBFB048FDACC52490687830" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FDACC52490687830" box="[614,816,582,609]" country="Mexico" latitude="15.866944" longLatPrecision="1" longitude="-97.09194" name="Puerto Escondido" pageId="27" pageNumber="228" stateProvince="Oaxaca">Puerto Escondido</location>
,
<quantity id="9769FCE6FFBFB048FCF6C52590217831" box="[828,889,582,609]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="228" unit="m" value="10.0">
<elevation id="DBBCB630FFBFB048FCF6C52590217831" box="[828,889,582,609]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="228" unit="m" value="10.0">10 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFBFB048FC4FC52597407831" box="[901,1048,582,609]" degrees="15.86694444" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="1" value="15.866944">15.86694444</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FBEBC52597987830" box="[1057,1216,582,608]" degrees="97.09194444" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="1" value="-97.09194">-97.09194444</geoCoordinate>
,
<date id="242F77C3FFBFB048FB06C52596087830" box="[1228,1360,582,608]" pageId="27" pageNumber="228" value="1990-09-01">
<collectingDate id="346B8E2BFFBFB048FB06C52596087830" box="[1228,1360,582,608]" pageId="27" pageNumber="228" value="1990-09-01">01/IX/1990</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBFB048FA96C52493AE78D4" pageId="27" pageNumber="228">L. M. Godínez</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FECFC50992E878D4" box="[261,432,618,644]" pageId="27" pageNumber="228">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFFBFB048FE7DC509928378D4" box="[439,475,618,644]" count="1" pageId="27" pageNumber="228" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FE38C509939E78F8" collectingDate="2018-12-18" collectionCode="ECOAB" collectorName="O. Caballero" country="Mexico" elevation="64" latitude="15.9487" location="San Pedro Tututepec" longLatPrecision="7" longitude="-97.2825" pageId="27" pageNumber="228" specimenCount="2" specimenCount-male="2" stateProvince="Oaxaca">
<location id="554E07D8FFBFB048FE38C509918578D5" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FE38C509918578D5" box="[498,733,618,645]" country="Mexico" latitude="15.9487" longLatPrecision="7" longitude="-97.2825" name="San Pedro Tututepec" pageId="27" pageNumber="228" stateProvince="Oaxaca">San Pedro Tututepec</location>
,
<quantity id="9769FCE6FFBFB048FD22C509907C78D5" box="[744,804,618,645]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="27" pageNumber="228" unit="m" value="64.0">
<elevation id="DBBCB630FFBFB048FD22C509907C78D5" box="[744,804,618,645]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="27" pageNumber="228" unit="m" value="64.0">64 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFBFB048FCFAC50990D378D5" box="[816,907,618,645]" degrees="15.9487" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="5" value="15.9487">15.9487</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FC58C50990A278D5" box="[914,1018,618,645]" degrees="97.2825" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="5" value="-97.2825">-97.2825</geoCoordinate>
,
<date id="242F77C3FFBFB048FBCFC50997CA78D4" box="[1029,1170,618,644]" pageId="27" pageNumber="228" value="2018-12-18">
<collectingDate id="346B8E2BFFBFB048FBCFC50997CA78D4" box="[1029,1170,618,644]" pageId="27" pageNumber="228" value="2018-12-18">18/XII/2018</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBFB048FB57C509967578D4" box="[1181,1325,618,644]" pageId="27" pageNumber="228">O. Caballero</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FAF6C50996C578D4" box="[1340,1437,618,644]" pageId="27" pageNumber="228">ECOAB</collectionCode>
<specimenCount id="46979A8AFFBFB048FF5DC5ED93E578F8" box="[151,189,654,680]" count="2" pageId="27" pageNumber="228" type="male">2♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBFB048FF1EC5ED979C78F8" box="[212,1220,654,681]" collectingDate="2012-01-28" collectionCode="ECOAB" collectorName="P. Sagot" country="Mexico" elevation="19" latitude="15.6668" location="Santa Maria Tonameca" longLatPrecision="7" longitude="-96.5406" pageId="27" pageNumber="228" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Oaxaca">
<location id="554E07D8FFBFB048FF1EC5ED928778F9" LSID="urn:lsid:plazi:treatment:D838E015FFBDB048FF5DC2E9925E7E2D:554E07D8FFBFB048FF1EC5ED928778F9" box="[212,479,654,681]" country="Mexico" latitude="15.6668" longLatPrecision="7" longitude="-96.5406" name="Santa Maria Tonameca" pageId="27" pageNumber="228" stateProvince="Oaxaca">Santa María Tonameca</location>
,
<quantity id="9769FCE6FFBFB048FE26C5ED917278F9" box="[492,554,654,681]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="27" pageNumber="228" unit="m" value="19.0">
<elevation id="DBBCB630FFBFB048FE26C5ED917278F9" box="[492,554,654,681]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="27" pageNumber="228" unit="m" value="19.0">19 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFBFB048FDF3C5ED91CC78F9" box="[569,660,654,681]" degrees="15.6668" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="5" value="15.6668">15.6668</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FD57C5ED905D78F9" box="[669,773,654,681]" degrees="96.5406" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="5" value="-96.5406">-96.5406</geoCoordinate>
,
<date id="242F77C3FFBFB048FCD9C5ED90DA78F8" box="[787,898,654,680]" pageId="27" pageNumber="228" value="2012-01-28">
<collectingDate id="346B8E2BFFBFB048FCD9C5ED90DA78F8" box="[787,898,654,680]" pageId="27" pageNumber="228" value="2012-01-28">28/I/2012</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBFB048FC5AC5EC90B678F8" box="[912,1006,654,681]" pageId="27" pageNumber="228">P. Sagot</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FC35C5ED973878F8" box="[1023,1120,654,680]" pageId="27" pageNumber="228">ECOAB</collectionCode>
<specimenCount id="46979A8AFFBFB048FBA3C5ED97D478F8" box="[1129,1164,654,680]" count="3" pageId="27" pageNumber="228" type="female">3♀</specimenCount>
<specimenCount id="46979A8AFFBFB048FB5CC5ED97E278F8" box="[1174,1210,654,680]" count="1" pageId="27" pageNumber="228" type="male">1♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFFBFB048FB19C5ED9123789C" collectingDate="2012-01-29" collectionCode="ECOAB" collectorName="Sagot" country="Mexico" elevation="30" latitude="15.67066" longLatPrecision="5" longitude="-96.538" pageId="27" pageNumber="228" specimenCount="2" specimenCount-female="2" stateProvince="Oaxaca">
<quantity id="9769FCE6FFBFB048FB19C5ED964978F9" box="[1235,1297,654,681]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="228" unit="m" value="30.0">
<elevation id="DBBCB630FFBFB048FB19C5ED964978F9" box="[1235,1297,654,681]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="228" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFBFB048FAEAC5ED96D178F9" box="[1312,1417,654,681]" degrees="15.67066" direction="north" orientation="latitude" pageId="27" pageNumber="228" precision="1" value="15.67066">15.67066</geoCoordinate>
<geoCoordinate id="35A537C4FFBFB048FA59C5EC93AD789D" degrees="96.5380" direction="west" orientation="longitude" pageId="27" pageNumber="228" precision="5" value="-96.538">- 96.5380</geoCoordinate>
,
<date id="242F77C3FFBFB048FECAC5D19228789C" box="[256,368,690,716]" pageId="27" pageNumber="228" value="2012-01-29">
<collectingDate id="346B8E2BFFBFB048FECAC5D19228789C" box="[256,368,690,716]" pageId="27" pageNumber="228" value="2012-01-29">29/I/2012</collectingDate>
</date>
, P.
