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<document id="B589A58F1E8D234EB958619F9E39B65E" ID-DOI="10.6620/ZS.2018.57-27" ID-ISSN="1810-522X" ID-PMC="PMC6517715" ID-PubMed="31966267" ID-Zenodo-Dep="13314991" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721916104468" checkinUser="felipe" docAuthor="Cai, Yixiong &amp; Ng, Peter Kee Lin" docDate="2018" docId="03998788FF85040CCA80FAB73FFEFD62" docLanguage="en" docName="ZoolStud.57.27.1-33.pdf" docOrigin="Zoological Studies 57 (27)" docStyle="DocumentStyle:CB122E4603BA284FA287D3847868CAF6.4:ZoolStud.2016-2018.journal_article" docStyleId="CB122E4603BA284FA287D3847868CAF6" docStyleName="ZoolStud.2016-2018.journal_article" docStyleVersion="4" docTitle="Caridina heterodactyla Liang and Yan 1985" docType="treatment" docVersion="5" lastPageNumber="29" masterDocId="FFA0FFF0FF9C0410CA0AFFEB3F02FFF5" masterDocTitle="Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="26" updateTime="1723578273083" updateUser="ExternalLinkService">
<mods:mods id="5E24F118DD8ACC7BB4498C6C4FFEE8F5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9E846EFC4C0E87C071D117DD2A70CA99">Freshwater Shrimps from Karst Caves of Southern China, with Descriptions of Seven New Species and the Identity of Typhlocaridina linyunensis Li and Luo, 2001 (Crustacea: Decapoda: Caridea)</mods:title>
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<mods:namePart id="FCEA6ACA7752FC24A04E97E28A14706A">Cai, Yixiong</mods:namePart>
<mods:affiliation id="91B454CEF24D25A3CD868BCCFD438630">National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore</mods:affiliation>
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<mods:namePart id="133B8296CB3A4B19922C2411D82569E4">Ng, Peter Kee Lin</mods:namePart>
<mods:affiliation id="08FE2D05B2C03DFDB3B3DBECBEC029B7">National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore &amp; Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of &amp; National Biodiversity Centre, National Parks Board, 1 Cluny Road, Singapore 259569, Republic of Singapore</mods:affiliation>
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<mods:typeOfResource id="273601E29440314EC22A3B73F8D2702B">text</mods:typeOfResource>
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<mods:title id="3BD4EA17D40C5817330B050E44BD52F0">Zoological Studies</mods:title>
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<mods:date id="88A495D2C5F6645045A3020F780E62AD">2018</mods:date>
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<mods:number id="3C81A36945C0BD9CF67E83E25D845011">2018-06-26</mods:number>
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<mods:number id="46DDB499222D61C39FF482329A8B7666">57</mods:number>
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<mods:number id="868FA693F8DB601832DC872141FD64AA">27</mods:number>
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<mods:identifier id="4E1753518236EFC9897D6713EB8DDEBB" type="DOI">10.6620/ZS.2018.57-27</mods:identifier>
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<treatment id="03998788FF85040CCA80FAB73FFEFD62" ID-DOI="http://doi.org/10.5281/zenodo.12826774" ID-Zenodo-Dep="12826774" LSID="urn:lsid:plazi:treatment:03998788FF85040CCA80FAB73FFEFD62" httpUri="http://treatment.plazi.org/id/03998788FF85040CCA80FAB73FFEFD62" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
<subSubSection id="C32A6515FF850409CA80FAB73DDDFA83" box="[138,735,1372,1398]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8B8F369EFF850409CA80FAB73DDDFA83" blockId="25.[138,735,1372,1430]" box="[138,735,1372,1398]" pageId="25" pageNumber="26">
<heading id="D0C781F2FF850409CA80FAB73DDDFA83" bold="true" box="[138,735,1372,1398]" centered="true" fontSize="10" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="4C304D1DFF850409CA80FAB73DDDFA83" ID-CoL="8V8G3" authority="Liang and Yan, 1985" authorityName="Liang and Yan" authorityYear="1985" box="[138,735,1372,1398]" class="Malacostraca" family="Atyidae" genus="Caridina" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="heterodactyla">
<emphasis id="B944EA8CFF850409CA80FAB73DDDFA83" bold="true" box="[138,735,1372,1398]" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409CA80FAB73FF7FA83" bold="true" box="[138,245,1372,1398]" italics="true" pageId="25" pageNumber="26">Caridina</emphasis>
aff.
<emphasis id="B944EA8CFF850409CB26FAB63EDBFA83" bold="true" box="[300,473,1373,1398]" italics="true" pageId="25" pageNumber="26">heterodactyla</emphasis>
<bibRefCitation id="EFA14B6FFF850409CBEAFAB63DDDFA83" author="Liang X &amp; Yan S." box="[480,735,1373,1398]" pageId="25" pageNumber="26" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">Liang and Yan, 1985</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A6515FF85040CCB6CFA963FFEFD62" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26" type="description">
<paragraph id="8B8F369EFF850409CB6CFA963D01FA63" blockId="25.[138,735,1372,1430]" box="[358,515,1405,1430]" pageId="25" pageNumber="26">
(
<figureCitation id="130B2A1BFF850409CB67FA963ED3FA63" box="[365,465,1405,1430]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Figs. 19</figureCitation>
,
<figureCitation id="130B2A1BFF850409CBD6FA963EFEFA63" box="[476,508,1405,1430]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">20</figureCitation>
)
</paragraph>
<paragraph id="8B8F369EFF850409CACEFA533FCEF9A4" blockId="25.[136,738,1464,1969]" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409CACEFA533EBDFA24" box="[196,447,1464,1489]" italics="true" pageId="25" pageNumber="26">Material examined</emphasis>
: Female, cl
<quantity id="4CC89B7BFF850409C879FA533DD9FA24" box="[627,731,1464,1489]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="25" pageNumber="26" unit="mm" value="7.5">7.5 mm</quantity>
, damaged with incomplete abdomen, ZRC Tonglin Cave, near Nan Cave, Kaiyuan County,
<collectingRegion id="49F4F87CFF850409C889FA133FFBF9C4" country="China" name="Yunnan" pageId="25" pageNumber="26">Yunnan Province</collectingRegion>
, southern
<collectingCountry id="F327760EFF850409CB75F9F33ECEF9C7" box="[383,460,1560,1586]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
, coll. B. Sket,
<date id="FF8E105EFF850409C88DF9F33FCAF9A4" pageId="25" pageNumber="26" value="1995-11-26">26 Nov 1995</date>
.
</paragraph>
<paragraph id="8B8F369EFF850409CACEF9B33DC3F8E4" blockId="25.[136,738,1464,1969]" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409CACEF9B33D63F984" box="[196,609,1624,1649]" italics="true" pageId="25" pageNumber="26">Comparative material examined</emphasis>
:
<taxonomicName id="4C304D1DFF850409C87CF9B33D51F964" authority="Liang and Yan, 1985" authorityName="Liang and Yan" authorityYear="1985" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="heteradactyla">
<emphasis id="B944EA8CFF850409C87CF9B33E34F964" italics="true" pageId="25" pageNumber="26">Caridina heteradactyla</emphasis>
<bibRefCitation id="EFA14B6FFF850409CB48F9933D51F964" author="Liang X &amp; Yan S." box="[322,595,1656,1681]" pageId="25" pageNumber="26" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">Liang and Yan, 1985</bibRefCitation>
</taxonomicName>
-
<typeStatus id="548B883CFF850409C86DF9933DDEF964" box="[615,732,1656,1681]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
: male, cl
<quantity id="4CC89B7BFF850409CAE1F9733E45F944" box="[235,327,1688,1713]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="25" pageNumber="26" unit="mm" value="5.6">5.6 mm</quantity>
, SFU, mountain stream at Sizhong County,
<collectingRegion id="49F4F87CFF850409CAE6F9533EBFF924" box="[236,445,1720,1745]" country="China" name="Yunnan" pageId="25" pageNumber="26">Yunnan Province</collectingRegion>
,
<collectingCountry id="F327760EFF850409CBC0F9533D16F927" box="[458,532,1720,1746]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
, 1983.