<collectorName id="FD6434D5FFBFB048FE5CC5D1928E789C" box="[406,470,690,716]" pageId="27" pageNumber="228">Sagot</collectorName>
(
<collectionCode id="3680C9C6FFBFB048FE2EC5D1911D789C" box="[484,581,690,716]" pageId="27" pageNumber="228">ECOAB</collectionCode>
<specimenCount id="46979A8AFFBFB048FD86C5D19128789C" box="[588,624,690,716]" count="2" pageId="27" pageNumber="228" type="female">2♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBFB048FF0DC5B5962A79D1" pageId="27" pageNumber="228" type="distribution">
<paragraph id="502E5103FFBFB048FF0DC5B5962A79D1" blockId="27.[151,1437,150,1149]" pageId="27" pageNumber="228">
<emphasis id="62E58D11FFBFB048FF0DC5B5923978A0" bold="true" box="[199,353,726,752]" pageId="27" pageNumber="228">Distribution.</emphasis>
This species was previously registered as present in
<collectingCountry id="28861193FFBFB048FC7BC5B5975278A0" box="[945,1034,726,752]" name="Panama" pageId="27" pageNumber="228">Panamá</collectingCountry>
and
<collectingCountry id="28861193FFBFB048FB8AC5B597E478A0" box="[1088,1212,726,752]" name="Costa Rica" pageId="27" pageNumber="228">Costa Rica</collectingCountry>
, but here we extend the distribution northwards to South Mexico, in the Pacific slope along the coast of Oaxaca and Chiapas, areas with tropical dry forest, basically in the Pacific Lowlands biogeographic province (
<bibRefCitation id="34002CF2FFBFB048FBC8C47D97B07968" author="Morrone, J. J. &amp; Escalante, T. &amp; Rodriguez-Tapia, G. &amp; Carmona, A. &amp; Arana, M. &amp; Mercado-Gomez, J. D." box="[1026,1256,798,825]" pageId="27" pageNumber="228" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. &amp; Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="62E58D11FFBFB048FBA7C47C97C67968" box="[1133,1182,798,824]" italics="true" pageId="27" pageNumber="228">et al</emphasis>
. 2022
</bibRefCitation>
), and also south to
<collectingCountry id="28861193FFBFB048FF79C4219279790C" box="[179,289,834,860]" name="Colombia" pageId="27" pageNumber="228">Colombia</collectingCountry>
with the synonymy of
<taxonomicName id="97912A80FFBFB048FDD5C42091CF790C" box="[543,663,835,860]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="62E58D11FFBFB048FDD5C42091CF790C" box="[543,663,835,860]" italics="true" pageId="27" pageNumber="228">H. tayrona</emphasis>
</taxonomicName>
. There are also records from
<collectingCountry id="28861193FFBFB048FC10C421974D790C" box="[986,1045,834,860]" name="Cuba" pageId="27" pageNumber="228">Cuba</collectingCountry>
and The Española Island, but these females registered likely are a new species yet to be described (
<figureCitation id="C8AA4D86FFBFB048FC94C40590E979D0" box="[862,945,870,896]" captionStart="FIGURE 14" captionStartId="25.[151,250,1098,1122]" captionTargetBox="[160,1428,187,1064]" captionTargetId="figure-18@25.[151,1436,179,1072]" captionTargetPageId="25" captionText="FIGURE 14. Known distribution records for Heriades currani." pageId="27" pageNumber="228">Fig. 14</figureCitation>
). Altitudinal range: sea level to
<quantity id="9769FCE6FFBFB048FADCC405963679D1" box="[1302,1390,870,897]" metricMagnitude="3" metricUnit="m" metricValue="1.395" pageId="27" pageNumber="228" unit="m" value="1395.0">1395 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBFB048FF0DC4E9925E7E2D" pageId="27" pageNumber="228" type="discussion">
<paragraph id="502E5103FFBFB048FF0DC4E9925E7E2D" blockId="27.[151,1437,150,1149]" pageId="27" pageNumber="228">
<emphasis id="62E58D11FFBFB048FF0DC4E9921779F4" bold="true" box="[199,335,906,932]" pageId="27" pageNumber="228">Comments.</emphasis>
The
<typeStatus id="8F2AEFA1FFBFB048FE42C4E992B279F4" box="[392,490,906,932]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="97912A80FFBFB048FDA8C4E8918079F4" authorityName="Michener" authorityYear="1943" box="[610,728,907,932]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="62E58D11FFBFB048FDA8C4E8918079F4" box="[610,728,907,932]" italics="true" pageId="27" pageNumber="228">H. currani</emphasis>
</taxonomicName>
has small tubercles on apical margin of clypeus, however there is variation in the size, as well as, on outer surface of mandible in females, and small differences in the tubercle on the basal point of outer ridge. Also, there are small differences in the width and length of the hairs in the tergal fasciae. In males, S5, S6 and the genital capsule are similar to
<taxonomicName id="97912A80FFBFB048FCABC494967D7E40" authority="Gonzalez &amp; Griswold, 2011" authorityName="Gonzalez &amp; Griswold" authorityYear="2011" box="[865,1317,1014,1040]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="62E58D11FFBFB048FCABC49490837E40" box="[865,987,1015,1040]" italics="true" pageId="27" pageNumber="228">H. tayrona</emphasis>
<bibRefCitation id="34002CF2FFBFB048FC29C495967D7E40" author="Gonzalez, V. H. &amp; Griswold, T." box="[995,1317,1014,1040]" pageId="27" pageNumber="228" pagination="255 - 259" refId="ref43127" refString="Gonzalez, V. H. &amp; Griswold, T. (2011) Heriades tayrona N. Sp., the First Osmiine Bee from South America (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society, 84 (4), 255 - 259. https: // doi. org / 10.2317 / JKES 110317.1" type="journal article" year="2011">Gonzalez &amp; Griswold, 2011</bibRefCitation>
</taxonomicName>
, described from northern
<collectingCountry id="28861193FFBFB048FEF5C37992E97E64" box="[319,433,1050,1076]" name="Colombia" pageId="27" pageNumber="228">Colombia</collectingCountry>
. As males from South Mexico have an intermediate ocelli size between
<taxonomicName id="97912A80FFBFB048FB38C37896337E64" authorityName="Michener" authorityYear="1943" box="[1266,1387,1051,1076]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="62E58D11FFBFB048FB38C37896337E64" box="[1266,1387,1051,1076]" italics="true" pageId="27" pageNumber="228">H. currani</emphasis>
</taxonomicName>
and
<taxonomicName id="97912A80FFBFB048FF5DC35C924A7E08" authorityName="Gonzalez &amp; Griswold" authorityYear="2011" box="[151,274,1087,1112]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="62E58D11FFBFB048FF5DC35C924A7E08" box="[151,274,1087,1112]" italics="true" pageId="27" pageNumber="228">H. tayrona</emphasis>
</taxonomicName>
, and this was the only character upholding the species status of
<emphasis id="62E58D11FFBFB048FBC9C35C97DD7E08" box="[1027,1157,1087,1112]" italics="true" pageId="27" pageNumber="228">
<taxonomicName id="97912A80FFBFB048FBC9C35C97D97E08" authorityName="Gonzalez &amp; Griswold" authorityYear="2011" box="[1027,1153,1087,1112]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="tayrona">H. tayrona</taxonomicName>
,
</emphasis>
it is here added as new synonym.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,678 @@
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<mods:title id="F65E0ABD93C3DE7B15A4CF848F67F396">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<paragraph id="502E5103FFBFB048FF5DC3AD902E7EB8" blockId="27.[151,886,1230,1292]" box="[151,886,1230,1257]" pageId="27" pageNumber="228">
<heading id="0B66E66FFFBFB048FF5DC3AD902E7EB8" bold="true" box="[151,886,1230,1257]" fontSize="11" level="1" pageId="27" pageNumber="228" reason="1">
<emphasis id="62E58D11FFBFB048FF5DC3AD902E7EB8" bold="true" box="[151,886,1230,1257]" pageId="27" pageNumber="228">
<taxonomicName id="97912A80FFBFB048FF5DC3AD90497EB8" authority="Rojas-Arias, Griswold &amp; Ayala." authorityName="Rojas-Arias, Griswold &amp; Ayala." authorityYear="2025" box="[151,785,1230,1257]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="danuncia" status="sp. nov.">
<emphasis id="62E58D11FFBFB048FF5DC3AD92267EB9" bold="true" box="[151,382,1230,1257]" italics="true" pageId="27" pageNumber="228">Heriades danuncia</emphasis>
Rojas-Arias, Griswold &amp; Ayala.
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<taxonomicNameLabel id="79D6306AFFBFB048FCD3C3AD902E7EB8" box="[793,886,1230,1256]" pageId="27" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="188B0288FFBFB048FF5DC391920D7F5C" box="[151,341,1266,1292]" pageId="27" pageNumber="228" type="description">
<paragraph id="502E5103FFBFB048FF5DC391920D7F5C" blockId="27.[151,886,1230,1292]" box="[151,341,1266,1292]" pageId="27" pageNumber="228">
(
<figureCitation id="C8AA4D86FFBFB048FF55C39193A27F5C" box="[159,250,1266,1292]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="27" pageNumber="228">Figs. 2J</figureCitation>
,
<figureCitation id="C8AA4D86FFBFB048FECCC39192157F5C" box="[262,333,1266,1292]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStartId-0="28.[151,250,1098,1122]" captionStartId-1="29.[151,250,1943,1967]" captionTargetBox-0="[160,1428,187,1064]" captionTargetBox-1="[169,1398,185,1905]" captionTargetId-0="figure-22@28.[151,1436,179,1072]" captionTargetId-1="figure-18@29.[168,1420,179,1917]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionText-0="FIGURE 16. Known distribution records for Heriades danuncia sp. nov." captionText-1="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="27" pageNumber="228">1617</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBFB048FF5DC259970A7CED" pageId="27" pageNumber="228" type="diagnosis">
<paragraph id="502E5103FFBFB048FF5DC259970A7CED" blockId="27.[151,1437,1338,2013]" pageId="27" pageNumber="228">
<emphasis id="62E58D11FFBFB048FF5DC259924A7F04" bold="true" box="[151,274,1338,1364]" pageId="27" pageNumber="228">Diagnosis.</emphasis>
Ocelli medium-sized (
<figureCitation id="C8AA4D86FFBFB048FDDBC25991307F04" box="[529,616,1338,1364]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="27" pageNumber="228">Fig. 3B</figureCitation>
); punctation large along the whole body; dorsolateral angles of pronotum with tuft of plumose, short, whitish to yellowish hairs (
<figureCitation id="C8AA4D86FFBFB048FCCBC23D90D27F28" box="[769,906,1374,1400]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="27" pageNumber="228">Fig. 17B, D</figureCitation>
); tergal fasciae with row of thick, whitish hairs (
<figureCitation id="C8AA4D86FFBFB048FF55C2E193B77FCC" box="[159,239,1410,1436]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="27" pageNumber="228">Fig. 3J</figureCitation>
); body with bare areas on frons, disc of scutum, mesepisternum and terga.
<emphasis id="62E58D11FFBFB048FBFFC2E197377FCC" box="[1077,1135,1410,1436]" italics="true" pageId="27" pageNumber="228">Male</emphasis>
: outer surface of mandible with strong outer and condylar ridges, from base to midpoint, where is strongly interrupted; clypeus with plumose, white to goldish, dense hairs, decreasing in length towards mid clypeus and absent on lower third (Fig.
<figureCitation id="C8AA4D86FFBFB048FAEDC2A996D67FB4" box="[1319,1422,1482,1508]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="27" pageNumber="228">Fig. 17A</figureCitation>
); S5 with two subtriangular lobes and one line of capitate hairs directed and increasing in length outward (
<figureCitation id="C8AA4D86FFBFB048FAE0C28D96D77C58" box="[1322,1423,1518,1544]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="27" pageNumber="228">Fig. 17E</figureCitation>
); S6 with one posterior tuft of simple, short hairs and one subapical, dense line of capitate hairs, gonostylus with quadranqular apex, and long hairs (
<figureCitation id="C8AA4D86FFBFB048FDE7C15591CA7C00" box="[557,658,1590,1616]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="27" pageNumber="228">Fig. 17F</figureCitation>
).
<emphasis id="62E58D11FFBFB048FD62C15591A47C00" box="[680,764,1590,1616]" italics="true" pageId="27" pageNumber="228">Female</emphasis>
: outer surface of mandible with slight elevation at base of outer ridge; apical margin of clypeus with strong emargination and more than five small tubercles or crenulated; upper area of clypeus with medium-sized punctures. This species is similar to
<taxonomicName id="97912A80FFBFB048FBE5C11C979D7CC8" box="[1071,1221,1662,1688]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="micheneri">
<emphasis id="62E58D11FFBFB048FBE5C11C979D7CC8" box="[1071,1221,1662,1688]" italics="true" pageId="27" pageNumber="228">H. micheneri</emphasis>
</taxonomicName>
(
<figureCitation id="C8AA4D86FFBFB048FB12C11D963D7CC8" box="[1240,1381,1662,1689]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="27" pageNumber="228">Fig. 25A, B</figureCitation>
) but different by having slightly larger mesepisternum punctures and slightly wider scape.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBFB04DFF0DC1A591BD7F91" lastPageId="30" lastPageNumber="231" pageId="27" pageNumber="228" type="description">
<paragraph id="502E5103FFBFB048FF0DC1A5964B7CB0" blockId="27.[151,1437,1338,2013]" box="[199,1299,1734,1761]" pageId="27" pageNumber="228">
<emphasis id="62E58D11FFBFB048FF0DC1A592FC7CB0" bold="true" box="[199,420,1734,1761]" pageId="27" pageNumber="228">
Description.
<emphasis id="62E58D11FFBFB048FEAAC1A592C37CB1" bold="true" box="[352,411,1734,1761]" italics="true" pageId="27" pageNumber="228">Male</emphasis>
:
</emphasis>
Body length,
<quantity id="9769FCE6FFBFB048FD89C1A591C17CB1" box="[579,665,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="27" pageNumber="228" unit="mm" value="5.9">5.9 mm</quantity>
(5.26.3, n=6); forewing length,
<quantity id="9769FCE6FFBFB048FBC5C1A5973D7CB1" box="[1039,1125,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="27" pageNumber="228" unit="mm" value="3.6">3.6 mm</quantity>
(3.43.9, n=6).
</paragraph>
<paragraph id="502E5103FFBFB04FFF0DC18991067F9A" blockId="27.[151,1437,1338,2013]" lastBlockId="28.[151,1437,1168,2023]" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228">
<emphasis id="62E58D11FFBFB048FF0DC189925C7D54" box="[199,260,1770,1796]" italics="true" pageId="27" pageNumber="228">Head</emphasis>
. Length
<quantity id="9769FCE6FFBFB048FEA6C189929C7D55" box="[364,452,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="27" pageNumber="228" unit="mm" value="1.6">1.6 mm</quantity>
(1.51.7, n=6), width
<quantity id="9769FCE6FFBFB048FD0CC18990467D55" box="[710,798,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="27" pageNumber="228" unit="mm" value="1.6">1.6 mm</quantity>
(1.51.8, n=6); eyes about 2.9 times longer than broad, 1.9 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FFBFB048FC91C06D90E97D79" box="[859,945,1806,1833]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="27" pageNumber="228" unit="mm" value="0.8">0.8 mm</quantity>
(0.70.9, n=6); upper interocular distance
<quantity id="9769FCE6FFBFB048FA44C06D939B7D1D" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="228" unit="mm" value="1.0">1 mm</quantity>
(0.91, n=6); interantennal distance 2 times antennocular distance; lateral ocelli medium-sized, 1.4 times DOP in interocellar area, median ocellus smaller; interocellar distance 3.3 times OD; ocellocular distance 3 times OD; OPR distance 3.5 times OD; labrum with basal, medial, wide elevation; outer surface of mandible with strong outer and condylar ridge, from base to mid distance, strongly interrupted (
<figureCitation id="C8AA4D86FFBFB048FC47C0FD90AC7DE8" box="[909,1012,1950,1976]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="27" pageNumber="228">Fig. 17A</figureCitation>
); clypeus 1.5 times broader than long, apical margin with shallow emargination; juxtantennal carina absent but strongly angulated; scape black, length 0.7 times alveolocellar distance, 2 times longer than wide; pedicel black, about 1.3 times longer than F1 and about 0.8 times longer than F2; flagellum black, first broader than long remaining flagellomeres 1.8 longer than first; F11 2 times longer than broad.
<emphasis id="62E58D11FFB8B04FFE7AC3BA91777EA2" box="[432,559,1241,1266]" italics="true" pageId="28" pageNumber="229">Mesosoma.</emphasis>
Scutum 2.4 times longer than scutellum, 2.9 times longer than tegula, and 3.4 longer than axilla; intertegular distance
<quantity id="9769FCE6FFB8B04FFD9FC39F91F37F47" box="[597,683,1276,1303]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="28" pageNumber="229" unit="mm" value="1.1">1.1 mm</quantity>
(1.11.3, n=6); metanotum with transverse ridge, in posterior view with two small medial rounded elevations on top; propodeum with small to middle pits (
<figureCitation id="C8AA4D86FFB8B04FFBA1C24397BE7F6A" box="[1131,1254,1312,1338]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" pageId="28" pageNumber="229">Fig. 3G, H</figureCitation>
),
<quantity id="9769FCE6FFB8B04FFB33C24396057F6B" box="[1273,1373,1312,1339]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="229" unit="mm" value="0.06">0.06 mm</quantity>
long; legs dark amber.
<emphasis id="62E58D11FFB8B04FFE93C226928E7F0E" box="[345,470,1349,1374]" italics="true" pageId="28" pageNumber="229">Metasoma.</emphasis>
T1
<quantity id="9769FCE6FFB8B04FFDC9C22791027F0F" box="[515,602,1348,1375]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="229" unit="mm" value="1.4">1.4 mm</quantity>
broad (1.21.5, n=6); T3
<quantity id="9769FCE6FFB8B04FFC4BC22790807F0F" box="[897,984,1348,1375]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="28" pageNumber="229" unit="mm" value="1.6">1.6 mm</quantity>
broad (1.41.7, n=6); T6 with posterior margin not projected; posterior margin of S1 truncate without emargination (
<figureCitation id="C8AA4D86FFB8B04FFBC1C20B97097FD2" box="[1035,1105,1384,1410]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="28" pageNumber="229">Fig. 2</figureCitation>
Jb), in profile, with posterior margin projected and strong process apex on one half of distance from apex to base (
<figureCitation id="C8AA4D86FFB8B04FFB85C2EF97CB7FF6" box="[1103,1171,1420,1446]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="28" pageNumber="229">Fig. 2</figureCitation>
Ja), process in posterior view with sub-quadrate shape (
<figureCitation id="C8AA4D86FFB8B04FFE3EC2D3916F7F9A" box="[500,567,1456,1482]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." pageId="28" pageNumber="229">Fig. 2</figureCitation>
Jc).
</paragraph>
<caption id="04EE018BFFB8B04FFF5DC32990267E32" box="[151,894,1098,1122]" pageId="28" pageNumber="229" startId="28.[151,250,1098,1122]" targetBox="[160,1428,187,1064]" targetPageId="28" targetType="figure">
<paragraph id="502E5103FFB8B04FFF5DC32990267E32" blockId="28.[151,894,1098,1122]" box="[151,894,1098,1122]" pageId="28" pageNumber="229">
<emphasis id="62E58D11FFB8B04FFF5DC32992787E32" bold="true" box="[151,288,1098,1122]" pageId="28" pageNumber="229">FIGURE 16.</emphasis>
Known distribution records for
<taxonomicName id="97912A80FFB8B04FFDADC32990707E32" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[615,808,1098,1122]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="danuncia" status="sp. nov.">
<emphasis id="62E58D11FFB8B04FFDADC32990707E32" box="[615,808,1098,1122]" italics="true" pageId="28" pageNumber="229">Heriades danuncia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFB8B04FFCE5C32890267E32" box="[815,894,1099,1122]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="502E5103FFB8B04FFF0DC2B797A57DB7" blockId="28.[151,1437,1168,2023]" pageId="28" pageNumber="229">
<emphasis id="62E58D11FFB8B04FFF0DC2B792097FBE" box="[199,337,1492,1518]" italics="true" pageId="28" pageNumber="229">Sculpturing.</emphasis>
Clypeus with small (at base 0.4 times OD), almost contiguous, coarse and equally sized punctures; supraclypeal area with punctures slightly denser than base of clypeus; frons with punctures notably bigger, coarser and denser than supraclypeal area; on frons in front of ocelli slightly smaller than frons; vertex with bigger, sparser punctures than on supraclypeal area, with micropunctures between punctures as on vertex; hypostomal area with punctation bigger than on clypeus; gena with bigger, coarser and sparser punctures than on hypostomal area; anterior surface of pronotum with punctures sparser than on clypeus, and smaller, less coarse on pronotal lobes; disc of scutum with bigger (0.7 times OD), denser punctures than on vertex, with micropunctures between punctures; rest of mesosoma as on scutum except: scutellum with punctures smaller and sparser, mesepisternum with equal DBP as on scutum; metepisternum with smaller punctures, less dense and coarse; T1 with smaller (0.6 times OD), slightly denser punctures than on scutum, without micropunctures between punctures; T2 with punctures smaller and sparser than on T1; T3 with bigger, denser and coarse punctures than on T1, slightly rounded, separated by less than one quarter of DOP; T4 and T5 with bigger, denser and as coarse punctures as on T3; dorsal surface of T6 with slightly smaller and sparser punctation as on T5, on posterior surface with notably smaller and slightly sparser punctation, DBP less than one third DOP; sides of S1 with punctation bigger and parser than T1, and in process smaller and sparser; sides of S2 with punctures smaller than on T1, disc of S2 microreticulate without punctures.
</paragraph>
<caption id="04EE018BFFB9B04EFF5DC0F490187D83" pageId="29" pageNumber="230" startId="29.[151,250,1943,1967]" targetBox="[169,1398,185,1905]" targetPageId="29" targetType="figure">
<paragraph id="502E5103FFB9B04EFF5DC0F490187D83" blockId="29.[151,1437,1943,2003]" pageId="29" pageNumber="230">
<emphasis id="62E58D11FFB9B04EFF5DC0F492787DFF" bold="true" box="[151,288,1943,1967]" pageId="29" pageNumber="230">FIGURE 17.</emphasis>
<taxonomicName id="97912A80FFB9B04EFEEDC0F492B07DFF" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[295,488,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="danuncia" status="sp. nov.">
<emphasis id="62E58D11FFB9B04EFEEDC0F492B07DFF" box="[295,488,1943,1967]" italics="true" pageId="29" pageNumber="230">Heriades danuncia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFB9B04EFE25C0FB91667DFF" box="[495,574,1944,1967]" pageId="29" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
Male:
<emphasis id="62E58D11FFB9B04EFD4CC0F491C77DFF" bold="true" box="[646,671,1943,1967]" pageId="29" pageNumber="230">A.</emphasis>
frontal view
<emphasis id="62E58D11FFB9B04EFCEDC0F490677DFF" bold="true" box="[807,831,1943,1967]" pageId="29" pageNumber="230">B.</emphasis>
dorsal view. Female:
<emphasis id="62E58D11FFB9B04EFBD5C0F4976F7DFF" bold="true" box="[1055,1079,1943,1967]" pageId="29" pageNumber="230">C.</emphasis>
frontal view
<emphasis id="62E58D11FFB9B04EFB0AC0F497817DFF" bold="true" box="[1216,1241,1943,1967]" pageId="29" pageNumber="230">D.</emphasis>
dorsal view. Male:
<emphasis id="62E58D11FFB9B04EFF5DC0D893F67D83" bold="true" box="[151,174,1979,2003]" pageId="29" pageNumber="230">E.</emphasis>
S5.