<typeStatus id="548B883CFF850409C864F9533DDFF924" box="[622,733,1720,1745]" pageId="25" pageNumber="26" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D36FD17FF850409CA82F9333FF9F904" box="[136,251,1752,1777]" count="1" pageId="25" pageNumber="26" type="female">1 female</specimenCount>
, cl
<quantity id="4CC89B7BFF850409CB20F9333E8DF904" box="[298,399,1752,1777]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="25" pageNumber="26" unit="mm" value="5.6">5.6 mm</quantity>
, SFU, mountain stream at Sizhong County,
<collectingRegion id="49F4F87CFF850409CB5BF9133D1DF8E4" box="[337,543,1784,1809]" country="China" name="Yunnan" pageId="25" pageNumber="26">Yunnan Province</collectingRegion>
,
<collectingCountry id="F327760EFF850409C823F9133D71F8E7" box="[553,627,1784,1810]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
, 1983.
</paragraph>
<paragraph id="8B8F369EFF850409CACEF8F33BA2FEA3" blockId="25.[136,738,1464,1969]" lastBlockId="25.[813,1414,253,1974]" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409CACEF8F33E4BF8C4" box="[196,329,1816,1841]" italics="true" pageId="25" pageNumber="26">Description</emphasis>
: Rostrum (
<figureCitation id="130B2A1BFF850409CBC6F8F33D31F8C4" box="[460,563,1816,1841]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19A</figureCitation>
) short, sharply downturned, reaching slightly beyond end of basal segment of antennular peduncle, dorsally with 24 teeth, of which, 12 are situated at carapace behind posterior margin of orbital margin, occupying 0.3 carapace length, armed ventrally with 7 small teeth. Antennal spine fused with suborbital angle. Pterygostomian angle rounded.
</paragraph>
<paragraph id="8B8F369EFF850409C963FEB63C8EFDE3" blockId="25.[813,1414,253,1974]" pageId="25" pageNumber="26">
Eyes well developed (
<figureCitation id="130B2A1BFF850409CE77FEB63BECFE83" box="[1149,1262,349,374]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19A</figureCitation>
). Stylocerite reaching slightly beyond end of basal segment of antennular peduncle. Antennular peduncle (
<figureCitation id="130B2A1BFF850409CF5FFE763C5CFE23" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20A</figureCitation>
) 0.43 times as long as carapace. Scaphocerite (
<figureCitation id="130B2A1BFF850409C93EFE363C9EFE03" box="[820,924,477,502]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20B</figureCitation>
) 2.7 times as long as wide, outer margin straight.
</paragraph>
<paragraph id="8B8F369EFF850409C963FDF63B03FBC3" blockId="25.[813,1414,253,1974]" pageId="25" pageNumber="26">
Mouthparts as figured. Mandible (
<figureCitation id="130B2A1BFF850409CF1BFDF63A7CFDC3" box="[1297,1406,541,566]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19B</figureCitation>
) with blunt teeth at extremity of incisor process. Maxillula (
<figureCitation id="130B2A1BFF850409C9B0FDB63B31FD82" box="[954,1075,605,631]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19C</figureCitation>
) with simple palp, lower lacinia broadly rounded, upper lacinia broadly elongated, inner edge straight, with dense setae and teeth. Maxilla (
<figureCitation id="130B2A1BFF850409CE28FD563B90FD23" box="[1058,1170,701,726]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19D</figureCitation>
) with slender palp, upper endite subdivided, scaphognathite tapering with numerous long hooked setae posteriorly. Palp of first maxilliped (
<figureCitation id="130B2A1BFF850409CE18FCF63B7CFCC3" box="[1042,1150,797,822]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19E</figureCitation>
) stout, terminating in triangular projection. Endopod of second maxilliped (
<figureCitation id="130B2A1BFF850409C93FFCB63CA9FC83" box="[821,939,861,886]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19F</figureCitation>
) with fused dactylus and propodal segments. Third maxilliped (
<figureCitation id="130B2A1BFF850409CE88FC963BEFFC62" box="[1154,1261,893,919]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20C</figureCitation>
) reaching to distal end of antennular peduncle, ending in single terminal claw; exopod reaching posterior quarter of penultimate segment, ultimate segment slightly shorter than pentultimate segment. Epipods on first 4 pereiopods.