<emphasis id="62E58D11FFB9B04EFF16C0D893B77D83" bold="true" box="[220,239,1979,2003]" pageId="29" pageNumber="230">F.</emphasis>
S6.
<emphasis id="62E58D11FFB9B04EFED7C0D8926F7D83" bold="true" box="[285,311,1979,2003]" pageId="29" pageNumber="230">G.</emphasis>
genital capsule in dorsal view and
<emphasis id="62E58D11FFB9B04EFD56C0D891EE7D83" bold="true" box="[668,694,1979,2003]" pageId="29" pageNumber="230">H.</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="502E5103FFBAB04DFF0DC7F496DE7BD9" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="231">
<emphasis id="62E58D11FFBAB04DFF0DC7F492687AE0" box="[199,304,151,176]" italics="true" pageId="30" pageNumber="231">Vestiture.</emphasis>
Clypeus with plumose, white to goldish, and dense hairs, decreasing in length towards mid clypeus, absent on lower third; supraclypeal area and frons with plumose, thick, white to goldish hairs directed medially and dorsally respectively, hairs similar in length (
<figureCitation id="C8AA4D86FFBAB04DFD45C7BC91AF7AA9" box="[655,759,223,249]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="30" pageNumber="231">Fig. 17A</figureCitation>
); hypostomal area without plumose, long, hairs; dorsolateral angles of pronotum with rounded tuft of a few, plumose, thick orange hairs; anterior and posterior margin of mesepisternum with plumose, thin, long, scattered, white hairs; disc of metepisternum without characteristic tuft of thick, short, white hairs; tergal fasciae with row of thick, short, white to golden hairs, present from T1 to T3; disc of T1 to T6 with simple, long, thin, white hairs in punctures increasing in length towards posterior terga (
<figureCitation id="C8AA4D86FFBAB04DFAD9C60C96217BD9" box="[1299,1401,367,393]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="30" pageNumber="231">Fig. 17B</figureCitation>
).
</paragraph>
<paragraph id="502E5103FFBAB04DFF0DC6F090C37831" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="231">
<emphasis id="62E58D11FFBAB04DFF0DC6F092117BFD" box="[199,329,403,429]" italics="true" pageId="30" pageNumber="231">
<taxonomicName id="97912A80FFBAB04DFF0DC6F0921D7BFD" box="[199,325,403,429]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="30" pageNumber="231" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
.
</emphasis>
S5 with two subtriangular lobes directed posteriorly, and one line of capitate hairs directed and increasing in length toward sides, and some simple short hairs towards sides, without subapical line (
<figureCitation id="C8AA4D86FFBAB04DFACFC6D496327B81" box="[1285,1386,439,465]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="30" pageNumber="231">Fig. 17E</figureCitation>
); S6 with acute, right-angled apex, with one posterior tuft of simple, short hairs and one subapical, dense line of capitate hairs (
<figureCitation id="C8AA4D86FFBAB04DFF2AC69C921E7849" box="[224,326,511,537]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="30" pageNumber="231">Fig. 17F</figureCitation>
); genital capsule in dorsal view
<quantity id="9769FCE6FFBAB04DFD0AC69C90417849" box="[704,793,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="30" pageNumber="231" unit="mm" value="1.1">1.1 mm</quantity>
length, and
<quantity id="9769FCE6FFBAB04DFC66C69D975D7849" box="[940,1029,510,537]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="231" unit="mm" value="0.5">0.5 mm</quantity>
width, 2.4 times longer than wide, gonobase
<quantity id="9769FCE6FFBAB04DFEC2C5409234786D" box="[264,364,547,573]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="231" unit="mm" value="0.09">0.09 mm</quantity>
length, 4.9 times wider than long, and gonocoxite
<quantity id="9769FCE6FFBAB04DFC6FC54090A3786D" box="[933,1019,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="30" pageNumber="231" unit="mm" value="0.4">0.4 mm</quantity>
length, gonostylus
<quantity id="9769FCE6FFBAB04DFB1CC540964F786D" box="[1238,1303,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="231" unit="mm" value="1.0">1 mm</quantity>
length with quadrate apex, and long hairs, penis valve
<quantity id="9769FCE6FFBAB04DFDBEC52491927831" box="[628,714,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="231" unit="mm" value="0.7">0.7 mm</quantity>
length (
<figureCitation id="C8AA4D86FFBAB04DFCECC52490D67831" box="[806,910,583,609]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="30" pageNumber="231">Fig. 17G</figureCitation>
).
</paragraph>
<paragraph id="502E5103FFBAB04DFF0DC50990F57999" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="231">
<emphasis id="62E58D11FFBAB04DFF0DC509927F78D5" bold="true" box="[199,295,618,645]" pageId="30" pageNumber="231">
<emphasis id="62E58D11FFBAB04DFF0DC509924678D5" bold="true" box="[199,286,618,645]" italics="true" pageId="30" pageNumber="231">Female</emphasis>
:
</emphasis>
Body length
<quantity id="9769FCE6FFBAB04DFE0BC509914F78D5" box="[449,535,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="30" pageNumber="231" unit="mm" value="5.9">5.9 mm</quantity>
(5.55.9, n=3); forewing length
<quantity id="9769FCE6FFBAB04DFC43C508908778D5" box="[905,991,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="30" pageNumber="231" unit="mm" value="3.7">3.7 mm</quantity>
(3.33.7, n=3). Similar to male except:
<emphasis id="62E58D11FFBAB04DFF5DC5EC938C78F9" box="[151,212,655,681]" italics="true" pageId="30" pageNumber="231">Head</emphasis>
. Length
<quantity id="9769FCE6FFBAB04DFEF6C5ED92CC78F9" box="[316,404,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="231" unit="mm" value="1.5">1.5 mm</quantity>
(1.51.6, n=3), width
<quantity id="9769FCE6FFBAB04DFD52C5EC91A878F9" box="[664,752,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="30" pageNumber="231" unit="mm" value="1.6">1.6 mm</quantity>
(1.61.7, n=3); eyes about 3.5 times longer than broad, 1.6 times wider than gena in profile; lower interocular distance
<quantity id="9769FCE6FFBAB04DFCE7C5D090DA789D" box="[813,898,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="231" unit="mm" value="0.8">0.8 mm</quantity>
(0.80.9, n=3); upper interocular distance
<quantity id="9769FCE6FFBAB04DFA96C5D096C5789D" box="[1372,1437,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="231" unit="mm" value="1.0">1 mm</quantity>
(11.1, n=3); interantennal distance 1.8 times antennocular distance; interocellar distance 3.3 times OD, ocellocular distance 2.8 times OD, OPR 3.5 times OD; labrum without wide, basal, medial tubercle; mandible on outer surface with outer and condylar ridge subparallel without tubercle; dorsal margin of mandible with carina; clypeus with shallow emargination on two fifths and crenulated apical margin (
<figureCitation id="C8AA4D86FFBAB04DFC45C42090A0790D" box="[911,1016,835,861]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="30" pageNumber="231">Fig. 17C</figureCitation>
); scape 2.6 times longer than broad; pedicel 1.2 longer than broad, about 2 times longer than F1 and F2, remaining flagellomeres 2 times longer than F1.
<emphasis id="62E58D11FFBAB04DFF5DC4E8925779F4" box="[151,271,907,932]" italics="true" pageId="30" pageNumber="231">Mesosoma</emphasis>
. Scutum 2.6 times longer than scutellum; intertegular distance
<quantity id="9769FCE6FFBAB04DFC01C4E8977879F5" box="[971,1056,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="231" unit="mm" value="1.2">1.2 mm</quantity>
(1.21.3, n=3).
<emphasis id="62E58D11FFBAB04DFB19C4E8961079F4" box="[1235,1352,907,932]" italics="true" pageId="30" pageNumber="231">Metasoma</emphasis>
. T1
<quantity id="9769FCE6FFBAB04DFAB3C4E8939B7999" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="30" pageNumber="231" unit="mm" value="1.3">1.3 mm</quantity>
broad (1.31.4, n=3); T3
<quantity id="9769FCE6FFBAB04DFE2FC4CD91637999" box="[485,571,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="231" unit="mm" value="1.5">1.5 mm</quantity>
broad (1.51.6, n=3) (
<figureCitation id="C8AA4D86FFBAB04DFCF2C4CC90F97999" box="[824,929,943,969]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="30" pageNumber="231">Fig. 17D</figureCitation>
).
</paragraph>
<paragraph id="502E5103FFBAB04DFF0DC4B096417F29" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="231">
<emphasis id="62E58D11FFBAB04DFF0DC4B0920979BD" box="[199,337,979,1005]" italics="true" pageId="30" pageNumber="231">Sculpturing.</emphasis>
Mandibles with smooth, microreticulate integument; clypeus punctures with medium-sized (0.5 times OD), contiguous and coarse, integument slightly dull and smooth between punctures; supraclypeal area with punctures slightly bigger and sparser than on base of clypeus; frons with bigger, coarser, and slightly denser punctation than on base of clypeus, with micropunctures between punctures; vertex with punctures bigger, coarser, and sparser than on clypeus with few micropunctures between punctures; gena evenly punctate, less coarse than on frons in front of ocelli; scutum with bigger and denser punctures than on vertex, smaller towards anterior margin; rest of mesosoma as on scutum except: scutellum with sparser punctures; mesepisternum with sparser punctures, smaller towards ventral half; metasomal terga bright with few micropunctures between punctures; T1 with smaller (0.4 times OD), punctures than on scutum, with rounded shape; T2 with smaller, sparser than on T1, semicircular shaped punctures; T3 with punctures bigger (0.7 times DOP), notably less coarse than on scutum, horseshoe-shaped, and slightly sparser (0.3 times DOP); dorsal surface of T6 with smaller, and more coarsely punctures than on T5; sterna with punctation smaller and slightly denser than on scutum, microreticulate on posterior margin.
</paragraph>
<paragraph id="502E5103FFBAB04DFF0DC2E091BD7F91" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="231">
<emphasis id="62E58D11FFBAB04DFF0DC2E092687FCC" box="[199,304,1411,1436]" italics="true" pageId="30" pageNumber="231">Vestiture.</emphasis>
Clypeus and frons with simple, short, whitish hairs on punctures (
<figureCitation id="C8AA4D86FFBAB04DFBFAC2E097C07FCD" box="[1072,1176,1411,1437]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="30" pageNumber="231">Fig. 17C</figureCitation>
); T1 to T4 with apical fasciae of one row of thick, goldish hairs (
<figureCitation id="C8AA4D86FFBAB04DFDBAC2C491807F91" box="[624,728,1447,1473]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." pageId="30" pageNumber="231">Fig. 17D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBAB04DFF0DC2A892D57D8D" pageId="30" pageNumber="231" type="materials_examined">
<paragraph id="502E5103FFBAB04DFF0DC2A890E17CED" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="231">
<emphasis id="62E58D11FFBAB04DFF0DC2A8922E7FB4" bold="true" box="[199,374,1482,1509]" pageId="30" pageNumber="231">Type material:</emphasis>
<materialsCitation id="E0F95B5EFFBAB04DFEB1C2A891097C59" collectingDate="1970-09-07" collectionCode="BBSL" collectorName="RM Bohart, T" country="Mexico" elevation="387" location="Alamos" pageId="30" pageNumber="231" specimenCode="BBSL 337774" specimenCount="387" stateProvince="Sonora" typeStatus="holotype">
<emphasis id="62E58D11FFBAB04DFEB1C2A8973F7FB4" bold="true" box="[379,1127,1482,1509]" pageId="30" pageNumber="231">
<typeStatus id="8F2AEFA1FFBAB04DFEB1C2A892B07FB5" box="[379,488,1483,1509]" pageId="30" pageNumber="231" type="holotype">Holotype</typeStatus>
male (
<specimenCode id="0037F978FFBAB04DFDFEC2A9918F7FB4" box="[564,727,1482,1508]" collectionCode="BBSL" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL 337774</specimenCode>
, SM0310945):
<collectingCountry id="28861193FFBAB04DFC4CC2A997597FB4" box="[902,1025,1482,1508]" name="Mexico" pageId="30" pageNumber="231">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFBAB04DFBC1C2A9973A7FB4" box="[1035,1122,1482,1508]" country="Mexico" name="Sonora" pageId="30" pageNumber="231">Sonora</collectingRegion>
,
</emphasis>
<location id="554E07D8FFBAB04DFBA7C2A8979F7FB5" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFBA7C2A8979F7FB5" box="[1133,1223,1483,1509]" country="Mexico" name="Alamos" pageId="30" pageNumber="231" stateProvince="Sonora">Alamos</location>
,
<specimenCount id="46979A8AFFBAB04DFB1AC2A896457FB5" box="[1232,1309,1483,1509]" count="387" pageId="30" pageNumber="231" type="generic">
<quantity id="9769FCE6FFBAB04DFB1AC2A896417FB5" box="[1232,1305,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="3.87" pageId="30" pageNumber="231" unit="m" value="387.0">
<elevation id="DBBCB630FFBAB04DFB1AC2A896417FB5" box="[1232,1305,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="3.87" pageId="30" pageNumber="231" unit="m" value="387.0">387 m</elevation>
</quantity>
.
</specimenCount>
<date id="242F77C3FFBAB04DFAE8C2A896C07FB5" box="[1314,1432,1483,1509]" pageId="30" pageNumber="231" value="1970-09-07">
<collectingDate id="346B8E2BFFBAB04DFAE8C2A896C07FB5" box="[1314,1432,1483,1509]" pageId="30" pageNumber="231" value="1970-09-07">7/IX/1970</collectingDate>
</date>
, GE,
<collectorName id="FD6434D5FFBAB04DFF04C28C922C7C59" box="[206,372,1519,1545]" pageId="30" pageNumber="231">RM Bohart, T</collectorName>
. Griswold (EBCh)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBAB04DFD97C28D91847C7D" collectingDate="1970-09-07" collectionCode="BBSL" collectorName="RM Bohart" country="Mexico" elevation="387" latitude="27.025555" location="Alamos" longLatPrecision="20" longitude="-108.94139" pageId="30" pageNumber="231" specimenCount="1" stateProvince="Sonora" typeStatus="allotype">
<emphasis id="62E58D11FFBAB04DFD97C28D97617C58" bold="true" box="[605,1081,1518,1544]" pageId="30" pageNumber="231">
<typeStatus id="8F2AEFA1FFBAB04DFD97C28D919A7C58" box="[605,706,1518,1544]" pageId="30" pageNumber="231" type="allotype">Allotype</typeStatus>
(LR0006):
<collectingCountry id="28861193FFBAB04DFC99C28D90967C58" box="[851,974,1518,1544]" name="Mexico" pageId="30" pageNumber="231">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFBAB04DFC17C28D976C7C58" box="[989,1076,1518,1544]" country="Mexico" name="Sonora" pageId="30" pageNumber="231">Sonora</collectingRegion>
,
</emphasis>
<location id="554E07D8FFBAB04DFB8EC28C97C67C59" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFB8EC28C97C67C59" box="[1092,1182,1519,1545]" country="Mexico" latitude="27.025555" longLatPrecision="20" longitude="-108.94139" name="Alamos" pageId="30" pageNumber="231" stateProvince="Sonora">Alamos</location>
,
<quantity id="9769FCE6FFBAB04DFB67C28C97A27C59" box="[1197,1274,1519,1545]" metricMagnitude="2" metricUnit="m" metricValue="3.87" pageId="30" pageNumber="231" unit="m" value="387.0">
<elevation id="DBBCB630FFBAB04DFB67C28C97A27C59" box="[1197,1274,1519,1545]" metricMagnitude="2" metricUnit="m" metricValue="3.87" pageId="30" pageNumber="231" unit="m" value="387.0">387 m</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FFBAB04DFADBC28C96C47C59" box="[1297,1436,1519,1545]" degrees="27" direction="north" minutes="01" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="32" value="27.025555">27°0132”N</geoCoordinate>
<geoCoordinate id="35A537C4FFBAB04DFF5DC171926F7C7D" box="[151,311,1554,1581]" degrees="108" direction="west" minutes="56" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="29" value="-108.94139">108°5629”W</geoCoordinate>
]
<date id="242F77C3FFBAB04DFE8EC17092E17C7D" box="[324,441,1555,1581]" pageId="30" pageNumber="231" value="1970-09-07">
<collectingDate id="346B8E2BFFBAB04DFE8EC17092E17C7D" box="[324,441,1555,1581]" pageId="30" pageNumber="231" value="1970-09-07">7/IX/1970</collectingDate>
</date>
, GE,
<collectorName id="FD6434D5FFBAB04DFE3CC170912E7C7D" box="[502,630,1555,1581]" pageId="30" pageNumber="231">RM Bohart</collectorName>
, (
<collectionCode id="3680C9C6FFBAB04DFD43C170918D7C7D" box="[649,725,1555,1581]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBAB04DFD2CC17092187C01" country="Mexico" elevation="387" latitude="27.025555" location="Alamos" longLatPrecision="20" longitude="-108.94139" pageId="30" pageNumber="231" specimenCount="1" stateProvince="Sonora" typeStatus="paratype">
<emphasis id="62E58D11FFBAB04DFD2CC17097177C7C" bold="true" box="[742,1103,1554,1581]" pageId="30" pageNumber="231">
<typeStatus id="8F2AEFA1FFBAB04DFD2CC170903A7C7D" box="[742,866,1555,1581]" pageId="30" pageNumber="231" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="28861193FFBAB04DFCA7C17190B07C7C" box="[877,1000,1554,1580]" name="Mexico" pageId="30" pageNumber="231">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFBAB04DFC38C17197117C7C" box="[1010,1097,1554,1580]" country="Mexico" name="Sonora" pageId="30" pageNumber="231">Sonora</collectingRegion>
,
</emphasis>
<location id="554E07D8FFBAB04DFB99C17097F57C7D" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFB99C17097F57C7D" box="[1107,1197,1555,1581]" country="Mexico" latitude="27.025555" longLatPrecision="20" longitude="-108.94139" name="Alamos" pageId="30" pageNumber="231" stateProvince="Sonora">Alamos</location>
,
<quantity id="9769FCE6FFBAB04DFB7DC17097A77C7D" box="[1207,1279,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="3.87" pageId="30" pageNumber="231" unit="m" value="387.0">
<elevation id="DBBCB630FFBAB04DFB7DC17097A77C7D" box="[1207,1279,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="3.87" pageId="30" pageNumber="231" unit="m" value="387.0">387 m</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FFBAB04DFADBC17096C47C7D" box="[1297,1436,1555,1581]" degrees="27" direction="north" minutes="01" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="32" value="27.025555">27°0132”N</geoCoordinate>
<geoCoordinate id="35A537C4FFBAB04DFF5DC15592627C01" box="[151,314,1590,1617]" degrees="108" direction="west" minutes="56" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="29" value="-108.94139">108°5629”W</geoCoordinate>
]
</materialsCitation>
,
<date id="242F77C3FFBAB04DFE85C154929D7C01" box="[335,453,1591,1617]" pageId="30" pageNumber="231" value="1991-09-03">3/IX/1991</date>
, J. L. Neff, T. Griswold, J. G. Rozen, (EBCh,
<specimenCount id="46979A8AFFBAB04DFC2EC15490A37C01" box="[996,1019,1591,1617]" count="1" pageId="30" pageNumber="231" type="female">1♀</specimenCount>
<specimenCount id="46979A8AFFBAB04DFC36C15497477C01" box="[1020,1055,1591,1617]" count="9" pageId="30" pageNumber="231" type="male">9♂</specimenCount>
),
<date id="242F77C3FFBAB04DFBE6C15497E87C01" box="[1068,1200,1591,1617]" pageId="30" pageNumber="231" value="1970-09-07">7/IX/1970</date>
, GE, RM Bohart,
<materialsCitation id="E0F95B5EFFBAB04DFA4CC15491567C25" collectingDate="1975-09-01" collectionCode="BBSL" collectorName="Villegas" country="Mexico" pageId="30" pageNumber="231" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<date id="242F77C3FFBAB04DFA4CC15493AF7C25" pageId="30" pageNumber="231" value="1975-09-01">
<collectingDate id="346B8E2BFFBAB04DFA4CC15493AF7C25" pageId="30" pageNumber="231" value="1975-09-01">1/ IX/1975</collectingDate>
</date>
, B.