</paragraph>
<paragraph id="8B8F369EFF850409C963FBD63C99F983" blockId="25.[813,1414,253,1974]" pageId="25" pageNumber="26">
First pereiopod (
<figureCitation id="130B2A1BFF850409CE5AFBD63BC8FBA3" box="[1104,1226,1085,1110]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20D</figureCitation>
) short, stout, reaching distal margin of eye, merus 2.1 times as long as broad; carpus strong concave anteriorly, 1.3 times as long as high, chela 2.6 times as long as broad, fingers slightly longer than palm. Second pereiopod (
<figureCitation id="130B2A1BFF850409CE2BFB363B8FFB03" box="[1057,1165,1245,1270]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20E</figureCitation>
) reaching almost to end of second segment of antennular peduncle; carpus 3.9 times as long as high, chela shorter than carpus, 2.3 times as long as broad, fingers as long as palm. Third pereiopod (
<figureCitation id="130B2A1BFF850409CF06FAB63C46FA62" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20F, G</figureCitation>
) reaching beyond distal end of scaphocrite, propodus 10 times as long as broad, 4.8 times as long as dactylus (claw included); dactylus ending as 2 claws, with 5 spines on flexor margin. Fifth pereiopod (
<figureCitation id="130B2A1BFF850409C9F3FA163B64F9E3" box="[1017,1126,1533,1558]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="25" pageNumber="26">Fig. 20H</figureCitation>
) reaching distal end of antennular peduncle, propodus 10 times as long as wide, dactylus missing. Eggs1.2 ×
<quantity id="4CC89B7BFF850409CF1BF9D63A67F9A3" box="[1297,1381,1597,1622]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="26" unit="mm" value="0.8">0.8mm</quantity>
in diameter.
</paragraph>
<paragraph id="8B8F369EFF850409C963F9963BCAF963" blockId="25.[813,1414,253,1974]" box="[873,1224,1661,1687]" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409C963F9963CBFF963" box="[873,957,1661,1686]" italics="true" pageId="25" pageNumber="26">Habitat</emphasis>
: Stream in karst cave.
</paragraph>
<paragraph id="8B8F369EFF85040CC963F9763E1DFDA3" blockId="25.[813,1414,253,1974]" lastBlockId="28.[174,776,253,663]" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
<emphasis id="B944EA8CFF850409C963F9763CD6F943" box="[873,980,1693,1718]" italics="true" pageId="25" pageNumber="26">Remarks</emphasis>
: The single female specimen found from the Tonglin Cave in Kaiyan is morphologically closest to
<taxonomicName id="4C304D1DFF850409C9A4F9363A87F903" authority="Liang and Yan, 1985" authorityName="Liang and Yan" authorityYear="1985" box="[942,1413,1757,1782]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="heterodactyla">
<emphasis id="B944EA8CFF850409C9A4F9363B7BF903" box="[942,1145,1757,1782]" italics="true" pageId="25" pageNumber="26">C. heterodactyla</emphasis>
<bibRefCitation id="EFA14B6FFF850409CE89F9363A87F903" author="Liang X &amp; Yan S." box="[1155,1413,1757,1782]" pageId="25" pageNumber="26" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">
Liang and Yan,
<quantity id="4CC89B7BFF850409CF43F9363A87F903" box="[1353,1413,1757,1782]" metricMagnitude="1" metricUnit="m" metricValue="5.0419" pageId="25" pageNumber="26" unit="in" value="1985.0">1985</quantity>
</bibRefCitation>
</taxonomicName>
in the downturned rostrum, the large number of postorbital teeth on carapace, the eggs and the long stylocerite, which reaches the end of basal segment of the antennular peduncle. It can be distinguished from
<taxonomicName id="4C304D1DFF850409CE1DF8963BE2F863" authorityName="Liang and Yan" authorityYear="1985" box="[1047,1248,1917,1942]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="heterodactyla">
<emphasis id="B944EA8CFF850409CE1DF8963BE2F863" box="[1047,1248,1917,1942]" italics="true" pageId="25" pageNumber="26">C. heterodactyla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F7FF850409CEE1F8963A2DF863" box="[1259,1327,1917,1942]" pageId="25" pageNumber="26" sensu="stricto">s. str.</taxonomicNameLabel>
by the much shorter length of rostrum (
<figureCitation id="130B2A1BFF850409CEB6F8763A28F843" box="[1212,1322,1949,1974]" captionStart="Fig" captionStartId="26.[174,208,1897,1917]" captionTargetBox="[290,1349,255,1859]" captionTargetId="figure-52@26.[272,1353,249,1871]" captionTargetPageId="26" captionText="Fig. 19. Caridina aff. heterodactyla Liang and Yan, 1985. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="25" pageNumber="26">Fig. 19A</figureCitation>
), which reaches only slightly beyond the end of the basal segment of antennular peduncle; and the shorter carpus of first pereiopod (1.3 times as long as high,
<figureCitation id="130B2A1BFF80040CCAA4FEB63E1EFE83" box="[174,284,349,374]" captionStart="Fig" captionStartId="27.[136,171,1897,1917]" captionTargetBox="[217,1364,251,1867]" captionTargetId="figure-6@27.[171,1379,220,1926]" captionTargetPageId="27" captionText="Fig. 20. Caridina aff. heterodactyla Liang and Yan, 1985. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="28" pageNumber="29">Fig. 20D</figureCitation>
) (vs. 2.0-2.3 times as long as high; cf.
<bibRefCitation id="EFA14B6FFF80040CCAA4FE963EA4FE63" author="Liang X &amp; Yan S." box="[174,422,381,406]" pageId="28" pageNumber="29" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">Liang and Yan, 1985</bibRefCitation>
: fig. 2(4)). We believe that the morphological differences indicate that the present specimen is probably an undescribed species. But with only a single incomplete female specimen available, we prefer to defer the naming of new species to a later stage when more specimens are available.
</paragraph>
<footnote id="E82B2A90FF850409CA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="25" pageNumber="26">
<paragraph id="8B8F369EFF850409CA82F8363E9AF81A" blockId="25.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="25" pageNumber="26">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF86040ACAA4F8823E39F840" ID-DOI="http://doi.org/10.5281/zenodo.13315031" ID-Zenodo-Dep="13315031" httpUri="https://zenodo.org/record/13315031/files/figure.png" pageId="26" pageNumber="27" startId="26.[174,208,1897,1917]" targetBox="[290,1349,255,1859]" targetPageId="26" targetType="figure">
<paragraph id="8B8F369EFF86040ACAA4F8823E39F840" blockId="26.[174,1451,1897,1973]" pageId="26" pageNumber="27">
<emphasis id="B944EA8CFF86040ACAA4F8823FFBF888" bold="true" box="[174,249,1897,1917]" pageId="26" pageNumber="27">Fig. 19.</emphasis>
<taxonomicName id="4C304D1DFF86040ACB0DF8823DD1F888" authority="Liang and Yan, 1985" authorityName="Liang and Yan" authorityYear="1985" box="[263,723,1897,1917]" class="Malacostraca" family="Atyidae" genus="Caridina" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="heterodactyla">
<emphasis id="B944EA8CFF86040ACB0DF8823E54F888" box="[263,342,1897,1917]" italics="true" pageId="26" pageNumber="27">Caridina</emphasis>
aff.
<emphasis id="B944EA8CFF86040ACB8BF8823D03F888" box="[385,513,1897,1917]" italics="true" pageId="26" pageNumber="27">heterodactyla</emphasis>
<bibRefCitation id="EFA14B6FFF86040AC803F8823DD1F888" author="Liang X &amp; Yan S." box="[521,723,1897,1917]" pageId="26" pageNumber="27" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">Liang and Yan, 1985</bibRefCitation>
</taxonomicName>
. (A) cephalothorax and cephalic appendages, lateral view, (B) mandible, (C) maxillula, (D) maxilla, (E) first maxilliped, (F) second maxilliped. Scale bars: A, D-F = 1.0 mm; B, C = 0.5 mm. (ovigerous female, cl 7.5 mm, ZRC).