<collectorName id="FD6434D5FFBAB04DFEEFC13892D97C25" box="[293,385,1627,1653]" pageId="30" pageNumber="231">Villegas</collectorName>
(
<collectionCode id="3680C9C6FFBAB04DFE5AC138928F7C25" box="[400,471,1627,1653]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBAB04DFE15C138915D7C25" box="[479,517,1627,1653]" count="2" pageId="30" pageNumber="231" type="male">2♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFFBAB04DFDD1C138972B7C25" box="[539,1139,1627,1653]" collectingDate="1970-09-07" collectionCode="BBSL" collectorName="RM Bohart, T &amp; Griswold" country="Mexico" pageId="30" pageNumber="231" specimenCount="8" specimenCount-male="8" typeStatus="paratype">
<date id="242F77C3FFBAB04DFDD1C13891C87C25" box="[539,656,1627,1653]" pageId="30" pageNumber="231" value="1970-09-07">
<collectingDate id="346B8E2BFFBAB04DFDD1C13891C87C25" box="[539,656,1627,1653]" pageId="30" pageNumber="231" value="1970-09-07">7/IX/1970</collectingDate>
</date>
, GE,
<collectorName id="FD6434D5FFBAB04DFD18C138902A7C25" box="[722,882,1627,1653]" pageId="30" pageNumber="231">RM Bohart, T</collectorName>
.
<collectorName id="FD6434D5FFBAB04DFCB4C13890BE7C25" box="[894,998,1627,1653]" pageId="30" pageNumber="231">Griswold</collectorName>
(
<collectionCode id="3680C9C6FFBAB04DFC3CC13897657C25" box="[1014,1085,1627,1653]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBAB04DFB8FC13897327C25" box="[1093,1130,1627,1653]" count="8" pageId="30" pageNumber="231" type="male">8♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="E0F95B5EFFBAB04DFB4BC13892267CC9" collectingDate="1977-09-07" collectionCode="KUNHM-ENT" collectorName="Brooks" country="Mexico" pageId="30" pageNumber="231" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="242F77C3FFBAB04DFB4BC13897AF7C25" box="[1153,1271,1627,1653]" pageId="30" pageNumber="231" value="1977-09-07">
<collectingDate id="346B8E2BFFBAB04DFB4BC13897AF7C25" box="[1153,1271,1627,1653]" pageId="30" pageNumber="231" value="1977-09-07">7/IX/1977</collectingDate>
</date>
, R. W.
<collectorName id="FD6434D5FFBAB04DFA81C13896C47C25" box="[1355,1436,1627,1653]" pageId="30" pageNumber="231">Brooks</collectorName>
(
<collectionCode id="3680C9C6FFBAB04DFF54C11C92117CC9" box="[158,329,1663,1689]" pageId="30" pageNumber="231">KUNHM-ENT</collectionCode>
<specimenCount id="46979A8AFFBAB04DFE9BC11C922D7CC9" box="[337,373,1663,1689]" count="1" pageId="30" pageNumber="231" type="male">1♂</specimenCount>
)
</materialsCitation>
,
<quantity id="9769FCE6FFBAB04DFE41C11C928C7CC9" box="[395,468,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="30" pageNumber="231" unit="m" value="440.0">440 m</quantity>
, [
<geoCoordinate id="35A537C4FFBAB04DFE2DC11C912A7CC9" box="[487,626,1663,1689]" degrees="27" direction="north" minutes="01" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="32" value="27.025555">27°0132”N</geoCoordinate>
<geoCoordinate id="35A537C4FFBAB04DFDB3C11D90447CC9" box="[633,796,1662,1689]" degrees="108" direction="west" minutes="56" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="29" value="-108.94139">108°5629”W</geoCoordinate>
], L. M. Godinez, R. Ayala (EBCh
<specimenCount id="46979A8AFFBAB04DFB56C11C979A7CC9" box="[1180,1218,1663,1689]" count="7" pageId="30" pageNumber="231" type="male">7♂</specimenCount>
).
<materialsCitation id="E0F95B5EFFBAB04DFB1DC11C928F7CED" country="Mexico" elevation="105" latitude="27.917778" location="Guaymas" longLatPrecision="20" longitude="-110.90889" pageId="30" pageNumber="231" specimenCount="1" typeStatus="paratype">
<location id="554E07D8FFBAB04DFB1DC11C961C7CC9" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFB1DC11C961C7CC9" box="[1239,1348,1663,1689]" country="Mexico" latitude="27.917778" longLatPrecision="20" longitude="-110.90889" name="Guaymas" pageId="30" pageNumber="231">Guaymas</location>
,
<quantity id="9769FCE6FFBAB04DFA84C11D96CF7CC9" box="[1358,1431,1662,1689]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="30" pageNumber="231" unit="m" value="105.0">
<elevation id="DBBCB630FFBAB04DFA84C11D96CF7CC9" box="[1358,1431,1662,1689]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="30" pageNumber="231" unit="m" value="105.0">105 m</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FFBAB04DFF54C1C192727CED" box="[158,298,1698,1725]" degrees="27" direction="north" minutes="55" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="04" value="27.917778">27°5504”N</geoCoordinate>
<geoCoordinate id="35A537C4FFBAB04DFEFBC1C192887CED" box="[305,464,1698,1725]" degrees="110" direction="west" minutes="54" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="32" value="-110.90889">110°5432”W</geoCoordinate>
]
</materialsCitation>
<materialsCitation id="E0F95B5EFFBAB04DFE17C1C090EC7CED" box="[477,948,1699,1725]" collectingDate="1966-09-26" collectionCode="BBSL" collectorName="Bohart" country="American Samoa" location="AS" pageId="30" pageNumber="231" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="242F77C3FFBAB04DFE17C1C0913A7CED" box="[477,610,1699,1725]" pageId="30" pageNumber="231" value="1966-09-26">
<collectingDate id="346B8E2BFFBAB04DFE17C1C0913A7CED" box="[477,610,1699,1725]" pageId="30" pageNumber="231" value="1966-09-26">26/IX/1966</collectingDate>
</date>
, CE &amp;
<collectingCountry id="28861193FFBAB04DFD78C1C0918E7CED" box="[690,726,1699,1725]" name="American Samoa" pageId="30" pageNumber="231">AS</collectingCountry>
<collectorName id="FD6434D5FFBAB04DFD17C1C090717CED" box="[733,809,1699,1725]" pageId="30" pageNumber="231">Bohart</collectorName>
(
<collectionCode id="3680C9C6FFBAB04DFCF2C1C090277CED" box="[824,895,1699,1725]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBAB04DFC4CC1C090FF7CED" box="[902,935,1699,1725]" count="1" pageId="30" pageNumber="231" type="female">1♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="502E5103FFBAB04DFF0DC1A592D57D8D" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="231">
<emphasis id="62E58D11FFBAB04DFF0DC1A591657CB0" bold="true" box="[199,573,1734,1760]" pageId="30" pageNumber="231">Additional material examined:</emphasis>
<materialsCitation id="E0F95B5EFFBAB04DFD82C1A4918C7D55" collectingDate="2011-07-06" collectingDateMax="2011-07-11" collectingDateMin="2011-07-06" collectionCode="BBSL" collectorName="M. E. Irwin" country="United States of America" location="Mosca" municipality="Alamosa Co. North Zapata Creek" pageId="30" pageNumber="231" specimenCount="1" specimenCount-male="1" stateProvince="Colorado">
<emphasis id="62E58D11FFBAB04DFD82C1A490E97CB0" bold="true" box="[584,945,1734,1761]" pageId="30" pageNumber="231">
<collectingCountry id="28861193FFBAB04DFD82C1A490737CB0" box="[584,811,1734,1761]" name="United States of America" pageId="30" pageNumber="231">UNITED STATES</collectingCountry>
:
<collectingRegion id="92559FE1FFBAB04DFCF0C1A590F47CB0" box="[826,940,1734,1760]" country="United States of America" name="Colorado" pageId="30" pageNumber="231">Colorado</collectingRegion>
,
</emphasis>
<collectingMunicipality id="B04ACB79FFBAB04DFC70C1A4961B7CB1" box="[954,1347,1735,1761]" pageId="30" pageNumber="231">Alamosa Co. North Zapata Creek</collectingMunicipality>
;
<location id="554E07D8FFBAB04DFA98C1A496C57CB1" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFA98C1A496C57CB1" box="[1362,1437,1735,1761]" country="United States of America" municipality="Alamosa Co. North Zapata Creek" name="Mosca" pageId="30" pageNumber="231" stateProvince="Colorado">Mosca</location>
25kmE,
<date id="242F77C3FFBAB04DFF31C18892C57D55" box="[251,413,1771,1797]" pageId="30" pageNumber="231" value="2011-07-06" valueMax="2011-07-11" valueMin="2011-07-06">
<collectingDate id="346B8E2BFFBAB04DFF31C18892C57D55" box="[251,413,1771,1797]" pageId="30" pageNumber="231" value="2011-07-06" valueMax="2011-07-11" valueMin="2011-07-06">6-11/VII/2011</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBAB04DFE67C18891637D55" box="[429,571,1771,1797]" pageId="30" pageNumber="231">M. E. Irwin</collectorName>
, (
<collectionCode id="3680C9C6FFBAB04DFD99C18891C27D55" box="[595,666,1771,1797]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBAB04DFD6FC18891927D55" box="[677,714,1771,1797]" count="1" pageId="30" pageNumber="231" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBAB04DFD2EC18997867D79" collectingDate="5 - IX- 1968, 31 / VIII / 1968" collectionCode="BBSL" collectorName="T. A. Sears &amp; R. C. Gardner &amp; C. S. Glaser" country="Mexico" elevation="1426" latitude="27.522223" location="Santo Nino" longLatPrecision="20" longitude="-104.83723" pageId="30" pageNumber="231" specimenCount="13" specimenCount-male="13" stateProvince="Chihuahua">
<emphasis id="62E58D11FFBAB04DFD2EC18990A47D54" bold="true" box="[740,1020,1770,1796]" pageId="30" pageNumber="231">
<collectingCountry id="28861193FFBAB04DFD2EC18990077D54" box="[740,863,1770,1796]" name="Mexico" pageId="30" pageNumber="231">MEXICO</collectingCountry>
:
<collectingRegion id="92559FE1FFBAB04DFCA5C18990AF7D54" box="[879,1015,1770,1796]" country="Mexico" name="Chihuahua" pageId="30" pageNumber="231">Chihuahua</collectingRegion>
,
</emphasis>
<location id="554E07D8FFBAB04DFBC2C18897D67D55" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFBC2C18897D67D55" box="[1032,1166,1771,1797]" country="Mexico" latitude="27.522223" longLatPrecision="20" longitude="-104.83723" name="Santo Nino" pageId="30" pageNumber="231" stateProvince="Chihuahua">Santo Nino</location>
, [
<quantity id="9769FCE6FFBAB04DFB6EC188965B7D55" box="[1188,1283,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.426" pageId="30" pageNumber="231" unit="m" value="1426.0">
<elevation id="DBBCB630FFBAB04DFB6EC188965B7D55" box="[1188,1283,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.426" pageId="30" pageNumber="231" unit="m" value="1426.0">1426 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFBAB04DFAD9C18896C47D55" box="[1299,1436,1771,1797]" degrees="27" direction="north" minutes="31" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="20" value="27.522223">27°3120”N</geoCoordinate>
<geoCoordinate id="35A537C4FFBAB04DFF5DC06D92617D79" box="[151,313,1806,1833]" degrees="104" direction="west" minutes="50" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="14" value="-104.83723">104°5014”W</geoCoordinate>
], 1/
<collectingDate id="346B8E2BFFBAB04DFE95C06D91257D79" box="[351,637,1806,1833]" isEnumeration="true" pageId="30" pageNumber="231">
<date id="242F77C3FFBAB04DFE95C06D92837D79" box="[351,475,1806,1833]" pageId="30" pageNumber="231" value="1968-09-05">5-IX-1968</date>
,
<date id="242F77C3FFBAB04DFE2CC06C91257D79" box="[486,637,1807,1833]" pageId="30" pageNumber="231" value="1968-08-31">31/VIII/1968</date>
</collectingDate>
,
<collectorName id="FD6434D5FFBAB04DFD42C06C905D7D79" box="[648,773,1807,1833]" pageId="30" pageNumber="231">T. A. Sears</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFCDBC06C90E97D79" box="[785,945,1807,1833]" pageId="30" pageNumber="231">R. C. Gardner</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFC76C06C971C7D79" box="[956,1092,1807,1833]" pageId="30" pageNumber="231">C. S. Glaser</collectorName>
(
<collectionCode id="3680C9C6FFBAB04DFB99C06C97C27D79" box="[1107,1178,1807,1833]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBAB04DFB6BC06C978D7D79" box="[1185,1237,1807,1833]" count="13" pageId="30" pageNumber="231" type="male">13♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBAB04DFB20C06D961A7D1D" collectingDate="1969-07-25" collectionCode="BBSL" collectorName="T. A. Sears &amp; R. C. Gardner &amp; C. S. Glaser" country="Mexico" elevation="240" latitude="26.708889" location="Choix" longLatPrecision="20" longitude="-108.32444" pageId="30" pageNumber="231" specimenCount="1" specimenCount-male="1" stateProvince="Sinaloa">
<emphasis id="62E58D11FFBAB04DFB20C06D96117D78" bold="true" box="[1258,1353,1806,1832]" pageId="30" pageNumber="231">
<collectingRegion id="92559FE1FFBAB04DFB20C06D961C7D78" box="[1258,1348,1806,1832]" country="Mexico" name="Sinaloa" pageId="30" pageNumber="231">Sinaloa</collectingRegion>
,
</emphasis>
<location id="554E07D8FFBAB04DFA9BC06C96C07D79" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFA9BC06C96C07D79" box="[1361,1432,1807,1833]" country="Mexico" latitude="26.708889" longLatPrecision="20" longitude="-108.32444" name="Choix" pageId="30" pageNumber="231" stateProvince="Sinaloa">Choix</location>
, [
<quantity id="9769FCE6FFBAB04DFF54C05093B47D1D" box="[158,236,1843,1869]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="30" pageNumber="231" unit="m" value="240.0">
<elevation id="DBBCB630FFBAB04DFF54C05093B47D1D" box="[158,236,1843,1869]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="30" pageNumber="231" unit="m" value="240.0">240 m</elevation>
</quantity>
,
<geoCoordinate id="35A537C4FFBAB04DFF31C05092DC7D1D" box="[251,388,1843,1869]" degrees="26" direction="north" minutes="42" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="32" value="26.708889">26°4232”N</geoCoordinate>
<geoCoordinate id="35A537C4FFBAB04DFE44C05091687D1D" box="[398,560,1843,1869]" degrees="108" direction="west" minutes="19" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="28" value="-108.32444">108°1928”W</geoCoordinate>
],
<date id="242F77C3FFBAB04DFD8CC051918B7D1D" box="[582,723,1842,1869]" pageId="30" pageNumber="231" value="1969-07-25">
<collectingDate id="346B8E2BFFBAB04DFD8CC051918B7D1D" box="[582,723,1842,1869]" pageId="30" pageNumber="231" value="1969-07-25">25/VII/1969</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBAB04DFD2BC050903B7D1D" box="[737,867,1843,1869]" pageId="30" pageNumber="231">T. A. Sears</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFCB8C050974F7D1D" box="[882,1047,1843,1869]" pageId="30" pageNumber="231">R. C. Gardner</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFBEEC05097E97D1D" box="[1060,1201,1843,1869]" pageId="30" pageNumber="231">C. S. Glaser</collectorName>
(
<collectionCode id="3680C9C6FFBAB04DFB09C05096527D1D" box="[1219,1290,1843,1869]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBAB04DFADEC05096607D1D" box="[1300,1336,1843,1869]" count="1" pageId="30" pageNumber="231" type="male">1♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBAB04DFA9BC05093AF7DC5" collectingDate="1968-08-31" collectionCode="BBSL" collectorName="T. A. Sears &amp; R. C. Gardner &amp; C. S. Glaser" country="Mexico" elevation="442" latitude="26.728611" location="Choix" longLatPrecision="20" longitude="-108.2975" pageId="30" pageNumber="231" specimenCount="22" specimenCount-female="4" specimenCount-male="18" stateProvince="Sinaloa">
<location id="554E07D8FFBAB04DFA9BC05096C07D1D" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFA9BC05096C07D1D" box="[1361,1432,1843,1869]" country="Mexico" latitude="26.728611" longLatPrecision="20" longitude="-108.2975" name="Choix" pageId="30" pageNumber="231" stateProvince="Sinaloa">Choix</location>
, 4miNW,
<quantity id="9769FCE6FFBAB04DFF36C034921D7D21" box="[252,325,1879,1905]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="30" pageNumber="231" unit="m" value="442.0">
<elevation id="DBBCB630FFBAB04DFF36C034921D7D21" box="[252,325,1879,1905]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="30" pageNumber="231" unit="m" value="442.0">442 m</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FFBAB04DFE9DC03492BB7D21" box="[343,483,1879,1905]" degrees="26" direction="north" minutes="43" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="43" value="26.728611">26°4343”N</geoCoordinate>
<geoCoordinate id="35A537C4FFBAB04DFE23C03591D37D21" box="[489,651,1878,1905]" degrees="108" direction="west" minutes="17" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="51" value="-108.2975">108°1751”W</geoCoordinate>
], 29, 27,
<date id="242F77C3FFBAB04DFD3BC03490DF7D21" box="[753,903,1879,1905]" pageId="30" pageNumber="231" value="1968-08-31">
<collectingDate id="346B8E2BFFBAB04DFD3BC03490DF7D21" box="[753,903,1879,1905]" pageId="30" pageNumber="231" value="1968-08-31">31/VIII/1968</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBAB04DFC5BC03497557D21" box="[913,1037,1879,1905]" pageId="30" pageNumber="231">T. A. Sears</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFBD2C03497EF7D21" box="[1048,1207,1879,1905]" pageId="30" pageNumber="231">R. C. Gardner</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFB0BC034961F7D21" box="[1217,1351,1879,1905]" pageId="30" pageNumber="231">C. S. Glaser</collectorName>
(
<collectionCode id="3680C9C6FFBAB04DFA9CC03496C57D21" box="[1366,1437,1879,1905]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBAB04DFF5DC01893E07DC5" box="[151,184,1915,1941]" count="4" pageId="30" pageNumber="231" type="female">4♀</specimenCount>
<specimenCount id="46979A8AFFBAB04DFF72C01893B67DC5" box="[184,238,1915,1941]" count="18" pageId="30" pageNumber="231" type="male">18♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBAB04DFEC8C01892157DE9" collectingDate="1968-09-05" collectionCode="BBSL" collectorName="T. A. Sears &amp; R. C. Gardner &amp; C. S. Glaser" country="Mexico" elevation="465" latitude="26.728056" location="Choix" longLatPrecision="20" longitude="-108.29611" pageId="30" pageNumber="231" specimenCount="14" specimenCount-female="2" specimenCount-male="12" stateProvince="Sinaloa">
<location id="554E07D8FFBAB04DFEC8C01892117DC5" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFEC8C01892117DC5" box="[258,329,1915,1941]" country="Mexico" latitude="26.728056" longLatPrecision="20" longitude="-108.29611" name="Choix" pageId="30" pageNumber="231" stateProvince="Sinaloa">Choix</location>
, 5.5miNW,
<quantity id="9769FCE6FFBAB04DFE04C019914E7DC5" box="[462,534,1914,1941]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="30" pageNumber="231" unit="m" value="465.0">
<elevation id="DBBCB630FFBAB04DFE04C019914E7DC5" box="[462,534,1914,1941]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="30" pageNumber="231" unit="m" value="465.0">465 m</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FFBAB04DFDE2C01891EC7DC5" box="[552,692,1915,1941]" degrees="26" direction="north" minutes="43" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="41" value="26.728056">26°4341”N</geoCoordinate>
<geoCoordinate id="35A537C4FFBAB04DFD70C01890027DC5" box="[698,858,1915,1941]" degrees="108" direction="west" minutes="17" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="46" value="-108.29611">108°1746”W</geoCoordinate>
]
<date id="242F77C3FFBAB04DFCADC01990857DC5" box="[871,989,1914,1941]" pageId="30" pageNumber="231" value="1968-09-05">
<collectingDate id="346B8E2BFFBAB04DFCADC01990857DC5" box="[871,989,1914,1941]" pageId="30" pageNumber="231" value="1968-09-05">5/IX/1968</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBAB04DFC2DC018973B7DC5" box="[999,1123,1915,1941]" pageId="30" pageNumber="231">T. A. Sears</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFBA4C01896547DC5" box="[1134,1292,1915,1941]" pageId="30" pageNumber="231">R. C. Gardner</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFADCC01896C47DC5" box="[1302,1436,1915,1941]" pageId="30" pageNumber="231">C. S. Glaser</collectorName>