</paragraph>
</caption>
<footnote id="E82B2A90FF86040ACE96F8363AA9F81A" box="[1180,1451,2013,2031]" pageId="26" pageNumber="27">
<paragraph id="8B8F369EFF86040ACE96F8363AA9F81A" blockId="26.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="26" pageNumber="27">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="DF4F6616FF87040BCA82F8823D5EF840" ID-DOI="http://doi.org/10.5281/zenodo.13315033" ID-Zenodo-Dep="13315033" httpUri="https://zenodo.org/record/13315033/files/figure.png" pageId="27" pageNumber="28" startId="27.[136,171,1897,1917]" targetBox="[217,1364,251,1867]" targetPageId="27" targetType="figure">
<paragraph id="8B8F369EFF87040BCA82F8823D5EF840" blockId="27.[136,1415,1897,1973]" pageId="27" pageNumber="28">
<emphasis id="B944EA8CFF87040BCA82F8823FD7F888" bold="true" box="[136,213,1897,1917]" pageId="27" pageNumber="28">Fig. 20.</emphasis>
<taxonomicName id="4C304D1DFF87040BCAEFF8823DC2F888" authority="Liang and Yan, 1985" authorityName="Liang and Yan" authorityYear="1985" box="[229,704,1897,1917]" class="Malacostraca" family="Atyidae" genus="Caridina" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="heterodactyla">
<emphasis id="B944EA8CFF87040BCAEFF8823E34F888" box="[229,310,1897,1917]" italics="true" pageId="27" pageNumber="28">Caridina</emphasis>
aff.
<emphasis id="B944EA8CFF87040BCB6FF8823EEAF888" box="[357,488,1897,1917]" italics="true" pageId="27" pageNumber="28">heterodactyla</emphasis>
<bibRefCitation id="EFA14B6FFF87040BCBFAF8823DC2F888" author="Liang X &amp; Yan S." box="[496,704,1897,1917]" pageId="27" pageNumber="28" pagination="164 - 174" refId="ref15980" refString="Liang X, Yan S. 1985. Study on Caridina (Decapoda, Caridea) from Yunnan, China. Oceanol Limnol Sinica 16 (2): 164 - 174. (in Chinese with English abstract)" type="journal article" year="1985">Liang and Yan, 1985</bibRefCitation>
</taxonomicName>
. (A) antennular peduncle, (B) scaphocerite, (C) third maxilliped, (D) first pereiopod, (E) second pereiopod, (F) third pereiopod, (G) dactylus of third pereiopod, (H) fifth pereiopod. Scale bars: A-F, H = 1.0 mm; G = 0.3 mm. (ovigerous female, cl 7.5 mm, ZRC).
</paragraph>
</caption>
<footnote id="E82B2A90FF87040BCA82F8363E9AF81A" box="[136,408,2013,2031]" pageId="27" pageNumber="28">
<paragraph id="8B8F369EFF87040BCA82F8363E9AF81A" blockId="27.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="27" pageNumber="28">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="8B8F369EFF80040CCAE0FDB63FFEFD62" blockId="28.[174,776,253,663]" pageId="28" pageNumber="29">
<emphasis id="B944EA8CFF80040CCAE0FDB63E6DFD83" box="[234,367,605,630]" italics="true" pageId="28" pageNumber="29">Distribution</emphasis>
:
<collectingRegion id="49F4F87CFF80040CCB75FDB63D48FD83" box="[383,586,605,630]" country="China" name="Yunnan" pageId="28" pageNumber="29">Yunnan Province</collectingRegion>
(Tonglin Cave),
<collectingCountry id="F327760EFF80040CCAA4FD963FFAFD62" box="[174,248,637,663]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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