(
<collectionCode id="3680C9C6FFBAB04DFF55C0FC93BE7DE9" box="[159,230,1951,1977]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBAB04DFF27C0FC92567DE9" box="[237,270,1951,1977]" count="2" pageId="30" pageNumber="231" type="female">2♀</specimenCount>
<specimenCount id="46979A8AFFBAB04DFEC7C0FC921C7DE9" box="[269,324,1951,1977]" count="12" pageId="30" pageNumber="231" type="male">12♂</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="E0F95B5EFFBAB04DFE93C0FC92D17D8D" collectingDate="1968-08-06" collectionCode="BBSL" collectorName="T. A. Sears &amp; R. C. Gardner &amp; C. S. Glaser" country="Mexico" elevation="442" latitude="26.728611" location="Choix" longLatPrecision="20" longitude="-108.2975" pageId="30" pageNumber="231" specimenCount="2" specimenCount-male="2" stateProvince="Sinaloa">
<location id="554E07D8FFBAB04DFE93C0FC92F87DE9" LSID="urn:lsid:plazi:treatment:D838E015FFBFB04CFF5DC3AD918C7C64:554E07D8FFBAB04DFE93C0FC92F87DE9" box="[345,416,1951,1977]" country="Mexico" latitude="26.728611" longLatPrecision="20" longitude="-108.2975" name="Choix" pageId="30" pageNumber="231" stateProvince="Sinaloa">Choix</location>
, 6miNW,
<quantity id="9769FCE6FFBAB04DFDD9C0FC91047DE9" box="[531,604,1951,1977]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="30" pageNumber="231" unit="m" value="442.0">
<elevation id="DBBCB630FFBAB04DFDD9C0FC91047DE9" box="[531,604,1951,1977]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="30" pageNumber="231" unit="m" value="442.0">442 m</elevation>
</quantity>
, [
<geoCoordinate id="35A537C4FFBAB04DFDBAC0FC91A37DE9" box="[624,763,1951,1977]" degrees="26" direction="north" minutes="43" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="43" value="26.728611">26°4343”N</geoCoordinate>
<geoCoordinate id="35A537C4FFBAB04DFCC9C0FD90FD7DE9" box="[771,933,1950,1977]" degrees="108" direction="west" minutes="17" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="51" value="-108.2975">108°1751”W</geoCoordinate>
],
<date id="242F77C3FFBAB04DFC72C0FC97187DE9" box="[952,1088,1951,1977]" pageId="30" pageNumber="231" value="1968-08-06">
<collectingDate id="346B8E2BFFBAB04DFC72C0FC97187DE9" box="[952,1088,1951,1977]" pageId="30" pageNumber="231" value="1968-08-06">6/VIII/1968</collectingDate>
</date>
,
<collectorName id="FD6434D5FFBAB04DFB86C0FC97927DE9" box="[1100,1226,1951,1977]" pageId="30" pageNumber="231">T. A. Sears</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFB1DC0FC962F7DE9" box="[1239,1399,1951,1977]" pageId="30" pageNumber="231">R. C. Gardner</collectorName>
,
<collectorName id="FD6434D5FFBAB04DFA49C0FC93A67D8D" pageId="30" pageNumber="231">C. S. Glaser</collectorName>
(
<collectionCode id="3680C9C6FFBAB04DFEC6C0A0920B7D8D" box="[268,339,1987,2013]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="30" pageNumber="231" type="Museum">BBSL</collectionCode>
<specimenCount id="46979A8AFFBAB04DFE90C0A092D87D8D" box="[346,384,1987,2013]" count="2" pageId="30" pageNumber="231" type="male">2♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="04EE018BFFBBB04CFF5DC329904E7E32" box="[151,790,1098,1122]" pageId="31" pageNumber="232" startId="31.[151,250,1098,1122]" targetBox="[159,1428,187,1063]" targetPageId="31" targetType="figure">
<paragraph id="502E5103FFBBB04CFF5DC329904E7E32" blockId="31.[151,790,1098,1122]" box="[151,790,1098,1122]" pageId="31" pageNumber="232">
<emphasis id="62E58D11FFBBB04CFF5DC32992787E32" bold="true" box="[151,288,1098,1122]" pageId="31" pageNumber="232">FIGURE 18.</emphasis>
Known distribution records for
<taxonomicName id="97912A80FFBBB04CFDADC32990487E32" authorityName="Crawford" authorityYear="1913" box="[615,784,1098,1122]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="leavitti">
<emphasis id="62E58D11FFBBB04CFDADC32990487E32" box="[615,784,1098,1122]" italics="true" pageId="31" pageNumber="232">Heriades leavitti</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="188B0288FFBBB04CFF0DC3D797C07F47" pageId="31" pageNumber="232" type="distribution">
<paragraph id="502E5103FFBBB04CFF0DC3D797C07F47" blockId="31.[151,1437,1204,1588]" pageId="31" pageNumber="232">
<emphasis id="62E58D11FFBBB04CFF0DC3D792397E9E" bold="true" box="[199,353,1204,1230]" pageId="31" pageNumber="232">Distribution.</emphasis>
This species is present mainly in the northern
<collectingCountry id="28861193FFBBB04CFCA1C3D7909B7E9E" box="[875,963,1204,1230]" name="Mexico" pageId="31" pageNumber="232">Mexico</collectingCountry>
, in the states of
<collectingRegion id="92559FE1FFBBB04CFBB3C3D797AA7E9E" box="[1145,1266,1204,1230]" country="Mexico" name="Chihuahua" pageId="31" pageNumber="232">Chihuahua</collectingRegion>
and
<collectingRegion id="92559FE1FFBBB04CFAE3C3D696237E9F" box="[1321,1403,1205,1231]" country="Mexico" name="Sonora" pageId="31" pageNumber="232">Sonora</collectingRegion>
, in the Sierra Madre Occidental and Chihuahuan biogeographic provinces (
<bibRefCitation id="34002CF2FFBBB04CFC67C3BB97D67EA2" author="Morrone, J. J. &amp; Escalante, T. &amp; Rodriguez-Tapia, G. &amp; Carmona, A. &amp; Arana, M. &amp; Mercado-Gomez, J. D." box="[941,1166,1240,1267]" pageId="31" pageNumber="232" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. &amp; Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
Morrone
<emphasis id="62E58D11FFBBB04CFBDCC3BA971D7EA2" box="[1046,1093,1240,1266]" italics="true" pageId="31" pageNumber="232">et al</emphasis>
. 2022
</bibRefCitation>
). There is a questionable record of
<specimenCount id="46979A8AFFBBB04CFECEC39E92317F46" box="[260,361,1276,1303]" count="1" pageId="31" pageNumber="232" type="male">one male</specimenCount>
from
<collectingRegion id="92559FE1FFBBB04CFE64C39F91417F46" box="[430,537,1276,1302]" country="United States of America" name="Colorado" pageId="31" pageNumber="232">Colorado</collectingRegion>
,
<collectingCountry id="28861193FFBBB04CFDEFC39F91E37F46" box="[549,699,1276,1302]" name="United States of America" pageId="31" pageNumber="232">United States</collectingCountry>
(
<figureCitation id="C8AA4D86FFBBB04CFD00C39F90457F46" box="[714,797,1276,1302]" captionStart="FIGURE 16" captionStartId="28.[151,250,1098,1122]" captionTargetBox="[160,1428,187,1064]" captionTargetId="figure-22@28.[151,1436,179,1072]" captionTargetPageId="28" captionText="FIGURE 16. Known distribution records for Heriades danuncia sp. nov." pageId="31" pageNumber="232">Fig. 16</figureCitation>
). Altitudinal range:
<quantity id="9769FCE6FFBBB04CFC37C39F97CC7F47" box="[1021,1172,1276,1303]" metricMagnitude="2" metricUnit="m" metricValue="7.65" metricValueMax="15.11" metricValueMin="0.19" pageId="31" pageNumber="232" unit="m" value="765.0" valueMax="1511.0" valueMin="19.0">19 to 1511 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBBB04CFF0DC243924A7FD1" pageId="31" pageNumber="232" type="etymology">
<paragraph id="502E5103FFBBB04CFF0DC243924A7FD1" blockId="31.[151,1437,1204,1588]" pageId="31" pageNumber="232">
<emphasis id="62E58D11FFBBB04CFF0DC24392167F6A" bold="true" box="[199,334,1312,1338]" pageId="31" pageNumber="232">Etymology.</emphasis>
This species is dedicated to Dra.
<taxonomicName id="97912A80FFBBB04CFD12C24390CE7F6A" authority="Urban" authorityName="Urban" box="[728,918,1312,1338]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="danuncia">Danuncia Urban</taxonomicName>
, taxonomist and melittologist from
<collectingCountry id="28861193FFBBB04CFAF9C24396227F6A" box="[1331,1402,1312,1338]" name="Brazil" pageId="31" pageNumber="232">Brazil</collectingCountry>
, at Universidade Federal do
<collectingRegion id="92559FE1FFBBB04CFE72C227915F7F0E" box="[440,519,1348,1374]" country="Brazil" name="Parana" pageId="31" pageNumber="232">Paraná</collectingRegion>
, who contributed to the knowledge of Neotropical bees, especially Eucerini and
<taxonomicName id="97912A80FFBBB04CFF5DC20492567FD1" authorityName="Ashmead" authorityYear="1899" box="[151,270,1383,1409]" class="Insecta" family="Megachilidae" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="tribe" tribe="Anthidiini">Anthidiini</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="188B0288FFBBB04CFF0DC2E8918C7C64" pageId="31" pageNumber="232" type="discussion">
<paragraph id="502E5103FFBBB04CFF0DC2E8918C7C64" blockId="31.[151,1437,1204,1588]" pageId="31" pageNumber="232">
<emphasis id="62E58D11FFBBB04CFF0DC2E892177FF5" bold="true" box="[199,335,1419,1445]" pageId="31" pageNumber="232">Comments.</emphasis>
In males, the DBP on frons can vary from 0.25 to 0.20 DOP, as well as in the DOP on T3. The males of this species are the most distinct ones among the genus by having tubercles on carinae on outer surface of mandible, and gonostylus, with subquadrate tips with tuft of simple and hooked hairs. Females of this species differ from
<taxonomicName id="97912A80FFBBB04CFF1FC29492307C40" box="[213,360,1526,1552]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="micheneri">
<emphasis id="62E58D11FFBBB04CFF1FC29492307C40" box="[213,360,1526,1552]" italics="true" pageId="31" pageNumber="232">H. micheneri</emphasis>
</taxonomicName>
as well by having usually more orangish hairs in tuft of dorsolateral angles of pronotum, while in
<taxonomicName id="97912A80FFBBB04CFF7EC178921E7C64" box="[180,326,1562,1588]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="micheneri">
<emphasis id="62E58D11FFBBB04CFF7EC178921E7C64" box="[180,326,1562,1588]" italics="true" pageId="31" pageNumber="232">H. micheneri</emphasis>
</taxonomicName>
hairs are usually whitish to yellow.
</paragraph>
</subSubSection>
</treatment>
</document>